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Taxonomy

all genera

Unionidae

Margaritiferidae

Hyriidae

Etheriidae

Mycetopodidae

Iridinidae


Species & Genus Richness by
Taxonomy

Geography

Summary

North America

Central America

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Species Richness & Endemism by
Geography

Data last updated
Mon 1 Dec 2025

Publication Wu et al. (2021): summary

publication summary | cladograms

Full Citation

Wu, R., J. An, R. Chen, S. Ouyang & X. Wu. 2021. Taxonomy, phylogeny and evolution of freshwater mussels (Unionoida: Unionoidea) in China revealed by multilocus phylogenetic analyses and mitochondrial phylogenomics. Journal of Lake Sciences 33(6): 1788-1804. Available electronically.

Attributed Nominal Species

Parreysia burmana Bolotov et al., 2017 = Radiatula myitkyinae

‘Blanford’ of Authors.
‘Blanford’ Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Radiatula burmana’].
‘Blanford’ Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Huang et al., 2019. MPE 130: Table S4.
Ortiz-Sepulveda et al., 2020. MPE 148: Table S1.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
(misidentified reference to Unio burmanus Blanford, 1869)

image of type lot

Pseudodon cambodjensis Pfeiffer & Graf, 2013 = Pseudodon cambodjensis

‘Petit’ of Authors.
Pfeiffer & Graf, 2013. J. Moll. Studies 79: 251.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Zieritz et al., 2016. Sci. Total Environ. 571: 1071.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Monodontina cambodjensis’].
Lopes-Lima et al., 2017. MPE 106: 177.
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Monodontina cambodjensis’].
Razak et al., 2018. Conserv. Genet. Resour. 11: 459.
Huang et al., 2019. MPE 130: Table S4 [as ‘Monodontina cambodjensis’].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Monodontina cambodjensis’].
Bolotov et al., 2020. Sci. Repts. 10: Table S1 [as ‘Monodontina cambodiensis’ (sic)].
Konopleva et al., 2021. Sci. Repts. 11: Table S1 [as ‘Monodontina cambodiensis’ (sic)].
Jeratthitikul et al., 2021. Eur. J. Taxonomy 775: 123 [as ‘Monodontina cambodiensis’ (sic)].
Pfeiffer et al., 2021. Invert. Syst. 35: 453 [as ‘Monodontina cambodjensis’].
Wu et al., 2021. ZooKeys 1054: 87 [as ‘Monodontina cambodjensis’].
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
Konopleva et al., 2023. Diversity 15(1), 10: Table S2 [as ‘Monodontina cambodjensis’].
Dai et al., 2023. Invert. Syst. 37(2): Table 1 [as ‘Monodontina cambodjensis’].
Dai et al., 2024. Zoosyst. Evol. 100(4): Table 2.
Dai et al., 2025. Zoosyst. Evol. 101(2): Table 1.
(reference to Monocondylea cambodjensis Petit de la Saussaye, 1865)
Type(s): SEQUENCED SPECIMEN UMMZ_304350, Pursat River, just downstream (north) of Highway 5 bridge, Pursat City, Cambodia.

Lasmigona compressa Graf & Ó Foighil, 2000 = Lasmigona costata

‘Lea’ of Authors.
Graf & Ó Foighil, 2000. J. Moll. Studies 66: 160.
Graf & Cummings, 2006. Zool. J. Linn. Soc. 148: 348.
Hoeh et al., 2009. Malacologia 51(2): 310.
Jadhav & Jamkhedkar, 2009. Res. J. BioTech. 4(1): 43.
Whelan et al., 2011. MPE 61: 506.
Lopes-Lima et al., 2017. MPE 106: 177.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2018. Sci. Repts. 8: Table S1.
Huang et al., 2019. MPE 130: Table S4.
Lopes-Lima et al., 2020. MPE 146: Table 1.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
Jeratthitikul et al., 2022. Sci. Repts. 12: Table S1.
Wu et al., 2023. ZooKeys 1166: 263, Table 1.
Liu et al., 2023. ZooKeys 1179: Table 1.
Dai et al., 2025. Zoosyst. Evol. 101(2): Table 1. [in part]
(misidentified reference to Symphynota compressa Lea, 1829)

Unio crassus Soroka, 2005 = Unio nanus

‘Philippson’ of Authors.
Soroka, 2005. Zool. Sci. 22: 1138, Table 1.
Soroka, 2010. Folia Malac. 18(4): Table 2. [in part]
Reis et al., 2013. Graellsia 69(1): 22, Table 2.
Prié & Puillandre, 2014. Hydrobiologia 735(1): online resource 1 [as ‘Unio crassus crassus’].
Douda et al., 2014. J. Appl. Ecol. 51: 1087. [in part]
Froufe et al., 2016. Org Divers Evol 16(3): 606, fig. 3. [in part]
Klishko et al., 2017. Zootaxa 4286(1): 98, Table 3.
Soroka & Burzynski, 2017. mtDNA B 2(2): 863.
Burzynski et al., 2017. MPE 107: 606.
Lopes-Lima et al., 2017. Biol. Rev. 92(1): Table S1 [as ‘Unio crassus crassus’]. [in part]
Araujo et al., 2018. J. Linn. Soc., Zool. 182(2): supplementary table. [in part]
Klishko et al., 2018. Systemat. Biodiv. 16(3): 291.
Burzynski & Soroka, 2018. PeerJ 6, e5573: 3.
Wang et al., 2018. Conserv. Genet. Resour. 10(4): 611.
Wang et al., 2018. Conserv. Genet. Resour. 11: 140.
Cha et al., 2018. mtDNA B 3(2): 946.
Aunins et al., 2018. mtDNA B 3(2): 1125.
Wu et al., 2018. PLoS One 13(8), e0200956: 6. [in part]
Klishko et al., 2019. Ruthenica 29(1): 57, Table 1.
Wu et al., 2019. ZooKeys 812: 29.
Huang et al., 2019. MPE 130: Table S5. [in part]
Fukata & Iigo, 2019. mtDNA B 4(2): 3489.
Guerra et al., 2019. BMC Evol. Biol. 19 (229): Table S1.
Froufe et al., 2019. Heredity 124: 185.
Soroka, 2020. J. Zool. Syst. Evol. Res. 58(2): Table 2.
Fukata & Iigo, 2020. mtDNA B 5(2): 1216.
Sano et al., 2020. Plankton Benthos Res. 15(2): 99.
Wu et al., 2020. Invert. Syst. 34: Table S2.
Kilikowska et al., 2020. Life 10(7): 1. [in part]
Lopes-Lima et al., 2021. MPE 163: fig. 6 [by voucher number]. [in part]
Sayenko et al., 2021. Systemat. Biodiv. 19: 58, Table 1.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 2.
Zieritz et al., 2021. Zool. J. Linn. Soc. 193(3): 1092.
Kim et al., 2021. mtDNA B 6(1): 80.
Pfarrer et al., 2022. Hydrobiologia 849: ESM1.
Wu et al., 2022. Zool. Scripta 51: Table S2.
Dai et al., 2023. Invert. Syst. 37(2): Table 2.
Chen et al., 2023. ZooKeys 1173: Table 1.
Wu et al., 2023. Org Divers Evol (2023): Table 2.
Dai et al., 2024. Zool. Scripta 53: Table 1.
Wu et al., 2024. J. Zool. Syst. Evol. Res. 2024, 1499508: Table 2.
Wu et al., 2024. Zool. Scripta 53(5): Table S3.
Dai et al., 2024. Invert. Syst. 38(6): Table 3. [in part]
Chen et al., 2025. Zoosyst. Evol. 101(4): Tables S2-S3.
Dai et al., 2025. Zoosyst. Evol. 101(2): Tables 1-2.
(misidentified reference to Unio crassus Retzius, 1788)

Anodontites elongata Whelan et al., 2011 = Tamsiella schroeteriana

‘Swainson’ of Authors.
Whelan et al., 2011. MPE 61: 506.
Graf et al., 2015. MPE 85: 3.
Bolotov et al., 2016. MPE 103: 108.
Lopes-Lima et al., 2017. MPE 106: 178.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 208.
Combosch et al., 2017. MPE 107: 195.
Bolotov et al., 2018. Sci. Repts. 8: Table S1.
Huang et al., 2018. MPE 120: 302.
Lopes-Lima et al., 2018. MPE 127: 103.
Huang et al., 2019. MPE 130: Table S4.
Bolotov et al., 2019. Sci. Repts. 9: Table S1.
Pfeiffer et al., 2019. MPE 137: 116.
Guerra et al., 2019. BMC Evol. Biol. 19 (229): 3.
Bolotov et al., 2020. Sci. Repts. 10: Table S1.
Sano et al., 2020. Plankton Benthos Res. 15(2): 101.
Wu et al., 2020. Invert. Syst. 34: 195.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
Chen et al., 2025. Zoosyst. Evol. 101(4): Table 1.
(misidentified reference to Anodon elongatus Swainson, 1823)

There are 12 nominal species attributed to this publication (none available).
image of type lot

Pilsbryoconcha exilis Pfeiffer & Graf, 2015 = Pilsbryoconcha carinifera

‘Lea’ of Authors.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Ng et al., 2016. PLoS One 11(8), e0161130: [5]. [in part]
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘P. aff. exilis sp. 1’].
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘P. aff. exilis sp. 1’].
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘P. aff. exilis sp. 1’].
Huang et al., 2019. MPE 130: Table S4 [as ‘P. aff. exilis sp. 1’].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘P. aff. exilis sp. 1’].
Lopes-Lima et al., 2020. MPE 146: Table 1 [as ‘P. aff. exilis sp. 1’].
Zieritz et al., 2020. Biodivers. Conserv. 29: 2244.
Wu et al., 2021. ZooKeys 1054: 87.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
Konopleva et al., 2023. Diversity 15(1), 10: Table S2 [as ‘P. aff. exilis’].
Dai et al., 2023. Invert. Syst. 37(2): Tables 1-2. [in part]
(misidentified reference to Anodonta exilis Lea, 1838)
Type(s): SEQUENCED SPECIMEN UMMZ_304647, Beung Dang Teuk Leach Community Fishery Pursat Province, Cambodia.

Lamellidens generosus Whelan et al., 2011 = Lamellidens savadiensis

‘Gould’ of Authors.
Whelan et al., 2011. MPE 61: 506.
Pfeiffer & Graf, 2013. J. Moll. Studies 79: 251.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Araujo et al., 2017. Zool. J. Linn. Soc. 179(2): 280.
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Pfeiffer et al., 2018. J. Moll. Studies 84(4): [3].
Pfeiffer et al., 2019. MPE 137: 117.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
(misidentified reference to Unio generosus Gould, 1847)

Chamberlainia hainesiana Kongim et al., 2015 = Chamberlainia somsakpanhai

‘Lea’ of Authors.
Kongim et al., 2015. ZooKeys 514: 98, fig. 2A.
Lopes-Lima et al., 2017. MPE 106: 177.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Froufe et al., 2019. Heredity 124: 185.
Huang et al., 2019. MPE 130: Table S4.
Chen et al., 2020. mtDNA B 5(3): 2800.
Teiga-Teixeira et al., 2020. mtDNA B 5(3): 2978.
Zieritz et al., 2020. Biodivers. Conserv. 29: 2244.
Bolotov et al., 2020. Sci. Repts. 10: Table S1.
Lopes-Lima et al., 2020. MPE 146: Table 1.
Zieritz et al., 2021. Zool. J. Linn. Soc. 193(3): Table 1.
Zieritz et al., 2021. Aquatic Conserv.: Mar. Freshw. Ecosyst. 31: 3176.
Jeratthitikul et al., 2021. Eur. J. Taxonomy 775: 125.
Wu et al., 2021. ZooKeys 1054: 87.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
Wu et al., 2022. Zool. Scripta 51: Table S2.
Pavan-Kumar et al., 2022. Mol. Biol. Rep. 49: Table S1.
Jeratthitikul et al., 2022. Sci. Repts. 12: Table S1.
Dai et al., 2023. Invert. Syst. 37(2): Tables 1-2.
Konopleva et al., 2023. Zoosyst. Evol. 99(1): Table S1.
Wu et al., 2023. Org Divers Evol (2023): Table 2.
Dai et al., 2024. Zool. Scripta 53: Table 1.
Dai et al., 2024. Invert. Syst. 38(6): Table 3.
Dai et al., 2024. Zoosyst. Evol. 100(4): Table 2.
Chen et al., 2025. Zoosyst. Evol. 101(4): Table S3.
Dai et al., 2025. Zoosyst. Evol. 101(2): Table 1.
Dai et al., 2025. Zoosyst. Evol. 101(2): Table 2.
(misidentified reference to Unio hainesianus Lea, 1856)

Cuneopsis heudei Lopes-Lima et al., 2017 = Cuneopsis szechenyii

‘Heude’ of Authors.
Liu et al., 2017. PLoS One 12(12 ), e0189737: Table S4.
Lopes-Lima et al., 2017. MPE 106: Table 2.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Wu et al., 2018. PLoS One 13(8), e0200956: 5.
Huang et al., 2019. MPE 130: Tables S1, S4. [in part]
Lopes-Lima et al., 2020. MPE 146: Table 1. [in part]
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1. [in part]
Wu et al., 2022. Zool. Scripta 51: Table S1. [in part]
Jeratthitikul et al., 2022. Sci. Repts. 12: Table S1.
Konopleva et al., 2023. Zoosyst. Evol. 99(1): Table S1.
Chen et al., 2023. ZooKeys 1173: Table 1.
Wu et al., 2023. ZooKeys 1166: Table 1.
Liu et al., 2023. ZooKeys 1179: Table 1.
Chen et al., 2025. Zoosyst. Evol. 101(4): Table S2. [in part]
Dai et al., 2025. Zoosyst. Evol. 101(2): Table 1. [in part]
(misidentified reference to Unio heudei Heude, 1874)

Schistodesmus lampreyanus Huang et al., 2002 = Schistodesmus xinyuensis

‘Baird & Adams’ of Authors.
Huang et al., 2002. J. Moll. Studies 68: Table 1.
Zhang et al., 2004. Acta Hydrobiol. Sinica 28(3): fig. 1.
Zhou et al., 2007. Acta Zool. Sinica 53(6): Table 1.
Szumowski et al., 2012. Amer. Malac. Bull. 30(1): Table S3.
Liu et al., 2017. PLoS One 12(12 ), e0189737: Table S4.
Lopes-Lima et al., 2017. MPE 106: Table 2.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Sano et al., 2017. J. Wat. Res. Prot. 9: Table 2.
Wu et al., 2018. PLoS One 13(8), e0200956: Table 1.
Wu et al., 2019. ZooKeys 812: 29.
Fukata & Iigo, 2019. mtDNA B 4(2): fig. 1.
Huang et al., 2019. MPE 130: Tables S1, S4. [in part]
Fukata & Iigo, 2020. mtDNA B 5(2): fig. 1.
Wu et al., 2020. Invert. Syst. 34: Table S2.
Wu et al., 2020. Malacologia 63(1): Table 1.
Lopes-Lima et al., 2020. MPE 146: Table 1.
Choi et al., 2020. Sci. Repts. 10: Table S5.
Sano et al., 2020. Plankton Benthos Res. 15(2): Table 1.
Kim et al., 2021. mtDNA B 6(1): fig. 1.
Wu et al., 2021. J. Lake Sci. 33(6): attached Tables 1-2.
Wu et al., 2022. Zool. Scripta 51: Tables S1-S2.
Pavan-Kumar et al., 2022. Mol. Biol. Rep. 49: Table S1.
Jeratthitikul et al., 2022. Sci. Repts. 12: Table S1.
Dai et al., 2023. Invert. Syst. 37(2): Table 2.
Wu et al., 2023. Org Divers Evol (2023): Table 2.
Wu et al., 2023. ZooKeys 1166: Table 1.
Liu et al., 2023. ZooKeys 1179: Table 1.
Bogan et al., 2023. Diversity 15(6), 710: Table 1.
Chen et al., 2023. ZooKeys 1173: Table 1.
Dai et al., 2024. Zool. Scripta 53: Tables 1, S2.
Wu et al., 2024. J. Zool. Syst. Evol. Res. 2024, 1499508: Table 2.
Wu et al., 2024. Zool. Scripta 53(5): Tables S2-S3.
Dai et al., 2024. Invert. Syst. 38(6): Tables 2-3.
Hou et al., 2024. Invert. Syst. 38, IS24071: Table 2.
Hou et al., 2025. Zoosyst. Evol. 101(1): Table S2.
Hou et al., 2025. Zoosyst. Evol. 101(1): Table S2.
Dai et al., 2025. Zoosyst. Evol. 101(2): Tables 1-2.
Chen et al., 2025. Zoosyst. Evol. 101(4): Table S3.
(misidentified reference to Unio (Dysnomia) lampreyanus Baird & Adams, 1867)

Arconaia lanceolate Zhou et al., 2017 = Lanceolaria lanceolata

Zhou et al., 2017. Conserv. Genet. Resour. 9: 235.
Wang et al., 2018. Conserv. Genet. Resour. 11: 140.
Wu et al., 2021. J. Lake Sci. 33(6): 1799.
(error for Triquetra lanceolata Lea, 1856)

Ensidens sagittarius Pfeiffer & Graf, 2015 = Ensidens telus

‘Lea’ of Authors.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Lopes-Lima et al., 2017. MPE 106: 177.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘E. aff. sagittarius sp. 1].
Huang et al., 2019. MPE 130: Table S4 [as ‘Ensidens aff. sagittarius sp. 1].
Zieritz et al., 2020. Biodivers. Conserv. 29: 2242.
Lopes-Lima et al., 2020. MPE 146: Table 1.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
(misidentified reference to Unio sagittarius Lea, 1856)

Taxonomic Opinions

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