Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae

tribe ANODONTINI Rafinesque, 1820: cladomics

3 subtribes, 2 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 99 (122 total)
number of subtaxa (rank: subtribe) = 5
analyzed in 366 cladograms in 208 publications
maximum number of terminals in an analysis = 80 (mean = 6.91, median = 4)
maximum number of subtaxa (subtribe) = 3 (mean = 1.81, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 312
maximum number of included terminals = 80, (mean = 7.94 median = 5)
maximum number of subtaxa (subtribe) = 3 (mean = 1.95, median = 2)
proportion of cladograms that support basic monophyly = 0.87

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 148
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (subtribe) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 80 (mean = 9.2, median = 7)
maximum number of subtaxa (subtribe) = 3 (mean = 2.61, median = 3)
proportion of cladograms that support rigorous monophyly = 0.78

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Anodontini cladogram distances
PCoA Ordination

NJ Tree of Anodontini cladogram distances
NJ Tree

Cladograms

Aunins et al., 2018 [Publ 3863], fig. 1

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
1.000 11 11 9 3 29 15 4 **
Cladomics cladogram LEPIDODESMINI UNIONINI ACUTICOSTINI LANCEOLARIINI CRISTARIINA ANODONTINA ALASMIDONTINA

Wu et al., 2018 [Publ 3836], fig. 2

cst iT tT bi riS oT roT roS iD
0.931 13 13 9 3 40 20 4 12.0
Cladomics cladogram LEPIDODESMINI UNIONINI ACUTICOSTINI LANCEOLARIINI CRISTARIINA Anodonta cygnea Anodonta anatina ALASMIDONTINA

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
0.912 20 20 13 3 181 28 4 15.0
Cladomics cladogram ACUTICOSTINI LEPIDODESMINI UNIONINI LANCEOLARIINI CRISTARIINA Alasmidonta Lasmigona Pseudanodonta Simpsonaias Strophitus Pyganodon Utterbackia Anodonta anatina Anodonta cygnea Brachyanodon

Huang et al., 2019 [Publ 3843], fig. 3

cst iT tT bi riS oT roT roS iD
0.926 10 10 8 3 46 16 4 5.0
Cladomics cladogram LEPIDODESMINI UNIONINI LANCEOLARIINI ACUTICOSTINI CRISTARIINA Anodonta cygnea Anodonta anatina ALASMIDONTINA

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
0.912 20 20 17 3 179 27 4 15.0
Cladomics cladogram ACUTICOSTINI UNIONINI LEPIDODESMINI LANCEOLARIINI CRISTARIINA Alasmidonta Lasmigona Anodonta anatina Anodonta cygnea Brachyanodon Pyganodon Utterbackia Pseudanodonta Simpsonaias Strophitus

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.765 20 48 25 3 181 28 4 46.0
Cladomics cladogram Chamberlainia Megalonaias Uniomerus QUADRULINA Sinohyriopsis LAMPROTULINI GONIDEINI PSEUDODONTINI Rectidentini Amblema Toxolasma PLEUROBEMINI Obliquaria Plectomerus Ellipsaria Potamilus leptodon Medionidus Potamilus alatus Ptychobranchus Truncilla Venustaconcha Cambarunio Leaunio Ortmanniana Lampsilis ACUTICOSTINI Alasmidonta Anodonta anatina Anemina Anodonta cygnea Cristaria Brachyanodon Lasmigona LEPIDODESMINI Pseudanodonta Pyganodon Simpsonaias Strophitus Utterbackia Schistodesmus UNIONINA ACULAMPROTULINA LANCEOLARIINI Pletholophus Sinanodonta Cuneopsis celtiformis Tchangsinaia Cuneopsis rufescens Nodularia Cuneopsis szechenyii Cuneopsis heudei

Wu et al., 2019 [Publ 3864], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 9 9 7 3 37 17 4 4.0
Cladomics cladogram LEPIDODESMINI UNIONINI ACUTICOSTINI LANCEOLARIINI CRISTARIINA ANODONTINA ALASMIDONTINA

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
0.949 33 33 25 3 89 38 4 30.0
Cladomics cladogram ACUTICOSTINI LEPIDODESMINI UNIONINI LANCEOLARIINI CRISTARIINA Pseudanodonta ALASMIDONTINA Anodonta cygnea Anodonta anatina Brachyanodon

Fukata & Iigo, 2020 [Publ 4106], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 3 44 18 4 1.0
Cladomics cladogram LEPIDODESMINI LANCEOLARIINI UNIONINI ACUTICOSTINI CRISTARIINA ANODONTINA ALASMIDONTINA

Wu et al., 2020 [Publ 4121], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 9 9 7 3 39 17 4 4.0
Cladomics cladogram LEPIDODESMINI UNIONINI ACUTICOSTINI LANCEOLARIINI CRISTARIINA ANODONTINA ALASMIDONTINA

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
0.931 10 10 8 3 81 16 4 9.0
Cladomics cladogram ACUTICOSTINI LEPIDODESMINI UNIONINI LANCEOLARIINI CRISTARIINA Lasmigona Anodonta Pseudanodonta Pyganodon Strophitus

Wu et al., 2021 [Publ 4366], fig. 3-BI

cst iT tT bi riS oT roT roS iD
0.935 13 13 11 3 60 20 4 4.0
Cladomics cladogram LEPIDODESMINI UNIONINI ACUTICOSTINI LANCEOLARIINI CRISTARIINA Anodonta cygnea Anodonta anatina ALASMIDONTINA

Wu et al., 2021 [Publ 4366], fig. 3-ML

cst iT tT bi riS oT roT roS iD
0.935 13 13 11 3 60 20 4 4.0
Cladomics cladogram LEPIDODESMINI UNIONINI ACUTICOSTINI LANCEOLARIINI CRISTARIINA Anodonta cygnea Anodonta anatina ALASMIDONTINA

Wu et al., 2021 [Publ 4366], fig. 4

cst iT tT bi riS oT roT roS iD
0.935 12 12 10 3 56 20 4 3.0
Cladomics cladogram LEPIDODESMINI UNIONINI ACUTICOSTINI LANCEOLARIINI CRISTARIINA Anodonta cygnea Anodonta anatina ALASMIDONTINA

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
0.897 10 10 8 3 81 16 4 11.0
Cladomics cladogram ACUTICOSTINI UNIONINI LANCEOLARIINI LEPIDODESMINI CRISTARIINA Anodonta cygnea Lasmigona Anodonta anatina Pseudanodonta Pyganodon Strophitus

Wu et al., 2022 [Publ 4238], fig. 2d

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 3 28 28 4 3.0
Cladomics cladogram LEPIDODESMINI UNIONINI LANCEOLARIINI ACUTICOSTINI CRISTARIINA ANODONTINA ALASMIDONTINA

Wu et al., 2022 [Publ 4238], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 3 63 25 4 3.0
Cladomics cladogram LEPIDODESMINI UNIONINI ACUTICOSTINI LANCEOLARIINI CRISTARIINA ANODONTINA ALASMIDONTINA

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 3 32 11 4 6.0
Cladomics cladogram LEPIDODESMINI UNIONINI LANCEOLARIINI ACUTICOSTINI CRISTARIINA ANODONTINA ALASMIDONTINA

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

cst iT tT bi riS oT roT roS iD
0.960 6 6 4 3 33 11 4 5.0
Cladomics cladogram CRISTARIINA ANODONTINA ALASMIDONTINA ACUTICOSTINI LANCEOLARIINI LEPIDODESMINI ACULAMPROTULINA UNIONINA OXYNAIINA

Dai et al., 2023 [Publ 4285], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 3 46 10 4 6.0
Cladomics cladogram LEPIDODESMINI UNIONINI ACUTICOSTINI LANCEOLARIINI CRISTARIINA ANODONTINA ALASMIDONTINA

Chen et al., 2023 [Publ 4302], fig. 1

cst iT tT bi riS oT roT roS iD
0.861 10 32 24 3 31 22 4 39.0
Cladomics cladogram COELATURINI INDOCHINELLINI PARREYSIINI Anemina Amuranodonta Pletholophus Sinanodonta Beringiana Lanceolaria gladiola ACUTICOSTINI LEPIDODESMINI Lanceolaria oxyrhyncha Cristaria UNIONINI ALASMIDONTINA ANODONTINA

Chen et al., 2023 [Publ 4302], fig. 2

cst iT tT bi riS oT roT roS iD
0.972 10 10 8 3 31 22 4 13.0
Cladomics cladogram ACUTICOSTINI LEPIDODESMINI UNIONINI LANCEOLARIINI CRISTARIINA ALASMIDONTINA ANODONTINA

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cst iT tT bi riS oT roT roS iD
1.000 18 18 16 3 77 28 4 9.0
Cladomics cladogram LEPIDODESMINI UNIONINI ACUTICOSTINI LANCEOLARIINI CRISTARIINA ANODONTINA ALASMIDONTINA

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cst iT tT bi riS oT roT roS iD
0.914 18 18 14 3 77 28 4 8.0
Cladomics cladogram LEPIDODESMINI UNIONINI ACUTICOSTINI LANCEOLARIINI CRISTARIINA Brachyanodon Pseudanodonta Anodonta cygnea ALASMIDONTINA Anodonta exulcerata Anodonta anatina

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cst iT tT bi riS oT roT roS iD
0.971 18 18 16 3 73 28 4 7.0
Cladomics cladogram LEPIDODESMINI UNIONINI ACUTICOSTINI LANCEOLARIINI CRISTARIINA Pseudanodonta Anodonta Brachyanodon ALASMIDONTINA

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cst iT tT bi riS oT roT roS iD
0.971 18 18 16 3 73 28 4 7.0
Cladomics cladogram LEPIDODESMINI UNIONINI ACUTICOSTINI LANCEOLARIINI CRISTARIINA Brachyanodon ALASMIDONTINA Pseudanodonta Anodonta

Dai et al., 2024 [Publ 4358], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 3 46 21 4 6.0
Cladomics cladogram LEPIDODESMINI UNIONINI LANCEOLARIINI ACUTICOSTINI CRISTARIINA ANODONTINA ALASMIDONTINA

Dai et al., 2025 [Publ 4440], fig. 1

cst iT tT bi riS oT roT roS iD
0.895 10 38 31 3 34 25 4 44.0
Cladomics cladogram LAMPSILINI QUADRULINI Anemina Amuranodonta PARREYSIINI COELATURINI INDOCHINELLINI Pletholophus Sinanodonta Beringiana Cristaria ALASMIDONTINA ANODONTINA LANCEOLARIINI LEPIDODESMINI ACUTICOSTINI UNIONINI

Dai et al., 2025 [Publ 4440], fig. 2

cst iT tT bi riS oT roT roS iD
0.868 10 35 29 3 34 25 4 41.0
Cladomics cladogram COELATURINI INDOCHINELLINI PARREYSIINI Anemina Amuranodonta Pletholophus Sinanodonta Beringiana ALASMIDONTINA ANODONTINA Lanceolaria gladiola Lanceolaria oxyrhyncha Cristaria ACUTICOSTINI LEPIDODESMINI UNIONINI

Dai et al., 2025 [Publ 4440], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 3 77 32 4 3.0
Cladomics cladogram LEPIDODESMINI UNIONINI ACUTICOSTINI LANCEOLARIINI CRISTARIINA ANODONTINA ALASMIDONTINA

Chen et al., 2025 [Publ 4448], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 11 11 9 3 74 32 4 4.0
Cladomics cladogram LEPIDODESMINI UNIONINI LANCEOLARIINI ACUTICOSTINI CRISTARIINA ANODONTINA ALASMIDONTINA

Chen et al., 2025 [Publ 4487], fig. 2

cst iT tT bi riS oT roT roS iD
0.973 10 10 8 3 31 22 4 13.0
Cladomics cladogram GLOBUNIONINA ACULAMPROTULINA UNIONINA OXYNAIINA ACUTICOSTINI LEPIDODESMINI LANCEOLARIINI CRISTARIINA ALASMIDONTINA ANODONTINA

Chen et al., 2025 [Publ 4487], fig. 3

cst iT tT bi riS oT roT roS iD
0.946 10 13 11 3 31 22 4 16.0
Cladomics cladogram UNIONINI ACUTICOSTINI ALASMIDONTINA ANODONTINA LEPIDODESMINI Cristaria LANCEOLARIINI Anemina Amuranodonta Pletholophus Sinanodonta Beringiana

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cst iT tT bi riS oT roT roS iD
0.947 17 17 15 3 123 51 4 8.0
Cladomics cladogram LEPIDODESMINI UNIONINI LANCEOLARIINI ACUTICOSTINI CRISTARIINA Anodonta cygnea Anodonta exulcerata Anodonta anatina ALASMIDONTINA Brachyanodon Pseudanodonta

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.818 80 156 164 3 553 76 4 161.0
Cladomics cladogram Lamellidens Balwantia Trapezidens LEOPARREYSIINI Pseudomulleria INDOCHINELLINI PARREYSIINI COELATURINI Amuranodonta Anemina Buldowskia Beringiana Pletholophus Mimo Acudonta Sinanodonta Cristaria LEPIDODESMINI LANCEOLARIINI Pseudoacuticosta retiaria 2 Acuticosta chinensis 2 Acuticosta chinensis 1 Pseudoacuticosta retiaria 1 Globunio ACULAMPROTULINA MIDDENDORFFINAIINA UNIONINA OXYNAIINA Pseudodontoideus Lasmigona Pterosyna Arcidens ANODONTINA Utterbackia Pyganodon Utterbackiana Simpsonaias Alasminota Alasmidonta Prolasmidonta Platynaias Anodontoides Strophitus

Wu et al., 2026 [Publ 4811], fig. 4

cst iT tT bi riS oT roT roS iD
0.977 26 26 24 3 69 39 4 15.0
Cladomics cladogram LEPIDODESMINI UNIONINI ACUTICOSTINI LANCEOLARIINI CRISTARIINA Anodonta Pseudanodonta ALASMIDONTINA

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 3 13 13 3 3.0
Cladomics cladogram CRISTARIINA ANODONTINA ALASMIDONTINA LANCEOLARIINI LEPIDODESMINI UNIONINI

Wang et al., 2018 [Publ 3865], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 3 24 10 3 6.0
Cladomics cladogram UNIONINI ACUTICOSTINI LANCEOLARIINI CRISTARIINA ANODONTINA ALASMIDONTINA

Cha et al., 2018 [Publ 3916], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 9 9 7 3 29 12 3 4.0
Cladomics cladogram LEPIDODESMINI UNIONINI LANCEOLARIINI CRISTARIINA ANODONTINA ALASMIDONTINA

Froufe et al., 2019 [Publ 3936], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 9 9 7 3 35 13 3 4.0
Cladomics cladogram LEPIDODESMINI UNIONINI LANCEOLARIINI CRISTARIINA ANODONTINA ALASMIDONTINA

Froufe et al., 2019 [Publ 3936], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 9 9 7 3 35 13 3 4.0
Cladomics cladogram LEPIDODESMINI UNIONINI LANCEOLARIINI CRISTARIINA ANODONTINA ALASMIDONTINA

Sano et al., 2020 [Publ 4138], fig. 2

cst iT tT bi riS oT roT roS iD
0.935 19 36 36 3 50 17 3 31.0
Cladomics cladogram LAMPSILINI CHAMBERLAINIINI Inversidens LAMPROTULINI Ptychorhynchus Obovalis Anemina Buldowskia Pletholophus Beringiana Sinanodonta Cristaria ANODONTINA ALASMIDONTINA LANCEOLARIINI LEPIDODESMINI UNIONINI

Sano et al., 2020 [Publ 4138], fig. 3

cst iT tT bi riS oT roT roS iD
0.935 19 36 37 3 50 17 3 32.0
Cladomics cladogram CHAMBERLAINIINI LAMPSILINI LAMPROTULINI GONIDEINI Anemina Buldowskia Sinanodonta Pletholophus Beringiana Cristaria ANODONTINA ALASMIDONTINA LANCEOLARIINI LEPIDODESMINI UNIONINI

Sano et al., 2020 [Publ 4138], fig. 4

cst iT tT bi riS oT roT roS iD
0.935 19 36 37 3 50 17 3 32.0
Cladomics cladogram LAMPSILINI CHAMBERLAINIINI Inversidens LAMPROTULINI Obovalis Ptychorhynchus Anemina Buldowskia Sinanodonta Pletholophus Beringiana Cristaria ANODONTINA ALASMIDONTINA LEPIDODESMINI LANCEOLARIINI UNIONINI

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

cst iT tT bi riS oT roT roS iD
1.000 9 9 7 3 39 13 3 4.0
Cladomics cladogram LEPIDODESMINI UNIONINI LANCEOLARIINI CRISTARIINA ANODONTINA ALASMIDONTINA

Wu et al., 2024 [Publ 4379], fig. 5

cst iT tT bi riS oT roT roS iD
0.970 18 18 16 3 35 33 3 7.0
Cladomics cladogram LEPIDODESMINI UNIONINI LANCEOLARIINI CRISTARIINA Brachyanodon ALASMIDONTINA Pseudanodonta Anodonta

Wu et al., 2024 [Publ 4388], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 3 46 11 3 6.0
Cladomics cladogram LEPIDODESMINI UNIONINI LANCEOLARIINI CRISTARIINA ANODONTINA ALASMIDONTINA

Hou et al., 2024 [Publ 4400], fig. 4

cst iT tT bi riS oT roT roS iD
0.971 18 18 16 3 57 53 3 7.0
Cladomics cladogram LEPIDODESMINI UNIONINI LANCEOLARIINI CRISTARIINA Pseudanodonta Anodonta Brachyanodon ALASMIDONTINA

Wang et al., 2025 [Publ 4488], fig. 3B

cst iT tT bi riS oT roT roS iD
0.974 26 26 24 3 38 36 3 15.0
Cladomics cladogram LEPIDODESMINI UNIONINI ACUTICOSTINI CRISTARIINA Anodonta Pseudanodonta ALASMIDONTINA

Wu et al., 2026 [Publ 4825], fig. 6B

cst iT tT bi riS oT roT roS iD
0.974 25 25 24 3 40 38 3 17.0
Cladomics cladogram LEPIDODESMINI UNIONINI LANCEOLARIINI CRISTARIINA Anodonta Pseudanodonta ALASMIDONTINA

Plazzi et al., 2011 [Publ 3676], fig. 5

cst iT tT bi riS oT roT roS iD
0.786 5 18 11 3 13 4 2 21.0
Cladomics cladogram CRISTARIINA LANCEOLARIINI Anodonta UNIONINI ALASMIDONTINA Pseudanodonta Alathyria LAMPROTULINI CHAMBERLAINIINI QUADRULINI PLEUROBEMINI LAMPSILINI

Fonseca et al., 2016 [Publ 3775], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 9 9 7 3 22 9 2 4.0
Cladomics cladogram LANCEOLARIINI UNIONINI CRISTARIINA ANODONTINA ALASMIDONTINA

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
0.966 14 14 11 3 67 15 2 12.0
Cladomics cladogram UNIONINI LANCEOLARIINI CRISTARIINA ANODONTINA ALASMIDONTINA

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
0.929 15 15 13 3 124 13 2 17.0
Cladomics cladogram UNIONINI LANCEOLARIINI CRISTARIINA Anodonta anatina ALASMIDONTINA Pseudanodonta Anodonta cygnea Brachyanodon

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
0.821 15 18 11 3 124 13 2 19.0
Cladomics cladogram UNIONINI LANCEOLARIINI CRISTARIINA Anodonta anatina Pseudanodonta ALASMIDONTINA Anodonta cygnea Brachyanodon

Bolotov et al., 2017 [Publ 3764], fig. S1

cst iT tT bi riS oT roT roS iD
0.500 15 18 0 3 110 13 2 22.0
Cladomics cladogram UNIONINI Alasmidonta Anodonta anatina Anodonta cygnea Brachyanodon Lasmigona Pseudanodonta Pyganodon Simpsonaias Strophitus Sinanodonta lauta Cristaria plicata Cristaria radiata Pletholophus Sinanodonta lucida Sinanodonta jourdyi Lanceolaria gladiola Lanceolaria grayii Lanceolaria danzhouensis

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 9 9 7 3 23 9 2 4.0
Cladomics cladogram LANCEOLARIINI UNIONINI CRISTARIINA ANODONTINA ALASMIDONTINA

Bolotov et al., 2018 [Publ 3826], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 3 72 7 2 13.0
Cladomics cladogram UNIONINI LANCEOLARIINI CRISTARIINA ANODONTINA ALASMIDONTINA

Wang et al., 2018 [Publ 3846], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 3 24 10 2 6.0
Cladomics cladogram LANCEOLARIINI UNIONINI CRISTARIINA ANODONTINA ALASMIDONTINA

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

cst iT tT bi riS oT roT roS iD
1.000 4 4 1 3 16 6 2 8.0
Cladomics cladogram LANCEOLARIINI UNIONINI CRISTARIINA ANODONTINA ALASMIDONTINA

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 6 6 2 7.0
Cladomics cladogram LANCEOLARIINI UNIONINI CRISTARIINA ANODONTINA ALASMIDONTINA

Huang et al., 2019 [Publ 3843], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 19 6 2 7.0
Cladomics cladogram LANCEOLARIINI UNIONINI CRISTARIINA ANODONTINA ALASMIDONTINA

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 53 4 2 7.0
Cladomics cladogram LANCEOLARIINI UNIONINI CRISTARIINA ANODONTINA ALASMIDONTINA

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 53 4 2 7.0
Cladomics cladogram LANCEOLARIINI UNIONINI CRISTARIINA ANODONTINA ALASMIDONTINA

Guerra et al., 2019 [Publ 4168], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 3 34 6 2 8.0
Cladomics cladogram LANCEOLARIINI UNIONINI CRISTARIINA ANODONTINA ALASMIDONTINA

Guerra et al., 2019 [Publ 4168], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 18 6 2 7.0
Cladomics cladogram LANCEOLARIINI UNIONINI CRISTARIINA ANODONTINA ALASMIDONTINA

Guerra et al., 2019 [Publ 4168], fig. 2

cst iT tT bi riS oT roT roS iD
0.947 7 7 5 3 34 6 2 8.0
Cladomics cladogram ACULAMPROTULINA OXYNAIINA UNIONINA LANCEOLARIINI CRISTARIINA ANODONTINA ALASMIDONTINA

Guerra et al., 2019 [Publ 4168], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 18 6 2 9.0
Cladomics cladogram LANCEOLARIINI UNIONINI ALASMIDONTINA ANODONTINA CRISTARIINA

Riccardi et al., 2020 [Publ 4089], fig. 2

cst iT tT bi riS oT roT roS iD
0.895 13 13 11 3 6 4 2 8.0
Cladomics cladogram LANCEOLARIINI UNIONINI CRISTARIINA Anodonta anatina Brachyanodon ALASMIDONTINA Pseudanodonta Anodonta exulcerata Anodonta cygnea

Chen et al., 2020 [Publ 4170], fig. 1

cst iT tT bi riS oT roT roS iD
0.714 11 34 32 3 23 4 2 39.0
Cladomics cladogram Prolasmidonta Alasmidonta Anemina PSEUDODONTINI Lamprotula caveata Chamberlainia OXYNAIINA LEPIDODESMINI ACULAMPROTULINA Platynaias Utterbackia imbecillis Pyganodon Sinanodonta Cristaria ANODONTINA Utterbackia peninsularis Potamilus alatus Lamprotula leaii Sinohyriopsis GONIDEINI Gibbosula QUADRULINI Toxolasma Lampsilis ornata Potamilus leptodon Lampsilis siliquoidea Lampsilis powellii

Wu et al., 2022 [Publ 4238], fig. 2e

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 14 14 2 7.0
Cladomics cladogram LANCEOLARIINI UNIONINI CRISTARIINA ANODONTINA ALASMIDONTINA

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

cst iT tT bi riS oT roT roS iD
0.964 14 14 12 3 62 12 2 14.0
Cladomics cladogram UNIONINI LANCEOLARIINI CRISTARIINA ANODONTINA ALASMIDONTINA

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

cst iT tT bi riS oT roT roS iD
0.929 14 14 12 3 62 12 2 14.0
Cladomics cladogram UNIONINI LANCEOLARIINI CRISTARIINA ANODONTINA ALASMIDONTINA

Klymus et al., 2025 [Publ 4814], fig. 3A

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 32 11 2 7.0
Cladomics cladogram LANCEOLARIINI UNIONINI CRISTARIINA ANODONTINA ALASMIDONTINA

Klymus et al., 2025 [Publ 4814], fig. 3B

cst iT tT bi riS oT roT roS iD
0.955 4 4 2 3 32 11 2 7.0
Cladomics cladogram CRISTARIINA ANODONTINA ALASMIDONTINA OXYNAIINA UNIONINA ACULAMPROTULINA LANCEOLARIINI

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 43 15 2 7.0
Cladomics cladogram LANCEOLARIINI UNIONINI CRISTARIINA ANODONTINA ALASMIDONTINA

Chong et al., 2008 [Publ 2668], fig. 3

cst iT tT bi riS oT roT roS iD
0.762 17 18 10 3 1 1 1 26.0
Cladomics cladogram Anodonta cygnea Pyganodon Alasminota Anodonta anatina UNIONINI CRISTARIINA Brachyanodon nuttalliana Brachyanodon californiensis Pseudodontoideus Alasmidonta Pterosyna Pseudanodonta Strophitus Prolasmidonta Platynaias Brachyanodon oregonensis 4 Brachyanodon oregonensis 3 Brachyanodon oregonensis 2 Brachyanodon oregonensis 1 Brachyanodon kennerlyi

Wu et al., 2016 [Publ 3711], fig. 1

cst iT tT bi riS oT roT roS iD
0.944 7 7 5 3 14 5 1 8.0
Cladomics cladogram UNIONINI Sinanodonta Anemina Cristaria ANODONTINA ALASMIDONTINA

Feng et al., 2016 [Publ 3800], fig. 2

cst iT tT bi riS oT roT roS iD
0.769 6 12 10 3 8 1 1 15.0
Cladomics cladogram Margaritifera ANODONTINA Utterbackia UNIONINI Pyganodon CRISTARIINA GONIDEINI QUADRULINI LAMPSILINI

Guerra et al., 2017 [Publ 3772], fig. 2a

cst iT tT bi riS oT roT roS iD
0.938 8 8 6 3 17 2 1 9.0
Cladomics cladogram UNIONINI Sinanodonta Cristaria ANODONTINA ALASMIDONTINA

Guerra et al., 2017 [Publ 3772], fig. 2b

cst iT tT bi riS oT roT roS iD
0.938 8 8 6 3 17 2 1 7.0
Cladomics cladogram UNIONINA ACULAMPROTULINA CRISTARIINA ANODONTINA ALASMIDONTINA

Huang et al., 2018 [Publ 3828], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 9 9 7 3 33 9 1 4.0
Cladomics cladogram UNIONINI CRISTARIINA ANODONTINA ALASMIDONTINA

Huang et al., 2018 [Publ 3828], fig. 6-M

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 13 4 1 7.0
Cladomics cladogram UNIONINI CRISTARIINA ANODONTINA ALASMIDONTINA

Chase et al., 2018 [Publ 3847], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 14 14 13 3 25 2 1 13.0
Cladomics cladogram UNIONINI CRISTARIINA ANODONTINA ALASMIDONTINA

Wu et al., 2018 [Publ 4143], fig. 2

cst iT tT bi riS oT roT roS iD
0.944 7 7 5 3 15 6 1 8.0
Cladomics cladogram UNIONINI Sinanodonta Anemina Cristaria ANODONTINA ALASMIDONTINA

Froufe et al., 2019 [Publ 3936], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 15 3 1 7.0
Cladomics cladogram UNIONINI CRISTARIINA ANODONTINA ALASMIDONTINA

Froufe et al., 2019 [Publ 3936], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 15 3 1 7.0
Cladomics cladogram UNIONINI CRISTARIINA ANODONTINA ALASMIDONTINA

Wu et al., 2020 [Publ 4147], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 3 23 11 1 4.0
Cladomics cladogram UNIONINI CRISTARIINA ANODONTINA ALASMIDONTINA

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 19 3 1 7.0
Cladomics cladogram UNIONINI CRISTARIINA ANODONTINA ALASMIDONTINA

Wu et al., 2023 [Publ 4339], fig. 5

cst iT tT bi riS oT roT roS iD
0.923 9 9 7 3 10 2 1 11.0
Cladomics cladogram LEPIDODESMINI CRISTARIINA ALASMIDONTINA ANODONTINA

Hou et al., 2025 [Publ 4445], fig. 5

cst iT tT bi riS oT roT roS iD
0.966 18 18 16 3 34 32 1 7.0
Cladomics cladogram UNIONINI CRISTARIINA Brachyanodon ALASMIDONTINA Pseudanodonta Anodonta

Soroka, 2008 [Publ 3695], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 1 0 0 8.0
Cladomics cladogram PLEUROBEMINI CRISTARIINA ANODONTINA ALASMIDONTINA

Soroka, 2008 [Publ 3695], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 1 0 0 8.0
Cladomics cladogram PLEUROBEMINI CRISTARIINA ANODONTINA ALASMIDONTINA

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

cst iT tT bi riS oT roT roS iD
0.529 20 20 0 3 85 0 0 26.0
Cladomics cladogram GONIDEINI Reginaia Amblema QUADRULINI PLEUROBEMINI POPENAIADINI Dromus Epioblasma Glebula Lampsilis ovata Lampsilis virescens Lemiox Obliquaria Obovaria olivaria Obovaria unicolor Plectomerus Potamilus alatus Potamilus capax Alasmidonta marginata Prolasmidonta Alasmidonta undulata CRISTARIINA Brachyanodon kennerlyi Brachyanodon nuttalliana Brachyanodon oregonensis Arcidens confragosus Arcidens wheeleri Pterosyna Platynaias compressa Lasmigona Platynaias decorata Platynaias subviridis Pyganodon grandis Pyganodon lacustris Pseudodontoideus Strophitus Utterbackia imbecillis Utterbackia peninsularis 1 Utterbackia peninsularis 2

Burzynski & Soroka, 2018 [Publ 3862], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 0 0 0 7.0
Cladomics cladogram CRISTARIINA ANODONTINA ALASMIDONTINA

Whelan et al., 2023 [Publ 4287], fig. 2

cst iT tT bi riS oT roT roS iD
0.917 41 41 42 3 0 0 0 37.0
Cladomics cladogram CRISTARIINA Anodonta cygnea Brachyanodon Pseudanodonta Anodonta anatina ALASMIDONTINA

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

cst iT tT bi riS oT roT roS iD
0.700 13 13 11 3 15 0 0 19.0
Cladomics cladogram Mutela Chambardia CRISTARIINA Anodonta exulcerata Anodonta cygnea Brachyanodon californiensis Brachyanodon nuttalliana Brachyanodon kennerlyi Brachyanodon oregonensis Anodonta anatina ALASMIDONTINA

Huang et al., 2002 [Publ 2657], fig. 1

cst iT tT bi riS oT roT roS iD
0.909 7 7 5 2 27 7 4 11.0
Cladomics cladogram CRISTARIINA ALASMIDONTINA ACUTICOSTINI Nodularia Tchangsinaia LANCEOLARIINI Schistodesmus LEPIDODESMINI

Huang et al., 2002 [Publ 2657], fig. 2

cst iT tT bi riS oT roT roS iD
0.818 7 14 5 2 27 7 4 18.0
Cladomics cladogram Strophitus Schistodesmus LEPIDODESMINI ACUTICOSTINI CRISTARIINA LANCEOLARIINI Nodularia Tchangsinaia Utterbackia Utterbackiana Alasmidonta Lasmigona Ptychorhynchus CHAMBERLAINIINI LAMPROTULINI Sinosolenaia Megalonaias Elliptoideus Uniomerus Pleurobema Potamilus fragilis Medionidus Amblema Obliquaria Plectomerus Villosa Ellipsaria Lampsilis Potamilus purpuratus Ptychobranchus

Konopleva et al., 2023 [Publ 4269], fig. S1_4

cst iT tT bi riS oT roT roS iD
0.966 16 16 14 2 40 17 4 19.0
Cladomics cladogram UNIONINI LEPIDODESMINI LANCEOLARIINI ACUTICOSTINI ANODONTINA CRISTARIINA

Dai et al., 2024 [Publ 4386], fig. 5-BI

cst iT tT bi riS oT roT roS iD
0.917 2 17 0 2 20 15 4 24.0
Cladomics cladogram LAMELLIDENTINI INDOCHINELLINI ACUTICOSTINI CRISTARIINA ANODONTINA LANCEOLARIINI ACULAMPROTULINA LEPIDODESMINI UNIONINA MIDDENDORFFINAIINA OXYNAIINA

Dai et al., 2024 [Publ 4386], fig. 5-ML

cst iT tT bi riS oT roT roS iD
0.958 2 17 0 2 20 15 4 24.0
Cladomics cladogram LAMELLIDENTINI INDOCHINELLINI ANODONTINA ACUTICOSTINI CRISTARIINA LEPIDODESMINI LANCEOLARIINI UNIONINI

Dai et al., 2024 [Publ 4386], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 38 24 4 8.0
Cladomics cladogram LEPIDODESMINI UNIONINI LANCEOLARIINI ACUTICOSTINI CRISTARIINA ALASMIDONTINA

Wu et al., 2023 [Publ 4296], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 2 19 17 3 12.0
Cladomics cladogram ACUTICOSTINI LANCEOLARIINI UNIONINI ANODONTINA ALASMIDONTINA

Liu et al., 2023 [Publ 4310], fig. 4

cst iT tT bi riS oT roT roS iD
0.917 4 4 2 2 20 18 3 12.0
Cladomics cladogram ACUTICOSTINI Lanceolaria lanceolata Lanceolaria grayii Lanceolaria gladiola UNIONINI ANODONTINA ALASMIDONTINA

Liu et al., 2023 [Publ 4310], fig. 5

cst iT tT bi riS oT roT roS iD
0.875 4 4 2 2 20 18 3 12.0
Cladomics cladogram ACUTICOSTINI Lanceolaria grayii Lanceolaria gladiola Lanceolaria lanceolata UNIONINI Lasmigona ANODONTINA Pyganodon Strophitus

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 2 26 6 2 10.0
Cladomics cladogram CRISTARIINA ALASMIDONTINA ACUTICOSTINI UNIONINI

Xue et al., 2016 [Publ 3659], fig. 1

cst iT tT bi riS oT roT roS iD
0.889 5 5 3 2 10 6 2 9.0
Cladomics cladogram LANCEOLARIINI UNIONINI Cristaria ALASMIDONTINA Anemina Sinanodonta

Zhou et al., 2016 [Publ 3710], fig. 1

cst iT tT bi riS oT roT roS iD
0.938 6 6 4 2 14 3 2 9.0
Cladomics cladogram LEPIDODESMINI UNIONINI Sinanodonta Cristaria ALASMIDONTINA

Wen et al., 2017 [Publ 3795], fig. 3

cst iT tT bi riS oT roT roS iD
0.929 3 3 1 2 13 4 2 8.0
Cladomics cladogram ANODONTINA CRISTARIINA ACULAMPROTULINA LEPIDODESMINI UNIONINA

Bolotov et al., 2020 [Publ 4084], fig. 1

cst iT tT bi riS oT roT roS iD
0.952 12 12 10 2 24 7 2 15.0
Cladomics cladogram UNIONINI LANCEOLARIINI ANODONTINA CRISTARIINA

Rosenberg et al., 1997 [Publ 2720], fig. 2A

cst iT tT bi riS oT roT roS iD
0.600 4 18 1 2 18 1 1 23.0
Cladomics cladogram None 1 ANODONTINA GONIDEINI Pyganodon grandis Utterbackia Pyganodon cataracta Lampsilis straminea Lampsilis teres Megalonaias Obliquaria Theliderma Quadrula Uniomerus Amblema Elliptio Fusconaia Plectomerus Pleurobema UNIONINI

Rosenberg et al., 1997 [Publ 2720], fig. 2B

cst iT tT bi riS oT roT roS iD
0.583 4 18 1 2 18 2 1 23.5
Cladomics cladogram Cumberlandia None 1 None 2 Margaritifera ANODONTINA GONIDEINI Pyganodon grandis Utterbackia Pyganodon cataracta Lampsilis straminea Lampsilis teres Megalonaias Obliquaria Theliderma Quadrula Uniomerus Amblema Elliptio Fusconaia Plectomerus ACULAMPROTULINA UNIONINA

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

cst iT tT bi riS oT roT roS iD
0.917 6 6 4 2 24 2 1 12.0
Cladomics cladogram ANODONTINA ALASMIDONTINA QUADRULINI PLEUROBEMINI AMBLEMINI LAMPSILINI

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

cst iT tT bi riS oT roT roS iD
0.917 6 6 4 2 24 2 1 12.0
Cladomics cladogram ANODONTINA ALASMIDONTINA QUADRULINI PLEUROBEMINI AMBLEMINI LAMPSILINI

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

cst iT tT bi riS oT roT roS iD
0.900 3 3 1 2 31 2 1 10.0
Cladomics cladogram ANODONTINA ALASMIDONTINA GONIDEINI UNIONINI Cumberlandia Margaritifera Toxolasma AMBLEMINI Glebula Cyrtonaias QUADRULINI PLEUROBEMINI Potamilus Ortmanniana Ligumia

Hoeh et al., 2001 [Publ 2652], fig. 14f2

cst iT tT bi riS oT roT roS iD
0.800 3 3 1 2 28 2 1 10.0
Cladomics cladogram Pyganodon ANODONTINA Strophitus Mutela Anodontites guanarensis Anodontites trigona Monocondylaea RHIPIDODONTINI Echyridella Lortiella CASTALIINI Velesunio

Hoeh et al., 2001 [Publ 2652], fig. 14f4

cst iT tT bi riS oT roT roS iD
0.700 3 3 1 2 28 2 1 10.0
Cladomics cladogram Unio pictorum Unio tumidus ANODONTINA ALASMIDONTINA

Hoeh et al., 2002 [Publ 2653], fig. 2

cst iT tT bi riS oT roT roS iD
0.700 3 3 1 2 28 2 1 10.0
Cladomics cladogram Unio pictorum Unio tumidus ANODONTINA ALASMIDONTINA

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

cst iT tT bi riS oT roT roS iD
0.900 3 3 1 2 28 2 1 10.0
Cladomics cladogram UNIONINI Pyganodon ANODONTINA Strophitus

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 28 2 1 10.0
Cladomics cladogram UNIONINI ANODONTINA ALASMIDONTINA

Manendo et al., 2008 [Publ 2669], fig. 1

cst iT tT bi riS oT roT roS iD
0.909 5 5 3 2 8 3 1 10.0
Cladomics cladogram UNIONINI Anodonta Pseudanodonta ALASMIDONTINA

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

cst iT tT bi riS oT roT roS iD
0.571 6 8 6 2 19 3 1 14.0
Cladomics cladogram OXYNAIINA Anodonta cygnea Unio pictorum 2 Unio pictorum 1 Unio caffer Anodonta ALASMIDONTINA

Soroka, 2010 [Publ 3404], fig. 2

cst iT tT bi riS oT roT roS iD
0.800 4 7 4 2 3 3 1 12.0
Cladomics cladogram CRISTARIINA UNIONINI ANODONTINA

Soroka, 2010 [Publ 3404], fig. 2-M

cst iT tT bi riS oT roT roS iD
0.889 2 5 0 2 3 3 1 12.0
Cladomics cladogram CRISTARIINA ANODONTINA UNIONINI

Soroka, 2010 [Publ 3404], fig. 3a

cst iT tT bi riS oT roT roS iD
0.800 5 9 7 2 4 4 1 14.0
Cladomics cladogram CRISTARIINA ANODONTINA UNIONINI

Soroka, 2010 [Publ 3404], fig. 3a-M

cst iT tT bi riS oT roT roS iD
0.889 3 6 4 2 3 3 1 13.0
Cladomics cladogram ANODONTINA CRISTARIINA UNIONINI

Soroka, 2010 [Publ 3404], fig. 3b

cst iT tT bi riS oT roT roS iD
0.700 5 9 5 2 4 4 1 15.0
Cladomics cladogram Anodonta anatina Anodonta cygnea Pseudanodonta CRISTARIINA UNIONINI

Soroka, 2010 [Publ 3404], fig. 3b-M

cst iT tT bi riS oT roT roS iD
0.889 3 6 4 2 3 3 1 13.0
Cladomics cladogram ANODONTINA CRISTARIINA UNIONINI

Soroka, 2010 [Publ 3404], fig. 4

cst iT tT bi riS oT roT roS iD
0.700 4 7 5 2 3 3 1 14.0
Cladomics cladogram Anodonta anatina UNIONINI CRISTARIINA Anodonta cygnea Pseudanodonta

Soroka, 2010 [Publ 3404], fig. 5a

cst iT tT bi riS oT roT roS iD
0.800 5 5 3 2 3 3 1 12.0
Cladomics cladogram UNIONINI Anodonta anatina CRISTARIINA Anodonta cygnea Pseudanodonta

Soroka, 2010 [Publ 3404], fig. 5b

cst iT tT bi riS oT roT roS iD
0.700 5 8 5 2 3 3 1 15.0
Cladomics cladogram Anodonta anatina CRISTARIINA UNIONINI Anodonta cygnea Pseudanodonta

Soroka, 2010 [Publ 3404], fig. 8a

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 3 3 1 8.0
Cladomics cladogram UNIONINI CRISTARIINA ANODONTINA

Soroka, 2010 [Publ 3404], fig. 8b

cst iT tT bi riS oT roT roS iD
0.889 3 6 4 2 3 3 1 11.0
Cladomics cladogram CRISTARIINA ANODONTINA UNIONINI

Zieritz et al., 2012 [Publ 3673], fig. 4

cst iT tT bi riS oT roT roS iD
0.700 4 8 4 2 4 3 1 13.0
Cladomics cladogram ANODONTINA CRISTARIINA Margaritifera UNIONINI

Huang et al., 2013 [Publ 3629], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 2 10 1 1 7.0
Cladomics cladogram UNIONINI CRISTARIINA ALASMIDONTINA

Fonseca et al., 2016 [Publ 3775], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 6 1 1 8.0
Cladomics cladogram UNIONINI ANODONTINA ALASMIDONTINA

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 10 3 1 8.0
Cladomics cladogram UNIONINI ANODONTINA ALASMIDONTINA

Burzynski et al., 2017 [Publ 3774], fig. 1-top

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 10 3 1 8.0
Cladomics cladogram UNIONINI ANODONTINA ALASMIDONTINA

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 7 1 1 8.0
Cladomics cladogram UNIONINI ANODONTINA ALASMIDONTINA

Froufe et al., 2017 [Publ 3802], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 2 6 3 1 9.0
Cladomics cladogram UNIONINI CRISTARIINA ANODONTINA

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 4 2 1 8.0
Cladomics cladogram UNIONINI ANODONTINA ALASMIDONTINA

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 4 2 1 8.0
Cladomics cladogram UNIONINI ANODONTINA ALASMIDONTINA

Araujo et al., 2018 [Publ 3779], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 15 14 1 9.0
Cladomics cladogram UNIONINI ANODONTINA ALASMIDONTINA

Chase et al., 2018 [Publ 3847], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 8 2 1 --
Cladomics cladogram UNIONINI ANODONTINA ALASMIDONTINA

Chase et al., 2018 [Publ 3861], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 8 2 1 8.0
Cladomics cladogram UNIONINI ANODONTINA ALASMIDONTINA

Chase et al., 2018 [Publ 3861], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 8 2 1 8.0
Cladomics cladogram UNIONINI ANODONTINA ALASMIDONTINA

Tomilova et al., 2020 [Publ 4097], fig. 3

cst iT tT bi riS oT roT roS iD
0.800 6 8 5 2 2 2 1 13.0
Cladomics cladogram UNIONINI CRISTARIINA ANODONTINA

Pfarrer et al., 2022 [Publ 4305], fig. 2-5

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 2 7 5 1 8.0
Cladomics cladogram UNIONINI CRISTARIINA ANODONTINA

Bogan et al., 2023 [Publ 4295], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 28 28 25 2 3 1 1 29.0
Cladomics cladogram UNIONINI ANODONTINA CRISTARIINA

Xiang et al., 2025 [Publ 4433], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 9 9 7 2 4 3 1 10.0
Cladomics cladogram LANCEOLARIINI ANODONTINA CRISTARIINA

Altaba et al., 2025 [Publ 4849], fig. 10

cst iT tT bi riS oT roT roS iD
0.867 8 8 6 2 4 4 1 11.0
Cladomics cladogram UNIONINI Anodonta anatina Anodonta cygnea CRISTARIINA

Altaba et al., 2025 [Publ 4849], fig. 11

cst iT tT bi riS oT roT roS iD
0.867 10 10 8 2 4 4 1 13.0
Cladomics cladogram UNIONINI CRISTARIINA ANODONTINA

Altaba et al., 2025 [Publ 4849], fig. 12

cst iT tT bi riS oT roT roS iD
0.714 9 9 8 2 4 4 1 15.0
Cladomics cladogram UNIONINI Amuranodonta Cristaria Beringiana Sinanodonta woodiana 2 Sinanodonta woodiana 1 Sinanodonta lauta Sinanodonta tumens ANODONTINA

Hoeh, 1990 [Publ 2717], fig. 2

cst iT tT bi riS oT roT roS iD
0.833 15 15 13 2 0 0 0 20.0
Cladomics cladogram Pterosyna Platynaias Lasmigona Utterbackia Pyganodon Anodonta Brachyanodon Utterbackiana

Hoeh, 1990 [Publ 2717], fig. 3

cst iT tT bi riS oT roT roS iD
0.583 15 15 2 2 0 0 0 22.0
Cladomics cladogram Pyganodon fragilis Utterbackia imbecillis Utterbackia peggyae Anodonta Brachyanodon Utterbackiana couperiana 1 Utterbackiana couperiana 2 Utterbackiana suborbiculata Lasmigona Utterbackiana implicata Pterosyna Platynaias Pyganodon cataracta Pyganodon gibbosa Pyganodon grandis Pyganodon lacustris

Hoeh, 1990 [Publ 2717], fig. 4

cst iT tT bi riS oT roT roS iD
0.833 15 15 13 2 0 0 0 20.0
Cladomics cladogram Pterosyna Platynaias Lasmigona Utterbackia Pyganodon Anodonta Brachyanodon Utterbackiana

Lydeard et al., 1996 [Publ 2656], fig. 4

cst iT tT bi riS oT roT roS iD
0.900 6 6 4 2 22 0 0 14.5
Cladomics cladogram Utterbackia Utterbackiana ANODONTINA Strophitus Lasmigona Alasmidonta Megalonaias QUADRULINA Amblema Plectomerus LAMPSILINI Elliptoideus Uniomerus Eurynia Pleurobema

Nagel et al., 1996 [Publ 3400], fig. 351

cst iT tT bi riS oT roT roS iD
0.714 5 5 4 2 0 0 0 12.0
Cladomics cladogram Pseudanodonta Anodonta cygnea CRISTARIINA Anodonta 2 Anodonta anatina Anodonta 1

Nagel et al., 1998 [Publ 1946], fig. 7

cst iT tT bi riS oT roT roS iD
0.714 5 5 4 2 0 0 0 12.0
Cladomics cladogram Pseudanodonta Anodonta cygnea CRISTARIINA Anodonta 2 Anodonta anatina Anodonta 1

Krebs et al., 2003 [Publ 2691], fig. 3

cst iT tT bi riS oT roT roS iD
0.923 10 10 5 2 23 0 0 18.5
Cladomics cladogram Utterbackiana Platynaias Alasmidonta Pterosyna Lasmigona Strophitus Utterbackia Pyganodon ANODONTINA Megalonaias QUADRULINA Elliptoideus Pleurobema Uniomerus Fusconaia Eurynia Amblema Toxolasma Plectomerus Obliquaria Ptychobranchus Potamilus fragilis Medionidus Villosa Ellipsaria Lampsilis teres Lampsilis siliquoidea Potamilus purpuratus Potamilus alatus

Mock et al., 2004 [Publ 2686], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 0 0 0 --
Cladomics cladogram ANODONTINA CRISTARIINA

Mock et al., 2004 [Publ 2686], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 0 0 0 --
Cladomics cladogram ANODONTINA CRISTARIINA

Campbell et al., 2005 [Publ 2660], fig. 1

cst iT tT bi riS oT roT roS iD
0.800 6 6 4 2 96 0 0 13.0
Cladomics cladogram CHAMBERLAINIINI Brachyanodon Anodonta Alasminota Pyganodon Lasmigona Pseudodontoideus GONIDEINI LAMPROTULINI QUADRULINI PLEUROBEMINI Plectomerus Reginaia LAMPSILINI POPENAIADINI Amblema

Campbell et al., 2005 [Publ 2660], fig. 2

cst iT tT bi riS oT roT roS iD
0.900 6 6 3 2 96 0 0 13.0
Cladomics cladogram CHAMBERLAINIINI Pyganodon Lasmigona Pseudodontoideus Alasminota ANODONTINA

Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 2 0 0 12.0
Cladomics cladogram GONIDEINI CRISTARIINA ANODONTINA

Walker et al., 2006 [Publ 2666], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 12 0 0 10.5
Cladomics cladogram ANODONTINA ALASMIDONTINA LAMPROTULINI GONIDEINI PSEUDODONTINI LAMPSILINI PLEUROBEMINI Cumberlandia

Walker et al., 2006 [Publ 2666], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 12 0 0 10.5
Cladomics cladogram ANODONTINA ALASMIDONTINA PLEUROBEMINI LAMPSILINI

Walker et al., 2006 [Publ 2666], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 12 0 0 10.5
Cladomics cladogram ANODONTINA ALASMIDONTINA PLEUROBEMINI LAMPSILINI

Zanatta et al., 2007 [Publ 2698], fig. 1A

cst iT tT bi riS oT roT roS iD
0.833 16 16 10 2 0 0 0 21.0
Cladomics cladogram Platynaias Lasmigona Pterosyna Anodonta Brachyanodon Utterbackiana Utterbackia Pyganodon

Zanatta et al., 2007 [Publ 2698], fig. 1B

cst iT tT bi riS oT roT roS iD
0.833 16 16 13 2 0 0 0 21.0
Cladomics cladogram Lasmigona Platynaias Pterosyna Utterbackia Pyganodon Anodonta Brachyanodon Utterbackiana

Cyr et al., 2007 [Publ 2745], fig. 2B

cst iT tT bi riS oT roT roS iD
1.000 5 5 4 2 0 0 0 13.0
Cladomics cladogram ANODONTINA ALASMIDONTINA

Cyr et al., 2007 [Publ 2745], fig. 2C

cst iT tT bi riS oT roT roS iD
1.000 5 5 4 2 0 0 0 13.0
Cladomics cladogram ANODONTINA ALASMIDONTINA

Soroka, 2010 [Publ 3404], fig. 4-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 0 0 0 --
Cladomics cladogram ANODONTINA CRISTARIINA

Soroka, 2010 [Publ 3404], fig. 5a-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 0 0 0 --
Cladomics cladogram ANODONTINA CRISTARIINA

Soroka, 2010 [Publ 3404], fig. 5b-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 0 0 0 --
Cladomics cladogram ANODONTINA CRISTARIINA

Breton et al., 2011 [Publ 3376], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 12 12 11 2 26 0 0 15.0
Cladomics cladogram CHAMBERLAINIINI CRISTARIINA ALASMIDONTINA

Chen et al., 2012 [Publ 3624], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 5 0 0 --
Cladomics cladogram ALASMIDONTINA CRISTARIINA CHAMBERLAINIINI LAMPROTULINI

Sheng et al., 2014 [Publ 3704], fig. 2

cst iT tT bi riS oT roT roS iD
0.667 2 8 0 2 6 0 0 15.0
Cladomics cladogram CRISTARIINA ALASMIDONTINA Lampsilis Venustaconcha QUADRULINI LAMPROTULINI CHAMBERLAINIINI

Mezhzherin et al., 2014 [Publ 3705], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 2 0 0 0 9.0
Cladomics cladogram CRISTARIINA ANODONTINA

Mezhzherin et al., 2014 [Publ 3705], fig. 4

cst iT tT bi riS oT roT roS iD
0.857 4 4 2 2 0 0 0 9.0
Cladomics cladogram CRISTARIINA ANODONTINA

Froufe et al., 2015 [Publ 3664], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 9 0 0 8.0
Cladomics cladogram LAMPSILINI QUADRULINI ANODONTINA ALASMIDONTINA

Froufe et al., 2015 [Publ 3664], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 5 0 0 8.0
Cladomics cladogram ANODONTINA ALASMIDONTINA LAMPSILINI QUADRULINI GONIDEINI LAMPROTULINI

Wei et al., 2016 [Publ 3665], fig. 0

cst iT tT bi riS oT roT roS iD
0.667 2 4 0 2 4 0 0 11.0
Cladomics cladogram CHAMBERLAINIINI CRISTARIINA LAMPSILINI ALASMIDONTINA QUADRULINI

Amaro et al., 2017 [Publ 3791], fig. 2-M

cst iT tT bi riS oT roT roS iD
0.875 4 4 1 2 6 0 0 11.0
Cladomics cladogram LAMPROTULINI GONIDEINI Pyganodon Lasmigona CRISTARIINA Utterbackia

Wen et al., 2017 [Publ 3795], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 3 0 0 8.0
Cladomics cladogram ANODONTINA ALASMIDONTINA QUADRULINI LAMPSILINI

Froufe et al., 2017 [Publ 3802], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 2 0 0 0 9.0
Cladomics cladogram CRISTARIINA ANODONTINA

Froufe et al., 2017 [Publ 3802], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 12 12 9 2 0 0 0 13.0
Cladomics cladogram ANODONTINA CRISTARIINA

Smith et al., 2018 [Publ 3790], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 18 18 17 2 0 0 0 24.0
Cladomics cladogram CRISTARIINA ALASMIDONTINA

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 26 0 0 --
Cladomics cladogram ANODONTINA ALASMIDONTINA AMBLEMINI LAMPSILINI PLEUROBEMINI QUADRULINI

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 27 0 0 --
Cladomics cladogram ANODONTINA ALASMIDONTINA Megalonaias Uniomerus QUADRULINA PLEUROBEMINI LAMPSILINI AMBLEMINI

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 24 0 0 --
Cladomics cladogram ANODONTINA ALASMIDONTINA QUADRULINI AMBLEMINI PLEUROBEMINI LAMPSILINI

Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 2 0 0 0 8.0
Cladomics cladogram ANODONTINA CRISTARIINA

Riccardi et al., 2020 [Publ 4089], fig. 1

cst iT tT bi riS oT roT roS iD
0.875 6 6 4 2 0 0 0 9.0
Cladomics cladogram CRISTARIINA ANODONTINA

Smith, 2021 [Publ 4331], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 17 0 0 8.0
Cladomics cladogram INDOCHINELLINI CRISTARIINA ALASMIDONTINA

Whelan et al., 2023 [Publ 4287], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 28 28 27 2 0 0 0 29.0
Cladomics cladogram ANODONTINA ALASMIDONTINA

Whelan et al., 2023 [Publ 4287], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 27 27 25 2 0 0 0 30.0
Cladomics cladogram ANODONTINA ALASMIDONTINA

Perkins et al., 2025 [Publ 4447], fig. 2_mtDNA-ITS

cst iT tT bi riS oT roT roS iD
0.952 36 36 37 2 0 0 0 38.0
Cladomics cladogram Anodonta Pseudanodonta ALASMIDONTINA

Sano et al., 2017 [Publ 3799], fig. 2

cst iT tT bi riS oT roT roS iD
0.900 9 9 7 1 24 12 4 16.0
Cladomics cladogram CRISTARIINA Tchangsinaia Nodularia Schistodesmus ACUTICOSTINI Inversiunio LEPIDODESMINI LANCEOLARIINI

Huang et al., 2019 [Publ 3843], fig. 1

cst iT tT bi riS oT roT roS iD
0.800 8 32 30 1 38 24 4 37.0
Cladomics cladogram CHAMBERLAINIINI Anemina ACUTICOSTINI Sinanodonta ACULAMPROTULINA Cristaria LANCEOLARIINI LEPIDODESMINI OXYNAIINA

Huang et al., 2019 [Publ 3843], fig. S1

cst iT tT bi riS oT roT roS iD
0.900 8 33 36 1 39 25 4 47.0
Cladomics cladogram Anemina Sinanodonta ACUTICOSTINI LEPIDODESMINI Cristaria LANCEOLARIINI UNIONINI CHAMBERLAINIINI Parvasolenaia rivularis Parvasolenaia triangularis Ptychorhynchus Sinosolenaia carinata LAMPROTULINI Sinosolenaia recognita Sinosolenaia oleivora 2 Sinosolenaia oleivora 1 Sinosolenaia iridinea Sinosolenaia oleivora 3

Sano et al., 2020 [Publ 4138], fig. 1

cst iT tT bi riS oT roT roS iD
0.565 10 22 5 1 24 12 4 39.0
Cladomics cladogram GONIDEINI LAMPROTULINI CHAMBERLAINIINI Lanceolaria oxyrhyncha 1 Lanceolaria oxyrhyncha 2 Cristaria plicata 1 Cristaria plicata 2 Sinanodonta lauta Beringiana japonica Beringiana fukuharai 1 Beringiana beringiana Beringiana fukuharai 2 Beringiana fukuharai 3 Buldowskia iwakawai 1 Buldowskia iwakawai 2 Sinanodonta tumens 1 Sinanodonta tumens 2 Pletholophus Anemina Lanceolaria lanceolata Cristaria plicata 3 Lanceolaria gladiola Sinanodonta angula LEPIDODESMINI ACULAMPROTULINA ACUTICOSTINI Cuneopsis Nodularia Schistodesmus Inversiunio

Zhang et al., 2004 [Publ 2735], fig. 1

cst iT tT bi riS oT roT roS iD
0.938 2 2 0 1 12 10 3 --
Cladomics cladogram CRISTARIINA UNIONINI ACUTICOSTINI LANCEOLARIINI

Zhou et al., 2007 [Publ 2658], fig. 1

cst iT tT bi riS oT roT roS iD
0.812 2 2 0 1 19 11 3 --
Cladomics cladogram CRISTARIINA CHAMBERLAINIINI Lamprotula leaii Lamprotula caveata Gibbosula Lamprotula cornuumlunae ACULAMPROTULINA Tchangsinaia Nodularia Schistodesmus LANCEOLARIINI ACUTICOSTINI

Ouyang et al., 2011 [Publ 3408], fig. 1

cst iT tT bi riS oT roT roS iD
0.800 3 3 1 1 10 6 3 11.0
Cladomics cladogram CRISTARIINA OXYNAIINA Lanceolaria grayii ACUTICOSTINI Lanceolaria gladiola ACULAMPROTULINA

Ouyang et al., 2015 [Publ 4493], fig. 1

cst iT tT bi riS oT roT roS iD
0.882 4 4 2 1 27 12 3 11.0
Cladomics cladogram UNIONINI LANCEOLARIINI ACUTICOSTINI CRISTARIINA

Ouyang et al., 2015 [Publ 4493], fig. 2

cst iT tT bi riS oT roT roS iD
0.882 4 4 2 1 27 12 3 11.0
Cladomics cladogram LANCEOLARIINI ACULAMPROTULINA ACUTICOSTINI CRISTARIINA OXYNAIINA

Liu et al., 2017 [Publ 3939], fig. 2

cst iT tT bi riS oT roT roS iD
0.933 1 1 0 1 14 13 3 --
Cladomics cladogram CRISTARIINA GONIDEINI

Wu et al., 2026 [Publ 4811], fig. 3

cst iT tT bi riS oT roT roS iD
0.933 2 2 0 1 15 8 3 --
Cladomics cladogram UNIONINA LANCEOLARIINI GLOBUNIONINA ACUTICOSTINI ANODONTINA

Zhou et al., 2007 [Publ 2658], fig. 2

cst iT tT bi riS oT roT roS iD
0.692 2 18 0 1 16 6 2 25.0
Cladomics cladogram Sinanodonta Cristaria LAMPROTULINI QUADRULINI ACULAMPROTULINA ACUTICOSTINI OXYNAIINA

Zhou et al., 2017 [Publ 3788], fig. 2

cst iT tT bi riS oT roT roS iD
0.917 1 1 0 1 15 5 2 --
Cladomics cladogram CRISTARIINA LANCEOLARIINI UNIONINI QUADRULINI LAMPSILINI CHAMBERLAINIINI LAMPROTULINI GONIDEINI

Sano et al., 2017 [Publ 3799], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 1 13 5 2 16.0
Cladomics cladogram CRISTARIINA LANCEOLARIINI UNIONINI

Kano et al., 2019 [Publ 4194], fig. 6

cst iT tT bi riS oT roT roS iD
0.917 3 3 1 1 7 3 2 13.0
Cladomics cladogram CRISTARIINA LANCEOLARIINI UNIONINI Inversidens Obovalis LAMPROTULINI

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

cst iT tT bi riS oT roT roS iD
0.667 5 5 5 1 7 3 2 32.0
Cladomics cladogram CRISTARIINA Nodularia LANCEOLARIINI Inversiunio

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 20 20 2 --
Cladomics cladogram UNIONINI LANCEOLARIINI ANODONTINA

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 20 20 2 --
Cladomics cladogram UNIONINI LANCEOLARIINI ANODONTINA

Dai et al., 2024 [Publ 4386], fig. 3

cst iT tT bi riS oT roT roS iD
0.941 1 1 0 1 13 12 2 --
Cladomics cladogram UNIONINA MIDDENDORFFINAIINA OXYNAIINA ACULAMPROTULINA ANODONTINA LEPIDODESMINI

Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5_7

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 3 3 1 --
Cladomics cladogram ANODONTINA UNIONINI

Rosenberg et al., 1994 [Publ 2719], fig. 3

cst iT tT bi riS oT roT roS iD
0.500 3 10 0 1 16 1 1 17.0
Cladomics cladogram Elliptio Fusconaia AMBLEMINI Pleurobema UNIONINI Pyganodon cataracta Pyganodon grandis Utterbackia Lampsilis teres Lampsilis straminea Megalonaias Obliquaria Theliderma Quadrula Uniomerus

Nagel et al., 1998 [Publ 1946], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 6 4 1 10.0
Cladomics cladogram ANODONTINA UNIONINI

Nagel et al., 1998 [Publ 1946], fig. 9

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 4 1 1 --
Cladomics cladogram UNIONINI ANODONTINA

Graf, 2000 [Publ 2647], fig. 1

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 16 1 1 --
Cladomics cladogram ALASMIDONTINA LAMPSILINI

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 1 1 --
Cladomics cladogram UNIONINI ALASMIDONTINA

Graf, 2002 [Publ 2651], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 16 2 1 --
Cladomics cladogram UNIONINI ALASMIDONTINA

Huff et al., 2004 [Publ 2697], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 1 1 --
Cladomics cladogram UNIONINI ANODONTINA

Huff et al., 2004 [Publ 2697], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 1 1 --
Cladomics cladogram UNIONINI ANODONTINA

Huff et al., 2004 [Publ 2697], fig. 5-6

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 11 1 1 --
Cladomics cladogram UNIONINI ANODONTINA LAMPSILINI

Källersjö et al., 2005 [Publ 2684], fig. 1A

cst iT tT bi riS oT roT roS iD
0.750 3 3 0 1 4 3 1 10.0
Cladomics cladogram Margaritifera ANODONTINA UNIONINI

Källersjö et al., 2005 [Publ 2684], fig. 1B-F

cst iT tT bi riS oT roT roS iD
0.750 3 3 1 1 4 3 1 10.0
Cladomics cladogram Margaritifera ANODONTINA UNIONINI

Graf & Cummings, 2006 [Publ 1968], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 43 2 1 12.0
Cladomics cladogram UNIONINI ALASMIDONTINA

Taylor et al., 2007 [Publ 2696], fig. 1-3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 1 1 --
Cladomics cladogram ANODONTINA UNIONINI

Araujo et al., 2009 [Publ 2589], fig. 7A

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 10 7 1 --
Cladomics cladogram OXYNAIINA Unio gibbus Unio tumidus ANODONTINA Unio pictorum 1 Unio pictorum 2 Unio mancus Unio tumidiformis Unio crassus

Araujo et al., 2009 [Publ 2589], fig. 7B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 7 1 --
Cladomics cladogram ANODONTINA UNIONINI

Araujo et al., 2009 [Publ 2589], fig. 7C

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 6 1 --
Cladomics cladogram ANODONTINA UNIONINI

Hoeh et al., 2009 [Publ 2716], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 43 2 1 12.0
Cladomics cladogram UNIONINI ALASMIDONTINA

Hoeh et al., 2009 [Publ 2716], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 42 2 1 12.0
Cladomics cladogram UNIONINI ALASMIDONTINA

Hoeh et al., 2009 [Publ 2716], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 42 2 1 12.0
Cladomics cladogram UNIONINI ALASMIDONTINA

Soroka, 2010 [Publ 3404], fig. 8a-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 2 2 1 --
Cladomics cladogram CRISTARIINA UNIONINI

Soroka, 2010 [Publ 3404], fig. 8b-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 2 2 1 --
Cladomics cladogram CRISTARIINA UNIONINI

Whelan et al., 2011 [Publ 3380], fig. 1-ML

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 34 2 1 --
Cladomics cladogram ALASMIDONTINA UNIONINI

Whelan et al., 2011 [Publ 3380], fig. 1-MP

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 34 2 1 --
Cladomics cladogram ALASMIDONTINA UNIONINI

Reis et al., 2013 [Publ 3580], fig. 4

cst iT tT bi riS oT roT roS iD
0.875 3 3 1 1 11 6 1 10.0
Cladomics cladogram ANODONTINA UNIONINI

Reis et al., 2013 [Publ 3580], fig. 5

cst iT tT bi riS oT roT roS iD
0.875 3 3 0 1 12 7 1 10.0
Cladomics cladogram ANODONTINA UNIONINI

Prié & Puillandre, 2014 [Publ 3529], fig. 2

cst iT tT bi riS oT roT roS iD
0.875 3 3 1 1 14 11 1 10.0
Cladomics cladogram ANODONTINA UNIONINI

Marshall et al., 2014 [Publ 3593], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 1 1 --
Cladomics cladogram ALASMIDONTINA UNIONINI

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 38 1 1 --
Cladomics cladogram UNIONINI ALASMIDONTINA

Bolotov et al., 2016 [Publ 3743], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 2 1 --
Cladomics cladogram ANODONTINA UNIONINI

Bolotov et al., 2016 [Publ 3743], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 2 1 --
Cladomics cladogram ANODONTINA UNIONINI

Lopes-Lima et al., 2017 [Publ 3734], fig. 1

cst iT tT bi riS oT roT roS iD
0.875 3 3 1 1 14 9 1 10.0
Cladomics cladogram ANODONTINA UNIONINI

Araujo et al., 2017 [Publ 3801], fig. 2

cst iT tT bi riS oT roT roS iD
0.909 1 1 0 1 13 3 1 --
Cladomics cladogram OXYNAIINA ACULAMPROTULINA ANODONTINA UNIONINA

Bogatov et al., 2018 [Publ 3859], fig. 1

cst iT tT bi riS oT roT roS iD
0.625 2 2 0 1 4 3 1 --
Cladomics cladogram Unio tumidus ANODONTINA Unio crassus Unio pictorum

Bogatov et al., 2018 [Publ 3859], fig. 2

cst iT tT bi riS oT roT roS iD
0.625 2 2 0 1 4 3 1 --
Cladomics cladogram Unio crassus Unio pictorum Unio tumidus ANODONTINA

Klishko et al., 2019 [Publ 3886], fig. 9

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 9 8 1 --
Cladomics cladogram ANODONTINA UNIONINI

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 1 1 --
Cladomics cladogram UNIONINI ANODONTINA

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 1 1 --
Cladomics cladogram UNIONINI ANODONTINA

Lopes-Lima et al., 2021 [Publ 4217], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 22 16 1 10.0
Cladomics cladogram ANODONTINA UNIONINI

Sonowal et al., 2021 [Publ 4221], fig. 3a

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 14 3 1 --
Cladomics cladogram ANODONTINA Unio ravoisieri Unio pictorum ACULAMPROTULINA

Sonowal et al., 2021 [Publ 4221], fig. 3b

cst iT tT bi riS oT roT roS iD
0.556 1 1 0 1 14 3 1 --
Cladomics cladogram ACULAMPROTULINA Lamprotula LAMPSILINI Potomida Unio pictorum ANODONTINA Unio ravoisieri LAMELLIDENTINI PLEUROBEMINI

Sonowal et al., 2021 [Publ 4221], fig. 4

cst iT tT bi riS oT roT roS iD
0.556 1 1 0 1 15 3 1 --
Cladomics cladogram ANODONTINA UNIONINI LAMELLIDENTINI

Lyubas et al., 2022 [Publ 4255], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 1 26 23 1 12.0
Cladomics cladogram ANODONTINA UNIONINI

Lyubas et al., 2022 [Publ 4255], fig. S1

cst iT tT bi riS oT roT roS iD
0.875 5 5 3 1 26 23 1 12.0
Cladomics cladogram ANODONTINA UNIONINI

Gomes-dos-Santos et al., 2023 [Publ 4463], fig. 2a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 20 1 1 --
Cladomics cladogram ANODONTINA UNIONINI

Gomes-dos-Santos et al., 2023 [Publ 4463], fig. 2b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 1 1 --
Cladomics cladogram ANODONTINA UNIONINI

Wu et al., 2024 [Publ 4379], fig. 4_6

cst iT tT bi riS oT roT roS iD
0.909 3 3 1 1 20 15 1 8.0
Cladomics cladogram ACULAMPROTULINA CRISTARIINA OXYNAIINA

Zieritz et al., 2024 [Publ 4381], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 44 1 1 --
Cladomics cladogram ANODONTINA UNIONINI

Wu et al., 2025 [Publ 4442], fig. 2

cst iT tT bi riS oT roT roS iD
0.929 33 33 31 1 2 2 1 36.0
Cladomics cladogram UNIONINI CRISTARIINA

Wu et al., 2025 [Publ 4442], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 15 15 13 1 2 2 1 18.0
Cladomics cladogram UNIONINI CRISTARIINA

Chen et al., 2026 [Publ 4805], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 33 33 31 1 1 1 1 36.0
Cladomics cladogram UNIONINI CRISTARIINA

Hoeh et al., 1995 [Publ 3391], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 7 7 4 1 0 0 0 12.0
Cladomics cladogram ALASMIDONTINA

Lydeard et al., 1996 [Publ 2656], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 1 21 0 0 14.0
Cladomics cladogram ALASMIDONTINA Megalonaias QUADRULINA Elliptoideus Eurynia Pleurobema Uniomerus Amblema Plectomerus Obliquaria Potamilus fragilis Ptychobranchus Medionidus Potamilus purpuratus Lampsilis Ellipsaria Villosa

Hoeh et al., 1996 [Publ 2694], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 2 0 0 --
Cladomics cladogram PLEUROBEMINI ALASMIDONTINA

Mulvey et al., 1997 [Publ 3390], fig. 1_3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 4 0 0 --
Cladomics cladogram ALASMIDONTINA QUADRULINI AMBLEMINI

Hoeh et al., 1998 [Publ 2695], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 4 0 0 --
Cladomics cladogram ANODONTINA Margaritifera AMBLEMINI

Hoeh et al., 1998 [Publ 2695], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 4 0 0 --
Cladomics cladogram Margaritifera ANODONTINA

King et al., 1999 [Publ 2699], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 9 9 7 1 0 0 0 16.0
Cladomics cladogram ALASMIDONTINA

King et al., 1999 [Publ 2699], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 6 6 3 1 0 0 0 15.0
Cladomics cladogram ALASMIDONTINA

Lydeard et al., 2000 [Publ 2654], fig. 2a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram ANODONTINA Cumberlandia LAMPSILINI AMBLEMINI QUADRULINI PLEUROBEMINI

Lydeard et al., 2000 [Publ 2654], fig. 2b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram ANODONTINA Cumberlandia LAMPSILINI AMBLEMINI QUADRULINI PLEUROBEMINI

Lydeard et al., 2000 [Publ 2654], fig. 2c

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram ANODONTINA Cumberlandia QUADRULINI LAMPSILINI AMBLEMINI PLEUROBEMINI

Lydeard et al., 2000 [Publ 2654], fig. 2d

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram ANODONTINA Cumberlandia AMBLEMINI LAMPSILINI QUADRULINI PLEUROBEMINI

Turner et al., 2000 [Publ 2722], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram QUADRULINI ALASMIDONTINA

Campbell, 2000 [Publ 2724], fig. 1-5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 2 0 0 --
Cladomics cladogram ALASMIDONTINA PLEUROBEMINI

Hoeh et al., 2002 [Publ 2673], fig. 1-F

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 7 0 0 --
Cladomics cladogram ALASMIDONTINA PLEUROBEMINI LAMPSILINI

Hoeh et al., 2002 [Publ 2673], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 7 0 0 --
Cladomics cladogram ALASMIDONTINA PLEUROBEMINI LAMPSILINI

Curole & Kocher, 2002 [Publ 2712], fig. 1a-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 0 0 --
Cladomics cladogram ALASMIDONTINA LAMPSILINI

Curole & Kocher, 2002 [Publ 2712], fig. 1b-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 0 0 --
Cladomics cladogram ALASMIDONTINA QUADRULINI

Krebs, 2004 [Publ 2692], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 3 0 0 --
Cladomics cladogram ALASMIDONTINA LAMPSILINI PLEUROBEMINI

Krebs, 2004 [Publ 2692], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 2 0 0 --
Cladomics cladogram ALASMIDONTINA LAMPSILINI PLEUROBEMINI

Huff et al., 2004 [Publ 2697], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 0 0 --
Cladomics cladogram ANODONTINA Margaritifera margaritifera Cumberlandia Pseudunio Margaritifera dahurica Margaritifera middendorffi 1 Margaritifera middendorffi 2 Margaritifera falcata 1 Margaritifera falcata 2 Margaritifera falcata 3 Margaritifera falcata 4

Curole & Kocher, 2005 [Publ 2682], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 0 0 --
Cladomics cladogram ALASMIDONTINA Margaritifera

Curole & Kocher, 2005 [Publ 2682], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 10 0 0 --
Cladomics cladogram ALASMIDONTINA QUADRULINI Plectomerus Amblema LAMPSILINI

Walker et al., 2006 [Publ 2666], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 34 0 0 --
Cladomics cladogram ALASMIDONTINA Cumberlandia Pseudunio Margaritifera GONIDEINI PSEUDODONTINI Cyrtonaias PLEUROBEMINI Potamilus Lampsilis Ligumia Ortmanniana

Walker et al., 2006 [Publ 2666], fig. 6-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 12 0 0 --
Cladomics cladogram ALASMIDONTINA PLEUROBEMINI LAMPSILINI

Zanatta & Murphy, 2006 [Publ 2672], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 55 0 0 --
Cladomics cladogram ALASMIDONTINA LAMPSILINI PLEUROBEMINI AMBLEMINI POPENAIADINI

Zanatta & Murphy, 2006 [Publ 2672], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 0 0 --
Cladomics cladogram ALASMIDONTINA Cumberlandia PLEUROBEMINI AMBLEMINI POPENAIADINI LAMPSILINI

Cyr et al., 2007 [Publ 2745], fig. 2A

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 1 0 0 13.0
Cladomics cladogram LAMPSILINI ALASMIDONTINA

Cyr et al., 2007 [Publ 2745], fig. 2D

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 0 0 0 13.0
Cladomics cladogram Pyganodon

Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 0 0 --
Cladomics cladogram CRISTARIINA LAMPSILINI CHAMBERLAINIINI LAMPROTULINI

Soroka, 2010 [Publ 3404], fig. 9a

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 1 0 0 0 14.0
Cladomics cladogram CRISTARIINA

Soroka, 2010 [Publ 3404], fig. 9b

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 0 0 0 13.0
Cladomics cladogram CRISTARIINA

Soroka, 2010 [Publ 3404], fig. 9c

cst iT tT bi riS oT roT roS iD
1.000 4 4 3 1 0 0 0 14.0
Cladomics cladogram CRISTARIINA

Jiang et al., 2010 [Publ 3438], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 0 0 --
Cladomics cladogram LAMPSILINI CRISTARIINA LAMPROTULINI CHAMBERLAINIINI

Boyer et al., 2011 [Publ 2718], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 6 6 2 1 29 0 0 14.0
Cladomics cladogram ALASMIDONTINA Reginaia PLEUROBEMINI QUADRULINI Amblema LAMPSILINI

Boyer et al., 2011 [Publ 2718], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 9 9 6 1 29 0 0 16.0
Cladomics cladogram ALASMIDONTINA QUADRULINA Megalonaias Toxolasma Amblema PLEUROBEMINI Reginaia Obliquaria Potamilus Ellipsaria Truncilla Lampsilis cardium Ligumia Obovaria Epioblasma Lampsilis teres Lampsilis siliquoidea Ortmanniana Lampsilis higginsii

Moyer & Díaz-Ferguson, 2012 [Publ 3375], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 3 0 0 13.0
Cladomics cladogram ALASMIDONTINA LAMPSILINI

Peng et al., 2012 [Publ 3689], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 2 0 0 --
Cladomics cladogram CRISTARIINA CHAMBERLAINIINI

Inoue et al., 2013 [Publ 3528], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 16 0 0 --
Cladomics cladogram GONIDEINI ALASMIDONTINA PLEUROBEMINI QUADRULINI Paetulunio AMBLEMINI Lampsilis Leaunio Obovaria Villosa Cambarunio

Zanatta & Harris, 2013 [Publ 3530], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 4 0 0 --
Cladomics cladogram ALASMIDONTINA LAMPSILINI

Hinzmann et al., 2013 [Publ 3541], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 0 0 0 10.0
Cladomics cladogram ANODONTINA

Inoue et al., 2013 [Publ 3579], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 0 0 --
Cladomics cladogram ALASMIDONTINA Cumberlandia Pseudunio Margaritifera

Huang et al., 2013 [Publ 3629], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 4 0 0 --
Cladomics cladogram ALASMIDONTINA GONIDEINI LAMPROTULINI QUADRULINI LAMPSILINI

Zhang et al., 2013 [Publ 4214], fig. 5

cst iT tT bi riS oT roT roS iD
0.400 2 4 0 1 2 0 0 12.0
Cladomics cladogram Cristaria Sinanodonta Margaritifera LAMPROTULINI

Froufe et al., 2014 [Publ 3592], fig. 2

cst iT tT bi riS oT roT roS iD
0.800 3 3 1 1 0 0 0 10.0
Cladomics cladogram ANODONTINA

Inoue et al., 2014 [Publ 3635], fig. 3A

cst iT tT bi riS oT roT roS iD
1.000 14 14 13 1 6 0 0 18.0
Cladomics cladogram GONIDEINI ALASMIDONTINA

Inoue et al., 2014 [Publ 3635], fig. 3B

cst iT tT bi riS oT roT roS iD
0.833 7 7 2 1 1 0 0 16.0
Cladomics cladogram AMBLEMINI ALASMIDONTINA

Klishko et al., 2014 [Publ 3666], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 0 0 0 11.0
Cladomics cladogram CRISTARIINA

Pfeiffer et al., 2016 [Publ 3660], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 37 0 0 --
Cladomics cladogram ALASMIDONTINA AMBLEMINI LAMPSILINI Elliptoideus QUADRULINI Elliptio Parvaspina Plethobasus Hemistena Eurynia Fusconaia Pleurobema riddellii Pleurobema sintoxia Pleurobema cordatum Pleuronaia Pleurobema pyriforme Pleurobema strodeanum Pleurobema clava

Xia et al., 2016 [Publ 3712], fig. 7B

cst iT tT bi riS oT roT roS iD
0.400 2 3 0 1 1 0 0 10.0
Cladomics cladogram Sinanodonta Cristaria CHAMBERLAINIINI

Bolotov et al., 2016 [Publ 3732], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 6 6 5 1 2 0 0 14.0
Cladomics cladogram CRISTARIINA Gibbosula Margaritifera

Klishko et al., 2016 [Publ 3733], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 0 0 0 11.0
Cladomics cladogram CRISTARIINA

Lopes-Lima et al., 2016 [Publ 3757], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 0 0 0 10.0
Cladomics cladogram ANODONTINA

Sayenko et al., 2017 [Publ 3760], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 0 0 0 13.0
Cladomics cladogram CRISTARIINA

Guerra et al., 2017 [Publ 3772], fig. 2a-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 6 0 0 --
Cladomics cladogram ALASMIDONTINA GONIDEINI LAMPROTULINI LAMPSILINI QUADRULINI

Guerra et al., 2017 [Publ 3772], fig. 2b-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 6 0 0 --
Cladomics cladogram ALASMIDONTINA GONIDEINI LAMPROTULINI LAMPSILINI QUADRULINI

Amaro et al., 2017 [Publ 3791], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 1 11 0 0 10.0
Cladomics cladogram ALASMIDONTINA LAMPSILINI QUADRULINI PLEUROBEMINI LAMPROTULINI GONIDEINI

Amaro et al., 2017 [Publ 3791], fig. 3A

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 8 0 0 9.0
Cladomics cladogram ALASMIDONTINA LAMPROTULINI GONIDEINI QUADRULINI LAMPSILINI

Amaro et al., 2017 [Publ 3791], fig. 3A-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 5 0 0 --
Cladomics cladogram LAMPSILINI ALASMIDONTINA

Amaro et al., 2017 [Publ 3791], fig. 3B

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 8 0 0 9.0
Cladomics cladogram ALASMIDONTINA LAMPSILINI QUADRULINI LAMPROTULINI GONIDEINI

Amaro et al., 2017 [Publ 3791], fig. 3B-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 5 0 0 --
Cladomics cladogram ALASMIDONTINA LAMPROTULINI GONIDEINI

Vikhrev et al., 2017 [Publ 3810], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 1 2 0 0 13.0
Cladomics cladogram CRISTARIINA Gibbosula Margaritifera

Hewitt et al., 2018 [Publ 3758], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 6 6 3 1 3 0 0 15.0
Cladomics cladogram PLEUROBEMINI ALASMIDONTINA

Bespalaya et al., 2018 [Publ 3811], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 8 8 7 1 2 0 0 14.0
Cladomics cladogram CRISTARIINA Gibbosula Margaritifera

Pieri et al., 2018 [Publ 3867], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 0 0 --
Cladomics cladogram ALASMIDONTINA QUADRULINI PLEUROBEMINI

Kondakov et al., 2018 [Publ 3914], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 1 0 0 0 13.0
Cladomics cladogram CRISTARIINA

Metzger et al., 2018 [Publ 3952], fig. 1C

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 2 0 0 --
Cladomics cladogram ALASMIDONTINA PLEUROBEMINI LAMPSILINI

Hewitt et al., 2019 [Publ 3962], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 0 0 --
Cladomics cladogram ALASMIDONTINA PLEUROBEMINI LAMPSILINI

Krebs et al., 2019 [Publ 4092], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 0 0 0 12.0
Cladomics cladogram Pyganodon

Inoue et al., 2020 [Publ 3941], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 41 0 0 --
Cladomics cladogram ALASMIDONTINA QUADRULINI PLEUROBEMINI AMBLEMINI LAMPSILINI

Inoue et al., 2020 [Publ 3941], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 0 0 --
Cladomics cladogram ALASMIDONTINA QUADRULINI Reginaia PLEUROBEMINI Amblema LAMPSILINI

Lopes-Lima et al., 2020 [Publ 4080], fig. 10

cst iT tT bi riS oT roT roS iD
1.000 12 12 8 1 2 0 0 19.0
Cladomics cladogram Margaritifera CRISTARIINA

Lopes-Lima et al., 2020 [Publ 4080], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 9 9 7 1 2 0 0 16.0
Cladomics cladogram Margaritifera CRISTARIINA

Lopes-Lima et al., 2020 [Publ 4080], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 1 2 0 0 15.0
Cladomics cladogram Margaritifera CRISTARIINA

Lopes-Lima et al., 2020 [Publ 4080], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 1 2 0 0 10.0
Cladomics cladogram Margaritifera CRISTARIINA

Lopes-Lima et al., 2020 [Publ 4080], fig. 9

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 2 0 0 12.0
Cladomics cladogram Margaritifera CRISTARIINA

Lopes-Lima et al., 2021 [Publ 4217], fig. 11

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 0 0 0 11.0
Cladomics cladogram Anodonta

Bauer et al., 2021 [Publ 4365], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 8 8 5 1 0 0 0 13.0
Cladomics cladogram CRISTARIINA

Stewart et al., 2021 [Publ 4451], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 10 10 9 1 6 0 0 15.0
Cladomics cladogram LAMPSILINI ALASMIDONTINA

Sano et al., 2022 [Publ 4311], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 12 12 12 1 0 0 0 20.0
Cladomics cladogram CRISTARIINA

Sano et al., 2022 [Publ 4311], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 18 18 16 1 0 0 0 23.0
Cladomics cladogram CRISTARIINA

Smith et al., 2023 [Publ 4291], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 1 0 0 0 17.0
Cladomics cladogram ALASMIDONTINA

Bogan et al., 2023 [Publ 4295], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 1 1 0 0 14.0
Cladomics cladogram Gibbosula CRISTARIINA

Bogan et al., 2023 [Publ 4295], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 1 1 0 0 14.0
Cladomics cladogram Gibbosula CRISTARIINA

Bogan et al., 2023 [Publ 4295], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 12 12 9 1 1 0 0 18.0
Cladomics cladogram Gibbosula CRISTARIINA

Bulakhova et al., 2023 [Publ 4325], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 0 0 0 14.0
Cladomics cladogram Beringiana

Douda et al., 2024 [Publ 4364], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 13 13 12 1 1 0 0 20.0
Cladomics cladogram Margaritifera CRISTARIINA

Dai et al., 2024 [Publ 4385], fig. 3-BI

cst iT tT bi riS oT roT roS iD
1.000 27 27 24 1 4 0 0 32.0
Cladomics cladogram CRISTARIINA LAMELLIDENTINI INDOCHINELLINI

Dai et al., 2024 [Publ 4385], fig. 3-ML

cst iT tT bi riS oT roT roS iD
1.000 27 27 25 1 4 0 0 32.0
Cladomics cladogram CRISTARIINA LAMELLIDENTINI INDOCHINELLINI

Dai et al., 2024 [Publ 4389], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 43 0 0 10.0
Cladomics cladogram CRISTARIINA Rectidentini CHAMBERLAINIINI LAMPROTULINI GONIDEINI SCHEPMANIINI PSEUDODONTINI

Kashiri et al., 2024 [Publ 4822], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 1 2 0 0 13.0
Cladomics cladogram CRISTARIINA Gibbosula Margaritifera

Xiang et al., 2025 [Publ 4433], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 27 27 25 1 1 0 0 32.0
Cladomics cladogram INDOCHINELLINI CRISTARIINA

Dai et al., 2025 [Publ 4441], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 44 0 0 10.0
Cladomics cladogram CRISTARIINA Rectidentini CHAMBERLAINIINI LAMPROTULINI GONIDEINI SCHEPMANIINI PSEUDODONTINI

Chen et al., 2025 [Publ 4663], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 45 0 0 10.0
Cladomics cladogram CRISTARIINA Rectidentini CHAMBERLAINIINI LAMPROTULINI GONIDEINI SCHEPMANIINI PSEUDODONTINI

Dai et al., 2026 [Publ 4757], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 30 30 29 1 4 0 0 37.0
Cladomics cladogram CRISTARIINA INDOCHINELLINI LAMELLIDENTINI

Itoh et al., 2026 [Publ 4842], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 1 0 0 0 15.0
Cladomics cladogram CRISTARIINA

Tumpeesuwan et al., 2026 [Publ 4845], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 1 0 0 0 18.0
Cladomics cladogram CRISTARIINA

Barraza et al., 2026 [Publ 4851], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 18 0 0 --
Cladomics cladogram CRISTARIINA Amblema Psoronaias Nephronaias Reginaia QUADRULINI PLEUROBEMINI Mesonaias Pachynaias Cyrtonaias Popenaias