Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini | Cristariina

genus Beringiana Starobogatov in Zatravkin, 1983: cladomics

4 Recent valid species (4 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 4 (4 total)
analyzed in 58 cladograms in 41 publications
maximum number of terminals in an analysis = 4 (mean = 1.97, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 22
maximum number of included terminals = 4, (mean = 3.55 median = 4)
proportion of cladograms that support basic monophyly = 0.95

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Beringiana cladogram distances
PCoA Ordination

NJ Tree of Beringiana cladogram distances
NJ Tree

Cladograms

Chen et al., 2026 [Publ 4805], fig. 4

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 30 29 8 **
Cladomics cladogram Cristaria Beringiana beringiana Beringiana japonica Beringiana gosannensis Beringiana fukuharai Mimo Acudonta Sinanodonta Pletholophus Anemina Amuranodonta Buldowskia

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.900 4 4 2 4 629 29 8 2.0
Cladomics cladogram Mimo Acudonta Sinanodonta Pletholophus Beringiana japonica Beringiana beringiana Beringiana fukuharai Beringiana gosannensis

Xiang et al., 2025 [Publ 4433], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 24 23 7 2.0
Cladomics cladogram Pletholophus Cristaria Anemina Amuranodonta Buldowskia Mimo Sinanodonta Beringiana beringiana Beringiana japonica Beringiana fukuharai Beringiana gosannensis

Wu et al., 2025 [Publ 4442], fig. 2

cst iT tT bi riS oT roT roS iD
0.889 4 4 2 4 31 29 7 4.0
Cladomics cladogram Cristaria Acudonta Sinanodonta Beringiana gosannensis Beringiana fukuharai Beringiana beringiana Beringiana japonica Pletholophus guangzhouensis Pletholophus tenuis Pletholophus reinianus Pletholophus honglinhensis Anemina Amuranodonta Buldowskia

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 118 20 6 2.0
Cladomics cladogram Cristaria Anemina Amuranodonta Buldowskia Pletholophus Sinanodonta Beringiana japonica Beringiana beringiana Beringiana fukuharai Beringiana gosannensis

Bogan et al., 2023 [Publ 4295], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 27 23 6 2.0
Cladomics cladogram Cristaria Amuranodonta Anemina Buldowskia Pletholophus Sinanodonta Beringiana japonica Beringiana beringiana Beringiana fukuharai Beringiana gosannensis

Dai et al., 2024 [Publ 4385], fig. 3-BI

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 27 23 6 2.0
Cladomics cladogram Amuranodonta Anemina Buldowskia Cristaria Pletholophus Sinanodonta Beringiana japonica Beringiana beringiana Beringiana fukuharai Beringiana gosannensis

Dai et al., 2024 [Publ 4385], fig. 3-ML

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 27 23 6 2.0
Cladomics cladogram Amuranodonta Anemina Buldowskia Cristaria Pletholophus Sinanodonta Beringiana fukuharai Beringiana gosannensis Beringiana beringiana Beringiana japonica

Dai et al., 2026 [Publ 4757], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 30 26 6 2.0
Cladomics cladogram Amuranodonta Anemina Buldowskia Cristaria Pletholophus Sinanodonta Beringiana japonica Beringiana beringiana Beringiana fukuharai Beringiana gosannensis

Sano et al., 2022 [Publ 4311], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 8 8 4 4.0
Cladomics cladogram Pletholophus Cristaria Buldowskia Sinanodonta Beringiana fukuharai Beringiana japonica Beringiana gosannensis Beringiana beringiana

Sano et al., 2022 [Publ 4311], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 14 14 4 4.0
Cladomics cladogram Pletholophus Cristaria Buldowskia Sinanodonta Beringiana fukuharai Beringiana gosannensis Beringiana japonica Beringiana beringiana

Lopes-Lima et al., 2020 [Publ 4080], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 4 2 1 4.0
Cladomics cladogram Sinanodonta Beringiana gosannensis Beringiana fukuharai Beringiana beringiana Beringiana japonica

Itoh et al., 2026 [Publ 4842], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 1 1 1 2.0
Cladomics cladogram Sinanodonta Beringiana japonica Beringiana beringiana Beringiana fukuharai Beringiana gosannensis

Bulakhova et al., 2023 [Publ 4325], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 0 0 0 2.0
Cladomics cladogram Beringiana beringiana Beringiana japonica Beringiana gosannensis Beringiana fukuharai

Sano et al., 2020 [Publ 4138], fig. 2

cst iT tT bi riS oT roT roS iD
0.857 3 3 1 3 66 8 5 3.0
Cladomics cladogram Pletholophus Sinanodonta Beringiana fukuharai Beringiana japonica Beringiana beringiana

Sano et al., 2020 [Publ 4138], fig. 1

cst iT tT bi riS oT roT roS iD
0.286 3 22 5 3 31 7 5 33.0
Cladomics cladogram Sinohyriopsis Lamprotula Pronodularia Sinosolenaia Inversidens Obovalis omiensis 1 Obovalis omiensis 2 Ptychorhynchus Lanceolaria oxyrhyncha 1 Lanceolaria oxyrhyncha 2 Cristaria plicata 1 Cristaria plicata 2 Sinanodonta lauta Beringiana japonica Beringiana fukuharai 1 Beringiana beringiana Beringiana fukuharai 2 Beringiana fukuharai 3 Buldowskia iwakawai 1 Buldowskia iwakawai 2 Sinanodonta tumens 1 Sinanodonta tumens 2 Pletholophus Anemina Lanceolaria lanceolata Cristaria plicata 3 Lanceolaria gladiola Sinanodonta angula Lepidodesma Scriptolamprotula Acuticosta Cuneopsis Nodularia Schistodesmus Inversiunio

Sano et al., 2020 [Publ 4138], fig. 3

cst iT tT bi riS oT roT roS iD
0.857 3 3 1 3 66 8 5 3.0
Cladomics cladogram Sinanodonta Pletholophus Beringiana fukuharai Beringiana japonica Beringiana beringiana

Sano et al., 2020 [Publ 4138], fig. 4

cst iT tT bi riS oT roT roS iD
0.857 3 3 1 3 66 8 5 3.0
Cladomics cladogram Sinanodonta Pletholophus Beringiana fukuharai Beringiana japonica Beringiana beringiana

Sano et al., 2017 [Publ 3799], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 18 5 4 3.0
Cladomics cladogram Cristaria Buldowskia Pletholophus Sinanodonta Beringiana fukuharai Beringiana japonica Beringiana beringiana

Sano et al., 2017 [Publ 3799], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 30 6 4 3.0
Cladomics cladogram Cristaria Buldowskia Pletholophus Sinanodonta Beringiana fukuharai Beringiana japonica Beringiana beringiana

Lopes-Lima et al., 2020 [Publ 4080], fig. 10

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 12 10 1 --
Cladomics cladogram Sinanodonta Beringiana japonica Beringiana beringiana

Douda et al., 2024 [Publ 4364], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 12 11 1 --
Cladomics cladogram Sinanodonta Beringiana beringiana Beringiana japonica

Wu et al., 2025 [Publ 4442], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 16 14 7 --
Cladomics cladogram Cristaria Amuranodonta Anemina Buldowskia Beringiana fukuharai Pletholophus Acudonta Sinanodonta

Wang et al., 2025 [Publ 4488], fig. 3B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 63 14 7 --
Cladomics cladogram Cristaria Amuranodonta Anemina Buldowskia Beringiana fukuharai Pletholophus Acudonta Sinanodonta

Wu et al., 2026 [Publ 4811], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 14 7 --
Cladomics cladogram Cristaria Amuranodonta Anemina Buldowskia Beringiana fukuharai Pletholophus Acudonta Sinanodonta

Wu et al., 2026 [Publ 4825], fig. 6B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 64 13 7 --
Cladomics cladogram Cristaria Amuranodonta Anemina Buldowskia Beringiana fukuharai Pletholophus Acudonta Sinanodonta

Bolotov et al., 2020 [Publ 4084], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 8 5 --
Cladomics cladogram Amuranodonta Anemina Buldowskia Cristaria Beringiana beringiana Sinanodonta

Konopleva et al., 2023 [Publ 4269], fig. S1_4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 55 12 5 --
Cladomics cladogram Anemina Amuranodonta Buldowskia Cristaria Beringiana beringiana Sinanodonta

Chen et al., 2023 [Publ 4302], fig. 1

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 40 5 5 --
Cladomics cladogram Pletholophus Sinanodonta Beringiana beringiana

Chen et al., 2023 [Publ 4302], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 5 5 --
Cladomics cladogram Cristaria Anemina Amuranodonta Pletholophus Sinanodonta Beringiana beringiana

Dai et al., 2025 [Publ 4440], fig. 1

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 43 5 5 --
Cladomics cladogram Pletholophus Sinanodonta Beringiana beringiana

Dai et al., 2025 [Publ 4440], fig. 2

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 43 5 5 --
Cladomics cladogram Pletholophus Sinanodonta Beringiana beringiana

Chen et al., 2025 [Publ 4487], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 5 5 --
Cladomics cladogram Cristaria Anemina Amuranodonta Pletholophus Sinanodonta Beringiana beringiana

Chen et al., 2025 [Publ 4487], fig. 3

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 40 5 5 --
Cladomics cladogram Anemina Amuranodonta Pletholophus Sinanodonta Beringiana beringiana

Xiang et al., 2025 [Publ 4433], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 12 6 4 --
Cladomics cladogram Cristaria Anemina Beringiana fukuharai Mimo Sinanodonta

Altaba et al., 2025 [Publ 4849], fig. 10

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 5 4 --
Cladomics cladogram Anemina Amuranodonta Cristaria Beringiana beringiana Sinanodonta

Froufe et al., 2017 [Publ 3802], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 9 3 --
Cladomics cladogram Anemina Cristaria Beringiana beringiana Sinanodonta

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 5 3 --
Cladomics cladogram Anemina Cristaria Beringiana fukuharai Sinanodonta

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 5 3 --
Cladomics cladogram Cristaria Anemina Beringiana fukuharai Sinanodonta

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 5 3 --
Cladomics cladogram Cristaria Anemina Beringiana fukuharai Sinanodonta

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 5 3 --
Cladomics cladogram Cristaria Anemina Beringiana fukuharai Sinanodonta

Wu et al., 2024 [Publ 4379], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 5 3 --
Cladomics cladogram Cristaria Anemina Beringiana fukuharai Sinanodonta

Wu et al., 2024 [Publ 4388], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 3 3 --
Cladomics cladogram Cristaria Anemina Beringiana fukuharai Sinanodonta

Hou et al., 2024 [Publ 4400], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 74 5 3 --
Cladomics cladogram Cristaria Anemina Beringiana fukuharai Sinanodonta

Hou et al., 2025 [Publ 4445], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 51 5 3 --
Cladomics cladogram Cristaria Anemina Beringiana fukuharai Sinanodonta

Altaba et al., 2025 [Publ 4849], fig. 11

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 3 3 --
Cladomics cladogram Cristaria Amuranodonta Sinanodonta Beringiana beringiana

Altaba et al., 2025 [Publ 4849], fig. 12

cst iT tT bi riS oT roT roS iD
0.600 1 1 0 1 12 6 3 --
Cladomics cladogram Beringiana beringiana Sinanodonta woodiana 2 Sinanodonta woodiana 1 Sinanodonta lauta Sinanodonta tumens Anodonta

Tumpeesuwan et al., 2026 [Publ 4845], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 9 3 --
Cladomics cladogram Beringiana beringiana Sinanodonta Buldowskia Cristaria

Soroka, 2010 [Publ 3404], fig. 9a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 4 4 2 --
Cladomics cladogram Buldowskia Beringiana beringiana Sinanodonta

Klishko et al., 2014 [Publ 3666], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 3 2 --
Cladomics cladogram Cristaria Beringiana beringiana Sinanodonta

Klishko et al., 2016 [Publ 3733], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 3 2 --
Cladomics cladogram Cristaria Sinanodonta Beringiana beringiana

Kano et al., 2019 [Publ 4194], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 2 2 --
Cladomics cladogram Sinanodonta Pletholophus Beringiana japonica

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

cst iT tT bi riS oT roT roS iD
0.500 1 3 3 1 11 4 2 28.0
Cladomics cladogram Cristaria Sinanodonta tumens Sinanodonta lauta 5 Sinanodonta lauta 6 Beringiana japonica 15 Beringiana japonica 16 Beringiana japonica 17 Beringiana japonica 18 Beringiana japonica 19 Beringiana japonica 20 Beringiana japonica 1 Beringiana japonica 2 Beringiana japonica 3 Beringiana japonica 4 Beringiana japonica 5 Sinanodonta lauta 1 Beringiana japonica 6 Sinanodonta lauta 2 Sinanodonta calipygos 1 Beringiana japonica 7 Sinanodonta lauta 4 Beringiana japonica 8 Beringiana japonica 9 Beringiana japonica 10 Beringiana japonica 11 Beringiana japonica 12 Beringiana japonica 13 Beringiana japonica 14 Sinanodonta lauta 3 Sinanodonta calipygos 2

Chong et al., 2008 [Publ 2668], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 1 1 --
Cladomics cladogram Beringiana beringiana Sinanodonta

Sayenko et al., 2017 [Publ 3760], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 3 1 --
Cladomics cladogram Beringiana beringiana Sinanodonta

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 27 3 1 --
Cladomics cladogram Beringiana beringiana Sinanodonta

Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 4 0 0 --
Cladomics cladogram Beringiana beringiana Brachyanodon

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 104 0 0 --
Cladomics cladogram Alasmidonta marginata Prolasmidonta Alasmidonta undulata Beringiana beringiana Brachyanodon kennerlyi Brachyanodon nuttalliana Brachyanodon oregonensis Arcidens confragosus Arcidens wheeleri Pterosyna Platynaias compressa Lasmigona Platynaias decorata Platynaias subviridis Pyganodon grandis Pyganodon lacustris Pseudodontoideus Strophitus Utterbackia imbecillis Utterbackia peninsularis 1 Utterbackia peninsularis 2