Bivalvia | Unionida | Etherioidea | Iridinidae

subfamily ASPATHARIINAE Modell, 1942: cladomics

2 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 22 (23 total)
number of subtaxa (rank: genus) = 2
analyzed in 48 cladograms in 28 publications
maximum number of terminals in an analysis = 4 (mean = 1.67, median = 2)
maximum number of subtaxa (genus) = 2 (mean = 1.6, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 30
maximum number of included terminals = 4, (mean = 2.07 median = 2)
maximum number of subtaxa (genus) = 2 (mean = 1.97, median = 2)
proportion of cladograms that support basic monophyly = 0.97

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 21
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 4 (mean = 2.1, median = 2)
maximum number of subtaxa (genus) = 2 (mean = 2, median = 2)
proportion of cladograms that support rigorous monophyly = 0.95

Cladograms

Graf & Cummings, 2006 [Publ 1968], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 45 2 1 ** 
  +----- Iridininae
--+                
  |  +-- Chambardia **
  +--+             
     +-- Aspatharia **

Hoeh et al., 2009 [Publ 2716], fig. 3

cstiTtTbiriSoTroTroSiD
0.833 2 4 0 2 45 2 1 0.0 
           +-- Chambardia **      
           |                   
  +--------+-- Aspatharia **      
  |        |                   
--+        +-- Iridininae      
  |                            
  |  +-------- Etheria         
  +--+                         
     |  +----- Mulleria        
     +--+                      
        |  +-- Monocondylaeinae
        +--+                   
           +-- Anodontitinae

Hoeh et al., 2009 [Publ 2716], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 44 2 1 0.0 
  +----- Iridininae
--+                
  |  +-- Chambardia **
  +--+             
     +-- Aspatharia **

Hoeh et al., 2009 [Publ 2716], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 44 2 1 0.0 
  +----- Iridininae
--+                
  |  +-- Chambardia **
  +--+             
     +-- Aspatharia **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

cstiTtTbiriSoTroTroSiD
0.833 2 2 0 2 34 1 1 0.0 
        +-- Aspatharia **      
  +-----+                   
  |     +-- Chambardia **      
--+                         
  |  +----- Iridininae      
  +--+                      
     |  +-- Mycetopodinae   
     +--+                   
        +-- Monocondylaeinae

Whelan et al., 2011 [Publ 3380], fig. 1-MP

cstiTtTbiriSoTroTroSiD
0.833 2 2 0 2 34 1 1 0.0 
        +-- Aspatharia **      
  +-----+                   
  |     +-- Chambardia **      
--+                         
  |  +----- Iridininae      
  +--+                      
     |  +-- Mycetopodinae   
     +--+                   
        +-- Monocondylaeinae

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cstiTtTbiriSoTroTroSiD
0.833 2 2 0 2 79 1 1 0.0 
        +-- Aspatharia **      
  +-----+                   
  |     +-- Chambardia **      
--+                         
  |  +----- Etheria         
  +--+                      
     |  +-- Monocondylaeinae
     +--+                   
        +-- Mycetopodinae

Combosch et al., 2017 [Publ 3806], fig. 1

cstiTtTbiriSoTroTroSiD
0.833 2 2 0 2 13 1 1 0.0 
           +-- Chambardia **      
  +--------+                   
  |        +-- Aspatharia **      
--+                            
  |  +-------- Etheria         
  +--+                         
     |  +----- Iridininae      
     +--+                      
        |  +-- Monocondylaeinae
        +--+                   
           +-- Anodontitinae

Combosch et al., 2017 [Publ 3806], fig. 2

cstiTtTbiriSoTroTroSiD
0.833 2 2 0 2 13 1 1 0.0 
           +-- Chambardia **      
  +--------+                   
  |        +-- Aspatharia **      
--+                            
  |  +-------- Etheria         
  +--+                         
     |  +----- Iridininae      
     +--+                      
        |  +-- Monocondylaeinae
        +--+                   
           +-- Anodontitinae

Huang et al., 2018 [Publ 3828], fig. 3_5

cstiTtTbiriSoTroTroSiD
0.833 2 2 0 2 27 1 1 0.0 
        +-- Aspatharia **      
  +-----+                   
  |     +-- Chambardia **      
--+                         
  |  +----- Etheria         
  +--+                      
     |  +-- Monocondylaeinae
     +--+                   
        +-- Anodontitinae

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

cstiTtTbiriSoTroTroSiD
0.833 2 2 0 2 26 1 1 0.0 
        +-- Aspatharia **      
  +-----+                   
  |     +-- Chambardia **      
--+                         
  |  +----- Etheria         
  +--+                      
     |  +-- Anodontitinae   
     +--+                   
        +-- Monocondylaeinae

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

cstiTtTbiriSoTroTroSiD
0.833 2 2 0 2 26 1 1 0.0 
        +-- Chambardia **      
  +-----+                   
  |     +-- Aspatharia **      
--+                         
  |  +----- Etheria         
  +--+                      
     |  +-- Monocondylaeinae
     +--+                   
        +-- Anodontitinae

Huang et al., 2019 [Publ 3843], fig. 2

cstiTtTbiriSoTroTroSiD
0.833 2 2 0 2 199 3 1 0.0 
  +----- Etheria   
--+                
  |  +-- Aspatharia **
  +--+             
     +-- Chambardia **

Huang et al., 2019 [Publ 3843], fig. S3

cstiTtTbiriSoTroTroSiD
0.833 2 2 0 2 197 3 1 0.0 
  +----- Etheria   
--+                
  |  +-- Aspatharia **
  +--+             
     +-- Chambardia **

Huang et al., 2019 [Publ 3843], fig. S4

cstiTtTbiriSoTroTroSiD
0.833 2 2 0 2 199 3 1 0.0 
  +-------- Etheria         
  |                         
  |     +-- Aspatharia **      
  +-----+                   
  |     +-- Chambardia **      
--+                         
  |     +-- Anodontites     
  +-----+                   
  |     +-- Iridininae      
  |                         
  |  +----- Mycetopodinae   
  +--+                      
     |  +-- Monocondylaeinae
     +--+                   
        +-- Lamproscapha

Wu et al., 2020 [Publ 4121], fig. 2

cstiTtTbiriSoTroTroSiD
0.833 2 2 0 2 32 1 1 0.0 
        +-- Aspatharia **      
  +-----+                   
  |     +-- Chambardia **      
--+                         
  |  +----- Etheria         
  +--+                      
     |  +-- Monocondylaeinae
     +--+                   
        +-- Anodontitinae

Wu et al., 2020 [Publ 4121], fig. S1

cstiTtTbiriSoTroTroSiD
0.833 2 2 0 2 32 1 1 0.0 
        +-- Aspatharia **      
  +-----+                   
  |     +-- Chambardia **      
--+                         
  |  +----- Etheria         
  +--+                      
     |  +-- Monocondylaeinae
     +--+                   
        +-- Anodontitinae

Sano et al., 2020 [Publ 4138], fig. 2

cstiTtTbiriSoTroTroSiD
0.833 2 2 0 2 67 1 1 0.0 
        +-- Aspatharia **      
  +-----+                   
  |     +-- Chambardia **      
--+                         
  |  +----- Etheria         
  +--+                      
     |  +-- Monocondylaeinae
     +--+                   
        +-- Anodontitinae

Sano et al., 2020 [Publ 4138], fig. 3

cstiTtTbiriSoTroTroSiD
0.833 2 2 0 2 67 1 1 0.0 
        +-- Aspatharia **      
  +-----+                   
  |     +-- Chambardia **      
--+                         
  |  +----- Etheria         
  +--+                      
     |  +-- Monocondylaeinae
     +--+                   
        +-- Anodontitinae

Sano et al., 2020 [Publ 4138], fig. 4

cstiTtTbiriSoTroTroSiD
0.833 2 2 0 2 67 1 1 0.0 
        +-- Aspatharia **      
  +-----+                   
  |     +-- Chambardia **      
--+                         
  |  +----- Etheria         
  +--+                      
     |  +-- Monocondylaeinae
     +--+                   
        +-- Anodontitinae

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cstiTtTbiriSoTroTroSiD
0.833 4 4 3 2 629 3 1 0.0 
                          +-- Aspatharia **             
  +-----------------------+                          
  |                       +-- Chambardia **             
--+                                                  
  |  +----------------------- Etheria                
  |  |                                               
  +--+  +-------------------- Anodontites_trapesialis
     |  |                                            
     +--+  +----------------- Mulleria               
        |  |                                         
        +--+  +-------------- Lamproscapha           
           |  |                                      
           +--+           +-- Monocondylaea          
              |  +--------+                          
              |  |        +-- Tamsiella              
              +--+                                   
                 |  +-------- Mycetopodinae          
                 |  |                                
                 +--+     +-- Anodontites_guanarensis
                    |  +--+                          
                    |  |  +-- Fossula                
                    +--+                             
                       |  +-- Anodontites_trigona    
                       +--+                          
                          +-- Anodontites_elongata

Bolotov et al., 2016 [Publ 3743], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 28 0 0 0.0 
  +----- Etheria   
--+                
  |  +-- Aspatharia **
  +--+             
     +-- Chambardia **

Bolotov et al., 2016 [Publ 3743], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 28 0 0 0.0 
  +----- Etheria   
--+                
  |  +-- Chambardia **
  +--+             
     +-- Aspatharia **

Bolotov et al., 2017 [Publ 3761], fig. S1

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 137 0 0 0.0 
  +----- Etheria   
--+                
  |  +-- Chambardia **
  +--+             
     +-- Aspatharia **

Bolotov et al., 2017 [Publ 3761], fig. S2

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 137 0 0 0.0 
     +-- Monocondylaeinae
  +--+                   
  |  +-- Etheria         
--+                      
  |  +-- Aspatharia **      
  +--+                   
     +-- Chambardia **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 120 0 0 0.0 
  +----- Etheria   
--+                
  |  +-- Aspatharia **
  +--+             
     +-- Chambardia **

Wu et al., 2021 [Publ 4366], fig. 2_I

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 89 0 0 0.0 
        +-- Aspatharia **      
  +-----+                   
  |     +-- Chambardia **      
--+                         
  |  +----- Etheria         
  +--+                      
     |  +-- Monocondylaeinae
     +--+                   
        +-- Anodontitinae

Wu et al., 2021 [Publ 4366], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 89 0 0 0.0 
        +-- Chambardia **      
  +-----+                   
  |     +-- Aspatharia **      
--+                         
  |  +----- Etheria         
  +--+                      
     |  +-- Monocondylaeinae
     +--+                   
        +-- Anodontitinae

Chen et al., 2025 [Publ 4455], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 24 0 0 0.0 
        +-- Aspatharia **      
  +-----+                   
  |     +-- Chambardia **      
--+                         
  |  +----- Etheria         
  +--+                      
     |  +-- Monocondylaeinae
     +--+                   
        +-- Anodontitinae

Walker et al., 2006 [Publ 2666], fig. 6

cstiTtTbiriSoTroTroSiD
0.800 1 1 0 1 35 2 1 0.0 
  +-------- Chambardia **      
--+                         
  |  +----- Mulleria        
  +--+                      
     |  +-- Anodontitinae   
     +--+                   
        +-- Monocondylaeinae

Guerra et al., 2019 [Publ 4168], fig. 1

cstiTtTbiriSoTroTroSiD
0.800 1 1 0 1 40 1 1 0.0 
  +-------------- Chambardia **             
--+                                      
  |  +----------- Anodontites_trapesialis
  +--+                                   
     |  +-------- Monocondylaea          
     +--+                                
        |  +----- Lamproscapha           
        +--+                             
           |  +-- Anodontites_elongata   
           +--+                          
              +-- Fossula

Guerra et al., 2019 [Publ 4168], fig. 2

cstiTtTbiriSoTroTroSiD
0.800 1 1 0 1 40 1 1 0.0 
  +----------------- Chambardia **             
--+                                         
  |  +-------------- Iridininae             
  +--+                                      
     |  +----------- Anodontites_trapesialis
     +--+                                   
        |  +-------- Monocondylaea          
        +--+                                
           |  +----- Lamproscapha           
           +--+                             
              |  +-- Anodontites_elongata   
              +--+                          
                 +-- Fossula

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 1 26 2 1 0.0 
  +-- Iridininae
--+             
  +-- Chambardia **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cstiTtTbiriSoTroTroSiD
0.800 1 1 0 1 139 1 1 0.0 
  +-------------- Chambardia **             
--+                                      
  |  +----------- Anodontites_trapesialis
  +--+                                   
     |  +-------- Monocondylaea          
     +--+                                
        |  +----- Lamproscapha           
        +--+                             
           |  +-- Anodontites_elongata   
           +--+                          
              +-- Fossula

Bieler et al., 2014 [Publ 3622], fig. 26_27_29_30

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 4 0 0 0.0 
  +-- Aspatharia **      
--+                   
  +-- Margaritiferinae

Bieler et al., 2014 [Publ 3622], fig. 28

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 4 0 0 0.0 
  +-- Aspatharia **      
--+                   
  +-- Margaritiferinae

Graf et al., 2015 [Publ 3619], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 19 0 0 0.0 
  +----- Chambardia **      
--+                      
  |  +-- Monocondylaeinae
  +--+                   
     +-- Etheria

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 39 0 0 0.0 
  +-- Anodontitinae
--+                
  +-- Chambardia **

González & Giribet, 2015 [Publ 3632], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 5 0 0 0.0 
  +-- Aspatharia **      
--+                   
  +-- Margaritiferinae

González & Giribet, 2015 [Publ 3632], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 5 0 0 0.0 
  +-- Aspatharia **      
--+                   
  +-- Margaritiferinae

González & Giribet, 2015 [Publ 3632], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 5 0 0 0.0 
  +-------- Aspatharia **      
--+                         
  |  +----- Velesunioninae  
  +--+                      
     |  +-- Unioninae       
     +--+                   
        +-- Margaritiferinae

González & Giribet, 2015 [Publ 3632], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 5 0 0 0.0 
  +-- Aspatharia **      
--+                   
  +-- Margaritiferinae

Elderkin et al., 2016 [Publ 3693], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 2 0 0 0.0 
  +-- Etheria       
  |                 
--+-- Aspatharia **    
  |                 
  +-- Velesunioninae

Konopleva et al., 2017 [Publ 3807], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 0 0 0.0 
  +-- Etheria   
--+             
  +-- Chambardia **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 56 0 0 0.0 
  +-- Etheria   
--+             
  +-- Aspatharia **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 56 0 0 0.0 
  +-- Etheria   
--+             
  +-- Aspatharia **

Lemer et al., 2019 [Publ 4093], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 5 0 0 0.0 
  +-- Etheria   
--+             
  +-- Aspatharia **

Smith, 2021 [Publ 4331], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 19 0 0 0.0 
  +-- Etheria   
--+             
  +-- Aspatharia **