Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini | Cristariina

genus Beringiana Starobogatov in Zatravkin, 1983: cladomics

4 Recent valid species (4 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 4 (4 total)
analyzed in 49 cladograms in 34 publications
maximum number of terminals in an analysis = 4 (mean = 1.96, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 19
maximum number of included terminals = 4, (mean = 3.47 median = 4)
proportion of cladograms that support basic monophyly = 0.95

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Beringiana cladogram distances
PCoA Ordination

NJ Tree of Beringiana cladogram distances
NJ Tree

Cladograms

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
1 3 0 20 11 4 2 0 
  +-------------- Cristaria               
  |                                       
  |  +----------- Sinanodonta_tumens      
  |  |                                    
  |  |        +-- Sinanodonta_lauta__5    
  |  |        |                           
--+  |        +-- Sinanodonta_lauta__6    
  |  |        |                           
  |  |        +-- Beringiana_japonica__15 ** 
  |  |        |                           
  |  |        +-- Beringiana_japonica__16 ** 
  |  |  +-----+                           
  |  |  |     +-- Beringiana_japonica__17 ** 
  +--+  |     |                           
     |  |     +-- Beringiana_japonica__18 ** 
     |  |     |                           
     |  |     +-- Beringiana_japonica__19 ** 
     |  |     |                           
     |  |     +-- Beringiana_japonica__20 ** 
     |  |                                 
     |  |  +----- Beringiana_japonica__1 **  
     |  |  |                              
     |  |  +----- Beringiana_japonica__2 **  
     |  |  |                              
     +--+  +----- Beringiana_japonica__3 **  
        |  |                              
        |  +----- Beringiana_japonica__4 **  
        |  |                              
        |  +----- Beringiana_japonica__5 **  
        |  |                              
        |  +----- Sinanodonta_lauta__1    
        |  |                              
        |  +----- Beringiana_japonica__6 **  
        |  |                              
        |  +----- Sinanodonta_lauta__2    
        |  |                              
        |  +----- Sinanodonta_calipygos__1
        |  |                              
        +--+----- Beringiana_japonica__7 **  
           |                              
           +----- Sinanodonta_lauta__4    
           |                              
           +----- Beringiana_japonica__8 **  
           |                              
           +----- Beringiana_japonica__9 **  
           |                              
           +----- Beringiana_japonica__10 ** 
           |                              
           +----- Beringiana_japonica__11 ** 
           |                              
           +----- Beringiana_japonica__12 ** 
           |                              
           +----- Beringiana_japonica__13 ** 
           |                              
           +----- Beringiana_japonica__14 ** 
           |                              
           |  +-- Sinanodonta_lauta__3    
           +--+                           
              +-- Sinanodonta_calipygos__2

Wu et al., 2025 [Publ 4442], fig. 2

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
4 4 2 4 31 29 7 28 
  +----------------- Cristaria                  
  |                                             
  |              +-- Acudonta                   
  |     +--------+                              
  |     |        +-- Sinanodonta                
--+  +--+                                       
  |  |  |  +-------- Beringiana_gosannensis **     
  |  |  +--+                                    
  |  |     |  +----- Beringiana_fukuharai **       
  |  |     +--+                                 
  |  |        |  +-- Beringiana_beringiana **      
  +--+        +--+                              
     |           +-- Beringiana_japonica **        
     |                                          
     |        +----- Pletholophus_guangzhouensis
     |  +-----+                                 
     |  |     |  +-- Pletholophus_tenuis        
     |  |     +--+                              
     +--+        +-- Pletholophus_reinianus     
        |                                       
        |  +-------- Pletholophus_honglinhensis 
        +--+                                    
           |  +----- Anemina                    
           +--+                                 
              |  +-- Amuranodonta               
              +--+                              
                 +-- Buldowskia

Sano et al., 2022 [Publ 4311], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 4 8 8 4 28 
           +----- Pletholophus          
  +--------+                            
  |        |  +-- Cristaria             
  |        +--+                         
--+           +-- Buldowskia            
  |                                     
  |  +----------- Sinanodonta           
  +--+                                  
     |  +-------- Beringiana_fukuharai **  
     +--+                               
        |  +----- Beringiana_japonica **   
        +--+                            
           |  +-- Beringiana_gosannensis **
           +--+                         
              +-- Beringiana_beringiana **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
4 4 2 4 118 20 6 28 
  +----------------- Cristaria             
  |                                        
  |           +----- Anemina               
  +-----------+                            
--+           |  +-- Amuranodonta          
  |           +--+                         
  |              +-- Buldowskia            
  |                                        
  |  +-------------- Pletholophus          
  +--+                                     
     |  +----------- Sinanodonta           
     +--+                                  
        |  +-------- Beringiana_japonica **   
        +--+                               
           |  +----- Beringiana_beringiana ** 
           +--+                            
              |  +-- Beringiana_fukuharai **  
              +--+                         
                 +-- Beringiana_gosannensis **

Dai et al., 2026 [Publ 4757], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 4 30 26 6 28 
                 +----- Amuranodonta          
  +--------------+                            
  |              |  +-- Anemina               
  |              +--+                         
--+                 +-- Buldowskia            
  |                                           
  |  +----------------- Cristaria             
  +--+                                        
     |  +-------------- Pletholophus          
     +--+                                     
        |  +----------- Sinanodonta           
        +--+                                  
           |  +-------- Beringiana_japonica **   
           +--+                               
              |  +----- Beringiana_beringiana ** 
              +--+                            
                 |  +-- Beringiana_fukuharai **  
                 +--+                         
                    +-- Beringiana_gosannensis **

Xiang et al., 2025 [Publ 4433], fig. 6

iTtTbiriSoTroTroSiD
4 4 2 4 24 23 7 28 
  +-------------- Pletholophus          
  |                                     
  |     +-------- Cristaria             
  |  +--+                               
--+  |  |  +----- Anemina               
  |  |  +--+                            
  |  |     |  +-- Amuranodonta          
  |  |     +--+                         
  +--+        +-- Buldowskia            
     |                                  
     |        +-- Mimo                  
     |  +-----+                         
     |  |     +-- Sinanodonta           
     |  |                               
     +--+     +-- Beringiana_beringiana ** 
        |  +--+                         
        |  |  +-- Beringiana_japonica **   
        +--+                            
           |  +-- Beringiana_fukuharai **  
           +--+                         
              +-- Beringiana_gosannensis **

Dai et al., 2024 [Publ 4385], fig. 3-BI

iTtTbiriSoTroTroSiD
4 4 2 4 27 23 6 28 
                 +----- Amuranodonta          
  +--------------+                            
  |              |  +-- Anemina               
  |              +--+                         
--+                 +-- Buldowskia            
  |                                           
  |  +----------------- Cristaria             
  +--+                                        
     |  +-------------- Pletholophus          
     +--+                                     
        |  +----------- Sinanodonta           
        +--+                                  
           |  +-------- Beringiana_japonica **   
           +--+                               
              |  +----- Beringiana_beringiana ** 
              +--+                            
                 |  +-- Beringiana_fukuharai **  
                 +--+                         
                    +-- Beringiana_gosannensis **

Sano et al., 2022 [Publ 4311], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 4 14 14 4 28 
           +----- Pletholophus          
  +--------+                            
  |        |  +-- Cristaria             
  |        +--+                         
--+           +-- Buldowskia            
  |                                     
  |  +----------- Sinanodonta           
  +--+                                  
     |  +-------- Beringiana_fukuharai **  
     +--+                               
        |  +----- Beringiana_gosannensis **
        +--+                            
           |  +-- Beringiana_japonica **   
           +--+                         
              +-- Beringiana_beringiana **

Dai et al., 2024 [Publ 4385], fig. 3-ML

iTtTbiriSoTroTroSiD
4 4 2 4 27 23 6 28 
              +----- Amuranodonta          
  +-----------+                            
  |           |  +-- Anemina               
  |           +--+                         
--+              +-- Buldowskia            
  |                                        
  |  +-------------- Cristaria             
  |  |                                     
  +--+  +----------- Pletholophus          
     |  |                                  
     +--+  +-------- Sinanodonta           
        |  |                               
        +--+     +-- Beringiana_fukuharai **  
           |  +--+                         
           |  |  +-- Beringiana_gosannensis **
           +--+                            
              |  +-- Beringiana_beringiana ** 
              +--+                         
                 +-- Beringiana_japonica **

Bulakhova et al., 2023 [Publ 4325], fig. 5

iTtTbiriSoTroTroSiD
4 4 2 4 0 0 0 28 
     +-- Beringiana_beringiana ** 
  +--+                         
  |  +-- Beringiana_japonica **   
--+                            
  |  +-- Beringiana_gosannensis **
  +--+                         
     +-- Beringiana_fukuharai **

Lopes-Lima et al., 2020 [Publ 4080], fig. 7

iTtTbiriSoTroTroSiD
4 4 2 4 4 2 1 28 
  +----------- Sinanodonta           
--+                                  
  |  +-------- Beringiana_gosannensis **
  +--+                               
     |  +----- Beringiana_fukuharai **  
     +--+                            
        |  +-- Beringiana_beringiana ** 
        +--+                         
           +-- Beringiana_japonica **

Bogan et al., 2023 [Publ 4295], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 4 27 23 6 28 
  +-------------------- Cristaria             
  |                                           
  |              +----- Amuranodonta          
--+  +-----------+                            
  |  |           |  +-- Anemina               
  |  |           +--+                         
  +--+              +-- Buldowskia            
     |                                        
     |  +-------------- Pletholophus          
     +--+                                     
        |  +----------- Sinanodonta           
        +--+                                  
           |  +-------- Beringiana_japonica **   
           +--+                               
              |  +----- Beringiana_beringiana ** 
              +--+                            
                 |  +-- Beringiana_fukuharai **  
                 +--+                         
                    +-- Beringiana_gosannensis **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 65 8 5 28 
  +----------- Sinanodonta          
--+                                 
  |  +-------- Pletholophus         
  +--+                              
     |  +----- Beringiana_fukuharai ** 
     +--+                           
        |  +-- Beringiana_japonica **  
        +--+                        
           +-- Beringiana_beringiana **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 65 8 5 28 
  +----------- Sinanodonta          
--+                                 
  |  +-------- Pletholophus         
  +--+                              
     |  +----- Beringiana_fukuharai ** 
     +--+                           
        |  +-- Beringiana_japonica **  
        +--+                        
           +-- Beringiana_beringiana **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 65 8 5 28 
  +----------- Pletholophus         
--+                                 
  |  +-------- Sinanodonta          
  +--+                              
     |  +----- Beringiana_fukuharai ** 
     +--+                           
        |  +-- Beringiana_japonica **  
        +--+                        
           +-- Beringiana_beringiana **

Sano et al., 2017 [Publ 3799], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 30 6 4 28 
           +-- Cristaria            
  +--------+                        
  |        +-- Buldowskia           
  |                                 
--+        +-- Pletholophus         
  |  +-----+                        
  |  |     +-- Sinanodonta          
  +--+                              
     |  +----- Beringiana_fukuharai ** 
     +--+                           
        |  +-- Beringiana_japonica **  
        +--+                        
           +-- Beringiana_beringiana **

Sano et al., 2017 [Publ 3799], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 18 5 4 28 
              +-- Cristaria            
  +-----------+                        
  |           +-- Buldowskia           
--+                                    
  |  +----------- Pletholophus         
  +--+                                 
     |  +-------- Sinanodonta          
     +--+                              
        |  +----- Beringiana_fukuharai ** 
        +--+                           
           |  +-- Beringiana_japonica **  
           +--+                        
              +-- Beringiana_beringiana **

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
3 22 5 5 31 7 5 24 
  +----- Lanceolaria_grayii__1  
  |                             
  +----- Lanceolaria_grayii__2  
  |                             
  +----- Cristaria_plicata__1   
  |                             
  +----- Cristaria_plicata__2   
  |                             
  +----- Sinanodonta_lauta      
  |                             
  +----- Beringiana_japonica **    
  |                             
  +----- Beringiana_fukuharai__1 **
  |                             
  +----- Beringiana_beringiana **  
  |                             
  +----- Beringiana_fukuharai__2 **
  |                             
  +----- Beringiana_fukuharai__3 **
  |                             
  +----- Buldowskia_iwakawai__1 
  |                             
  +----- Buldowskia_iwakawai__2 
  |                             
  +----- Sinanodonta_tumens__1  
--+                             
  +----- Sinanodonta_tumens__2  
  |                             
  +----- Pletholophus           
  |                             
  +----- Anemina                
  |                             
  +----- Lanceolaria_lanceolata 
  |                             
  +----- Cristaria_plicata__3   
  |                             
  +----- Lanceolaria_gladiola   
  |                             
  +----- Sinanodonta_angula     
  |                             
  +----- Lepidodesma            
  |                             
  |  +-- Aculamprotula          
  +--+                          
  |  +-- Acuticosta             
  |                             
  |  +-- Cuneopsis              
  +--+                          
  |  +-- Nodularia              
  |                             
  |  +-- Schistodesmus          
  +--+                          
     +-- Inversiunio

Douda et al., 2024 [Publ 4364], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 12 11 1 -- 
  +----- Sinanodonta          
--+                           
  |  +-- Beringiana_beringiana **
  +--+                        
     +-- Beringiana_japonica **

Lopes-Lima et al., 2020 [Publ 4080], fig. 10

iTtTbiriSoTroTroSiD
2 2 0 2 12 10 1 -- 
  +----- Sinanodonta          
--+                           
  |  +-- Beringiana_japonica **  
  +--+                        
     +-- Beringiana_beringiana **

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 5 5 -- 
  +----------- Cristaria            
  |                                 
--+        +-- Anemina              
  |  +-----+                        
  |  |     +-- Amuranodonta         
  +--+                              
     |  +----- Pletholophus         
     +--+                           
        |  +-- Sinanodonta          
        +--+                        
           +-- Beringiana_beringiana **

Sayenko et al., 2017 [Publ 3760], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 3 3 1 -- 
  +-- Beringiana_beringiana **
--+                        
  +-- Sinanodonta

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
1 1 0 1 91 5 3 -- 
  +-------- Cristaria           
--+                             
  |  +----- Anemina             
  +--+                          
     |  +-- Beringiana_fukuharai **
     +--+                       
        +-- Sinanodonta

Wu et al., 2025 [Publ 4442], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 16 14 7 -- 
  +----------- Cristaria           
  |                                
  |     +----- Amuranodonta        
--+  +--+                          
  |  |  |  +-- Anemina             
  |  |  +--+                       
  |  |     +-- Buldowskia          
  +--+                             
     |     +-- Beringiana_fukuharai **
     |  +--+                       
     |  |  +-- Pletholophus        
     +--+                          
        |  +-- Acudonta            
        +--+                       
           +-- Sinanodonta

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iTtTbiriSoTroTroSiD
1 1 0 1 103 0 0 -- 
-- Beringiana_beringiana **

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 5 5 -- 
  +----- Pletholophus         
--+                           
  |  +-- Sinanodonta          
  +--+                        
     +-- Beringiana_beringiana **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
1 1 0 1 55 12 5 -- 
     +----- Anemina              
  +--+                           
  |  |  +-- Amuranodonta         
  |  +--+                        
--+     +-- Buldowskia           
  |                              
  |  +----- Cristaria            
  +--+                           
     |  +-- Beringiana_beringiana **
     +--+                        
        +-- Sinanodonta

Chong et al., 2008 [Publ 2668], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 17 1 1 -- 
  +-- Beringiana_beringiana **
--+                        
  +-- Sinanodonta

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 53 5 3 -- 
  +-------- Cristaria           
--+                             
  |  +----- Anemina             
  +--+                          
     |  +-- Beringiana_fukuharai **
     +--+                       
        +-- Sinanodonta

Hou et al., 2025 [Publ 4445], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 5 3 -- 
  +-------- Cristaria           
--+                             
  |  +----- Anemina             
  +--+                          
     |  +-- Beringiana_fukuharai **
     +--+                       
        +-- Sinanodonta

Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Beringiana_beringiana **

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 44 5 5 -- 
  +----- Pletholophus         
--+                           
  |  +-- Sinanodonta          
  +--+                        
     +-- Beringiana_beringiana **

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 35 8 5 -- 
     +----- Amuranodonta         
  +--+                           
  |  |  +-- Anemina              
  |  +--+                        
--+     +-- Buldowskia           
  |                              
  |  +----- Cristaria            
  +--+                           
     |  +-- Beringiana_beringiana **
     +--+                        
        +-- Sinanodonta

Xiang et al., 2025 [Publ 4433], fig. 7

iTtTbiriSoTroTroSiD
1 1 0 1 12 6 4 -- 
  +----------- Cristaria           
--+                                
  |  +-------- Anemina             
  +--+                             
     |  +----- Beringiana_fukuharai **
     +--+                          
        |  +-- Mimo                
        +--+                       
           +-- Sinanodonta

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
1 1 0 1 91 5 3 -- 
  +-------- Cristaria           
--+                             
  |  +----- Anemina             
  +--+                          
     |  +-- Beringiana_fukuharai **
     +--+                       
        +-- Sinanodonta

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 27 3 1 -- 
  +-- Beringiana_beringiana **
--+                        
  +-- Sinanodonta

Klishko et al., 2014 [Publ 3666], fig. 8

iTtTbiriSoTroTroSiD
1 1 0 1 3 3 2 -- 
  +----- Cristaria            
--+                           
  |  +-- Beringiana_beringiana **
  +--+                        
     +-- Sinanodonta

Soroka, 2010 [Publ 3404], fig. 9a

iTtTbiriSoTroTroSiD
1 1 0 1 4 4 2 -- 
  +----- Buldowskia           
--+                           
  |  +-- Beringiana_beringiana **
  +--+                        
     +-- Sinanodonta

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 75 5 3 -- 
  +-------- Cristaria           
--+                             
  |  +----- Anemina             
  +--+                          
     |  +-- Beringiana_fukuharai **
     +--+                       
        +-- Sinanodonta

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
1 1 0 1 95 5 3 -- 
  +-------- Cristaria           
--+                             
  |  +----- Anemina             
  +--+                          
     |  +-- Beringiana_fukuharai **
     +--+                       
        +-- Sinanodonta

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 44 5 5 -- 
  +----- Pletholophus         
--+                           
  |  +-- Sinanodonta          
  +--+                        
     +-- Beringiana_beringiana **

Wang et al., 2025 [Publ 4488], fig. 3B

iTtTbiriSoTroTroSiD
1 1 0 1 64 14 7 -- 
  +----------- Cristaria           
  |                                
  |     +----- Amuranodonta        
--+  +--+                          
  |  |  |  +-- Anemina             
  |  |  +--+                       
  |  |     +-- Buldowskia          
  +--+                             
     |     +-- Beringiana_fukuharai **
     |  +--+                       
     |  |  +-- Pletholophus        
     +--+                          
        |  +-- Acudonta            
        +--+                       
           +-- Sinanodonta

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 52 3 3 -- 
  +-------- Cristaria           
--+                             
  |  +----- Anemina             
  +--+                          
     |  +-- Beringiana_fukuharai **
     +--+                       
        +-- Sinanodonta

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 40 5 5 -- 
        +-- Anemina              
  +-----+                        
  |     +-- Amuranodonta         
--+                              
  |  +----- Pletholophus         
  +--+                           
     |  +-- Sinanodonta          
     +--+                        
        +-- Beringiana_beringiana **

Kano et al., 2019 [Publ 4194], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 9 2 2 -- 
  +----- Sinanodonta        
--+                         
  |  +-- Pletholophus       
  +--+                      
     +-- Beringiana_japonica **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
1 1 0 1 95 5 3 -- 
     +-- Anemina             
  +--+                       
  |  +-- Cristaria           
--+                          
  |  +-- Beringiana_fukuharai **
  +--+                       
     +-- Sinanodonta

Klishko et al., 2016 [Publ 3733], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 3 3 2 -- 
  +----- Cristaria            
--+                           
  |  +-- Sinanodonta          
  +--+                        
     +-- Beringiana_beringiana **

Froufe et al., 2017 [Publ 3802], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 11 9 3 -- 
  +-------- Anemina              
--+                              
  |  +----- Cristaria            
  +--+                           
     |  +-- Beringiana_beringiana **
     +--+                        
        +-- Sinanodonta

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 5 5 -- 
  +----------- Cristaria            
  |                                 
--+        +-- Anemina              
  |  +-----+                        
  |  |     +-- Amuranodonta         
  +--+                              
     |  +----- Pletholophus         
     +--+                           
        |  +-- Sinanodonta          
        +--+                        
           +-- Beringiana_beringiana **