Bivalvia | Unionida | Etherioidea | Iridinidae

subfamily ASPATHARIINAE Modell, 1942: cladomics

2 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 22 (23 total)
number of subtaxa (rank: genus) = 2
analyzed in 52 cladograms in 28 publications
maximum number of terminals in an analysis = 4 (mean = 1.62, median = 2)
maximum number of subtaxa (genus) = 2 (mean = 1.56, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 30
maximum number of included terminals = 4, (mean = 2.07 median = 2)
maximum number of subtaxa (genus) = 2 (mean = 1.97, median = 2)
proportion of cladograms that support basic monophyly = 0.97

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 21
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 4 (mean = 2.1, median = 2)
maximum number of subtaxa (genus) = 2 (mean = 2, median = 2)
proportion of cladograms that support rigorous monophyly = 0.95

Cladograms

Graf & Cummings, 2006 [Publ 1968], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 45 2 1 **
Cladomics cladogram IRIDININAE Chambardia Aspatharia

Hoeh et al., 2009 [Publ 2716], fig. 3

cst iT tT bi riS oT roT roS iD
0.833 2 4 0 2 45 2 1 0.0
Cladomics cladogram Chambardia Aspatharia IRIDININAE Etheria Mulleria MONOCONDYLAEINAE ANODONTITINAE

Hoeh et al., 2009 [Publ 2716], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 44 2 1 0.0
Cladomics cladogram IRIDININAE Chambardia Aspatharia

Hoeh et al., 2009 [Publ 2716], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 44 2 1 0.0
Cladomics cladogram IRIDININAE Chambardia Aspatharia

Whelan et al., 2011 [Publ 3380], fig. 1-ML

cst iT tT bi riS oT roT roS iD
0.833 2 2 0 2 34 1 1 0.0
Cladomics cladogram Aspatharia Chambardia IRIDININAE MYCETOPODINAE MONOCONDYLAEINAE

Whelan et al., 2011 [Publ 3380], fig. 1-MP

cst iT tT bi riS oT roT roS iD
0.833 2 2 0 2 34 1 1 0.0
Cladomics cladogram Aspatharia Chambardia IRIDININAE MYCETOPODINAE MONOCONDYLAEINAE

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
0.833 2 2 0 2 79 1 1 0.0
Cladomics cladogram Aspatharia Chambardia Etheria MONOCONDYLAEINAE MYCETOPODINAE

Combosch et al., 2017 [Publ 3806], fig. 1

cst iT tT bi riS oT roT roS iD
0.833 2 2 0 2 211 1 1 0.0
Cladomics cladogram Chambardia Aspatharia Etheria IRIDININAE MONOCONDYLAEINAE ANODONTITINAE

Combosch et al., 2017 [Publ 3806], fig. 2

cst iT tT bi riS oT roT roS iD
0.833 2 2 0 2 212 1 1 0.0
Cladomics cladogram Chambardia Aspatharia Etheria IRIDININAE MONOCONDYLAEINAE ANODONTITINAE

Huang et al., 2018 [Publ 3828], fig. 3_5

cst iT tT bi riS oT roT roS iD
0.833 2 2 0 2 27 1 1 0.0
Cladomics cladogram Aspatharia Chambardia Etheria MONOCONDYLAEINAE ANODONTITINAE

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

cst iT tT bi riS oT roT roS iD
0.833 2 2 0 2 26 1 1 0.0
Cladomics cladogram Aspatharia Chambardia Etheria ANODONTITINAE MONOCONDYLAEINAE

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

cst iT tT bi riS oT roT roS iD
0.833 2 2 0 2 26 1 1 0.0
Cladomics cladogram Chambardia Aspatharia Etheria MONOCONDYLAEINAE ANODONTITINAE

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
0.833 2 2 0 2 199 3 1 0.0
Cladomics cladogram Etheria Aspatharia Chambardia

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
0.833 2 2 0 2 197 3 1 0.0
Cladomics cladogram Etheria Aspatharia Chambardia

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.833 2 2 0 2 199 3 1 0.0
Cladomics cladogram Etheria Aspatharia Chambardia Anodontites IRIDININAE MYCETOPODINAE MONOCONDYLAEINAE Lamproscapha

Wu et al., 2020 [Publ 4121], fig. 2

cst iT tT bi riS oT roT roS iD
0.833 2 2 0 2 32 1 1 0.0
Cladomics cladogram Aspatharia Chambardia Etheria MONOCONDYLAEINAE ANODONTITINAE

Wu et al., 2020 [Publ 4121], fig. S1

cst iT tT bi riS oT roT roS iD
0.833 2 2 0 2 32 1 1 0.0
Cladomics cladogram Aspatharia Chambardia Etheria MONOCONDYLAEINAE ANODONTITINAE

Sano et al., 2020 [Publ 4138], fig. 2

cst iT tT bi riS oT roT roS iD
0.833 2 2 0 2 67 1 1 0.0
Cladomics cladogram Aspatharia Chambardia Etheria MONOCONDYLAEINAE ANODONTITINAE

Sano et al., 2020 [Publ 4138], fig. 3

cst iT tT bi riS oT roT roS iD
0.833 2 2 0 2 67 1 1 0.0
Cladomics cladogram Aspatharia Chambardia Etheria MONOCONDYLAEINAE ANODONTITINAE

Sano et al., 2020 [Publ 4138], fig. 4

cst iT tT bi riS oT roT roS iD
0.833 2 2 0 2 67 1 1 0.0
Cladomics cladogram Aspatharia Chambardia Etheria MONOCONDYLAEINAE ANODONTITINAE

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.833 4 4 3 2 629 3 1 0.0
Cladomics cladogram Aspatharia Chambardia Etheria Anodontites trapesialis Mulleria Lamproscapha Monocondylaea Tamsiella MYCETOPODINAE Anodontites guanarensis Fossula Anodontites trigona Anodontites elongata

Bolotov et al., 2016 [Publ 3743], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 28 0 0 0.0
Cladomics cladogram Etheria Aspatharia Chambardia

Bolotov et al., 2016 [Publ 3743], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 28 0 0 0.0
Cladomics cladogram Etheria Chambardia Aspatharia

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 137 0 0 0.0
Cladomics cladogram Etheria Chambardia Aspatharia

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 137 0 0 0.0
Cladomics cladogram MONOCONDYLAEINAE Etheria Aspatharia Chambardia

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 120 0 0 0.0
Cladomics cladogram Etheria Aspatharia Chambardia

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 89 0 0 0.0
Cladomics cladogram Aspatharia Chambardia Etheria MONOCONDYLAEINAE ANODONTITINAE

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 89 0 0 0.0
Cladomics cladogram Chambardia Aspatharia Etheria MONOCONDYLAEINAE ANODONTITINAE

Chen et al., 2025 [Publ 4455], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 24 0 0 0.0
Cladomics cladogram Aspatharia Chambardia Etheria MONOCONDYLAEINAE ANODONTITINAE

Walker et al., 2006 [Publ 2666], fig. 6

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 35 2 1 0.0
Cladomics cladogram Chambardia Mulleria ANODONTITINAE MONOCONDYLAEINAE

Guerra et al., 2019 [Publ 4168], fig. 1

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 40 1 1 0.0
Cladomics cladogram Chambardia Anodontites trapesialis Monocondylaea Lamproscapha Anodontites elongata Fossula

Guerra et al., 2019 [Publ 4168], fig. 2

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 40 1 1 0.0
Cladomics cladogram Chambardia IRIDININAE Anodontites trapesialis Monocondylaea Lamproscapha Anodontites elongata Fossula

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 26 2 1 0.0
Cladomics cladogram IRIDININAE Chambardia

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 139 1 1 0.0
Cladomics cladogram Chambardia Anodontites trapesialis Monocondylaea Lamproscapha Anodontites elongata Fossula

Bieler et al., 2014 [Publ 3622], fig. 26

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 0.0
Cladomics cladogram Aspatharia MARGARITIFERINAE

Bieler et al., 2014 [Publ 3622], fig. 27

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 0.0
Cladomics cladogram Aspatharia MARGARITIFERINAE

Bieler et al., 2014 [Publ 3622], fig. 28

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 0.0
Cladomics cladogram Aspatharia MARGARITIFERINAE

Bieler et al., 2014 [Publ 3622], fig. 29

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 0.0
Cladomics cladogram Aspatharia MARGARITIFERINAE

Bieler et al., 2014 [Publ 3622], fig. 30

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 0.0
Cladomics cladogram Aspatharia MARGARITIFERINAE

Bieler et al., 2014 [Publ 3622], fig. 31

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 0.0
Cladomics cladogram MARGARITIFERINAE Aspatharia

Graf et al., 2015 [Publ 3619], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 0 0 0.0
Cladomics cladogram Chambardia MONOCONDYLAEINAE Etheria

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 39 0 0 0.0
Cladomics cladogram ANODONTITINAE Chambardia

González & Giribet, 2015 [Publ 3632], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 0.0
Cladomics cladogram Aspatharia MARGARITIFERINAE

González & Giribet, 2015 [Publ 3632], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 0.0
Cladomics cladogram Aspatharia MARGARITIFERINAE

González & Giribet, 2015 [Publ 3632], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 0.0
Cladomics cladogram Aspatharia VELESUNIONINAE UNIONINAE MARGARITIFERINAE

González & Giribet, 2015 [Publ 3632], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 0.0
Cladomics cladogram Aspatharia MARGARITIFERINAE

Elderkin et al., 2016 [Publ 3693], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 2 0 0 0.0
Cladomics cladogram Etheria Aspatharia VELESUNIONINAE

Konopleva et al., 2017 [Publ 3807], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 0.0
Cladomics cladogram Etheria Chambardia

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 0 0 0.0
Cladomics cladogram Etheria Aspatharia

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 0 0 0.0
Cladomics cladogram Etheria Aspatharia

Lemer et al., 2019 [Publ 4093], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 0.0
Cladomics cladogram Etheria Aspatharia

Smith, 2021 [Publ 4331], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 0 0 0.0
Cladomics cladogram Etheria Aspatharia