Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae | Pseudodontini | Pseudodontina

genus Bineurus Simpson, 1900: cladomics

5 Recent valid species (5 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 5 (5 total)
analyzed in 42 cladograms in 29 publications
maximum number of terminals in an analysis = 5 (mean = 2.71, median = 3)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 36
maximum number of included terminals = 5, (mean = 3 median = 3)
proportion of cladograms that support basic monophyly = 0.92

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Bineurus cladogram distances
PCoA Ordination

NJ Tree of Bineurus cladogram distances
NJ Tree

Cladograms

Jeratthitikul et al., 2025 [Publ 4423], fig. 1

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
5 5 3 5 46 34 9 0 
                          +-- Lannanaia           
  +-----------------------+                       
  |                       +-- Pilsbryoconcha      
  |                                               
--+  +----------------------- Songkhlanaia        
  |  |                                            
  |  |              +-------- Sundadontina        
  +--+  +-----------+                             
     |  |           |  +----- Isannaia            
     |  |           +--+                          
     |  |              |  +-- Namkongnaia         
     +--+              +--+                       
        |                 +-- Pseudodon           
        |                                         
        |  +----------------- Nyeinchanconcha     
        +--+                                      
           |  +-------------- Thaiconcha          
           +--+                                   
              |  +----------- Bineurus_loeiensis **  
              +--+                                
                 |  +-------- Bineurus_exilis **     
                 +--+                             
                    |  +----- Bineurus_mouhotii **   
                    +--+                          
                       |  +-- Bineurus_anodontinum **
                       +--+                       
                          +-- Bineurus_panhai **

Konopleva et al., 2025 [Publ 4446], fig. 4

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
4 4 2 4 41 36 9 3 
  +-------------------------- Lannanaia           
  |                                               
--+  +----------------------- Pilsbryoconcha      
  |  |                                            
  +--+  +-------------------- Songkhlanaia        
     |  |                                         
     |  |           +-------- Sundadontina        
     +--+  +--------+                             
        |  |        |  +----- Pseudodon           
        |  |        +--+                          
        |  |           |  +-- Namkongnaia         
        +--+           +--+                       
           |              +-- Isannaia            
           |                                      
           |  +-------------- Nyeinchanconcha     
           +--+                                   
              |  +----------- Thaiconcha          
              +--+                                
                 |  +-------- Bineurus_loeiensis **  
                 +--+                             
                    |  +----- Bineurus_anodontinum **
                    +--+                          
                       |  +-- Bineurus_exilis **     
                       +--+                       
                          +-- Bineurus_mouhotii **

Jeratthitikul & Sutcharit, 2023 [Publ 4289], fig. 1

iTtTbiriSoTroTroSiD
5 5 3 5 8 6 6 6 
     +----------- Bineurus_mouhotii **   
  +--+                                
  |  |  +-------- Bineurus_loeiensis **  
  |  +--+                             
  |     |  +----- Bineurus_panhai **     
  |     +--+                          
  |        |  +-- Bineurus_anodontinum **
--+        +--+                       
  |           +-- Bineurus_exilis **     
  |                                   
  |        +----- Namkongnaia         
  |     +--+                          
  |     |  |  +-- Sundadontina        
  |     |  +--+                       
  +-----+     +-- Nyeinchanconcha     
        |                             
        |  +----- Pseudodon           
        +--+                          
           |  +-- Pilsbryoconcha      
           +--+                       
              +-- Thaiconcha

Bolotov et al., 2023 [Publ 4312], fig. S2

iTtTbiriSoTroTroSiD
5 5 3 5 36 31 7 4 
  +-------------------------- Songkhlanaia        
  |                                               
  |                       +-- Nyeinchanconcha     
--+  +--------------------+                       
  |  |                    +-- Sundadontina        
  |  |                                            
  +--+                    +-- Pseudodon           
     |  +-----------------+                       
     |  |                 +-- Pilsbryoconcha      
     +--+                                         
        |  +----------------- Namkongnaia         
        +--+                                      
           |  +-------------- Thaiconcha          
           +--+                                   
              |  +----------- Bineurus_loeiensis **  
              +--+                                
                 |  +-------- Bineurus_panhai **     
                 +--+                             
                    |  +----- Bineurus_anodontinum **
                    +--+                          
                       |  +-- Bineurus_exilis **     
                       +--+                       
                          +-- Bineurus_mouhotii **

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iTtTbiriSoTroTroSiD
5 5 3 5 55 32 9 4 
  +-------------------------- Pseudodon           
  |                                               
  |                       +-- Isannaia            
  |                 +-----+                       
--+                 |     +-- Pilsbryoconcha      
  |  +--------------+                             
  |  |              |  +----- Namkongnaia         
  |  |              +--+                          
  |  |                 |  +-- Lannanaia           
  +--+                 +--+                       
     |                    +-- Songkhlanaia        
     |                                            
     |  +-------------------- Sundadontina        
     +--+                                         
        |  +----------------- Nyeinchanconcha     
        +--+                                      
           |  +-------------- Thaiconcha          
           +--+                                   
              |  +----------- Bineurus_loeiensis **  
              +--+                                
                 |  +-------- Bineurus_panhai **     
                 +--+                             
                    |  +----- Bineurus_anodontinum **
                    +--+                          
                       |  +-- Bineurus_mouhotii **   
                       +--+                       
                          +-- Bineurus_exilis **

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
4 4 2 4 79 22 7 3 
  +----------------------------- Pilsbryoconcha      
--+                                                  
  |  +-------------------------- Songkhlanaia        
  +--+                                               
     |  +----------------------- Namkongnaia         
     +--+                                            
        |  +-------------------- Pseudodon           
        +--+                                         
           |  +----------------- Sundadontina        
           +--+                                      
              |  +-------------- Nyeinchanconcha     
              +--+                                   
                 |  +----------- Thaiconcha          
                 +--+                                
                    |  +-------- Bineurus_loeiensis **  
                    +--+                             
                       |  +----- Bineurus_mouhotii **   
                       +--+                          
                          |  +-- Bineurus_anodontinum **
                          +--+                       
                             +-- Bineurus_exilis **

Bolotov et al., 2023 [Publ 4312], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 4 35 30 7 3 
  +-------------------------- Pilsbryoconcha      
  |                                               
--+  +----------------------- Songkhlanaia        
  |  |                                            
  +--+                    +-- Namkongnaia         
     |  +-----------------+                       
     |  |                 +-- Pseudodon           
     +--+                                         
        |  +----------------- Sundadontina        
        +--+                                      
           |  +-------------- Nyeinchanconcha     
           +--+                                   
              |  +----------- Thaiconcha          
              +--+                                
                 |  +-------- Bineurus_loeiensis **  
                 +--+                             
                    |  +----- Bineurus_mouhotii **   
                    +--+                          
                       |  +-- Bineurus_anodontinum **
                       +--+                       
                          +-- Bineurus_exilis **

Bolotov et al., 2023 [Publ 4312], fig. S1

iTtTbiriSoTroTroSiD
4 4 2 4 35 30 7 3 
  +-------------------- Pilsbryoconcha      
  |                                         
  |           +-------- Songkhlanaia        
--+  +--------+                             
  |  |        |  +----- Sundadontina        
  |  |        +--+                          
  |  |           |  +-- Namkongnaia         
  +--+           +--+                       
     |              +-- Pseudodon           
     |                                      
     |  +-------------- Nyeinchanconcha     
     +--+                                   
        |  +----------- Thaiconcha          
        +--+                                
           |  +-------- Bineurus_loeiensis **  
           +--+                             
              |  +----- Bineurus_mouhotii **   
              +--+                          
                 |  +-- Bineurus_anodontinum **
                 +--+                       
                    +-- Bineurus_exilis **

Konopleva et al., 2021 [Publ 4213], fig. 1

iTtTbiriSoTroTroSiD
4 4 2 4 23 15 6 3 
                 +----- Pilsbryoconcha_exilis__2
  +--------------+                              
  |              |  +-- Pilsbryoconcha_exilis__1
  |              +--+                           
  |                 +-- Namkongnaia             
--+                                             
  |                 +-- Pseudodon               
  |  +--------------+                           
  |  |              +-- Sundadontina            
  +--+                                          
     |  +-------------- Nyeinchanconcha         
     +--+                                       
        |  +----------- Thaiconcha              
        +--+                                    
           |  +-------- Bineurus_loeiensis **      
           +--+                                 
              |  +----- Bineurus_mouhotii **       
              +--+                              
                 |  +-- Bineurus_anodontinum **    
                 +--+                           
                    +-- Bineurus_exilis **

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iTtTbiriSoTroTroSiD
3 3 0 3 32 14 6 3 
           +----- Pilsbryoconcha_exilis       
  +--------+                                  
  |        |  +-- Pilsbryoconcha_linguaeformis
  |        +--+                               
  |           +-- Namkongnaia_lemeslei__2     
  |                                           
--+     +-------- Sundadontina                
  |  +--+                                     
  |  |  |  +----- Pseudodon                   
  |  |  +--+                                  
  |  |     |  +-- Namkongnaia_lemeslei__1     
  +--+     +--+                               
     |        +-- Namkongnaia_inkhavilayi     
     |                                        
     |  +-------- Nyeinchanconcha             
     |  |                                     
     +--+  +----- Thaiconcha                  
        |  |                                  
        +--+  +-- Bineurus_panhai **             
           |  |                               
           +--+-- Bineurus_exilis **             
              |                               
              +-- Bineurus_mouhotii **

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
3 3 1 3 109 14 5 2 
        +----- Bineurus_loeiensis **
  +-----+                        
  |     |  +-- Bineurus_exilis **   
  |     +--+                     
--+        +-- Bineurus_mouhotii ** 
  |                              
  |  +-------- Nyeinchanconcha   
  |  |                           
  +--+     +-- Thaiconcha        
     |  +--+                     
     |  |  +-- Pilsbryoconcha    
     +--+                        
        |  +-- Pseudodon         
        +--+                     
           +-- Sundadontina

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
3 3 1 3 136 9 5 4 
  +----------------- Pilsbryoconcha       
  |                                       
  |              +-- Sundadontina_tumida  
--+  +-----------+                        
  |  |           +-- Pseudodon            
  |  |                                    
  +--+           +-- Nyeinchanconcha      
     |  +--------+                        
     |  |        +-- Sundadontina_cumingii
     +--+                                 
        |  +-------- Thaiconcha           
        +--+                              
           |  +----- Bineurus_exilis **      
           +--+                           
              |  +-- Bineurus_mouhotii **    
              +--+                        
                 +-- Bineurus_loeiensis **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
3 3 1 3 136 10 5 4 
  +-------- Thaiconcha        
--+                           
  |  +----- Bineurus_exilis **   
  +--+                        
     |  +-- Bineurus_mouhotii ** 
     +--+                     
        +-- Bineurus_loeiensis **

Konopleva et al., 2019 [Publ 3900], fig. 1

iTtTbiriSoTroTroSiD
3 8 0 3 49 5 2 8 
  +-- Bineurus_mouhotii **         
  |                             
  +-- Pseudodon_cambodjensis    
  |                             
  +-- Pseudodon_mekongi         
  |                             
  +-- Pilsbryoconcha_exilis     
--+                             
  +-- Pilsbryoconcha_carinifera 
  |                             
  +-- Bineurus_loeiensis **        
  |                             
  +-- Bineurus_exilis **           
  |                             
  +-- Pseudodon_vondembuschianus

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 115 14 5 2 
        +----- Bineurus_loeiensis **
  +-----+                        
  |     |  +-- Bineurus_exilis **   
  |     +--+                     
--+        +-- Bineurus_mouhotii ** 
  |                              
  |  +-------- Nyeinchanconcha   
  |  |                           
  +--+     +-- Thaiconcha        
     |  +--+                     
     |  |  +-- Pilsbryoconcha    
     +--+                        
        |  +-- Pseudodon         
        +--+                     
           +-- Sundadontina

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
3 8 0 3 80 5 2 8 
  +-- Bineurus_mouhotii **         
  |                             
  +-- Pseudodon_cambodjensis    
  |                             
  +-- Pseudodon_mekongi         
  |                             
  +-- Pilsbryoconcha_exilis     
--+                             
  +-- Pilsbryoconcha_carinifera 
  |                             
  +-- Bineurus_loeiensis **        
  |                             
  +-- Bineurus_exilis **           
  |                             
  +-- Pseudodon_vondembuschianus

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
3 3 1 3 42 11 5 2 
     +-------- Nyeinchanconcha   
  +--+                           
  |  |  +----- Bineurus_loeiensis **
  |  +--+                        
  |     |  +-- Bineurus_mouhotii ** 
  |     +--+                     
--+        +-- Bineurus_exilis **   
  |                              
  |        +-- Thaiconcha        
  |     +--+                     
  |     |  +-- Sundadontina      
  +-----+                        
        |  +-- Pilsbryoconcha    
        +--+                     
           +-- Pseudodon

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
3 3 1 3 196 9 5 2 
  +----------------- Pilsbryoconcha       
  |                                       
  |              +-- Sundadontina_tumida  
--+  +-----------+                        
  |  |           +-- Pseudodon            
  |  |                                    
  +--+           +-- Nyeinchanconcha      
     |  +--------+                        
     |  |        +-- Sundadontina_cumingii
     +--+                                 
        |  +-------- Thaiconcha           
        +--+                              
           |  +----- Bineurus_loeiensis **   
           +--+                           
              |  +-- Bineurus_mouhotii **    
              +--+                        
                 +-- Bineurus_exilis **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 198 9 5 2 
  +----------------- Pilsbryoconcha    
  |                                    
--+              +-- Pseudodon         
  |  +-----------+                     
  |  |           +-- Sundadontina      
  +--+                                 
     |  +----------- Nyeinchanconcha   
     +--+                              
        |  +-------- Thaiconcha        
        +--+                           
           |  +----- Bineurus_loeiensis **
           +--+                        
              |  +-- Bineurus_mouhotii ** 
              +--+                     
                 +-- Bineurus_exilis **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
3 3 1 3 198 9 5 2 
  +-------- Nyeinchanconcha       
  |                               
  +-------- Thaiconcha            
  |                               
  +-------- Sundadontina_cumingii 
  |                               
  +-------- Sundadontina_tumida__1
--+                               
  +-------- Sundadontina_tumida__2
  |                               
  +-------- Pilsbryoconcha        
  |                               
  +-------- Pseudodon             
  |                               
  |  +----- Bineurus_loeiensis **    
  +--+                            
     |  +-- Bineurus_mouhotii **     
     +--+                         
        +-- Bineurus_exilis **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
3 3 1 3 122 9 5 4 
        +-- Nyeinchanconcha      
  +-----+                        
  |     +-- Sundadontina_cumingii
  |                              
  |     +-- Sundadontina_tumida  
  +-----+                        
  |     +-- Pseudodon            
--+                              
  |     +-- Thaiconcha           
  +-----+                        
  |     +-- Pilsbryoconcha       
  |                              
  |  +----- Bineurus_exilis **      
  +--+                           
     |  +-- Bineurus_mouhotii **    
     +--+                        
        +-- Bineurus_loeiensis **

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 33 15 6 4 
  +----------------- Pilsbryoconcha       
  |                                       
  |              +-- Namkongnaia          
  |        +-----+                        
  |        |     +-- Pseudodon            
--+  +-----+                              
  |  |     |  +----- Sundadontina_tumida  
  |  |     +--+                           
  |  |        |  +-- Sundadontina_brandti 
  |  |        +--+                        
  +--+           +-- Sundadontina_taskaevi
     |                                    
     |           +-- Nyeinchanconcha      
     |  +--------+                        
     |  |        +-- Sundadontina_cumingii
     +--+                                 
        |  +-------- Thaiconcha           
        +--+                              
           |  +----- Bineurus_mouhotii **    
           +--+                           
              |  +-- Bineurus_panhai **      
              +--+                        
                 +-- Bineurus_exilis **

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
3 3 1 3 42 11 5 2 
           +-- Thaiconcha        
  +--------+                     
  |        +-- Sundadontina      
  |                              
  |        +-- Pilsbryoconcha    
--+--------+                     
  |        +-- Pseudodon         
  |                              
  |  +-------- Nyeinchanconcha   
  +--+                           
     |  +----- Bineurus_loeiensis **
     +--+                        
        |  +-- Bineurus_mouhotii ** 
        +--+                     
           +-- Bineurus_exilis **

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
2 2 0 2 35 4 3 -- 
  +----------- Pilsbryoconcha      
--+                                
  |  +-------- Pseudodon           
  +--+                             
     |  +----- Thaiconcha          
     +--+                          
        |  +-- Bineurus_anodontinum **
        +--+                       
           +-- Bineurus_loeiensis **

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 35 4 3 -- 
  +----------- Pilsbryoconcha      
--+                                
  |  +-------- Pseudodon           
  +--+                             
     |  +----- Thaiconcha          
     +--+                          
        |  +-- Bineurus_anodontinum **
        +--+                       
           +-- Bineurus_loeiensis **

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 30 5 4 -- 
  +-------------- Pilsbryoconcha      
--+                                   
  |  +----------- Pseudodon           
  +--+                                
     |  +-------- Sundadontina        
     +--+                             
        |  +----- Thaiconcha          
        +--+                          
           |  +-- Bineurus_loeiensis **  
           +--+                       
              +-- Bineurus_anodontinum **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
2 2 0 2 74 13 6 -- 
        +----- Pilsbryoconcha   
  +-----+                       
  |     |  +-- Namkongnaia      
  |     +--+                    
  |        +-- Pseudodon        
--+                             
  |        +-- Nyeinchanconcha  
  |  +-----+                    
  |  |     +-- Sundadontina     
  +--+                          
     |  +----- Thaiconcha       
     +--+                       
        |  +-- Bineurus_panhai **  
        +--+                    
           +-- Bineurus_mouhotii **

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 46 10 5 -- 
  +----------------- Pilsbryoconcha    
--+                                    
  |  +-------------- Pseudodon         
  +--+                                 
     |  +----------- Sundadontina      
     +--+                              
        |  +-------- Nyeinchanconcha   
        +--+                           
           |  +----- Thaiconcha        
           +--+                        
              |  +-- Bineurus_loeiensis **
              +--+                     
                 +-- Bineurus_mouhotii **

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 44 6 4 -- 
           +-- Pilsbryoconcha      
  +--------+                       
  |        +-- Pseudodon           
--+                                
  |  +-------- Sundadontina        
  +--+                             
     |  +----- Thaiconcha          
     +--+                          
        |  +-- Bineurus_loeiensis **  
        +--+                       
           +-- Bineurus_anodontinum **

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
2 2 0 2 35 4 3 -- 
  +----------- Pilsbryoconcha      
--+                                
  |  +-------- Pseudodon           
  +--+                             
     |  +----- Thaiconcha          
     +--+                          
        |  +-- Bineurus_anodontinum **
        +--+                       
           +-- Bineurus_loeiensis **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
2 2 0 2 74 13 6 -- 
        +----- Pilsbryoconcha   
  +-----+                       
  |     |  +-- Namkongnaia      
  |     +--+                    
  |        +-- Pseudodon        
--+                             
  |        +-- Nyeinchanconcha  
  |  +-----+                    
  |  |     +-- Sundadontina     
  +--+                          
     |  +----- Thaiconcha       
     +--+                       
        |  +-- Bineurus_panhai **  
        +--+                    
           +-- Bineurus_mouhotii **

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
2 5 0 2 35 4 3 6 
  +-------- Pilsbryoconcha      
  |                             
--+  +----- Bineurus_anodontinum **
  |  |                          
  +--+----- Bineurus_loeiensis **  
     |                          
     |  +-- Thaiconcha          
     +--+                       
        +-- Pseudodon

Chen et al., 2025 [Publ 4663], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 47 10 5 -- 
  +----------------- Pilsbryoconcha    
--+                                    
  |  +-------------- Pseudodon         
  +--+                                 
     |  +----------- Sundadontina      
     +--+                              
        |  +-------- Nyeinchanconcha   
        +--+                           
           |  +----- Thaiconcha        
           +--+                        
              |  +-- Bineurus_loeiensis **
              +--+                     
                 +-- Bineurus_mouhotii **

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 35 4 3 -- 
  +----------- Pilsbryoconcha      
--+                                
  |  +-------- Pseudodon           
  +--+                             
     |  +----- Thaiconcha          
     +--+                          
        |  +-- Bineurus_anodontinum **
        +--+                       
           +-- Bineurus_loeiensis **

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 45 10 5 -- 
  +----------------- Pilsbryoconcha    
--+                                    
  |  +-------------- Pseudodon         
  +--+                                 
     |  +----------- Sundadontina      
     +--+                              
        |  +-------- Nyeinchanconcha   
        +--+                           
           |  +----- Thaiconcha        
           +--+                        
              |  +-- Bineurus_loeiensis **
              +--+                     
                 +-- Bineurus_mouhotii **

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
2 2 0 2 35 4 3 -- 
  +-------- Pilsbryoconcha      
  |                             
--+     +-- Bineurus_anodontinum **
  |  +--+                       
  |  |  +-- Bineurus_loeiensis **  
  +--+                          
     |  +-- Thaiconcha          
     +--+                       
        +-- Pseudodon

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 3 3 -- 
  +-------- Pilsbryoconcha      
--+                             
  |  +----- Bineurus_anodontinum **
  +--+                          
     |  +-- Pseudodon           
     +--+                       
        +-- Sundadontina

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 30 5 4 -- 
  +-------- Sundadontina   
  |                        
--+     +-- Nyeinchanconcha
  |  +--+                  
  |  |  +-- Bineurus_exilis **
  +--+                     
     |  +-- Pilsbryoconcha 
     +--+                  
        +-- Pseudodon

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
1 1 0 1 79 4 3 -- 
  +-------- Pilsbryoconcha 
--+                        
  |  +----- Pseudodon      
  +--+                     
     |  +-- Bineurus_exilis **
     +--+                  
        +-- Nyeinchanconcha

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
1 1 0 1 121 5 4 -- 
  +-------- Pilsbryoconcha   
  |                          
--+  +----- Pseudodon        
  |  |                       
  +--+  +-- Nyeinchanconcha  
     |  |                    
     +--+-- Bineurus_mouhotii **
        |                    
        +-- Sundadontina

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 3 3 -- 
  +-------- Bineurus_anodontinum **
--+                             
  |  +----- Pilsbryoconcha      
  +--+                          
     |  +-- Pseudodon           
     +--+                       
        +-- Sundadontina

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 39 5 4 -- 
  +----------- Pilsbryoconcha 
--+                           
  |  +-------- Pseudodon      
  +--+                        
     |  +----- Sundadontina   
     +--+                     
        |  +-- Nyeinchanconcha
        +--+                  
           +-- Bineurus_exilis **