Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini

subtribe ANODONTINA Rafinesque, 1820: cladomics

3 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 11 (24 total)
number of subtaxa (rank: genus) = 3
analyzed in 224 cladograms in 121 publications
maximum number of terminals in an analysis = 10 (mean = 2.03, median = 1)
maximum number of subtaxa (genus) = 3 (mean = 1.43, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 110
maximum number of included terminals = 10, (mean = 3.09 median = 3)
maximum number of subtaxa (genus) = 3 (mean = 1.88, median = 2)
proportion of cladograms that support basic monophyly = 0.6

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 59
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 10 (mean = 3.63, median = 3)
maximum number of subtaxa (genus) = 3 (mean = 2.37, median = 2)
proportion of cladograms that support rigorous monophyly = 0.37

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Anodontina cladogram distances
PCoA Ordination

NJ Tree of Anodontina cladogram distances
NJ Tree

Cladograms

Whelan et al., 2023 [Publ 4287], fig. 2

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
4 37 38 3 37 37 2 0 
  +----------- Cristariina     
  |                            
--+  +-------- Anodonta_cygnea ** 
  |  |                         
  +--+     +-- Brachyanodon **    
     |  +--+                   
     |  |  +-- Pseudanodonta **   
     +--+                      
        |  +-- Anodonta_anatina **
        +--+                   
           +-- Alasmidontina

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
10 10 9 3 621 69 2 47 
  +----------------------- Anodonta_anatina **           
  |                                                   
--+  +-------------------- Pseudanodonta **              
  |  |                                                
  +--+  +----------------- Brachyanodon_oregonensis__2 **
     |  |                                             
     |  |           +----- Anodonta_cygnea **            
     +--+  +--------+                                 
        |  |        |  +-- Anodonta_seddonae **          
        |  |        +--+                              
        +--+           +-- Anodonta_exulcerata **        
           |                                          
           |  +----------- Brachyanodon_impura **        
           |  |                                       
           +--+  +-------- Brachyanodon **               
              |  |                                    
              +--+  +----- Brachyanodon_nuttalliana__1 **
                 |  |                                 
                 +--+----- Brachyanodon_nuttalliana__2 **
                    |                                 
                    |  +-- Brachyanodon_californiensis **
                    +--+                              
                       +-- Brachyanodon_oregonensis__1 **

Chong et al., 2008 [Publ 2668], fig. 3

iTtTbiriSoTroTroSiD
7 18 10 3 11 10 2 46 
  +----------------- Anodonta_cygnea **            
  |                                             
  +----------------- Pyganodon                  
  |                                             
  +----------------- Alasminota                 
  |                                             
  +----------------- Anodonta_anatina **           
  |                                             
  +----------------- Unionina                   
  |                                             
  +----------------- Cristariina                
  |                                             
  |              +-- Brachyanodon_nuttalliana **   
--+--------------+                              
  |              +-- Brachyanodon_californiensis **
  |                                             
  |              +-- Pseudodontoideus           
  |              |                              
  +--------------+-- Alasmidonta                
  |              |                              
  |              +-- Pterosyna                  
  |                                             
  |  +-------------- Pseudanodonta **              
  |  |                                          
  |  |        +----- Strophitus                 
  |  +--------+                                 
  +--+        |  +-- Prolasmidonta              
     |        +--+                              
     |           +-- Platynaias                 
     |                                          
     |  +----------- Brachyanodon_oregonensis__4 **
     +--+                                       
        |  +-------- Brachyanodon_oregonensis__3 **
        +--+                                    
           |  +----- Brachyanodon_oregonensis__2 **
           +--+                                 
              |  +-- Brachyanodon_oregonensis__1 **
              +--+                              
                 +-- Brachyanodon_kennerlyi **

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

iTtTbiriSoTroTroSiD
7 9 7 2 21 6 2 39 
  +-------------- Cristariina                
  |                                          
  |           +-- Anodonta_exulcerata **        
--+  +--------+                              
  |  |        +-- Anodonta_cygnea **            
  |  |                                       
  +--+        +-- Brachyanodon_californiensis **
     |  +-----+                              
     |  |     +-- Brachyanodon_nuttalliana **   
     |  |                                    
     +--+     +-- Brachyanodon_kennerlyi **     
        |  +--+                              
        |  |  +-- Brachyanodon_oregonensis **   
        +--+                                 
           |  +-- Anodonta_anatina **           
           +--+                              
              +-- Alasmidontina

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
5 13 11 3 136 12 2 35 
  +-------------- Cristariina        
  |                                  
--+           +-- Anodonta_cygnea **    
  |  +--------+                      
  |  |        +-- Anodonta_exulcerata **
  +--+                               
     |  +-------- Anodonta_anatina **   
     +--+                            
        |  +----- Alasmidontina      
        +--+                         
           |  +-- Brachyanodon **       
           +--+                      
              +-- Pseudanodonta **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
5 5 3 3 91 13 2 39 
  +----------- Cristariina  
--+                         
  |  +-------- Alasmidontina
  +--+                      
     |  +----- Anodonta **     
     +--+                   
        |  +-- Brachyanodon ** 
        +--+                
           +-- Pseudanodonta **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
5 12 10 3 87 13 2 38 
  +----------- Cristariina  
--+                         
  |  +-------- Brachyanodon ** 
  +--+                      
     |  +----- Alasmidontina
     +--+                   
        |  +-- Pseudanodonta **
        +--+                
           +-- Anodonta **

Lyubas et al., 2022 [Publ 4255], fig. S1

iTtTbiriSoTroTroSiD
5 5 3 2 26 0 0 41 
  +----------- Anodonta_anatina **   
--+                               
  |  +-------- Pseudanodonta **      
  +--+                            
     |  +----- Anodonta_exulcerata **
     +--+                         
        |  +-- Anodonta_seddonae **  
        +--+                      
           +-- Anodonta_cygnea **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
5 12 8 3 91 13 2 36 
  +----------- Cristariina        
  |                               
--+  +-------- Brachyanodon **       
  |  |                            
  |  +-------- Pseudanodonta **      
  +--+                            
     |  +----- Anodonta_cygnea **    
     |  |                         
     +--+----- Alasmidontina      
        |                         
        |  +-- Anodonta_exulcerata **
        +--+                      
           +-- Anodonta_anatina **

Hou et al., 2025 [Publ 4445], fig. 5

iTtTbiriSoTroTroSiD
5 12 10 3 48 13 2 38 
  +----------- Cristariina  
--+                         
  |  +-------- Brachyanodon ** 
  +--+                      
     |  +----- Alasmidontina
     +--+                   
        |  +-- Pseudanodonta **
        +--+                
           +-- Anodonta **

Lyubas et al., 2022 [Publ 4255], fig. 7

iTtTbiriSoTroTroSiD
5 5 3 2 26 0 0 41 
  +-- Pseudanodonta **
--+                
  +-- Anodonta **

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
5 12 10 3 71 13 2 38 
  +-------- Cristariina  
  |                      
--+     +-- Pseudanodonta **
  |  +--+                
  |  |  +-- Anodonta **     
  +--+                   
     |  +-- Brachyanodon ** 
     +--+                
        +-- Alasmidontina

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
5 12 10 3 87 13 2 38 
  +-------- Cristariina  
  |                      
--+     +-- Pseudanodonta **
  |  +--+                
  |  |  +-- Anodonta **     
  +--+                   
     |  +-- Brachyanodon ** 
     +--+                
        +-- Alasmidontina

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
5 12 10 3 49 13 2 38 
  +----------- Cristariina  
--+                         
  |  +-------- Brachyanodon ** 
  +--+                      
     |  +----- Alasmidontina
     +--+                   
        |  +-- Pseudanodonta **
        +--+                
           +-- Anodonta **

Riccardi et al., 2020 [Publ 4089], fig. 2

iTtTbiriSoTroTroSiD
5 9 7 3 14 8 2 39 
  +-------------- Cristariina        
  |                                  
--+  +----------- Anodonta_anatina **   
  |  |                               
  +--+        +-- Brachyanodon **       
     |  +-----+                      
     |  |     +-- Alasmidontina      
     +--+                            
        |  +----- Pseudanodonta **      
        +--+                         
           |  +-- Anodonta_exulcerata **
           +--+                      
              +-- Anodonta_cygnea **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
4 4 2 3 72 10 2 40 
  +-------------- Cristariina     
--+                               
  |  +----------- Alasmidontina   
  +--+                            
     |  +-------- Pseudanodonta **   
     +--+                         
        |  +----- Anodonta_anatina **
        +--+                      
           |  +-- Brachyanodon **    
           +--+                   
              +-- Anodonta_cygnea **

Nagel et al., 1998 [Publ 1946], fig. 7

iTtTbiriSoTroTroSiD
4 5 4 2 1 1 1 41 
  +-------------- Pseudanodonta **   
--+                               
  |  +----------- Anodonta_cygnea ** 
  +--+                            
     |  +-------- Cristariina     
     +--+                         
        |  +----- Anodonta__2 **     
        +--+                      
           |  +-- Anodonta_anatina **
           +--+                   
              +-- Anodonta__1 **

Lopes-Lima et al., 2021 [Publ 4217], fig. 11

iTtTbiriSoTroTroSiD
4 4 2 1 0 0 0 40 
1

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
4 11 4 3 196 16 2 39 
  +----------- Cristariina     
  |                            
  |  +-------- Alasmidonta     
  |  |                         
--+  +-------- Lasmigona       
  |  |                         
  |  +-------- Pseudanodonta **   
  |  |                         
  |  +-------- Simpsonaias     
  +--+                         
     +-------- Strophitus      
     |                         
     |     +-- Pyganodon       
     +-----+                   
     |     +-- Utterbackia     
     |                         
     |  +----- Anodonta_anatina **
     +--+                      
        |  +-- Anodonta_cygnea ** 
        +--+                   
           +-- Brachyanodon **

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
4 11 9 2 91 21 2 37 
  +-------- Cristariina  
--+                      
  |  +----- Anodonta **     
  +--+                   
     |  +-- Pseudanodonta **
     +--+                
        +-- Alasmidontina

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
4 9 3 3 134 11 2 40 
  +----- Anodonta_anatina **
  |                      
  +----- Pseudanodonta **   
--+                      
  +----- Alasmidontina   
  |                      
  |  +-- Anodonta_cygnea ** 
  +--+                   
     +-- Brachyanodon **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
4 17 0 3 120 11 2 46 
  +-- Alasmidonta          
  |                        
  +-- Anodonta_anatina **     
  |                        
  +-- Anodonta_cygnea **      
  |                        
  +-- Brachyanodon **         
  |                        
  +-- Lasmigona            
  |                        
  +-- Pseudanodonta **        
  |                        
  +-- Pyganodon            
  |                        
  +-- Simpsonaias          
  |                        
  +-- Strophitus           
--+                        
  +-- Sinanodonta_lauta    
  |                        
  +-- Cristaria_plicata    
  |                        
  +-- Cristaria_radiata    
  |                        
  +-- Pletholophus         
  |                        
  +-- Sinanodonta_lucida   
  |                        
  +-- Sinanodonta_jourdyi  
  |                        
  +-- Lanceolaria_gladiola 
  |                        
  +-- Lanceolaria_grayii__1
  |                        
  +-- Lanceolaria_grayii__2

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
4 11 8 3 194 16 2 42 
  +-------------- Cristariina     
  |                               
  |           +-- Alasmidonta     
  |  +--------+                   
--+  |        +-- Lasmigona       
  |  |                            
  |  |     +----- Anodonta_anatina **
  |  +-----+                      
  |  |     |  +-- Anodonta_cygnea ** 
  +--+     +--+                   
     |        +-- Brachyanodon **    
     |                            
     |        +-- Pyganodon       
     |  +-----+                   
     |  |     +-- Utterbackia     
     +--+                         
        |  +----- Pseudanodonta **   
        +--+                      
           |  +-- Simpsonaias     
           +--+                   
              +-- Strophitus

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
4 9 4 3 118 29 2 41 
  +----------- Cristariina     
  |                            
--+  +-------- Pseudanodonta **   
  |  |                         
  +--+-------- Alasmidontina   
     |                         
     |  +----- Anodonta_cygnea ** 
     +--+                      
        |  +-- Anodonta_anatina **
        +--+                   
           +-- Brachyanodon **

Riccardi et al., 2020 [Publ 4089], fig. 1

iTtTbiriSoTroTroSiD
4 4 2 2 2 2 1 38 
  +-------- Cristariina        
  |                            
--+     +-- Anodonta_anatina **   
  |  +--+                      
  |  |  +-- Pseudanodonta **      
  +--+                         
     |  +-- Anodonta_exulcerata **
     +--+                      
        +-- Anodonta_cygnea **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
4 47 24 3 196 16 2 73 
  +-------- Acuticosta           
  |                              
  +-------- Alasmidonta          
  |                              
  +-------- Anodonta_anatina **     
  |                              
  +-------- Anemina              
  |                              
  +-------- Anodonta_cygnea **      
  |                              
  +-------- Cristaria            
  |                              
  +-------- Brachyanodon **         
  |                              
  +-------- Lasmigona            
  |                              
  +-------- Lepidodesma          
  |                              
  +-------- Pseudanodonta **        
  |                              
  +-------- Pyganodon            
  |                              
--+-------- Simpsonaias          
  |                              
  +-------- Strophitus           
  |                              
  +-------- Utterbackia          
  |                              
  +-------- Lanceolaria          
  |                              
  +-------- Schistodesmus        
  |                              
  +-------- Unionina             
  |                              
  +-------- Aculamprotulina      
  |                              
  |     +-- Pletholophus         
  +-----+                        
  |     +-- Sinanodonta          
  |                              
  |  +----- Cuneopsis_celtiformis
  |  |                           
  |  +----- Tchangsinaia         
  |  |                           
  +--+----- Cuneopsis_rufescens  
     |                           
     +----- Nodularia            
     |                           
     |  +-- Cuneopsis_szechenyii 
     +--+                        
        +-- Cuneopsis_heudei

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
4 4 2 3 72 10 2 40 
  +----------- Cristariina  
--+                         
  |  +-------- Alasmidontina
  +--+                      
     |  +----- Pseudanodonta **
     +--+                   
        |  +-- Brachyanodon ** 
        +--+                
           +-- Anodonta **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
4 4 1 3 76 10 2 39 
  +----------- Cristariina     
  |                            
--+  +-------- Alasmidontina   
  |  |                         
  +--+  +----- Pseudanodonta **   
     |  |                      
     +--+  +-- Anodonta_anatina **
        |  |                   
        +--+-- Anodonta_cygnea ** 
           |                   
           +-- Brachyanodon **

Nagel et al., 1996 [Publ 3400], fig. 351

iTtTbiriSoTroTroSiD
4 5 4 2 1 1 1 41 
  +-------------- Pseudanodonta **   
--+                               
  |  +----------- Anodonta_cygnea ** 
  +--+                            
     |  +-------- Cristariina     
     +--+                         
        |  +----- Anodonta__2 **     
        +--+                      
           |  +-- Anodonta_anatina **
           +--+                   
              +-- Anodonta__1 **

Tomilova et al., 2020 [Publ 4097], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 2 4 2 1 40 
  +-------- Anodonta_anatina **   
--+                            
  |  +----- Pseudanodonta **      
  +--+                         
     |  +-- Anodonta_cygnea **    
     +--+                      
        +-- Anodonta_exulcerata **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
4 9 7 3 134 11 2 43 
  +-------------- Cristariina     
--+                               
  |  +----------- Anodonta_anatina **
  +--+                            
     |  +-------- Alasmidontina   
     +--+                         
        |  +----- Pseudanodonta **   
        +--+                      
           |  +-- Anodonta_cygnea ** 
           +--+                   
              +-- Brachyanodon **

Wang et al., 2025 [Publ 4488], fig. 3B

iTtTbiriSoTroTroSiD
4 11 9 2 61 22 2 37 
  +-------- Cristariina  
--+                      
  |  +----- Anodonta **     
  +--+                   
     |  +-- Pseudanodonta **
     +--+                
        +-- Alasmidontina

Wu et al., 2023 [Publ 4339], fig. 5

iTtTbiriSoTroTroSiD
3 3 1 2 16 6 2 39 
  +----------- Cristariina     
--+                            
  |  +-------- Alasmidontina   
  +--+                         
     |  +----- Anodonta_anatina **
     +--+                      
        |  +-- Pseudanodonta **   
        +--+                   
           +-- Anodonta_cygnea **

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iTtTbiriSoTroTroSiD
3 20 0 1 102 17 2 42 
  +-- Alasmidonta_marginata      
  |                              
  +-- Prolasmidonta              
  |                              
  +-- Alasmidonta_undulata       
  |                              
  +-- Cristariina                
  |                              
  +-- Brachyanodon_kennerlyi **     
  |                              
  +-- Brachyanodon_nuttalliana **   
  |                              
  +-- Brachyanodon_oregonensis **   
  |                              
  +-- Arcidens_confragosus       
  |                              
  +-- Arcidens_wheeleri          
  |                              
  +-- Pterosyna                  
  |                              
--+-- Platynaias_compressa       
  |                              
  +-- Lasmigona                  
  |                              
  +-- Platynaias_decorata        
  |                              
  +-- Platynaias_subviridis      
  |                              
  +-- Pyganodon_grandis          
  |                              
  +-- Pyganodon_lacustris        
  |                              
  +-- Pseudodontoideus           
  |                              
  +-- Strophitus                 
  |                              
  +-- Utterbackia_imbecillis     
  |                              
  +-- Utterbackia_peninsularis__1
  |                              
  +-- Utterbackia_peninsularis__2

Plazzi et al., 2011 [Publ 3676], fig. 5

iTtTbiriSoTroTroSiD
3 17 10 2 15 2 2 52 
  +----------------- Cristariina  
  |                               
  |  +-------------- Lanceolaria  
  |  |                            
--+  +-------------- Anodonta **     
  |  |                            
  |  +-------------- Unionina     
  |  |                            
  |  |           +-- Alasmidontina
  +--+-----------+                
     |           +-- Pseudanodonta **
     |                            
     |           +-- Alathyria    
     |           |                
     |  +--------+-- Pronodularia 
     |  |        |                
     |  |        +-- Sinohyriopsis
     +--+                         
        |  +-------- Quadrulina   
        |  |                      
        +--+-------- Parvaspina   
           |                      
           |  +----- Lampsilis    
           +--+                   
              |  +-- Epioblasma   
              +--+                
                 +-- Venustaconcha

Manendo et al., 2008 [Publ 2669], fig. 1

iTtTbiriSoTroTroSiD
3 5 3 2 10 2 1 39 
  +----- Anodonta **     
--+                   
  |  +-- Pseudanodonta **
  +--+                
     +-- Alasmidontina

Soroka, 2010 [Publ 3404], fig. 5a

iTtTbiriSoTroTroSiD
3 5 3 2 5 2 1 41 
  +-------- Anodonta_anatina **
--+                         
  |  +----- Cristariina     
  +--+                      
     |  +-- Anodonta_cygnea ** 
     +--+                   
        +-- Pseudanodonta **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
3 3 1 2 53 13 1 39 
  +-------- Cristariina     
--+                         
  |  +----- Anodonta_anatina **
  +--+                      
     |  +-- Pseudanodonta **   
     +--+                   
        +-- Anodonta_cygnea **

Soroka, 2010 [Publ 3404], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 2 4 1 1 37 
  +----- Anodonta_cygnea ** 
--+                      
  |  +-- Anodonta_anatina **
  +--+                   
     +-- Pseudanodonta **

Pfarrer et al., 2022 [Publ 4305], fig. 2-5

iTtTbiriSoTroTroSiD
3 3 1 1 9 2 1 39 
  +-- Cristariina
--+              
  +-- Anodonta **

Froufe et al., 2017 [Publ 3802], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 1 1 1 1 39 
  +-- Cristariina
--+              
  +-- Anodonta **

Soroka, 2010 [Publ 3404], fig. 3a

iTtTbiriSoTroTroSiD
3 3 1 2 6 2 1 39 
  +----- Anodonta_anatina **
--+                      
  |  +-- Anodonta_cygnea ** 
  +--+                   
     +-- Pseudanodonta **

Nagel et al., 1998 [Publ 1946], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 2 6 0 0 39 
  +-- Pseudanodonta **
--+                
  +-- Anodonta **

Reis et al., 2013 [Publ 3580], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 2 11 0 0 37 
  +----- Anodonta_cygnea ** 
--+                      
  |  +-- Anodonta_anatina **
  +--+                   
     +-- Pseudanodonta **

Soroka, 2010 [Publ 3404], fig. 3b

iTtTbiriSoTroTroSiD
3 9 5 2 6 2 1 44 
  +----- Anodonta_anatina **
  |                      
  +----- Anodonta_cygnea ** 
--+                      
  +----- Pseudanodonta **   
  |                      
  |  +-- Cristariina     
  +--+                   
     +-- Unionina

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 2 33 9 1 39 
  +-------- Cristariina     
--+                         
  |  +----- Anodonta_anatina **
  +--+                      
     |  +-- Pseudanodonta **   
     +--+                   
        +-- Anodonta_cygnea **

Mezhzherin et al., 2014 [Publ 3705], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 2 1 1 1 39 
  +----- Cristariina  
--+                   
  |  +-- Pseudanodonta **
  +--+                
     +-- Anodonta **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
3 6 4 2 88 7 2 40 
  +-------------- Cristariina     
  |                               
--+  +----------- Anodonta_cygnea ** 
  |  |                            
  +--+        +-- Lasmigona       
     |  +-----+                   
     |  |     +-- Anodonta_anatina **
     +--+                         
        |  +----- Pseudanodonta **   
        +--+                      
           |  +-- Pyganodon       
           +--+                   
              +-- Strophitus

Froufe et al., 2017 [Publ 3802], fig. 7

iTtTbiriSoTroTroSiD
3 3 1 1 7 1 1 39 
  +-- Cristariina
--+              
  +-- Anodonta **

Perkins et al., 2025 [Publ 4447], fig. 2_mtDNA-ITS

iTtTbiriSoTroTroSiD
3 36 37 2 33 33 1 32 
  +----- Anodonta **     
--+                   
  |  +-- Pseudanodonta **
  +--+                
     +-- Alasmidontina

Lopes-Lima et al., 2016 [Publ 3757], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 2 0 0 0 39 
  +-- Pseudanodonta **
--+                
  +-- Anodonta **

Froufe et al., 2014 [Publ 3592], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 2 0 0 0 39 
  +----- Anodonta_anatina **
--+                      
  |  +-- Pseudanodonta **   
  +--+                   
     +-- Anodonta_cygnea **

Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8

iTtTbiriSoTroTroSiD
3 3 1 1 3 3 1 39 
  +-- Anodonta **   
--+              
  +-- Cristariina

Källersjö et al., 2005 [Publ 2684], fig. 1B-F

iTtTbiriSoTroTroSiD
3 3 1 2 4 0 0 39 
  +----- Anodonta_anatina **
--+                      
  |  +-- Pseudanodonta **   
  +--+                   
     +-- Anodonta_cygnea **

Soroka, 2010 [Publ 3404], fig. 5b

iTtTbiriSoTroTroSiD
3 8 5 2 5 2 1 44 
  +-------- Anodonta_anatina **
  |                         
--+  +----- Cristariina     
  |  |                      
  +--+----- Unionina        
     |                      
     |  +-- Anodonta_cygnea ** 
     +--+                   
        +-- Pseudanodonta **

Källersjö et al., 2005 [Publ 2684], fig. 1A

iTtTbiriSoTroTroSiD
3 3 0 2 4 0 0 38 
  +-- Anodonta_anatina **
  |                   
--+-- Pseudanodonta **   
  |                   
  +-- Anodonta_cygnea **

Lopes-Lima et al., 2021 [Publ 4217], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 1 22 0 0 39 
1

Soroka, 2010 [Publ 3404], fig. 4

iTtTbiriSoTroTroSiD
3 7 5 2 4 1 1 43 
        +-- Anodonta_anatina **
  +-----+                   
  |     +-- Unionina        
--+                         
  |  +----- Cristariina     
  +--+                      
     |  +-- Anodonta_cygnea ** 
     +--+                   
        +-- Pseudanodonta **

Mezhzherin et al., 2014 [Publ 3705], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 2 1 1 1 39 
  +-------- Cristariina     
--+                         
  |  +----- Anodonta_anatina **
  +--+                      
     |  +-- Anodonta_cygnea ** 
     +--+                   
        +-- Pseudanodonta **

Lopes-Lima et al., 2017 [Publ 3734], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 2 14 0 0 39 
  +----- Anodonta_anatina **
--+                      
  |  +-- Pseudanodonta **   
  +--+                   
     +-- Anodonta_cygnea **

Prié & Puillandre, 2014 [Publ 3529], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 2 14 0 0 39 
  +----- Anodonta_anatina **
--+                      
  |  +-- Pseudanodonta **   
  +--+                   
     +-- Anodonta_cygnea **

Hinzmann et al., 2013 [Publ 3541], fig. 5

iTtTbiriSoTroTroSiD
3 3 1 2 0 0 0 39 
  +-- Pseudanodonta **
--+                
  +-- Anodonta **

Reis et al., 2013 [Publ 3580], fig. 5

iTtTbiriSoTroTroSiD
3 3 0 2 12 0 0 38 
  +-- Anodonta_anatina **
  |                   
--+-- Anodonta_cygnea ** 
  |                   
  +-- Pseudanodonta **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
3 5 3 2 88 7 2 39 
  +-------------- Cristariina  
--+                            
  |  +----------- Lasmigona    
  +--+                         
     |  +-------- Anodonta **     
     +--+                      
        |  +----- Pseudanodonta **
        +--+                   
           |  +-- Pyganodon    
           +--+                
              +-- Strophitus

Zieritz et al., 2012 [Publ 3673], fig. 4

iTtTbiriSoTroTroSiD
3 3 0 2 5 1 1 38 
  +-- Pseudanodonta **   
  |                   
--+-- Anodonta_anatina **
  |                   
  +-- Anodonta_cygnea **

Xiang et al., 2025 [Publ 4433], fig. 7

iTtTbiriSoTroTroSiD
2 2 0 1 11 7 1 -- 
  +-- Anodonta **   
--+              
  +-- Cristariina

Nagel et al., 1998 [Publ 1946], fig. 9

iTtTbiriSoTroTroSiD
2 2 0 2 4 0 0 -- 
  +-- Anodonta **     
--+                
  +-- Pseudanodonta **

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
2 7 0 1 51 11 2 38 
  +-------- Cristariina     
--+                         
  |  +----- Anodonta_cygnea ** 
  +--+                      
     |  +-- Anodonta_anatina **
     +--+                   
        +-- Alasmidontina

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 39 8 2 -- 
  +-------- Cristariina  
--+                      
  |  +----- Alasmidontina
  +--+                   
     |  +-- Anodonta **     
     +--+                
        +-- Pseudanodonta **

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 39 8 2 -- 
  +-------- Cristariina  
--+                      
  |  +----- Alasmidontina
  +--+                   
     |  +-- Anodonta **     
     +--+                
        +-- Pseudanodonta **

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

iTtTbiriSoTroTroSiD
2 2 0 1 20 0 0 -- 
1

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 66 17 2 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 3 1 1 -- 
  +-- Cristariina 
--+               
  +-- Brachyanodon **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
2 9 0 1 72 11 2 33 
  +-------- Cristariina     
--+                         
  |  +----- Anodonta_cygnea ** 
  +--+                      
     |  +-- Anodonta_anatina **
     +--+                   
        +-- Alasmidontina

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 21 8 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 39 8 2 -- 
  +----- Alasmidontina
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Pseudanodonta **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 72 8 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Froufe et al., 2017 [Publ 3802], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 1 10 10 1 -- 
  +-- Anodonta **   
--+              
  +-- Cristariina

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 66 17 2 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Soroka, 2010 [Publ 3404], fig. 8b

iTtTbiriSoTroTroSiD
2 2 0 1 4 1 1 -- 
1

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

iTtTbiriSoTroTroSiD
2 2 0 1 20 0 0 -- 
1

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
2 6 0 1 54 8 2 34 
  +-------- Cristariina     
--+                         
  |  +----- Anodonta_cygnea ** 
  +--+                      
     |  +-- Anodonta_anatina **
     +--+                   
        +-- Alasmidontina

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 52 8 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Zanatta et al., 2007 [Publ 2698], fig. 1A

iTtTbiriSoTroTroSiD
2 13 0 2 14 14 1 40 
           +-- Platynaias   
           |                
  +--------+-- Lasmigona    
  |        |                
  |        +-- Pterosyna    
--+                         
  |  +-------- Anodonta **     
  |  |                      
  +--+     +-- Brachyanodon ** 
     |  +--+                
     |  |  +-- Utterbackiana
     +--+                   
        |  +-- Utterbackia  
        +--+                
           +-- Pyganodon

Campbell et al., 2005 [Publ 2660], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 100 4 1 -- 
  +-------- Pyganodon       
  |                         
  |     +-- Lasmigona       
--+-----+                   
  |     +-- Pseudodontoideus
  |                         
  |  +----- Alasminota      
  +--+                      
     |  +-- Anodonta **        
     +--+                   
        +-- Brachyanodon **

Soroka, 2010 [Publ 3404], fig. 8a

iTtTbiriSoTroTroSiD
2 2 0 1 4 1 1 -- 
  +-- Cristariina
--+              
  +-- Anodonta **

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 43 8 2 -- 
  +----- Alasmidontina
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Pseudanodonta **

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iTtTbiriSoTroTroSiD
2 8 0 1 23 4 1 41 
  +----------- Anodonta_cygnea ** 
  |                            
--+  +-------- Unio_pictorum__2
  |  |                         
  +--+     +-- Unio_pictorum__1
     |  +--+                   
     |  |  +-- Unio_caffer     
     +--+                      
        |  +-- Anodonta **        
        +--+                   
           +-- Alasmidontina

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 38 9 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
2 9 0 1 72 11 2 33 
  +-------- Cristariina     
--+                         
  |  +----- Anodonta_cygnea ** 
  +--+                      
     |  +-- Anodonta_anatina **
     +--+                   
        +-- Alasmidontina

Hoeh, 1990 [Publ 2717], fig. 4

iTtTbiriSoTroTroSiD
2 5 0 2 13 13 1 39 
           +-- Pterosyna    
           |                
  +--------+-- Platynaias   
  |        |                
  |        +-- Lasmigona    
--+                         
  |        +-- Utterbackia  
  |  +-----+                
  |  |     +-- Pyganodon    
  +--+                      
     |  +----- Anodonta **     
     +--+                   
        |  +-- Brachyanodon ** 
        +--+                
           +-- Utterbackiana

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 39 8 2 -- 
  +----- Alasmidontina
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Pseudanodonta **

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 32 9 2 -- 
  +-------- Alasmidontina
--+                      
  |  +----- Sinanodonta  
  +--+                   
     |  +-- Cristaria    
     +--+                
        +-- Anodonta **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 1 66 17 2 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Wu et al., 2026 [Publ 4811], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 15 0 0 -- 
1

Bogatov et al., 2018 [Publ 3859], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 4 0 0 -- 
  +-- Anodonta **     
--+                
  +-- Pseudanodonta **

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 43 8 2 -- 
  +----- Alasmidontina
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Pseudanodonta **

Chase et al., 2018 [Publ 3847], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 1 37 12 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Whelan et al., 2023 [Publ 4287], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 1 25 25 1 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
2 8 0 1 67 10 2 34 
  +-------- Cristariina     
--+                         
  |  +----- Anodonta_cygnea ** 
  +--+                      
     |  +-- Anodonta_anatina **
     +--+                   
        +-- Alasmidontina

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 86 8 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5_7

iTtTbiriSoTroTroSiD
2 2 0 2 3 0 0 -- 
  +-- Pseudanodonta **
--+                
  +-- Anodonta **

Campbell et al., 2005 [Publ 2660], fig. 1

iTtTbiriSoTroTroSiD
2 6 0 2 100 4 1 41 
  +----------- Brachyanodon **    
  |                            
--+        +-- Anodonta **        
  |  +-----+                   
  |  |     +-- Alasminota      
  +--+                         
     |  +----- Pyganodon       
     +--+                      
        |  +-- Lasmigona       
        +--+                   
           +-- Pseudodontoideus

Hoeh, 1990 [Publ 2717], fig. 2

iTtTbiriSoTroTroSiD
2 5 0 2 13 13 1 39 
           +-- Pterosyna    
           |                
  +--------+-- Platynaias   
  |        |                
  |        +-- Lasmigona    
--+                         
  |        +-- Utterbackia  
  |  +-----+                
  |  |     +-- Pyganodon    
  +--+                      
     |  +----- Anodonta **     
     +--+                   
        |  +-- Brachyanodon ** 
        +--+                
           +-- Utterbackiana

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 84 9 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Klishko et al., 2019 [Publ 3886], fig. 9

iTtTbiriSoTroTroSiD
2 2 0 1 9 0 0 -- 
1

Bogatov et al., 2018 [Publ 3859], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 4 0 0 -- 
  +-- Anodonta **     
--+                
  +-- Pseudanodonta **

Zanatta et al., 2007 [Publ 2698], fig. 1B

iTtTbiriSoTroTroSiD
2 6 0 2 14 14 1 39 
           +-- Lasmigona    
           |                
  +--------+-- Platynaias   
  |        |                
  |        +-- Pterosyna    
--+                         
  |        +-- Utterbackia  
  |  +-----+                
  |  |     +-- Pyganodon    
  +--+                      
     |  +----- Anodonta **     
     +--+                   
        |  +-- Brachyanodon ** 
        +--+                
           +-- Utterbackiana

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
2 2 0 1 37 8 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Hoeh, 1990 [Publ 2717], fig. 3

iTtTbiriSoTroTroSiD
2 15 0 2 13 13 1 46 
  +----- Pyganodon_fragilis         
  |                                 
  +----- Utterbackia_imbecillis     
  |                                 
  +----- Utterbackia_peggyae        
  |                                 
  +----- Anodonta **                   
  |                                 
  +----- Brachyanodon **               
  |                                 
  +----- Utterbackiana_couperiana__1
  |                                 
--+----- Utterbackiana_couperiana__2
  |                                 
  +----- Utterbackiana_suborbiculata
  |                                 
  |  +-- Lasmigona                  
  +--+                              
  |  +-- Utterbackiana_implicata    
  |                                 
  |  +-- Pterosyna                  
  |  |                              
  |  +-- Platynaias                 
  |  |                              
  |  +-- Pyganodon_cataracta        
  +--+                              
     +-- Pyganodon_gibbosa          
     |                              
     +-- Pyganodon_grandis          
     |                              
     +-- Pyganodon_lacustris

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
1 1 0 1 22 3 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
1 1 0 1 21 1 1 -- 
-- Anodonta_anatina **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 48 8 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 43 8 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Hoeh et al., 1998 [Publ 2695], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Anodonta_cygnea **

Huff et al., 2004 [Publ 2697], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Anodonta **

Soroka, 2010 [Publ 3404], fig. 3b-M

iTtTbiriSoTroTroSiD
1 1 0 1 5 2 1 -- 
-- Anodonta_anatina **

Froufe et al., 2015 [Publ 3664], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 7 2 1 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Anodonta_cygnea **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iTtTbiriSoTroTroSiD
1 1 0 1 29 5 1 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Huff et al., 2004 [Publ 2697], fig. 5-6

iTtTbiriSoTroTroSiD
1 1 0 1 11 0 0 -- 
-- Anodonta **

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 3 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Lydeard et al., 2000 [Publ 2654], fig. 2d

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Anodonta_cygnea **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 81 9 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Araujo et al., 2009 [Publ 2589], fig. 7C

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Anodonta_cygnea **

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Anodonta_anatina **

Lydeard et al., 2000 [Publ 2654], fig. 2b

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Anodonta_cygnea **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 45 8 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Burzynski & Soroka, 2018 [Publ 3862], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 3 3 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Araujo et al., 2009 [Publ 2589], fig. 7B

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Anodonta_cygnea **

Mock et al., 2004 [Publ 2686], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 1 1 1 -- 
  +-- Anodonta **   
--+              
  +-- Cristariina

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Anodonta_cygnea **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
1 1 0 1 38 5 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Soroka, 2008 [Publ 3695], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 3 2 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 43 8 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 6 2 1 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Soroka, 2010 [Publ 3404], fig. 5a-M

iTtTbiriSoTroTroSiD
1 1 0 1 1 1 1 -- 
  +-- Anodonta **   
--+              
  +-- Cristariina

Soroka, 2010 [Publ 3404], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 4 1 1 -- 
-- Anodonta_anatina **

Cyr et al., 2007 [Publ 2745], fig. 2B

iTtTbiriSoTroTroSiD
1 1 0 1 4 4 1 -- 
  +-- Brachyanodon ** 
--+                
  +-- Alasmidontina

Bolotov et al., 2016 [Publ 3743], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Anodonta_anatina **

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 25 1 1 -- 
  +-- Brachyanodon ** 
--+                
  +-- Alasmidontina

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 23 3 1 -- 
  +-------- Lasmigona    
--+                      
  |  +----- Pseudanodonta **
  +--+                   
     |  +-- Pyganodon    
     +--+                
        +-- Strophitus

Wu et al., 2022 [Publ 4238], fig. 2e

iTtTbiriSoTroTroSiD
1 1 0 1 17 3 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 29 6 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
1 1 0 1 19 3 2 -- 
  +-- Cristariina  
  |                
--+-- Anodonta **     
  |                
  +-- Alasmidontina

Walker et al., 2006 [Publ 2666], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 14 2 1 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 6 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Lydeard et al., 2000 [Publ 2654], fig. 2a

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Anodonta_cygnea **

Chase et al., 2018 [Publ 3847], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 9 1 1 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Feng et al., 2016 [Publ 3800], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 13 5 2 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Wen et al., 2017 [Publ 3795], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 5 2 1 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 44 0 0 -- 
-- Anodonta_cygnea **

Dai et al., 2024 [Publ 4386], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Anodonta_anatina **

Chase et al., 2018 [Publ 3861], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 10 2 1 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iTtTbiriSoTroTroSiD
1 1 0 1 12 2 1 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 37 8 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Rosenberg et al., 1997 [Publ 2720], fig. 2A

iTtTbiriSoTroTroSiD
1 1 0 1 21 3 1 -- 
-- Anodonta_anatina **

Soroka, 2010 [Publ 3404], fig. 3a-M

iTtTbiriSoTroTroSiD
1 1 0 1 5 2 1 -- 
-- Anodonta_anatina **

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
1 1 0 1 24 7 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 36 4 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Cyr et al., 2007 [Publ 2745], fig. 2C

iTtTbiriSoTroTroSiD
1 1 0 1 4 4 1 -- 
  +-- Pseudanodonta **
--+                
  +-- Alasmidontina

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
1 1 0 1 30 2 1 -- 
  +----- Pyganodon 
--+                
  |  +-- Anodonta **  
  +--+             
     +-- Strophitus

Hoeh et al., 1998 [Publ 2695], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Anodonta_cygnea **

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 6 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iTtTbiriSoTroTroSiD
1 1 0 1 12 2 1 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Lydeard et al., 2000 [Publ 2654], fig. 2c

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Anodonta_cygnea **

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Anodonta_anatina **

Araujo et al., 2009 [Publ 2589], fig. 7A

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Anodonta_cygnea **

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
1 1 0 1 21 1 1 -- 
-- Anodonta_anatina **

Fonseca et al., 2016 [Publ 3775], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 8 2 1 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Soroka, 2010 [Publ 3404], fig. 4-M

iTtTbiriSoTroTroSiD
1 1 0 1 1 1 1 -- 
  +-- Anodonta **   
--+              
  +-- Cristariina

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 3 2 -- 
  +----- Alasmidontina
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Cristariina

Huff et al., 2004 [Publ 2697], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 7 0 0 -- 
-- Anodonta **

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 41 8 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Froufe et al., 2015 [Publ 3664], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 11 2 1 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 31 8 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 46 3 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
1 1 0 1 30 2 1 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Froufe et al., 2019 [Publ 3936], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 18 3 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
1 1 0 1 30 2 1 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Anodonta_anatina **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iTtTbiriSoTroTroSiD
1 1 0 1 9 3 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iTtTbiriSoTroTroSiD
1 1 0 1 21 3 1 -- 
-- Anodonta **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
1 1 0 1 30 2 1 -- 
  +----- Pyganodon 
--+                
  |  +-- Anodonta **  
  +--+             
     +-- Strophitus

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 23 3 1 -- 
  +-- Pseudanodonta **
--+                
  +-- Alasmidontina

Froufe et al., 2019 [Publ 3936], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 18 3 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Taylor et al., 2007 [Publ 2696], fig. 1-3

iTtTbiriSoTroTroSiD
1 1 0 1 3 0 0 -- 
-- Anodonta_cygnea **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 3 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 6 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 27 1 1 -- 
  +-- Brachyanodon ** 
--+                
  +-- Alasmidontina

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 21 6 2 -- 
  +----------- Sinanodonta  
--+                         
  |  +-------- Anemina      
  +--+                      
     |  +----- Cristaria    
     +--+                   
        |  +-- Anodonta **     
        +--+                
           +-- Alasmidontina

Huff et al., 2004 [Publ 2697], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 11 0 0 -- 
-- Anodonta **

Krebs et al., 2003 [Publ 2691], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 32 9 1 -- 
  +----------- Utterbackiana
  |                         
  +----------- Platynaias   
  |                         
  +----------- Alasmidonta  
--+                         
  |        +-- Pterosyna    
  +--------+                
  |        +-- Lasmigona    
  |                         
  |  +-------- Strophitus   
  +--+                      
     |  +----- Utterbackia  
     +--+                   
        |  +-- Pyganodon    
        +--+                
           +-- Anodonta **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 52 6 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 15 0 0 -- 
-- Anodonta_cygnea **

Chase et al., 2018 [Publ 3861], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 10 2 1 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 20 6 2 -- 
  +----------- Sinanodonta  
--+                         
  |  +-------- Anemina      
  +--+                      
     |  +----- Cristaria    
     +--+                   
        |  +-- Anodonta **     
        +--+                
           +-- Alasmidontina

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 9 2 1 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Bolotov et al., 2016 [Publ 3743], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Anodonta_anatina **

Bogan et al., 2023 [Publ 4295], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 30 27 1 -- 
  +-- Anodonta **   
--+              
  +-- Cristariina

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 28 1 1 -- 
  +-- Brachyanodon ** 
--+                
  +-- Alasmidontina

Wen et al., 2017 [Publ 3795], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 15 2 1 -- 
  +-- Anodonta **   
--+              
  +-- Cristariina

Walker et al., 2006 [Publ 2666], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 14 2 1 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Mock et al., 2004 [Publ 2686], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 1 1 1 -- 
  +-- Anodonta **   
--+              
  +-- Cristariina

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 6 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Soroka, 2008 [Publ 3695], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 3 2 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Soroka, 2010 [Publ 3404], fig. 5b-M

iTtTbiriSoTroTroSiD
1 1 0 1 1 1 1 -- 
  +-- Anodonta **   
--+              
  +-- Cristariina

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 52 6 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 2 1 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 6 2 1 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
1 1 0 1 24 7 2 -- 
  +-------- Sinanodonta  
--+                      
  |  +----- Cristaria    
  +--+                   
     |  +-- Anodonta **     
     +--+                
        +-- Alasmidontina

Whelan et al., 2023 [Publ 4287], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 27 27 1 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iTtTbiriSoTroTroSiD
1 1 0 1 29 5 1 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 6 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 22 3 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Walker et al., 2006 [Publ 2666], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 2 1 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 3 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
1 1 0 1 47 8 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
1 1 0 1 16 3 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 22 3 1 -- 
  +-- Pseudanodonta **
--+                
  +-- Alasmidontina

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 30 8 2 -- 
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Lydeard et al., 1996 [Publ 2656], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 27 5 1 -- 
     +----- Utterbackia  
  +--+                   
  |  |  +-- Utterbackiana
  |  +--+                
--+     +-- Anodonta **     
  |                      
  |  +----- Strophitus   
  +--+                   
     |  +-- Lasmigona    
     +--+                
        +-- Alasmidonta

Araujo et al., 2018 [Publ 3779], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 17 2 1 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 2 1 -- 
  +-- Anodonta **     
--+                
  +-- Alasmidontina