Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Unionini

subtribe ACULAMPROTULINA Huang & Wu in Huang et al., 2019: cladomics

1 genus

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 9 (9 total)
number of subtaxa (rank: genus) = 1
analyzed in 119 cladograms in 67 publications
maximum number of terminals in an analysis = 6 (mean = 2.5, median = 2)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 70
maximum number of included terminals = 6, (mean = 3.56 median = 4)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)
proportion of cladograms that support basic monophyly = 1

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Aculamprotulina cladogram distances
PCoA Ordination

NJ Tree of Aculamprotulina cladogram distances
NJ Tree

Cladograms

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
5 5 3 1 625 61 4 0 
  +----------- Globunionina      
--+                              
  |  +-------- Scriptolamprotula ** 
  +--+                           
     |  +----- Middendorffinaiina
     +--+                        
        |  +-- Unionina          
        +--+                     
           +-- Oxynaiina

Chen et al., 2026 [Publ 4827], fig. 2

iTtTbiriSoTroTroSiD
6 6 4 1 6 2 1 1 
  +-- Globunionina     
--+                    
  +-- Scriptolamprotula **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
5 5 3 1 117 23 3 4 
  +-------- Scriptolamprotula ** 
--+                           
  |  +----- Middendorffinaiina
  +--+                        
     |  +-- Unionina          
     +--+                     
        +-- Oxynaiina

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
5 5 3 1 196 16 2 2 
1

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
5 5 3 1 194 15 2 2 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
5 5 3 1 48 9 2 4 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Lopes-Lima et al., 2020 [Publ 4080], fig. 5

iTtTbiriSoTroTroSiD
5 5 3 1 4 2 1 0 
  +-- Oxynaiina        
--+                    
  +-- Scriptolamprotula **

Huang et al., 2019 [Publ 3843], fig. S1

iTtTbiriSoTroTroSiD
5 5 3 1 42 12 1 0 
  +-- Scriptolamprotula **
--+                    
  +-- Oxynaiina

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
5 5 3 1 41 12 1 4 
1

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
5 5 2 1 196 16 2 3 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
4 4 2 1 64 11 2 3 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
4 4 2 1 34 19 2 3 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
4 4 2 1 30 1 1 3 
  +-- Unionina         
--+                    
  +-- Scriptolamprotula **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 1 83 23 3 3 
  +-------- Globunionina     
--+                          
  |  +----- Scriptolamprotula **
  +--+                       
     |  +-- Unionina         
     +--+                    
        +-- Oxynaiina

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 1 40 17 4 3 
  +-------------------- Globunionina      
  |                                       
--+  +----------------- Scriptolamprotula ** 
  |  |                                    
  |  |              +-- Middendorffinaiina
  +--+  +-----------+                     
     |  |           +-- Nodularia         
     |  |                                 
     +--+  +----------- Cuneopsis         
        |  |                              
        |  |        +-- Diaurora          
        +--+--------+                     
           |        +-- Schistodesmus     
           |                              
           |  +-------- Pseudocuneopsis   
           +--+                           
              |  +----- Inversiunio       
              +--+                        
                 |  +-- Tchangsinaia      
                 +--+                     
                    +-- Unionina

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 1 81 23 2 3 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Wu et al., 2020 [Publ 4147], fig. 4

iTtTbiriSoTroTroSiD
4 4 2 1 10 1 1 3 
  +-- Unionina         
--+                    
  +-- Scriptolamprotula **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
4 4 2 1 69 11 2 3 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 1 44 9 2 3 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
4 4 2 1 91 18 2 3 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
4 4 2 1 136 42 4 3 
  +----------- Scriptolamprotula ** 
--+                              
  |  +-------- Globunionina      
  +--+                           
     |  +----- Oxynaiina         
     +--+                        
        |  +-- Middendorffinaiina
        +--+                     
           +-- Unionina

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
4 4 2 1 69 11 2 3 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 1 30 1 1 3 
  +-- Unionina         
--+                    
  +-- Scriptolamprotula **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 1 42 9 2 3 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
4 4 1 1 40 17 4 2 
  +----------------------- Globunionina      
  |                                          
--+  +-------------------- Scriptolamprotula ** 
  |  |                                       
  +--+  +----------------- Middendorffinaiina
     |  |                                    
     +--+  +-------------- Nodularia         
        |  |                                 
        +--+  +----------- Cuneopsis         
           |  |                              
           |  |        +-- Diaurora          
           +--+--------+                     
              |        +-- Schistodesmus     
              |                              
              |  +-------- Pseudocuneopsis   
              +--+                           
                 |  +----- Unionina          
                 +--+                        
                    |  +-- Tchangsinaia      
                    +--+                     
                       +-- Inversiunio

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
4 4 2 1 91 28 4 3 
  +----------- Globunionina      
--+                              
  |  +-------- Scriptolamprotula ** 
  +--+                           
     |  +----- Unionina          
     +--+                        
        |  +-- Middendorffinaiina
        +--+                     
           +-- Oxynaiina

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 1 69 16 2 3 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Wang et al., 2025 [Publ 4488], fig. 3B

iTtTbiriSoTroTroSiD
4 4 2 1 60 28 3 3 
  +-------- Scriptolamprotula ** 
--+                           
  |  +----- Unionina          
  +--+                        
     |  +-- Middendorffinaiina
     +--+                     
        +-- Oxynaiina

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
4 4 2 1 49 25 2 3 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Hou et al., 2025 [Publ 4445], fig. 5

iTtTbiriSoTroTroSiD
4 4 2 1 48 28 2 3 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 1 52 7 2 3 
  +----- Scriptolamprotula **
--+                       
  |  +-- Oxynaiina        
  +--+                    
     +-- Unionina

Wu et al., 2026 [Publ 4825], fig. 6B

iTtTbiriSoTroTroSiD
4 4 2 1 61 25 3 3 
  +-------- Scriptolamprotula ** 
--+                           
  |  +----- Unionina          
  +--+                        
     |  +-- Middendorffinaiina
     +--+                     
        +-- Oxynaiina

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 1 26 7 2 3 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
4 4 2 1 91 18 2 3 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
4 4 2 1 87 18 2 3 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
4 4 2 1 52 8 3 3 
  +-------- Middendorffinaiina
--+                           
  |  +----- Scriptolamprotula ** 
  +--+                        
     |  +-- Unionina          
     +--+                     
        +-- Oxynaiina

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 1 18 2 2 3 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
4 4 2 1 37 17 2 3 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
4 4 2 1 71 44 3 3 
  +-------- Scriptolamprotula ** 
--+                           
  |  +----- Unionina          
  +--+                        
     |  +-- Middendorffinaiina
     +--+                     
        +-- Oxynaiina

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
4 4 2 1 87 18 2 3 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 1 20 7 2 2 
  +----- Scriptolamprotula **
--+                       
  |  +-- Oxynaiina        
  +--+                    
     +-- Unionina

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 1 35 7 2 2 
  +----- Scriptolamprotula **
--+                       
  |  +-- Oxynaiina        
  +--+                    
     +-- Unionina

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 1 51 10 2 2 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 1 37 8 2 2 
  +----- Scriptolamprotula **
--+                       
  |  +-- Oxynaiina        
  +--+                    
     +-- Unionina

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 1 41 7 2 2 
  +----- Scriptolamprotula **
--+                       
  |  +-- Oxynaiina        
  +--+                    
     +-- Unionina

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 1 66 10 2 2 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 1 66 10 2 2 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Wu et al., 2024 [Publ 4379], fig. 4_6

iTtTbiriSoTroTroSiD
3 3 1 1 20 12 1 2 
1

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
3 3 1 1 41 7 2 2 
  +----- Scriptolamprotula **
--+                       
  |  +-- Oxynaiina        
  +--+                    
     +-- Unionina

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 1 18 2 2 2 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Zhou et al., 2007 [Publ 2658], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 1 15 2 1 2 
1

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
3 3 1 1 88 8 2 2 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 1 18 4 1 2 
  +-- Oxynaiina        
--+                    
  +-- Scriptolamprotula **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
3 3 1 1 88 8 2 2 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
3 3 1 1 39 6 2 2 
  +----- Scriptolamprotula **
--+                       
  |  +-- Oxynaiina        
  +--+                    
     +-- Unionina

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
3 3 1 1 45 7 2 2 
  +----- Scriptolamprotula **
--+                       
  |  +-- Oxynaiina        
  +--+                    
     +-- Unionina

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 1 28 2 2 2 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 1 66 10 2 2 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 30 5 2 -- 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Wen et al., 2017 [Publ 3795], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 14 1 1 -- 
1

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 1 51 16 2 -- 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Zhou et al., 2017 [Publ 3788], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 14 2 1 -- 
  +-- Oxynaiina        
--+                    
  +-- Scriptolamprotula **

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 1 21 11 2 -- 
  +----- Unionina         
--+                       
  |  +-- Scriptolamprotula **
  +--+                    
     +-- Oxynaiina

Wu et al., 2026 [Publ 4825], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 1 17 5 1 -- 
1

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 1 22 12 2 -- 
  +-- Unionina         
  |                    
--+-- Scriptolamprotula **
  |                    
  +-- Oxynaiina

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 32 1 1 -- 
1

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 1 35 5 2 -- 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 29 5 2 -- 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
2 2 0 1 37 5 2 -- 
1

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 1 22 12 2 -- 
  +----- Unionina         
--+                       
  |  +-- Scriptolamprotula **
  +--+                    
     +-- Oxynaiina

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 15 2 1 -- 
-- Scriptolamprotula_tientsinensis **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 124 9 2 -- 
  +-- Unio_crassus         
  |                        
  +-- Unio_pictorum        
  |                        
  +-- Unio_tumidus         
  |                        
  +-- Scriptolamprotula **    
  |                        
  +-- Cuneopsis_szechenyii 
--+                        
  +-- Tchangsinaia         
  |                        
  +-- Cuneopsis_rufescens  
  |                        
  +-- Nodularia_douglasiae 
  |                        
  +-- Nodularia_nuxpersicae
  |                        
  +-- Schistodesmus

Kim et al., 2021 [Publ 4197], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 17 11 2 -- 
-- Scriptolamprotula_tientsinensis **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
1 1 0 1 19 4 1 -- 
  +-- Scriptolamprotula **
--+                    
  +-- Unionina

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 7 2 -- 
  +----- Scriptolamprotula **
--+                       
  |  +-- Oxynaiina        
  +--+                    
     +-- Unionina

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
1 1 0 1 80 10 2 -- 
  +----- Unionina         
--+                       
  |  +-- Scriptolamprotula **
  +--+                    
     +-- Oxynaiina

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
1 1 0 1 138 9 2 -- 
  +----------- Unionina            
  |                                
--+  +-------- Scriptolamprotula **   
  |  |                             
  |  +-------- Schistodesmus       
  +--+                             
     +-------- Nodularia           
     |                             
     |  +----- Cuneopsis_szechenyii
     +--+                          
        |  +-- Tchangsinaia        
        +--+                       
           +-- Cuneopsis_rufescens

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
1 1 0 1 75 9 2 -- 
  +----- Unionina         
--+                       
  |  +-- Scriptolamprotula **
  +--+                    
     +-- Oxynaiina

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 4 2 -- 
  +----- Scriptolamprotula **
--+                       
  |  +-- Oxynaiina        
  +--+                    
     +-- Unionina

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
1 1 0 1 21 11 3 -- 
  +-------- Scriptolamprotula ** 
--+                           
  |  +----- Unionina          
  +--+                        
     |  +-- Middendorffinaiina
     +--+                     
        +-- Oxynaiina

Ouyang et al., 2015 [Publ 4493], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 30 7 1 -- 
           +-- Schistodesmus        
  +--------+                        
  |        +-- Scriptolamprotula **    
  |                                 
--+        +-- Tchangsinaia         
  |  +-----+                        
  |  |     +-- Cuneopsis_rufescens  
  +--+                              
     |  +----- Nodularia            
     +--+                           
        |  +-- Cuneopsis_szechenyii 
        +--+                        
           +-- Cuneopsis_celtiformis

Ouyang et al., 2011 [Publ 3408], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 12 2 1 -- 
-- Scriptolamprotula_tortuosa **

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
1 1 0 1 24 1 1 -- 
  +-- Scriptolamprotula **
--+                    
  +-- Unionina

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 4 2 -- 
  +----- Oxynaiina        
--+                       
  |  +-- Scriptolamprotula **
  +--+                    
     +-- Unionina

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 17 3 -- 
  +----------------- Scriptolamprotula ** 
  |                                    
  |              +-- Middendorffinaiina
--+  +-----------+                     
  |  |           +-- Nodularia         
  |  |                                 
  +--+  +----------- Cuneopsis         
     |  |                              
     |  |        +-- Diaurora          
     +--+--------+                     
        |        +-- Schistodesmus     
        |                              
        |  +-------- Pseudocuneopsis   
        +--+                           
           |  +----- Inversiunio       
           +--+                        
              |  +-- Tchangsinaia      
              +--+                     
                 +-- Unionina

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
1 1 0 1 75 9 2 -- 
  +----- Unionina         
--+                       
  |  +-- Scriptolamprotula **
  +--+                    
     +-- Oxynaiina

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 22 4 2 -- 
  +----- Oxynaiina        
--+                       
  |  +-- Scriptolamprotula **
  +--+                    
     +-- Unionina

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 17 3 -- 
  +----------------- Scriptolamprotula ** 
  |                                    
  |              +-- Middendorffinaiina
--+  +-----------+                     
  |  |           +-- Nodularia         
  |  |                                 
  +--+  +----------- Cuneopsis         
     |  |                              
     |  |        +-- Diaurora          
     +--+--------+                     
        |        +-- Schistodesmus     
        |                              
        |  +-------- Pseudocuneopsis   
        +--+                           
           |  +----- Inversiunio       
           +--+                        
              |  +-- Tchangsinaia      
              +--+                     
                 +-- Unionina

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iTtTbiriSoTroTroSiD
1 1 0 1 21 1 1 -- 
-- Scriptolamprotula_tortuosa **

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iTtTbiriSoTroTroSiD
1 1 0 1 12 2 1 -- 
  +-- Scriptolamprotula **
--+                    
  +-- Unionina

Ouyang et al., 2015 [Publ 4493], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 7 1 -- 
-- Scriptolamprotula_tortuosa **

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 16 12 2 -- 
-- Scriptolamprotula_tientsinensis **

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 40 17 3 -- 
  +-------- Globunionina     
--+                          
  |  +----- Scriptolamprotula **
  +--+                       
     |  +-- Unionina         
     +--+                    
        +-- Oxynaiina

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 6 1 -- 
-- Scriptolamprotula_tortuosa **

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 17 3 -- 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 6 1 1 -- 
  +-- Unionina         
--+                    
  +-- Scriptolamprotula **

Wu et al., 2022 [Publ 4238], fig. 2a_S1

iTtTbiriSoTroTroSiD
1 1 0 1 11 11 1 -- 
  +-- Scriptolamprotula **
--+                    
  +-- Oxynaiina

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
1 1 0 1 21 11 3 -- 
-- Scriptolamprotula_scripta **

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 6 1 1 -- 
  +-- Unionina         
--+                    
  +-- Scriptolamprotula **

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
1 1 0 1 24 1 1 -- 
-- Scriptolamprotula_tortuosa **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 4 2 -- 
-- Scriptolamprotula_tortuosa **

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iTtTbiriSoTroTroSiD
1 1 0 1 12 2 1 -- 
  +-- Scriptolamprotula **
--+                    
  +-- Unionina

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 4 2 -- 
  +----- Scriptolamprotula **
--+                       
  |  +-- Oxynaiina        
  +--+                    
     +-- Unionina

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
1 1 0 1 16 3 1 -- 
  +-- Scriptolamprotula **
--+                    
  +-- Unionina

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 138 9 2 -- 
  +----- Unionina         
--+                       
  |  +-- Scriptolamprotula **
  +--+                    
     +-- Oxynaiina

Dai et al., 2024 [Publ 4386], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 13 10 3 -- 
-- Scriptolamprotula_scripta **

Klymus et al., 2025 [Publ 4814], fig. 3B

iTtTbiriSoTroTroSiD
1 1 0 1 35 9 2 -- 
-- Scriptolamprotula_tortuosa **

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
1 1 0 1 14 2 1 -- 
-- Scriptolamprotula_tientsinensis **

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 13 2 2 -- 
  +-- Oxynaiina        
--+                    
  +-- Scriptolamprotula **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 6 2 -- 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Wu et al., 2022 [Publ 4238], fig. 2e

iTtTbiriSoTroTroSiD
1 1 0 1 17 12 2 -- 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Klymus et al., 2025 [Publ 4814], fig. 3A

iTtTbiriSoTroTroSiD
1 1 0 1 35 9 2 -- 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 46 13 2 -- 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
1 1 0 1 14 2 1 -- 
-- Scriptolamprotula_tientsinensis **

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
1 1 0 1 28 1 1 -- 
  +-- Unionina         
--+                    
  +-- Scriptolamprotula **

Xue et al., 2016 [Publ 3659], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 14 3 2 -- 
  +-------- Scriptolamprotula **
--+                          
  |  +----- Schistodesmus    
  +--+                       
     |  +-- Nodularia        
     +--+                    
        +-- Unionina

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iTtTbiriSoTroTroSiD
1 1 0 1 9 4 1 -- 
  +-- Scriptolamprotula **
--+                    
  +-- Unionina

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 52 8 2 -- 
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 6 2 -- 
  +----- Scriptolamprotula **
--+                       
  |  +-- Oxynaiina        
  +--+                    
     +-- Unionina