MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
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Geography
Species Richness & Endemism of
freshwater mussels
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Data last updated
Fri 13 February 2026
Bivalvia | ... | Unionoidea | Unionidae | Ambleminae | Quadrulini | Quadrulina | Pustulosa
species Pustulosa kieneriana (Lea, 1852): synonymy
3 specimens + 13 GenBank vouchers
Synonymy
Unio keinerianus Lea, 1852
Lea, 1852. Proc. Amer. Phil. Soc. 5: 251. [nomen nudum]
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 281 [37], pl. 23, fig. 40.
(incorrect original spelling of Unio kienerianus Lea, 1852)
Type(s): HOLOTYPE USNM_84273, Coosawattee R., Ga.
Quadrula (sec. Theliderma) pustulosa var. kieneriana — Simpson (1914)
Cyclonaias kieneriana — Bieler (2021)
Unio kienerianus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 281 [37], pl. 23, fig. 40 [as ‘keineriana’].
(correct name for Unio keinerianus Lea, 1852)
Quadrula (sec. Theliderma) pustulosa var. keineriana [sic] — Simpson (1900)
Quadrula (sec. Theliderma) pustulosa var. kieneriana — Simpson (1914)
Quadrula (Bullata) pustulosa-keineriana [sic] — Frierson (1927)
Quadrula (Pustulosa) pustulosa keineriana [sic] — Haas (1969)
Quadrula pustulosa — R.I. Johnson (1980), Howells et al. (1996)
Quadrula keineriana [sic] — Serb et al. (2003)
Quadrula kieneriana — Campbell et al. (2005), Williams et al. (2008), Bogan (2010), Szumowski et al. (2012)
Amphinaias keineriana [sic] — ITIS (2013)
Cyclonaias kieneriana — Williams et al. (2017), Lopes-Lima et al. (2019), Graf & Cummings (2021), Bieler (2021), FMCS (2021), Hopper et al. (2023), Hauck et al. (2023)
Pustulosa kieneriana — Neemuchwala et al. (2023), FMCS (2025)
Unio asperatus Lea, 1861
Lea, 1861. Proc. Acad. Nat. Sci. 13: 41.
Lea, 1862. Jour. Acad. Nat. Sci. 5 [O. 8]: 68 [72], pl. 7, fig. 218.
Type(s): HOLOTYPE USNM_84272, Claiborne, Ala.
Quadrula (sec. Theliderma) pustulosa var. pernodosa — Simpson (1900), Simpson (1914)
Quadrula pustulosa asperata — Utterback (1915)
Quadrula asperata — Ortmann (1923), Turgeon et al. (1988), Williams et al. (1993), Turgeon et al. (1998), Lydeard et al. (1999), ... Williams et al. (2008), Bogan (2010), ITIS (2012), Szumowski et al. (2012), Haag (2012)
Quadrula (Bullata) pustulosa-asperata — Frierson (1927)
Quadrula (Pustulosa) pustulosa pernodosa — Haas (1969)
Quadrula pustulosa — Burch (1975), R.I. Johnson (1980)
Amphinaias asperata — IUCN (1996), Graf & Cummings (2007), Cummings & Graf (2010)
Cyclonaias asperata — Williams et al. (2017), Johnson et al. (2018), Hauck et al. (2023)
Cyclonaias kieneriana — Lopes-Lima et al. (2019), Graf & Cummings (2021), FMCS (2021)
Pustulosa kieneriana — FMCS (2025)
Quadrula archeri Frierson, 1905
Frierson, 1905. Nautilus 19(2): 13, pl. 1, figs. 1-2.
Type(s): HOLOTYPE UMMZ_113538, Tallapoosa River, Tallassee, Elmore, Alabama.
Quadrula (sec. Theliderma) archeri — Simpson (1914)
Quadrula (Bullata) archeri — Frierson (1927)
Quadrula (Pustulosa) archeri — Haas (1969)
Quadrula archeri — Burch (1975)
Amphinaias archeri — Graf & Cummings (2007), Cummings & Graf (2010)
Quadrula asperata — Williams et al. (2008)
Cyclonaias archeri — Williams et al. (2017), FMCS (2021), Hopper et al. (2023)
Cyclonaias pustulosa — Lopes-Lima et al. (2019), Graf & Cummings (2021)
Pustulosa kieneriana — FMCS (2025)
Unio asperatus Sowerby, 1868
‘Lea’ Sowerby, 1868. Conch. Icon. 16: pl. 85, sp. 450.
(reference to Unio asperatus Lea, 1861)
Quadrula (sec. Theliderma) pustulosa var. pernodosa — Simpson (1900)
Quadrula (Striata) striata — Frierson (1927)
Taxonomic History (since 1900)
Quadrula (sec. Theliderma) pustulosa var. keineriana [sic] — Simpson (1900)
Quadrula (sec. Theliderma) pustulosa var. pernodosa — Simpson (1900), Simpson (1914)
Quadrula (sec. Theliderma) pustulosa var. kieneriana — Simpson (1914)
Quadrula (sec. Theliderma) archeri — Simpson (1914)
Quadrula pustulosa asperata — Utterback (1915)
Quadrula asperata — Ortmann (1923), Turgeon et al. (1988), Williams et al. (1993), Turgeon et al. (1998), Lydeard et al. (1999), ... Williams et al. (2008), Bogan (2010), ITIS (2012), Szumowski et al. (2012), Haag (2012)
Quadrula (Bullata) pustulosa-keineriana [sic] — Frierson (1927)
Quadrula (Bullata) pustulosa-asperata — Frierson (1927)
Quadrula (Striata) striata — Frierson (1927)
Quadrula (Bullata) archeri — Frierson (1927)
Quadrula (Pustulosa) pustulosa keineriana [sic] — Haas (1969)
Quadrula (Pustulosa) pustulosa pernodosa — Haas (1969)
Quadrula (Pustulosa) archeri — Haas (1969)
Quadrula pustulosa — Burch (1975), R.I. Johnson (1980), Howells et al. (1996)
Quadrula archeri — Burch (1975)
Amphinaias asperata — IUCN (1996), Graf & Cummings (2007), Cummings & Graf (2010)
Quadrula keineriana [sic] — Serb et al. (2003)
Quadrula kieneriana — Campbell et al. (2005), Williams et al. (2008), Bogan (2010), Szumowski et al. (2012)
Amphinaias archeri — Graf & Cummings (2007), Cummings & Graf (2010)
Amphinaias keineriana [sic] — ITIS (2013)
Cyclonaias kieneriana — Williams et al. (2017), Lopes-Lima et al. (2019), Bieler (2021), Graf & Cummings (2021), FMCS (2021), Hopper et al. (2023), Hauck et al. (2023)
Cyclonaias asperata — Williams et al. (2017), Johnson et al. (2018), Hauck et al. (2023)
Cyclonaias archeri — Williams et al. (2017), FMCS (2021), Hopper et al. (2023)
Cyclonaias pustulosa — Lopes-Lima et al. (2019), Graf & Cummings (2021)
Pustulosa kieneriana — Neemuchwala et al. (2023), FMCS (2025)
