Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae | Lamproscaphini | Lamproscaphina

genus Hyriopsis Conrad, 1853: cladomics

10 Recent valid species (10 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 10 (10 total)
analyzed in 62 cladograms in 40 publications
maximum number of terminals in an analysis = 9 (mean = 2.6, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 28
maximum number of included terminals = 9, (mean = 4.54 median = 5)
proportion of cladograms that support basic monophyly = 0.32

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Hyriopsis cladogram distances
PCoA Ordination

NJ Tree of Hyriopsis cladogram distances
NJ Tree

Cladograms

Konopleva et al., 2023 [Publ 4262], fig. 8

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
0.750 9 10 8 9 73 7 2 **
Cladomics cladogram Ensidens Hyriopsis desowitzi Hyriopsis delaportei Rectidens Hyriopsis panhai Hyriopsis myersiana Hyriopsis khoratensis Hyriopsis bialata Hyriopsis sakhonensis Hyriopsis phuphaniensis Hyriopsis kratiensis

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.750 9 11 9 9 624 8 2 7.0
Cladomics cladogram Ensidens Rectidens Hyriopsis delaportei Hyriopsis desowitzi Hyriopsis panhai Hyriopsis bialata Hyriopsis khoratensis Hyriopsis kratiensis Hyriopsis phuphaniensis Hyriopsis myersiana Hyriopsis sakhonensis

Pfeiffer et al., 2021 [Publ 4126], fig. 27

cst iT tT bi riS oT roT roS iD
1.000 9 9 5 9 2 1 1 7.0
Cladomics cladogram Rectidens Hyriopsis delaportei Hyriopsis desowitzi Hyriopsis khoratensis Hyriopsis kratiensis Hyriopsis phuphaniensis Hyriopsis sakhonensis Hyriopsis myersiana Hyriopsis panhai Hyriopsis bialata

Jeratthitikul et al., 2021 [Publ 4212], fig. 4

cst iT tT bi riS oT roT roS iD
0.750 8 9 7 8 13 3 2 7.0
Cladomics cladogram Ensidens Hyriopsis desowitzi Hyriopsis delaportei Rectidens Hyriopsis myersiana Hyriopsis bialata Hyriopsis panhai Hyriopsis khoratensis Hyriopsis phuphaniensis Hyriopsis sakhonensis

Pfeiffer et al., 2021 [Publ 4126], fig. 9

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 6 23 7 2 6.0
Cladomics cladogram Ensidens Rectidens Hyriopsis delaportei Hyriopsis myersiana Hyriopsis bialata Hyriopsis khoratensis Hyriopsis kratiensis Hyriopsis sakhonensis

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
0.750 5 6 4 5 134 6 2 10.0
Cladomics cladogram Ensidens Hyriopsis bialata Hyriopsis khoratensis Hyriopsis panhai Hyriopsis myersiana Rectidens Hyriopsis desowitzi

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
0.750 5 11 7 5 134 6 2 14.0
Cladomics cladogram Lens Physunio Yaukthwa Hyriopsis desowitzi Rectidens Ensidens Hyriopsis panhai Hyriopsis khoratensis Hyriopsis myersiana Hyriopsis bialata

Bolotov et al., 2017 [Publ 3764], fig. S1

cst iT tT bi riS oT roT roS iD
0.750 5 6 2 5 120 6 2 6.0
Cladomics cladogram Ensidens Hyriopsis desowitzi Rectidens Hyriopsis panhai Hyriopsis myersiana Hyriopsis bialata Hyriopsis khoratensis

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
0.750 5 11 5 5 196 6 2 12.0
Cladomics cladogram Physunio Yaukthwa Lens Hyriopsis desowitzi Rectidens Ensidens Hyriopsis bialata Hyriopsis myersiana Hyriopsis panhai Hyriopsis khoratensis

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
0.750 5 6 4 5 194 6 2 8.0
Cladomics cladogram Ensidens Hyriopsis desowitzi Rectidens Hyriopsis panhai Hyriopsis khoratensis Hyriopsis bialata Hyriopsis myersiana

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.750 5 11 6 5 196 6 2 12.0
Cladomics cladogram Physunio modelli Physunio Physunio superbus Yaukthwa Lens Hyriopsis desowitzi Hyriopsis myersiana Hyriopsis panhai Hyriopsis bialata Hyriopsis khoratensis Rectidens Ensidens

Konopleva et al., 2019 [Publ 3900], fig. 1

cst iT tT bi riS oT roT roS iD
0.750 5 6 3 5 46 6 2 5.0
Cladomics cladogram Ensidens Hyriopsis desowitzi Rectidens Hyriopsis panhai Hyriopsis myersiana Hyriopsis khoratensis Hyriopsis bialata

Konopleva et al., 2019 [Publ 4041], fig. 3

cst iT tT bi riS oT roT roS iD
0.500 5 12 0 5 29 7 2 15.0
Cladomics cladogram Yaukthwa Lens Trapezoideus Physunio Ensidens ingallsianus Ensidens telus Ensidens dugasti Hyriopsis khoratensis Hyriopsis bialata Hyriopsis myersiana Hyriopsis desowitzi Rectidens Hyriopsis panhai Ensidens jaculus 1 Ensidens jaculus 2 Ensidens spiculus Ensidens sagittarius

Bolotov et al., 2020 [Publ 4100], fig. 1

cst iT tT bi riS oT roT roS iD
0.750 5 6 4 5 112 7 2 6.0
Cladomics cladogram Ensidens Hyriopsis desowitzi Rectidens Hyriopsis khoratensis Hyriopsis bialata Hyriopsis myersiana Hyriopsis panhai

Bolotov et al., 2020 [Publ 4100], fig. S1

cst iT tT bi riS oT roT roS iD
0.750 5 6 4 5 106 7 2 4.0
Cladomics cladogram Ensidens Hyriopsis desowitzi Rectidens Hyriopsis panhai Hyriopsis myersiana Hyriopsis khoratensis Hyriopsis bialata

Zieritz et al., 2020 [Publ 4102], fig. 4

cst iT tT bi riS oT roT roS iD
0.750 5 6 2 5 26 6 2 6.0
Cladomics cladogram Ensidens Hyriopsis desowitzi Rectidens Hyriopsis panhai Hyriopsis khoratensis Hyriopsis bialata Hyriopsis myersiana

Zieritz et al., 2016 [Publ 3804], fig. S2-1

cst iT tT bi riS oT roT roS iD
1.000 5 5 2 5 1 1 1 8.0
Cladomics cladogram Rectidens Hyriopsis desowitzi Hyriopsis khoratensis Hyriopsis myersiana Hyriopsis panhai Hyriopsis bialata

Muanta et al., 2019 [Publ 3921], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 5 5 1 8.0
Cladomics cladogram Ensidens Hyriopsis desowitzi Hyriopsis myersiana Hyriopsis khoratensis Hyriopsis panhai

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
0.750 3 7 2 3 78 4 2 10.0
Cladomics cladogram Lens Physunio Hyriopsis desowitzi Rectidens Ensidens Hyriopsis khoratensis Hyriopsis myersiana

Bolotov et al., 2018 [Publ 3826], fig. 2

cst iT tT bi riS oT roT roS iD
0.333 3 8 0 3 79 5 1 13.0
Cladomics cladogram Ensidens telus Ensidens ingallsianus Ensidens jaculus Hyriopsis desowitzi Hyriopsis myersiana Hyriopsis khoratensis Ensidens spiculus Ensidens sagittarius Lens eximius Physunio modelli 1 Yaukthwa nesemanni Yaukthwa panhai Lens comptus Lens novoselovi Physunio modelli 2 Physunio Yaukthwa inlenensis Lens contradens Lens micropterus

Zieritz et al., 2021 [Publ 4218], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 29 2 2 --
Cladomics cladogram Ensidens Rectidens Hyriopsis bialata Hyriopsis desowitzi

Wu et al., 2024 [Publ 4350], fig. 5a

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 34 2 2 --
Cladomics cladogram Ensidens Rectidens Hyriopsis bialata Hyriopsis desowitzi

Wu et al., 2024 [Publ 4350], fig. 5b

cst iT tT bi riS oT roT roS iD
0.500 2 13 0 2 34 2 2 19.0
Cladomics cladogram Hyriopsis bialata Rectidens Khairuloconcha Ensidens Hyriopsis desowitzi Yaukthwa elongatula Trapezoideus Physunio Yaukthwa inlenensis Yaukthwa paiensis Lens Inversidens Lamprotula Chamberlainia Potomida Sinosolenaia Gonidea Microcondylaea Koreosolenaia Parvasolenaia Postolata Ptychorhynchus

Wu et al., 2024 [Publ 4350], fig. 5c

cst iT tT bi riS oT roT roS iD
0.500 2 13 0 2 34 2 2 18.0
Cladomics cladogram Ensidens Hyriopsis desowitzi Rectidens Hyriopsis bialata Khairuloconcha Physunio Yaukthwa paiensis Yaukthwa inlenensis Trapezoideus Yaukthwa elongatula Lens Inversidens Lamprotula Chamberlainia Potomida Sinosolenaia Gonidea Microcondylaea Koreosolenaia Parvasolenaia Postolata Ptychorhynchus

Wu et al., 2024 [Publ 4350], fig. 6-7

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 34 2 2 --
Cladomics cladogram Ensidens Rectidens Hyriopsis bialata Hyriopsis desowitzi

Liu et al., 2024 [Publ 4380], fig. 5

cst iT tT bi riS oT roT roS iD
0.750 2 3 0 2 34 2 2 9.0
Cladomics cladogram Ensidens Hyriopsis bialata Hyriopsis desowitzi Rectidens

Zieritz et al., 2024 [Publ 4381], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 43 3 2 --
Cladomics cladogram Ensidens Rectidens Hyriopsis bialata Hyriopsis desowitzi

Wu et al., 2024 [Publ 4388], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 34 2 2 --
Cladomics cladogram Ensidens Rectidens Hyriopsis bialata Hyriopsis desowitzi

Pfeiffer et al., 2018 [Publ 3827], fig. 1

cst iT tT bi riS oT roT roS iD
0.750 1 3 0 1 6 2 2 11.0
Cladomics cladogram Physunio Trapezoideus Lens Ensidens Hyriopsis 1 Hyriopsis 2 Rectidens

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 2 2 --
Cladomics cladogram Ensidens Hyriopsis bialata Rectidens

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 2 2 --
Cladomics cladogram Ensidens Hyriopsis bialata Rectidens

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 2 2 --
Cladomics cladogram Ensidens Hyriopsis bialata Rectidens

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 34 2 2 --
Cladomics cladogram Ensidens Hyriopsis bialata Rectidens

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 75 2 2 --
Cladomics cladogram Ensidens Hyriopsis bialata Rectidens

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 75 2 2 --
Cladomics cladogram Ensidens Rectidens Hyriopsis bialata

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 58 2 2 --
Cladomics cladogram Ensidens Hyriopsis bialata Rectidens

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 1 1 --
Cladomics cladogram Hyriopsis bialata Rectidens

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 1 1 --
Cladomics cladogram Hyriopsis bialata Rectidens

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 3 1 --
Cladomics cladogram Hyriopsis desowitzi Ensidens

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 3 1 --
Cladomics cladogram Hyriopsis desowitzi Ensidens

Dai et al., 2023 [Publ 4285], fig. 3-BI

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 44 1 1 --
Cladomics cladogram Hyriopsis bialata Ensidens

Dai et al., 2023 [Publ 4285], fig. 3-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 44 1 1 --
Cladomics cladogram Hyriopsis bialata Ensidens

Dai et al., 2023 [Publ 4285], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 1 1 --
Cladomics cladogram Hyriopsis bialata Rectidens

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 1 1 --
Cladomics cladogram Hyriopsis bialata Rectidens

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 1 1 --
Cladomics cladogram Hyriopsis bialata Rectidens

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 1 1 --
Cladomics cladogram Hyriopsis bialata Rectidens

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 1 1 --
Cladomics cladogram Hyriopsis bialata Rectidens

Matos et al., 2024 [Publ 4383], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 1 1 --
Cladomics cladogram Hyriopsis bialata Rectidens

Matos et al., 2024 [Publ 4384], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 1 1 --
Cladomics cladogram Hyriopsis bialata Rectidens

Wu et al., 2024 [Publ 4388], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 1 1 --
Cladomics cladogram Hyriopsis bialata Rectidens

Dai et al., 2024 [Publ 4389], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 45 1 1 --
Cladomics cladogram Ensidens Hyriopsis bialata

Zhang et al., 2024 [Publ 4397], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 26 1 1 --
Cladomics cladogram Hyriopsis bialata Rectidens

Dai et al., 2025 [Publ 4440], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 86 1 1 --
Cladomics cladogram Hyriopsis bialata Rectidens

Dai et al., 2025 [Publ 4441], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 46 1 1 --
Cladomics cladogram Ensidens Hyriopsis bialata

Chen et al., 2025 [Publ 4448], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 84 1 1 --
Cladomics cladogram Hyriopsis bialata Rectidens

Chen et al., 2025 [Publ 4663], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 1 1 --
Cladomics cladogram Ensidens Hyriopsis bialata

Klymus et al., 2025 [Publ 4814], fig. 3A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 1 1 --
Cladomics cladogram Hyriopsis bialata Rectidens

Klymus et al., 2025 [Publ 4814], fig. 3B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 1 1 --
Cladomics cladogram Hyriopsis bialata Rectidens

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 139 1 1 --
Cladomics cladogram Hyriopsis bialata Rectidens

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 46 1 1 --
Cladomics cladogram Hyriopsis bialata Rectidens

Wu et al., 2026 [Publ 4811], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 1 1 --
Cladomics cladogram Hyriopsis bialata Rectidens

Zieritz et al., 2026 [Publ 4812], fig. 7a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 3 1 --
Cladomics cladogram Hyriopsis bialata Rectidens