species Pyganodon grandis (Say, 1829): synonymy

Anodonta grandis Say, 1829

Say, 1829. New Harm. Dissem. 2(22): 341, sp. 3.
Say, 1858. [in] Binney, Compl. Writings T. Say: 139.
Type(s): TYPE LOST XXX_UNKNOWN, Fox River of the Wabash.

Anodonta grandis — Simpson (1900), Ortmann (1909), Ortmann (1909), Ortmann (1909), Ortmann (1910), ... Rosenberg et al. (1994), Howells et al. (1996), Rosenberg et al. (1997), Kongim et al. (2015), Soliman et al. (2016)
Unio (Anodonta) grandis — Call (1900)
Anodonta (Pyganodon) grandis — Frierson (1927), Kat (1983), R.I. Johnson (1999)
Anodonta (s.s.) grandis — Haas (1930)
Pyganodon grandis — Modell (1945), Hoeh (1990), Cummings & Mayer (1992), Williams et al. (1993), Vidrine (1993), ... Chen et al. (2025), FMCS (2025), Cummings et al. (2025), Hou et al. (2025), Wang et al. (2025)
Anodonta (Pyganodon) grandis grandis — Haas (1969)
Anodonta grandis grandis — Clarke (1973), Burch (1975), R.I. Johnson (1980), Mackie et al. (1980), Oesch (1984), Liu et al. (1996), ITIS (2012)
Pyganodon grandis grandis — Sayenko et al. (2005), Watters et al. (2009)
P. grandis — Sheng et al. (2014), Marshall et al. (2018)

Anodonta lugubris Say, 1829

Say, 1829. New Harm. Dissem. 2(22): 340, sp. 2.
Say, 1858. [in] Binney, Compl. Writings T. Say: 138.

Anodonta lugubris — Simpson (1900), Simpson (1914)
Anodonta (Pyganodon) grandis-lugubris — Frierson (1927)
Pyganodon lugubris — Modell (1945)
Anodonta (Pyganodon) grandis lugubris — Haas (1969)
Anodonta grandis — Howells et al. (1996)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008)
Anodonta (Pyganodon) grandis — R.I. Johnson (1999)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta inflata Rafinesque, 1831

Rafinesque, 1831. Continuation: 6.
Rafinesque, 1864. [in] Binney & Tryon, reprint: 81.

Anodonta inflata — Simpson (1900), Simpson (1914), Burch (1975)
Anodonta (Pyganodon) grandis — Frierson (1927)
Anodonta (Pyganodon) grandis grandis — Haas (1969)

Anodonta inflata var. fuscata Rafinesque, 1831

Rafinesque, 1831. Continuation: 6.
Rafinesque, 1864. [in] Binney & Tryon, reprint: 82.
(intro'd as intra-sp. of Anodonta inflata Rafinesque, 1831)

Anodonta inflata var. viridis Rafinesque, 1831

Rafinesque, 1831. Continuation: 6.
Rafinesque, 1864. [in] Binney & Tryon, reprint: 82.
(homonym of Anodonta (Lastena) ohiensis var. viridis Rafinesque, 1820; intro'd as intra-sp. of Anodonta inflata Rafinesque, 1831)

Anodonta inflata var. zonalis Rafinesque, 1831

Rafinesque, 1831. Continuation: 6.
Rafinesque, 1864. [in] Binney & Tryon, reprint: 82.
(intro'd as intra-sp. of Anodonta inflata Rafinesque, 1831)

Anodonta corpulenta Cooper, 1834

Cooper, 1834. Narr. Exped. Upper Miss., Appendix: 154.
Type(s): UNKNOWN USNM_86677, [no locality available - North America].

Anodonta corpulenta — Simpson (1900), Simpson (1914), Utterback (1915), F.C. Baker (1928), Henderson (1935), R.I. Johnson (1980)
Anodonta (Pyganodon) grandis — Frierson (1927)
Anodonta (Pyganodon) grandis grandis — Haas (1969)
Anodonta grandis corpulenta — Burch (1975), Gordon et al. (1979), Oesch (1984), ITIS (2012)
Anodonta (Pyganodon) corpulenta — Heard (1975)
Anodonta grandis — Mathiak (1979), Howells et al. (1996)
Pyganodon grandis — Vidrine (1993), Parmalee & Bogan (1998), Williams et al. (2008), IUCN (2012)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta declivis Conrad, 1834

Conrad, 1834. Amer. J. Sci. 25(2): 341, pl. 1, fig. 11.
Conrad, 1853. Proc. Acad. Nat. Sci. 6: 263.

Anodonta grandis var. gigantea — Simpson (1900), Simpson (1914)
Anodonta (Pyganodon) grandis-lugubris — Frierson (1927)
Anodonta grandis plana — F.C. Baker (1928)
Anodonta (Pyganodon) grandis lugubris — Haas (1969)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta plana Lea, 1834

Lea, 1834. Trans. Amer. Phil. Soc. 5 [O. 1]: 48 [as ‘palna’], pl. 7, fig. 18.
(correct name for Anodonta palna Lea, 1834)
Type(s): HOLOTYPE USNM_86626, Bear Grass Creek, near Louisville, Ky.

Anodonta grandis var. gigantea — Simpson (1900), Simpson (1914)
Unio (Anodonta) grandis — Call (1900)
Anodonta (Pyganodon) grandis-lugubris — Frierson (1927)
Anodonta grandis plana — F.C. Baker (1928)
Anodonta (Pyganodon) grandis lugubris — Haas (1969)
Anodonta grandis grandis — Burch (1975), R.I. Johnson (1980)
Pyganodon grandis — Strayer & Jirka (1997), Parmalee & Bogan (1998), Williams et al. (2008)

Anodonta stewartiana Lea, 1834

Lea, 1834. Trans. Amer. Phil. Soc. 5 [O. 1]: 47 [159], pl. 6, fig. 17.
Type(s): HOLOTYPE USNM_86664, Bayou Teche, Louisiana.

Anodonta stewartiana — Simpson (1900), Simpson (1914)
Anodonta (Pyganodon) grandis — Frierson (1927)
Anodonta (Pyganodon) grandis grandis — Haas (1969)
Anodonta grandis grandis — R.I. Johnson (1980)
Pyganodon grandis — Vidrine (1993), Parmalee & Bogan (1998), Williams et al. (2008)
Anodonta grandis — Howells et al. (1996)
Pyganodon grandis grandis — Watters et al. (2009)

Symphynota benedictensis Lea, 1834

Lea, 1834. Trans. Amer. Phil. Soc. 5 [O. 1]: 104, pl. 16, fig. 48.
(renamed Margaron (Anodonta) benedictii Lea, 1870)
Type(s): HOLOTYPE USNM_86381, Lake Champlain, Vermont.

Anodonta grandis var. benedictensis — Simpson (1900), Ortmann (1909), Simpson (1914)
Anodonta (Pyganodon) grandis-benedictensis — Frierson (1927)
Anodonta (Pyganodon) grandis benedictensis — Haas (1969)
Pyganodon grandis — Strayer & Jirka (1997), Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta decora Lea, 1838

Lea, 1836. Synopsis Fam. Naiades: 52. [nomen nudum]
Lea, 1838. Trans. Amer. Phil. Soc. 6 [O. 2]: 64, pl. 20, fig. 63.
Type(s): HOLOTYPE USNM_86633, canal, near Cincinnati, Ohio.

Anodonta grandis var. gigantea — Simpson (1900), Simpson (1914)
Unio (Anodonta) grandis — Call (1900)
Anodonta (Pyganodon) grandis-lugubris — Frierson (1927)
Anodonta (Pyganodon) grandis lugubris — Haas (1969)
Pyganodon grandis — Strayer & Jirka (1997), Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta gigantea Lea, 1838

Lea, 1836. Synopsis Fam. Naiades: 52. [nomen nudum]
Lea, 1838. Trans. Amer. Phil. Soc. 6 [O. 2]: 1 [1], pl. 1, fig. 1.
Type(s): HOLOTYPE (MONOTYPY) USNM_86659, Port Gibson, Mississippi.

Anodonta grandis var. gigantea — Simpson (1900), Ortmann (1909), Simpson (1914)
Unio (Anodonta) grandis — Call (1900)
Anodonta grandis gigantea — Ortmann (1918)
Anodonta grandis — Ortmann (1925), Howells et al. (1996)
Anodonta (Pyganodon) grandis — Frierson (1927)
Anodonta gigantea — F.C. Baker (1928)
Anodonta (Pyganodon) grandis grandis — Haas (1969)
Anodonta grandis grandis — Burch (1975), R.I. Johnson (1980)
Pyganodon grandis — Vidrine (1993), Strayer & Jirka (1997), Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta ovata Lea, 1838

Lea, 1836. Synopsis Fam. Naiades: 52. [nomen nudum]
Lea, 1838. Trans. Amer. Phil. Soc. 6 [O. 2]: 2, pl. 2, fig. 2.
(homonym of Unio ovata Stark, 1828)
Type(s): HOLOTYPE USNM_86615, Marietta, Ohio.

Anodonta grandis — Simpson (1900), Simpson (1914)
Anodonta (Pyganodon) grandis-lugubris — Frierson (1927)
Anodonta grandis plana — F.C. Baker (1928)
Anodonta (Pyganodon) grandis lugubris — Haas (1969)
Anodonta grandis grandis — Burch (1975)
Pyganodon grandis — Strayer & Jirka (1997), Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta pepinianus Lea, 1838

Lea, 1838. Trans. Amer. Phil. Soc. 6 [O. 2]: 96, pl. 16, fig. 51.
Type(s): HOLOTYPE USNM_86440, Lake Pepin, Ohio.

Anodonta pepiniana — Simpson (1900), Simpson (1914)
Anodonta (Pyganodon) grandis-pepiniana — Frierson (1927)
Anodonta (Pyganodon) grandis pepiniana — Haas (1969)
Pyganodon grandis — Strayer & Jirka (1997), Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta salmonia Lea, 1838

Lea, 1836. Synopsis Fam. Naiades: 51. [nomen nudum]
Lea, 1838. Trans. Amer. Phil. Soc. 6 [O. 2]: 45, pl. 14, fig. 41.
Type(s): HOLOTYPE USNM_86540, Poland, Ohio.

Anodonta grandis — Simpson (1900), Simpson (1914)
Unio (Anodonta) salmonia — Call (1900)
Anodonta grandis var. salmonia — Ortmann (1909)
Anodonta (Pyganodon) grandis-lugubris — Frierson (1927)
Anodonta grandis plana — F.C. Baker (1928)
Anodonta (Pyganodon) grandis lugubris — Haas (1969)
Anodonta grandis grandis — Burch (1975)
Pyganodon grandis — Strayer & Jirka (1997), Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta footiana Lea, 1840

Lea, 1840. Proc. Amer. Phil. Soc. 1: 289.
Lea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 225 [63], pl. 20, fig. 44.
Type(s): HOLOTYPE USNM_86468, Fort Winnebago.

Anodonta grandis var. footiana — Simpson (1900), Simpson (1914)
Unio (Anodonta) footiana — Call (1900)
Anodonta grandis footiana — Ortmann (1909), F.C. Baker (1928)
Anodonta (Pyganodon) grandis-lugubris — Frierson (1927)
Pyganodon grandis footianus — Modell (1945)
Anodonta (Pyganodon) grandis lugubris — Haas (1969)
Anodonta grandis grandis — Burch (1975), R.I. Johnson (1980)
Anodonta grandis f. footiana — Heard (1975)
Pyganodon grandis — Strayer & Jirka (1997), Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta harpethensis Lea, 1840

Lea, 1840. Proc. Amer. Phil. Soc. 1: 289.
Lea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 224 [62], pl. 19, fig. 42.
Type(s): HOLOTYPE USNM_86657, Harpeth R., Tenn.

Anodonta grandis var. gigantea — Simpson (1900), Simpson (1914)
Anodonta (Pyganodon) grandis-lugubris — Frierson (1927)
Anodonta (Pyganodon) grandis lugubris — Haas (1969)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta maryattana Lea, 1840

Lea, 1840. Proc. Amer. Phil. Soc. 1: 289 [as ‘maryattiana’].
Lea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 226, pl. 20, fig. 45.
(correct name for Anodonta maryattiana Lea, 1840)

Anodonta grandis var. footiana — Simpson (1900), Simpson (1914)
Anodonta (Pyganodon) grandis-lugubris — Frierson (1927)
Anodonta grandis footiana — F.C. Baker (1928)
Anodonta (Pyganodon) grandis lugubris — Haas (1969)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008), Bieler (2021)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta maryattiana Lea, 1840

Lea, 1840. Proc. Amer. Phil. Soc. 1: 289.
(incorrect original spelling of Anodonta maryattana Lea, 1840)

Anodonta grandis var. footiana — Simpson (1900), Simpson (1914)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008), Bieler (2021)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta nilssonii Küster, 1842

Küster, 1842. Conch. Cab. Anodonta 9: 61, pl. 17, figs. 3-4. [in part]
(misidentified reference to Anodonta nilssonii Küster, 1842)

Anodonta grandis var. footiana — Simpson (1900), Simpson (1914)
Anodonta grandis grandis — Burch (1975)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta linnaeana Lea, 1852

Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 289 [45], pl. 27, fig. 51 [as ‘linnæana’].
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252 [as ‘linnæana’]. [nomen nudum]
(correct name for Anodonta linnaena Lea, 1852)
Type(s): FIGURED SYNTYPE USNM_86702, Lake Concordia, Louisiana.

Anodonta linneana [sic] — Simpson (1900), Simpson (1914)
Anodonta (Pyganodon) grandis — Frierson (1927)
Anodonta (Pyganodon) grandis grandis — Haas (1969)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008), Bieler (2021)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta linnaena Lea, 1852

Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 289 [45] [as ‘linnæna’].
(incorrect original spelling of Anodonta linnaeana Lea, 1852)

Pyganodon grandis — Bieler (2021)

Anodonta opaca Lea, 1852

Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 285, pl. 25, fig. 46.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
Type(s): HOLOTYPE USNM_128835, New Orleans.

Anodonta opaca — Simpson (1900), Simpson (1914)
Anodonta grandis — Utterback (1915), Howells et al. (1996)
Anodonta (Pyganodon) grandis — Frierson (1927)
Anodonta (Pyganodon) grandis grandis — Haas (1969)
Anodonta grandis grandis — R.I. Johnson (1980), Oesch (1984)
Pyganodon grandis — Vidrine (1993), Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta virens Lea, 1852

Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 290 [46], pl. 28, fgi. 53.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
Type(s): HOLOTYPE USNM_86666, Red R., Alexandria, Louisiana.

Anodonta stewartiana — Simpson (1900), Simpson (1914)
Anodonta (Pyganodon) grandis — Frierson (1927)
Anodonta (Pyganodon) grandis grandis — Haas (1969)
Anodonta grandis grandis — R.I. Johnson (1980)
Anodonta grandis — Howells et al. (1996)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta linneana ‘Lea’ of Authors

‘Lea’ of Authors.
Conrad, 1853. Proc. Acad. Nat. Sci. 6: 264.
Hanley, 1856. Illustr. Descr. Cat. Rec. Bivalve Shells: 387.
Sowerby, 1870. Conch. Icon. 17: pl. 35, sp. 144.
Simpson, 1900. Proc. U.S. Nat. Mus. 22: 646.
Simpson, 1914. Descr. Cat. Naiades: 431.
Frierson, 1927. Check List N. Amer. Naiades: 14.
Haas, 1969. Das Tierreich 88: 364.
Parmalee & Bogan, 1998. The Freshwater Mussels of Tennessee: 206.
Watters et al., 2009. Freshw. Mussels OH: 260.
(error for Anodonta linnaeana Lea, 1852)

Pyganodon grandis — Bieler (2021)

Anodonta marryattana ‘Lea’ of Authors

‘Lea’ of Authors.
Conrad, 1853. Proc. Acad. Nat. Sci. 6: 264.
Sowerby, 1870. Conch. Icon. 17: pl. 28, sp. 111.
Simpson, 1900. Proc. U.S. Nat. Mus. 22: 642.
Simpson, 1914. Descr. Cat. Naiades: 422.
F.C. Baker, 1928. Bull. Wisconsin Geol. Nat. Hist. Survey 70(2): 157.
Haas, 1969. Das Tierreich 88: 365.
(error for Anodonta maryattana Lea, 1840)

Anodonta grandis var. footiana — Simpson (1900), Simpson (1914)
Anodonta grandis grandis — Burch (1975)
Pyganodon grandis — Parmalee & Bogan (1998)

Anodonta lewisii Lea, 1857

Lea, 1857. Proc. Acad. Nat. Sci. 9: 84.
Lea, 1860. Jour. Acad. Nat. Sci. 4 [O. 8]: 362 [44], pl. 62, fig. 187.
Type(s): HOLOTYPE USNM_86595, Mohawk, N.Y.

Anodonta grandis — Simpson (1900), Simpson (1914)
Anodonta (Pyganodon) grandis-lugubris — Frierson (1927)
Anodonta (Pyganodon) grandis lugubris — Haas (1969)
Anodonta grandis grandis — Burch (1975)
Pyganodon grandis — Strayer & Jirka (1997), Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta danielsii Lea, 1858

Lea, 1858. Proc. Acad. Nat. Sci. 10: 139.
Lea, 1860. Jour. Acad. Nat. Sci. 4 [O. 8]: 365 [47], pl. 63, fig. 190.
Type(s): HOLOTYPE USNM_86426, Topeka, Kas.

Anodonta danielsii — Simpson (1900), Simpson (1914), Utterback (1915)
Anodonta (Pyganodon) grandis-lugubris — Frierson (1927)
Anodonta grandis — Murray & Leonard (1962), Howells et al. (1996)
Anodonta (Pyganodon) grandis lugubris — Haas (1969)
Anodonta grandis grandis — R.I. Johnson (1980), Oesch (1984)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta gesnerii Lea, 1858

Lea, 1858. Proc. Acad. Nat. Sci. 10: 139.
Lea, 1859. Jour. Acad. Nat. Sci. 4 [O. 7]: 231 [49], pl. 31, fig. 109.
Type(s): TYPE USNM_86427, Uphaupee Cr., ..., Ala.

Anodonta hallenbeckii — Simpson (1900), Simpson (1914)
Anodonta (Pyganodon) cataracta-hallenbeckii — Frierson (1927)
Anodonta hallenbecki — Clench & Turner (1956)
Anodonta (Pyganodon) cataracta hallenbeckii — Haas (1969)
Anodonta (Pyganodon) cataracta cataracta — R.I. Johnson (1970)
Anodonta cataracta — Burch (1975)
Pyganodon grandis — Brim Box & Williams (2000), Williams et al. (2008)

Anodonta texasensis Lea, 1859

Lea, 1859. Proc. Acad. Nat. Sci. 11: 113.
Lea, 1860. Jour. Acad. Nat. Sci. 4 [O. 8]: 366 [48], pl. 63, fig. 191.
Type(s): HOLOTYPE (MONOTYPY) USNM_86680, Texas.

Anodonta texasensis — Simpson (1900), Simpson (1914)
Anodonta danielsii — Utterback (1915)
Anodonta (Pyganodon) grandis-lugubris — Frierson (1927)
Anodonta (Pyganodon) grandis lugubris — Haas (1969)
Anodonta grandis grandis — R.I. Johnson (1980)
Anodonta grandis — Howells et al. (1996)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008), Cummings et al. (2025)
Toxolasma texasense — Lee (2006)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta kennicottii Lea, 1861

Lea, 1861. Proc. Acad. Nat. Sci. 13: 56.
Lea, 1862. Jour. Acad. Nat. Sci. 5 [O. 9]: 214 [36], pl. 33, fig. 283.
Type(s): HOLOTYPE USNM_86442, N End of Lake Winnipeg.

Anodonta kennicottii — Simpson (1900), Simpson (1914), F.C. Baker (1928)
Anodonta (Pyganodon) grandis-pepiniana — Frierson (1927)
Anodonta (Pyganodon) grandis pepiniana — Haas (1969)
Anodonta grandis simpsoniana — Clarke (1973), Burch (1975)
Pyganodon grandis — Strayer & Jirka (1997), Parmalee & Bogan (1998)

Anodonta simpsoniana Lea, 1861

Lea, 1861. Proc. Acad. Nat. Sci. 13: 56.
Lea, 1862. Jour. Acad. Nat. Sci. 5 [O. 9]: 212 [34], pl. 32, fig. 281.
Type(s): HOLOTYPE USNM_86434, Fort Rae, Gt. Slave Lake.

Anodonta kennicottii — Simpson (1900), Simpson (1914), F.C. Baker (1928)
Anodonta (Anodontoides) ferussaciana — Frierson (1927)
Anodontoides ferussacianus ferussacianus — Haas (1969)
Anodonta grandis simpsoniana — Clarke (1973), Burch (1975), R.I. Johnson (1980), Mackie et al. (1980), ITIS (2012)
Pyganodon grandis — Strayer & Jirka (1997), Parmalee & Bogan (1998), Williams et al. (2008), IUCN (2012)
Pyganodon simpsonaia [sic] — Cyr et al. (2007)
Pyganodon grandis grandis — Watters et al. (2009)
Pyganodon simpsoniana — Doucet-Beaupré et al. (2012)

Anodonta leonensis Lea, 1862

Lea, 1862. Proc. Acad. Nat. Sci. 14: 169.
Lea, 1866. Jour. Acad. Nat. Sci. 6 [O. 11]: 25 [29], pl. 9, fig. 24.
Type(s): LECTOTYPE [AS HOLOTYPE] USNM_87806 (by R.I. Johnson, 1974: 83), Leon, Texas.

Anodonta grandis var. leonensis — Simpson (1900), Simpson (1914)
Anodonta (Pyganodon) grandis-lugubris — Frierson (1927)
Anodonta (Pyganodon) grandis lugubris — Haas (1969)
Anodonta grandis grandis — Burch (1975), R.I. Johnson (1980)
Anodonta grandis — Howells et al. (1996)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008), Cummings et al. (2025)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta bealei Lea, 1863

Lea, 1863. Proc. Acad. Nat. Sci. 15: 194.
Lea, 1866. Jour. Acad. Nat. Sci. 6 [O. 11]: 26 [30], pl. 9, fig. 25.
Type(s): LECTOTYPE [AS HOLOTYPE] USNM_86690 (by R.I. Johnson, 1974: 19), Leon Co., Texas.

Anodonta bealei — Simpson (1900), Simpson (1914), Utterback (1915)
Anodonta (Pyganodon) grandis-bealei — Frierson (1927)
Anodonta (Pyganodon) grandis bealei — Haas (1969)
Anodonta grandis grandis — R.I. Johnson (1980)
Anodonta grandis — Howells et al. (1996)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008), Cummings et al. (2025)
Anodonta (Pyganodon) grandis — R.I. Johnson (1999)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta dallasiana Lea, 1863

Lea, 1863. Proc. Acad. Nat. Sci. 15: 190.
Lea, 1866. Jour. Acad. Nat. Sci. 6 [O. 11]: 29 [33], pl. 11, fig. 28.
Type(s): HOLOTYPE USNM_86443, Mo. of Saskatchewan R., L. Winnipeg.

Anodonta kennicottii — Simpson (1900)
Anodonta pepiniana — Simpson (1914)
Anodonta (Pyganodon) grandis-pepiniana — Frierson (1927)
Anodonta (Pyganodon) grandis pepiniana — Haas (1969)
Anodonta grandis grandis — Clarke (1973), Burch (1975)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodon imbricata J.G. Anthony, 1865

J.G. Anthony, 1865. Amer. J. Conch. 1(2): 159, pl. 14, fig. 1.
Type(s): HOLOTYPE MCZ_161861, Camp Lake, Michigan.

Anodonta grandis var. footiana — Simpson (1900), Simpson (1914)
Anodonta (Pyganodon) grandis-lugubris — Frierson (1927)
Anodonta (Pyganodon) grandis lugubris — Haas (1969)
Anodonta grandis grandis — Burch (1975)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodon opalina J.G. Anthony, 1865

J.G. Anthony, 1865. Amer. J. Conch. 1(2): 159, pl. 14, fig. 2.
Type(s): HOLOTYPE MCZ_150633, Shears Lake, Michigan.

Anodonta grandis var. gigantea — Simpson (1900), Simpson (1914)
Anodonta (Pyganodon) grandis-pepiniana — Frierson (1927)
Anodonta grandis — F.C. Baker (1928)
Anodonta (Pyganodon) grandis pepiniana — Haas (1969)
Anodonta grandis grandis — Burch (1975)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodon subangulata J.G. Anthony, 1865

J.G. Anthony, 1865. Amer. J. Conch. 1(2): 158, pl. 13, fig. 1.
Type(s): HOLOTYPE MCZ_150645, Michigan.

Anodonta grandis var. gigantea — Simpson (1900), Simpson (1914)
Anodonta (Pyganodon) grandis-lugubris — Frierson (1927)
Anodonta (Pyganodon) grandis lugubris — Haas (1969)
Anodonta grandis grandis — Burch (1975)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodon subinflata J.G. Anthony, 1865

J.G. Anthony, 1865. Amer. J. Conch. 1(2): 160, pl. 15, fig. 1 [as ‘sub-inflata’].
(homonym of Anodonta variabilis subinflata Moquin-Tandon, 1855)
Type(s): HOLOTYPE MCZ_161863, [Shears Lake, Kent Co.], Michigan.

Anodonta grandis var. footiana — Simpson (1900), Simpson (1914)
Anodonta (Pyganodon) grandis-lugubris — Frierson (1927)
Anodonta (Pyganodon) grandis lugubris — Haas (1969)
Anodonta grandis grandis — Burch (1975)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodon inornata J.G. Anthony, 1866

J.G. Anthony, 1866. Amer. J. Conch. 2(2): 145, pl. 7, fig. 1.
Type(s): HOLOTYPE MCZ_161867, Slawsons Lake, Michigan.

Anodonta grandis var. gigantea — Simpson (1900), Simpson (1914)
Anodonta grandis grandis — Burch (1975)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodon mcnielii J.G. Anthony, 1866

J.G. Anthony, 1866. Amer. J. Conch. 2(2): 144, pl. 6, fig. 1.
Type(s): HOLOTYPE MCZ_150644, Sand Lake, Michigan.

Anodonta grandis var. gigantea — Simpson (1900), Simpson (1914)
Anodonta (Pyganodon) grandis-lugubris — Frierson (1927)
Anodonta (Pyganodon) grandis lugubris — Haas (1969)
Anodonta grandis grandis — Burch (1975)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodon subgibbosa J.G. Anthony, 1866

J.G. Anthony, 1866. Amer. J. Conch. 2(2): 144, pl. 6, fig. 2.

Anodonta grandis var. gigantea — Simpson (1900)
Anodonta subgibbosa — Simpson (1914)
Anodonta (Pyganodon) grandis-lugubris — Frierson (1927)
Anodonta (Pyganodon) grandis lugubris — Haas (1969)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta salmonea ‘Lea’ of Authors

‘Lea’ of Authors.
Sowerby, 1867. Conch. Icon. 17: pl. 19, sp. 74.
Frierson, 1927. Check List N. Amer. Naiades: 15.
F.C. Baker, 1928. Bull. Wisconsin Geol. Nat. Hist. Survey 70(2): 155.
(error for Anodonta salmonia Lea, 1838)

Anodonta grandis — Simpson (1900)
Pyganodon grandis — Parmalee & Bogan (1998)

Anodonta corpulentoides Lea, 1868

Lea, 1868. Proc. Acad. Nat. Sci. 20: 164.

Anodonta corpulenta — Henderson (1935)

Anodonta grandioides Lea, 1868

Lea, 1868. Proc. Acad. Nat. Sci. 20: 164.

Anodonta grandis — Henderson (1935)

Margaron (Anodonta) benedictii Lea, 1870

Lea, 1870. Synopsis Fam. Naiades: 75.
(new name for Symphynota benedictensis Lea, 1834)

Anodonta grandis var. benedictensis — Simpson (1900), Simpson (1914)
Anodonta (Pyganodon) grandis-benedictensis — Frierson (1927)
Anodonta (Pyganodon) grandis benedictensis — Haas (1969)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Margaron (Anodonta) marryatana ‘Lea’ of Authors

‘Lea’ of Authors.
Lea, 1870. Synopsis Fam. Naiades: 82.
B.H. Wright, 1888. Check List N. Amer. Unionidae etc.: [7] [as ‘Anodonta marryatana’].
Frierson, 1927. Check List N. Amer. Naiades: 15.
(error for Anodonta maryattana Lea, 1840)

Pyganodon grandis — Bieler (2021)

Anodonta houghtonensis Currier in DeCamp, 1881

Currier, 1867. Amer. J. Conch. 3(2): 14, pl. 1, fig. 2.
Currier in DeCamp, 1881. Misc. Publ. Kent Sci. Inst. (5): 14, pl. 1, fig. 2.

Anodonta grandis var. footiana — Simpson (1900), Simpson (1914)
Anodonta (Pyganodon) grandis-pepiniana — Frierson (1927)
Anodonta (Pyganodon) grandis pepiniana — Haas (1969)
Anodonta grandis grandis — Burch (1975)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta bealii ‘Lea’ of Authors

‘Lea’ of Authors.
Singley, 1893. Geol. Surv. Texas (1892): 316.
R.I. Johnson, 1980. Bull. MCZ 149(2): 92.
(error for Anodonta bealei Lea, 1863)

Pyganodon grandis — Parmalee & Bogan (1998)

Anodonta implicata Chadwick, 1906

‘Say’ Chadwick, 1906. unknown publication: 94.
(misidentified reference to Anodonta implicata Say, 1829)

Anodonta grandis plana — F.C. Baker (1928)
Pyganodon grandis — Parmalee & Bogan (1998)

Anodonta dakota Frierson, 1910

Frierson, 1910. Nautilus 23(9): 113, pl. 10.
Type(s): TYPE UMMZ_104010, Clear Lake, Ulvers Pt., Deuel Co., South Dakota.

Anodonta dakota — Simpson (1914)
Anodonta (Pyganodon) grandis-dakota — Frierson (1927)
Anodonta (Pyganodon) grandis dakota — Haas (1969)
Anodonta grandis grandis — R.I. Johnson (1980)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta dakotana Utterback, 1915

‘Frierson’ Utterback, 1915. Amer. Midl. Nat. 4(6): 265, pl. 24, figs. 77a-b.

Anodonta (Pyganodon) grandis-dakotana — Frierson (1927)
Anodonta (Pyganodon) grandis dakotana — Haas (1969)
Pyganodon grandis — Parmalee & Bogan (1998)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta palna Lea, 1834

Lea, 1834. Trans. Amer. Phil. Soc. 5 [O. 1]: 48.
(incorrect original spelling of Anodonta plana Lea, 1834)
Type(s): HOLOTYPE USNM_86626, Bear Grass Creek, near Louisville, Ky.

Anodonta grandis var. gigantea — Simpson (1900)
Anodonta grandis plana — F.C. Baker (1928)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta inflata Lea, 1836

‘Major Le Conte’s Cabinet’ Lea, 1836. Synopsis Fam. Naiades: 52. [nomen nudum]
‘Major Le Conte’s Cabinet’ Lea, 1838. Synopsis Fam. Naiades: 31. [nomen nudum]
‘Major Le Conte’s Cabinet’ Lea, 1852. Synopsis Fam. Naiades: 51. [nomen nudum]

Anodonta sulcata Küster, 1842

‘Lamarck’ Küster, 1842. Conch. Cab. Anodonta 9: 62, pl. 18, fig. 1.
(misidentified reference to Anodonta sulcata Lamarck, 1819)

Anodonta grandis var. footiana — Simpson (1900), Simpson (1914)
Anodonta grandis grandis — Burch (1975)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodon (Pseudodon) stewartii Gould, 1844

Gould, 1844. Proc. Boston Soc. Nat. Hist. 1: 161.
(error for Anodonta stewartiana Lea, 1834)

Anodonta cultrata Lea, 1852

‘Gould’ Lea, 1852. Synopsis Fam. Naiades: 47.
(intro'd as syn of Anodonta cultrata Lea, 1852)

Anodonta grandis var. benedictensis — Simpson (1900), Simpson (1914)

Anodonta harpenthis Conrad, 1853

‘Lea’ Conrad, 1853. Proc. Acad. Nat. Sci. 6: 264.
(error for Anodonta harpethensis Lea, 1840)

Anodonta fluviatilis Lapham, 1860

‘Lea’ Lapham, 1860. Proc. Acad. Nat. Sci. 12: 156.
(misidentified reference to Anodonta fluviatilis ‘Gmelin’ of Authors)

Anodonta grandis — F.C. Baker (1928)
Pyganodon grandis — Parmalee & Bogan (1998)

Anodon edentulus Sowerby, 1867

‘Lea, MS’ Sowerby, 1867. Conch. Icon. 17: pl. 17, sp. 60.
(misidentified reference to Alasmodonta edentula Say, 1829)

Anodonta grandis var. footiana — Simpson (1900), Simpson (1914)
Pyganodon grandis — Parmalee & Bogan (1998)
Pyganodon grandis grandis — Watters et al. (2009)

Anodon ovata Sowerby, 1869

‘Lea’ Sowerby, 1869. Conch. Icon. 17: pl. 22, sp. 85.
(reference to Anodonta ovata Lea, 1838)

Anodonta grandis var. gigantea — Simpson (1900), Simpson (1914)
Anodonta grandis grandis — Burch (1975)
Pyganodon grandis — Parmalee & Bogan (1998)

Margaron (Anodonta) subglobosa Lea, 1870

‘Anth.’ Lea, 1870. Synopsis Fam. Naiades: 81.
(error for Anodon subgibbosa J.G. Anthony, 1866)

Anodonta grandis var. gigantea — Simpson (1900)
Anodonta subgibbosa — Simpson (1914)
Pyganodon grandis — Parmalee & Bogan (1998)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta pepiniana Clessin, 1874

‘Lea’ Clessin, 1874. Conchyl. Cab.: 158, pl. 53, figs. 1-2.
(reference to Anodonta pepinianus Lea, 1838)

Anodonta pepiniana — Simpson (1900)

Anodonta hockingensis Call, 1880

‘Moores MSS’ Call, 1880. Amer. Natural. 14: 530.
(intro'd as syn of Anodonta grandis Say, 1829)

Anodonta hockingensis — Simpson (1900), Simpson (1914), Haas (1969)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta somersii Call, 1880

‘Moores MSS’ Call, 1880. Amer. Natural. 14: 530.
(intro'd as syn of Anodonta grandis Say, 1829)

Anodonta somersi — Simpson (1900), Simpson (1914), Haas (1969)
Pyganodon grandis — Parmalee & Bogan (1998), Williams et al. (2008)
Pyganodon grandis grandis — Watters et al. (2009)

Anodonta gessneri Paetel, 1890

‘Lea’ Paetel, 1890. Cat. Conch.-Samml. 3: 179.
(error for Anodonta gesnerii Lea, 1858)

Anodonta hallenbeckii — Simpson (1900), Simpson (1914)

Pyganodon simpsonaia Cyr et al., 2007

‘Lea’ Cyr et al., 2007. Can. J. Zool. 85: 1217.
(error for Anodonta simpsoniana Lea, 1861)