MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Mon 29 June 2026
Bivalvia | Unionida | Unionoidea | Margaritiferidae | Margaritiferinae
genus Margaritifera Schumacher, 1815: cladomics
7 Recent valid species (26 total species)
Cladomics pages are experimental — even more so than the rest of this site!
The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.
The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.
Following the summary, the available published clades representing the taxon are listed.
For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!
Cladomics Summary
number of species = 7 (26 total)
analyzed in 233 cladograms in 136 publications
maximum number of terminals in an analysis = 7 (mean = 2.21, median = 1)
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 113
maximum number of included terminals = 7, (mean = 3.5 median = 2)
proportion of cladograms that support basic monophyly = 0.96
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Cladograms
Araujo et al., 2017 [Publ 3746], fig. 1_2b
** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 7 | 7 | 5 | 7 | 6 | 3 | 2 | ** |
+-- Cumberlandia
+-----------+
| +-- Pseudunio
|
--+ +-- Margaritifera_dahurica **
| +--------+
| | +-- Margaritifera_margaritifera **
+--+
| +-------- Margaritifera_falcata **
| |
+--+ +-- Margaritifera_middendorffi **
| +--+
| | +-- Margaritifera_laevis **
+--+
| +-- Margaritifera_marrianae **
+--+
+-- Margaritifera_hembeli **
Araujo et al., 2017 [Publ 3746], fig. 2a
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 7 | 7 | 5 | 7 | 4 | 3 | 2 | 0.0 |
+-------------- Gibbosula | --+ +-- Margaritifera_dahurica ** | +--------+ | | +-- Margaritifera_margaritifera ** +--+ | +-------- Margaritifera_falcata ** | | +--+ +-- Margaritifera_laevis ** | +--+ | | +-- Margaritifera_middendorffi ** +--+ | +-- Margaritifera_marrianae ** +--+ +-- Margaritifera_hembeli **
Huang et al., 2018 [Publ 3828], fig. 3_5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 7 | 7 | 5 | 7 | 22 | 3 | 2 | 4.0 |
+-- Cumberlandia
+--------------+
| +-- Pseudunio
|
--+ +-- Margaritifera_margaritifera **
| +-----------+
| | +-- Margaritifera_dahurica **
+--+
| +----------- Margaritifera_falcata **
+--+
| +-------- Margaritifera_laevis **
+--+
| +----- Margaritifera_hembeli **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_marrianae **
Lopes-Lima et al., 2018 [Publ 3829], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 7 | 7 | 5 | 7 | 21 | 4 | 2 | 0.0 |
+-- Cumberlandia
+-----------+
| +-- Pseudunio
|
--+ +-- Margaritifera_dahurica **
| +--------+
| | +-- Margaritifera_margaritifera **
+--+
| +-------- Margaritifera_falcata **
| |
+--+ +-- Margaritifera_middendorffi **
| +--+
| | +-- Margaritifera_laevis **
+--+
| +-- Margaritifera_hembeli **
+--+
+-- Margaritifera_marrianae **
Lopes-Lima et al., 2018 [Publ 3829], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 7 | 7 | 5 | 7 | 21 | 4 | 2 | 2.0 |
+-- Cumberlandia
+--------------+
| +-- Pseudunio
|
--+ +-- Margaritifera_margaritifera **
| +-----------+
| | +-- Margaritifera_dahurica **
+--+
| +----------- Margaritifera_falcata **
+--+
| +-------- Margaritifera_laevis **
+--+
| +----- Margaritifera_middendorffi **
+--+
| +-- Margaritifera_hembeli **
+--+
+-- Margaritifera_marrianae **
Huang et al., 2019 [Publ 3843], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 7 | 7 | 4 | 7 | 194 | 4 | 2 | 3.0 |
+----------- Cumberlandia | +----------- Pseudunio | --+ +-- Margaritifera_dahurica ** | +-----+ | | +-- Margaritifera_margaritifera ** | | +--+ +----- Margaritifera_laevis ** | | | | +-- Margaritifera_falcata ** +--+--+ | +-- Margaritifera_middendorffi ** | | +-- Margaritifera_hembeli ** +--+ +-- Margaritifera_marrianae **
Huang et al., 2019 [Publ 3843], fig. S3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 7 | 7 | 5 | 7 | 192 | 4 | 2 | 0.0 |
+-- Cumberlandia
+-----------+
| +-- Pseudunio
|
--+ +-- Margaritifera_dahurica **
| +--------+
| | +-- Margaritifera_margaritifera **
+--+
| +-------- Margaritifera_falcata **
| |
+--+ +-- Margaritifera_hembeli **
| +--+
| | +-- Margaritifera_marrianae **
+--+
| +-- Margaritifera_laevis **
+--+
+-- Margaritifera_middendorffi **
Huang et al., 2019 [Publ 3843], fig. S4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 7 | 7 | 3 | 7 | 194 | 4 | 2 | 2.0 |
+-------------- Cumberlandia | --+ +----------- Pseudunio | | | | +-------- Margaritifera_falcata ** +--+ | | | +-- Margaritifera_dahurica ** | +-----+ +--+ +-- Margaritifera_margaritifera ** | | +----- Margaritifera_laevis ** | | +--+----- Margaritifera_middendorffi ** | | +-- Margaritifera_hembeli ** +--+ +-- Margaritifera_marrianae **
Lyubas et al., 2019 [Publ 3963], fig. 10
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 7 | 7 | 5 | 7 | 9 | 6 | 2 | 2.0 |
+-- Cumberlandia
+--------------+
| +-- Pseudunio
|
--+ +-- Margaritifera_margaritifera **
| +-----------+
| | +-- Margaritifera_dahurica **
+--+
| +----------- Margaritifera_falcata **
+--+
| +-------- Margaritifera_laevis **
+--+
| +----- Margaritifera_middendorffi **
+--+
| +-- Margaritifera_marrianae **
+--+
+-- Margaritifera_hembeli **
Wu et al., 2020 [Publ 4121], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 7 | 7 | 5 | 7 | 27 | 3 | 2 | 0.0 |
+-- Gibbosula
+-----------+
| +-- Pseudunio
|
--+ +-- Margaritifera_dahurica **
| +--------+
| | +-- Margaritifera_margaritifera **
+--+
| +-------- Margaritifera_falcata **
| |
+--+ +-- Margaritifera_hembeli **
| +--+
| | +-- Margaritifera_marrianae **
+--+
| +-- Margaritifera_laevis **
+--+
+-- Margaritifera_middendorffi **
Wu et al., 2020 [Publ 4121], fig. S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 7 | 7 | 5 | 7 | 27 | 3 | 2 | 0.0 |
+-- Cumberlandia
+-----------+
| +-- Pseudunio
|
--+ +-- Margaritifera_dahurica **
| +--------+
| | +-- Margaritifera_margaritifera **
+--+
| +-------- Margaritifera_falcata **
| |
+--+ +-- Margaritifera_hembeli **
| +--+
| | +-- Margaritifera_marrianae **
+--+
| +-- Margaritifera_laevis **
+--+
+-- Margaritifera_middendorffi **
Sano et al., 2020 [Publ 4138], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 7 | 7 | 5 | 7 | 62 | 3 | 2 | 2.0 |
+-- Gibbosula
+--------------+
| +-- Pseudunio
|
--+ +-- Margaritifera_dahurica **
| +-----------+
| | +-- Margaritifera_margaritifera **
+--+
| +----------- Margaritifera_falcata **
+--+
| +-------- Margaritifera_middendorffi **
+--+
| +----- Margaritifera_laevis **
+--+
| +-- Margaritifera_hembeli **
+--+
+-- Margaritifera_marrianae **
Sano et al., 2020 [Publ 4138], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 7 | 7 | 5 | 7 | 62 | 3 | 2 | 2.0 |
+----------------- Gibbosula | --+ +-- Margaritifera_margaritifera ** | +-----------+ | | +-- Margaritifera_dahurica ** +--+ | +----------- Margaritifera_falcata ** +--+ | +-------- Margaritifera_middendorffi ** +--+ | +----- Margaritifera_laevis ** +--+ | +-- Margaritifera_hembeli ** +--+ +-- Margaritifera_marrianae **
Sano et al., 2020 [Publ 4138], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 7 | 7 | 5 | 7 | 62 | 3 | 2 | 0.0 |
+-------------- Gibbosula | --+ +-- Margaritifera_margaritifera ** | +--------+ | | +-- Margaritifera_dahurica ** +--+ | +-------- Margaritifera_falcata ** | | +--+ +-- Margaritifera_laevis ** | +--+ | | +-- Margaritifera_middendorffi ** +--+ | +-- Margaritifera_hembeli ** +--+ +-- Margaritifera_marrianae **
Chen et al., 2025 [Publ 4455], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.500 | 7 | 7 | 5 | 7 | 19 | 4 | 2 | 2.0 |
+----------------- Cumberlandia | --+ +-- Margaritifera_dahurica ** | +-----------+ | | +-- Margaritifera_margaritifera ** +--+ | +----------- Margaritifera_falcata ** +--+ | +-------- Margaritifera_middendorffi ** +--+ | +----- Margaritifera_laevis ** +--+ | +-- Margaritifera_hembeli ** +--+ +-- Margaritifera_marrianae **
Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 7 | 7 | 5 | 7 | 133 | 4 | 2 | 2.0 |
+-------------------- Cumberlandia | --+ +----------------- Pseudunio | | +--+ +-- Margaritifera_dahurica ** | +-----------+ | | +-- Margaritifera_margaritifera ** +--+ | +----------- Margaritifera_falcata ** +--+ | +-------- Margaritifera_laevis ** +--+ | +----- Margaritifera_middendorffi ** +--+ | +-- Margaritifera_hembeli ** +--+ +-- Margaritifera_marrianae **
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 7 | 7 | 5 | 7 | 626 | 4 | 2 | 2.0 |
+----------------- Cumberlandia | --+ +-- Margaritifera_dahurica ** | +-----------+ | | +-- Margaritifera_margaritifera ** +--+ | +----------- Margaritifera_falcata ** +--+ | +-------- Margaritifera_middendorffi ** +--+ | +----- Margaritifera_laevis ** +--+ | +-- Margaritifera_hembeli ** +--+ +-- Margaritifera_marrianae **
Bolotov et al., 2015 [Publ 3646], fig. 2A
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 6 | 8 | 5 | 6 | 3 | 2 | 2 | 7.0 |
+-------------- Unio | | +----------- Margaritifera_margaritifera ** --+ | | | +----- Margaritifera_dahurica__1 ** | | | | +-----+----- Margaritifera_dahurica__2 ** +--+ | | | +-- Pseudunio | +--+ | +-- Cumberlandia | | +-------- Margaritifera_falcata ** +--+ | +----- Margaritifera_laevis ** +--+ | +-- Margaritifera_middendorffi ** +--+ +-- Margaritifera_marrianae **
Takeuchi et al., 2015 [Publ 3661], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 6 | 6 | 4 | 6 | 4 | 2 | 2 | 3.0 |
+----------------- Pseudunio_auricularius__1 | +----------------- Pseudunio_auricularius__2 --+ | +-------------- Cumberlandia | | +--+ +-- Margaritifera_margaritifera ** | +--------+ | | +-- Margaritifera_dahurica ** +--+ | +-------- Margaritifera_falcata ** +--+ | +----- Margaritifera_laevis ** +--+ | +-- Margaritifera_middendorffi ** +--+ +-- Margaritifera_marrianae **
Bolotov et al., 2016 [Publ 3743], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 6 | 6 | 4 | 6 | 24 | 3 | 2 | 3.0 |
+-- Gibbosula
+-----------+
| +-- Cumberlandia
|
--+ +-- Margaritifera_dahurica **
| +--------+
| | +-- Margaritifera_margaritifera **
+--+
| +-------- Margaritifera_falcata **
+--+
| +----- Margaritifera_laevis **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_marrianae **
Bolotov et al., 2016 [Publ 3743], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 6 | 6 | 4 | 6 | 24 | 3 | 2 | 3.0 |
+-- Gibbosula
+-----------+
| +-- Cumberlandia
|
--+ +-- Margaritifera_margaritifera **
| +--------+
| | +-- Margaritifera_dahurica **
+--+
| +-------- Margaritifera_falcata **
+--+
| +----- Margaritifera_laevis **
+--+
| +-- Margaritifera_marrianae **
+--+
+-- Margaritifera_middendorffi **
Bolotov et al., 2017 [Publ 3761], fig. S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 6 | 6 | 4 | 6 | 133 | 3 | 2 | 3.0 |
+-- Gibbosula
+-----------+
| +-- Pseudunio
|
--+ +-- Margaritifera_dahurica **
| +--------+
| | +-- Margaritifera_margaritifera **
+--+
| +-------- Margaritifera_falcata **
+--+
| +----- Margaritifera_middendorffi **
+--+
| +-- Margaritifera_laevis **
+--+
+-- Margaritifera_marrianae **
Bolotov et al., 2017 [Publ 3761], fig. S2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 6 | 6 | 4 | 6 | 133 | 3 | 2 | 3.0 |
+-------------- Gibbosula | +-------------- Pseudunio --+ | +-- Margaritifera_dahurica ** | +--------+ | | +-- Margaritifera_margaritifera ** +--+ | +-------- Margaritifera_falcata ** +--+ | +----- Margaritifera_middendorffi ** +--+ | +-- Margaritifera_laevis ** +--+ +-- Margaritifera_marrianae **
Vikhrev et al., 2018 [Publ 3812], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 6 | 6 | 4 | 6 | 7 | 4 | 2 | 3.0 |
+-- Cumberlandia
+-----------+
| +-- Pseudunio
|
--+ +-- Margaritifera_margaritifera **
| +--------+
| | +-- Margaritifera_dahurica **
+--+
| +-------- Margaritifera_falcata **
+--+
| +----- Margaritifera_laevis **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_marrianae **
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 6 | 6 | 4 | 6 | 70 | 3 | 2 | 3.0 |
+-------------- Pseudunio | --+ +-- Margaritifera_margaritifera ** | +--------+ | | +-- Margaritifera_dahurica ** +--+ | +-------- Margaritifera_falcata ** +--+ | +----- Margaritifera_middendorffi ** +--+ | +-- Margaritifera_laevis ** +--+ +-- Margaritifera_marrianae **
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 6 | 6 | 4 | 6 | 70 | 3 | 2 | 1.0 |
+-- Gibbosula
+-----------+
| +-- Pseudunio
|
--+ +-- Margaritifera_dahurica **
| +--------+
| | +-- Margaritifera_margaritifera **
+--+
| +-------- Margaritifera_falcata **
+--+
| +----- Margaritifera_marrianae **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_laevis **
Huff et al., 2004 [Publ 2697], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 5 | 7 | 3 | 5 | 7 | 2 | 2 | 6.0 |
+----- Lampsilis
+-----+
| | +-- Unio
| +--+
| +-- Anodonta
|
--+ +-------- Margaritifera_dahurica **
| |
| +-------- Margaritifera_margaritifera__1 **
| |
| +-------- Margaritifera_margaritifera__2 **
| |
+--+-------- Margaritifera_margaritifera__3 **
|
| +-- Pseudunio
+-----+
| +-- Cumberlandia
|
| +----- Margaritifera_falcata **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_marrianae **
Huff et al., 2004 [Publ 2697], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 5 | 6 | 2 | 2 | 2.0 |
+----------- Pseudunio | +----------- Cumberlandia --+ | +-- Margaritifera_dahurica ** | +-----+ | | +-- Margaritifera_margaritifera ** +--+ | +----- Margaritifera_falcata ** +--+ | +-- Margaritifera_middendorffi ** +--+ +-- Margaritifera_marrianae **
Huff et al., 2004 [Publ 2697], fig. 5-6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 5 | 7 | 2 | 2 | 2.0 |
+-- Pseudunio
+--------+
| +-- Cumberlandia
|
--+ +-- Margaritifera_dahurica **
| +-----+
| | +-- Margaritifera_margaritifera **
+--+
| +----- Margaritifera_falcata **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_marrianae **
Inoue et al., 2013 [Publ 3579], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 5 | 4 | 3 | 2 | 2.0 |
+-------------- Cumberlandia | --+ +----------- Pseudunio | | +--+ +-- Margaritifera_dahurica ** | +-----+ | | +-- Margaritifera_margaritifera ** +--+ | +----- Margaritifera_falcata ** +--+ | +-- Margaritifera_laevis ** +--+ +-- Margaritifera_marrianae **
Bolotov et al., 2015 [Publ 3646], fig. 2B
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 5 | 4 | 3 | 2 | 2.0 |
+-------------- Cumberlandia | --+ +----------- Pseudunio | | +--+ +-- Margaritifera_dahurica ** | +-----+ | | +-- Margaritifera_margaritifera ** +--+ | +----- Margaritifera_falcata ** +--+ | +-- Margaritifera_middendorffi ** +--+ +-- Margaritifera_laevis **
Takeuchi et al., 2015 [Publ 3661], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 5 | 5 | 3 | 5 | 5 | 3 | 2 | 2.0 |
+----------------- Pseudunio_marocanus | --+ +-------------- Pseudunio_auricularius | | +--+ +----------- Cumberlandia | | +--+ +-- Margaritifera_dahurica ** | +-----+ | | +-- Margaritifera_margaritifera ** +--+ | +----- Margaritifera_falcata ** +--+ | +-- Margaritifera_middendorffi ** +--+ +-- Margaritifera_laevis **
Huff et al., 2004 [Publ 2697], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 4 | 6 | 2 | 4 | 4 | 2 | 2 | 11.0 |
+----------- Anodonta | --+ +-------- Margaritifera_margaritifera ** | | | | +----- Cumberlandia +--+ | | | +-- Pseudunio | | | +--+ +-- Margaritifera_dahurica ** | | | +-- Margaritifera_middendorffi__1 ** | | | +-- Margaritifera_middendorffi__2 ** +--+ +-- Margaritifera_falcata__1 ** | +-- Margaritifera_falcata__2 ** | +-- Margaritifera_falcata__3 ** | +-- Margaritifera_falcata__4 **
Walker et al., 2006 [Publ 2666], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 32 | 2 | 2 | 3.0 |
+----------- Cumberlandia | --+ +-------- Pseudunio | | +--+ +-- Margaritifera_margaritifera ** | +--+ | | +-- Margaritifera_dahurica ** +--+ | +-- Margaritifera_falcata ** +--+ +-- Margaritifera_middendorffi **
Araujo et al., 2009 [Publ 2395], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 4 | 3 | 2 | 3.0 |
+----------- Cumberlandia | --+ +-------- Pseudunio | | +--+ +-- Margaritifera_margaritifera ** | +--+ | | +-- Margaritifera_dahurica ** +--+ | +-- Margaritifera_falcata ** +--+ +-- Margaritifera_middendorffi **
Wu et al., 2021 [Publ 4366], fig. 2_I
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 87 | 2 | 2 | 3.0 |
+-- Pseudunio
+-----+
| +-- Cumberlandia
|
--+ +-- Margaritifera_middendorffi **
| +--+
| | +-- Margaritifera_laevis **
+--+
| +-- Margaritifera_dahurica **
+--+
+-- Margaritifera_margaritifera **
Wu et al., 2021 [Publ 4366], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 87 | 2 | 2 | 3.0 |
+-- Cumberlandia
+-----+
| +-- Pseudunio
|
--+ +-- Margaritifera_middendorffi **
| +--+
| | +-- Margaritifera_laevis **
+--+
| +-- Margaritifera_dahurica **
+--+
+-- Margaritifera_margaritifera **
Bolotov et al., 2020 [Publ 4084], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 0 | 4 | 32 | 0 | 0 | 5.0 |
+-- Margaritifera_dahurica **
|
+-- Margaritifera_margaritifera **
+--------------------+
| +-- Margaritifera_laevis **
| |
| +-- Margaritifera_middendorffi **
|
| +-- Amblema
| |
| +-- Ortmanniana
| |
| +-- Lampsilis
| |
+--------------------+-- Cambarunio
| |
| +-- Elliptio
| |
--+ +-- Pleurobema
| |
| +-- Quadrula
|
| +-- Lamprotula
| |
| +-- Potomida
| |
| +-- Pronodularia
| +-----------------+
| | +-- Gonidea
| | |
| | +-- Leguminaia
| | |
| | +-- Microcondylaea
+--+
| +----- Middendorffinaia
| +-----------+
| | | +-- Nodularia
| | +--+
| | +-- Unio
+--+
| +-------------- Lanceolaria
| |
| | +----- Anodonta_anatina
+--+ +-----+
| | | +-- Pseudanodonta
| | +--+
| | +-- Anodonta_cygnea
+--+
| +----- Amuranodonta
| +--+
| | | +-- Anemina
| | +--+
+--+ +-- Buldowskia
|
| +----- Cristaria
+--+
| +-- Beringiana
+--+
+-- Sinanodonta
Bolotov et al., 2020 [Publ 4100], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.000 | 4 | 5 | 0 | 4 | 113 | 0 | 0 | 6.0 |
+-- Gibbosula
|
+-- Margaritifera_dahurica **
|
+-----------------------------+-- Margaritifera_margaritifera **
| |
| +-- Margaritifera_laevis **
| |
| +-- Margaritifera_middendorffi **
|
| +-- Trapezidens
| +-----------------------+
| | +-- Lamellidens
| |
| +--+ +-- Parreysia
| | | +--------------------+
| | | | +-- Leoparreysia
| | | |
| | +--+ +-- Coelatura
| | | +-----------------+
--+ | | | +-- Nitia
| | +--+
| | | +----------------- Indonaia
| | +--+
| | | +-------------- Indochinella
| | +--+
| | | +----------- Radiatula
| | +--+
| | | +-------- Scabiellus
| | +--+
| | | +----- Scabies
| | +--+
| | | +-- Unionetta
| | +--+
| | +-- Harmandia
| |
+--+ +-- Trapezoideus
| +--+
| | +-- Physunio
| +-----------+
| | | +-- Yaukthwa
| | +--+
| +--+ +-- Lens
| | |
| | | +-------------- Ensidens
| | | |
| | +--+ +----------- Hyriopsis_desowitzi
| | | |
| | +--+ +-------- Rectidens
| | | |
| | +--+ +-- Hyriopsis_khoratensis
| | | +--+
| | | | +-- Hyriopsis_bialata
| | +--+
| | | +-- Hyriopsis_myersiana
| | +--+
| | +-- Hyriopsis_panhai
| |
+--------+ +-------------- Indopseudodon
| |
| +--+ +----------- Bineurus
| | | |
| | +--+ +-------- Nyeinchanconcha
| | | |
| | +--+ +-- Thaiconcha
| | | +--+
| | | | +-- Pilsbryoconcha
| | +--+
| | | +-- Pseudodon
| | +--+
| | +-- Sundadontina
| |
| | +-- Sinosolenaia_iridinea
| | |
+--+ +-- Sinosolenaia_recognita__1
| |
| +-- Sinosolenaia_carinata
| |
| +-- Leguminaia
| |
| +-- Microcondylaea
| |
| +-- Gonidea
| |
| +-- Ptychorhynchus
| |
| +-- Parvasolenaia
| |
+--------------+-- Lamprotula_leaii
|
+-- Lamprotula
|
+-- Lamprotula_caveata
|
+-- Potomida
|
+-- Pronodularia
|
+-- Sinohyriopsis
|
+-- Chamberlainia
|
+-- Sinosolenaia_recognita__2
|
+-- Sinosolenaia_oleivora
Huang et al., 2018 [Publ 3828], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 39 | 2 | 2 | 4.0 |
+----------- Cumberlandia --+ | +-------- Pseudunio +--+ | +----- Margaritifera_falcata ** +--+ | +-- Margaritifera_margaritifera ** +--+ +-- Margaritifera_dahurica **
Huang et al., 2019 [Publ 3843], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 53 | 2 | 2 | 4.0 |
+----------- Cumberlandia --+ | +-------- Pseudunio +--+ | +----- Margaritifera_falcata ** +--+ | +-- Margaritifera_margaritifera ** +--+ +-- Margaritifera_dahurica **
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 7 | 2 | 2 | 4.0 |
+----------- Cumberlandia --+ | +-------- Pseudunio +--+ | +----- Margaritifera_falcata ** +--+ | +-- Margaritifera_margaritifera ** +--+ +-- Margaritifera_dahurica **
Guerra et al., 2019 [Publ 4168], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 38 | 3 | 2 | 4.0 |
+----------- Cumberlandia --+ | +-------- Pseudunio +--+ | +----- Margaritifera_falcata ** +--+ | +-- Margaritifera_dahurica ** +--+ +-- Margaritifera_margaritifera **
Guerra et al., 2019 [Publ 4168], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 38 | 3 | 2 | 4.0 |
+----------- Cumberlandia --+ | +-------- Pseudunio +--+ | +----- Margaritifera_falcata ** +--+ | +-- Margaritifera_dahurica ** +--+ +-- Margaritifera_margaritifera **
Wu et al., 2021 [Publ 4366], fig. 3-BI
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 70 | 2 | 2 | 4.0 |
+----------- Cumberlandia --+ | +-------- Pseudunio +--+ | +----- Margaritifera_falcata ** +--+ | +-- Margaritifera_margaritifera ** +--+ +-- Margaritifera_dahurica **
Wu et al., 2021 [Publ 4366], fig. 3-ML
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 70 | 2 | 2 | 4.0 |
+----------- Cumberlandia --+ | +-------- Pseudunio +--+ | +----- Margaritifera_falcata ** +--+ | +-- Margaritifera_margaritifera ** +--+ +-- Margaritifera_dahurica **
Wu et al., 2021 [Publ 4366], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 65 | 2 | 2 | 4.0 |
+----------- Cumberlandia --+ | +-------- Pseudunio +--+ | +----- Margaritifera_falcata ** +--+ | +-- Margaritifera_margaritifera ** +--+ +-- Margaritifera_dahurica **
Wu et al., 2022 [Publ 4238], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 70 | 2 | 2 | 4.0 |
+----------- Cumberlandia --+ | +-------- Pseudunio +--+ | +----- Margaritifera_falcata ** +--+ | +-- Margaritifera_dahurica ** +--+ +-- Margaritifera_margaritifera **
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 92 | 2 | 2 | 4.0 |
+----------- Cumberlandia --+ | +-------- Pseudunio +--+ | +----- Margaritifera_falcata ** +--+ | +-- Margaritifera_dahurica ** +--+ +-- Margaritifera_margaritifera **
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 92 | 2 | 2 | 4.0 |
+----------- Cumberlandia --+ | +-------- Pseudunio +--+ | +----- Margaritifera_falcata ** +--+ | +-- Margaritifera_dahurica ** +--+ +-- Margaritifera_margaritifera **
Dai et al., 2025 [Publ 4440], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 84 | 2 | 2 | 4.0 |
+----------- Cumberlandia --+ | +-------- Pseudunio +--+ | +----- Margaritifera_falcata ** +--+ | +-- Margaritifera_dahurica ** +--+ +-- Margaritifera_margaritifera **
Chen et al., 2025 [Publ 4448], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 82 | 2 | 2 | 4.0 |
+----------- Cumberlandia --+ | +-------- Pseudunio +--+ | +----- Margaritifera_falcata ** +--+ | +-- Margaritifera_dahurica ** +--+ +-- Margaritifera_margaritifera **
Breton et al., 2011 [Publ 3376], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 35 | 1 | 1 | 4.0 |
+-------- Cumberlandia --+ | +----- Margaritifera_margaritifera ** +--+ | +-- Margaritifera_falcata ** +--+ +-- Margaritifera_marrianae **
Chase et al., 2018 [Publ 3847], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 36 | 1 | 1 | 6.0 |
+-------- Cumberlandia --+ | +----- Margaritifera_falcata ** +--+ | +-- Margaritifera_marrianae ** +--+ +-- Margaritifera_margaritifera **
Mock et al., 2013 [Publ 3598], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 0 | 0 | 0 | 4.0 |
+----- Margaritifera_margaritifera ** --+ | +-- Margaritifera_falcata ** +--+ +-- Margaritifera_laevis **
Stewart et al., 2021 [Publ 4451], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 13 | 0 | 0 | 4.0 |
+----- Margaritifera_margaritifera **
+--------+
| | +-- Margaritifera_falcata **
| +--+
| +-- Margaritifera_marrianae **
--+
| +----------- Toxolasma
| |
| | +-- Alasmidonta
+--+ +-----+
| | +-- Pterosyna
| |
+--+ +-- Strophitus
| +--+
| | +-- Platynaias
+--+
| +-- Pyganodon
+--+
+-- Utterbackia
Rosenberg et al., 1994 [Publ 2719], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.667 | 2 | 2 | 0 | 2 | 17 | 1 | 1 | -- |
+----------- Cumberlandia | | +-- Margaritifera_falcata ** +--------+ | +-- Margaritifera_margaritifera ** | --+ +-------- Gonidea | | | | +-- Elliptio | | | | | +-- Fusconaia | | | +--+ +--+-- Plectomerus | | | | | +-- Pleurobema | | | | | +-- Unio | | | | +-- Pyganodon_cataracta +--+ | | +-- Pyganodon_grandis | | | +-- Utterbackia | | | +-- Lampsilis_teres | | | +-- Lampsilis_straminea +--+ +-- Megalonaias | +-- Obliquaria | +-- Theliderma | +-- Quadrula | +-- Uniomerus
Lydeard et al., 1996 [Publ 2656], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 25 | 1 | 1 | -- |
+----- Cumberlandia --+ | +-- Margaritifera_margaritifera ** +--+ +-- Margaritifera_falcata **
Lydeard et al., 1996 [Publ 2656], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 26 | 1 | 1 | -- |
+----- Cumberlandia --+ | +-- Margaritifera_margaritifera ** +--+ +-- Margaritifera_falcata **
Rosenberg et al., 1997 [Publ 2720], fig. 2A
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.667 | 2 | 2 | 0 | 2 | 20 | 1 | 1 | -- |
+----------- Cumberlandia | +----------- None__2 | | +-- Margaritifera_falcata ** +--------+ --+ +-- Margaritifera_margaritifera ** | | +-------- None__1 | | | | +----- Anodonta | | | | | | +-- Gonidea +--+ | | | | +-- Pyganodon_grandis | | | | | +-- Utterbackia | | | | | +-- Pyganodon_cataracta +--+ | | +-- Lampsilis_straminea | | | +-- Lampsilis_teres | | | +-- Megalonaias | | | +-- Obliquaria | | +--+-- Theliderma | +-- Quadrula | +-- Uniomerus | +-- Amblema | +-- Elliptio | +-- Fusconaia | +-- Plectomerus | +-- Pleurobema | +-- Unio
Rosenberg et al., 1997 [Publ 2720], fig. 2B
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.667 | 2 | 2 | 0 | 2 | 20 | 1 | 1 | -- |
+-------- Cumberlandia | +-------- None__1 | +-------- None__2 | | +-- Margaritifera_falcata ** +-----+ | +-- Margaritifera_margaritifera ** --+ | +----- Anodonta | | | | +-- Gonidea | | | | | +-- Pyganodon_grandis | | | | | +-- Utterbackia | | | | | +-- Pyganodon_cataracta +--+ | | +-- Lampsilis_straminea | | | +-- Lampsilis_teres | | | +-- Megalonaias | | | +-- Obliquaria | | +--+-- Theliderma | +-- Quadrula | +-- Uniomerus | +-- Amblema | +-- Elliptio | +-- Fusconaia | +-- Plectomerus | +-- Scriptolamprotula | +-- Unio
Walker et al., 2006 [Publ 2666], fig. 6-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 12 | 1 | 1 | -- |
+----- Cumberlandia | --+ +-- Margaritifera_margaritifera ** | | +--+-- Margaritifera_dahurica__1 ** | +-- Margaritifera_dahurica__2 **
Lopes-Lima et al., 2017 [Publ 3792], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 30 | 1 | 1 | -- |
+----- Pseudunio --+ | +-- Margaritifera_falcata ** +--+ +-- Margaritifera_dahurica **
Aunins et al., 2018 [Publ 3863], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 38 | 1 | 1 | -- |
+----- Cumberlandia --+ | +-- Margaritifera_falcata ** +--+ +-- Margaritifera_dahurica **
Cha et al., 2018 [Publ 3916], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 36 | 1 | 1 | -- |
+----- Pseudunio --+ | +-- Margaritifera_falcata ** +--+ +-- Margaritifera_dahurica **
Wu et al., 2019 [Publ 3864], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 44 | 1 | 1 | -- |
+----- Cumberlandia --+ | +-- Margaritifera_dahurica ** +--+ +-- Margaritifera_falcata **
Froufe et al., 2019 [Publ 3936], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 42 | 1 | 1 | -- |
+----- Pseudunio --+ | +-- Margaritifera_dahurica ** +--+ +-- Margaritifera_falcata **
Froufe et al., 2019 [Publ 3936], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 42 | 1 | 1 | -- |
+----- Pseudunio --+ | +-- Margaritifera_dahurica ** +--+ +-- Margaritifera_falcata **
Wu et al., 2020 [Publ 4121], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 46 | 1 | 1 | -- |
+----- Cumberlandia --+ | +-- Margaritifera_falcata ** +--+ +-- Margaritifera_dahurica **
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 46 | 1 | 1 | -- |
+----- Pseudunio --+ | +-- Margaritifera_falcata ** +--+ +-- Margaritifera_dahurica **
Krebs et al., 2003 [Publ 2691], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 31 | 0 | 0 | -- |
+-- Margaritifera_margaritifera **
+-----------------------------+
| +-- Margaritifera_falcata **
|
| +----------- Utterbackiana
| |
| +----------- Platynaias
| |
| +----------- Alasmidonta
| +-----------------+
--+ | | +-- Pterosyna
| | +--------+
| | | +-- Lasmigona
| | |
| | | +-------- Strophitus
| | +--+
| | | +----- Utterbackia
| | +--+
| | | +-- Pyganodon
+--+ +--+
| +-- Anodonta
|
| +-------- Megalonaias
| +-----------------+
| | | +----- Quadrula_quadrula__2
| | +--+
| | | +-- Quadrula_quadrula__1
| | +--+
| | +-- Theliderma
| |
+--+ +----------- Elliptoideus
| +-----------+
| | | +-------- Pleurobema
| | +--+
| | | +----- Uniomerus
| | +--+
| | | +-- Fusconaia
+--+ +--+
| +-- Eurynia
|
| +-------------------- Amblema
| |
| | +----- Toxolasma
| | +-----------+
+--+ | | +-- Plectomerus
| | +--+
| | +-- Obliquaria
| |
+--+ +----- Ptychobranchus
| +--------+
| | | +-- Potamilus_fragilis
| | +--+
| | +-- Medionidus
+--+
| +-- Villosa
| +--------+
| | +-- Ellipsaria
+--+
| +-------- Lampsilis_teres
+--+
| +----- Lampsilis_siliquoidea
+--+
| +-- Potamilus_purpuratus
+--+
+-- Potamilus_alatus
Fonseca et al., 2016 [Publ 3775], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 29 | 0 | 0 | -- |
+-- Margaritifera_falcata **
+-----------------------+
| +-- Margaritifera_dahurica **
|
| +-------- Quadrula
| +--------------+
| | | +----- Toxolasma
--+ | +--+
| | | +-- Lampsilis
| | +--+
| | +-- Venustaconcha
| |
| | +----------- Sinohyriopsis
+--+ +--------+
| | | +-------- Sinosolenaia
| | +--+
| | | +----- Potomida
| | +--+
| | | +-- Lamprotula
| | +--+
| | +-- Pronodularia
+--+
| +-------------- Lanceolaria
| +--+
| | | +----------- Scriptolamprotula
| | +--+
| | | +-------- Unio
| | +--+
| | | +----- Schistodesmus
| | +--+
+--+ | +-- Nodularia
| +--+
| +-- Tchangsinaia
|
| +----- Cristaria
| +-----+
| | | +-- Anemina
| | +--+
+-----+ +-- Sinanodonta
|
| +-------- Anodonta
+--+
| +----- Platynaias
+--+
| +-- Pyganodon
+--+
+-- Utterbackia
Feng et al., 2016 [Publ 3800], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 12 | 0 | 0 | -- |
+-- Margaritifera_dahurica **
+--------------+
| +-- Margaritifera_falcata **
|
| +-- Anodonta
--+ +-----------+
| | +-- Utterbackia
| |
| | +-------- Tchangsinaia
+--+ +--+
| | | +----- Pyganodon
| | +--+
| | | +-- Anemina
+--+ +--+
| +-- Sinanodonta
|
| +-------- Sinosolenaia
+--+
| +----- Quadrula
+--+
| +-- Potamilus
+--+
+-- Lampsilis
Amaro et al., 2017 [Publ 3791], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.000 | 2 | 15 | 15 | 2 | 14 | 0 | 0 | 20.0 |
+----------------------- Neotrigonia | | +-------------------- Margaritifera_margaritifera__2 ** --+ | | | +-- Margaritifera_margaritifera__1 ** | | +--------------+ +--+ | +-- Margaritifera_falcata ** | | | | +-- Lasmigona | | +--+ +--+ | +-- Platynaias | +--------+ | | | +-- Pyganodon | | +--+ | | +-- Utterbackia +--+ | +----- Fusconaia | +-----+ | | | +-- Pronodularia | | +--+ +--+ +-- Sinosolenaia | | +-------- Quadrula +--+ | +----- Toxolasma +--+ | +-- Venustaconcha +--+ +-- Lampsilis
Amaro et al., 2017 [Publ 3791], fig. 3A
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 10 | 0 | 0 | -- |
+-- Margaritifera_falcata **
+-----------+
| +-- Margaritifera_margaritifera **
|
| +----- Platynaias
--+ +-----+
| | | +-- Pyganodon
| | +--+
| | +-- Utterbackia
+--+
| +-- Pronodularia
| +-----+
| | +-- Sinosolenaia
+--+
| +----- Quadrula
+--+
| +-- Toxolasma
+--+
+-- Venustaconcha
Amaro et al., 2017 [Publ 3791], fig. 3B
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 10 | 0 | 0 | -- |
+-- Margaritifera_falcata **
+-----------+
| +-- Margaritifera_margaritifera **
|
| +----- Platynaias
--+ +-----+
| | | +-- Pyganodon
| | +--+
| | +-- Utterbackia
+--+
| +-- Toxolasma
| +-----+
| | +-- Venustaconcha
+--+
| +----- Quadrula
+--+
| +-- Pronodularia
+--+
+-- Sinosolenaia
Wen et al., 2017 [Publ 3795], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 14 | 0 | 0 | -- |
+-- Margaritifera_dahurica **
+--------------+
| +-- Margaritifera_falcata **
|
| +----------- Quadrula
| +--+
--+ | | +-------- Toxolasma
| | +--+
| | | +----- Potamilus
| | +--+
| | | +-- Lampsilis
+--+ +--+
| +-- Venustaconcha
|
| +----------- Lamprotula
| |
| | +----- Anodonta
+--+ +--+
| | | +-- Sinanodonta
| | +--+
+--+ +-- Cristaria
|
| +----- Scriptolamprotula
+--+
| +-- Lepidodesma
+--+
+-- Unio
Sano et al., 2017 [Publ 3799], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 2 | 2 | 19 | 0 | 0 | -- |
+-------- Margaritifera_laevis__2 **
+--------+
| | +----- Margaritifera_laevis__1 **
| +--+
| | +-- Margaritifera_middendorffi **
| +--+
| +-- Margaritifera_laevis__3 **
|
--+ +-------- Obovalis
| +-----+
| | | +----- Inversidens
| | +--+
| | | +-- Pronodularia
| | +--+
| | +-- Sinohyriopsis
+--+
| +----- Lanceolaria
| +-----+
| | | +-- Nodularia
| | +--+
| | +-- Inversiunio
+--+
| +-- Cristaria
| +-----+
| | +-- Buldowskia
+--+
| +----- Pletholophus
+--+
| +-- Sinanodonta
+--+
+-- Beringiana
Sano et al., 2017 [Publ 3799], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 31 | 0 | 0 | -- |
+-- Margaritifera_laevis **
+-----------------+
| +-- Margaritifera_middendorffi **
|
| +-- Obovalis
| +--------+
| | +-- Ptychorhynchus
| |
| +-----+ +-- Inversidens
--+ | | +-----+
| | | | +-- Sinosolenaia
| | +--+
| | | +----- Sinohyriopsis
| | +--+
| | | +-- Pronodularia
| | +--+
| | +-- Lamprotula
+--+
| +-- Cristaria
| +-----+
| | +-- Buldowskia
| +-----+
| | | +----- Beringiana
| | +--+
| | | +-- Pletholophus
| | +--+
+--+ +-- Sinanodonta
|
| +-- Tchangsinaia
| +--------+
| | +-- Nodularia
| |
+--+ +-- Schistodesmus
| +-----+
| | +-- Acuticosta
+--+
| +----- Inversiunio
+--+
| +-- Lepidodesma
+--+
+-- Lanceolaria
Konopleva et al., 2017 [Publ 3807], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 29 | 0 | 0 | -- |
+-- Margaritifera_dahurica **
+-----------------+
| +-- Margaritifera_laevis **
|
| +-- Trapezidens
| +-----------+
| | +-- Lamellidens
--+ +--+
| | | +----------- Leoparreysia
| | +--+
| | | +-------- Coelatura
| | +--+
| | | +----- Indochinella
| | +--+
+--+ | +-- Radiatula
| +--+
| +-- Scabies
|
| +----- Ensidens
| +-----+
| | | +-- Lens
| | +--+
+-----+ +-- Physunio
|
| +-------- Sundadontina
| |
+--+ +-- Nyeinchanconcha
| +--+
| | +-- Bineurus
+--+
| +-- Pilsbryoconcha
+--+
+-- Pseudodon
Wu et al., 2018 [Publ 3836], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 51 | 0 | 0 | -- |
+-- Margaritifera_dahurica **
+-----------------------------+
| +-- Margaritifera_falcata **
|
| +----------------------------- Quadrula
| |
| | +-- Amblema
| +--------------------------+
--+ | +-- Obovaria
| |
| | +-------- Pleurobema
| +--------------------+
| | | +----- Fusconaia
| | +--+
| | | +-- Elliptio
| | +--+
+--+ +-- Eurynia
|
| +-- Potomida
| +--------+
| | +-- Lamprotula
| +--------------+
| | | +-------- Sinosolenaia_carinata
| | | |
| | +--+ +----- Gonidea
| | | |
| | +--+----- Sinosolenaia_oleivora
| | |
| | | +-- Parvasolenaia
+--+ +--+
| +-- Ptychorhynchus
|
| +----------------------- Sinohyriopsis
| |
| | +-------------------- Lepidodesma
| | |
| | | +----------- Scriptolamprotula
+--+ | |
| +--------+ +-------- Unio
| | | |
| | +--+ +-- Nodularia
| | | +--+
| | | | +-- Schistodesmus
+--+ +--+
| | +-- Tchangsinaia
| +--+
| +-- Cuneopsis
|
| +-- Acuticosta
| +--------------+
| | +-- Lanceolaria
| |
| | +-- Anemina
+--+ |
| +-----------+-- Cristaria
| | |
| | +-- Sinanodonta
+--+
| +----------- Anodonta_cygnea
| |
+--+ +-------- Anodonta_anatina
| |
+--+ +----- Strophitus
| |
+--+ +-- Platynaias
| |
+--+-- Alasmidonta
|
+-- Pyganodon
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 55 | 0 | 0 | -- |
+-- Margaritifera_margaritifera **
+-----------------------------+
| +-- Margaritifera_hembeli **
|
| +-- Quadrula
| +--------------------+
| | +-- Uniomerus
| |
| | +----- Elliptoideus
| +-----+ +--------------+
| | | | | +-- Elliptio
| | | | +--+
| | | | +-- Pleurobema
| | | |
| | +--+ +----- Reginaia
| | | +-----------+
| | | | | +-- Plectomerus
--+ | | | +--+
| | | | +-- Amblema
| | +--+
| | | +-- Psoronaias
| | | +-----------+
| | | | +-- Popenaias
| | +--+
| | | +----------- Obliquaria
| | +--+
| | | +-------- Cyrtonaias
| | +--+
| | | +----- Leaunio
| | +--+
| | | +-- Lampsilis
| | +--+
| | +-- Pachynaias
| |
+--+ +-- Chamberlainia
| +--------------------+
| | +-- Sinohyriopsis
| |
| | +----------------- Lamprotula
| | +--+
| | | | +-------------- Gonidea
| +--+ | +--+
| | | | | +----------- Indopseudodon
| | | | +--+
| | | | | +-------- Bineurus
| | | | +--+
| | | | | +----- Pilsbryoconcha
| | | | +--+
| | +--+ | +-- Pseudodon
| | | +--+
| | | +-- Sundadontina
| | |
| | | +----- Ensidens
| | | +-----+
| | | | | +-- Hyriopsis
| | | | +--+
| | | | +-- Rectidens
| | +--------+
| | | +-- Trapezoideus
| | | +-----+
+--+ | | +-- Physunio
| +--+
| | +----- Yaukthwa
| +--+
| | +-- Lens
| +--+
| +-- Solenaia
|
| +----------- Lanceolaria
| |
| | +----- Unio
| +--------+ +--+
| | | | | +-- Nodularia
| | | | +--+
| | | | +-- Tchangsinaia
| | +--+
| | | +-- Cristaria
| | | +--+
| | | | +-- Sinanodonta
+-----+ +--+
| | +-- Anodonta
| +--+
| +-- Pyganodon
|
| +-- Trapezidens
| +--------------+
| | +-- Lamellidens
+--+
| +-------------- Leoparreysia
| |
+--+ +-- Coelatura
| +--------+
| | +-- Prisodontopsis
+--+
| +-------- Indonaia
+--+
| +----- Indochinella
+--+
| +-- Scabies
+--+
+-- Harmandia
Pfeiffer et al., 2019 [Publ 3937], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 55 | 0 | 0 | -- |
+----- Hyridella --+ | +-- Margaritifera_margaritifera ** +--+ +-- Margaritifera_hembeli **
Lopes-Lima et al., 2020 [Publ 4080], fig. 10
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 12 | 0 | 0 | -- |
+-- Margaritifera_middendorffi **
+--+
| +-- Margaritifera_laevis **
--+
| +-- Beringiana
+--+
+-- Sinanodonta
Lopes-Lima et al., 2020 [Publ 4080], fig. 11
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 10 | 0 | 0 | -- |
+-- Margaritifera_laevis **
+--+
| +-- Margaritifera_middendorffi **
--+
| +-- Nodularia
+--+
+-- Lanceolaria
Lopes-Lima et al., 2020 [Publ 4080], fig. 12
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 6 | 0 | 0 | -- |
+-- Margaritifera_laevis **
+--+
| +-- Margaritifera_middendorffi **
--+
| +-- Nodularia
+--+
+-- Inversiunio
Lopes-Lima et al., 2020 [Publ 4080], fig. 13
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 8 | 0 | 0 | -- |
+-- Margaritifera_laevis **
+--+
| +-- Margaritifera_middendorffi **
--+
| +-- Middendorffinaia
+--+
+-- Nodularia
Lopes-Lima et al., 2020 [Publ 4080], fig. 14
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 4 | 0 | 0 | -- |
+-- Margaritifera_laevis **
+-----+
| +-- Margaritifera_middendorffi **
--+
| +----- Lamprotula
+--+
| +-- Parvasolenaia
+--+
+-- Koreosolenaia
Lopes-Lima et al., 2020 [Publ 4080], fig. 16
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 6 | 0 | 0 | -- |
+-- Margaritifera_laevis **
+--------+
| +-- Margaritifera_middendorffi **
--+
| +-------- Chamberlainia
+--+
| +----- Sinohyriopsis
+--+
| +-- Pronodularia
+--+
+-- Lamprotula
Lopes-Lima et al., 2020 [Publ 4080], fig. 17
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 4 | 0 | 0 | -- |
+-- Margaritifera_laevis **
+--+
| +-- Margaritifera_middendorffi **
--+
| +-- Chamberlainia
+--+
+-- Sinohyriopsis
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 120 | 0 | 0 | -- |
+-- Margaritifera_laevis **
+--------------------------------+
| +-- Margaritifera_middendorffi **
|
| +-- Lamproscapha
| +-----+
| | +-- Mycetopoda
| +-----------------------+
| | | +----- Etheria
| | +--+
| | | +-- Aspatharia
--+ | +--+
| | +-- Chambardia
| |
| | +-- Coelatura
| | |
| | +--------------------------+-- Leoparreysia
| | | |
| | | +-- Radiatula
| | |
| | | +-------------------- Acuticosta
| | | |
+--+ | +-------------------- Lepidodesma
| | |
| | | +----------------- Scriptolamprotula
| | | |
| | +--+ +-------------- Middendorffinaia
| | | | |
| | | +--+ +----------- Unio
| | | | |
| | | +--+ +-------- Nodularia
| | | | |
| | | +--+ +----- Schistodesmus
| | +-----+ | |
| | | | +--+----- Cuneopsis
| | | | |
| | | | | +-- Tchangsinaia
+--+ | | +--+
| | | +-- Inversiunio
| | |
| | | +-------------- Lanceolaria
| | | |
| | | | +-------- Cristaria
| | | | |
| | | | | +----- Anemina
| | | | +--+
| | +-----+ +--+ | +-- Amuranodonta
| | | | | +--+
| | | | | +-- Buldowskia
| | | | |
| | | | | +----- Pletholophus
| | | | +--+
| | | | | +-- Beringiana
| | +--+ +--+
| | | +-- Sinanodonta
| | |
| | | +-------- Pseudanodonta
| | | |
| | | | +----- Anodonta_cygnea
+--+ | +--+
| | | | +-- Anodonta_anatina
| +--+ +--+
| | +-- Brachyanodon
| |
| | +----- Pyganodon
| | |
| | +----- Simpsonaias
| +--+
| +----- Strophitus
| |
| | +-- Alasmidonta
| +--+
| +-- Lasmigona
|
| +-- Lens
| |
| +-- Ensidens
| |
| +--------------------+-- Physunio
| | |
| | +-- Rectidens
| | |
| | +-- Yaukthwa
| |
| | +-- Amblema
| | |
+--+ +-- Elliptio
| +-----------------+
| | +-- Lampsilis
| | |
| | +-- Quadrula
| |
| | +-- Chamberlainia
| | +-----+
| | | +-- Sinohyriopsis
+--+ +--------+
| | | +----- Pronodularia
| | +--+
| | | +-- Potomida
| | +--+
| | +-- Lamprotula
| |
| | +----------- Indopseudodon
+--+ |
| +--+ +-------- Pilsbryoconcha
| | | |
| | +--+ +----- Pseudodon
| | | |
| | +--+ +-- Nyeinchanconcha
| | | |
| | +--+-- Bineurus
+--+ |
| +-- Sundadontina
|
| +-------- Sinosolenaia
| +--+
| | | +----- Gonidea
| | +--+
| | | +-- Leguminaia
+--+ +--+
| +-- Microcondylaea
|
| +-------- Inversidens
| |
+--+ +-- Obovalis
| +--+
| | +-- Ptychorhynchus
+--+
| +-- Parvasolenaia
+--+
+-- Koreosolenaia
Lopes-Lima et al., 2020 [Publ 4080], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 7 | 0 | 0 | -- |
+-- Margaritifera_laevis **
+--+
| +-- Margaritifera_middendorffi **
--+
| +-- Nodularia
+--+
+-- Scriptolamprotula
Lopes-Lima et al., 2020 [Publ 4080], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 9 | 0 | 0 | -- |
+-- Margaritifera_laevis **
+--------+
| +-- Margaritifera_middendorffi **
--+
| +-------- Sinanodonta
+--+
| +----- Amuranodonta
+--+
| +-- Anemina
+--+
+-- Buldowskia
Lopes-Lima et al., 2020 [Publ 4080], fig. 7
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 6 | 0 | 0 | -- |
+-- Margaritifera_laevis **
+--+
| +-- Margaritifera_dahurica **
--+
| +-- Sinanodonta
+--+
+-- Beringiana
Lopes-Lima et al., 2020 [Publ 4080], fig. 8
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 5 | 0 | 0 | -- |
+-- Margaritifera_laevis **
+--+
| +-- Margaritifera_middendorffi **
--+
| +-- Sinanodonta
+--+
+-- Cristaria
Lopes-Lima et al., 2020 [Publ 4080], fig. 9
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 4 | 0 | 0 | -- |
+-- Margaritifera_laevis **
+-----+
| +-- Margaritifera_middendorffi **
--+
| +----- Cristaria
+--+
| +-- Sinanodonta
+--+
+-- Pletholophus
Sano et al., 2020 [Publ 4138], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 32 | 0 | 0 | -- |
+----- Gibbosula --+ | +-- Margaritifera_middendorffi ** +--+ +-- Margaritifera_laevis **
Wu et al., 2020 [Publ 4147], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 12 | 0 | 0 | -- |
+----- Gibbosula --+ | +-- Margaritifera_margaritifera ** +--+ +-- Margaritifera_falcata **
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 35 | 0 | 0 | -- |
+-- Margaritifera_dahurica **
+-----------------------------+
| +-- Margaritifera_falcata **
|
| +-------------- Quadrula
| |
| +--------------+ +----------- Toxolasma
| | | |
| | +--+ +-------- Potamilus
--+ | | |
| | +--+ +-- Lampsilis_cardium
| | | +--+
| | | | +-- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_siliquoidea
| | +--+
| | +-- Venustaconcha
| |
+--+ +-- Chamberlainia
| +-----------+
| | +-- Sinohyriopsis
| |
| +-----------+ +-- Lamprotula
| | | +--------+
| | | | +-- Potomida
| | | |
| | +--+ +-- Pseudodon
| | | +-----+
| | | | +-- Pilsbryoconcha
| | +--+
+--+ | +----- Parvasolenaia
| +--+
| | +-- Microcondylaea
| +--+
| +-- Sinosolenaia
|
| +----------------------- Lamellidens
| |
| | +-------------------- Lepidodesma
+--+ |
| | +----------- Scriptolamprotula
| | +-----+
+--+ | | +-------- Unio
| | +--+
| | | +----- Schistodesmus
| | +--+
+--+ | +-- Nodularia
| +--+
| +-- Tchangsinaia
|
| +-------------- Lanceolaria
| |
+--+ +----------- Acuticosta
| |
+--+ +-- Anodonta
| +-----+
| | +-- Pyganodon
+--+
| +----- Cristaria
+--+
| +-- Sinanodonta
+--+
+-- Anemina
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 37 | 0 | 0 | -- |
+-- Margaritifera_dahurica **
+-----------------------+
| +-- Margaritifera_falcata **
|
| +-------------------- Lamellidens
| |
| | +-- Anodonta
| | +-----+
| | | +-- Pyganodon
| +--+ +--------+
| | | | | +----- Cristaria
| | | | +--+
| | | | | +-- Anemina
--+ | | | +--+
| | +--+ +-- Sinanodonta
| | |
| | | +-- Acuticosta
| | | +-----------+
| | | | +-- Lanceolaria
| | | |
| | +--+ +-- Lepidodesma
| | | +--------+
| | | | +-- Scriptolamprotula
| | +--+
| | | +-------- Unio
+--+ +--+
| | +----- Schistodesmus
| +--+
| | +-- Nodularia
| +--+
| +-- Tchangsinaia
|
| +-------------- Quadrula
| |
| +--+ +----------- Toxolasma
| | | |
| | +--+ +-------- Potamilus
| | | |
| | +--+ +-- Lampsilis_cardium
| | | +--+
| | | | +-- Lampsilis_ornata
| | +--+
+-----+ | +-- Lampsilis_siliquoidea
| +--+
| +-- Venustaconcha
|
| +-- Chamberlainia
| +-----------+
| | +-- Sinohyriopsis
| |
| | +----- Sinosolenaia
+--+ +-----+
| | | +-- Parvasolenaia
| | +--+
| | +-- Microcondylaea
+--+
| +-- Pseudodon
| +-----+
| | +-- Pilsbryoconcha
+--+
| +----- Potomida
+--+
| +-- Lamprotula
+--+
+-- Pronodularia
Wu et al., 2023 [Publ 4339], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 17 | 0 | 0 | -- |
+-- Margaritifera_margaritifera **
+-----------------+
| +-- Margaritifera_dahurica **
|
| +-- Elliptio
| +--------+
| | +-- Quadrula
--+ +-----+
| | | +-------- Obliquaria
| | +--+
| | | +----- Truncilla
| | +--+
| | | +-- Lampsilis
+--+ +--+
| +-- Potamilus
|
| +-------------- Lepidodesma
| |
| | +----- Anemina
+--+ +-----+
| | | +-- Cristaria
| | +--+
+--+ +-- Sinanodonta
|
| +-------- Platynaias
+--+
| +----- Anodonta_anatina
+--+
| +-- Pseudanodonta
+--+
+-- Anodonta_cygnea
Wu et al., 2024 [Publ 4350], fig. 5a
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 34 | 0 | 0 | -- |
+-- Margaritifera_dahurica **
+--------------------------+
| +-- Margaritifera_margaritifera **
|
| +-------------------------- Chamberlainia
| |
--+ | +----------- Khairuloconcha
| | |
| | | +----- Ensidens
| | +-----------+ +--+
| | | | | | +-- Rectidens
+--+ | | | +--+
| | | | +-- Hyriopsis
| | +--+
| | | +-- Physunio
| | | +--+
| | | | +-- Trapezoideus
| | +--+
| | | +-- Lens
+--+ +--+
| +-- Yaukthwa
|
| +-------------- Schepmania
| +-----+
| | | +----------- Indopseudodon
| | +--+
| | | +-------- Pilsbryoconcha
| | +--+
| | | +----- Pseudodon
| | +--+
+--+ | +-- Thaiconcha
| +--+
| +-- Bineurus
|
| +----- Inversidens
| +-----------+
| | | +-- Potomida
| | +--+
+--+ +-- Lamprotula
|
| +-------------- Sinosolenaia
| |
+--+ +-- Gonidea
| +--------+
| | +-- Microcondylaea
+--+
| +-------- Parvasolenaia
+--+
| +----- Koreosolenaia
+--+
| +-- Postolata
+--+
+-- Ptychorhynchus
Wu et al., 2024 [Publ 4350], fig. 5b
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 34 | 0 | 0 | -- |
+-- Margaritifera_dahurica **
+--------------------------------+
| +-- Margaritifera_margaritifera **
|
| +-------------------------------- Schepmania
| |
--+ | +----------- Indopseudodon
| | +-----------------+
| | | | +-------- Pilsbryoconcha
| | | +--+
| | | | +----- Bineurus
+--+ | +--+
| | | +-- Pseudodon
| | +--+
| | +-- Thaiconcha
| |
| | +----------------------- Hyriopsis_bialata
| | |
+--+ +--+ +-- Rectidens
| | | +-----------------+
| | | | +-- Khairuloconcha
| | +--+
| | | +----------------- Ensidens
| | | |
| | +--+ +-------------- Hyriopsis_desowitzi
| | | |
| | +--+ +----------- Yaukthwa_elongatula
| | | |
| | +--+ +-------- Trapezoideus
+--+ | |
| +--+ +-- Physunio
| | +--+
| | | +-- Yaukthwa_inlenensis
| +--+
| | +-- Yaukthwa_paiensis
| +--+
| +-- Lens
|
| +-------------------- Inversidens
| |
| | +----- Lamprotula
+-----+ +-----------+
| | | +-- Chamberlainia
| | +--+
+--+ +-- Potomida
|
| +-------------- Sinosolenaia
| |
+--+ +-- Gonidea
| +--------+
| | +-- Microcondylaea
+--+
| +-------- Koreosolenaia
+--+
| +----- Parvasolenaia
+--+
| +-- Postolata
+--+
+-- Ptychorhynchus
Wu et al., 2024 [Publ 4350], fig. 5c
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 34 | 0 | 0 | -- |
+-- Margaritifera_dahurica **
+-----------------------------+
| +-- Margaritifera_margaritifera **
|
| +----------------------------- Schepmania_nieuwenhuisi__1
| |
| +----------------------------- Schepmania_nieuwenhuisi__2
--+ |
| | +----------- Indopseudodon
| | |
| | +--------------+ +-------- Pilsbryoconcha
| | | | |
| | | +--+ +----- Bineurus_anodontinum
+--+ | | |
| | +--+----- Bineurus_loeiensis
| | |
| | | +-- Pseudodon
| | +--+
| | +-- Thaiconcha
| |
| | +----- Ensidens
| | |
| | +----- Hyriopsis_desowitzi
+--+ |
| +----- Rectidens
| |
| +----- Hyriopsis_bialata
| +-----------------+
| | +----- Khairuloconcha
| | |
| | | +-- Physunio
| | | |
| | | +-- Yaukthwa_paiensis
| | | |
| | | +-- Yaukthwa_inlenensis
| | +--+
+--+ +-- Trapezoideus
| |
| +-- Yaukthwa_elongatula
| |
| +-- Lens
|
| +-------------------- Inversidens
| |
| | +----- Lamprotula
+--+ +-----------+
| | | +-- Chamberlainia
| | +--+
+--+ +-- Potomida
|
| +-------------- Sinosolenaia
| |
+--+ +----------- Gonidea
| |
+--+----------- Microcondylaea
|
| +-------- Koreosolenaia
+--+
| +----- Parvasolenaia
+--+
| +-- Postolata
+--+
+-- Ptychorhynchus
Wu et al., 2024 [Publ 4350], fig. 6-7
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 34 | 0 | 0 | -- |
+-- Margaritifera_dahurica **
+-----------------------------+
| +-- Margaritifera_margaritifera **
|
| +----------------------------- Chamberlainia
| |
--+ | +----------- Khairuloconcha
| | |
| | | +----- Ensidens
| | +--------------+ +--+
| | | | | | +-- Rectidens
+--+ | | | +--+
| | | | +-- Hyriopsis
| | +--+
| | | +-- Physunio
| | | +--+
| | | | +-- Trapezoideus
| | +--+
| | | +-- Lens
+--+ +--+
| +-- Yaukthwa
|
| +-------------- Schepmania
| +--------+
| | | +----------- Indopseudodon
| | +--+
| | | +-------- Pilsbryoconcha
| | +--+
| | | +----- Pseudodon
| | +--+
+--+ | +-- Thaiconcha
| +--+
| +-- Bineurus
|
| +-- Potomida
| +-----------------+
| | +-- Lamprotula
+--+
| +----------------- Inversidens
| |
+--+ +-------------- Sinosolenaia
| |
+--+ +-- Gonidea
| +--------+
| | +-- Microcondylaea
+--+
| +-------- Koreosolenaia
+--+
| +----- Parvasolenaia
+--+
| +-- Postolata
+--+
+-- Ptychorhynchus
Wu et al., 2024 [Publ 4350], fig. 8a_S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 89 | 0 | 0 | -- |
+-- Margaritifera_dahurica **
+--------------------------------------+
| +-- Margaritifera_margaritifera **
|
| +-- Quadrula
| +--------------------------+
| | +-- Uniomerus
| |
| +--------+ +-- Elliptio
| | | +-----------------------+
| | | | +-- Pleurobema
| | +--+
| | | +----------------------- Amblema
| | | |
| | +--+ +-------------------- Popenaias
| | | |
--+ | +--+ +----------------- Toxolasma
| | | |
| | +--+ +-------------- Potamilus
| | | |
| | +--+ +----------- Leaunio
| | | |
| | | | +----- Lampsilis_ornata
| | +--+ +--+
| | | | | +-- Lampsilis_cardium
| | | | +--+
| | +--+ +-- Lampsilis_cariosa
| | |
| | | +----- Venustaconcha
| | +--+
| | | +-- Lampsilis_powellii
+--+ +--+
| +-- Lampsilis_siliquoidea
|
| +-- Chamberlainia
| +-----------------+
| | +-- Sinohyriopsis
| |
| | +-- Hyriopsis
| | +--+
| +--------------+ | +-- Rectidens
| | | +-----------+
| | | | | +-- Lens
| | | | +--+
| | | | +-- Physunio
| | | |
| | +--+ +----- Potomida
| | | +--------+
| | | | | +-- Pronodularia
| | | | +--+
| | | | +-- Lamprotula
| | +--+
| | | +-- Pseudodon
| | | +--------+
| | | | +-- Pilsbryoconcha
+--+ +--+
| | +-------- Inversidens
| | |
| +--+ +-- Parvasolenaia
| | +--+
| | | +-- Ptychorhynchus
| +--+
| | +-- Microcondylaea
| +--+
| +-- Sinosolenaia
|
| +-------------------------------- Lamellidens
| |
| | +----------------------------- Lepidodesma
| | |
+--+ | +-------------- Scriptolamprotula
| | |
| | +-----------+ +----------- Unio
| | | | |
+--+ | +--+ +-------- Nodularia
| | | |
| | +--+-------- Pseudocuneopsis
| | |
| | | +----- Schistodesmus
| | +--+
+--+ | +-- Tchangsinaia
| +--+
| +-- Cuneopsis
|
| +-- Acuticosta
| +--------------------+
| | +-- Lanceolaria
| |
| | +-------- Cristaria
+--+ +-----------+
| | | +----- Anemina
| | +--+
| | | +-- Beringiana
| | +--+
+--+ +-- Sinanodonta
|
| +-- Pseudanodonta
| +--------------+
| | +-- Anodonta
+--+
| +-------------- Brachyanodon
| |
+--+ +-- Pyganodon
| +--------+
| | +-- Utterbackia
+--+
| +-------- Pterosyna
+--+
| +----- Prolasmidonta
+--+
| +-- Alasmidonta
+--+
+-- Platynaias
Wu et al., 2024 [Publ 4379], fig. 4_6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 21 | 0 | 0 | -- |
+-- Margaritifera_dahurica **
+-----------+
| +-- Margaritifera_margaritifera **
|
--+ +-- Potomida
| +--------+
| | +-- Lamprotula
+--+
| +-------- Scriptolamprotula
| |
+--+ +-- Cristaria
| +--+
| | +-- Sinanodonta
+--+
| +-- Schistodesmus
+--+
+-- Nodularia
Wu et al., 2024 [Publ 4350], fig. 8b_S2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 89 | 0 | 0 | -- |
+-- Margaritifera_dahurica **
+--------------------------------------+
| +-- Margaritifera_margaritifera **
|
| +-- Quadrula
| +--------------------------+
| | +-- Uniomerus
| |
| +--------+ +-- Elliptio
| | | +-----------------------+
| | | | +-- Pleurobema
| | +--+
| | | +----------------------- Amblema
| | | |
| | +--+ +-------------------- Popenaias
| | | |
--+ | +--+ +----------------- Toxolasma
| | | |
| | +--+ +-------------- Potamilus
| | | |
| | +--+ +----------- Leaunio
| | | |
| | +--+ +-------- Venustaconcha
| | | |
| | | | +-- Lampsilis_powellii
| | +--+-----+
| | | +-- Lampsilis_siliquoidea
| | |
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_cardium
+--+ +--+
| +-- Lampsilis_cariosa
|
| +-- Chamberlainia
| +-----------------+
| | +-- Sinohyriopsis
| |
| | +-- Hyriopsis
| | +--+
| +--------------+ | +-- Rectidens
| | | +-----------+
| | | | | +-- Lens
| | | | +--+
| | | | +-- Physunio
| | | |
| | +--+ +----- Potomida
| | | +--------+
| | | | | +-- Pronodularia
| | | | +--+
| | | | +-- Lamprotula
| | +--+
| | | +-- Pseudodon
| | | +--------+
| | | | +-- Pilsbryoconcha
+--+ +--+
| | +-------- Inversidens
| | |
| +--+ +-- Parvasolenaia
| | +--+
| | | +-- Ptychorhynchus
| +--+
| | +-- Microcondylaea
| +--+
| +-- Sinosolenaia
|
| +-------------------------------- Lamellidens
| |
| | +----------------------------- Lepidodesma
| | |
+--+ | +-------------- Scriptolamprotula
| | |
| | +-----------+ +----------- Unio
| | | | |
+--+ | +--+ +-------- Nodularia
| | | |
| | +--+-------- Pseudocuneopsis
| | |
| | | +----- Schistodesmus
| | +--+
+--+ | +-- Tchangsinaia
| +--+
| +-- Cuneopsis
|
| +-- Acuticosta
| +--------------------+
| | +-- Lanceolaria
| |
| | +-------- Cristaria
+--+ +-----------+
| | | +----- Anemina
| | +--+
| | | +-- Beringiana
| | +--+
+--+ +-- Sinanodonta
|
| +----------------- Brachyanodon
| |
| | +-- Pseudanodonta
+--+ +-----------+
| | +-- Anodonta
| |
+--+ +-- Pyganodon
| +--------+
| | +-- Utterbackia
+--+
| +-------- Pterosyna
+--+
| +----- Prolasmidonta
+--+
| +-- Alasmidonta
+--+
+-- Platynaias
Liu et al., 2024 [Publ 4380], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 34 | 0 | 0 | -- |
+-- Margaritifera_dahurica **
+-----------------------+
| +-- Margaritifera_margaritifera **
|
| +-------------- Schepmania
| +--------+
| | | +----------- Indopseudodon
| | +--+
--+ | | +-------- Pilsbryoconcha
| | +--+
| | | +----- Pseudodon
| | +--+
| | | +-- Thaiconcha
| | +--+
| | +-- Bineurus
| |
+--+ +-------------- Sinosolenaia
| |
| | +-- Gonidea
| +-----+ +--------+
| | | | +-- Microcondylaea
| | | |
| | +--+ +-- Koreosolenaia
| | | +-----+
| | | | +-- Parvasolenaia
| | +--+
| | | +----- Postolata
| | +--+
+--+ | +-- Obovalis
| +--+
| +-- Ptychorhynchus
|
| +----- Chamberlainia
| +-----------+
| | | +-- Potomida
| | +--+
| | +-- Lamprotula
+--+
| +-------------- Khairuloconcha
| |
| | +-------- Ensidens
| | +--+
+--+ | | +----- Hyriopsis_bialata
| | +--+
| | | +-- Hyriopsis_desowitzi
| | +--+
+--+ +-- Rectidens
|
| +-- Physunio
| +--+
| | +-- Trapezoideus
+-----+
| +-- Lens
+--+
+-- Yaukthwa
Zhang et al., 2024 [Publ 4397], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 25 | 0 | 0 | -- |
+-- Margaritifera_dahurica **
+-----------------------------+
| +-- Margaritifera_margaritifera **
|
| +-- Chamberlainia
--+ +--------------------------+
| | +-- Sinohyriopsis
| |
| | +-- Hyriopsis
| | +--+
+--+ | +-- Rectidens
| +--------------------+
| | | +-- Lens
| | +--+
| | +-- Physunio
| |
+--+ +----- Potomida
| +-----------------+
| | | +-- Pronodularia
| | +--+
| | +-- Lamprotula
+--+
| +-- Pilsbryoconcha
| +-----------------+
| | +-- Pseudodon
+--+
| +----------------- Inversidens
| |
+--+ +-- Microcondylaea
| +-----------+
| | +-- Sinosolenaia
+--+
| +----------- Parvasolenaia
+--+
| +-------- Koreosolenaia
+--+
| +----- Ptychorhynchus
+--+
| +-- Cosmopseudodon
+--+
+-- Postolata
Wu et al., 2024 [Publ 4388], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 34 | 0 | 0 | -- |
+-- Margaritifera_dahurica **
+--------------------------------+
| +-- Margaritifera_margaritifera **
|
| +-------------------------------- Chamberlainia
| |
--+ | +----------- Khairuloconcha
| | |
| | | +----- Ensidens
| | +-----------------+ +--+
| | | | | | +-- Rectidens
+--+ | | | +--+
| | | | +-- Hyriopsis
| | +--+
| | | +-- Physunio
| | | +--+
| | | | +-- Trapezoideus
| | +--+
+--+ | +-- Lens
| +--+
| +-- Yaukthwa
|
| +-- Potomida
| +-----------------------+
| | +-- Lamprotula
| |
| | +-------------- Schepmania
| | +--------+
+--+ | | +----------- Indopseudodon
| | +--+
| | | +-------- Pilsbryoconcha
| | +--+
| | | +----- Pseudodon
+--+ +--+
| | +-- Thaiconcha
| +--+
| +-- Bineurus
|
| +-------------------- Sinosolenaia
| |
+--+ +-- Gonidea
| +--------------+
| | +-- Microcondylaea
+--+
| +-------------- Koreosolenaia
+--+
| +----------- Parvasolenaia
+--+
| +-------- Postolata
+--+
| +----- Cosmopseudodon
+--+
| +-- Ptychorhynchus
+--+
+-- Obovalis
Wu et al., 2024 [Publ 4388], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 51 | 0 | 0 | -- |
+-- Margaritifera_dahurica **
+--------------------------------+
| +-- Margaritifera_margaritifera **
|
| +-- Quadrula
| +-----------------+
| | +-- Uniomerus
| |
| +-----------+ +-- Elliptio
| | | +--------------+
| | | | +-- Pleurobema
| | +--+
| | | +-------------- Popenaias
--+ | +--+
| | | +----------- Toxolasma
| | +--+
| | | +-------- Potamilus
| | +--+
| | | +----- Leaunio
| | +--+
| | | +-- Lampsilis
| | +--+
| | +-- Venustaconcha
| |
| | +-------------------------- Lamellidens
| | |
+--+ | +----------------------- Lepidodesma
| +--+ |
| | | | +----------------- Scriptolamprotula
| | | | +--+
| | | | | | +-------------- Unio
| | +--+ | +--+
| | | | | +----------- Pseudocuneopsis
| | | | +--+
| | | | | +-------- Nodularia
| | | | +--+
| | | | | +----- Tchangsinaia
| | +--+ +--+
| | | | +-- Cuneopsis
| | | +--+
| | | +-- Schistodesmus
| | |
| | | +-------------- Lanceolaria
| | | |
| | | | +----- Anodonta
+--+ +-----+ +-----+
| | | | +-- Alasmidonta
| | | +--+
| +--+ +-- Utterbackia
| |
| | +-------- Cristaria
| +--+
| | +----- Anemina
| +--+
| | +-- Beringiana
| +--+
| +-- Sinanodonta
|
| +-- Chamberlainia
| +--------------------+
| | +-- Sinohyriopsis
| |
| | +-- Hyriopsis
| | +--+
+-----+ | +-- Rectidens
| +--------------+
| | | +-- Lens
| | +--+
| | +-- Physunio
| |
+--+ +----- Potomida
| +-----------+
| | | +-- Pronodularia
| | +--+
| | +-- Lamprotula
+--+
| +-- Pseudodon
| +-----------+
| | +-- Pilsbryoconcha
| |
+--+ +-- Microcondylaea
| +--------+
| | +-- Sinosolenaia
+--+
| +-------- Parvasolenaia
+--+
| +----- Ptychorhynchus
+--+
| +-- Cosmopseudodon
+--+
+-- Postolata
Hou et al., 2024 [Publ 4400], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 73 | 0 | 0 | -- |
+-- Margaritifera_dahurica **
+--------------------------------+
| +-- Margaritifera_margaritifera **
|
| +-- Lampsilis
--+ +-----------------------------+
| | +-- Quadrula
| |
| | +----------------------------- Lepidodesma
+--+ |
| | +-------------------- Scriptolamprotula
| | +-----+
| | | | +----------------- Unio
| | | +--+
+--+ | | +-------------- Middendorffinaia
| | +--+
| | | +----------- Pseudocuneopsis
| | +--+
| | | +-------- Nodularia
| | +--+
+--+ | +----- Schistodesmus
| +--+
| | +-- Tchangsinaia
| +--+
| +-- Cuneopsis
|
| +----------------------- Lanceolaria
| |
| | +-------- Cristaria
| | +-----------+
+--+ | | +----- Anemina
| | +--+
| | | +-- Beringiana
| | +--+
+--+ +-- Sinanodonta
|
| +-- Pseudanodonta
| +--------------+
| | +-- Anodonta
+--+
| +-------------- Brachyanodon
| |
+--+ +-- Pyganodon
| +--------+
| | +-- Utterbackia
+--+
| +-------- Pterosyna
+--+
| +----- Prolasmidonta
+--+
| +-- Alasmidonta
+--+
+-- Platynaias
Machordom et al., 2003 [Publ 2688], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 3 | 2 | 2 | -- |
+----- Cumberlandia --+ | +-- Margaritifera_margaritifera ** +--+ +-- Pseudunio
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 5 | 2 | 2 | -- |
+----- Margaritifera_margaritifera ** --+ | +-- Pseudunio +--+ +-- Cumberlandia
Guerra et al., 2019 [Publ 4168], fig. 1-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 21 | 2 | 2 | -- |
+----- Cumberlandia --+ | +-- Margaritifera_margaritifera ** +--+ +-- Pseudunio
Guerra et al., 2019 [Publ 4168], fig. 2-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 21 | 2 | 2 | -- |
+----- Pseudunio --+ | +-- Margaritifera_margaritifera ** +--+ +-- Cumberlandia
Campbell & Lydeard, 2026 [Publ 4810], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 46 | 2 | 2 | -- |
+----- Pseudunio --+ | +-- Cumberlandia +--+ +-- Margaritifera_margaritifera **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 29 | 1 | 1 | -- |
+-- Margaritifera_margaritifera ** --+ +-- Cumberlandia
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 29 | 1 | 1 | -- |
+-- Margaritifera_margaritifera ** --+ +-- Cumberlandia
Bogan & Hoeh, 2000 [Publ 2650], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 33 | 1 | 1 | -- |
+-- Cumberlandia --+ +-- Margaritifera_margaritifera **
Hoeh et al., 2001 [Publ 2652], fig. 14f2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 30 | 1 | 1 | -- |
+-- Cumberlandia --+ +-- Margaritifera_margaritifera **
Hoeh et al., 2001 [Publ 2652], fig. 14f4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 30 | 1 | 1 | -- |
+-- Cumberlandia --+ +-- Margaritifera_margaritifera **
Hoeh et al., 2002 [Publ 2653], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 30 | 1 | 1 | -- |
+-- Margaritifera_margaritifera ** --+ +-- Cumberlandia
Huang et al., 2002 [Publ 2657], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.667 | 1 | 1 | 0 | 1 | 33 | 1 | 1 | -- |
+----------------------------------------------- Cumberlandia | +----------------------------------------------- Margaritifera_margaritifera ** | | +----- Acuticosta | +--+ | | | +-- Nodularia | | +--+ | +--------+ +-- Tchangsinaia | | | | | | +----- Lanceolaria --+ | +--+ | | | +-- Schistodesmus | +--------------------------+ +--+ | | | +-- Lepidodesma | | | | | | +-- Sinanodonta | | | +-----------+ | | | | +-- Cristaria | | +--+ | | | +----------- Utterbackiana | | +--+ +--+ | +-------- Utterbackia | +--+ | | +----- Strophitus | +--+ | | +-- Alasmidonta | +--+ | +-- Lasmigona | | +----------------------------------------- Sinosolenaia | | +--+ +-------------------------------------- Lamprotula | | +--+ +----------------------------------- Sinohyriopsis | | +--+ +-------------------------------- Ptychorhynchus | | +--+ +----------------------------- Megalonaias | | | | +----- Elliptoideus +--+ +--------------------+ | | | +-- Uniomerus | | +--+ +--+ +-- Pleurobema | | +----------------------- Amblema | | +--+ +-- Obliquaria | +-----------------+ | | +-- Plectomerus +--+ | +----------------- Medionidus +--+ | +-------------- Potamilus_fragilis +--+ | +----------- Ptychobranchus +--+ | +-------- Potamilus_purpuratus +--+ | +----- Lampsilis +--+ | +-- Villosa +--+ +-- Ellipsaria
Huang et al., 2002 [Publ 2657], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.667 | 1 | 1 | 0 | 1 | 33 | 1 | 1 | -- |
+-------------------- Cumberlandia | +-------------------- Margaritifera_margaritifera ** | | +-------- Strophitus | | | +-------- Schistodesmus | | | +-------- Lepidodesma | | | +-------- Acuticosta | | --+ +-------- Lanceolaria | | | +--------+ +-- Sinanodonta | | +-----+ | | | +-- Cristaria | | | | | | +-- Nodularia | | +-----+ | | | +-- Tchangsinaia | | | | | | +----- Utterbackia | | | | | | +--+----- Utterbackiana +--+ | | | +-- Alasmidonta | +--+ | +-- Lasmigona | | +-------------- Ptychorhynchus | | | +-------------- Sinohyriopsis | | | +-------------- Lamprotula | | +--+-------------- Sinosolenaia | | +----------- Megalonaias | | | | +----- Elliptoideus | | +--+ +--+ | | +-- Uniomerus | | +--+ | | +-- Pleurobema | | | | +----- Potamilus_fragilis +--+ | | +----- Medionidus | | | +----- Amblema | | | | +-- Obliquaria +--+--+ | +-- Plectomerus | | +-- Villosa | | | +-- Ellipsaria | | +--+-- Lampsilis | +-- Potamilus_purpuratus | +-- Ptychobranchus
Roe & Hoeh, 2003 [Publ 2655], fig. 4f2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 30 | 1 | 1 | -- |
+-- Cumberlandia --+ +-- Margaritifera_margaritifera **
Roe & Hoeh, 2003 [Publ 2655], fig. 4f3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 30 | 1 | 1 | -- |
+-- Cumberlandia --+ +-- Margaritifera_margaritifera **
Machordom et al., 2003 [Publ 2688], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 2 | 1 | 1 | -- |
+-- Margaritifera_margaritifera ** --+ +-- Pseudunio
Campbell et al., 2005 [Publ 2660], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 101 | 1 | 1 | -- |
+-- Margaritifera_margaritifera ** --+ +-- Cumberlandia
Campbell et al., 2005 [Publ 2660], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 101 | 1 | 1 | -- |
+-- Cumberlandia --+ +-- Margaritifera_margaritifera **
Graf & Cummings, 2006 [Publ 1968], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 46 | 1 | 1 | -- |
+-- Margaritifera_margaritifera ** --+ +-- Cumberlandia
Zhou et al., 2007 [Publ 2658], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 20 | 1 | 1 | -- |
+-- Margaritifera_margaritifera ** --+ +-- Cumberlandia
Hoeh et al., 2009 [Publ 2716], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 46 | 1 | 1 | -- |
+-- Margaritifera_margaritifera ** --+ +-- Cumberlandia
Hoeh et al., 2009 [Publ 2716], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 45 | 1 | 1 | -- |
+-- Margaritifera_margaritifera ** --+ +-- Cumberlandia
Hoeh et al., 2009 [Publ 2716], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 45 | 1 | 1 | -- |
+-- Margaritifera_margaritifera ** --+ +-- Cumberlandia
Whelan et al., 2011 [Publ 3380], fig. 1-ML
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 35 | 1 | 1 | -- |
+-- Margaritifera_margaritifera ** --+ +-- Cumberlandia
Whelan et al., 2011 [Publ 3380], fig. 1-MP
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 35 | 1 | 1 | -- |
+-- Margaritifera_margaritifera ** --+ +-- Cumberlandia
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 39 | 1 | 1 | -- |
+-- Margaritifera_margaritifera ** --+ +-- Cumberlandia
Lopes-Lima et al., 2017 [Publ 3734], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 16 | 1 | 1 | -- |
+-- Pseudunio --+ +-- Margaritifera_margaritifera **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 80 | 1 | 1 | -- |
+-- Cumberlandia --+ +-- Margaritifera_margaritifera **
Guerra et al., 2017 [Publ 3772], fig. 2a
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 24 | 1 | 1 | -- |
+-- Margaritifera_falcata ** --+ +-- Cumberlandia
Guerra et al., 2017 [Publ 3772], fig. 2b
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 24 | 1 | 1 | -- |
+-- Margaritifera_falcata ** --+ +-- Cumberlandia
Välilä et al., 2017 [Publ 3793], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 2 | 1 | 1 | -- |
+-- Margaritifera_margaritifera ** --+ +-- Pseudunio
Huang et al., 2019 [Publ 3843], fig. 3-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 22 | 1 | 1 | -- |
+-- Pseudunio --+ +-- Margaritifera_margaritifera **
Lopes-Lima et al., 2019 [Publ 3901], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 27 | 1 | 1 | -- |
+-- Margaritifera_falcata ** --+ +-- Cumberlandia
Lopes-Lima et al., 2019 [Publ 3901], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 28 | 1 | 1 | -- |
+-- Cumberlandia --+ +-- Margaritifera_falcata **
Lopes-Lima et al., 2019 [Publ 3901], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 25 | 1 | 1 | -- |
+-- Margaritifera_falcata ** --+ +-- Cumberlandia
Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 18 | 1 | 1 | -- |
+-- Margaritifera_falcata ** --+ +-- Cumberlandia
Zieritz et al., 2021 [Publ 4218], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 30 | 1 | 1 | -- |
+-- Margaritifera_margaritifera ** --+ +-- Pseudunio
Zieritz et al., 2024 [Publ 4381], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 44 | 1 | 1 | -- |
+-- Margaritifera_margaritifera ** --+ +-- Pseudunio
Nagel et al., 1998 [Publ 1946], fig. 9
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- |
+-------------- Margaritifera ** --+ | +----------- Potomida +--+ | +-------- Microcondylaea +--+ | +----- Unio +--+ | +-- Anodonta +--+ +-- Pseudanodonta
Hoeh et al., 1998 [Publ 2695], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 4 | 0 | 0 | -- |
+-- Margaritifera_margaritifera ** --+ +-- Amblema
Hoeh et al., 1998 [Publ 2695], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 4 | 0 | 0 | -- |
+-- Margaritifera_margaritifera ** --+ +-- Anodonta
Graf, 2000 [Publ 2647], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 16 | 0 | 0 | -- |
+----------------------- Margaritifera ** | | +-------- Unio --+ +-----------+ | | | +----- Elliptio | | +--+ | | | +-- Pyganodon +--+ +--+ | +-- Lampsilis | | +----------------- Parreysia | | +--+ +-------------- Grandidieria | | | | +-------- Castalia +--+ +--+ | | | +----- Diplodon | | +--+ | | | +-- Hyridella +--+ +--+ | +-- Velesunio | | +-------- Mutela | | +--+ +----- Anodontites | | +--+----- Mycetopoda | | +-- Etheria +--+ +-- Mulleria
Turner et al., 2000 [Publ 2722], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- |
+----------------------------- Margaritifera_margaritifera ** | | +-- Quadrula --+ +-----------------------+ | | +-- Utterbackiana | | +--+ +-- Amblema | +--------------------+ | | +-- Eurynia +--+ | +-------------------- Medionidus | | +--+ +----------------- Lampsilis_teres | | | +----------------- Villosa +--+ | +-- Lampsilis_satura | +-----------+ | | +-- Lampsilis_cardium +--+ | +----------- Lampsilis_reeveiana | | +--+ +-------- Lampsilis_hydiana__1 | | +--+-------- Lampsilis_hydiana__2 | | +----- Lampsilis_siliquoidea__2 +--+ | +-- Lampsilis_siliquoidea__1 +--+ +-- Lampsilis_powellii
Campbell, 2000 [Publ 2724], fig. 1-5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 2 | 0 | 0 | -- |
+----- Margaritifera_margaritifera ** --+ | +-- Utterbackia +--+ +-- Elliptio
Curole & Kocher, 2002 [Publ 2712], fig. 1-F
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 3 | 0 | 0 | -- |
+-------- Margaritifera_hembeli ** --+ | +----- Lampsilis +--+ | +-- Quadrula +--+ +-- Pustulosa
Giribet & Distel, 2003 [Publ 3670], fig. 3f5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- |
+----- Margaritifera_margaritifera ** | --+ +-- Elliptio | | +--+-- Lampsilis | +-- Unio
Curole & Kocher, 2005 [Publ 2682], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 10 | 0 | 0 | -- |
+-- Pyganodon --+ +-- Margaritifera_hembeli **
Curole & Kocher, 2005 [Publ 2682], fig. 1-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 11 | 0 | 0 | -- |
+-------------------- Margaritifera_hembeli ** | | +-- Utterbackiana --+ +--------------+ | | +-- Pyganodon | | +--+ +-- Quadrula | +-----------+ | | +-- Pustulosa +--+ | +----------- Plectomerus | | +--+ +-------- Amblema | | +--+ +-- Potamilus | +--+ | | +-- Glebula +--+ | +-- Leaunio +--+ +-- Lampsilis
Källersjö et al., 2005 [Publ 2684], fig. 1A
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 6 | 0 | 0 | -- |
+----- Margaritifera_margaritifera ** | +----- Unio --+ | +-- Anodonta_anatina | | +--+-- Pseudanodonta | +-- Anodonta_cygnea
Källersjö et al., 2005 [Publ 2684], fig. 1B-F
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 6 | 0 | 0 | -- |
+-------- Margaritifera_margaritifera ** | --+-------- Unio | | +----- Anodonta_anatina +--+ | +-- Pseudanodonta +--+ +-- Anodonta_cygnea
Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 4 | 0 | 0 | -- |
+-------- Margaritifera_falcata ** --+ | +----- Gonidea +--+ | +-- Beringiana +--+ +-- Brachyanodon
Taylor et al., 2007 [Publ 2696], fig. 1-3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 3 | 0 | 0 | -- |
+----- Margaritifera_margaritifera ** --+ | +-- Anodonta +--+ +-- Unio
Zieritz et al., 2012 [Publ 3673], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 7 | 0 | 0 | -- |
+-- Sinanodonta | --+-- Margaritifera_margaritifera ** | +-- Unio
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 30 | 0 | 0 | -- |
+-------------------------- Margaritifera_margaritifera ** | +-------------------------- Velesunio | | +-- Lamellidens | +--------------------+ | | +-- Coelatura --+ | | | +-------- Acuticosta | | +--+ | | | | +----- Scriptolamprotula | | | +--+ | | | | +-- Unio | | | +--+ +--+ +--------+ +-- Nodularia | | | | | | +-- Cristaria | | | +-----+ | | | | +-- Sinanodonta | | +--+ | | | +----- Strophitus | | +--+ | | | +-- Platynaias +--+ +--+ | +-- Pyganodon | | +----------- Quadrula | +-----+ | | | +-------- Eurynia | | +--+ | | | +----- Amblema | | +--+ | | | +-- Lampsilis +--+ +--+ | +-- Truncilla | | +-------------- Sinohyriopsis | | | | +----- Gonidea +--+ +-----+ | | | +-- Pseudodon | | +--+ +--+ +-- Sundadontina | | +-------- Physunio +--+ | +----- Pronodularia +--+ | +-- Potomida +--+ +-- Lamprotula
Reis et al., 2013 [Publ 3580], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- |
+----------------- Margaritifera_margaritifera ** | --+ +-- Amblema | +-----------+ | | +-- Quadrula +--+ | +----------- Potomida +--+ | +-------- Unio +--+ | +----- Anodonta_cygnea +--+ | +-- Anodonta_anatina +--+ +-- Pseudanodonta
Reis et al., 2013 [Publ 3580], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- |
+-------------- Margaritifera_margaritifera ** | --+ +-- Amblema | +--------+ | | +-- Theliderma +--+ | +-------- Potomida | | +--+ +----- Unio | | +--+ +-- Anodonta_anatina | | +--+-- Anodonta_cygnea | +-- Pseudanodonta
Huang et al., 2013 [Publ 3629], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 15 | 0 | 0 | -- |
+----------------- Margaritifera_falcata ** | | +----------- Unio | | | +--+ +-- Sinanodonta --+ | | +-----+ | | | | +-- Cristaria | | +--+ | | | +----- Platynaias | | +--+ | | | +-- Pyganodon +--+ +--+ | +-- Utterbackia | | +----- Sinohyriopsis | +-----+ | | | +-- Sinosolenaia | | +--+ +--+ +-- Pronodularia | | +-------- Quadrula +--+ | +----- Toxolasma +--+ | +-- Lampsilis +--+ +-- Venustaconcha
Zhang et al., 2013 [Publ 4214], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 3 | 0 | 0 | -- |
+-- Margaritifera_margaritifera ** --+ +-- Lamprotula
Bieler et al., 2014 [Publ 3622], fig. 26_27_29_30
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 4 | 0 | 0 | -- |
+-- Aspatharia --+ +-- Margaritifera_margaritifera **
Bieler et al., 2014 [Publ 3622], fig. 28
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 4 | 0 | 0 | -- |
+-- Aspatharia --+ +-- Margaritifera_margaritifera **
Graf et al., 2014 [Publ 3591], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 10 | 0 | 0 | -- |
+-------------------------- Margaritifera_margaritifera ** | +-------------------------- Unio --+ | +----------------------- Leoparreysia | | +--+ +-------------------- Nitia | | +--+ +----------------- Coelatura_aegyptiaca__2 | | +--+ +-------------- Coelatura_aegyptiaca__1 | | +--+ +----------- Coelatura_gabonensis | | +--+ +-------- Coelatura_kunenensis | | | | +----- Prisodontopsis +--+ | | | +-- Coelatura_choziensis__1 | | | +--+ +-- Coelatura_choziensis__2 | | | +-- Coelatura_luapulaensis__1 | | +--+-- Coelatura_luapulaensis__2 | +-- Coelatura_luapulaensis__3 | +-- Coelatura_luapulaensis__4 | +-- Coelatura_luapulaensis__5
Graf et al., 2015 [Publ 3619], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 19 | 0 | 0 | -- |
+-- Unio --+ +-- Margaritifera_margaritifera **
González & Giribet, 2015 [Publ 3632], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- |
+-- Aspatharia --+ +-- Margaritifera_margaritifera **
González & Giribet, 2015 [Publ 3632], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- |
+-- Aspatharia --+ +-- Margaritifera_margaritifera **
González & Giribet, 2015 [Publ 3632], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- |
+-- Unio --+ +-- Margaritifera_margaritifera **
González & Giribet, 2015 [Publ 3632], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- |
+-- Aspatharia --+ +-- Margaritifera_margaritifera **
Zhou et al., 2016 [Publ 3710], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 19 | 0 | 0 | -- |
+-------------- Margaritifera_falcata ** | | +-------- Quadrula --+ +--+ | | | +----- Toxolasma | | +--+ | | | +-- Venustaconcha +--+ +--+ | +-- Lampsilis | | +-------- Sinohyriopsis +--+ | +----- Sinosolenaia +--+ | +-- Lamprotula +--+ +-- Pronodularia
Bolotov et al., 2016 [Publ 3732], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 7 | 0 | 0 | -- |
+-- Gibbosula --+ +-- Margaritifera_dahurica **
Amaro et al., 2017 [Publ 3791], fig. 3A-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 6 | 0 | 0 | -- |
+-------------- Margaritifera_margaritifera ** | --+ +----------- Quadrula | | +--+ +-- Pronodularia | +-----+ | | +-- Sinosolenaia +--+ | +----- Venustaconcha +--+ | +-- Pyganodon +--+ +-- Utterbackia
Amaro et al., 2017 [Publ 3791], fig. 3B-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 6 | 0 | 0 | -- |
+-------------- Margaritifera_margaritifera ** | --+ +----------- Quadrula | | +--+ +-------- Venustaconcha | | +--+ +-- Utterbackia | +--+ | | +-- Pyganodon +--+ | +-- Pronodularia +--+ +-- Sinosolenaia
Froufe et al., 2017 [Publ 3802], fig. 7
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 9 | 0 | 0 | -- |
+----------- Margaritifera_margaritifera ** | --+ +-- Microcondylaea | +-----+ | | +-- Potomida +--+ | +----- Unio +--+ | +-- Sinanodonta +--+ +-- Anodonta
Combosch et al., 2017 [Publ 3806], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- |
+-------------- Margaritifera_margaritifera ** | --+ +-- Chambardia | +--------+ | | +-- Aspatharia +--+ | +-------- Etheria +--+ | +----- Mutela +--+ | +-- Tamsiella +--+ +-- Lamproscapha
Combosch et al., 2017 [Publ 3806], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- |
+-------- Margaritifera_margaritifera ** --+ | +----- Unio +--+ | +-- Scabies +--+ +-- Lampsilis
Vikhrev et al., 2017 [Publ 3810], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 9 | 0 | 0 | -- |
+-- Gibbosula --+ +-- Margaritifera_dahurica **
Hewitt et al., 2018 [Publ 3758], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 8 | 0 | 0 | -- |
+-------------- Margaritifera_margaritifera ** | | +-- Fusconaia --+ +--------+ | | +-- Pleurobema | | +--+ +-------- Pyganodon | | | | +-- Platynaias +--+-----+ | +-- Strophitus | | +----- Lasmigona +--+ | +-- Arcidens +--+ +-- Pterosyna
Bespalaya et al., 2018 [Publ 3811], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 9 | 0 | 0 | -- |
+-- Gibbosula --+ +-- Margaritifera_dahurica **
Pfeiffer et al., 2018 [Publ 3827], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 27 | 0 | 0 | -- |
+----------------------------------- Margaritifera_margaritifera ** | --+ +-- Trapezidens | +-----------------------------+ | | +-- Lamellidens +--+ | +----------------------------- Parreysia | | +--+ +-------------------------- Leoparreysia | | | | +----- Nitia +--+ +-----------------+ | | | +-- Coelatura | | +--+ +--+ +-- Prisodontopsis | | +-------------------- Indonaia +--+ | +----------------- Indochinella +--+ | +-------------- Radiatula +--+ | +----------- Scabiellus +--+ | +-------- Unionetta_fabagina__2 +--+ | +----- Harmandia +--+ | +-- Unionetta_fabagina__1 +--+ +-- Scabies
Bogatov et al., 2018 [Publ 3859], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- |
+----------- Margaritifera_margaritifera ** | --+ +-------- Unio_tumidus | | +--+ +-- Anodonta | +--+ | | +-- Pseudanodonta +--+ | +-- Unio_crassus +--+ +-- Unio_pictorum
Bogatov et al., 2018 [Publ 3859], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- |
+-------------- Margaritifera_margaritifera ** --+ | +----------- Unio_crassus +--+ | +-------- Unio_pictorum +--+ | +----- Unio_tumidus +--+ | +-- Anodonta +--+ +-- Pseudanodonta
Huang et al., 2019 [Publ 3843], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 45 | 0 | 0 | -- |
+-- Margaritifera_dahurica ** --+ +-- Gibbosula
Huang et al., 2019 [Publ 3843], fig. S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 46 | 0 | 0 | -- |
+-- Gibbosula --+ +-- Margaritifera_dahurica **
Klishko et al., 2019 [Publ 3886], fig. 9
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 10 | 0 | 0 | -- |
+----------- Margaritifera_dahurica ** --+ | +-------- Anodonta +--+ | +----- Unio +--+ | +-- Middendorffinaia +--+ +-- Nodularia
Konopleva et al., 2019 [Publ 3900], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 50 | 0 | 0 | -- |
+-- Gibbosula --+ +-- Margaritifera_dahurica **
Hewitt et al., 2019 [Publ 3962], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 9 | 0 | 0 | -- |
+----------------------------- Margaritifera_margaritifera ** | --+ +-------------------------- Pyganodon | | +--+ +----------------------- Fusconaia | | +--+ +-------------------- Lampsilis_siliquoidea | | | | +----------------- Lampsilis_fasciola +--+ | | | +-- Epioblasma | +--------------+ +--+ +-- Lampsilis_radiata | | +-------------- Lampsilis_ornata | | +--+ +-- Lampsilis_cardium__1 | +--------+ | | +-- Lampsilis_ovata__1 +--+ | +-------- Lampsilis_cardium__6 | | +--+ +----- Lampsilis_cardium__5 | | +--+ +-- Lampsilis_cardium__2 | | | +-- Lampsilis_ovata__2 +--+ +-- Lampsilis_cardium__3 | +-- Lampsilis_cardium__4
Lemer et al., 2019 [Publ 4093], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- |
+-- Margaritifera_margaritifera ** --+ +-- Lampsilis
Kano et al., 2019 [Publ 4194], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 9 | 0 | 0 | -- |
+-------------- Margaritifera_falcata ** | | +----- Sinanodonta --+ +-----+ | | | +-- Pletholophus | | +--+ | | +-- Beringiana +--+ | +----- Inversidens | +--+ | | | +-- Obovalis | | +--+ +--+ +-- Pronodularia | | +----- Lanceolaria +--+ | +-- Inversiunio +--+ +-- Nodularia
Wu et al., 2020 [Publ 4147], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 29 | 0 | 0 | -- |
+----------------------------- Margaritifera_falcata ** | | +----- Quadrula | +--------------------+ --+ | | +-- Toxolasma | | +--+ | | +-- Venustaconcha | | | | +-------- Sinohyriopsis +--+ +--------------+ | | | +----- Sinosolenaia | | +--+ | | | +-- Pronodularia | | +--+ | | +-- Lamprotula | | +--+ +----- Cristaria | +--+ | | | +-- Sinanodonta | | +--+ | +-----------+ +-- Anemina | | | | | | +----- Anodonta | | +--+ +--+ | +-- Pyganodon | +--+ | +-- Utterbackia | | +----------------- Scriptolamprotula +--+ | +-------------- Unio +--+ | +----------- Pseudocuneopsis +--+ | +-------- Nodularia +--+ | +----- Schistodesmus +--+ | +-- Cuneopsis +--+ +-- Tchangsinaia
Klunzinger et al., 2021 [Publ 4086], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- |
+-- Margaritifera_margaritifera ** --+ +-- Unio
Smith, 2021 [Publ 4331], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 19 | 0 | 0 | -- |
+-------------------- Margaritifera_margaritifera ** | | +----- Sinohyriopsis | +-----+ | | | +-- Lens | | +--+ --+ +-----+ +-- Lamprotula | | | | | | +-------- Scabies | | +--+ | | | +----- Cristaria | | +--+ +--+ | +-- Pyganodon | +--+ | +-- Utterbackia | | +-- Uniomerus | +-----------+ | | +-- Megalonaias +--+ | +----------- Elliptio +--+ | +-------- Amblema +--+ | +----- Potamilus +--+ | +-- Venustaconcha +--+ +-- Lampsilis
Bolotov et al., 2022 [Publ 4250], fig. 8
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 48 | 0 | 0 | -- |
+-- Gibbosula | +-- Margaritifera_dahurica ** | +-- Sinosolenaia | --+-- Leguminaia | +-- Gonidea | +-- Lamprotula | +-- Potomida
Dai et al., 2023 [Publ 4285], fig. 3-BI
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 44 | 0 | 0 | -- |
+-- Gibbosula --+ +-- Margaritifera_margaritifera **
Dai et al., 2023 [Publ 4285], fig. 3-ML
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 44 | 0 | 0 | -- |
+-- Gibbosula --+ +-- Margaritifera_margaritifera **
Dai et al., 2023 [Publ 4285], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 52 | 0 | 0 | -- |
+-- Gibbosula --+ +-- Margaritifera_dahurica **
Bogan et al., 2023 [Publ 4295], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 30 | 0 | 0 | -- |
+-- Gibbosula --+ +-- Margaritifera_laevis **
Chen et al., 2023 [Publ 4302], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 40 | 0 | 0 | -- |
+-- Gibbosula --+ +-- Margaritifera_dahurica **
Chen et al., 2023 [Publ 4302], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 40 | 0 | 0 | -- |
+-- Gibbosula --+ +-- Margaritifera_dahurica **
Dai et al., 2024 [Publ 4358], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 52 | 0 | 0 | -- |
+-- Gibbosula --+ +-- Margaritifera_dahurica **
Douda et al., 2024 [Publ 4364], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- |
+----- Margaritifera_middendorffi ** --+ | +-- Beringiana +--+ +-- Sinanodonta
Wu et al., 2024 [Publ 4379], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 52 | 0 | 0 | -- |
+-- Gibbosula --+ +-- Margaritifera_margaritifera **
Matos et al., 2024 [Publ 4383], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 19 | 0 | 0 | -- |
+----------------------- Margaritifera_margaritifera ** | | +-------------------- Amblema --+ | | | +-- Chamberlainia | | +--------------+ +--+ | +-- Sinohyriopsis | | | | +-- Physunio | | +--+ +--+ | +-- Lens | +--------+ | | | +-- Hyriopsis | | +--+ | | +-- Rectidens | | +--+ +----- Potomida | +-----+ | | | +-- Pronodularia | | +--+ | | +-- Lamprotula +--+ | +-- Pseudodon | +-----+ | | +-- Pilsbryoconcha +--+ | +----- Parvasolenaia +--+ | +-- Microcondylaea +--+ +-- Sinosolenaia
Matos et al., 2024 [Publ 4384], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 19 | 0 | 0 | -- |
+----------------------- Margaritifera_margaritifera ** | | +-------------------- Amblema --+ | | | +-- Chamberlainia | | +--------------+ +--+ | +-- Sinohyriopsis | | | | +-- Physunio | | +--+ +--+ | +-- Lens | +--------+ | | | +-- Hyriopsis | | +--+ | | +-- Rectidens | | +--+ +----- Potomida | +-----+ | | | +-- Pronodularia | | +--+ | | +-- Lamprotula +--+ | +-- Pseudodon | +-----+ | | +-- Pilsbryoconcha +--+ | +----- Parvasolenaia +--+ | +-- Microcondylaea +--+ +-- Sinosolenaia
Dai et al., 2024 [Publ 4385], fig. 3-BI
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 30 | 0 | 0 | -- |
+-- Gibbosula --+ +-- Margaritifera_margaritifera **
Dai et al., 2024 [Publ 4385], fig. 3-ML
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 30 | 0 | 0 | -- |
+-- Gibbosula --+ +-- Margaritifera_margaritifera **
Dai et al., 2024 [Publ 4386], fig. 5-BI
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 21 | 0 | 0 | -- |
+-- Margaritifera_margaritifera ** --+ +-- Gibbosula
Dai et al., 2024 [Publ 4386], fig. 5-ML
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 21 | 0 | 0 | -- |
+-- Margaritifera_margaritifera ** --+ +-- Gibbosula
Dai et al., 2024 [Publ 4389], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 45 | 0 | 0 | -- |
+-- Margaritifera_margaritifera ** --+ +-- Gibbosula
Kashiri et al., 2024 [Publ 4822], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 9 | 0 | 0 | -- |
+-- Gibbosula --+ +-- Margaritifera_dahurica **
Dai et al., 2025 [Publ 4440], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 43 | 0 | 0 | -- |
+-- Gibbosula --+ +-- Margaritifera_dahurica **
Dai et al., 2025 [Publ 4440], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 43 | 0 | 0 | -- |
+-- Gibbosula --+ +-- Margaritifera_dahurica **
Dai et al., 2025 [Publ 4441], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 46 | 0 | 0 | -- |
+-- Gibbosula --+ +-- Margaritifera_margaritifera **
Chen et al., 2025 [Publ 4487], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 40 | 0 | 0 | -- |
+-- Margaritifera_dahurica ** --+ +-- Gibbosula
Chen et al., 2025 [Publ 4487], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 40 | 0 | 0 | -- |
+-- Margaritifera_dahurica ** --+ +-- Gibbosula
Wang et al., 2025 [Publ 4488], fig. 3B
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 63 | 0 | 0 | -- |
+-- Gibbosula --+ +-- Margaritifera_margaritifera **
Chen et al., 2025 [Publ 4663], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 47 | 0 | 0 | -- |
+-- Gibbosula --+ +-- Margaritifera_margaritifera **
Dai et al., 2026 [Publ 4757], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 33 | 0 | 0 | -- |
+-- Gibbosula --+ +-- Margaritifera_margaritifera **
Wu et al., 2026 [Publ 4811], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 16 | 0 | 0 | -- |
+-- Margaritifera_margaritifera ** --+ +-- Gibbosula
Wu et al., 2026 [Publ 4811], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 94 | 0 | 0 | -- |
+-- Gibbosula --+ +-- Margaritifera_margaritifera **
Wu et al., 2026 [Publ 4825], fig. 6B
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 64 | 0 | 0 | -- |
+-- Gibbosula --+ +-- Margaritifera_margaritifera **
Prié et al., 2026 [Publ 4844], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 22 | 0 | 0 | -- |
+-------------------- Margaritifera_margaritifera ** | | +----------------- Echyridella | | | +----------------- Westralunio --+ | | | +-- Hyridella | +--------------+ | | +-- Cucumerunio | | | | +-- Velesunio +--+ +--+ | | +-- Alathyria +-----------+ | | +-- Lortiella | +--+ | +-- Microdontia | | +-------------- Diplodon | | +--+ +----------- Guyanhyrida | | | | +----- Iridea +--+ +--+ | | | +-- Prisodon | | +--+ | | +-- Triplodon +--+ | +-- Castalia_cordata | +--+ | | +-- Callonaia +--+ | +-- Castalia_ambigua +--+ +-- Castaliella
Prié et al., 2026 [Publ 4844], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- |
+-- Castalia --+ +-- Margaritifera_margaritifera **
