Bivalvia | Unionida | Etherioidea | Iridinidae

subfamily ASPATHARIINAE Modell, 1942: cladomics

2 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 22 (23 total)
number of subtaxa (rank: genus) = 2
analyzed in 48 cladograms in 28 publications
maximum number of terminals in an analysis = 4 (mean = 1.67, median = 2)
maximum number of subtaxa (genus) = 2 (mean = 1.6, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 30
maximum number of included terminals = 4, (mean = 2.07 median = 2)
maximum number of subtaxa (genus) = 2 (mean = 1.97, median = 2)
proportion of cladograms that support basic monophyly = 0.97

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 21
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 4 (mean = 2.1, median = 2)
maximum number of subtaxa (genus) = 2 (mean = 2, median = 2)
proportion of cladograms that support rigorous monophyly = 0.95

Cladograms

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

  +-- Aspatharia **
--+             
  +-- Chambardia **

Wu et al., 2021 [Publ 4366], fig. 2_I

  +-- Aspatharia **
--+             
  +-- Chambardia **

Wu et al., 2021 [Publ 4366], fig. 6

  +-- Chambardia **
--+             
  +-- Aspatharia **

Bolotov et al., 2017 [Publ 3761], fig. S2

  +-- Aspatharia **
--+             
  +-- Chambardia **

Bolotov et al., 2016 [Publ 3743], fig. 2

  +-- Aspatharia **
--+             
  +-- Chambardia **

Huang et al., 2019 [Publ 3843], fig. S4

  +-- Aspatharia **
--+             
  +-- Chambardia **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

  +-- Aspatharia **
--+             
  +-- Chambardia **

Bolotov et al., 2016 [Publ 3743], fig. 4

  +-- Chambardia **
--+             
  +-- Aspatharia **

Combosch et al., 2017 [Publ 3806], fig. 1

  +-- Chambardia **
--+             
  +-- Aspatharia **

Bolotov et al., 2017 [Publ 3761], fig. S1

  +-- Chambardia **
--+             
  +-- Aspatharia **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

  +-- Aspatharia **
--+             
  +-- Chambardia **

Sano et al., 2020 [Publ 4138], fig. 3

  +-- Aspatharia **
--+             
  +-- Chambardia **

Chen et al., 2025 [Publ 4455], fig. 1

  +-- Aspatharia **
--+             
  +-- Chambardia **

Wu et al., 2020 [Publ 4121], fig. 2

  +-- Aspatharia **
--+             
  +-- Chambardia **

Hoeh et al., 2009 [Publ 2716], fig. 3

  +-- Chambardia **
  |             
--+-- Aspatharia **
  |             
  +-- Iridininae

Whelan et al., 2011 [Publ 3380], fig. 1-MP

  +-- Aspatharia **
--+             
  +-- Chambardia **

Combosch et al., 2017 [Publ 3806], fig. 2

  +-- Chambardia **
--+             
  +-- Aspatharia **

Huang et al., 2019 [Publ 3843], fig. S3

  +-- Aspatharia **
--+             
  +-- Chambardia **

Huang et al., 2019 [Publ 3843], fig. 2

  +-- Aspatharia **
--+             
  +-- Chambardia **

Graf & Cummings, 2006 [Publ 1968], fig. 2

  +----- Iridininae
--+                
  |  +-- Chambardia **
  +--+             
     +-- Aspatharia **

Hoeh et al., 2009 [Publ 2716], fig. 5

  +----- Iridininae
--+                
  |  +-- Chambardia **
  +--+             
     +-- Aspatharia **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

  +-- Aspatharia **
--+             
  +-- Chambardia **

Sano et al., 2020 [Publ 4138], fig. 2

  +-- Aspatharia **
--+             
  +-- Chambardia **

Huang et al., 2018 [Publ 3828], fig. 3_5

  +-- Aspatharia **
--+             
  +-- Chambardia **

Sano et al., 2020 [Publ 4138], fig. 4

  +-- Aspatharia **
--+             
  +-- Chambardia **

Wu et al., 2020 [Publ 4121], fig. S1

  +-- Aspatharia **
--+             
  +-- Chambardia **

Hoeh et al., 2009 [Publ 2716], fig. 4

  +----- Iridininae
--+                
  |  +-- Chambardia **
  +--+             
     +-- Aspatharia **

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

  +-- Aspatharia **
--+             
  +-- Chambardia **

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

  +-- Iridininae
--+             
  +-- Chambardia **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

  +-- Chambardia **
--+             
  +-- Aspatharia **

Bieler et al., 2014 [Publ 3622], fig. 28

-- Aspatharia_pfeifferiana **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

-- Chambardia_wahlbergi **

Konopleva et al., 2017 [Publ 3807], fig. 6

-- Chambardia_wahlbergi **

Guerra et al., 2019 [Publ 4168], fig. 2

-- Chambardia_rubens **

Graf et al., 2015 [Publ 3619], fig. 1

-- Chambardia_wahlbergi **

González & Giribet, 2015 [Publ 3632], fig. 3

-- Aspatharia_pfeifferiana **

González & Giribet, 2015 [Publ 3632], fig. 6

-- Aspatharia_pfeifferiana **

Walker et al., 2006 [Publ 2666], fig. 6

-- Chambardia_rubens **

González & Giribet, 2015 [Publ 3632], fig. 2

-- Aspatharia_pfeifferiana **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

-- Aspatharia_pfeifferiana **

González & Giribet, 2015 [Publ 3632], fig. 4

-- Aspatharia_pfeifferiana **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

-- Chambardia_rubens **

Elderkin et al., 2016 [Publ 3693], fig. 2

-- Aspatharia_pfeifferiana **

Smith, 2021 [Publ 4331], fig. 1

-- Aspatharia_pfeifferiana **

Guerra et al., 2019 [Publ 4168], fig. 1

-- Chambardia_rubens **

Lemer et al., 2019 [Publ 4093], fig. 1

-- Aspatharia_pfeifferiana **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

-- Aspatharia_pfeifferiana **

Bieler et al., 2014 [Publ 3622], fig. 26_27_29_30

-- Aspatharia_pfeifferiana **