Lopes-Lima, M., J. Geist, S. Egg, L. Beran, A. Bikashvili et al. 2024. Integrative phylogenetic, phylogeographic and morphological characterisation of the Unio crassus species complex reveals cryptic diversity with important conservation implications. Molecular Phylogenetics and Evolution 195. 108046 (27 pp.). https://doi.org/10.1016/j.ympev.2024.108046.
(click the figure number to see the cladogram)
+-- Unio_pictorum
+--------------------------+
| +-- Unio_delphinus
--+
| +-------------------------- Unio_tumidiformis
| |
+--+ +----------------------- Unio_sesirmensis
| |
+--+ +-------------------- Unio_desectus
| |
+--+ +----------------- Unio_damascensis
| |
| | +-------------- Unio_ionicus
+--+ |
| | +-- Unio_nanus
| | +--------+
+--+ | +-- Unio_carneus
| |
| | +-- Unio_gontierii
| | |
| | +-- Unio_crassus__1
+--+--------+
| +-- Unio_crassus__2
| |
| +-- Unio_crassus__3
|
| +-------- Unio_vicarius
| |
+--+ +----- Unio_bruguierianus__1
| |
+--+----- Unio_bruguierianus__2
|
| +-- Unio_bruguierianus__3
+--+
+-- Unio_mardinensis
+----- Lanceolaria_lanceolata
+--------------------------+
| | +-- Anodonta_anatina
| +--+
| +-- Anodonta_cygnea
--+
| +-- Tchangsinaia_piscicula
| +--------------------------+
| | +-- Nodularia_douglasiae
+--+
| +-------------------------- Unio_gibbus
| |
+--+ +----------------------- Unio_tumidus
| |
| | +----- Unio_terminalis
+--+ +--------------+
| | | +-- Unio_delphinus
| | +--+
| | +-- Unio_pictorum
+--+
| +----------------- Unio_tumidiformis
| |
| | +-- Unio_ionicus
+--+ +-----------+
| | +-- Unio_carneus
| |
| | +-- Unio_damascensis
+--+ +--------+
| | +-- Unio_sesirmensis
| |
| | +----- Unio_bruguierianus
+--+ +--+
| | | +-- Unio_gontierii
| | +--+
| | +-- Unio_mardinensis
+--+
| +-- Unio_crassus
| +--+
| | +-- Unio_nanus
+--+
| +-- Unio_vicarius
+--+
+-- Unio_desectus
+----- Lanceolaria_lanceolata
+--------------------------------+
| | +-- Anodonta_anatina
| +--+
| +-- Anodonta_cygnea
--+
| +-- Tchangsinaia_piscicula
| +--------------------------------+
| | +-- Nodularia_dualobtusa
| |
+--+ +-- Unio_gibbus
| +-----------------------------+
| | +-- Unio_tumidus
| |
+--+ +----- Unio_terminalis
| +-----------------------+
| | | +-- Unio_delphinus
| | +--+
+--+ +-- Unio_pictorum
|
| +-------------------------- Unio_tumidiformis
| |
+--+ +----------------------- Unio_sesirmensis
| |
+--+ +-------------------- Unio_ionicus
| |
+--+ +----------------- Unio_damascensis
| |
+--+ +-------------- Unio_desectus
| |
| | +-- Unio_crassus
+--+ +--------+
| | +-- Unio_gontierii
| |
+--+ +-- Unio_bruguierianus
| +-----+
| | +-- Unio_mardinensis
+--+
| +----- Unio_vicarius
+--+
| +-- Unio_carneus
+--+
+-- Unio_nanus