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Taxonomy

all genera

Unionidae

Margaritiferidae

Hyriidae

Etheriidae

Mycetopodidae

Iridinidae


Species & Genus Richness by
Taxonomy

Geography

Summary

North America

Central America

South America

Afrotropics

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East Asia

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Species Richness & Endemism by
Geography

Data last updated
Mon 1 Dec 2025

Publication Konopleva et al. (2017): summary

publication summary | cladograms

Full Citation

Konopleva, E.S., I.N. Bolotov, I.V. Vikhrev, M.Y. Gofarov & A.V. Kondakov. 2017. An integrative approach underscores the taxonomic status of Lamellidens exolescens, a freshwater mussel from the Oriental tropics (Bivalvia: Unionidae). Systematics and Biodiversity 15(3): 204-217. Available electronically.

Attributed Nominal Species

Radiatula bonneaudii Whelan et al., 2011 = Radiatula chaudhurii

‘Eydoux’ of Authors.
Whelan et al., 2011. MPE 61: 506.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): [4].
Bolotov et al., 2017. Sci. Repts. 7: 10, Table 3 [as ‘sp. 1’].
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2017. Sci. Repts. 7: [5].
Bolotov et al., 2018. Sci. Repts. 8: Table S1.
Pfeiffer et al., 2018. J. Moll. Studies 84(4): [4]. [in part]
Jeratthitikul et al., 2019. Tropical Nat. Hist. 19(1): 25.
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Radiatula bonneaudii sp. 1’].
Huang et al., 2019. MPE 130: Table S4.
Ortiz-Sepulveda et al., 2020. MPE 148: Table S1.
(misidentified reference to Unio bonneaudii Eydoux, 1838)

image of type lot

Pseudodon cambodjensis Pfeiffer & Graf, 2013 = Pseudodon cambodjensis

‘Petit’ of Authors.
Pfeiffer & Graf, 2013. J. Moll. Studies 79: 251.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Zieritz et al., 2016. Sci. Total Environ. 571: 1071.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Monodontina cambodjensis’].
Lopes-Lima et al., 2017. MPE 106: 177.
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Monodontina cambodjensis’].
Razak et al., 2018. Conserv. Genet. Resour. 11: 459.
Huang et al., 2019. MPE 130: Table S4 [as ‘Monodontina cambodjensis’].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Monodontina cambodjensis’].
Bolotov et al., 2020. Sci. Repts. 10: Table S1 [as ‘Monodontina cambodiensis’ (sic)].
Konopleva et al., 2021. Sci. Repts. 11: Table S1 [as ‘Monodontina cambodiensis’ (sic)].
Jeratthitikul et al., 2021. Eur. J. Taxonomy 775: 123 [as ‘Monodontina cambodiensis’ (sic)].
Pfeiffer et al., 2021. Invert. Syst. 35: 453 [as ‘Monodontina cambodjensis’].
Wu et al., 2021. ZooKeys 1054: 87 [as ‘Monodontina cambodjensis’].
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
Konopleva et al., 2023. Diversity 15(1), 10: Table S2 [as ‘Monodontina cambodjensis’].
Dai et al., 2023. Invert. Syst. 37(2): Table 1 [as ‘Monodontina cambodjensis’].
Dai et al., 2024. Zoosyst. Evol. 100(4): Table 2.
Dai et al., 2025. Zoosyst. Evol. 101(2): Table 1.
(reference to Monocondylea cambodjensis Petit de la Saussaye, 1865)
Type(s): SEQUENCED SPECIMEN UMMZ_304350, Pursat River, just downstream (north) of Highway 5 bridge, Pursat City, Cambodia.

Contradens contradens Pfeiffer & Graf, 2015 = Lens micropterus

‘Lea’ of Authors.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Konopleva et al., 2019. Ecologica Montenegrina 24: Appendix 1 [as ‘Contradens aff. contradens sp. 1’].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Huang et al., 2019. MPE 130: Table S4 [as ‘Contradens aff. contradens sp. 1’].
Bolotov et al., 2020. Sci. Repts. 10: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Konopleva et al., 2023. Diversity 15(1), 10: Table S2 [as ‘Lens aff. contradens’].
(misidentified reference to Unio contradens Lea, 1838)

image of type lot

Lamellidens corrianus Whelan et al., 2011 = Trapezidens dolichorhynchus

‘Lea’ of Authors.
Whelan et al., 2011. MPE 61: 506.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Lamellidens corrianus sp. 1].
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Pfeiffer et al., 2018. J. Moll. Studies 84(4): [3].
Pfeiffer et al., 2019. MPE 137: 117.
Lopes-Lima et al., 2020. MPE 146: Table 1.
(misidentified reference to Unio corrianus Lea, 1834)
Type(s): SEQUENCED SPECIMEN UMMZ_304642, Ayeryarwaddy River, mud bank on E side of river, Mandalay Division, Burma. 21.48187, 95.30501.

image of type lot

Scabies crispata Pfeiffer & Graf, 2015 = Scabies mandarinus

‘Gould’ of Authors.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): [4].
Ng et al., 2016. PLoS One 11(8), e0161130: [5].
Bolotov et al., 2017. Sci. Repts. 7: [6], Table S1 [as ‘Scabies aff. crispata’].
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
(misidentified reference to Unio crispata Gould, 1843)
Type(s): SEQUENCED SPECIMEN UMMZ_304646, Beung Dang Teuk Leach Community Fishery Pursat Province, Cambodia.

There are 16 nominal species attributed to this publication (none available).
image of type lot

Pseudodon cumingii Pfeiffer & Graf, 2015 = Nyeinchanconcha nyeinchani

‘Lea’ of Authors.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Lopes-Lima et al., 2017. MPE 106: 177.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘”Bineurus” cumingii’].
Huang et al., 2019. MPE 130: Table S4 [as ‘”Bineurus” cumingii’].
Lopes-Lima et al., 2020. MPE 146: Table 1.
(misidentified reference to Anodonta cumingii Lea, 1851)
Type(s): SPECIMEN UMMZ_304648, Nam Phiat, Phonethong, ca. 2 km from confluence, Bolikhamxai Province, Laos. 18.08389, 104.97806.

Anodontites elongata Whelan et al., 2011 = Tamsiella schroeteriana

‘Swainson’ of Authors.
Whelan et al., 2011. MPE 61: 506.
Graf et al., 2015. MPE 85: 3.
Bolotov et al., 2016. MPE 103: 108.
Lopes-Lima et al., 2017. MPE 106: 178.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 208.
Combosch et al., 2017. MPE 107: 195.
Bolotov et al., 2018. Sci. Repts. 8: Table S1.
Huang et al., 2018. MPE 120: 302.
Lopes-Lima et al., 2018. MPE 127: 103.
Huang et al., 2019. MPE 130: Table S4.
Bolotov et al., 2019. Sci. Repts. 9: Table S1.
Pfeiffer et al., 2019. MPE 137: 116.
Guerra et al., 2019. BMC Evol. Biol. 19 (229): 3.
Bolotov et al., 2020. Sci. Repts. 10: Table S1.
Sano et al., 2020. Plankton Benthos Res. 15(2): 101.
Wu et al., 2020. Invert. Syst. 34: 195.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
Chen et al., 2025. Zoosyst. Evol. 101(4): Table 1.
(misidentified reference to Anodon elongatus Swainson, 1823)

image of type lot

Pilsbryoconcha exilis Pfeiffer & Graf, 2015 = Pilsbryoconcha carinifera

‘Lea’ of Authors.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Ng et al., 2016. PLoS One 11(8), e0161130: [5]. [in part]
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘P. aff. exilis sp. 1’].
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘P. aff. exilis sp. 1’].
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘P. aff. exilis sp. 1’].
Huang et al., 2019. MPE 130: Table S4 [as ‘P. aff. exilis sp. 1’].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘P. aff. exilis sp. 1’].
Lopes-Lima et al., 2020. MPE 146: Table 1 [as ‘P. aff. exilis sp. 1’].
Zieritz et al., 2020. Biodivers. Conserv. 29: 2244.
Wu et al., 2021. ZooKeys 1054: 87.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
Konopleva et al., 2023. Diversity 15(1), 10: Table S2 [as ‘P. aff. exilis’].
Dai et al., 2023. Invert. Syst. 37(2): Tables 1-2. [in part]
(misidentified reference to Anodonta exilis Lea, 1838)
Type(s): SEQUENCED SPECIMEN UMMZ_304647, Beung Dang Teuk Leach Community Fishery Pursat Province, Cambodia.

Lamellidens generosus Whelan et al., 2011 = Lamellidens savadiensis

‘Gould’ of Authors.
Whelan et al., 2011. MPE 61: 506.
Pfeiffer & Graf, 2013. J. Moll. Studies 79: 251.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Araujo et al., 2017. Zool. J. Linn. Soc. 179(2): 280.
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Pfeiffer et al., 2018. J. Moll. Studies 84(4): [3].
Pfeiffer et al., 2019. MPE 137: 117.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
(misidentified reference to Unio generosus Gould, 1847)

Pseudodon inoscularis Pfeiffer & Graf, 2013 = Sundadontina tumida

‘Gould’ of Authors.
Pfeiffer & Graf, 2013. J. Moll. Studies 79: 251.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Pseudodon cf. inoscularis’].
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘ “Monodontina” cf. inoscularis sp. 2’].
Huang et al., 2019. MPE 130: Table S4 [as ‘ “Monodontina” cf. inoscularis sp. 2’].
Lopes-Lima et al., 2020. MPE 146: Table 1 [as ‘ “Monodontina” cf. inoscularis sp. 2’].
(misidentified reference to Anodon inoscularis Gould, 1844)

Lamellidens marginalis Bolotov et al., 2017 = Lamellidens jenkinsianus

‘Lamarck’ of Authors.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Lamellidens marginalis sp. 4’].
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207. [in part]
Pavan-Kumar et al., 2022. Mol. Biol. Rep. 49: Table S1.
Wu et al., 2023. Org Divers Evol (2023): Table 2.
Wu et al., 2024. J. Zool. Syst. Evol. Res. 2024, 1499508: Table 2.
(misidentified reference to Unio marginalis Lamarck, 1819)

Lamellidens marginalis Bolotov et al., 2017 = Lamellidens mainwaringi

‘Lamarck’ of Authors.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Lamellidens marginalis sp. 2’ and ‘sp. 3’].
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Lamellidens marginalis sp. 2’ and ‘sp. 3’].
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207. [in part]
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Lamellidens marginalis sp. 3’].
Huang et al., 2019. MPE 130: Table S4 [as ‘Lamellidens marginalis sp. 2’ and ‘sp. 3’].
Jeratthitikul et al., 2019. Tropical Nat. Hist. 19(1): 25.
Konopleva et al., 2020. Ecologica Montenegrina 27: 47.
Bolotov et al., 2020. Sci. Repts. 10: Table S1.
(misidentified reference to Unio marginalis Lamarck, 1819)

Pseudodon mouhotii Pfeiffer & Graf, 2015 = Bineurus exilis

‘Lea’ of Authors.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Lopes-Lima et al., 2017. MPE 106: 177.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Pseudodon aff. mouhotii sp. 3’].
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Bineurus aff. mouhotii sp. 3’].
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Bineurus aff. mouhotii sp. 3’].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Bineurus aff. mouhotii sp. 3’].
Huang et al., 2019. MPE 130: Table S4 [as ‘Bineurus aff. mouhotii sp. 3’].
(misidentified reference to Monocondyloea mouhotii Lea, 1863)

Ensidens sagittarius Pfeiffer & Graf, 2015 = Ensidens telus

‘Lea’ of Authors.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Lopes-Lima et al., 2017. MPE 106: 177.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘E. aff. sagittarius sp. 1].
Huang et al., 2019. MPE 130: Table S4 [as ‘Ensidens aff. sagittarius sp. 1].
Zieritz et al., 2020. Biodivers. Conserv. 29: 2242.
Lopes-Lima et al., 2020. MPE 146: Table 1.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
(misidentified reference to Unio sagittarius Lea, 1856)

Pseudodon vondembuschianus Pfeiffer & Graf, 2015 = Pseudodon vondembuschianus

‘Lea’ of Authors.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310. [in part]
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Pseudodon aff. vondembuschianus sp. 2’]. [in part]
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207. [in part]
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Huang et al., 2019. MPE 130: Table S4 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
(reference to Margaritana vondembuschiana Lea, 1840)

Pseudodon vondembuschianus Graf, 2002 = Pseudodon vondembuschianus

‘Lea’ of Authors.
Graf, 2002. J. Moll. Studies 68: 66.
Graf & Cummings, 2006. Zool. J. Linn. Soc. 148: 348.
Hoeh et al., 2009. Malacologia 51(2): 310.
Whelan et al., 2011. MPE 61: 506. [in part]
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310. [in part]
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Pseudodon aff. vondembuschianus sp. 2’]. [in part]
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207. [in part]
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Huang et al., 2019. MPE 130: Table S4 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
(reference to Margaritana vondembuschiana Lea, 1840)

Taxonomic Opinions

Anodontites elongata (Swainson, 1823)

Castalia ambigua Lamarck, 1819

Chambardia wahlbergi (Krauss, 1848)

Coelatura aegyptiaca (Cailliaud, 1827)

Contradens contradens (Lea, 1838) = Lens contradens

Diplodon suavidicus (Lea, 1856)

Ensidens sagittarius (Lea, 1856)

Etheria elliptica Lamarck, 1807

Lamellidens corrianus (Lea, 1834)

Lamellidens exolescens (Gould, 1843) = Trapezidens exolescens

Lamellidens generosus (Gould, 1847)

Lamellidens marginalis (Lamarck, 1819)

Margaritifera dahurica (Middendorff, 1850)

Margaritifera laevis (Haas, 1910)

Neotrigonia lamarckii (Gray, 1838)

Neotrigonia margaritacea (Lamarck, 1804)

Oxynaia pugio (Benson, 1862) = Indochinella pugio

Parreysia olivacea Prashad, 1930 = Leoparreysia olivacea

Parreysia tavoyensis (Gould, 1843) = Leoparreysia tavoyensis

Pilsbryoconcha exilis (Lea, 1838)

Pseudodon cambodjensis (Petit de la Saussaye, 1865)

Pseudodon cumingii (Lea, 1850) = Sundadontina cumingii

Pseudodon inoscularis (Gould, 1844)

Pseudodon mouhotii (Lea, 1863) = Bineurus mouhotii

Pseudodon vondembuschianus (Lea, 1840)

Radiatula bonneaudii (Eydoux, 1838) = Indonaia bonneaudii

Scabies crispata (Gould, 1843)

Trapezoideus exolescens sensu Brandt (1974)

Triplodon corrugatus (Lamarck, 1819)

 
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