Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini

subtribe ANODONTINA Rafinesque, 1820: cladomics

3 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 11 (24 total)
number of subtaxa (rank: genus) = 3
analyzed in 232 cladograms in 125 publications
maximum number of terminals in an analysis = 10 (mean = 2.03, median = 1)
maximum number of subtaxa (genus) = 3 (mean = 1.43, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 114
maximum number of included terminals = 10, (mean = 3.11 median = 3)
maximum number of subtaxa (genus) = 3 (mean = 1.87, median = 2)
proportion of cladograms that support basic monophyly = 0.6

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 61
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 10 (mean = 3.67, median = 4)
maximum number of subtaxa (genus) = 3 (mean = 2.36, median = 2)
proportion of cladograms that support rigorous monophyly = 0.38

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Anodontina cladogram distances
PCoA Ordination

NJ Tree of Anodontina cladogram distances
NJ Tree

Cladograms

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 72 10 2 **
Cladomics cladogram CRISTARIINA ALASMIDONTINA Pseudanodonta Brachyanodon Anodonta

Chong et al., 2008 [Publ 2668], fig. 3

cst iT tT bi riS oT roT roS iD
0.706 7 18 10 3 11 10 2 19.0
Cladomics cladogram Anodonta cygnea Pyganodon Alasminota Anodonta anatina UNIONINA CRISTARIINA Brachyanodon nuttalliana Brachyanodon californiensis Pseudodontoideus Alasmidonta Pterosyna Pseudanodonta Strophitus Prolasmidonta Platynaias Brachyanodon oregonensis 4 Brachyanodon oregonensis 3 Brachyanodon oregonensis 2 Brachyanodon oregonensis 1 Brachyanodon kennerlyi

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
0.938 4 4 1 3 77 10 2 1.0
Cladomics cladogram CRISTARIINA ALASMIDONTINA Pseudanodonta Anodonta anatina Anodonta cygnea Brachyanodon

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
0.867 4 9 7 3 135 11 2 9.0
Cladomics cladogram CRISTARIINA Anodonta anatina ALASMIDONTINA Pseudanodonta Anodonta cygnea Brachyanodon

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
0.800 4 9 3 3 135 11 2 8.0
Cladomics cladogram Lanceolaria CRISTARIINA Anodonta anatina Pseudanodonta ALASMIDONTINA Anodonta cygnea Brachyanodon

Bolotov et al., 2017 [Publ 3764], fig. S1

cst iT tT bi riS oT roT roS iD
0.533 4 18 0 3 121 11 2 16.0
Cladomics cladogram Unio crassus Unio pictorum Unio tumidus ACULAMPROTULINA Cuneopsis szechenyii Tchangsinaia Cuneopsis rufescens Nodularia douglasiae Nodularia nuxpersicae Schistodesmus Alasmidonta Anodonta anatina Anodonta cygnea Brachyanodon Lasmigona Pseudanodonta Pyganodon Simpsonaias Strophitus Sinanodonta lauta Cristaria plicata Cristaria radiata Pletholophus Sinanodonta lucida Sinanodonta jourdyi Lanceolaria gladiola Lanceolaria grayii Lanceolaria danzhouensis

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
0.824 4 11 4 3 197 16 2 9.0
Cladomics cladogram CRISTARIINA Alasmidonta Lasmigona Pseudanodonta Simpsonaias Strophitus Pyganodon Utterbackia Anodonta anatina Anodonta cygnea Brachyanodon

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
0.824 4 11 8 3 195 16 2 9.0
Cladomics cladogram CRISTARIINA Alasmidonta Lasmigona Anodonta anatina Anodonta cygnea Brachyanodon Pyganodon Utterbackia Pseudanodonta Simpsonaias Strophitus

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.706 4 48 25 3 197 16 2 46.0
Cladomics cladogram Chamberlainia Megalonaias Uniomerus QUADRULINA Sinohyriopsis INDOPSEUDODONTINA PSEUDODONTINA Rectidentina CONTRADENTINA Lamprotula Potomida Pronodularia Sinosolenaia carinata Gonidea Leguminaia Microcondylaea Ptychorhynchus Parvasolenaia Koreosolenaia Sinosolenaia recognita Sinosolenaia iridinea Sinosolenaia oleivora Amblema Toxolasma Obliquaria Plectomerus Elliptoideus Elliptio Eurynia Fusconaia Pleurobema Ellipsaria Potamilus leptodon Medionidus Potamilus alatus Ptychobranchus Truncilla Venustaconcha Cambarunio Leaunio Ortmanniana Lampsilis Acuticosta Alasmidonta Anodonta anatina Anemina Anodonta cygnea Cristaria Brachyanodon Lasmigona Lepidodesma Pseudanodonta Pyganodon Simpsonaias Strophitus Utterbackia Schistodesmus UNIONINA ACULAMPROTULINA Lanceolaria Pletholophus Sinanodonta Cuneopsis celtiformis Tchangsinaia Cuneopsis rufescens Nodularia Cuneopsis szechenyii Cuneopsis heudei

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
0.895 4 9 4 3 118 29 2 7.0
Cladomics cladogram CRISTARIINA Pseudanodonta ALASMIDONTINA Anodonta cygnea Anodonta anatina Brachyanodon

Riccardi et al., 2020 [Publ 4089], fig. 2

cst iT tT bi riS oT roT roS iD
0.846 5 9 7 3 14 8 2 9.0
Cladomics cladogram CRISTARIINA Anodonta anatina Brachyanodon ALASMIDONTINA Pseudanodonta Anodonta exulcerata Anodonta cygnea

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

cst iT tT bi riS oT roT roS iD
0.933 4 4 2 3 72 10 2 2.0
Cladomics cladogram CRISTARIINA ALASMIDONTINA Pseudanodonta Anodonta anatina Brachyanodon Anodonta cygnea

Whelan et al., 2023 [Publ 4287], fig. 2

cst iT tT bi riS oT roT roS iD
0.913 4 37 38 3 37 37 2 40.0
Cladomics cladogram CRISTARIINA Anodonta cygnea Brachyanodon Pseudanodonta Anodonta anatina ALASMIDONTINA

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 3 90 13 2 3.0
Cladomics cladogram CRISTARIINA ALASMIDONTINA Anodonta Brachyanodon Pseudanodonta

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cst iT tT bi riS oT roT roS iD
0.882 5 12 8 3 90 13 2 9.0
Cladomics cladogram CRISTARIINA Brachyanodon Pseudanodonta Anodonta cygnea ALASMIDONTINA Anodonta exulcerata Anodonta anatina

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cst iT tT bi riS oT roT roS iD
0.941 5 12 10 3 86 13 2 10.0
Cladomics cladogram CRISTARIINA Pseudanodonta Anodonta Brachyanodon ALASMIDONTINA

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cst iT tT bi riS oT roT roS iD
0.941 5 12 10 3 86 13 2 10.0
Cladomics cladogram CRISTARIINA Brachyanodon ALASMIDONTINA Pseudanodonta Anodonta

Wu et al., 2024 [Publ 4379], fig. 5

cst iT tT bi riS oT roT roS iD
0.941 5 12 10 3 48 13 2 10.0
Cladomics cladogram CRISTARIINA Brachyanodon ALASMIDONTINA Pseudanodonta Anodonta

Hou et al., 2024 [Publ 4400], fig. 4

cst iT tT bi riS oT roT roS iD
0.941 5 12 10 3 70 13 2 10.0
Cladomics cladogram CRISTARIINA Pseudanodonta Anodonta Brachyanodon ALASMIDONTINA

Hou et al., 2025 [Publ 4445], fig. 5

cst iT tT bi riS oT roT roS iD
0.941 5 12 10 3 47 13 2 10.0
Cladomics cladogram CRISTARIINA Brachyanodon ALASMIDONTINA Pseudanodonta Anodonta

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cst iT tT bi riS oT roT roS iD
0.875 5 13 11 3 135 12 2 13.0
Cladomics cladogram CRISTARIINA Anodonta cygnea Anodonta exulcerata Anodonta anatina ALASMIDONTINA Brachyanodon Pseudanodonta

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.833 10 10 9 3 623 70 2 12.0
Cladomics cladogram Pseudodontoideus Lasmigona Pterosyna Arcidens Utterbackia Pyganodon Utterbackiana Simpsonaias Alasminota Alasmidonta Prolasmidonta Platynaias Anodontoides Strophitus Anodonta anatina Pseudanodonta Brachyanodon oregonensis 2 Anodonta cygnea Anodonta seddonae Anodonta exulcerata Brachyanodon impura Brachyanodon Brachyanodon nuttalliana 1 Brachyanodon nuttalliana 2 Brachyanodon californiensis Brachyanodon oregonensis 1

Plazzi et al., 2011 [Publ 3676], fig. 5

cst iT tT bi riS oT roT roS iD
0.750 3 17 10 2 15 2 2 15.0
Cladomics cladogram CRISTARIINA Lanceolaria Anodonta UNIONINA ALASMIDONTINA Pseudanodonta Alathyria Pronodularia Sinohyriopsis QUADRULINA Parvaspina Lampsilis Epioblasma Venustaconcha

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
0.833 3 5 3 2 88 7 2 3.0
Cladomics cladogram CRISTARIINA Lasmigona Anodonta Pseudanodonta Pyganodon Strophitus

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
0.750 3 6 4 2 88 7 2 6.0
Cladomics cladogram CRISTARIINA Anodonta cygnea Lasmigona Anodonta anatina Pseudanodonta Pyganodon Strophitus

Chen et al., 2023 [Publ 4302], fig. 1

cst iT tT bi riS oT roT roS iD
0.857 2 2 0 2 39 8 2 --
Cladomics cladogram ALASMIDONTINA Anodonta Pseudanodonta

Chen et al., 2023 [Publ 4302], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 39 8 2 --
Cladomics cladogram CRISTARIINA ALASMIDONTINA Anodonta Pseudanodonta

Wu et al., 2023 [Publ 4339], fig. 5

cst iT tT bi riS oT roT roS iD
0.900 3 3 1 2 16 6 2 3.0
Cladomics cladogram CRISTARIINA ALASMIDONTINA Anodonta anatina Pseudanodonta Anodonta cygnea

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

cst iT tT bi riS oT roT roS iD
0.667 7 9 7 2 21 6 2 9.0
Cladomics cladogram CRISTARIINA Anodonta exulcerata Anodonta cygnea Brachyanodon californiensis Brachyanodon nuttalliana Brachyanodon kennerlyi Brachyanodon oregonensis Anodonta anatina ALASMIDONTINA

Dai et al., 2025 [Publ 4440], fig. 1

cst iT tT bi riS oT roT roS iD
0.857 2 2 0 2 42 8 2 --
Cladomics cladogram ALASMIDONTINA Anodonta Pseudanodonta

Dai et al., 2025 [Publ 4440], fig. 2

cst iT tT bi riS oT roT roS iD
0.857 2 2 0 2 42 8 2 --
Cladomics cladogram ALASMIDONTINA Anodonta Pseudanodonta

Chen et al., 2025 [Publ 4487], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 39 8 2 --
Cladomics cladogram CRISTARIINA ALASMIDONTINA Anodonta Pseudanodonta

Chen et al., 2025 [Publ 4487], fig. 3

cst iT tT bi riS oT roT roS iD
0.857 2 2 0 2 39 8 2 --
Cladomics cladogram ALASMIDONTINA Anodonta Pseudanodonta

Wang et al., 2025 [Publ 4488], fig. 3B

cst iT tT bi riS oT roT roS iD
0.950 4 11 9 2 60 22 2 9.0
Cladomics cladogram CRISTARIINA Anodonta Pseudanodonta ALASMIDONTINA

Wu et al., 2026 [Publ 4811], fig. 4

cst iT tT bi riS oT roT roS iD
0.950 4 11 9 2 91 22 2 9.0
Cladomics cladogram CRISTARIINA Anodonta Pseudanodonta ALASMIDONTINA

Wu et al., 2026 [Publ 4825], fig. 6B

cst iT tT bi riS oT roT roS iD
0.950 4 11 9 2 61 21 2 9.0
Cladomics cladogram CRISTARIINA Anodonta Pseudanodonta ALASMIDONTINA

Hoeh, 1990 [Publ 2717], fig. 2

cst iT tT bi riS oT roT roS iD
0.818 2 5 0 2 13 13 1 7.0
Cladomics cladogram Pterosyna Platynaias Lasmigona Utterbackia Pyganodon Anodonta Brachyanodon Utterbackiana

Hoeh, 1990 [Publ 2717], fig. 3

cst iT tT bi riS oT roT roS iD
0.545 2 15 0 2 13 13 1 17.0
Cladomics cladogram Pyganodon fragilis Utterbackia imbecillis Utterbackia peggyae Anodonta Brachyanodon Utterbackiana couperiana 1 Utterbackiana couperiana 2 Utterbackiana suborbiculata Lasmigona Utterbackiana implicata Pterosyna Platynaias Pyganodon cataracta Pyganodon gibbosa Pyganodon grandis Pyganodon lacustris

Hoeh, 1990 [Publ 2717], fig. 4

cst iT tT bi riS oT roT roS iD
0.818 2 5 0 2 13 13 1 7.0
Cladomics cladogram Pterosyna Platynaias Lasmigona Utterbackia Pyganodon Anodonta Brachyanodon Utterbackiana

Nagel et al., 1996 [Publ 3400], fig. 351

cst iT tT bi riS oT roT roS iD
0.667 4 5 4 2 1 1 1 4.0
Cladomics cladogram Pseudanodonta Anodonta cygnea CRISTARIINA Anodonta 2 Anodonta anatina Anodonta 1

Nagel et al., 1998 [Publ 1946], fig. 7

cst iT tT bi riS oT roT roS iD
0.667 4 5 4 2 1 1 1 4.0
Cladomics cladogram Pseudanodonta Anodonta cygnea CRISTARIINA Anodonta 2 Anodonta anatina Anodonta 1

Campbell et al., 2005 [Publ 2660], fig. 1

cst iT tT bi riS oT roT roS iD
0.778 2 6 0 2 100 4 1 7.0
Cladomics cladogram Sinohyriopsis Brachyanodon Anodonta Alasminota Pyganodon Lasmigona Pseudodontoideus Gonidea Potomida Pronodularia QUADRULINA Megalonaias Uniomerus Elliptoideus Hemistena Plethobasus Elliptio Pleuronaia Fusconaia Pleurobema Eurynia Parvaspina Plectomerus Reginaia Popenaias Amblema Toxolasma Obliquaria Cyrtonaias Glebula Potamilus Medionidus acutissimus Lemiox Dromus Cyprogenia Sagittunio Medionidus conradicus Ptychobranchus Venustaconcha ellipsiformis 2 Ellipsaria Truncilla Villosa Obovaria olivaria Venustaconcha ellipsiformis 1 Epioblasma Obovaria unicolor Obovaria arkansasensis Obovaria subrotunda Hamiota Lampsilis cardium Lampsilis ovata Ortmanniana pectorosa Lampsilis ornata Ligumia Cambarunio Lampsilis teres Ortmanniana ligamentina Lampsilis siliquoidea Leaunio

Campbell et al., 2005 [Publ 2660], fig. 2

cst iT tT bi riS oT roT roS iD
0.889 2 2 0 2 100 4 1 --
Cladomics cladogram Pyganodon Lasmigona Pseudodontoideus Alasminota Anodonta Brachyanodon

Zanatta et al., 2007 [Publ 2698], fig. 1A

cst iT tT bi riS oT roT roS iD
0.818 2 13 0 2 14 14 1 14.0
Cladomics cladogram Platynaias Lasmigona Pterosyna Anodonta Brachyanodon Utterbackiana Utterbackia Pyganodon

Zanatta et al., 2007 [Publ 2698], fig. 1B

cst iT tT bi riS oT roT roS iD
0.818 2 6 0 2 14 14 1 7.0
Cladomics cladogram Lasmigona Platynaias Pterosyna Utterbackia Pyganodon Anodonta Brachyanodon Utterbackiana

Manendo et al., 2008 [Publ 2669], fig. 1

cst iT tT bi riS oT roT roS iD
0.857 3 5 3 2 10 2 1 3.0
Cladomics cladogram UNIONINA Anodonta Pseudanodonta ALASMIDONTINA

Soroka, 2010 [Publ 3404], fig. 2

cst iT tT bi riS oT roT roS iD
0.667 3 3 1 2 4 1 1 3.0
Cladomics cladogram UNIONINA Anodonta cygnea Anodonta anatina Pseudanodonta

Soroka, 2010 [Publ 3404], fig. 3a

cst iT tT bi riS oT roT roS iD
0.667 3 3 1 2 6 2 1 3.0
Cladomics cladogram UNIONINA Anodonta anatina Anodonta cygnea Pseudanodonta

Soroka, 2010 [Publ 3404], fig. 3b

cst iT tT bi riS oT roT roS iD
0.500 3 9 5 2 6 2 1 8.0
Cladomics cladogram Anodonta anatina Anodonta cygnea Pseudanodonta CRISTARIINA UNIONINA

Soroka, 2010 [Publ 3404], fig. 4

cst iT tT bi riS oT roT roS iD
0.500 3 7 5 2 4 1 1 7.0
Cladomics cladogram Anodonta anatina UNIONINA CRISTARIINA Anodonta cygnea Pseudanodonta

Soroka, 2010 [Publ 3404], fig. 5a

cst iT tT bi riS oT roT roS iD
0.667 3 5 3 2 5 2 1 5.0
Cladomics cladogram UNIONINA Anodonta anatina CRISTARIINA Anodonta cygnea Pseudanodonta

Soroka, 2010 [Publ 3404], fig. 5b

cst iT tT bi riS oT roT roS iD
0.500 3 8 5 2 5 2 1 8.0
Cladomics cladogram Anodonta anatina CRISTARIINA UNIONINA Anodonta cygnea Pseudanodonta

Zieritz et al., 2012 [Publ 3673], fig. 4

cst iT tT bi riS oT roT roS iD
0.667 3 3 0 2 5 1 1 2.0
Cladomics cladogram Pseudanodonta Anodonta anatina Anodonta cygnea CRISTARIINA Margaritifera UNIONINA

Mezhzherin et al., 2014 [Publ 3705], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 1 1 1 1.0
Cladomics cladogram CRISTARIINA Pseudanodonta Anodonta

Mezhzherin et al., 2014 [Publ 3705], fig. 4

cst iT tT bi riS oT roT roS iD
0.833 3 3 1 2 1 1 1 3.0
Cladomics cladogram CRISTARIINA Anodonta anatina Anodonta cygnea Pseudanodonta

Bolotov et al., 2020 [Publ 4084], fig. 1

cst iT tT bi riS oT roT roS iD
0.909 3 3 1 2 33 9 1 3.0
Cladomics cladogram CRISTARIINA Anodonta anatina Pseudanodonta Anodonta cygnea

Riccardi et al., 2020 [Publ 4089], fig. 1

cst iT tT bi riS oT roT roS iD
0.857 4 4 2 2 2 2 1 4.0
Cladomics cladogram CRISTARIINA Anodonta anatina Pseudanodonta Anodonta exulcerata Anodonta cygnea

Tomilova et al., 2020 [Publ 4097], fig. 3

cst iT tT bi riS oT roT roS iD
0.667 4 4 2 2 4 2 1 4.0
Cladomics cladogram UNIONINA CRISTARIINA Anodonta anatina Pseudanodonta Anodonta cygnea Anodonta exulcerata

Konopleva et al., 2023 [Publ 4269], fig. S1_4

cst iT tT bi riS oT roT roS iD
0.909 3 3 1 2 53 13 1 3.0
Cladomics cladogram CRISTARIINA Anodonta anatina Pseudanodonta Anodonta cygnea

Perkins et al., 2025 [Publ 4447], fig. 2_mtDNA-ITS

cst iT tT bi riS oT roT roS iD
0.950 3 36 37 2 33 33 1 37.0
Cladomics cladogram Anodonta Pseudanodonta ALASMIDONTINA

Altaba et al., 2025 [Publ 4849], fig. 11

cst iT tT bi riS oT roT roS iD
0.778 6 6 4 2 8 4 1 7.0
Cladomics cladogram CRISTARIINA Brachyanodon kennerlyi Brachyanodon oregonensis Anodonta cygnea Anodonta exulcerata Anodonta anatina Brachyanodon californiensis

Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5_7

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 3 0 0 --
Cladomics cladogram UNIONINA Pseudanodonta Anodonta

Nagel et al., 1998 [Publ 1946], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 6 0 0 1.0
Cladomics cladogram UNIONINA Pseudanodonta Anodonta

Nagel et al., 1998 [Publ 1946], fig. 9

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 4 0 0 --
Cladomics cladogram UNIONINA Anodonta Pseudanodonta

Källersjö et al., 2005 [Publ 2684], fig. 1A

cst iT tT bi riS oT roT roS iD
0.750 3 3 0 2 4 0 0 2.0
Cladomics cladogram Margaritifera UNIONINA Anodonta anatina Pseudanodonta Anodonta cygnea

Källersjö et al., 2005 [Publ 2684], fig. 1B-F

cst iT tT bi riS oT roT roS iD
0.750 3 3 1 2 4 0 0 3.0
Cladomics cladogram Margaritifera UNIONINA Anodonta anatina Pseudanodonta Anodonta cygnea

Hinzmann et al., 2013 [Publ 3541], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 0 0 0 1.0
Cladomics cladogram Pseudanodonta Anodonta

Reis et al., 2013 [Publ 3580], fig. 4

cst iT tT bi riS oT roT roS iD
0.750 3 3 1 2 11 0 0 3.0
Cladomics cladogram UNIONINA Anodonta cygnea Anodonta anatina Pseudanodonta

Reis et al., 2013 [Publ 3580], fig. 5

cst iT tT bi riS oT roT roS iD
0.750 3 3 0 2 12 0 0 2.0
Cladomics cladogram UNIONINA Anodonta anatina Anodonta cygnea Pseudanodonta

Prié & Puillandre, 2014 [Publ 3529], fig. 2

cst iT tT bi riS oT roT roS iD
0.750 3 3 1 2 14 0 0 3.0
Cladomics cladogram UNIONINA Anodonta anatina Pseudanodonta Anodonta cygnea

Froufe et al., 2014 [Publ 3592], fig. 2

cst iT tT bi riS oT roT roS iD
0.750 3 3 1 2 0 0 0 3.0
Cladomics cladogram Anodonta anatina Pseudanodonta Anodonta cygnea

Lopes-Lima et al., 2016 [Publ 3757], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 0 0 0 1.0
Cladomics cladogram Pseudanodonta Anodonta

Lopes-Lima et al., 2017 [Publ 3734], fig. 1

cst iT tT bi riS oT roT roS iD
0.750 3 3 1 2 14 0 0 3.0
Cladomics cladogram UNIONINA Anodonta anatina Pseudanodonta Anodonta cygnea

Bogatov et al., 2018 [Publ 3859], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 4 0 0 --
Cladomics cladogram UNIONINA Anodonta Pseudanodonta

Bogatov et al., 2018 [Publ 3859], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 4 0 0 --
Cladomics cladogram UNIONINA Anodonta Pseudanodonta

Lyubas et al., 2022 [Publ 4255], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 2 26 0 0 3.0
Cladomics cladogram UNIONINA Pseudanodonta Anodonta

Lyubas et al., 2022 [Publ 4255], fig. S1

cst iT tT bi riS oT roT roS iD
0.750 5 5 3 2 26 0 0 5.0
Cladomics cladogram UNIONINA Anodonta anatina Pseudanodonta Anodonta exulcerata Anodonta seddonae Anodonta cygnea

Soroka, 2008 [Publ 3695], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 2 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Soroka, 2008 [Publ 3695], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 2 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Wu et al., 2016 [Publ 3711], fig. 1

cst iT tT bi riS oT roT roS iD
0.900 1 1 0 1 20 6 2 --
Cladomics cladogram Sinanodonta Anemina Cristaria Anodonta ALASMIDONTINA

Fonseca et al., 2016 [Publ 3775], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 8 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Feng et al., 2016 [Publ 3800], fig. 2

cst iT tT bi riS oT roT roS iD
0.778 1 1 0 1 13 5 2 --
Cladomics cladogram Anodonta ALASMIDONTINA

Guerra et al., 2017 [Publ 3772], fig. 2a

cst iT tT bi riS oT roT roS iD
0.900 1 1 0 1 24 7 2 --
Cladomics cladogram Sinanodonta Cristaria Anodonta ALASMIDONTINA

Guerra et al., 2017 [Publ 3772], fig. 2b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 24 7 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 31 8 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 21 8 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Bolotov et al., 2018 [Publ 3826], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 81 9 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Huang et al., 2018 [Publ 3828], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 41 8 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Huang et al., 2018 [Publ 3828], fig. 6-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 16 3 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Wu et al., 2018 [Publ 3836], fig. 2

cst iT tT bi riS oT roT roS iD
0.833 2 7 0 1 51 11 2 7.0
Cladomics cladogram CRISTARIINA Anodonta cygnea Anodonta anatina ALASMIDONTINA

Wang et al., 2018 [Publ 3846], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 6 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Chase et al., 2018 [Publ 3847], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 37 12 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

cst iT tT bi riS oT roT roS iD
0.500 3 20 0 1 102 17 2 23.0
Cladomics cladogram Alasmidonta marginata Prolasmidonta Alasmidonta undulata CRISTARIINA Brachyanodon kennerlyi Brachyanodon nuttalliana Brachyanodon oregonensis Arcidens confragosus Arcidens wheeleri Pterosyna Platynaias compressa Lasmigona Platynaias decorata Platynaias subviridis Pyganodon grandis Pyganodon lacustris Pseudodontoideus Strophitus Utterbackia imbecillis Utterbackia peninsularis 1 Utterbackia peninsularis 2 Gonidea Reginaia Amblema Popenaias Dromus Epioblasma Glebula Lampsilis ovata Lampsilis virescens Lemiox Obliquaria Obovaria olivaria Obovaria unicolor Plectomerus Potamilus alatus Potamilus capax Uniomerus QUADRULINA Elliptoideus Parvaspina Plethobasus Eurynia Hemistena Elliptio Pleuronaia Pleurobema cordatum Fusconaia Pleurobema plenum Pleurobema stabile Pleurobema pyriforme Pleurobema georgianum 2 Pleurobema athearni Pleurobema decisum Pleurobema georgianum 1 Pleurobema hanleyianum 1 Pleurobema hanleyianum 2 Pleurobema beadleianum 2 Pleurobema strodeanum 2 Pleurobema rubellum Pleurobema beadleianum 1 Pleurobema taitianum Pleurobema perovatum Pleurobema strodeanum 1 Pleurobema oviforme 6 Pleurobema clava 3 Pleurobema oviforme 5 Pleurobema clava 2 Pleurobema oviforme 2 Pleurobema oviforme 3 Pleurobema oviforme 4 Pleurobema clava 1 Pleurobema oviforme 1 Pleurobema riddellii Pleurobema sintoxia 34 Pleurobema sintoxia 31 Pleurobema sintoxia 32 Pleurobema sintoxia 33 Pleurobema 4 Pleurobema 5 Pleurobema sintoxia 2 Pleurobema sintoxia 3 Pleurobema 8 Pleurobema 9 Pleurobema 10 Pleurobema sintoxia 5 Pleurobema sintoxia 6 Pleurobema sintoxia 7 Pleurobema sintoxia 8 Pleurobema sintoxia 9 Pleurobema sintoxia 10 Pleurobema sintoxia 11 Pleurobema sintoxia 12 Pleurobema sintoxia 13 Pleurobema sintoxia 14 Pleurobema sintoxia 15 Pleurobema sintoxia 16 Pleurobema sintoxia 17 Pleurobema sintoxia 18 Pleurobema sintoxia 19 Pleurobema sintoxia 20 Pleurobema sintoxia 21 Pleurobema sintoxia 22 Pleurobema sintoxia 23 Pleurobema sintoxia 24 Pleurobema sintoxia 25 Pleurobema sintoxia 26 Pleurobema sintoxia 27 Pleurobema sintoxia 28 Pleurobema sintoxia 29 Pleurobema sintoxia 30 Pleurobema 6 Pleurobema sintoxia 1 Pleurobema 7 Pleurobema sintoxia 4 Pleurobema 3 Pleurobema 1 Pleurobema 2

Burzynski & Soroka, 2018 [Publ 3862], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 3 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 3 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 3 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Aunins et al., 2018 [Publ 3863], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 38 9 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Wang et al., 2018 [Publ 3865], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 6 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Cha et al., 2018 [Publ 3916], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 37 8 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Wu et al., 2018 [Publ 4143], fig. 2

cst iT tT bi riS oT roT roS iD
0.900 1 1 0 1 21 6 2 --
Cladomics cladogram Sinanodonta Anemina Cristaria Anodonta ALASMIDONTINA

Huang et al., 2019 [Publ 3843], fig. 3

cst iT tT bi riS oT roT roS iD
0.818 2 6 0 1 54 8 2 6.0
Cladomics cladogram CRISTARIINA Anodonta cygnea Anodonta anatina ALASMIDONTINA

Huang et al., 2019 [Publ 3843], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 3 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Wu et al., 2019 [Publ 3864], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 45 8 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Froufe et al., 2019 [Publ 3936], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 43 8 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Froufe et al., 2019 [Publ 3936], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 18 3 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Froufe et al., 2019 [Publ 3936], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 18 3 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Froufe et al., 2019 [Publ 3936], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 43 8 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 3 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 3 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Guerra et al., 2019 [Publ 4168], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 6 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Guerra et al., 2019 [Publ 4168], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 3 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Guerra et al., 2019 [Publ 4168], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 6 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Guerra et al., 2019 [Publ 4168], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 3 2 --
Cladomics cladogram ALASMIDONTINA Anodonta CRISTARIINA

Fukata & Iigo, 2020 [Publ 4106], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 52 8 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Wu et al., 2020 [Publ 4121], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 8 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Sano et al., 2020 [Publ 4138], fig. 2

cst iT tT bi riS oT roT roS iD
0.875 2 2 0 1 67 17 2 --
Cladomics cladogram Anodonta ALASMIDONTINA

Sano et al., 2020 [Publ 4138], fig. 3

cst iT tT bi riS oT roT roS iD
0.875 2 2 0 1 67 17 2 --
Cladomics cladogram Anodonta ALASMIDONTINA

Sano et al., 2020 [Publ 4138], fig. 4

cst iT tT bi riS oT roT roS iD
0.875 2 2 0 1 67 17 2 --
Cladomics cladogram Anodonta ALASMIDONTINA

Wu et al., 2020 [Publ 4147], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 6 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Chen et al., 2020 [Publ 4170], fig. 1

cst iT tT bi riS oT roT roS iD
0.692 2 2 0 1 32 9 2 --
Cladomics cladogram ALASMIDONTINA Sinanodonta Cristaria Anodonta

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 8 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 3 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Wu et al., 2021 [Publ 4366], fig. 3-BI

cst iT tT bi riS oT roT roS iD
0.846 2 9 0 1 71 11 2 9.0
Cladomics cladogram CRISTARIINA Anodonta cygnea Anodonta anatina ALASMIDONTINA

Wu et al., 2021 [Publ 4366], fig. 3-ML

cst iT tT bi riS oT roT roS iD
0.846 2 9 0 1 71 11 2 9.0
Cladomics cladogram CRISTARIINA Anodonta cygnea Anodonta anatina ALASMIDONTINA

Wu et al., 2021 [Publ 4366], fig. 4

cst iT tT bi riS oT roT roS iD
0.846 2 8 0 1 66 10 2 8.0
Cladomics cladogram CRISTARIINA Anodonta cygnea Anodonta anatina ALASMIDONTINA

Wu et al., 2022 [Publ 4238], fig. 2d

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 36 8 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Wu et al., 2022 [Publ 4238], fig. 2e

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 3 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Wu et al., 2022 [Publ 4238], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 71 8 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 36 4 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 38 5 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Dai et al., 2023 [Publ 4285], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 6 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Dai et al., 2024 [Publ 4358], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 6 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Wu et al., 2024 [Publ 4388], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 6 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Dai et al., 2025 [Publ 4440], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 85 8 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Chen et al., 2025 [Publ 4448], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 83 9 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Klymus et al., 2025 [Publ 4814], fig. 3A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 3 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Klymus et al., 2025 [Publ 4814], fig. 3B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 3 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 46 3 2 --
Cladomics cladogram CRISTARIINA Anodonta ALASMIDONTINA

Lydeard et al., 1996 [Publ 2656], fig. 4

cst iT tT bi riS oT roT roS iD
0.889 1 1 0 1 27 5 1 --
Cladomics cladogram Utterbackia Utterbackiana Anodonta Strophitus Lasmigona Alasmidonta

Rosenberg et al., 1997 [Publ 2720], fig. 2A

cst iT tT bi riS oT roT roS iD
0.500 1 1 0 1 21 3 1 --
Cladomics cladogram Anodonta Gonidea Pyganodon grandis Utterbackia Pyganodon cataracta Lampsilis straminea Lampsilis teres Megalonaias Obliquaria Theliderma Quadrula Uniomerus Amblema Elliptio Fusconaia Plectomerus Pleurobema UNIONINA

Rosenberg et al., 1997 [Publ 2720], fig. 2B

cst iT tT bi riS oT roT roS iD
0.500 1 1 0 1 21 3 1 --
Cladomics cladogram Anodonta Gonidea Pyganodon grandis Utterbackia Pyganodon cataracta Lampsilis straminea Lampsilis teres Megalonaias Obliquaria Theliderma Quadrula Uniomerus Amblema Elliptio Fusconaia Plectomerus ACULAMPROTULINA UNIONINA

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 5 1 --
Cladomics cladogram Anodonta ALASMIDONTINA

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 5 1 --
Cladomics cladogram Anodonta ALASMIDONTINA

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 2 1 --
Cladomics cladogram Anodonta ALASMIDONTINA

Hoeh et al., 2001 [Publ 2652], fig. 14f2

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 30 2 1 --
Cladomics cladogram Pyganodon Anodonta Strophitus

Hoeh et al., 2001 [Publ 2652], fig. 14f4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 2 1 --
Cladomics cladogram Anodonta ALASMIDONTINA

Hoeh et al., 2002 [Publ 2653], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 2 1 --
Cladomics cladogram Anodonta ALASMIDONTINA

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 30 2 1 --
Cladomics cladogram Pyganodon Anodonta Strophitus

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 2 1 --
Cladomics cladogram Anodonta ALASMIDONTINA

Krebs et al., 2003 [Publ 2691], fig. 3

cst iT tT bi riS oT roT roS iD
0.917 1 1 0 1 32 9 1 --
Cladomics cladogram Utterbackiana Platynaias Alasmidonta Pterosyna Lasmigona Strophitus Utterbackia Pyganodon Anodonta

Mock et al., 2004 [Publ 2686], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 1 1 1 --
Cladomics cladogram Anodonta CRISTARIINA

Mock et al., 2004 [Publ 2686], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 1 1 1 --
Cladomics cladogram Anodonta CRISTARIINA

Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 3 1 1 --
Cladomics cladogram CRISTARIINA Brachyanodon

Walker et al., 2006 [Publ 2666], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 2 1 --
Cladomics cladogram Anodonta ALASMIDONTINA

Walker et al., 2006 [Publ 2666], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 2 1 --
Cladomics cladogram Anodonta ALASMIDONTINA

Walker et al., 2006 [Publ 2666], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 2 1 --
Cladomics cladogram Anodonta ALASMIDONTINA

Cyr et al., 2007 [Publ 2745], fig. 2B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 4 4 1 --
Cladomics cladogram Brachyanodon ALASMIDONTINA

Cyr et al., 2007 [Publ 2745], fig. 2C

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 4 4 1 --
Cladomics cladogram Pseudanodonta ALASMIDONTINA

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

cst iT tT bi riS oT roT roS iD
0.625 2 8 0 1 23 4 1 11.0
Cladomics cladogram OXYNAIINA Anodonta cygnea Unio pictorum 2 Unio pictorum 1 Unio caffer Anodonta ALASMIDONTINA

Soroka, 2010 [Publ 3404], fig. 2-M

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 4 1 1 --
Cladomics cladogram Anodonta UNIONINA

Soroka, 2010 [Publ 3404], fig. 3a-M

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 5 2 1 --
Cladomics cladogram Anodonta CRISTARIINA UNIONINA

Soroka, 2010 [Publ 3404], fig. 3b-M

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 5 2 1 --
Cladomics cladogram Anodonta CRISTARIINA UNIONINA

Soroka, 2010 [Publ 3404], fig. 4-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 1 1 1 --
Cladomics cladogram Anodonta CRISTARIINA

Soroka, 2010 [Publ 3404], fig. 5a-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 1 1 1 --
Cladomics cladogram Anodonta CRISTARIINA

Soroka, 2010 [Publ 3404], fig. 5b-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 1 1 1 --
Cladomics cladogram Anodonta CRISTARIINA

Soroka, 2010 [Publ 3404], fig. 8a

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 4 1 1 --
Cladomics cladogram CRISTARIINA Anodonta

Soroka, 2010 [Publ 3404], fig. 8b

cst iT tT bi riS oT roT roS iD
0.800 2 2 0 1 4 1 1 --
Cladomics cladogram Anodonta UNIONINA

Froufe et al., 2015 [Publ 3664], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 2 1 --
Cladomics cladogram Anodonta ALASMIDONTINA

Froufe et al., 2015 [Publ 3664], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 2 1 --
Cladomics cladogram Anodonta ALASMIDONTINA

Fonseca et al., 2016 [Publ 3775], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 2 1 --
Cladomics cladogram Anodonta ALASMIDONTINA

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 12 2 1 --
Cladomics cladogram Anodonta ALASMIDONTINA

Burzynski et al., 2017 [Publ 3774], fig. 1-top

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 12 2 1 --
Cladomics cladogram Anodonta ALASMIDONTINA

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 2 1 --
Cladomics cladogram Anodonta ALASMIDONTINA

Wen et al., 2017 [Publ 3795], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 15 2 1 --
Cladomics cladogram Anodonta CRISTARIINA

Wen et al., 2017 [Publ 3795], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 2 1 --
Cladomics cladogram Anodonta ALASMIDONTINA

Froufe et al., 2017 [Publ 3802], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 1 1 1 3.0
Cladomics cladogram CRISTARIINA Anodonta

Froufe et al., 2017 [Publ 3802], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 10 10 1 --
Cladomics cladogram Anodonta CRISTARIINA

Froufe et al., 2017 [Publ 3802], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 7 1 1 3.0
Cladomics cladogram CRISTARIINA Anodonta

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 6 2 1 --
Cladomics cladogram Anodonta ALASMIDONTINA

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 6 2 1 --
Cladomics cladogram Anodonta ALASMIDONTINA

Araujo et al., 2018 [Publ 3779], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 2 1 --
Cladomics cladogram Anodonta ALASMIDONTINA

Chase et al., 2018 [Publ 3847], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 1 1 --
Cladomics cladogram Anodonta ALASMIDONTINA

Chase et al., 2018 [Publ 3861], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 2 1 --
Cladomics cladogram Anodonta ALASMIDONTINA

Chase et al., 2018 [Publ 3861], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 2 1 --
Cladomics cladogram Anodonta ALASMIDONTINA

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 27 1 1 --
Cladomics cladogram Brachyanodon ALASMIDONTINA

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 28 1 1 --
Cladomics cladogram Brachyanodon ALASMIDONTINA

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 25 1 1 --
Cladomics cladogram Brachyanodon ALASMIDONTINA

Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 3 3 1 3.0
Cladomics cladogram Anodonta CRISTARIINA

Pfarrer et al., 2022 [Publ 4305], fig. 2-5

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 9 2 1 3.0
Cladomics cladogram CRISTARIINA Anodonta

Whelan et al., 2023 [Publ 4287], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 27 27 1 --
Cladomics cladogram Anodonta ALASMIDONTINA

Whelan et al., 2023 [Publ 4287], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 25 25 1 --
Cladomics cladogram Anodonta ALASMIDONTINA

Bogan et al., 2023 [Publ 4295], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 27 1 --
Cladomics cladogram Anodonta CRISTARIINA

Wu et al., 2023 [Publ 4296], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 3 1 --
Cladomics cladogram Pseudanodonta ALASMIDONTINA

Liu et al., 2023 [Publ 4310], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 23 3 1 --
Cladomics cladogram Pseudanodonta ALASMIDONTINA

Liu et al., 2023 [Publ 4310], fig. 5

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 23 3 1 --
Cladomics cladogram Lasmigona Pseudanodonta Pyganodon Strophitus

Dai et al., 2024 [Publ 4386], fig. 5-BI

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 21 1 1 --
Cladomics cladogram Acuticosta CRISTARIINA Anodonta Lanceolaria ACULAMPROTULINA Lepidodesma UNIONINA MIDDENDORFFINAIINA OXYNAIINA

Dai et al., 2024 [Publ 4386], fig. 5-ML

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 21 1 1 --
Cladomics cladogram Anodonta Acuticosta CRISTARIINA Lepidodesma Lanceolaria ACULAMPROTULINA UNIONINA MIDDENDORFFINAIINA OXYNAIINA

Xiang et al., 2025 [Publ 4433], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 11 7 1 --
Cladomics cladogram Anodonta CRISTARIINA

Altaba et al., 2025 [Publ 4849], fig. 10

cst iT tT bi riS oT roT roS iD
0.778 2 8 0 1 10 6 1 8.0
Cladomics cladogram OXYNAIINA UNIONINA Anodonta anatina Anodonta cygnea CRISTARIINA

Altaba et al., 2025 [Publ 4849], fig. 12

cst iT tT bi riS oT roT roS iD
0.750 2 2 0 1 11 7 1 --
Cladomics cladogram Amuranodonta Cristaria Beringiana Sinanodonta woodiana 2 Sinanodonta woodiana 1 Sinanodonta lauta Sinanodonta tumens Anodonta

Hoeh et al., 1998 [Publ 2695], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 4 0 0 --
Cladomics cladogram Anodonta Margaritifera Amblema

Hoeh et al., 1998 [Publ 2695], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 4 0 0 --
Cladomics cladogram Margaritifera Anodonta

Lydeard et al., 2000 [Publ 2654], fig. 2a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram Anodonta Cumberlandia Reginaia Lampsilis Obovaria QUADRULINA Pleurobema Fusconaia

Lydeard et al., 2000 [Publ 2654], fig. 2b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram Anodonta Cumberlandia Lampsilis Obovaria Reginaia QUADRULINA Pleurobema Fusconaia

Lydeard et al., 2000 [Publ 2654], fig. 2c

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram Anodonta Cumberlandia QUADRULINA Obovaria unicolor Obovaria olivaria Lampsilis Reginaia Pleurobema Fusconaia

Lydeard et al., 2000 [Publ 2654], fig. 2d

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram Anodonta Cumberlandia Reginaia Lampsilis Obovaria QUADRULINA Pleurobema Fusconaia

Huff et al., 2004 [Publ 2697], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 0 0 --
Cladomics cladogram UNIONINA Anodonta

Huff et al., 2004 [Publ 2697], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 0 0 --
Cladomics cladogram Anodonta Margaritifera margaritifera Cumberlandia Pseudunio Margaritifera dahurica Margaritifera middendorffi 1 Margaritifera middendorffi 2 Margaritifera falcata 1 Margaritifera falcata 2 Margaritifera falcata 3 Margaritifera falcata 4

Huff et al., 2004 [Publ 2697], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 0 0 --
Cladomics cladogram UNIONINA Anodonta

Huff et al., 2004 [Publ 2697], fig. 5-6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 0 0 --
Cladomics cladogram UNIONINA Anodonta Lampsilis

Taylor et al., 2007 [Publ 2696], fig. 1-3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 0 0 --
Cladomics cladogram Anodonta UNIONINA

Araujo et al., 2009 [Publ 2589], fig. 7A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 0 0 --
Cladomics cladogram Unio gibbus Unio tumidus Anodonta

Araujo et al., 2009 [Publ 2589], fig. 7B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 0 0 --
Cladomics cladogram Anodonta UNIONINA

Araujo et al., 2009 [Publ 2589], fig. 7C

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 0 0 --
Cladomics cladogram Anodonta UNIONINA

Bolotov et al., 2016 [Publ 3743], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 0 0 --
Cladomics cladogram Anodonta UNIONINA

Bolotov et al., 2016 [Publ 3743], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 0 0 --
Cladomics cladogram Anodonta UNIONINA

Araujo et al., 2017 [Publ 3801], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram Anodonta UNIONINA

Klishko et al., 2019 [Publ 3886], fig. 9

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 9 0 0 --
Cladomics cladogram Anodonta UNIONINA MIDDENDORFFINAIINA OXYNAIINA

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 0 0 --
Cladomics cladogram UNIONINA Anodonta

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram UNIONINA Anodonta

Lopes-Lima et al., 2021 [Publ 4217], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 22 0 0 3.0
Cladomics cladogram Anodonta UNIONINA

Lopes-Lima et al., 2021 [Publ 4217], fig. 11

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 0 0 0 4.0
Cladomics cladogram Anodonta

Sonowal et al., 2021 [Publ 4221], fig. 3a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram Anodonta Unio ravoisieri Unio pictorum ACULAMPROTULINA

Sonowal et al., 2021 [Publ 4221], fig. 3b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram ACULAMPROTULINA Lamprotula Lampsilis Potomida Unio pictorum Anodonta Unio ravoisieri Lamellidens Elliptio

Sonowal et al., 2021 [Publ 4221], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 15 0 0 --
Cladomics cladogram Anodonta UNIONINA Lamellidens

Gomes-dos-Santos et al., 2023 [Publ 4463], fig. 2a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 20 0 0 --
Cladomics cladogram Anodonta OXYNAIINA

Gomes-dos-Santos et al., 2023 [Publ 4463], fig. 2b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 0 0 --
Cladomics cladogram Anodonta OXYNAIINA

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 20 0 0 --
Cladomics cladogram Lanceolaria Anodonta

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 20 0 0 --
Cladomics cladogram Lanceolaria Anodonta

Zieritz et al., 2024 [Publ 4381], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 44 0 0 --
Cladomics cladogram Anodonta UNIONINA

Dai et al., 2024 [Publ 4386], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram Anodonta Lepidodesma

Wu et al., 2026 [Publ 4811], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 15 0 0 --
Cladomics cladogram Acuticosta Brachyanodon