species Pleurobema oviforme (Conrad, 1834): synonymy

Unio oviformis Conrad, 1834

Conrad, 1834. New Fresh-Water Shells: 46, pl. 3, fig. 6.
Conrad, 1853. Proc. Acad. Nat. Sci. 6: 254.

Pleurobema (s.s.) oviformis — Simpson (1900), Frierson (1927)
Pleurobema (s.s.) oviforme — Simpson (1914)
Pleurobema oviforme — Ortmann (1918), Ortmann (1924), Ortmann (1925), Haas (1969), Burch (1975), ... Wu et al. (2024), Dai et al. (2025), Chen et al. (2025), FMCS (2025), Schilling et al. (2025)
Pleurobema oviformis — R.I. Johnson (1980), Lopes-Lima et al. (2019)

Unio ravenelianus Lea, 1834

Lea, 1834. Trans. Amer. Phil. Soc. 5 [O. 1]: 32 [144], pl. 3, fig. 5.
(renamed Unio rudis Conrad, 1837)
Type(s): HOLOTYPE USNM_84783, French Broad R., near Ashville, N. Carolina.

Pleurobema (s.s.) raveneliana — Simpson (1900)
Pleurobema (s.s.) ravenelianum — Simpson (1914)
Pleurobema oviforme — Ortmann (1918), Parmalee & Bogan (1998), Williams et al. (2008), Schilling et al. (2025)
Quadrula (Pleuronaia) raveneliana — Frierson (1927), Haas (1969)

Unio rudis Conrad, 1837

Conrad, 1837. Monogr. Fam. Unionidae 1(8): 76, pl. 43, fig. 1.
Conrad, 1853. Proc. Acad. Nat. Sci. 6: 257.
(new name for Unio ravenelianus Lea, 1834)

Pleurobema (s.s.) rubella — Simpson (1900)
Pleurobema (s.s.) rubellum — Simpson (1914), Frierson (1927)
Pleurobema rubellum — Haas (1969), Parmalee & Bogan (1998)
Pleurobema oviforme — Parmalee & Bogan (1998), Williams et al. (2008), Schilling et al. (2025)

Unio holstonensis Lea, 1840

Lea, 1840. Proc. Amer. Phil. Soc. 1: 288.
Lea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 212 [50], pl. 15, fig. 27.
Type(s): HOLOTYPE USNM_84641, Holston R., Tenn.

Pleurobema (s.s.) holstonensis — Simpson (1900)
Pleurobema (s.s.) holstonense — Simpson (1914)
Pleurobema oviforme holstonense — Ortmann (1918), Ortmann (1924), Ortmann (1925)
Pleurobema (s.s.) oviformis — Frierson (1927)
Pleurobema oviforme — Haas (1969), Burch (1975), Parmalee & Bogan (1998), Williams et al. (2008), Schilling et al. (2025)
Pleurobema oviformis — R.I. Johnson (1980)

Unio argenteus Lea, 1841

Lea, 1841. Proc. Amer. Phil. Soc. 2: 82.
Lea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 242 [80], pl. 25, fig. 57.
Type(s): HOLOTYPE USNM_84778, Holston R., Tenn.

Pleurobema (s.s.) argentea — Simpson (1900)
Pleurobema (s.s.) argenteum — Simpson (1914)
Pleurobema oviforme argenteum — Ortmann (1918), Ortmann (1924), Ortmann (1925)
Quadrula (Pleuronaia) argentea — Frierson (1927), Haas (1969)
Pleurobema oviforme — Burch (1975), Parmalee & Bogan (1998), Schilling et al. (2025)
Pleurobema oviformis — R.I. Johnson (1980)

Unio mundus Lea, 1857

Lea, 1857. Proc. Acad. Nat. Sci. 9: 83.
Lea, 1866. Jour. Acad. Nat. Sci. 6 [O. 11]: 40 [44], pl. 14, fig. 38.
Type(s): HOLOTYPE USNM_84767, Tuscumbia, Ala.

Pleurobema (s.s.) holstonensis — Simpson (1900)
Pleurobema (s.s.) holstonense — Simpson (1914)
Pleurobema oviforme holstonense — Ortmann (1918)
Quadrula (Pleuronaia) munda — Frierson (1927), Haas (1969)
Pleurobema oviforme — Burch (1975), Parmalee & Bogan (1998), Williams et al. (2008), Schilling et al. (2025)

Unio lesleyi Lea, 1860

Lea, 1860. Proc. Acad. Nat. Sci. 12: 306.
Lea, 1860. Jour. Acad. Nat. Sci. 4 [O. 8]: 352, pl. 58, fig. 177.
Type(s): HOLOTYPE USNM_84759, Kentucky.

Pleurobema (s.s.) lesleyi — Simpson (1900), Simpson (1914)
Pleurobema oviforme — Ortmann (1918), Ortmann (1924), Parmalee & Bogan (1998), Williams et al. (2008), Schilling et al. (2025)
Quadrula (Pleuronaia) argentea — Frierson (1927), Haas (1969)

Unio tesserulae Lea, 1861

Lea, 1861. Proc. Acad. Nat. Sci. 13: 392 [as ‘tesserulæ’].
Lea, 1866. Jour. Acad. Nat. Sci. 6 [O. 11]: 40 [44], pl. 15, fig. 39 [as ‘tesserulæ’].
Type(s): HOLOTYPE USNM_84426, Nolachucky R., Tenn.

Pleurobema (s.s.) tesserulae — Simpson (1900), Simpson (1914)
Pleurobema oviforme holstonense — Ortmann (1918)
Quadrula (Pleuronaia) tesserulae — Frierson (1927), Haas (1969)
Pleurobema oviforme — Parmalee & Bogan (1998), Williams et al. (2008), Schilling et al. (2025)

Unio striatissimus J.G. Anthony, 1865

J.G. Anthony, 1865. Amer. J. Conch. 1(2): 156, pl. 12, fig. 1.

Pleurobema (s.s.) estabrookiana — Simpson (1900)
Pleurobema (s.s.) estabrookianum — Simpson (1914)
Pleurobema oviforme argenteum — Ortmann (1918)
Quadrula (Pleuronaia) striatissima — Frierson (1927), Haas (1969)
Fusconaia barnesiana — Parmalee & Bogan (1998)
Pleurobema oviforme — Parmalee & Bogan (1998), Williams et al. (2008), Schilling et al. (2025)
Pleuronaia barnesiana — Williams et al. (2008)

Unio clinchensis Lea, 1867

Lea, 1867. Proc. Acad. Nat. Sci. 19: 81.
Lea, 1868. Jour. Acad. Nat. Sci. 6 [O. 12]: 278 [38], pl. 37, fig. 91.
Type(s): HOLOTYPE USNM_84596, Clinch R., Tenn.

Pleurobema (s.s.) cuneolus — Simpson (1900)
Pleurobema (s.s.) clinchense — Simpson (1914)
Pleurobema oviforme — Ortmann (1918), Ortmann (1924), Parmalee & Bogan (1998), Williams et al. (2008), Schilling et al. (2025)
Quadrula (Pleuronaia) cuneola — Frierson (1927)
Quadrula (Pleuronaia) cuneolus — Haas (1969)
Fusconaia cuneolus — Burch (1975)

Unio planior Lea, 1868

Lea, 1868. Proc. Acad. Nat. Sci. 20: 145.
Lea, 1868. Jour. Acad. Nat. Sci. 6 [O. 12]: 316 [76], pl. 50, fig. 129.
Type(s): FIGURED SYNTYPE USNM_84391, Tennessee.

Pleurobema (s.s.) planior — Simpson (1900)
Pleurobema (s.s.) planius — Simpson (1914)
Pleurobema oviforme argenteum — Ortmann (1918)
Quadrula (Pleuronaia) striatissima — Frierson (1927), Haas (1969)
Pleurobema oviforme — Parmalee & Bogan (1998), Williams et al. (2008), Schilling et al. (2025)

Unio acuens Lea, 1871

Lea, 1871. Proc. Acad. Nat. Sci. 23: 190.
Lea, 1874. Jour. Acad. Nat. Sci. 8 [O. 13]: 27 [31], pl. 8, fig. 24.
Type(s): FIGURED SYNTYPE USNM_84599, Holston R., near Concord, E. Tenn.

Pleurobema (s.s.) acuens — Simpson (1900), Simpson (1914)
Pleurobema oviforme holstonense — Ortmann (1918)
Quadrula (Pleuronaia) tesserulae — Frierson (1927), Haas (1969)
Pleurobema oviforme — Parmalee & Bogan (1998), Williams et al. (2008), Schilling et al. (2025)

Unio lawii Lea, 1871

Lea, 1871. Proc. Acad. Nat. Sci. 23: 189.
Lea, 1874. Jour. Acad. Nat. Sci. 8 [O. 13]: 8 [12], pl. 2, fig. 4.
Type(s): FIGURED SYNTYPE USNM_84757, Holston R., Tenn.

Pleurobema (s.s.) holstonensis — Simpson (1900)
Pleurobema (s.s.) holstonense — Simpson (1914)
Pleurobema oviforme holstonense — Ortmann (1918)
Quadrula (Pleuronaia) munda — Frierson (1927), Haas (1969)
Pleurobema oviforme — Burch (1975), Parmalee & Bogan (1998), Williams et al. (2008), Schilling et al. (2025)

Unio pattinoides Lea, 1871

Lea, 1871. Proc. Acad. Nat. Sci. 23: 193.
Lea, 1874. Jour. Acad. Nat. Sci. 8 [O. 13]: 16 [20], pl. 4, fig. 12.
Type(s): FIGURED SYNTYPE USNM_84763, Holston R., Tenn.

Pleurobema (s.s.) holstonensis — Simpson (1900)
Pleurobema (s.s.) holstonense — Simpson (1914)
Pleurobema oviforme holstonense — Ortmann (1918)
Quadrula (Pleuronaia) munda — Frierson (1927), Haas (1969)
Pleurobema oviforme — Parmalee & Bogan (1998), Williams et al. (2008), Schilling et al. (2025)

Unio bellulus Lea, 1872

Lea, 1872. Proc. Acad. Nat. Sci. 24: 161.
Lea, 1874. Jour. Acad. Nat. Sci. 8 [O. 13]: 50 [54], pl. 17, fig. 48.
Type(s): FIGURED SYNTYPE USNM_84772, Mussel Shoals, Tenn R., Alabama.

Pleurobema (s.s.) holstonensis — Simpson (1900)
Pleurobema (s.s.) holstonense — Simpson (1914)
Pleurobema oviforme holstonense — Ortmann (1918)
Quadrula (Pleuronaia) munda — Frierson (1927), Haas (1969)
Pleurobema oviforme — Burch (1975), Parmalee & Bogan (1998), Williams et al. (2008), Schilling et al. (2025)

Unio brevis Lea, 1872

Lea, 1872. Proc. Acad. Nat. Sci. 24: 157.
Lea, 1874. Jour. Acad. Nat. Sci. 8 [O. 13]: 35 [39], pl. 12, fig. 32.
Type(s): FIGURED SYNTYPE USNM_84546, Conasauga R., E. Tenn.

Pleurobema (s.s.) brevis — Simpson (1900)
Pleurobema (s.s.) breve — Simpson (1914)
Pleurobema oviforme argenteum — Ortmann (1918)
Quadrula (Pleuronaia) striatissima — Frierson (1927), Haas (1969)
Pleurobema oviforme — Parmalee & Bogan (1998), Williams et al. (2008), Schilling et al. (2025)

Unio conasaugaensis Lea, 1872

Lea, 1872. Proc. Acad. Nat. Sci. 24: 155.
Lea, 1874. Jour. Acad. Nat. Sci. 8 [O. 13]: 33 [37], pl. 10, fig. 30.
Type(s): HOLOTYPE (MONOTYPY) USNM_85565, Conasauga Cr., Monroe Co., E. Tenn.

Pleurobema (s.s.) conasaugaensis — Simpson (1900)
Pleurobema (s.s.) conasaugaense — Simpson (1914)
Pleurobema oviforme — Ortmann (1918), Parmalee & Bogan (1998), Williams et al. (2008), Schilling et al. (2025)
Quadrula (Pleuronaia) striatissima — Frierson (1927), Haas (1969)

Unio swordianus S.H. Wright, 1897

S.H. Wright, 1897. Nautilus 11: 4.
Type(s): LECTOTYPE USNM_149654, Br. of Clinch River, Va.

Pleurobema (s.s.) swordiana — Simpson (1900)
Pleurobema swordiana — Simpson (1900)
Pleurobema (s.s.) swordianum — Simpson (1914)
Pleurobema oviforme argenteum — Ortmann (1918)
Quadrula (Pleuronaia) striatissima — Frierson (1927), Haas (1969)
Pleurobema oviforme — Parmalee & Bogan (1998), Williams et al. (2008), Schilling et al. (2025)

Unio patulus Conrad, 1838

‘Lea’ Conrad, 1838. Monogr. Fam. Unionidae 1(10): 92, pl. 50, fig. 2.
(misidentified reference to Unio patulus Lea, 1829)

Pleurobema (s.s.) clava — Simpson (1900)
Pleurobema oviforme — Ortmann (1918), Ortmann (1924)
Pleurobema (s.s.) oviformis — Frierson (1927)

Unio decisus Küster, 1852

‘Lea’ Küster, 1852. Conchyl. Cab. 9(2): 41, pl. 8, fig. 1. [in part]
(misidentified reference to Unio decisus Lea, 1831)

Pleurobema (s.s.) raveneliana — Simpson (1900)
Pleurobema (s.s.) ravenelianum — Simpson (1914)
Quadrula (Pleuronaia) argentea — Frierson (1927)
Pleurobema oviforme — Parmalee & Bogan (1998)

Unio clavus Call, 1885

‘Lamarck’ Call, 1885. unknown publication: ??.
(misidentified reference to Unio clava Lamarck, 1819)

Pleurobema oviforme — Ortmann (1924)

Unio conasangœnsis B.H. Wright, 1888

‘Lea’ B.H. Wright, 1888. Check List N. Amer. Unionidae etc.: [2].
(error for Unio conasaugaensis Lea, 1872)

Unio accuens Paetel, 1890

‘Lea’ Paetel, 1890. Cat. Conch.-Samml. 3: 143.
(error for Unio acuens Lea, 1871)

Pleurobema swordiana Simpson, 1900

‘Wright’ Simpson, 1900. Proc. Acad. Nat. Sci. 52: 81, pl. 4, fig. 4.
(reference to Unio swordianus S.H. Wright, 1897)

Pleurobema (s.s.) swordiana — Simpson (1900)
Quadrula (Pleuronaia) valida — Frierson (1927)

Pleurobema fassinans Ortmann, 1913

‘Lea’ Ortmann, 1913. Proc. Amer. Phil. Soc. 52(210): 310.
‘Lea’ Ortmann, 1914. Nautilus 28(3): 31.
(misidentified reference to Unio fassinans Lea, 1868)

Pleurobema oviforme argenteum — Ortmann (1918)
Pleurobema oviforme — Parmalee & Bogan (1998)