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Data last updated
Mon 1 Dec 2025

Bivalvia | Unionoida | Unionoidea | Unionidae | Gonideinae | Rectidentini

genus Hyriopsis Conrad, 1853: cladomics

10 Recent valid species (10 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 10 (10 total)
analyzed in 54 cladograms in 35 publications
maximum number of terminals in an analysis = 9 (mean = 2.69, median = 1.5)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 27
maximum number of included terminals = 9, (mean = 4.37 median = 5)
proportion of cladograms that support basic monophyly = 0.33

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Hyriopsis cladogram distances
PCoA Ordination

NJ Tree of Hyriopsis cladogram distances
NJ Tree

Cladograms

Konopleva et al., 2023 [Publ 4262], fig. 8

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
9 10 8 9 73 6 2 0 
  +----------------------- Ensidens               
  |                                               
--+                    +-- Hyriopsis_desowitzi **    
  |  +-----------------+                          
  |  |                 +-- Hyriopsis_delaportei **   
  +--+                                            
     |  +----------------- Rectidens              
     |  |                                         
     +--+  +-------------- Hyriopsis_panhai **       
        |  |                                      
        +--+  +----------- Hyriopsis_myersiana **    
           |  |                                   
           +--+        +-- Hyriopsis_khoratensis **  
              |  +-----+                          
              |  |     +-- Hyriopsis_bialata **      
              +--+                                
                 |  +----- Hyriopsis_sakhonensis **  
                 +--+                             
                    |  +-- Hyriopsis_phuphaniensis **
                    +--+                          
                       +-- Hyriopsis_kratiensis **

Pfeiffer et al., 2021 [Publ 4126], fig. 9

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
6 6 4 6 23 7 2 6 
  +-------------------- Ensidens             
--+                                          
  |  +----------------- Rectidens            
  +--+                                       
     |  +-------------- Hyriopsis_delaportei ** 
     +--+                                    
        |  +----------- Hyriopsis_myersiana **  
        +--+                                 
           |  +-------- Hyriopsis_bialata **    
           +--+                              
              |  +----- Hyriopsis_khoratensis **
              +--+                           
                 |  +-- Hyriopsis_kratiensis ** 
                 +--+                        
                    +-- Hyriopsis_sakhonensis **

Pfeiffer et al., 2021 [Publ 4126], fig. 27

iTtTbiriSoTroTroSiD
9 9 5 9 2 1 1 7 
  +-------------- Rectidens              
  |                                      
  |           +-- Hyriopsis_delaportei **   
--+  +--------+                          
  |  |        +-- Hyriopsis_desowitzi **    
  |  |                                   
  +--+  +-------- Hyriopsis_khoratensis **  
     |  |                                
     |  |     +-- Hyriopsis_kratiensis **   
     |  +-----+                          
     +--+     +-- Hyriopsis_phuphaniensis **
        |                                
        |  +----- Hyriopsis_sakhonensis **  
        |  |                             
        +--+  +-- Hyriopsis_myersiana **    
           |  |                          
           +--+-- Hyriopsis_panhai **       
              |                          
              +-- Hyriopsis_bialata **

Jeratthitikul et al., 2021 [Publ 4212], fig. 4

iTtTbiriSoTroTroSiD
8 9 7 8 13 3 2 7 
  +----------------- Ensidens               
  |                                         
  |              +-- Hyriopsis_desowitzi **    
--+  +-----------+                          
  |  |           +-- Hyriopsis_delaportei **   
  |  |                                      
  +--+  +----------- Rectidens              
     |  |                                   
     |  |     +----- Hyriopsis_myersiana **    
     +--+  +--+                             
        |  |  |  +-- Hyriopsis_bialata **      
        |  |  +--+                          
        +--+     +-- Hyriopsis_panhai **       
           |                                
           |  +----- Hyriopsis_khoratensis **  
           +--+                             
              |  +-- Hyriopsis_phuphaniensis **
              +--+                          
                 +-- Hyriopsis_sakhonensis **

Zieritz et al., 2016 [Publ 3804], fig. S2-1

iTtTbiriSoTroTroSiD
5 5 2 5 1 1 1 8 
  +----------- Rectidens            
  |                                 
--+  +-------- Hyriopsis_desowitzi **  
  |  |                              
  +--+  +----- Hyriopsis_khoratensis **
     |  |                           
     +--+  +-- Hyriopsis_myersiana **  
        |  |                        
        +--+-- Hyriopsis_panhai **     
           |                        
           +-- Hyriopsis_bialata **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
5 6 4 5 194 6 2 8 
  +-------------- Ensidens             
  |                                    
--+  +----------- Hyriopsis_desowitzi **  
  |  |                                 
  +--+  +-------- Rectidens            
     |  |                              
     +--+     +-- Hyriopsis_panhai **     
        |  +--+                        
        |  |  +-- Hyriopsis_khoratensis **
        +--+                           
           |  +-- Hyriopsis_bialata **    
           +--+                        
              +-- Hyriopsis_myersiana **

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
5 6 4 5 107 7 2 4 
  +----------------- Ensidens             
--+                                       
  |  +-------------- Hyriopsis_desowitzi **  
  +--+                                    
     |  +----------- Rectidens            
     +--+                                 
        |  +-------- Hyriopsis_panhai **     
        +--+                              
           |  +----- Hyriopsis_myersiana **  
           +--+                           
              |  +-- Hyriopsis_khoratensis **
              +--+                        
                 +-- Hyriopsis_bialata **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
5 6 4 5 134 6 2 10 
  +-------------- Ensidens             
  |                                    
--+           +-- Hyriopsis_bialata **    
  |  +--------+                        
  |  |        +-- Hyriopsis_khoratensis **
  +--+                                 
     |  +-------- Hyriopsis_panhai **     
     +--+                              
        |  +----- Hyriopsis_myersiana **  
        +--+                           
           |  +-- Rectidens            
           +--+                        
              +-- Hyriopsis_desowitzi **

Konopleva et al., 2019 [Publ 3900], fig. 1

iTtTbiriSoTroTroSiD
5 6 3 5 47 6 2 5 
  +-------------- Ensidens             
  |                                    
--+  +----------- Hyriopsis_desowitzi **  
  |  |                                 
  +--+----------- Rectidens            
     |                                 
     |  +-------- Hyriopsis_panhai **     
     +--+                              
        |  +----- Hyriopsis_myersiana **  
        +--+                           
           |  +-- Hyriopsis_khoratensis **
           +--+                        
              +-- Hyriopsis_bialata **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
5 11 7 5 134 6 2 14 
        +-- Hyriopsis_desowitzi **  
        |                        
  +-----+-- Rectidens            
  |     |                        
  |     +-- Ensidens             
--+                              
  |  +----- Hyriopsis_panhai **     
  |  |                           
  +--+----- Hyriopsis_khoratensis **
     |                           
     |  +-- Hyriopsis_myersiana **  
     +--+                        
        +-- Hyriopsis_bialata **

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
5 6 4 5 113 7 2 6 
  +-------------- Ensidens             
  |                                    
--+  +----------- Hyriopsis_desowitzi **  
  |  |                                 
  +--+  +-------- Rectidens            
     |  |                              
     +--+     +-- Hyriopsis_khoratensis **
        |  +--+                        
        |  |  +-- Hyriopsis_bialata **    
        +--+                           
           |  +-- Hyriopsis_myersiana **  
           +--+                        
              +-- Hyriopsis_panhai **

Zieritz et al., 2020 [Publ 4102], fig. 4

iTtTbiriSoTroTroSiD
5 6 2 5 26 6 2 6 
  +----------- Ensidens             
  |                                 
--+  +-------- Hyriopsis_desowitzi **  
  |  |                              
  |  +-------- Rectidens            
  +--+                              
     |  +----- Hyriopsis_panhai **     
     |  |                           
     +--+  +-- Hyriopsis_khoratensis **
        |  |                        
        +--+-- Hyriopsis_bialata **    
           |                        
           +-- Hyriopsis_myersiana **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
5 11 5 5 196 6 2 12 
  +----- Hyriopsis_desowitzi **  
  |                           
  +----- Rectidens            
  |                           
--+----- Ensidens             
  |                           
  |  +-- Hyriopsis_bialata **    
  |  |                        
  |  +-- Hyriopsis_myersiana **  
  +--+                        
     +-- Hyriopsis_panhai **     
     |                        
     +-- Hyriopsis_khoratensis **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
5 11 6 5 196 6 2 12 
  +----- Hyriopsis_desowitzi **  
  |                           
  +----- Hyriopsis_myersiana **  
  |                           
  +----- Hyriopsis_panhai **     
--+                           
  |  +-- Hyriopsis_bialata **    
  +--+                        
  |  +-- Hyriopsis_khoratensis **
  |                           
  |  +-- Rectidens            
  +--+                        
     +-- Ensidens

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
5 6 2 5 120 6 2 6 
  +----------- Ensidens             
  |                                 
--+  +-------- Hyriopsis_desowitzi **  
  |  |                              
  |  +-------- Rectidens            
  +--+                              
     |  +----- Hyriopsis_panhai **     
     |  |                           
     +--+  +-- Hyriopsis_myersiana **  
        |  |                        
        +--+-- Hyriopsis_bialata **    
           |                        
           +-- Hyriopsis_khoratensis **

Konopleva et al., 2019 [Publ 4041], fig. 3

iTtTbiriSoTroTroSiD
5 12 0 5 29 7 2 15 
  +-- Ensidens_ingallsianus
  |                        
  +-- Ensidens_telus       
  |                        
  +-- Ensidens_dugasti     
  |                        
  +-- Hyriopsis_khoratensis **
  |                        
  +-- Hyriopsis_bialata **    
  |                        
  +-- Hyriopsis_myersiana **  
  |                        
--+-- Hyriopsis_desowitzi **  
  |                        
  +-- Rectidens            
  |                        
  +-- Hyriopsis_panhai **     
  |                        
  +-- Ensidens_jaculus__1  
  |                        
  +-- Ensidens_jaculus__2  
  |                        
  +-- Ensidens_spiculus    
  |                        
  +-- Ensidens_sagittarius

Muanta et al., 2019 [Publ 3921], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 4 5 5 1 8 
  +----------- Ensidens             
--+                                 
  |  +-------- Hyriopsis_desowitzi **  
  +--+                              
     |  +----- Hyriopsis_myersiana **  
     +--+                           
        |  +-- Hyriopsis_khoratensis **
        +--+                        
           +-- Hyriopsis_panhai **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
3 8 0 3 80 5 1 13 
  +-- Ensidens_telus       
  |                        
  +-- Ensidens_ingallsianus
  |                        
  +-- Ensidens_jaculus     
  |                        
  +-- Hyriopsis_desowitzi **  
--+                        
  +-- Hyriopsis_myersiana **  
  |                        
  +-- Hyriopsis_khoratensis **
  |                        
  +-- Ensidens_spiculus    
  |                        
  +-- Ensidens_sagittarius

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
3 7 2 3 77 4 2 10 
  +----- Hyriopsis_desowitzi **  
  |                           
  +----- Rectidens            
--+                           
  +----- Ensidens             
  |                           
  |  +-- Hyriopsis_khoratensis **
  +--+                        
     +-- Hyriopsis_myersiana **

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 36 2 2 -- 
  +-------- Ensidens           
--+                            
  |  +----- Rectidens          
  +--+                         
     |  +-- Hyriopsis_bialata **  
     +--+                      
        +-- Hyriopsis_desowitzi **

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
2 2 0 2 35 2 2 -- 
  +-------- Ensidens           
--+                            
  |  +----- Rectidens          
  +--+                         
     |  +-- Hyriopsis_bialata **  
     +--+                      
        +-- Hyriopsis_desowitzi **

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 30 2 2 -- 
  +-------- Ensidens           
--+                            
  |  +----- Rectidens          
  +--+                         
     |  +-- Hyriopsis_bialata **  
     +--+                      
        +-- Hyriopsis_desowitzi **

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
2 2 0 2 35 2 2 -- 
  +-------- Ensidens           
--+                            
  |  +----- Rectidens          
  +--+                         
     |  +-- Hyriopsis_bialata **  
     +--+                      
        +-- Hyriopsis_desowitzi **

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
2 13 0 2 35 2 2 19 
  +----------------------- Hyriopsis_bialata **  
  |                                           
--+                    +-- Rectidens          
  |  +-----------------+                      
  |  |                 +-- Khairuloconcha     
  +--+                                        
     |  +----------------- Ensidens           
     |  |                                     
     +--+  +-------------- Hyriopsis_desowitzi **
        |  |                                  
        +--+  +----------- Yaukthwa_elongatula
           |  |                               
           +--+  +-------- Trapezoideus       
              |  |                            
              +--+     +-- Physunio           
                 |  +--+                      
                 |  |  +-- Yaukthwa_inlenensis
                 +--+                         
                    |  +-- Yaukthwa_paiensis  
                    +--+                      
                       +-- Lens

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 44 3 2 -- 
  +-------- Ensidens           
--+                            
  |  +----- Rectidens          
  +--+                         
     |  +-- Hyriopsis_bialata **  
     +--+                      
        +-- Hyriopsis_desowitzi **

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
2 3 0 2 35 2 2 9 
  +-------- Ensidens           
--+                            
  |  +----- Hyriopsis_bialata **  
  +--+                         
     |  +-- Hyriopsis_desowitzi **
     +--+                      
        +-- Rectidens

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
2 13 0 2 35 2 2 18 
  +----- Ensidens           
  |                         
  +----- Hyriopsis_desowitzi **
  |                         
  +----- Rectidens          
  |                         
  +----- Hyriopsis_bialata **  
--+                         
  +----- Khairuloconcha     
  |                         
  |  +-- Physunio           
  |  |                      
  |  +-- Yaukthwa_paiensis  
  |  |                      
  |  +-- Yaukthwa_inlenensis
  +--+                      
     +-- Trapezoideus       
     |                      
     +-- Yaukthwa_elongatula
     |                      
     +-- Lens

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 57 2 2 -- 
  +----- Ensidens         
--+                       
  |  +-- Hyriopsis_bialata **
  +--+                    
     +-- Rectidens

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
1 1 0 1 47 1 1 -- 
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 85 1 1 -- 
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
1 1 0 1 91 1 1 -- 
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Matos et al., 2024 [Publ 4384], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 19 1 1 -- 
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 53 1 1 -- 
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
1 1 0 1 75 2 2 -- 
  +-- Ensidens         
  |                    
--+-- Hyriopsis_bialata **
  |                    
  +-- Rectidens

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
1 1 0 1 95 1 1 -- 
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
1 1 0 1 44 1 1 -- 
  +-- Hyriopsis_bialata **
--+                    
  +-- Ensidens

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
1 1 0 1 90 3 1 -- 
  +-- Hyriopsis_desowitzi **
--+                      
  +-- Ensidens

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 35 2 2 -- 
  +----- Ensidens         
--+                       
  |  +-- Hyriopsis_bialata **
  +--+                    
     +-- Rectidens

Matos et al., 2024 [Publ 4383], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 19 1 1 -- 
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 47 1 1 -- 
  +-- Ensidens         
--+                    
  +-- Hyriopsis_bialata **

Pfeiffer et al., 2018 [Publ 3827], fig. 1

iTtTbiriSoTroTroSiD
1 3 0 2 6 2 2 11 
  +-- Ensidens    
  |               
  +-- Hyriopsis__1 **
--+               
  +-- Hyriopsis__2 **
  |               
  +-- Rectidens

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
1 1 0 1 44 1 1 -- 
  +-- Hyriopsis_bialata **
--+                    
  +-- Ensidens

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 2 2 -- 
  +----- Ensidens         
--+                       
  |  +-- Hyriopsis_bialata **
  +--+                    
     +-- Rectidens

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 2 2 -- 
  +----- Ensidens         
--+                       
  |  +-- Hyriopsis_bialata **
  +--+                    
     +-- Rectidens

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 87 1 1 -- 
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 90 3 1 -- 
  +-- Hyriopsis_desowitzi **
--+                      
  +-- Ensidens

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
1 1 0 1 91 1 1 -- 
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
1 1 0 1 95 1 1 -- 
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 1 1 -- 
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
1 1 0 1 22 1 1 -- 
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Zhang et al., 2024 [Publ 4397], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 27 1 1 -- 
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 46 1 1 -- 
  +-- Ensidens         
--+                    
  +-- Hyriopsis_bialata **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
1 1 0 1 75 2 2 -- 
  +----- Ensidens         
--+                       
  |  +-- Rectidens        
  +--+                    
     +-- Hyriopsis_bialata **

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 35 2 2 -- 
  +----- Ensidens         
--+                       
  |  +-- Hyriopsis_bialata **
  +--+                    
     +-- Rectidens
 
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