Bivalvia
| Unionida
| Unionoidea
| Unionidae
subfamily AMBLEMINAE Rafinesque, 1820: cladomics
5 tribes
Cladomics pages are experimental — even more so than the rest of this site!
The purpose of the cladomics pages for each genus and family-group level taxon is to
provide statistics and references indicating the extent to which published phylogenetic
work supports the classification used here.
The Cladomics Summary provides an
overview of the number of studies that have included terminal species representing this
taxon as well as an indication of how frequently it has been recovered as monophyletic.
Following the summary, the available published clades representing the taxon are listed.
For more information, visit the
Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various
statistics are explained on the FARQ!
Cladomics Summary
number of species = 308 (393 total)
number of subtaxa (rank: tribe) = 5
analyzed in 319 cladograms in 190 publications
maximum number of terminals in an analysis = 191 (mean = 11.3, median = 6)
maximum number of subtaxa (tribe) = 5 (mean = 2.6, median = 2)
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 275
maximum number of included terminals = 191, (mean = 12.95 median = 7)
maximum number of subtaxa (tribe) = 5 (mean = 2.85, median = 2)
proportion of cladograms that support basic monophyly = 0.88
Rigorous Monophyly
number of cladograms that test rigorous monophyly = 192
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (tribe) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 191 (mean = 11.46, median = 6)
maximum number of subtaxa (tribe) = 5 (mean = 3.05, median = 2.5)
proportion of cladograms that support rigorous monophyly = 0.83
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome.
See the Cladmoics FARQ for more information.
Cladograms
** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **
iT| tT | bi | riS | oT | roT | roS | iD |
| 191 |
582 |
651 |
5 |
441 |
376 |
3 |
0 |
+-------------------------------- Lamellidens
|
+-----+ +-- Balwantia
| | +--------------------------+
| | | +-- Trapezidens
| +--+
| | +-------------------------- Leoparreysiini
| | |
| +--+ +----------------------- Pseudomulleria
| | |
| | | +-- Parreysiini
| +--+ +-----------------+
| | | +-- Coelaturini
| | |
| +--+ +-- Pseudoscabies
| | +--------------+
| | | +-- Indonaia
| +--+
| | +-------------- Indochinella
| | |
| +--+ +----------- Radiatula
+--------------------------------------------------+ | |
| | +--+ +-- Songkhlanaia_songkhramensis__2
| | | +-----+
| | | | +-- Unionetta_fabagina__2
| | +--+
| | | +----- Harmandia
| | +--+
| | | +-- Unionetta_fabagina__1
| | +--+
| | +-- Scabies
| |
| | +----- Amuranodonta
| | +-----------------------------+
| | | | +-- Anemina
| | | +--+
| | | +-- Buldowskia
| | |
| | | +-- Beringiana
| | | +-----+
| | | | +-- Pletholophus
| +--+ +-----------------------+
| | | | +----- Mimo
| | | +--+
| | | | +-- Acudonta
| | | +--+
| | | +-- Sinanodonta
| | |
| | | +----------------------------- Cristaria
| +--+ |
| | | +----------------- Lepidodesmini
| | | |
| | | | +----------- Lanceolariini
| | | | +--+
| | | +--------+ | | +-------- Pseudoacuticosta_retiaria__2
| | | | | | +--+
| | | | | | | +----- Acuticosta_chinensis__2
| +--+ | | | +--+
| | | +--+ | +-- Acuticosta_chinensis__1
| | | | +--+
| | | | +-- Pseudoacuticosta_retiaria__1
| | | |
| | | | +----------- Globunio
| | | +--+
| | | | +-------- Aculamprotulina
| | | +--+
| +--+ | +----- Middendorffinaiina
| | +--+
| | | +-- Unionina
| | +--+
| | +-- Oxynaiina
| |
| | +-------- Pseudodontoideus
--+ | +--------------+
| | | | +----- Lasmigona
| | | +--+
| | | | +-- Pterosyna
| | | +--+
| +--+ +-- Arcidens
| |
| | +-------------------- Anodontina
| | |
| | | +-------------- Utterbackia
| | | +--+
| +--+ | | +----------- Pyganodon_grandis
| | | +--+
| | | | +-------- Pyganodon_lacustris
| | | +--+
| | | | +----- Pyganodon_cataracta__2
| +--+ +--+
| | | +-- Pyganodon_cataracta__1
| | +--+
| | +-- Potamilus_fragilis__3 **
| |
| | +-------------- Utterbackiana
| | |
| +--+ +----------- Simpsonaias
| | |
| | | +-- Alasminota
| +--+ +-----+
| | | +-- Alasmidonta
| | |
| +--+ +-- Prolasmidonta
| | +--+
| | | +-- Platynaias
| +--+
| | +-- Anodontoides
| +--+
| +-- Strophitus
|
| +-------------------------------------------------------------------------------------- Rectidentini
| |
| | +-------- Pseudunio
| | +--------------------------------------------------------------------------+
| | | | +----- Gibbosulinae
| | | +--+
| | | | +-- Cumberlandia
| | | +--+
| | | +-- Margaritifera
| | |
| | | +-------------------- Chamberlainiini
| | | |
| | | +--+ +----------------- Parvasolenaia
| | | | | |
| | | | +--+ +-------------- Pseudopostolata
| | | | | |
+--+ | | +--+ +----------- Koreosolenaia
| | | | |
| | | +--+ +-- Cosmopseudodon
| | | | +-----+
| | | | | +-- Guiunio
| | | +--+
| | +--------+ | +----- Ptychorhynchus
| | | | +--+
| | | | | +-- Obovalis
| | | | +--+
| | | | +-- Postolata
| | | |
| | | | +-- Caudiculatus
| | | | +--------------+
| | | | | +-- Lamprotulini
| | | | |
| | | +-----+ +-------------- Sinosolenaia_carinata
+--+ | | |
| | | | +----- Gonidea
| | +--+ +-----+
| +-----------------------------------------------+ | | | +-- Microcondylaea
| | | | | +--+
| | | +--+ +-- Leguminaia
| | | |
| | | | +-------- Inversidens
| | | +--+
| | | | +----- Sinosolenaia_recognita
| | | +--+
| | | | +-- Sinosolenaia_iridinea
| | | +--+
| | | +-- Sinosolenaia_oleivora
| | |
| | | +----------------------------- Indopseudodontina
| | | |
| | | | +-- Lannanaia
| | +--+ +-----------------------+
| | | | +-- Pilsbryoconcha
| | | |
| | | | +----- Songkhlanaia_tamodienica
| | +--+ +-----------------+
| | | | | +-- Songkhlanaia_moreleti
| | | | +--+
| | | | +-- Songkhlanaia_songkhramensis__1
| | | |
| | +--+ +-- Nyeinchanconcha
| | | +-----------------+
| | | | +-- Sundadontina
| | | |
+--+ | | +-- Isannaia
| +--+ +--+
| | | +-- Namkongnaia
| | +-----------+
| | | | +-- Thaiconcha
| | | +--+
| +--+ +-- Bineurus
| |
| | +-------------- Pseudodon_walpolei
| | |
| +--+ +----------- Pseudodon_vagulus
| | |
| | | +----- Pseudodon_vondembuschianus
| +--+ +--+
| | | | +-- Pseudodon_mekongi
| | | +--+
| +--+ +-- Schepmaniini
| |
| | +----- Pseudodon_lenyanensis__2
| +--+
| | +-- Pseudodon_inoscularis
| +--+
| +-- Pseudodon_lenyanensis__1
|
| +-- Quadrulini **
| +--------------------------------------------------------------------------+
| | +-- Pleurobemini **
| |
| | +----- Amblema **
+--+ +--------------------------------------------------------------------+
| | | +-- Nephronaias **
| | +--+
| | +-- Psoronaias **
| |
+--+ +----------------------------------------------------------------------- Reginaia **
| |
| | +----- Popenaias **
| | +--------------------------------------------------------------+
| | | | +-- Cyrtonaias **
+--+ | +--+
| | +-- Glebula **
| |
| | +----------------------------------------------------------------- Paetulunio **
| | |
+--+ | +----------------- Toxolasma_parvum **
| | |
| | | +----- Toxolasma_lividum__2 **
| | | +--------+
| | +--------------------------------------------+ | | +-- Toxolasma_cylindrellus **
| | | | | +--+
| | | | | +-- Toxolasma_lividum__1 **
| | | | |
+--+ | +--+ +-------- Toxolasma_texasiense__2 **
| | | +--+
| | | | | +----- Toxolasma_glans **
| | | | +--+
| | | | | +-- Toxolasma_paulum **
| | +--+ +--+
| | | +-- Toxolasma_texasiense__1 **
| | |
| | | +-------- Toxolasma__4 **
| | +--+
+--+ | +----- Toxolasma_granulatum **
| +--+
| | +-- Toxolasma__2 **
| +--+
| +-- Toxolasma__3 **
|
| +----- Toxolasma_pullus **
| +-----------------------------------------------------+
| | | +-- Toxolasma_germanum **
| | +--+
| | +-- Toxolasma__1 **
| |
| | +-- Obliquaria **
| | +-----------------------------------------------------+
| | | +-- Plectomerus **
+--+ |
| | +-------- Potamilus_inflatus **
| | +--------+
| | | | +----- Potamilus_ohiensis **
| | | +--+
| | | | +-- Potamilus_amphichaenus **
| | | +--+
| | +-----------------------------------+ +-- Potamilus_streckersoni **
| | | |
+--+ | | +----- Potamilus_fragilis__2 **
| | | +--------+
| | | | | +-- Potamilus_leptodon **
| | | | +--+
| | +--+ +-- Potamilus_fragilis__1 **
| | |
| | | +----------- Potamilus_metnecktayi **
| | | |
| | +--+ +-------- Potamilus_alatus__3 **
| | | |
| | +--+ +----- Potamilus_alatus__2 **
+--+ | |
| +--+----- Potamilus_purpuratus__2 **
| |
| | +-- Potamilus_alatus__1 **
| +--+
| +-- Potamilus_purpuratus__1 **
|
| +-- Villosa_villosa **
| +-----------------------------------------------+
| | +-- Atlanticoncha **
| |
| | +-- Cambarunio_nebulosus **
| | +--------------------------------------------+
+--+ | +-- Ligumia **
| |
| | +-------- Leaunio_umbrans **
| | +-----------------------------------+
| | | | +----- Cambarunio_taeniatus **
+--+ | +--+
| | | +-- Cambarunio_iris **
| | +--+
| | +-- Cambarunio_simus **
| |
| | +----------------------------------------- Hamiota **
| | |
+--+ | +----------------------- Pachynaias **
| | |
| | +--------------+ +-------------------- Sagittunio **
| | | | |
| | | | | +----- Ellipsaria **
| | | +--+ +-----------+
| | | | | | +-- Potamilus_capax **
| | | | | +--+
| | | | | +-- Truncilla **
+--+ | +--+
| | | +----- Medionidus_acutissimus **
| | | +--------+
| | | | | +-- Lemiox **
| | | | +--+
| | +--+ +-- Medionidus_penicillatus **
| | |
| | | +----------- Ptychobranchus **
| | | |
| | +--+ +-- Cyprogenia_aberti__2 **
| | | +-----+
+--+ | | +-- Dromus **
| +--+
| | +----- Cyprogenia_stegaria__2 **
| +--+
| | +-- Cyprogenia_aberti__1 **
| +--+
| +-- Cyprogenia_stegaria__1 **
|
| +-- Villosa_delumbis **
| +--------------------------------+
| | +-- Villosa_vibex **
| |
| | +-------------- Obovaria_olivaria **
| | |
| | +-----------------+ +-- Venustaconcha **
| | | | +--------+
| | | | | +-- Epioblasma **
| | | +--+
| | | | +-------- Obovaria_retusa **
+--+ | +--+
| | | +----- Obovaria_choctawensis **
| | +--+
| | | +-- Obovaria_unicolor **
| | +--+
| | +-- Obovaria_subrotunda **
| |
| | +-- Mesonaias **
| | +-----------------------+
| | | +-- Obovaria_arkansasensis **
| | |
+--+ +--+ +-- Lampsilis_splendida **
| | | +--------------------+
| | | | +-- Lampsilis_bracteata **
| | | |
| | +--+ +-- Lampsilis_fasciola **
| | | +-----------------+
| | | | +-- Ortmanniana_pectorosa **
| | +--+
| | | +----------------- Lampsilis_ornata **
| | | |
| | +--+ +-------------- Lampsilis_ovata__2 **
| | | |
| | +--+ +----------- Lampsilis_dolabraeformis **
| | | |
| | | | +----- Lampsilis_cardium__2 **
| | +--+ +--+
| | | | | +-- Lampsilis_cardium__1 **
+--+ | | +--+
| +--+ +-- Lampsilis_satura **
| |
| | +----- Lampsilis_cariosa__2 **
| +--+
| | +-- Lampsilis_ovata__1 **
| +--+
| +-- Lampsilis_cariosa__1 **
|
| +----------- Ortmanniana_ligamentina__2 **
| +--------------+
| | | +-------- Lampsilis_abrupta **
| | +--+
| | | +----- Lampsilis_higginsii__2 **
| | +--+
| | | +-- Ortmanniana_ligamentina__1 **
| | +--+
+--+ +-- Lampsilis_higginsii__1 **
|
| +----- Medionidus_conradicus **
| +-----------------+
| | | +-- Leaunio_lienosus **
| | +--+
| | +-- Leaunio_vanuxemensis **
| |
+--+ +-- Lampsilis_floridensis **
| +-----------------+
| | +-- Lampsilis_teres **
| |
| | +-- Lampsilis_siliquoidea__1 **
+--+ |
| +-- Lampsilis_siliquoidea__2 **
| +--------------+
| | +-- Lampsilis_powellii__1 **
| | |
+--+ +-- Lampsilis_powellii__2 **
|
| +-------------- Lampsilis_sietmani **
| |
+--+ +-- Lampsilis_straminea__2 **
| +--------+
| | +-- Lampsilis_virescens **
+--+
| +-------- Lampsilis_hydiana__2 **
+--+
| +----- Lampsilis_hydiana__1 **
+--+
| +-- Lampsilis_bergmanni **
+--+
+-- Lampsilis_straminea__1 **
iT| tT | bi | riS | oT | roT | roS | iD |
| 90 |
90 |
92 |
5 |
12 |
10 |
2 |
703 |
+-------------------- Chamberlainiini
|
+-------------------- Unioninae
--+
| +-- Gonideini
| +--------------+
| | +-- Lamprotulini
+--+
| +-------------- Quadrulini **
| |
+--+ +----------- Pleurobemini **
| |
+--+ +-- Plectomerus **
| +-----+
| | +-- Reginaia **
+--+
| +----- Lampsilini **
+--+
| +-- Popenaiadini **
+--+
+-- Amblema **
iT| tT | bi | riS | oT | roT | roS | iD |
| 90 |
90 |
81 |
5 |
12 |
10 |
2 |
683 |
+-- Chamberlainiini
+-----------------------------+
| +-- Unioninae
|
--+ +-- Gonideini
| +--------------------------+
| | +-- Lamprotulini
| |
+--+ +-------------------------- Quadrulini **
| |
| | +----------------------- Pleurobemini **
+--+ |
| | +----- Popenaiadini **
| | +--------------+
| | | | +-- Amblema **
+--+ | +--+
| | +-- Reginaia **
| |
| | +-- Plectomerus **
| | +--------------+
+--+ | +-- Toxolasma **
| |
| | +----- Obliquaria **
| | +--------+
| | | | +-- Cyrtonaias **
| | | +--+
| | | +-- Glebula **
| | |
+--+ | +----------- Sagittunio **
| | |
| | +----------- Leaunio **
| | |
| | +----------- Potamilus **
| | |
| | +----------- Hamiota **
| | |
| | | +-- Ellipsaria **
+--+ +--------+
| | +-- Truncilla **
| |
| | +----- Lampsilis_teres **
| +-----+
| | | +-- Ortmanniana_ligamentina **
| | +--+
| | +-- Lampsilis_siliquoidea **
| |
| | +----- Villosa **
| +-----+
| | | +-- Ligumia **
| | +--+
| | +-- Cambarunio **
| |
+--+ +-------- Ortmanniana_pectorosa **
+--+
| | +----- Lampsilis_ornata **
| +--+
| | +-- Lampsilis_cardium **
| +--+
| +-- Lampsilis_ovata **
|
| +-------- Obovaria_olivaria **
| |
| +-------- Venustaconcha_ellipsiformis__2 **
+--+
| +-------- Epioblasma **
| |
| | +----- Obovaria_unicolor **
| +--+
| | +-- Obovaria_arkansasensis **
| +--+
| +-- Obovaria_subrotunda **
|
| +-- Ptychobranchus **
| +-----+
| | +-- Venustaconcha_ellipsiformis__1 **
| |
+--+ +----- Medionidus_conradicus **
| |
| | +-- Lemiox **
+--+--+
| +-- Medionidus_acutissimus **
|
| +-- Dromus **
+--+
+-- Cyprogenia **
iT| tT | bi | riS | oT | roT | roS | iD |
| 83 |
83 |
59 |
5 |
22 |
21 |
2 |
640 |
+-------------- Unioninae
|
| +----------- Gonideinae
--+ |
| | +-------- Reginaia **
| | |
+--+ +-------- Amblema **
| |
| | +-- Quadrulini **
| +-----+
| | +-- Pleurobemini **
+--+
| +----- Popenaiadini **
| |
| | +-- Dromus **
| | |
| | +-- Epioblasma **
| | |
+--+ +-- Glebula **
| |
| +-- Lampsilis_ovata **
| |
| +-- Lampsilis_virescens **
| |
| +-- Lemiox **
+--+
+-- Obliquaria **
|
+-- Obovaria_olivaria **
|
+-- Obovaria_unicolor **
|
+-- Plectomerus **
|
+-- Potamilus_alatus **
|
+-- Potamilus_capax **
iT| tT | bi | riS | oT | roT | roS | iD |
| 73 |
73 |
63 |
5 |
0 |
0 |
0 |
691 |
+-------------- Quadrulini **
|
--+ +----------- Pleurobemini **
| |
+--+ +-- Plectomerus **
| +-----+
| | +-- Reginaia **
+--+
| +----- Lampsilini **
+--+
| +-- Popenaiadini **
+--+
+-- Amblema **
iT| tT | bi | riS | oT | roT | roS | iD |
| 73 |
73 |
67 |
5 |
0 |
0 |
0 |
681 |
+----------------------------------- Quadrulini **
|
--+ +-------------------------------- Pleurobemini **
| |
| | +-- Amblema **
+--+ +--------------------------+
| | +-- Reginaia **
| |
+--+ +-------------------------- Popenaiadini **
| |
| | +-- Glebula **
+--+ +--------------------+
| | +-- Paetulunio **
| |
+--+ +-- Toxolasma **
| +-----------------+
| | +-- Plectomerus **
| |
+--+ +----------------- Obliquaria **
| |
| | +----- Potamilus **
+--+ +--------+
| | | +-- Dromus **
| | +--+
+--+ +-- Lemiox **
|
| +----------- Lampsilis_ovata **
| |
+--+ +-------- Lampsilis_virescens **
| |
+--+ +-- Epioblasma **
| +--+
| | +-- Obovaria_arkansasensis **
+--+
| +-- Obovaria_olivaria **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 55 |
55 |
47 |
4 |
2 |
1 |
1 |
688 |
+----- Pleurobemini **
|
--+ +-- Amblemini **
| |
+--+-- Popenaiadini **
|
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 54 |
54 |
49 |
4 |
2 |
1 |
1 |
692 |
+----------- Unioninae
--+
| +-------- Lampsilini **
+--+
| +----- Pleurobemini **
+--+
| +-- Amblemini **
+--+
+-- Popenaiadini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 43 |
43 |
42 |
5 |
0 |
0 |
0 |
685 |
+-------------------------------- Quadrulini **
|
| +-------------- Elliptoideus **
| +--------------+
| | | +----------- Plectomerus **
--+ | +--+
| | | +-------- Obovaria_unicolor **
| | +--+
| | | +----- Obovaria_olivaria **
| | +--+
| | | +-- Lampsilis **
+--+ +--+
| +-- Lemiox **
|
| +-------- Cyrtonaias **
| +-----------------+
| | | +----- Reginaia **
| | +--+
| | | +-- Popenaiadini **
+--+ +--+
| +-- Amblema **
|
| +----------------------- Elliptio **
| |
+--+ +-------------------- Eurynia **
| |
| | +----------------- Parvaspina **
+--+ |
| | +-- Hemistena **
| +--------------+
+--+ +-- Plethobasus **
|
| +-------------- Pleuronaia **
| |
| | +-- Pleurobema_cordatum **
+--+ +--------+
| | +-- Fusconaia **
| |
+--+ +-- Pleurobema_stabile **
| +-----+
| | +-- Pleurobema_sintoxia **
+--+
| +----- Pleurobema_decisum **
+--+
| +-- Pleurobema_clava **
+--+
+-- Pleurobema_pyriforme **
iT| tT | bi | riS | oT | roT | roS | iD |
| 42 |
42 |
36 |
5 |
0 |
0 |
0 |
672 |
+-------------------- Quadrulini **
|
--+ +----------------- Pleurobemini **
| |
+--+ +-- Reginaia **
| +-----------+
| | +-- Amblema **
+--+
| +----------- Popenaiadini **
| |
+--+ +-------- Cyrtonaias **
| |
+--+-------- Plectomerus **
|
| +----- Lemiox **
+--+
| +-- Lampsilis **
+--+
+-- Obovaria **
iT| tT | bi | riS | oT | roT | roS | iD |
| 42 |
42 |
40 |
5 |
0 |
0 |
0 |
680 |
+----------------- Quadrulini **
|
--+ +-------------- Pleurobemini **
| |
| | +-- Popenaiadini **
+--+ +--------+
| | +-- Amblema **
| |
+--+ +-------- Plectomerus **
| |
| | +-- Cyrtonaias **
+--+-----+
| +-- Reginaia **
|
| +----- Lemiox **
+--+
| +-- Lampsilis **
+--+
+-- Obovaria **
iT| tT | bi | riS | oT | roT | roS | iD |
| 42 |
42 |
41 |
5 |
0 |
0 |
0 |
681 |
+-------------------- Quadrulini **
|
--+ +----------------- Pleurobemini **
| |
| | +-- Popenaiadini **
+--+ +-----------+
| | +-- Amblema **
| |
+--+ +-- Cyrtonaias **
| +--------+
| | +-- Reginaia **
+--+
| +-------- Plectomerus **
+--+
| +----- Lemiox **
+--+
| +-- Lampsilis **
+--+
+-- Obovaria **
iT| tT | bi | riS | oT | roT | roS | iD |
| 41 |
41 |
40 |
4 |
0 |
0 |
0 |
694 |
+-------------------------------------- Quadrulini **
|
| +-- Amblema **
--+ +--------------------------------+
| | +-- Reginaia **
| |
+--+ +-------------------------------- Plectomerus **
| |
| | +----------------------------- Toxolasma **
+--+ |
| | +----- Popenaiadini **
| | +--------------------+
+--+ | | +-- Glebula **
| | +--+
| | +-- Cyrtonaias **
| |
+--+ +----------------------- Obliquaria **
| |
| | +-- Medionidus **
| | +--+
| | | +-- Lemiox **
+--+ +--------------+
| | | +-- Cyprogenia **
| | +--+
| | +-- Ptychobranchus **
| |
+--+ +-------- Potamilus **
| +--------+
| | | +----- Atlanticoncha **
| | +--+
| | | +-- Ellipsaria **
| | +--+
+--+ +-- Truncilla **
|
| +-- Hamiota **
| +-----------+
| | +-- Villosa **
| |
+--+ +----------- Ligumia **
| |
| | +----- Venustaconcha **
+--+ +--+
| | | +-- Obovaria **
| | +--+
+--+ +-- Epioblasma **
|
| +----- Ortmanniana_pectorosa **
+--+
| +-- Ortmanniana_ligamentina **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 41 |
41 |
41 |
4 |
1 |
1 |
1 |
670 |
+----------- Unioninae
--+
| +-------- Quadrulini **
+--+
| +----- Pleurobemini **
+--+
| +-- Amblemini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 41 |
41 |
42 |
4 |
0 |
0 |
0 |
694 |
+----------- Amblema **
|
--+ +-- Lampsilis **
| +-----+
| | +-- Plectomerus **
+--+
| +----- Popenaiadini **
+--+
| +-- Cyrtonaias **
+--+
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 40 |
40 |
40 |
4 |
1 |
1 |
1 |
682 |
+-------------- Unioninae
--+
| +----------- Quadrulini **
+--+
| +-------- Reginaia **
+--+
| +----- Pleurobemini **
+--+
| +-- Amblema **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 37 |
37 |
34 |
2 |
0 |
0 |
0 |
694 |
+-- Amblema **
|
--+-- Plectomerus **
|
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 37 |
37 |
33 |
4 |
1 |
1 |
1 |
681 |
+-------------------------------- Unioninae
|
--+ +----------------------------- Amblemini **
| |
+--+ +-------------------------- Lampsilini **
| |
+--+ +----------------------- Elliptoideus **
| |
+--+ +-------------------- Quadrulini **
| |
+--+ +----------------- Elliptio **
| |
| | +-------------- Parvaspina **
+--+ |
| +-------------- Plethobasus **
| |
| | +-- Hemistena **
+--+-----------+
| +-- Eurynia **
|
| +----------- Fusconaia **
| |
| | +----- Pleurobema_riddellii **
+--+ +--+
| | | +-- Pleurobema_sintoxia **
| | +--+
+--+ +-- Pleurobema_cordatum **
|
| +----- Pleuronaia **
| |
+--+ +-- Pleurobema_pyriforme **
| |
+--+-- Pleurobema_strodeanum **
|
+-- Pleurobema_clava **
iT| tT | bi | riS | oT | roT | roS | iD |
| 36 |
36 |
26 |
2 |
0 |
0 |
0 |
693 |
+-- Amblemini **
--+
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 35 |
35 |
30 |
1 |
0 |
0 |
0 |
686 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 33 |
33 |
21 |
5 |
0 |
0 |
0 |
684 |
+-------------------------- Quadrulini **
|
| +----------------------- Plectomerus **
--+ |
| | +-------------------- Popenaiadini **
| | |
| | +-------------------- Pleurobemini **
+--+ |
| | +-- Cyrtonaias **
| | +--+
| | | +-- Glebula **
| +--------------+
| | | +-- Reginaia **
+--+ +--+
| +-- Obliquaria **
|
| +----------------- Toxolasma **
| |
| | +-------------- Potamilus **
| | |
| | | +-- Ellipsaria **
| | +-----------+
+--+ | +-- Truncilla **
| |
| | +----- Medionidus **
| +--------+
| | | +-- Lemiox **
| | +--+
+--+ +-- Dromus **
|
| +----------- Cambarunio **
| |
| +----------- Leaunio **
| |
| +----------- Villosa **
| |
| | +-- Sagittunio **
| +--------+
+--+ +-- Lampsilis_ovata **
|
| +----- Venustaconcha **
+-----+
| | +-- Epioblasma **
| +--+
| +-- Obovaria **
|
| +-------- Lampsilis_teres **
+--+
| +----- Lampsilis_siliquoidea **
+--+
| +-- Lampsilis_hydiana **
+--+
+-- Lampsilis_straminea **
iT| tT | bi | riS | oT | roT | roS | iD |
| 30 |
30 |
31 |
3 |
0 |
0 |
0 |
688 |
+-------------------------------- Lampsilini **
|
--+ +----------------------------- Parvaspina **
| |
| | +-------------------------- Amblemini **
+--+ |
| | +----- Eurynia **
| | +-----------------+
+--+ | | +-- Plethobasus **
| | +--+
| | +-- Elliptio **
| |
| | +-------------------- Pleurobema_plenum **
+--+ |
| +-------------------- Pleuronaia **
| |
| | +-- Pleurobema_cordatum__1 **
| | +-----+
| | | +-- Fusconaia **
+--+ +--------+
| | | +----- Pleurobema_sintoxia__1 **
| | | |
| | +--+ +-- Pleurobema_cordatum__2 **
| | | |
| | +--+-- Pleurobema_sintoxia__2 **
| | |
+--+ +-- Pleurobema_sintoxia__3 **
|
| +-------------- Pleurobema_stabile **
| |
| | +-- Pleurobema_rubellum **
| | +-----+
| | | +-- Pleurobema_strodeanum **
+--+ +--+
| | | +----- Pleurobema_beadleianum **
| | +--+
| | | +-- Pleurobema_perovatum **
| | +--+
+--+ +-- Pleurobema_taitianum **
|
| +-- Pleurobema_clava **
| +-----+
| | +-- Pleurobema_oviforme **
| |
+--+ +-- Pleurobema_decisum **
| +--+
| | +-- Pleurobema_hanleyianum **
+--+
| +-- Pleurobema_pyriforme **
+--+
+-- Pleurobema_georgianum **
iT| tT | bi | riS | oT | roT | roS | iD |
| 30 |
30 |
29 |
5 |
1 |
1 |
1 |
690 |
+-------------- Gonideinae
--+
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 29 |
29 |
20 |
4 |
0 |
0 |
0 |
689 |
+----------- Lampsilis **
|
| +-------- Obliquaria **
| |
| +-------- Amblema **
| |
--+ +-------- Reginaia **
| |
| +-------- Potamilus **
| |
| +-------- Quadrulini **
| |
+--+ +-- Elliptoideus **
+-----+
| +-- Plectomerus **
|
| +----- Medionidus **
+--+
| | +-- Dromus **
| +--+
| +-- Cyprogenia **
|
| +----- Hemistena **
+--+
| +-- Elliptio **
+--+
+-- Fusconaia **
iT| tT | bi | riS | oT | roT | roS | iD |
| 29 |
29 |
25 |
4 |
9 |
9 |
1 |
688 |
+----------------------------------- Unioninae
|
--+ +-------------------------------- Quadrulina **
| |
+--+ +----------------------------- Megalonaias **
| |
+--+ +-------------------------- Toxolasma **
| |
+--+ +----------------------- Amblema **
| |
| | +-------------------- Pleurobemini **
+--+ |
| | +-- Reginaia **
| +-----------------+
+--+ +-- Obliquaria **
|
| +----------------- Potamilus **
| |
| | +-- Ellipsaria **
+--+ +-----------+
| | +-- Truncilla **
| |
+--+ +----------- Lampsilis_cardium **
| |
| | +----- Ligumia **
| +-----+
+--+ | +-- Obovaria **
| +--+
| +-- Epioblasma **
|
| +-------- Lampsilis_teres **
+--+
| +----- Lampsilis_siliquoidea **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis_higginsii **
iT| tT | bi | riS | oT | roT | roS | iD |
| 29 |
29 |
27 |
4 |
0 |
0 |
0 |
694 |
+-------------------------- Lampsilis **
|
--+ +----------------------- Potamilus **
| |
| | +----- Medionidus **
+--+ +--------------+
| | | +-- Dromus **
| | +--+
+--+ +-- Cyprogenia **
|
| +----------------- Obliquaria **
| |
+--+ +-------------- Amblema **
| |
| | +----- Reginaia **
+--+ +-----+
| | | +-- Elliptoideus **
| | +--+
+--+ +-- Plectomerus **
|
| +-------- Quadrulini **
+--+
| +----- Hemistena **
+--+
| +-- Elliptio **
+--+
+-- Fusconaia **
iT| tT | bi | riS | oT | roT | roS | iD |
| 29 |
29 |
22 |
4 |
6 |
6 |
1 |
683 |
+----------- Unioninae
|
--+ +-------- Reginaia **
| |
+--+ +-- Pleurobemini **
| +--+
| | +-- Quadrulini **
+--+
| +-- Amblema **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 28 |
28 |
26 |
5 |
0 |
0 |
0 |
693 |
+----------- Quadrulini **
--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 28 |
28 |
26 |
5 |
0 |
0 |
0 |
693 |
+----------- Quadrulini **
--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 26 |
26 |
23 |
3 |
0 |
0 |
0 |
694 |
+-- Lampsilini **
|
--+-- Amblemini **
|
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 26 |
26 |
18 |
3 |
0 |
0 |
0 |
692 |
+-- Lampsilini **
|
--+-- Amblemini **
|
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 26 |
26 |
24 |
5 |
1 |
1 |
1 |
689 |
+-------------- Gonideinae
--+
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Lampsilini **
+--+
+-- Popenaiadini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 25 |
25 |
9 |
4 |
0 |
0 |
0 |
694 |
+----------------------- Quadrulini **
|
+----------------------- Amblemini **
|
| +-------------------- Lampsilini **
| |
| +-------------------- Eurynia **
| |
| +-------------------- Fusconaia_cerina **
--+ |
| +-------------------- Fusconaia_flava **
| |
| +-------------------- Hemistena **
| |
| +-------------------- Plethobasus **
| |
| +-------------------- Parvaspina **
| |
+--+-------------------- Pleurobema_cordatum **
|
+-------------------- Pleuronaia **
|
+-------------------- Pleurobema_oviforme__2 **
|
+-------------------- Pleurobema_stabile **
|
+-------------------- Elliptio **
|
| +----------------- Pleurobema_decisum **
| |
| +----------------- Pleurobema_sintoxia **
| |
| +----------------- Pleurobema_clava **
+--+
+----------------- Pleurobema_oviforme__1 **
|
| +-------------- Pleurobema_beadleianum **
| |
+--+ +----------- Pleurobema_pyriforme **
| |
+--+ +-------- Pleurobema_rubellum **
| |
| +-------- Pleurobema_hanleyianum__2 **
+--+
+-------- Pleurobema_georgianum__2 **
|
| +----- Pleurobema_hanleyianum__1 **
+--+
| +-- Pleurobema_georgianum__1 **
+--+
+-- Pleurobema_athearni **
iT| tT | bi | riS | oT | roT | roS | iD |
| 25 |
25 |
20 |
4 |
175 |
136 |
3 |
682 |
+-------------------------------------- Parreysiinae
|
--+ +----------------------------------- Unioninae
| |
+--+ +-------------------------------- Rectidentini
| |
| | +-------- Chamberlainiini
+--+ +--------------------+
| | | +----- Lamprotulini
| | +--+
| | | +-- Gonideini
+--+ +--+
| +-- Pseudodontini
|
| +-------------------------- Quadrulini **
| |
+--+ +----------------------- Pleurobemini **
| |
+--+ +-------------------- Amblema **
| |
| | +----- Toxolasma **
+--+ +-----------+
| | | +-- Obliquaria **
| | +--+
| | +-- Plectomerus **
+--+
| +-------------- Potamilus_leptodon **
| |
| | +----------- Potamilus_alatus **
+--+ |
| | +-------- Venustaconcha **
| | |
+--+ +-------- Leaunio **
| |
| | +-- Ellipsaria **
| +-----+
+--+ +-- Truncilla **
|
| +-- Medionidus **
+-----+
| +-- Ptychobranchus **
|
| +----- Cambarunio **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 25 |
25 |
23 |
4 |
4 |
2 |
1 |
686 |
+-------------- Unioninae
|
--+ +----------- Megalonaias **
| |
+--+ +-- Uniomerus **
| +-----+
| | +-- Quadrulina **
+--+
| +----- Pleurobemini **
+--+
| +-- Lampsilini **
+--+
+-- Amblemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 25 |
25 |
6 |
4 |
0 |
0 |
0 |
695 |
+----------- Quadrulini **
|
+----------- Amblemini **
|
| +-- Elliptio_arctata **
| +-----+
| | +-- Elliptio_crassidens **
| |
| | +----- Pleurobema_decisum **
--+ | |
| | +----- Pleurobema_rubellum **
| | |
| | +----- Pleurobema_georgianum__2 **
| | |
| | +----- Pleurobema_stabile **
| | |
| | +----- Pleurobema_pyriforme **
| | |
+--+ +----- Pleurobema_oviforme__2 **
| |
| +----- Pleurobema_hanleyianum__2 **
| |
| +----- Pleuronaia **
| |
| +----- Pleurobema_georgianum__3 **
| |
| +----- Pleurobema_cordatum **
| |
| +----- Pleurobema_beadleianum **
| |
| +----- Hemistena **
+--+
+----- Fusconaia_flava **
|
+----- Fusconaia_cerina **
|
+----- Eurynia **
|
+----- Elliptio_arca **
|
+----- Lampsilini **
|
| +-- Plethobasus **
+--+
| +-- Parvaspina **
|
| +-- Pleurobema_sintoxia **
| |
+--+-- Pleurobema_oviforme__1 **
| |
| +-- Pleurobema_clava **
|
| +-- Pleurobema_athearni **
| |
+--+-- Pleurobema_georgianum__1 **
|
+-- Pleurobema_hanleyianum__1 **
iT| tT | bi | riS | oT | roT | roS | iD |
| 25 |
127 |
72 |
4 |
175 |
136 |
3 |
642 |
+-------------- Parreysiinae
|
| +----------- Chamberlainia
| |
| +----------- Megalonaias **
| |
| +----------- Uniomerus **
| |
--+ +----------- Quadrulina **
| |
| +----------- Sinohyriopsis
| |
| +----------- Lamprotulini
| |
| +----------- Gonideini
| |
+--+----------- Pseudodontini
|
+----------- Rectidentini
|
+----------- Unioninae
|
| +-------- Amblema **
| |
| +-------- Toxolasma **
| |
| +-------- Pleurobemini **
| |
| | +-- Obliquaria **
| +-----+
| | +-- Plectomerus **
+--+
| +----- Ellipsaria **
| |
| +----- Potamilus_leptodon **
| |
| +----- Medionidus **
| |
| +----- Potamilus_alatus **
| |
| +----- Ptychobranchus **
+--+
+----- Truncilla **
|
+----- Venustaconcha **
|
+----- Cambarunio **
|
+----- Leaunio **
|
| +-- Ortmanniana **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 25 |
25 |
23 |
4 |
173 |
134 |
3 |
683 |
+----------------------------------------------------- Parreysiinae
|
--+ +-------------------------------------------------- Pseudodontini
| |
+--+ +----------------------------------------------- Rectidentini
| |
+--+ +-------------------------------------------- Unioninae
| |
+--+ +----------------------------------------- Chamberlainiini
| |
+--+ +-- Lamprotulini
| +-----------------------------------+
| | +-- Gonideini
+--+
| +----------------------------------- Quadrulini **
| |
+--+ +-------------------------------- Pleurobemini **
| |
+--+ +----------------------------- Amblema **
| |
+--+ +-------------------------- Toxolasma **
| |
+--+ +-- Obliquaria **
| +--------------------+
| | +-- Plectomerus **
+--+
| +-------------------- Potamilus_leptodon **
| |
+--+ +----------------- Potamilus_alatus **
| |
| | +-- Medionidus **
+--+ +-----------+
| | +-- Ptychobranchus **
| |
+--+ +-- Ellipsaria **
| +--------+
| | +-- Truncilla **
+--+
| +-------- Leaunio **
| |
+--+ +-- Venustaconcha **
| +--+
| | +-- Cambarunio **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 24 |
24 |
10 |
3 |
0 |
0 |
0 |
690 |
+-- Amblemini **
|
--+-- Quadrulini **
|
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 24 |
24 |
21 |
4 |
4 |
2 |
1 |
672 |
+----------- Unioninae
--+
| +-------- Amblemini **
+--+
| +----- Lampsilini **
+--+
| +-- Pleurobemini **
+--+
+-- Quadrulini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 23 |
23 |
16 |
4 |
0 |
0 |
0 |
682 |
+----------- Quadrulini **
--+
| +-------- Reginaia **
+--+
| +----- Amblema **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 22 |
22 |
24 |
4 |
0 |
0 |
0 |
690 |
+----------- Quadrulini **
--+
| +-------- Reginaia **
+--+
| +----- Amblema **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 22 |
22 |
20 |
4 |
4 |
2 |
1 |
681 |
+----------- Unioninae
--+
| +-------- Quadrulini **
+--+
| +----- Amblemini **
+--+
| +-- Pleurobemini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 21 |
21 |
20 |
4 |
12 |
10 |
1 |
696 |
+----------------------------- Unioninae
|
| +-- Megalonaias **
| +-----------------------+
--+ | +-- Quadrulina **
| |
| | +----------- Elliptoideus **
| | +-----------+
+--+ | | +-------- Pleurobema **
| | +--+
| | | +----- Uniomerus **
| | +--+
| | | +-- Fusconaia **
+--+ +--+
| +-- Eurynia **
|
| +-------------------- Amblema **
| |
| | +----- Toxolasma **
| | +-----------+
+--+ | | +-- Plectomerus **
| | +--+
| | +-- Obliquaria **
| |
+--+ +----- Ptychobranchus **
| +--------+
| | | +-- Potamilus_fragilis **
| | +--+
| | +-- Medionidus **
+--+
| +-- Villosa **
| +--------+
| | +-- Ellipsaria **
+--+
| +-------- Lampsilis_teres **
+--+
| +----- Lampsilis_siliquoidea **
+--+
| +-- Potamilus_purpuratus **
+--+
+-- Potamilus_alatus **
iT| tT | bi | riS | oT | roT | roS | iD |
| 20 |
20 |
18 |
5 |
1 |
1 |
1 |
683 |
+----------------- Gonideinae
--+
| +-------------- Quadrulini **
+--+
| +----------- Pleurobemini **
+--+
| +-------- Amblema **
+--+
| +----- Plectomerus **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 20 |
20 |
13 |
2 |
0 |
0 |
0 |
678 |
+-- Quadrulini **
--+
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 20 |
20 |
18 |
5 |
1 |
1 |
1 |
683 |
+----------------- Gonideinae
--+
| +-------------- Quadrulini **
+--+
| +----------- Pleurobemini **
+--+
| +-------- Amblema **
+--+
| +----- Plectomerus **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 19 |
19 |
22 |
2 |
1 |
1 |
1 |
677 |
+----- Unioninae
--+
| +-- Quadrulini **
+--+
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 19 |
33 |
32 |
4 |
19 |
14 |
2 |
686 |
+-------------- Quadrulina **
|
| +----- Lamprotulini
--+ +-----+
| | | +-- Chamberlainiini
| | +--+
+--+ +-- Unioninae
|
| +-------- Megalonaias **
+--+
| +----- Pleurobemini **
+--+
| +-- Amblemini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 18 |
18 |
16 |
2 |
0 |
0 |
0 |
684 |
+-- Amblemini **
--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 18 |
18 |
13 |
2 |
0 |
0 |
0 |
683 |
+-------------------- Obliquaria **
|
+-------------------- Amblemini **
|
| +----------------- Lampsilis_teres **
| |
| +----------------- Obovaria **
--+ |
| | +-- Leaunio **
| +--------------+
| | +-- Cambarunio **
| |
| | +-------- Ortmanniana **
| | |
+--+--------+ +-- Lampsilis_hydiana **
| | +--+
| | | +-- Lampsilis_virescens **
| +--+
| | +-- Lampsilis_siliquoidea **
| +--+
| +-- Lampsilis_radiata **
|
| +-------------- Lampsilis_bracteata **
| |
+--+ +----------- Lampsilis_ornata **
| |
+--+ +-------- Lampsilis_ovata **
| |
+--+ +----- Lampsilis_cardium__3 **
| |
+--+ +-- Lampsilis_satura **
| |
+--+-- Lampsilis_cardium__1 **
|
+-- Lampsilis_cardium__2 **
iT| tT | bi | riS | oT | roT | roS | iD |
| 18 |
18 |
16 |
4 |
10 |
6 |
1 |
692 |
+----------------- Unioninae
|
| +-- Megalonaias **
--+ +-----------+
| | +-- Quadrulina **
| |
+--+ +----- Amblema **
| +-----+
| | | +-- Plectomerus **
| | +--+
+--+ +-- Lampsilini **
|
| +-------- Elliptoideus **
+--+
| +----- Uniomerus **
+--+
| +-- Eurynia **
+--+
+-- Pleurobema **
iT| tT | bi | riS | oT | roT | roS | iD |
| 18 |
18 |
16 |
5 |
123 |
96 |
3 |
683 |
+----- Gonideinae
+--------+
| | +-- Parreysiinae
| +--+
--+ +-- Unioninae
|
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Lampsilini **
+--+
| +-- Amblemini **
+--+
+-- Popenaiadini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 18 |
19 |
17 |
4 |
21 |
16 |
1 |
684 |
+----------- Unioninae
|
--+ +-------- Lampsilini **
| |
+--+ +-- Hyriinae
| +--+
| | +-- Amblemini **
+--+
| +-- Pleurobemini **
+--+
+-- Quadrulini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 18 |
18 |
14 |
4 |
9 |
6 |
1 |
690 |
+----------------------------- Unioninae
|
| +-- Megalonaias **
--+ +-----------------------+
| | +-- Quadrulina **
| |
| | +----- Elliptoideus **
+--+ |
| +-----------------+ +-- Eurynia **
| | | |
| | +--+-- Pleurobema **
| | |
+--+ +-- Uniomerus **
|
| +-------------------- Amblema **
| |
+--+ +-- Plectomerus **
| +--------------+
| | +-- Obliquaria **
+--+
| +-------------- Potamilus_fragilis **
| |
+--+ +----------- Ptychobranchus **
| |
+--+ +-------- Medionidus **
| |
+--+ +----- Potamilus_purpuratus **
| |
+--+----- Lampsilis **
|
| +-- Ellipsaria **
+--+
+-- Villosa **
iT| tT | bi | riS | oT | roT | roS | iD |
| 17 |
17 |
15 |
5 |
79 |
68 |
3 |
680 |
+----- Gonideinae
+--------+
| | +-- Parreysiinae
| +--+
--+ +-- Unioninae
|
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 17 |
17 |
14 |
5 |
75 |
69 |
3 |
679 |
+----- Gonideinae
+--------+
| | +-- Parreysiinae
| +--+
--+ +-- Unioninae
|
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 17 |
17 |
11 |
4 |
13 |
8 |
1 |
684 |
+----------- Unionini
|
--+ +-------- Anodontini
| |
+--+ +-- Quadrulini **
| +--+
| | +-- Pleurobemini **
+--+
| +-- Amblemini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 17 |
17 |
13 |
2 |
0 |
0 |
0 |
686 |
+-- Amblemini **
--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 17 |
17 |
15 |
5 |
79 |
68 |
3 |
680 |
+----- Gonideinae
+--------+
| | +-- Parreysiinae
| +--+
--+ +-- Unioninae
|
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 17 |
17 |
15 |
5 |
75 |
69 |
3 |
680 |
+----- Gonideinae
+--------+
| | +-- Parreysiinae
| +--+
--+ +-- Unioninae
|
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 17 |
17 |
15 |
4 |
13 |
8 |
1 |
688 |
+----------- Unionini
|
--+ +-------- Anodontini
| |
+--+ +-- Quadrulini **
| +--+
| | +-- Pleurobemini **
+--+
| +-- Amblemini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 16 |
16 |
12 |
2 |
0 |
0 |
0 |
689 |
+-- Amblemini **
--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 16 |
16 |
14 |
3 |
0 |
0 |
0 |
681 |
+----- Amblemini **
--+
| +-- Pleurobemini **
+--+
+-- Quadrulini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 16 |
16 |
14 |
1 |
0 |
0 |
0 |
683 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 16 |
16 |
14 |
5 |
72 |
63 |
2 |
681 |
+-- Gonideinae
+-----------+
| +-- Unioninae
--+
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 15 |
15 |
13 |
5 |
59 |
50 |
2 |
680 |
+-- Gonideinae
+-----------+
| +-- Unioninae
--+
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 15 |
15 |
13 |
5 |
38 |
36 |
2 |
680 |
+-- Unioninae
+-----------+
| +-- Gonideinae
--+
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 15 |
15 |
13 |
5 |
0 |
0 |
0 |
684 |
+----------- Toxolasma **
+--+
| | +-------- Popenaiadini **
| +--+
| | +----- Amblemini **
| +--+
--+ | +-- Quadrulini **
| +--+
| +-- Pleurobemini **
|
| +----------- Potamilus **
| |
+--+ +-- Lampsilis_cardium **
| +-----+
| | +-- Lampsilis_ornata **
+--+
| +----- Venustaconcha **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Lampsilis_powellii **
iT| tT | bi | riS | oT | roT | roS | iD |
| 15 |
15 |
13 |
5 |
42 |
35 |
3 |
690 |
+----- Gonideinae
+--------+
| | +-- Unioninae
| +--+
--+ +-- Parreysiinae
|
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 15 |
15 |
13 |
5 |
59 |
48 |
2 |
680 |
+-- Gonideinae
+-----------+
| +-- Unioninae
--+
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 15 |
15 |
13 |
5 |
42 |
35 |
3 |
692 |
+----------- Gonideinae
|
| +-- Unioninae
+--------+
--+ +-- Parreysiinae
|
| +-------- Quadrulini **
| |
+--+ +-- Pleurobemini **
| +--+
| | +-- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 15 |
15 |
12 |
4 |
2 |
2 |
2 |
687 |
+----------------- Gonideinae
|
+----------------- Unioninae
|
--+ +-- Pleurobemini **
| +-----------+
| | +-- Quadrulini **
| |
+--+ +-- Paetulunio **
| +--------+
| | +-- Amblemini **
+--+
| +-------- Lampsilis **
| |
+--+ +----- Leaunio **
| |
+--+----- Obovaria **
|
| +-- Villosa **
+--+
+-- Cambarunio **
iT| tT | bi | riS | oT | roT | roS | iD |
| 15 |
15 |
13 |
5 |
59 |
48 |
2 |
680 |
+-- Gonideinae
+-----------+
| +-- Unioninae
--+
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 15 |
15 |
12 |
4 |
0 |
0 |
0 |
684 |
+-------- Quadrula_quadrula **
|
--+-------- Quadrula_verrucosa **
|
| +----- Pleurobemini **
+--+
| +-- Amblemini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 15 |
15 |
13 |
5 |
71 |
62 |
2 |
680 |
+-- Gonideinae
+-----------+
| +-- Unioninae
--+
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 14 |
14 |
12 |
5 |
5 |
3 |
2 |
679 |
+-- Unioninae
+-----------+
| +-- Gonideinae
--+
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 14 |
14 |
12 |
4 |
20 |
18 |
2 |
692 |
+-------------------------------------------- Unioninae
|
--+ +----------------------------------------- Sinosolenaia
| |
+--+ +-------------------------------------- Lamprotulini
| |
+--+ +----------------------------------- Chamberlainiini
| |
+--+ +-------------------------------- Ptychorhynchus
| |
+--+ +----------------------------- Megalonaias **
| |
| | +----- Elliptoideus **
+--+ +--------------------+
| | | +-- Uniomerus **
| | +--+
+--+ +-- Pleurobema **
|
| +----------------------- Amblema **
| |
+--+ +-- Obliquaria **
| +-----------------+
| | +-- Plectomerus **
+--+
| +----------------- Medionidus **
+--+
| +-------------- Potamilus_fragilis **
+--+
| +----------- Ptychobranchus **
+--+
| +-------- Potamilus_purpuratus **
+--+
| +----- Lampsilis **
+--+
| +-- Villosa **
+--+
+-- Ellipsaria **
iT| tT | bi | riS | oT | roT | roS | iD |
| 14 |
14 |
12 |
1 |
0 |
0 |
0 |
691 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 14 |
14 |
6 |
4 |
20 |
18 |
2 |
690 |
+----------------- Unioninae
|
| +-------------- Ptychorhynchus
| |
--+ +-------------- Chamberlainiini
| |
| +-------------- Lamprotulini
| |
+--+-------------- Sinosolenaia
|
| +----------- Megalonaias **
| |
| | +----- Elliptoideus **
| | +--+
+--+ | | +-- Uniomerus **
| | +--+
| | +-- Pleurobema **
| |
| | +----- Potamilus_fragilis **
+--+ |
| +----- Medionidus **
| |
| +----- Amblema **
| |
| | +-- Obliquaria **
+--+--+
| +-- Plectomerus **
|
| +-- Villosa **
| |
| +-- Ellipsaria **
| |
+--+-- Lampsilis **
|
+-- Potamilus_purpuratus **
|
+-- Ptychobranchus **
iT| tT | bi | riS | oT | roT | roS | iD |
| 14 |
14 |
10 |
4 |
33 |
11 |
3 |
685 |
+----------------- Pseudodontini
|
--+ +-------------- Parreysiinae
| |
| +-------------- Rectidentini
+--+
+-------------- Unioninae
|
| +----------- Gonideini
+--+
| +-------- Quadrulini **
+--+
| +----- Amblemini **
+--+
| +-- Pleurobemini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 14 |
14 |
12 |
4 |
33 |
11 |
3 |
691 |
+----------------------------- Unioninae
|
--+ +-------------------------- Rectidentini
| |
| | +-- Gonideini
+--+ +--------------------+
| | +-- Pseudodontini
| |
| | +-------- Amblemini **
+--+ +-----------+
| | | +----- Pleurobemini **
| | +--+
| | | +-- Obliquaria **
+--+ +--+
| +-- Quadrulini **
|
| +----------------- Ptychobranchus **
+--+
| +-------------- Truncilla **
+--+
| +----------- Ortmanniana **
+--+
| +-------- Lampsilis **
+--+
| +----- Cambarunio **
+--+
| +-- Ligumia **
+--+
+-- Epioblasma **
iT| tT | bi | riS | oT | roT | roS | iD |
| 13 |
13 |
10 |
3 |
33 |
11 |
3 |
691 |
+-------------------------------- Pseudodontini
|
--+ +----------------------------- Rectidentini
| |
+--+ +-------------------------- Unioninae
| |
+--+ +----------------------- Gonideini
| |
| | +----- Pleurobemini **
+--+ +--------------+
| | | +-- Obliquaria **
| | +--+
+--+ +-- Quadrulini **
|
| +----------------- Ptychobranchus **
+--+
| +-------------- Truncilla **
+--+
| +----------- Ortmanniana **
+--+
| +-------- Lampsilis **
+--+
| +----- Cambarunio **
+--+
| +-- Ligumia **
+--+
+-- Epioblasma **
iT| tT | bi | riS | oT | roT | roS | iD |
| 13 |
13 |
7 |
4 |
33 |
11 |
3 |
686 |
+----------------------- Pseudodontini
|
--+ +-------------------- Rectidentini
| |
+--+ +----------------- Unioninae
| |
+--+ +-------------- Gonideini
| |
| | +-- Obliquaria **
+--+ +--------+
| | +-- Quadrulini **
| |
+--+ +-------- Amblemini **
| |
| +-------- Pleurobemini **
+--+
| +----- Truncilla **
| |
| +----- Ptychobranchus **
+--+
| +-- Ortmanniana **
| |
| +-- Cambarunio **
+--+
+-- Ligumia **
|
+-- Epioblasma **
iT| tT | bi | riS | oT | roT | roS | iD |
| 13 |
14 |
12 |
4 |
12 |
12 |
3 |
697 |
+-------------------------------- Parreysiinae
|
--+ +----------------------------- Unioninae
| |
+--+ +-------------------------- Lamprotulini
| |
+--+ +----------------------- Quadrulini **
| |
+--+ +-------------------- Gonideini
| |
| | +----- Eurynia **
+--+ +-----------+
| | | +-- Cambarunio **
| | +--+
| | +-- Ptychobranchus **
+--+
| +-- Amblemini **
| +-----------+
| | +-- Pleurobema **
+--+
| +----------- Truncilla **
| |
+--+ +-- Lampsilis **
| +-----+
| | +-- Epioblasma **
+--+
| +----- Ligumia **
+--+
| +-- Obliquaria **
+--+
+-- Ortmanniana **
iT| tT | bi | riS | oT | roT | roS | iD |
| 13 |
13 |
12 |
4 |
0 |
0 |
0 |
689 |
+-------- Quadrulini **
--+
| +----- Pleurobemini **
+--+
| +-- Amblemini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 12 |
12 |
10 |
5 |
0 |
0 |
0 |
689 |
+-- Popenaiadini **
+-----+
| +-- Lampsilini **
--+
| +----- Amblemini **
+--+
| +-- Pleurobemini **
+--+
+-- Quadrulini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 12 |
12 |
11 |
4 |
2 |
1 |
1 |
682 |
+-------- Lampsilini **
--+
| +----- Amblemini **
+--+
| +-- Quadrulini **
+--+
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 12 |
13 |
11 |
4 |
2 |
1 |
1 |
689 |
+-- Quadrulini **
+-----+
| +-- Unioninae
--+
| +----- Lampsilini **
+--+
| +-- Amblemini **
+--+
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 12 |
12 |
10 |
5 |
57 |
46 |
2 |
681 |
+-- Gonideinae
+-----------+
| +-- Unioninae
--+
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 12 |
12 |
10 |
5 |
41 |
39 |
2 |
681 |
+-- Gonideinae
+-----------+
| +-- Unioninae
--+
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 12 |
12 |
10 |
1 |
0 |
0 |
0 |
679 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 12 |
14 |
9 |
4 |
2 |
1 |
1 |
683 |
+----- Unioninae
|
+----- Margaritiferinae
|
--+----- Quadrulini **
|
+----- Lampsilini **
|
| +-- Amblemini **
+--+
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 12 |
12 |
8 |
5 |
0 |
0 |
0 |
687 |
+----- Amblemini **
+--+
| | +-- Pleurobemini **
| +--+
| +-- Quadrulini **
--+
| +----- Obliquaria **
| |
| +----- Toxolasma **
+--+
+----- Popenaiadini **
|
| +-- Glebula **
+--+
+-- Cyrtonaias **
iT| tT | bi | riS | oT | roT | roS | iD |
| 12 |
14 |
11 |
4 |
2 |
1 |
1 |
680 |
+----- Unioninae
|
+----- Margaritiferinae
|
--+----- Lampsilini **
|
+----- Amblemini **
|
| +-- Quadrulini **
+--+
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 12 |
17 |
0 |
4 |
10 |
6 |
2 |
687 |
+----- Anodontina
|
| +-- Gonideinae
| |
| +-- Pyganodon_grandis
| |
| +-- Utterbackia
| |
| +-- Pyganodon_cataracta
--+ |
| +-- Lampsilis_straminea **
| |
| +-- Lampsilis_teres **
| |
| +-- Megalonaias **
| |
| +-- Obliquaria **
| |
+--+-- Theliderma **
|
+-- Quadrula **
|
+-- Uniomerus **
|
+-- Amblema **
|
+-- Elliptio **
|
+-- Fusconaia **
|
+-- Plectomerus **
|
+-- Pleurobema **
|
+-- Unionini
iT| tT | bi | riS | oT | roT | roS | iD |
| 12 |
12 |
11 |
4 |
2 |
1 |
1 |
681 |
+-------- Amblemini **
--+
| +----- Lampsilini **
+--+
| +-- Quadrulini **
+--+
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 11 |
16 |
12 |
4 |
20 |
7 |
3 |
693 |
+----------------------------- Gonideinae
|
--+ +-------------------------- Parreysiinae
| |
+--+ +----------------------- Fusconaia **
| |
| +----------------------- Amblemini **
+--+
| +-------------------- Quadrulini **
| |
+--+ +----------------- Pleurobema **
| |
+--+ +-------------- Eurynia **
| |
| | +-- Cyrtonaias **
+--+-----------+
| +-- Unioninae
|
| +----------- Ortmanniana **
+--+
| +-------- Glebula **
+--+
| +----- Potamilus **
+--+
| +-- Ligumia **
+--+
+-- Toxolasma **
iT| tT | bi | riS | oT | roT | roS | iD |
| 11 |
17 |
0 |
4 |
11 |
7 |
2 |
687 |
+----- Anodontina
|
| +-- Gonideinae
| |
| +-- Pyganodon_grandis
| |
| +-- Utterbackia
| |
| +-- Pyganodon_cataracta
--+ |
| +-- Lampsilis_straminea **
| |
| +-- Lampsilis_teres **
| |
| +-- Megalonaias **
| |
| +-- Obliquaria **
| |
+--+-- Theliderma **
|
+-- Quadrula **
|
+-- Uniomerus **
|
+-- Amblema **
|
+-- Elliptio **
|
+-- Fusconaia **
|
+-- Plectomerus **
|
+-- Aculamprotulina
|
+-- Unionina
iT| tT | bi | riS | oT | roT | roS | iD |
| 11 |
11 |
9 |
4 |
20 |
7 |
3 |
680 |
+-------------- Gonideinae
--+
| +----------- Unioninae
+--+
| +-------- Amblemini **
+--+
| +----- Lampsilini **
+--+
| +-- Quadrulini **
+--+
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 11 |
28 |
16 |
4 |
20 |
7 |
3 |
701 |
+-------------------- Amblemini **
|
+-------------------- Parreysiinae
|
+-------------------- Eurynia **
|
+-------------------- Fusconaia **
|
+-------------------- Gonideinae
|
+-------------------- Pleurobema **
|
+-------------------- Quadrulini **
--+
| +----------- Ortmanniana **
+--------+
| | +-------- Glebula **
| +--+
| | +----- Potamilus **
| +--+
| | +-- Ligumia **
| +--+
| +-- Toxolasma **
|
| +----------------- Cyrtonaias **
| |
+--+ +-------------- Unionini
| |
| | +----------- Anodontini
+--+ |
| | +----- Iridininae
| | |
+--+ +--+ +-- Anodontites_guanarensis
| | | |
| | +--+-- Anodontites_trigona
| | |
+--+ +-- Monocondylaeinae
|
| +----- Rhipidodontini
| |
| +----- Echyridella
+--+
+----- Lortiella
|
| +-- Castaliini
+--+
+-- Velesunio
iT| tT | bi | riS | oT | roT | roS | iD |
| 11 |
11 |
6 |
4 |
0 |
0 |
0 |
686 |
+-------- Quadrula **
|
--+-------- Pustulosa **
|
| +----- Pleurobemini **
+--+
| +-- Amblemini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 11 |
11 |
9 |
3 |
43 |
40 |
2 |
684 |
+----------- Unioninae
--+
| +-------- Gonideinae
+--+
| +----- Quadrulini **
+--+
| +-- Pleurobemini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 11 |
11 |
5 |
4 |
23 |
7 |
3 |
682 |
+-------- Toxolasma **
|
+-------- Amblemini **
|
| +-- Glebula **
+-----+
--+ +-- Cyrtonaias **
|
| +-- Quadrulini **
+-----+
| +-- Pleurobemini **
|
| +----- Potamilus **
+--+
| +-- Ortmanniana **
+--+
+-- Ligumia **
iT| tT | bi | riS | oT | roT | roS | iD |
| 11 |
11 |
9 |
5 |
0 |
0 |
0 |
688 |
+-- Popenaiadini **
+-----+
| +-- Lampsilini **
--+
| +----- Amblemini **
+--+
| +-- Pleurobemini **
+--+
+-- Quadrulini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 11 |
11 |
9 |
2 |
0 |
0 |
0 |
688 |
+-- Lampsilini **
--+
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 11 |
11 |
9 |
4 |
20 |
7 |
3 |
682 |
+----------------- Parreysiinae
--+
| +-------------- Unioninae
+--+
| +----------- Gonideinae
+--+
| +-------- Amblemini **
+--+
| +----- Lampsilini **
+--+
| +-- Quadrulini **
+--+
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 11 |
11 |
8 |
4 |
20 |
7 |
3 |
683 |
+-- Gonideinae
+-----------+
| +-- Unioninae
|
--+ +-- Glebula **
| +--------+
| | +-- Cyrtonaias **
| |
+--+ +-------- Toxolasma **
| |
| | +----- Potamilus **
| +--+
+--+ | +-- Ligumia **
| +--+
| +-- Ortmanniana **
|
| +----- Amblemini **
+--+
| +-- Quadrulini **
+--+
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 11 |
11 |
9 |
1 |
0 |
0 |
0 |
689 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 11 |
11 |
9 |
1 |
0 |
0 |
0 |
690 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 11 |
15 |
2 |
4 |
8 |
5 |
2 |
689 |
+-------- Gonideinae
|
| +-- Elliptio **
| |
| +-- Fusconaia **
| |
--+ +--+-- Amblemini **
| | |
| | +-- Pleurobema **
| | |
| | +-- Unionini
| |
| | +-- Pyganodon_cataracta
+--+ |
| +-- Pyganodon_grandis
| |
| +-- Utterbackia
| |
| +-- Lampsilis_teres **
| |
| +-- Lampsilis_straminea **
+--+
+-- Megalonaias **
|
+-- Obliquaria **
|
+-- Theliderma **
|
+-- Quadrula **
|
+-- Uniomerus **
iT| tT | bi | riS | oT | roT | roS | iD |
| 10 |
10 |
8 |
3 |
46 |
36 |
2 |
683 |
+----------- Unioninae
--+
| +-------- Gonideinae
+--+
| +----- Quadrulini **
+--+
| +-- Pleurobemini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 10 |
10 |
8 |
4 |
43 |
41 |
3 |
683 |
+----- Gonideinae
+-----+
| | +-- Parreysiinae
| +--+
--+ +-- Unioninae
|
| +-------- Quadrulini **
+--+
| +----- Pleurobemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 10 |
10 |
9 |
2 |
0 |
0 |
0 |
693 |
+-- Lampsilini **
--+
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 10 |
10 |
5 |
2 |
0 |
0 |
0 |
684 |
+-------------- Pleurobemini **
|
+-------------- Obliquaria **
--+
| +----------- Potamilus_capax **
| |
| +----------- Lampsilis **
+--+
| +-------- Potamilus_fragilis **
| |
| | +----- Potamilus_inflatus **
+--+ |
| | +-- Potamilus_ohiensis **
| +--+
+--+ +-- Potamilus_amphichaenus **
|
| +-- Potamilus_purpuratus__1 **
| |
+--+-- Potamilus_purpuratus__2 **
|
+-- Potamilus_alatus **
iT| tT | bi | riS | oT | roT | roS | iD |
| 10 |
10 |
8 |
1 |
0 |
0 |
0 |
685 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 10 |
10 |
7 |
1 |
0 |
0 |
0 |
688 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 10 |
10 |
8 |
3 |
0 |
0 |
0 |
683 |
+----- Lampsilini **
--+
| +-- Amblemini **
+--+
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 10 |
10 |
8 |
1 |
0 |
0 |
0 |
687 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 9 |
9 |
7 |
4 |
129 |
104 |
3 |
682 |
+-------------------- Parreysiinae
|
--+ +----------------- Unioninae
| |
| | +-------------- Rectidentini
+--+ |
| | +-------- Pseudodontini
| | +--+
+--+ | | +----- Gonideini
| | +--+
| | | +-- Chamberlainiini
| | +--+
+--+ +-- Lamprotulini
|
| +-- Amblemini **
| +--+
| | +-- Lampsilini **
+-----+
| +-- Quadrulini **
+--+
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 9 |
9 |
7 |
4 |
11 |
7 |
3 |
682 |
+----- Gonideinae
+-----+
| | +-- Parreysiinae
| +--+
--+ +-- Unioninae
|
| +-------- Quadrulini **
+--+
| +----- Pleurobemini **
+--+
| +-- Amblemini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 9 |
9 |
7 |
2 |
7 |
0 |
0 |
686 |
+-- Popenaias **
+--------+
| +-- Lampsilini **
--+
| +-------- Nephronaias_hjalmarsoni **
+--+
| +----- Psoronaias **
+--+
| +-- Nephronaias_goascoranensis **
+--+
+-- Nephronaias_tempisquensis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 9 |
9 |
7 |
4 |
71 |
60 |
3 |
680 |
+-------------------------- Parreysiinae
|
--+ +----------------------- Unioninae
| |
| | +-------------------- Rectidentini
+--+ |
| | +-- Amblemini **
| | +--+
+--+ | +-- Lampsilini **
| +-----------+
| | | +-- Quadrulini **
| | +--+
| | +-- Pleurobemini **
+--+
| +-- Chamberlainiini
| +-----------+
| | +-- Lamprotulini
| |
+--+ +-- Parvasolenaia
| +--------+
| | +-- Pseudodontini
+--+
| +-------- Sinosolenaia
+--+
| +----- Gonidea
+--+
| +-- Microcondylaea
+--+
+-- Leguminaia
iT| tT | bi | riS | oT | roT | roS | iD |
| 9 |
9 |
7 |
3 |
2 |
1 |
1 |
688 |
+----------------- Glebula **
|
--+ +-- Amblema **
| +-----------+
| | +-- Quadrulini **
+--+
| +----------- Plectomerus **
+--+
| +-------- Leaunio **
+--+
| +----- Lampsilis_straminea **
+--+
| +-- Lampsilis_teres **
+--+
+-- Potamilus **
iT| tT | bi | riS | oT | roT | roS | iD |
| 9 |
9 |
7 |
2 |
22 |
22 |
2 |
684 |
+----------- Unioninae
|
--+ +-------- Gonideinae
| |
+--+ +-- Epioblasma **
| +--+
| | +-- Lampsilis **
+--+
| +-- Cyrtonaias **
+--+
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 9 |
9 |
7 |
4 |
67 |
57 |
2 |
684 |
+-------------------- Pseudodontini
|
--+ +----------------- Rectidentini
| |
+--+ +-------------- Gonideini
| |
+--+ +-- Chamberlainiini
| +--------+
| | +-- Lamprotulini
+--+
| +-------- Unioninae
| |
+--+ +-- Amblemini **
| +--+
| | +-- Lampsilini **
+--+
| +-- Quadrulini **
+--+
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 9 |
9 |
7 |
3 |
0 |
0 |
0 |
686 |
+-- Amblemini **
|
--+-- Quadrulini **
|
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 9 |
51 |
43 |
4 |
44 |
42 |
2 |
676 |
+-------- Quadrulini **
|
+-------- Pleurobemini **
|
| +-- Amblemini **
--+-----+
| +-- Lampsilini **
|
| +-- Lamprotulini
| +--+
| | +-- Gonideini
+--+
| +-- Chamberlainiini
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 9 |
9 |
0 |
4 |
115 |
115 |
3 |
687 |
+----------------- Parreysiinae
|
| +-------------- Pseudodontini
--+ |
| | +----------- Rectidentini
| | |
+--+ | +-------- Unioninae
| | |
| | | +-- Amblemini **
| | | |
+--+ | +-- Ortmanniana **
| | |
| | +-- Lampsilis **
| | |
| | +-- Cambarunio **
| | |
+--+ +--+-- Elliptio **
| | |
| | +-- Eurynia **
| | |
| | +-- Pleurobema **
| | |
| | +-- Quadrula_quadrula **
| | |
| | +-- Quadrula_verrucosa **
| |
| | +-- Chamberlainia
+--+ |
| +-- Sinohyriopsis
| |
| +-- Lamprotula_caveata
| |
| +-- Lamprotula_leaii
| |
| +-- Potomida
| |
| +-- Pronodularia
| |
+--+-- Gonidea
|
+-- Leguminaia_wheatleyi
|
+-- Leguminaia
|
+-- Microcondylaea
|
+-- Sinosolenaia_carinata
|
+-- Sinosolenaia_oleivora
|
+-- Sinosolenaia_recognita
iT| tT | bi | riS | oT | roT | roS | iD |
| 9 |
9 |
7 |
2 |
22 |
22 |
2 |
680 |
+-------------- Unioninae
--+
| +----------- Chamberlainiini
+--+
| +-------- Lamprotulini
+--+
| +----- Gonideini
+--+
| +-- Lampsilini **
+--+
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 9 |
9 |
7 |
4 |
129 |
104 |
3 |
684 |
+-------------------- Parreysiinae
|
--+ +----------------- Rectidentini
| |
+--+ +-------------- Pseudodontini
| |
+--+ +----------- Unioninae
| |
| | +----- Gonideini
+--+ +--+
| | | +-- Chamberlainiini
| | +--+
| | +-- Lamprotulini
+--+
| +-- Amblemini **
| +--+
| | +-- Lampsilini **
+--+
| +-- Quadrulini **
+--+
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 9 |
9 |
7 |
4 |
67 |
57 |
2 |
682 |
+-------------------- Pseudodontini
|
--+ +----------------- Rectidentini
| |
+--+ +-------------- Gonideini
| |
+--+ +-- Chamberlainiini
| +--------+
| | +-- Lamprotulini
+--+
| +-------- Unioninae
| |
+--+ +-- Amblemini **
| +--+
| | +-- Lampsilini **
+--+
| +-- Quadrulini **
+--+
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 9 |
9 |
7 |
3 |
3 |
2 |
1 |
686 |
+----------- Unioninae
--+
| +-------- Quadrulini **
+--+
| +----- Plectomerus **
+--+
| +-- Amblema **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 8 |
8 |
9 |
2 |
2 |
1 |
1 |
685 |
+----- Unioninae
--+
| +-- Pleurobemini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
13 |
0 |
4 |
75 |
75 |
3 |
687 |
+-------------- Parreysiinae
|
--+ +----------- Pseudodontini
| |
| | +-------- Rectidentini
+--+ |
| | +----- Unioninae
| | |
+--+ | +-- Lamprotula
| | |
| | +-- Potomida
| | |
| | +-- Pronodularia
+--+ |
| +-- Gonidea
| |
| +-- Leguminaia
| |
| +-- Microcondylaea
| |
+--+-- Amblemini **
|
+-- Ortmanniana **
|
+-- Lampsilis **
|
+-- Cambarunio **
|
+-- Elliptio **
|
+-- Pleurobema **
|
+-- Quadrulini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
7 |
7 |
1 |
0 |
0 |
0 |
685 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
7 |
5 |
2 |
32 |
30 |
3 |
684 |
+-- Parreysiinae
+-----+
| +-- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
7 |
3 |
1 |
0 |
0 |
0 |
687 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
7 |
5 |
1 |
0 |
0 |
0 |
688 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
7 |
10 |
1 |
0 |
0 |
0 |
684 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
7 |
0 |
4 |
29 |
25 |
2 |
687 |
+-- Amblemini **
|
+-- Ortmanniana **
|
+-- Lampsilis **
|
--+-- Cambarunio **
|
+-- Elliptio **
|
+-- Pleurobema **
|
+-- Quadrulini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
7 |
8 |
1 |
0 |
0 |
0 |
683 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
7 |
5 |
2 |
30 |
28 |
3 |
684 |
+-- Quadrulini **
+-----+
| +-- Lampsilini **
--+
| +----- Gonideinae
+--+
| +-- Parreysiinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
7 |
5 |
1 |
0 |
0 |
0 |
686 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
7 |
1 |
1 |
0 |
0 |
0 |
688 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
7 |
5 |
1 |
11 |
11 |
2 |
693 |
+-------------- Sinanodonta
|
--+ +----------- Cristaria
| |
+--+ +-- Gonideinae
| +-----+
| | +-- Quadrulini **
+--+
| +----- Aculamprotulina
+--+
| +-- Acuticostini
+--+
+-- Oxynaiina
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
7 |
8 |
1 |
0 |
0 |
0 |
683 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
16 |
14 |
2 |
27 |
26 |
2 |
691 |
+----- Prolasmidonta
+--+
| | +-- Alasmidonta
| +--+
+-----------------+ +-- Anemina
| |
| | +----- Pseudodontini
| +--+
| | +-- Lamprotula_caveata
| +--+
| +-- Chamberlainia
|
| +----- Oxynaiina
| +-----+
| | | +-- Lepidodesmini
--+ | +--+
| | +-- Aculamprotulina
| +-----------+
| | | +----- Platynaias
| | | +--+
| | | | | +-- Utterbackia_imbecillis
| | | | +--+
| | +--+ +-- Pyganodon
| | |
| | | +----- Sinanodonta
| | +--+
| | | +-- Cristaria
+--+ +--+
| +-- Anodontina
|
| +-- Utterbackia_peninsularis
| +-----+
| | +-- Potamilus_alatus **
| +-----------+
| | | +----- Lamprotula_leaii
| | +--+
| | | +-- Sinohyriopsis
+--+ +--+
| +-- Gonideini
|
| +----------------- Gibbosulinae
+--+
| +-------------- Quadrulini **
+--+
| +----------- Toxolasma **
+--+
| +-------- Lampsilis_ornata **
+--+
| +----- Potamilus_leptodon **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Lampsilis_powellii **
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
7 |
9 |
1 |
0 |
0 |
0 |
683 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
7 |
5 |
2 |
40 |
34 |
2 |
682 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
2 |
0 |
0 |
0 |
687.5 |
+-- Elliptio **
|
--+-- Quadrulini **
|
+-- Pleurobema **
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
6 |
1 |
0 |
0 |
0 |
687 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
8 |
7 |
3 |
10 |
7 |
2 |
686 |
+-------- Unioninae
|
--+ +-- Pleurobemini **
| +--+
| | +-- Gonideinae
+--+
| +-- Lampsilini **
+--+
+-- Quadrulini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
2 |
38 |
35 |
2 |
683 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
5 |
2 |
0 |
0 |
0 |
685 |
+-- Lampsilini **
--+
+-- Popenaiadini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
2 |
25 |
25 |
2 |
683 |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
2 |
8 |
4 |
2 |
685 |
+-------- Gonideinae
--+
| +----- Unioninae
+--+
| +-- Pleurobemini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
2 |
25 |
25 |
2 |
683 |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
2 |
10 |
8 |
2 |
683 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
2 |
9 |
6 |
2 |
686 |
+-------------- Pseudodontini
--+
| +----------- Gonidea
+--+
| +-------- Inversidens
+--+
| +----- Unioninae
+--+
| +-- Pleurobemini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
2 |
38 |
35 |
2 |
683 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
3 |
13 |
11 |
1 |
685 |
+-------- Unioninae
--+
| +----- Lampsilini **
+--+
| +-- Pleurobemini **
+--+
+-- Quadrulini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
3 |
2 |
30 |
4 |
2 |
686 |
+-------------- Gonideinae
|
--+ +----------- Cyrtonaias **
| |
+--+----------- Pleurobemini **
|
| +-------- Potamilus **
+--+
| +----- Lampsilis **
+--+
| +-- Ligumia **
+--+
+-- Ortmanniana **
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
7 |
5 |
2 |
9 |
6 |
2 |
689 |
+-------------- Unioninae
|
--+ +----------- Lamprotulini
| |
+--+ +-- Gonideini
| +-----+
| | +-- Pseudodontini
+--+
| +----- Lampsilini **
+--+
| +-- Pleurobemini **
+--+
+-- Margaritiferinae
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
2 |
3 |
2 |
1 |
687 |
+----- Unioninae
--+
| +-- Pleurobemini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
2 |
3 |
2 |
1 |
685 |
+----- Unioninae
--+
| +-- Pleurobemini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
2 |
42 |
39 |
2 |
683 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
2 |
40 |
34 |
2 |
683 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
2 |
36 |
26 |
2 |
683 |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
2 |
0 |
0 |
0 |
683 |
+-- Quadrulini **
--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
2 |
30 |
29 |
2 |
683 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
2 |
30 |
29 |
2 |
683 |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
3 |
85 |
68 |
3 |
685 |
+-------------------- Parreysiinae
|
--+ +----------------- Unioninae
| |
+--+ +-------------- Rectidentini
| |
| | +----- Quadrulini **
+--+ +-----+
| | | +-- Pleurobemini **
| | +--+
+--+ +-- Lampsilini **
|
| +-------- Chamberlainiini
+--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
2 |
43 |
35 |
2 |
683 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
2 |
9 |
6 |
2 |
688 |
+-------------- Pseudodontini
--+
| +----------- Gonidea
+--+
| +-------- Inversidens
+--+
| +----- Unioninae
+--+
| +-- Pleurobemini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
3 |
85 |
68 |
3 |
683 |
+-------------------- Parreysiinae
|
--+ +----------------- Unioninae
| |
+--+ +-------------- Rectidentini
| |
| | +----- Lampsilini **
+--+ +-----+
| | | +-- Quadrulini **
| | +--+
+--+ +-- Pleurobemini **
|
| +-------- Chamberlainiini
+--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
686 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
4 |
31 |
19 |
3 |
686 |
+----------------------- Parreysiinae
|
--+ +-- Pseudodontini
| +-----------------+
| | +-- Rectidentini
+--+
| +----------------- Unioninae
+--+
| +-------------- Gonideini
+--+
| +----------- Lamprotulini
+--+
| +-------- Pleurobemini **
+--+
| +----- Quadrulini **
+--+
| +-- Amblemini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
4 |
1 |
0 |
0 |
0 |
687 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
688 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
2 |
4 |
31 |
19 |
3 |
685 |
+----------------------- Parreysiinae
|
--+ +-------------------- Rectidentini
| |
+--+ +----------------- Pseudodontini
| |
+--+ +-------------- Unioninae
| |
+--+ +----------- Gonideini
| |
+--+ +-------- Lamprotulini
| |
+--+ +----- Quadrulini **
| |
+--+----- Pleurobemini **
|
| +-- Amblemini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
687 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
1 |
1 |
0 |
0 |
0 |
687 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
3 |
35 |
25 |
3 |
684 |
+-------------- Parreysiinae
--+
| +----------- Unioninae
+--+
| +-------- Gonideinae
+--+
| +----- Quadrulini **
+--+
| +-- Amblemini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
2 |
3 |
13 |
12 |
2 |
685 |
+-- Quadrulini **
|
--+-- Pleurobemini **
|
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
4 |
26 |
24 |
3 |
686 |
+----------------- Parreysiinae
--+
| +-------------- Unioninae
+--+
| +----------- Gonideinae
+--+
| +-------- Quadrulini **
+--+
| +----- Pleurobemini **
+--+
| +-- Amblemini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
3 |
8 |
8 |
1 |
686 |
+-------- Unioninae
--+
| +----- Lampsilini **
+--+
| +-- Amblemini **
+--+
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
2 |
33 |
30 |
2 |
684 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
2 |
2 |
0 |
0 |
0 |
685 |
+-- Lampsilini **
--+
+-- Amblemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
0 |
4 |
118 |
107 |
3 |
687 |
+----------------- Parreysiinae
|
--+ +-------------- Unioninae
| |
| | +----------- Rectidentini
+--+ |
| | +-- Amblemini **
| | |
+--+ +-- Pleurobemini **
| +-----+
| | +-- Lampsilini **
| | |
| | +-- Quadrulini **
+--+
| +-- Chamberlainiini
| +--+
| | +-- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
2 |
3 |
14 |
8 |
2 |
687 |
+----------------- Pseudodontini
--+
| +-------------- Rectidentini
+--+
| +----------- Unioninae
+--+
| +-------- Gonideini
+--+
| +----- Lampsilini **
+--+
| +-- Pleurobemini **
+--+
+-- Amblemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
687 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
2 |
2 |
37 |
17 |
2 |
685 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
2 |
2 |
38 |
38 |
2 |
685 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
685 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
2 |
2 |
27 |
25 |
2 |
685 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
2 |
2 |
36 |
33 |
2 |
685 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
2 |
2 |
16 |
15 |
2 |
685 |
+----- Gonideinae
--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
2 |
1 |
1 |
1 |
1 |
685 |
+-- Unioninae
--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
2 |
2 |
37 |
17 |
2 |
685 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
2 |
2 |
28 |
25 |
2 |
685 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
2 |
2 |
12 |
11 |
2 |
685 |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
2 |
2 |
1 |
1 |
1 |
687 |
+----- Unioninae
--+
| +-- Quadrulini **
+--+
+-- Amblemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
19 |
16 |
4 |
16 |
15 |
2 |
680 |
+-------- Amblemini **
|
+-------- Pleurobemini **
--+
+-------- Lampsilini **
|
| +----- Quadrulini **
+--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
2 |
3 |
6 |
6 |
2 |
685 |
+-- Gonideinae
+-----+
| +-- Unioninae
--+
| +----- Lampsilini **
+--+
| +-- Pleurobemini **
+--+
+-- Quadrulini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
687 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
2 |
2 |
21 |
15 |
2 |
685 |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
2 |
2 |
21 |
15 |
2 |
685 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
1 |
13 |
10 |
1 |
688 |
+-- Lampsilini **
--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
2 |
4 |
4 |
2 |
686 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Unioninae
+--+
+-- Gonideinae
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
2 |
13 |
13 |
2 |
686 |
+----------- Anodontini
|
--+ +-- Lanceolariini
| +-----+
| | +-- Unionini
+--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
2 |
19 |
14 |
2 |
686 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
2 |
17 |
15 |
2 |
686 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
6 |
4 |
2 |
5 |
5 |
2 |
687 |
+-------------- Cristariina
--+
| +----------- Alasmidontina
+--+
| +-------- Lampsilis **
+--+
| +----- Gonideinae
+--+
| +-- Venustaconcha **
+--+
+-- Quadrulini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
1 |
7 |
5 |
2 |
686 |
+----- Gonideinae
--+
| +-- Lampsilini **
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
5 |
3 |
3 |
2 |
2 |
1 |
688 |
+-------- Quadrulini **
--+
| +----- Unioninae
+--+
| +-- Lampsilini **
+--+
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
2 |
20 |
17 |
2 |
686 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
2 |
8 |
6 |
2 |
686 |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
2 |
19 |
19 |
2 |
686 |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
2 |
16 |
15 |
2 |
686 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Unioninae
+--+
+-- Gonideinae
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
2 |
20 |
19 |
2 |
686 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Unioninae
+--+
+-- Gonideinae
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
2 |
16 |
15 |
2 |
686 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Unioninae
+--+
+-- Gonideinae
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
2 |
14 |
12 |
2 |
686 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
2 |
11 |
9 |
2 |
686 |
+-- Anodontina
+--------+
| +-- Utterbackia
|
--+ +----- Unionini
| +--+
| | | +-- Pyganodon
| | +--+
+--+ +-- Cristariina
|
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
2 |
1 |
3 |
3 |
1 |
688 |
+-- Unioninae
--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
2 |
27 |
26 |
2 |
686 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
4 |
2 |
2 |
1 |
1 |
1 |
687 |
+----- Lampsilini **
--+
| +-- Unioninae
+--+
+-- Quadrulini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
1 |
0 |
0 |
0 |
686 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
2 |
8 |
6 |
2 |
686 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
2 |
18 |
18 |
2 |
686 |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
4 |
2 |
2 |
1 |
1 |
1 |
687 |
+----- Quadrulini **
--+
| +-- Unioninae
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
5 |
3 |
2 |
9 |
6 |
2 |
686 |
+-------- Unioninae
--+
| +----- Lampsilini **
+--+
| +-- Quadrulini **
+--+
+-- Gonideinae
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
2 |
9 |
6 |
2 |
686 |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
2 |
3 |
3 |
1 |
686 |
+----- Unioninae
--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
2 |
19 |
14 |
2 |
686 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
2 |
1 |
0 |
0 |
686 |
+-- Lampsilini **
--+
+-- Quadrulini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
7 |
0 |
2 |
14 |
8 |
3 |
690 |
+-------------------------- Lamellidens_mainwaringi
--+
| +----------------------- Lamprotula
+--+
| +-------------------- Potomida
+--+
| +----------------- Lampsilini **
+--+
| +-------------- Pleurobemini **
+--+
| +----------- Aculamprotulina
+--+
| +-------- Unio_pictorum
+--+
| +----- Anodontini
+--+
| +-- Unio_ravoisieri
+--+
+-- Lamellidens_corrianus
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
11 |
11 |
2 |
-- |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
6 |
0 |
2 |
5 |
4 |
2 |
687 |
+-------- Quadrulini **
--+
| +----- Gonideinae
+--+
| +-- Lampsilini **
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
43 |
41 |
3 |
-- |
+-- Parreysiinae
+-----+
| +-- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Lampsilini **
+--+
+-- Quadrulini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
8 |
7 |
2 |
-- |
+-- Lampsilini **
+--+
| +-- Quadrulini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
39 |
37 |
3 |
-- |
+-- Parreysiinae
+-----+
| +-- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Lampsilini **
+--+
+-- Quadrulini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
39 |
37 |
3 |
-- |
+-- Parreysiinae
+-----+
| +-- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Lampsilini **
+--+
+-- Quadrulini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
4 |
4 |
2 |
-- |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
6 |
0 |
2 |
13 |
7 |
3 |
686 |
+-- Pleurobemini **
|
--+-- Lampsilini **
|
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
39 |
37 |
3 |
-- |
+-- Parreysiinae
+-----+
| +-- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
6 |
4 |
2 |
-- |
+----- Gonideinae
|
--+----- Unioninae
|
| +-- Lampsilini **
+--+
+-- Quadrulini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
13 |
11 |
2 |
-- |
+-- Amblemini **
+--+
| +-- Quadrulini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
11 |
11 |
2 |
-- |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
1 |
1 |
1 |
-- |
+----- Unioninae
--+
| +-- Pleurobemini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
29 |
27 |
1 |
-- |
+----- Gonideinae
--+
| +-- Amblemini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
39 |
37 |
3 |
-- |
+-- Parreysiinae
+-----+
| +-- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Lampsilini **
+--+
+-- Quadrulini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
51 |
51 |
1 |
-- |
+----- Unioninae
--+
| +-- Lampsilini **
+--+
+-- Quadrulini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
10 |
10 |
2 |
-- |
+-- Gonideini
+--------+
| +-- Lamprotulini
--+
| +-------- Chamberlainiini
+--+
| +----- Unioninae
+--+
| +-- Lampsilini **
+--+
+-- Quadrulini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
3 |
0 |
2 |
4 |
1 |
1 |
687 |
+-- Pleurobemini **
|
--+-- Lampsilini **
|
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
43 |
41 |
3 |
-- |
+-- Amblemini **
+--------------+
| +-- Lampsilini **
|
| +-------- Parreysiinae
--+ +-----+
| | | +----- Unioninae
| | +--+
| | | +-- Chamberlainiini
+--+ +--+
| +-- Discomya
|
| +----------- Rectidentini
| |
+--+ +-- Schepmaniini
| +-----+
| | +-- Pseudodontini
+--+
| +----- Caudiculatus
+--+
| +-- Potomida
+--+
+-- Gonideini
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
9 |
0 |
2 |
13 |
7 |
3 |
687 |
+-- Aculamprotulina
|
+-- Lamprotula
|
+-- Lampsilini **
|
+-- Potomida
|
--+-- Unio_pictorum
|
+-- Anodontini
|
+-- Unio_ravoisieri
|
+-- Parreysiinae
|
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
9 |
2 |
1 |
-- |
+----- Unioninae
--+
| +-- Amblemini **
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
54 |
53 |
3 |
-- |
+----------- Parreysiinae
--+
| +-------- Unioninae
+--+
| +----- Gonideinae
+--+
| +-- Lampsilini **
+--+
+-- Amblemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
5 |
5 |
1 |
-- |
+----- Unioninae
--+
| +-- Lampsilini **
+--+
+-- Quadrulini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
1 |
13 |
13 |
2 |
-- |
+----- Unioninae
--+
| +-- Gonideinae
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
4 |
0 |
2 |
4 |
4 |
2 |
687 |
+----- Gonideinae
|
--+ +-- Cristariina
| |
| +-- Lampsilini **
+--+
+-- Alasmidontina
|
+-- Quadrulini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
6 |
0 |
2 |
5 |
4 |
2 |
687 |
+-------- Quadrulini **
--+
| +----- Lampsilini **
+--+
| +-- Unioninae
+--+
+-- Gonideinae
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
7 |
7 |
2 |
-- |
+-- Lampsilini **
+--+
| +-- Quadrulini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
3 |
0 |
2 |
15 |
4 |
2 |
689.5 |
+-------- Unionini
--+
| +----- Pleurobemini **
+--+
| +-- Anodontini
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
12 |
11 |
3 |
-- |
+-------- Parreysiinae
|
--+ +-- Amblemini **
| +--+
| | +-- Lampsilini **
+--+
| +-- Unioninae
+--+
+-- Gonideinae
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
5 |
5 |
1 |
-- |
+----- Unioninae
--+
| +-- Lampsilini **
+--+
+-- Quadrulini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
74 |
72 |
1 |
-- |
+----- Unioninae
--+
| +-- Lampsilini **
+--+
+-- Quadrulini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
1 |
7 |
6 |
1 |
-- |
+-- Pleurobemini **
--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
2 |
2 |
1 |
-- |
+----- Unioninae
--+
| +-- Lampsilini **
+--+
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
6 |
6 |
2 |
-- |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Lampsilini **
+--+
+-- Quadrulini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
43 |
41 |
3 |
-- |
+----------------------- Gonideinae
|
| +-- Lampsilini **
--+ +-----------------+
| | +-- Quadrulini **
| |
| | +----------------- Parreysiinae
+--+ |
| | +-- Anemina
| +--------------+
| | +-- Amuranodonta
+--+
| +----- Pletholophus
| +--------+
| | | +-- Sinanodonta
| | +--+
| | +-- Beringiana
+--+
| +----------- Cristaria
| |
| | +-- Alasmidontina
+--+--------+
| +-- Anodontina
|
| +-------- Lanceolariini
+--+
| +----- Lepidodesmini
+--+
| +-- Acuticostini
+--+
+-- Unionini
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
12 |
10 |
2 |
-- |
+-- Amblemini **
+--+
| +-- Quadrulini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
6 |
4 |
2 |
-- |
+----- Gonideinae
|
--+----- Unioninae
|
| +-- Lampsilini **
+--+
+-- Quadrulini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
0 |
0 |
0 |
-- |
+-- Quadrulini **
--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
7 |
0 |
0 |
-- |
-- Epioblasma_triquetra **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
19 |
18 |
1 |
-- |
+-- Amblemini **
--+
+-- Gonideinae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
3 |
2 |
1 |
-- |
+-- None__1
+--+
| +-- None__2
--+
| +-- Pleurobemini **
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
4 |
4 |
1 |
-- |
+-- Lampsilini **
--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
11 |
2 |
1 |
-- |
+-- Lampsilini **
--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
4 |
1 |
1 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
13 |
10 |
2 |
-- |
+----- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
16 |
14 |
2 |
-- |
+----- Amblemini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
3 |
3 |
2 |
-- |
+-- Lampsilini **
|
--+-- Unioninae
|
+-- Gonideinae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
16 |
15 |
2 |
-- |
+----- Amblemini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
33 |
10 |
3 |
-- |
+-- Lampsilini **
+--+
| +-- Parreysiinae
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
67 |
44 |
3 |
-- |
+-------------- Parreysiinae
--+
| +----------- Unioninae
+--+
| +-------- Chamberlainiini
+--+
| +----- Lampsilini **
+--+
| +-- Lamprotulini
+--+
+-- Gonideini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
3 |
3 |
1 |
-- |
+-- Pleurobemini **
--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
4 |
1 |
1 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
13 |
2 |
1 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
19 |
18 |
1 |
-- |
+-- Amblemini **
--+
+-- Gonideinae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
14 |
2 |
2 |
-- |
+----- Unioninae
--+
| +-- Lampsilini **
+--+
+-- Parreysiinae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
28 |
5 |
3 |
-- |
+-------- Gonideinae
--+
| +----- Unioninae
+--+
| +-- Lampsilini **
+--+
+-- Parreysiinae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
7 |
7 |
1 |
-- |
+-- Amblemini **
--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
9 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
10 |
2 |
1 |
-- |
+-- Lampsilini **
--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
11 |
2 |
1 |
-- |
+-- Unionini
|
--+-- Anodontini
|
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
8 |
0 |
0 |
-- |
-- Epioblasma_triquetra **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
25 |
1 |
1 |
-- |
+-- Lampsilini **
--+
+-- Gonideinae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
67 |
44 |
3 |
-- |
+-------- Parreysiinae
--+
| +----- Unioninae
+--+
| +-- Lampsilini **
+--+
+-- Gonideinae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
29 |
3 |
1 |
-- |
+-- Lampsilini **
--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
12 |
12 |
1 |
-- |
+-- Quadrulini **
--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
9 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
14 |
2 |
2 |
-- |
+----- Unioninae
--+
| +-- Parreysiinae
+--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
33 |
10 |
3 |
-- |
+-- Lampsilini **
+--+
| +-- Parreysiinae
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
12 |
1 |
1 |
-- |
+-- Unioninae
--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
10 |
9 |
2 |
-- |
+----- Gonideinae
--+
| +-- Amblemini **
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
2 |
1 |
1 |
-- |
+-- Unioninae
--+
+-- Pleurobemini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
67 |
44 |
3 |
-- |
+-------- Parreysiinae
--+
| +----- Unioninae
+--+
| +-- Lampsilini **
+--+
+-- Gonideinae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
29 |
3 |
1 |
-- |
+-- Lampsilini **
--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
10 |
9 |
2 |
-- |
+----- Amblemini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
1 |
1 |
1 |
-- |
+-- Gonideinae
--+
+-- Lampsilini **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
27 |
2 |
2 |
-- |
+----- Gonideinae
--+
| +-- Lampsilini **
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
9 |
8 |
2 |
-- |
+----- Gonideinae
--+
| +-- Amblemini **
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
3 |
3 |
2 |
-- |
+-- Unioninae
|
--+-- Lampsilini **
|
+-- Gonideinae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
10 |
10 |
2 |
-- |
+----- Amblemini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
27 |
2 |
2 |
-- |
+----- Gonideinae
--+
| +-- Lampsilini **
+--+
+-- Unioninae
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
3 |
3 |
1 |
-- |
+-- Pleurobemini **
--+
+-- Unioninae
