Page last updated
Wed 1 October 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Bivalvia | Unionoida | Etherioidea | Iridinidae

subfamily ASPATHARIINAE Modell, 1942: cladomics
     2 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 22 (23 total)
number of subtaxa (rank: genus) = 2
analyzed in 46 cladograms in 27 publications
maximum number of terminals in an analysis = 2 (mean = 1.63, median = 2)
maximum number of subtaxa (genus) = 2 (mean = 1.61, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 29
maximum number of included terminals = 2, (mean = 2 median = 2)
maximum number of subtaxa (genus) = 2 (mean = 1.97, median = 2)
proportion of cladograms that support basic monophyly = 0.97

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 20
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 2 (mean = 2, median = 2)
maximum number of subtaxa (genus) = 2 (mean = 2, median = 2)
proportion of cladograms that support rigorous monophyly = 0.95

Huang et al., 2019 [Publ 3843], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

  +-- Aspatharia **
--+             
  +-- Chambardia **

Bolotov et al., 2016 [Publ 3743], fig. 4

  +-- Chambardia **
--+             
  +-- Aspatharia **

Wu et al., 2020 [Publ 4121], fig. 2

  +-- Aspatharia **
--+             
  +-- Chambardia **

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

  +-- Aspatharia **
--+             
  +-- Chambardia **

Combosch et al., 2017 [Publ 3806], fig. 2

  +-- Chambardia **
--+             
  +-- Aspatharia **

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

  +-- Iridininae
--+             
  +-- Chambardia **

Huang et al., 2018 [Publ 3828], fig. 3_5

  +-- Aspatharia **
--+             
  +-- Chambardia **

Wu et al., 2021 [Publ 4366], fig. 6

  +-- Chambardia **
--+             
  +-- Aspatharia **

Graf & Cummings, 2006 [Publ 1968], fig. 2

  +----- Iridininae
--+                
  |  +-- Chambardia **
  +--+             
     +-- Aspatharia **

Huang et al., 2019 [Publ 3843], fig. S4

  +-- Aspatharia **
--+             
  +-- Chambardia **

Bolotov et al., 2017 [Publ 3761], fig. S2

  +-- Aspatharia **
--+             
  +-- Chambardia **

Sano et al., 2020 [Publ 4138], fig. 2

  +-- Aspatharia **
--+             
  +-- Chambardia **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

  +-- Aspatharia **
--+             
  +-- Chambardia **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

  +-- Chambardia **
--+             
  +-- Aspatharia **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

  +-- Aspatharia **
--+             
  +-- Chambardia **

Combosch et al., 2017 [Publ 3806], fig. 1

  +-- Chambardia **
--+             
  +-- Aspatharia **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

  +-- Aspatharia **
--+             
  +-- Chambardia **

Bolotov et al., 2017 [Publ 3761], fig. S1

  +-- Chambardia **
--+             
  +-- Aspatharia **

Chen et al., 2025 [Publ 4455], fig. 1

  +-- Aspatharia **
--+             
  +-- Chambardia **

Bolotov et al., 2016 [Publ 3743], fig. 2

  +-- Aspatharia **
--+             
  +-- Chambardia **

Wu et al., 2021 [Publ 4366], fig. 2_I

  +-- Aspatharia **
--+             
  +-- Chambardia **

Sano et al., 2020 [Publ 4138], fig. 4

  +-- Aspatharia **
--+             
  +-- Chambardia **

Hoeh et al., 2009 [Publ 2716], fig. 4

  +----- Iridininae
--+                
  |  +-- Chambardia **
  +--+             
     +-- Aspatharia **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

  +-- Aspatharia **
--+             
  +-- Chambardia **

Hoeh et al., 2009 [Publ 2716], fig. 5

  +----- Iridininae
--+                
  |  +-- Chambardia **
  +--+             
     +-- Aspatharia **

Sano et al., 2020 [Publ 4138], fig. 3

  +-- Aspatharia **
--+             
  +-- Chambardia **

Hoeh et al., 2009 [Publ 2716], fig. 3

  +-- Chambardia **
  |             
--+-- Aspatharia **
  |             
  +-- Iridininae

Huang et al., 2019 [Publ 3843], fig. S3

  +-- Aspatharia **
--+             
  +-- Chambardia **

Wu et al., 2020 [Publ 4121], fig. S1

  +-- Aspatharia **
--+             
  +-- Chambardia **

Guerra et al., 2019 [Publ 4168], fig. 1

-- Chambardia_rubens **

Smith, 2021 [Publ 4331], fig. 1

-- Aspatharia_pfeifferiana **

Lemer et al., 2019 [Publ 4093], fig. 1

-- Aspatharia_pfeifferiana **

González & Giribet, 2015 [Publ 3632], fig. 2

-- Aspatharia_pfeifferiana **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

-- Aspatharia_pfeifferiana **

Konopleva et al., 2017 [Publ 3807], fig. 6

-- Chambardia_wahlbergi **

Bieler et al., 2014 [Publ 3622], fig. 28

-- Aspatharia_pfeifferiana **

Graf et al., 2015 [Publ 3619], fig. 1

-- Chambardia_wahlbergi **

Elderkin et al., 2016 [Publ 3693], fig. 2

-- Aspatharia_pfeifferiana **

González & Giribet, 2015 [Publ 3632], fig. 4

-- Aspatharia_pfeifferiana **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

-- Chambardia_wahlbergi **

González & Giribet, 2015 [Publ 3632], fig. 3

-- Aspatharia_pfeifferiana **

Walker et al., 2006 [Publ 2666], fig. 6

-- Chambardia_rubens **

González & Giribet, 2015 [Publ 3632], fig. 6

-- Aspatharia_pfeifferiana **

Guerra et al., 2019 [Publ 4168], fig. 2

-- Chambardia_rubens **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

-- Aspatharia_pfeifferiana **

Bieler et al., 2014 [Publ 3622], fig. 26_27_29_30

-- Aspatharia_pfeifferiana **