Publication Graf & Cummings (2021): species taxonomic opinions

Graf, D.L. & K.S. Cummings. 2021. A ‘big data’ approach to global freshwater mussel diversity (Bivalvia: Unionoida), with an updated checklist of genera and species. Journal of Molluscan Studies 87(1). eyaa034 (36 pp.). https://doi.org/10.1093/mollus/eyaa034.

Elliptio dariensis (Lea, 1842)

Elliptio downiei (Lea, 1858)

Elliptio fisheriana (Lea, 1838)

Elliptio folliculata (Lea, 1838)

Elliptio fraterna (Lea, 1852)

Elliptio fumata (Lea, 1857)

Elliptio hopetonensis (Lea, 1838)

Elliptio icterina (Conrad, 1834)

+ Elliptio cylindraceus Frierson, 1927

+ Unio errans Lea, 1857

+ Unio hepaticus Lea, 1859

+ Unio raveneli Conrad, 1834

Elliptio jayensis (Lea, 1838)

+ Unio buckleyi Lea, 1843

Elliptio lanceolata (Lea, 1828) = Parvaspina lanceolata

Elliptio marsupiobesa Fuller, 1972

Elliptio mcmichaeli Clench & Turner, 1956

Elliptio monroensis (Lea, 1843)

Elliptio nigella (Lea, 1852)

Elliptio occulta (Lea, 1843)

Elliptio producta (Conrad, 1836)

Elliptio pullata (Lea, 1857)

Elliptio purpurella (Lea, 1857)

Elliptio roanokensis (Lea, 1838)

+ Elliptio judithae Clarke, 1986

Elliptio shepardiana (Lea, 1834)

Elliptio spinosa (Lea, 1836)

Elliptoideus sloatianus (Lea, 1840)

Elongaria orientalis (Lea, 1840) = Rectidens orientalis

Elongaria trompi (Drouet & Chaper, 1892) = Nannonaia trompi

Ensidens ingallsianus (Lea, 1852)

Ensidens pazii (Lea, 1862) = Ensidens ingallsianus

Ensidens sagittarius (Lea, 1856)

Epioblasma ahlstedti Jones & Neves, 2010

Epioblasma arcaeformis (Lea, 1831)

Epioblasma aureola Jones & Neves, 2010

Epioblasma biemarginata (Lea, 1857)

Epioblasma brevidens (Lea, 1831)

Epioblasma capsaeformis (Lea, 1834)

Epioblasma cincinnatiensis (Lea, 1840)

Epioblasma curtisii (Frierson & Utterback in Utterback, 1916)

Epioblasma flexuosa (Rafinesque, 1820)

Epioblasma florentina (Lea, 1857)

Epioblasma gubernaculum (Reeve, 1865)

Epioblasma haysiana (Lea, 1834)

Epioblasma lenior (Lea, 1842)

Epioblasma lewisii (Walker, 1910)

Epioblasma metastriata (Conrad, 1838)

Epioblasma obliquata (Rafinesque, 1820)

Epioblasma othcaloogensis (Lea, 1857)

Epioblasma penit a [sic] (Conrad, 1834) = Epioblasma penita

Epioblasma perobliqua (Conrad, 1836)

Epioblasma personata (Say, 1829)

Epioblasma propinqua (Lea, 1857)

Epioblasma rangiana (Lea, 1838)

+ Epioblasma biloba Rafinesque, 1831

Epioblasma sampsonii (Lea, 1862)

Epioblasma stewardsonii (Lea, 1852)

Epioblasma torulosa (Rafinesque, 1820)

Epioblasma triquetra (Rafinesque, 1820)

Epioblasma turgidula (Lea, 1858)

Epioblasma walkeri (Wilson & Clark, 1914)

Etheria elliptica Lamarck, 1807

+ Etheria semilunata Lamarck, 1807

Eurynia dilatata (Rafinesque, 1820)

Fossula fossiculifera (d’Orbigny, 1835)

Friersonia iridella (Pilsbry & Frierson, 1908)

Fusconaia burkei (Walker in Ortmann & Walker, 1922)

Fusconaia cerina (Conrad, 1838)

+ Quadrula rubidula Frierson, 1905

Fusconaia chunii (Lea, 1862) = Fusconaia chunii + Fusconaia askewi + Fusconaia lananensis

+ Unio askewi Marsh, 1896

+ Quadrula lananensis Frierson, 1901

Fusconaia cor (Conrad, 1834)

Fusconaia cuneolus (Lea, 1840)

Fusconaia escambia Clench & Turner, 1956

Fusconaia flava (Rafinesque, 1820)

+ Unio hebetatus Conrad, 1854

+ Unio trigonus Lea, 1831

Fusconaia masoni (Conrad, 1834)

+ Unio subplanus Conrad, 1837

+ Unio utriculus Lea, 1845

Fusconaia mitchelli (Simpson in Dall, 1896)

Fusconaia ozarkensis (Call, 1887)

Fusconaia subrotunda (Lea, 1831)

Germainaia geayi (Germain, 1911)

Gibbosula confragosa Frierson, 1928

Gibbosula crassa (Wood, 1815)

+ Unio (Quadrula) mansuyi Dautzenberg & Fischer, 1908

Gibbosula laosensis (Lea, 1863)

Gibbosula nanningensis Qian, Fang & He, 2015 = Lamprotula nanningensis

Gibbosula polysticta (Heude, 1877) = Scriptolamprotula polysticta

+ Quadrula (Lamprotula) similaris Simpson, 1900

Gibbosula rochechouartii (Heude, 1875)

Glebula rotundata (Lamarck, 1819)

Gonidea angulata (Lea, 1838)

+ Anodonta randalli Trask, 1854

Grandidieria burtoni (Woodward, 1859)

Haasica balzani (Ihering, 1893)

Haasodonta fannyae (R.I. Johnson, 1948)

Haasodonta vanheurni McMichael & Hiscock, 1958

Hamiota altilis (Conrad, 1834)

Hamiota australis (Simpson, 1900)

Hamiota perovalis (Conrad, 1834)

Hamiota subangulata (Lea, 1840)

Harmandia somboriensis Rochebrune, 1882

Hemistena lata (Rafinesque, 1820)

Hyridella australis (Lamarck, 1819)

Hyridella depressa (Lamarck, 1819)

Hyridella drapeta (Iredale, 1934)

Hyridella glenelgensis (Dennant, 1898)

Hyridella guppyi (E.A. Smith, 1885)

Hyridella misoolensis (Schepman, 1897)

Hyridella narracanensis (Cotton & Gabriel, 1932)

Hyriopsis bialata Simpson, 1900

+ Unio delphinus Gruner, 1841

Hyriopsis delaportei (Crosse & Fischer, 1876)

Hyriopsis desowitzi Brandt, 1974

Hyriopsis gracilis Haas, 1910 = Hyriopsis bialata

Hyriopsis myersiana (Lea, 1856)

Hyriopsis velthuizeni (Schepman, 1896)

Iheringella isocardioides (Lea, 1856) = Iheringella isocardiodes

Iheringella semisulcata (H. Adams, 1870)

Indochinella pugio (Benson, 1862)

Indonaia andersoniana (Nevill, 1877)

Indonaia caerulea (Lea, 1831)

Indonaia involuta (Hanley, 1856) = Assamnaia involuta

Indonaia khadakvaslaensis Ray, 1966 = Indonaia shurtleffiana

Indonaia lima (Simpson, 1900) = Pseudoscabies scobina

Indonaia nuttalliana (Lea, 1856) = Indonaia caerulea

Indonaia occata (Lea, 1860) = Indonaia gratiosa

Indonaia olivaria (Lea, 1831)

Indonaia pachysoma (Benson, 1862) = Indonaia corbis

Indonaia rectangularis (Tapparone Canefri, 1889) = Yaukthwa rectangularis

Indonaia shurtleffiana (Lea, 1856)

Indonaia subclathrata (Martens, 1899)

Indonaia substriata (Lea, 1856)

+ Nodularia (s.s.) pecten Preston, 1912

Indonaia theobaldi (Preston, 1912)

Inversidens brandtii (Kobelt, 1879)

Inversidens pantoensis (Neumayr, 1899)

Inversiunio jokohamensis (Ihering, 1893)

Inversiunio reinianus (Kobelt, 1879)

Inversiunio verrucosus Kondo, Hyun & Seung-Ho, 2007

Inversiunio yanagawensis (Kondo, 1982)

Koreosolenaia sitgyensis Lee, Kim, Lopes-Lima & Bogan in Lopes-Lima et al., 2020

Lamellidens brandti Bolotov, Konopleva & Vikhrev in Bolotov et al., 2017 = Lamellidens generosus

Lamellidens consobrinus (Lea, 1860) = Lamellidens generosus

Lamellidens corrianus (Lea, 1834)

Lamellidens ferrugineus (Annandale, 1918)

Lamellidens generosus (Gould, 1847) = Lamellidens generosus + Lamellidens ferrugineus

+ Physunio micropteroides Annandale, 1918

Lamellidens jenkinsianus (Benson, 1862)

Lamellidens lamellatus (Lea, 1838)

Lamellidens mainwaringi Preston, 1912

Lamellidens marginalis (Lamarck, 1819)

Lamellidens narainporensis Preston, 1912 = Lamellidens nongyangensis

Lamellidens phenchooganjensis Preston, 1912 = Lamellidens mainwaringi

Lamellidens rhadinaeus Annandale & Prashad, 1921 = Lamellidens candaharicus

Lamellidens savadiensis (Nevill, 1877)

Lamellidens theca (Benson, 1862) = Lamellidens corrianus

Lamellidens unioides Nesemann & Sharma in Nesemann et al., 2007

Lamproscapha ensiformis (Spix in Wagner, 1827)

Lamproscapha falsa (Simpson, 1900)

Lamprotula bazini (Heude, 1877)

Lamprotula blaisei (Dautzenberg & Fischer, 1905)

Lamprotula caveata (Heude, 1877)

Lamprotula chiai Chow, 1958

Lamprotula cornuumlunae (Heude, 1883)

Lamprotula divergens (Benson in Cantor, 1842)

Lamprotula elongata Liu, Zhang & Wang, 1980

Lamprotula kouangensis (Simpson, 1900)

Lamprotula leaii (Gray in Griffith & Pidgeon, 1833)

+ Unio gottschei Martens, 1894

Lamprotula linae He & Zhuang, 2013

Lamprotula microsticta (Heude, 1877)

Lamprotula paschalis (Heude, 1883)

Lamprotula plumbea (Dillwyn, 1817)

Lamprotula salaputium (Martens, 1902)

Lamprotula seomjinensis Kondo, Hyun & Seung-Ho, 2007)

Lamprotula triclava (Heude, 1877)

Lampsilis abrupta (Say, 1831)

Lampsilis bergmanni Inoue & Randklev in Inoue et al., 2020

Lampsilis binominata Simpson, 1900

Lampsilis bracteata (Gould, 1855)

Lampsilis brittsi Simpson, 1900

Lampsilis cardium Rafinesque, 1820

Lampsilis cariosa (Say, 1817)

Lampsilis dolabraeformis (Lea, 1838)

Lampsilis fasciola Rafinesque, 1820

Lampsilis floridensis (Lea, 1852)

Lampsilis higginsii (Lea, 1857)

Lampsilis hydiana (Lea, 1838)

Lampsilis ornata (Conrad, 1835)

Lampsilis ovata (Say, 1817)

Lampsilis powellii (Lea, 1852)

Lampsilis radiata (Gmelin, 1791)

+ Lampsilis (s.s.) fullerkati R.I. Johnson, 1984

Lampsilis rafinesqueana Frierson, 1927

Lampsilis reeveiana (Lea, 1852)

Lampsilis satura (Lea, 1852)

Lampsilis sietmani Keogh & Simons, 2019

Lampsilis siliquoidea (Barnes, 1823)

Lampsilis splendida (Lea, 1838)

Lampsilis straminea (Conrad, 1834)

Lampsilis streckeri Frierson, 1927

Lampsilis teres (Rafinesque, 1820)

Lampsilis virescens (Lea, 1858)

Lanceolaria acrorrhyncha (Martens, 1894)

Lanceolaria bilirata (Martens, 1902)

Lanceolaria bogani Thach, 2016 = Lanceolaria fruhstorferi

Lanceolaria cylindrica (Simpson, 1900)

Lanceolaria eucylindrica Lin, 1962

Lanceolaria fruhstorferi (Dautzenberg, 1900)

Lanceolaria gladiola (Heude, 1877)

Lanceolaria grayii (Griffith & Pidgeon, 1833)

+ Unio grayanus Lea, 1834

Lanceolaria kihirai Kondo & Hattori, 2019

Lanceolaria laevis (Martens, 1902)

Lanceolaria lanceolata (Lea, 1856)

+ Unio contortus var. muticus Heude, 1877

+ Arconaia huaihensis Huang & Li, 2003

Lanceolaria oxyrhyncha (Martens, 1861)

Lanceolaria triformis (Heude, 1877)

Lasmigona alabamensis Clarke, 1985 = Pterosyna alabamensis

Lasmigona complanata (Barnes, 1823) = Pterosyna complanata

Lasmigona compressa (Lea, 1829) = Platynaias compressa

Lasmigona costata (Rafinesque, 1820)

Lasmigona decorata (Lea, 1852) = Platynaias decorata

Lasmigona etowaensis (Conrad, 1849) = Alasminota etowaensis

Lasmigona holstonia (Lea, 1838) = Alasminota holstonia

Lasmigona subviridis (Conrad, 1835) = Platynaias subviridis

Leaunio lienosus (Conrad, 1834)

Leaunio ortmanni (Walker, 1925)

Leaunio pataecus Watters, 2018

Leaunio umbrans (Lea, 1857)

Leaunio vanuxemensis (Lea, 1838)

Leguminaia saulcyi (Bourguignat, 1852)

Leguminaia wheatleyi (Lea, 1862)

+ Monocondyloea mardinensis Lea, 1865

Leila blainvilliana (Lea, 1834)

Leila esula (d’Orbigny, 1835)

Lemiox rimosus (Rafinesque, 1831)

Leoparreysia bhamoensis (Theobald, 1873) = Leoparreysia tavoyensis

+ Unio mandelayensis Theobald, 1873

Leoparreysia burmana (Blanford, 1869)

Leoparreysia canefrii Vikhrev, Bolotov & Kondakov in Bolotov et al., 2017

Leoparreysia choprae (Prashad, 1930) = Leoparreysia tavoyensis

Leoparreysia feae (Tapparone Canefri, 1889) = Leoparreysia tavoyensis

Leoparreysia feddeni (Theobald, 1873) = Parreysia corrugata

Leoparreysia houngdarauica (Tapparone Canefri, 1889) = Leoparreysia tavoyensis

+ Unio houngdaranicus ‘Tapparone Canefri’ of Authors

Leoparreysia olivacea (Prashad, 1930)

Leoparreysia perconvexa (Preston, 1912)

Leoparreysia tavoyensis (Gould, 1843)

Leoparreysia vulcanus (Hanley, 1876)

Leoparreysia whitteni Bolotov, Konopleva, Vikrev, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Gofarov, Tomilova & Kondakov, 2019

Lepidodesma aligera (Heude, 1877)

Lepidodesma languilati (Heude, 1874)

Leptodea fragilis (Rafinesque, 1820) = Potamilus fragilis

+ Medionidus mcglameriae van der Schalie, 1939

Leptodea leptodon (Rafinesque, 1820) = Potamilus leptodon

Leptodea ochracea (Say, 1817) = Atlanticoncha ochracea

Ligumia recta (Lamarck, 1819)

Lortiella froggatti Iredale, 1934

Lortiella opertanea Ponder & Bayer, 2004

Lortiella rugata (Sowerby, 1868)

Margaritifera dahurica (Middendorff, 1850)

Margaritifera falcata (Gould, 1850)

Margaritifera hembeli (Conrad, 1838)

Margaritifera laevis (Haas, 1910)

Margaritifera margaritifera (Linnaeus, 1758)

Margaritifera marrianae R.I. Johnson, 1983

Margaritifera middendorffi (Rosén, 1926)

+ Margaritifera togakushiensis Kondo & Kobayashi, 2005

Martensnaias rubicunda (Martens, 1900) = Psoronaias rubicunda

Medionidus acutissimus (Lea, 1831)

Medionidus conradicus (Lea, 1834)

Medionidus parvulus (Lea, 1860)

Medionidus penicillatus (Lea, 1857)

Medionidus simpsonianus Walker, 1905

Medionidus walkeri (B.H. Wright, 1897)

Megalonaias nervosa (Rafinesque, 1820)

+ Unio boykinianus Lea, 1840

+ Unio heros Say, 1829

Megalonaias nickliniana (Lea, 1834)

Microcondylaea bonellii (Férussac, 1827)

Microdontia anodontaeformis (Tapparone Canefri, 1883)

Micronaias arata (Lea, 1843)

Micronaias fallaciosa (Simpson, 1914)

Micronaias granadensis (Lea, 1868)

Middendorffinaia mongolica (Middendorff, 1851)

Modellnaia siamensis Brandt, 1974

Moncetia anceyi Bourguignat, 1886

Monocondylaea corrientesensis (d’Orbigny, 1835)

Monocondylaea costulata (Moricand, 1858)

Monocondylaea franciscana (Moricand, 1837)

Monocondylaea guarayana (d’Orbigny, 1835)

Monocondylaea jaspidea (Hupé, 1857)

Monocondylaea minuana (d’Orbigny, 1835)

Monocondylaea paraguayana (d’Orbigny, 1835)

Monocondylaea parchappii (d’Orbigny, 1835)

Monodontina aeneola (Drouet & Chaper, 1892) = Cosmopseudodon aeneolus

Monodontina cambodjensis (Petit, 1865) = Pseudodon vondembuschianus

Monodontina laosica Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuang-pak, Tumpeesuwan & Kondakov, 2020 = Pseudodon mekongi

Monodontina lenyanensis Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuang-pak, Tumpeesuwan & Kondakov, 2020 = Pseudodon lenyanensis

Monodontina mekongi Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuang-pak, Tumpeesuwan & Kondakov, 2020 = Pseudodon mekongi

Monodontina vondembuschiana (Lea, 1840) = Pseudodon vondembuschianus

+ Microcondylaea hageni Strubell, 1897

Monodontina walpolei (Hanley, 1871) = Pseudodon walpolei

+ Pseudodon crassus Drouët in Drouët & Chaper, 1892

Mutela alata (Lea, 1864)

Mutela alluaudi Germain, 1909

Mutela bourguignati Bourguignat, 1886

Mutela dubia (Gmelin, 1791)

Mutela franci Daget, 1964

Mutela hargeri E.A. Smith, 1908

Mutela joubini (Germain, 1904)

Mutela langi Pilsbry & Bequaert, 1927

Mutela legumen (Rochebrune, 1886)

Mutela mabilli (Rochebrune, 1886)

Mutela rostrata (Rang, 1835)

Mutela soleniformis Bourguignat, 1886

Mutela wistarmorrisi Graf & Cummings, 2006

Mutela zambesiensis Mandahl-Barth, 1988

Mycetopoda legumen (Martens, 1888)

Mycetopoda pittieri Marshall, 1927

Mycetopoda siliquosa (Spix in Wagner, 1827)

Mycetopoda soleniformis d’Orbigny, 1835

Mycetopodella falcata (Higgins, 1868)

Nephritica haricotti (Frierson, 1927)

Nephritica poeyana (Lea, 1857)

Nephronaias aeruginosa (Morelet, 1849)

Nephronaias aztecorum (Philippi, 1847)

Nephronaias calamitarum (Morelet, 1849)