Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini

subtribe CRISTARIINA Lopes-Lima, Bogan & Froufe in Lopes-Lima et al., 2017: cladomics

10 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 39 (43 total)
number of subtaxa (rank: genus) = 10
analyzed in 210 cladograms in 125 publications
maximum number of terminals in an analysis = 33 (mean = 5.17, median = 4)
maximum number of subtaxa (genus) = 9 (mean = 2.8, median = 3)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 153
maximum number of included terminals = 33, (mean = 6.73 median = 4)
maximum number of subtaxa (genus) = 9 (mean = 3.46, median = 3)
proportion of cladograms that support basic monophyly = 0.85

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 94
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 33 (mean = 5.82, median = 4)
maximum number of subtaxa (genus) = 9 (mean = 3.69, median = 3)
proportion of cladograms that support rigorous monophyly = 0.82

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Cristariina cladogram distances
PCoA Ordination

NJ Tree of Cristariina cladogram distances
NJ Tree

Cladograms

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
0.833 33 156 164 9 600 47 2 **
Cladomics cladogram Amuranodonta Anemina Buldowskia Beringiana Pletholophus Mimo Acudonta Sinanodonta Cristaria Lepidodesma Lanceolaria Pseudoacuticosta retiaria 2 Acuticosta chinensis 2 Acuticosta chinensis 1 Pseudoacuticosta retiaria 1 Globunio ACULAMPROTULINA MIDDENDORFFINAIINA UNIONINA OXYNAIINA Pseudodontoideus Lasmigona Pterosyna Arcidens ANODONTINA Utterbackia Pyganodon Utterbackiana Simpsonaias Alasminota Alasmidonta Prolasmidonta Platynaias Anodontoides Strophitus Lamellidens Balwantia Trapezidens Leoparreysia Pseudomulleria Pseudoscabies Indonaia Indochinella Radiatula Scabiellus Unionetta fabagina 2 Harmandia Unionetta fabagina 1 Scabies Assamnaia Parreysia Nitia monceti 2 Nitia teretiuscula Nitia monceti 1 Grandidieria Pseudospatha tanganyicensis 2 Moncetia Pseudospatha tanganyicensis 1 Coelatura leopoldvillensis 2 Coelatura leopoldvillensis 1 Coelatura mesafricana Coelatura aegyptiaca 3 Coelatura hauttecoeuri 2 Nitia acuminata Coelatura aegyptiaca 2 Coelatura alluaudi 2 Coelatura hauttecoeuri 1 Coelatura stuhlmanni 2 Coelatura aegyptiaca 1 Coelatura stuhlmanni 1 Coelatura alluaudi 1 Coelatura bakeri 1 Coelatura bakeri 2 Coelatura horei 3 Coelatura ratidota Coelatura 2 Coelatura horei 1 Coelatura 1 Coelatura horei 2 Coelatura gabonensis Coelatura stanleyvillensis Coelatura kunenensis Nyassunio Coelatura hypsiprymna 2 Coelatura mossambicensis 1 Coelatura hypsiprymna 3 Coelatura mossambicensis 2 Coelatura luapulaensis 2 Coelatura choziensis 2 Coelatura hypsiprymna 1 Coelatura choziensis 1 Coelatura luapulaensis 1 Coelatura briarti Prisodontopsis aviculaeformis 1 Prisodontopsis mweruensis 1 Prisodontopsis aviculaeformis 2 Prisodontopsis aviculaeformis 3 Prisodontopsis mweruensis 2

Chen et al., 2026 [Publ 4805], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 33 33 31 9 1 0 0 153.0
Cladomics cladogram OXYNAIINA Cristaria Beringiana Mimo Acudonta Sinanodonta Pletholophus Anemina Amuranodonta Buldowskia

Wang et al., 2025 [Publ 4488], fig. 3B

cst iT tT bi riS oT roT roS iD
0.950 15 15 13 8 49 11 2 157.0
Cladomics cladogram Anodonta Pseudanodonta ALASMIDONTINA Cristaria Amuranodonta Anemina Buldowskia Beringiana Pletholophus Acudonta Sinanodonta

Wu et al., 2026 [Publ 4811], fig. 4

cst iT tT bi riS oT roT roS iD
0.950 15 15 13 8 80 11 2 157.0
Cladomics cladogram Anodonta Pseudanodonta ALASMIDONTINA Cristaria Amuranodonta Anemina Buldowskia Beringiana Pletholophus Acudonta Sinanodonta

Wu et al., 2026 [Publ 4825], fig. 6B

cst iT tT bi riS oT roT roS iD
0.950 14 14 13 8 51 11 2 159.0
Cladomics cladogram Anodonta Pseudanodonta ALASMIDONTINA Cristaria Amuranodonta Anemina Buldowskia Beringiana Pletholophus Acudonta Sinanodonta

Xiang et al., 2025 [Publ 4433], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 27 27 25 8 1 0 0 155.0
Cladomics cladogram Scabies Pletholophus Beringiana Mimo Sinanodonta Cristaria Anemina Amuranodonta Buldowskia

Wu et al., 2025 [Publ 4442], fig. 2

cst iT tT bi riS oT roT roS iD
0.900 33 33 31 8 2 0 0 159.0
Cladomics cladogram OXYNAIINA Cristaria Beringiana Acudonta Sinanodonta Pletholophus guangzhouensis Pletholophus tenuis Pletholophus reinianus Pletholophus honglinhensis Anemina Amuranodonta Buldowskia

Wu et al., 2025 [Publ 4442], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 15 15 13 8 2 0 0 157.0
Cladomics cladogram UNIONINA Cristaria Amuranodonta Anemina Buldowskia Beringiana Pletholophus Acudonta Sinanodonta

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
0.895 24 24 19 7 98 9 2 151.0
Cladomics cladogram Pseudanodonta ALASMIDONTINA Anodonta cygnea Anodonta anatina Brachyanodon Cristaria Anemina Amuranodonta Buldowskia Pletholophus Beringiana Sinanodonta

Bogan et al., 2023 [Publ 4295], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 27 27 24 7 4 1 1 148.0
Cladomics cladogram ANODONTINA Cristaria Amuranodonta Anemina Buldowskia Pletholophus Beringiana Sinanodonta

Dai et al., 2024 [Publ 4385], fig. 3-BI

cst iT tT bi riS oT roT roS iD
1.000 27 27 24 7 4 0 0 150.0
Cladomics cladogram Trapezidens Scabies Amuranodonta Anemina Buldowskia Cristaria Pletholophus Beringiana Sinanodonta

Dai et al., 2024 [Publ 4385], fig. 3-ML

cst iT tT bi riS oT roT roS iD
1.000 27 27 25 7 4 0 0 151.0
Cladomics cladogram Trapezidens Scabies Amuranodonta Anemina Buldowskia Cristaria Pletholophus Beringiana Sinanodonta

Dai et al., 2026 [Publ 4757], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 30 30 29 7 4 0 0 151.0
Cladomics cladogram Scabies Trapezidens Amuranodonta Anemina Buldowskia Cristaria Pletholophus Beringiana Sinanodonta

Sano et al., 2020 [Publ 4138], fig. 2

cst iT tT bi riS oT roT roS iD
0.875 11 36 36 6 58 8 2 145.0
Cladomics cladogram Lampsilis Sinohyriopsis Inversidens Ptychorhynchus Obovalis Lamprotula Pronodularia Potomida Anemina Buldowskia Pletholophus Beringiana Sinanodonta Cristaria ANODONTINA ALASMIDONTINA Lanceolaria Lepidodesma ACULAMPROTULINA UNIONINA OXYNAIINA

Sano et al., 2020 [Publ 4138], fig. 3

cst iT tT bi riS oT roT roS iD
0.875 11 36 37 6 58 8 2 143.0
Cladomics cladogram Sinohyriopsis Lampsilis Lamprotula Potomida Pronodularia Inversidens Obovalis Ptychorhynchus Anemina Buldowskia Sinanodonta Pletholophus Beringiana Cristaria ANODONTINA ALASMIDONTINA Lanceolaria Lepidodesma ACULAMPROTULINA UNIONINA OXYNAIINA

Sano et al., 2020 [Publ 4138], fig. 4

cst iT tT bi riS oT roT roS iD
0.875 11 36 37 6 58 8 2 141.0
Cladomics cladogram Lampsilis Sinohyriopsis Inversidens Obovalis Ptychorhynchus Lamprotula Pronodularia Potomida Anemina Buldowskia Sinanodonta Pletholophus Beringiana Cristaria ANODONTINA ALASMIDONTINA Lepidodesma Lanceolaria ACULAMPROTULINA UNIONINA OXYNAIINA

Chen et al., 2023 [Publ 4302], fig. 1

cst iT tT bi riS oT roT roS iD
0.857 6 32 24 6 35 4 2 162.0
Cladomics cladogram Coelatura Indonaia Parreysia Anemina Amuranodonta Pletholophus Sinanodonta Beringiana Lanceolaria gladiola Acuticosta Lepidodesma Lanceolaria oxyrhyncha Cristaria ALASMIDONTINA ANODONTINA ACULAMPROTULINA MIDDENDORFFINAIINA Nodularia Cuneopsis Diaurora Schistodesmus Pseudocuneopsis Inversiunio Tchangsinaia UNIONINA

Chen et al., 2023 [Publ 4302], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 6 35 4 2 166.0
Cladomics cladogram ALASMIDONTINA ANODONTINA Cristaria Anemina Amuranodonta Pletholophus Sinanodonta Beringiana

Dai et al., 2025 [Publ 4440], fig. 1

cst iT tT bi riS oT roT roS iD
0.857 6 38 31 6 38 4 2 159.0
Cladomics cladogram Lampsilis QUADRULINA Anemina Amuranodonta Parreysia Coelatura Indonaia Pletholophus Sinanodonta Beringiana Cristaria ALASMIDONTINA ANODONTINA Lanceolaria Lepidodesma Acuticosta GLOBUNIONINA ACULAMPROTULINA MIDDENDORFFINAIINA Nodularia Cuneopsis Diaurora Schistodesmus Pseudocuneopsis UNIONINA Tchangsinaia Inversiunio

Dai et al., 2025 [Publ 4440], fig. 2

cst iT tT bi riS oT roT roS iD
0.857 6 35 29 6 38 4 2 163.0
Cladomics cladogram Coelatura Indonaia Parreysia Anemina Amuranodonta Pletholophus Sinanodonta Beringiana ALASMIDONTINA ANODONTINA Lanceolaria gladiola Lanceolaria oxyrhyncha Cristaria Acuticosta Lepidodesma GLOBUNIONINA ACULAMPROTULINA MIDDENDORFFINAIINA Nodularia Cuneopsis Diaurora Schistodesmus Pseudocuneopsis Inversiunio Tchangsinaia UNIONINA

Chen et al., 2025 [Publ 4487], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 6 35 4 2 166.0
Cladomics cladogram ALASMIDONTINA ANODONTINA Cristaria Anemina Amuranodonta Pletholophus Sinanodonta Beringiana

Chen et al., 2025 [Publ 4487], fig. 3

cst iT tT bi riS oT roT roS iD
0.857 6 9 7 6 35 4 2 165.0
Cladomics cladogram ALASMIDONTINA ANODONTINA Lepidodesma Cristaria Lanceolaria Anemina Amuranodonta Pletholophus Sinanodonta Beringiana

Bolotov et al., 2020 [Publ 4084], fig. 1

cst iT tT bi riS oT roT roS iD
0.909 9 9 7 6 27 3 1 159.0
Cladomics cladogram ANODONTINA Amuranodonta Anemina Buldowskia Cristaria Beringiana Sinanodonta

Konopleva et al., 2023 [Publ 4269], fig. S1_4

cst iT tT bi riS oT roT roS iD
0.909 13 13 11 6 43 3 1 157.0
Cladomics cladogram ANODONTINA Anemina Amuranodonta Buldowskia Cristaria Beringiana Sinanodonta

Sano et al., 2020 [Publ 4138], fig. 1

cst iT tT bi riS oT roT roS iD
0.250 10 22 5 6 24 0 0 175.0
Cladomics cladogram Sinohyriopsis Lamprotula Pronodularia Sinosolenaia Inversidens Obovalis omiensis 1 Obovalis omiensis 2 Ptychorhynchus Lanceolaria oxyrhyncha 1 Lanceolaria oxyrhyncha 2 Cristaria plicata 1 Cristaria plicata 2 Sinanodonta lauta Beringiana japonica Beringiana fukuharai 1 Beringiana beringiana Beringiana fukuharai 2 Beringiana fukuharai 3 Buldowskia iwakawai 1 Buldowskia iwakawai 2 Sinanodonta tumens 1 Sinanodonta tumens 2 Pletholophus Anemina Lanceolaria lanceolata Cristaria plicata 3 Lanceolaria gladiola Sinanodonta angula Lepidodesma ACULAMPROTULINA Acuticosta Cuneopsis Nodularia Schistodesmus Inversiunio

Xiang et al., 2025 [Publ 4433], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 5 6 2 1 163.0
Cladomics cladogram ANODONTINA Cristaria Anemina Beringiana Mimo Sinanodonta

Altaba et al., 2025 [Publ 4849], fig. 10

cst iT tT bi riS oT roT roS iD
0.778 6 6 4 5 6 2 1 164.0
Cladomics cladogram Anodonta anatina Anodonta cygnea Anemina Amuranodonta Cristaria Beringiana Sinanodonta

Sano et al., 2017 [Publ 3799], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 5 13 0 0 166.0
Cladomics cladogram Lanceolaria OXYNAIINA Cristaria Buldowskia Pletholophus Sinanodonta Beringiana

Sano et al., 2017 [Publ 3799], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 9 9 7 5 24 0 0 167.0
Cladomics cladogram Cristaria Buldowskia Beringiana Pletholophus Sinanodonta Tchangsinaia Nodularia Schistodesmus Acuticosta Inversiunio Lepidodesma Lanceolaria

Sano et al., 2022 [Publ 4311], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 12 12 12 5 0 0 0 170.0
Cladomics cladogram Beringiana Sinanodonta Pletholophus Cristaria Buldowskia

Sano et al., 2022 [Publ 4311], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 18 18 16 5 0 0 0 162.0
Cladomics cladogram Beringiana Sinanodonta Pletholophus Cristaria Buldowskia

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
0.938 5 5 3 4 76 9 2 165.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Anemina Pletholophus Sinanodonta

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
0.824 9 9 7 4 192 11 2 159.0
Cladomics cladogram Anemina Cristaria Pletholophus Sinanodonta Alasmidonta Lasmigona Pseudanodonta Simpsonaias Strophitus Pyganodon Utterbackia Anodonta anatina Anodonta cygnea Brachyanodon

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
0.824 9 9 7 4 190 11 2 159.0
Cladomics cladogram Anemina Cristaria Pletholophus Sinanodonta Alasmidonta Lasmigona Anodonta anatina Anodonta cygnea Brachyanodon Pyganodon Utterbackia Pseudanodonta Simpsonaias Strophitus

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.706 9 48 25 4 192 11 2 153.0
Cladomics cladogram Chamberlainia Megalonaias Uniomerus QUADRULINA Sinohyriopsis INDOPSEUDODONTINA PSEUDODONTINA LIMNOSCAPHINA CONTRADENTINA Lamprotula Potomida Pronodularia Sinosolenaia carinata Gonidea Leguminaia Microcondylaea Ptychorhynchus Parvasolenaia Koreosolenaia Sinosolenaia recognita Sinosolenaia iridinea Sinosolenaia oleivora Amblema Toxolasma Obliquaria Plectomerus Elliptoideus Elliptio Eurynia Fusconaia Pleurobema Ellipsaria Potamilus leptodon Medionidus Potamilus alatus Ptychobranchus Truncilla Venustaconcha Cambarunio Leaunio Ortmanniana Lampsilis Acuticosta Alasmidonta Anodonta anatina Anemina Anodonta cygnea Cristaria Brachyanodon Lasmigona Lepidodesma Pseudanodonta Pyganodon Simpsonaias Strophitus Utterbackia Schistodesmus UNIONINA ACULAMPROTULINA Lanceolaria Pletholophus Sinanodonta Cuneopsis celtiformis Tchangsinaia Cuneopsis rufescens Nodularia Cuneopsis szechenyii Cuneopsis heudei

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 4 89 12 2 164.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Anemina Cristaria Beringiana Sinanodonta

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cst iT tT bi riS oT roT roS iD
0.882 6 6 4 4 89 12 2 164.0
Cladomics cladogram Cristaria Anemina Beringiana Sinanodonta Brachyanodon Pseudanodonta Anodonta cygnea ALASMIDONTINA Anodonta exulcerata Anodonta anatina

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cst iT tT bi riS oT roT roS iD
0.941 6 6 4 4 85 12 2 164.0
Cladomics cladogram Cristaria Anemina Beringiana Sinanodonta Pseudanodonta Anodonta Brachyanodon ALASMIDONTINA

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cst iT tT bi riS oT roT roS iD
0.941 6 6 4 4 85 12 2 164.0
Cladomics cladogram Cristaria Anemina Beringiana Sinanodonta Brachyanodon ALASMIDONTINA Pseudanodonta Anodonta

Wu et al., 2024 [Publ 4379], fig. 5

cst iT tT bi riS oT roT roS iD
0.941 6 6 4 4 47 12 2 164.0
Cladomics cladogram Cristaria Anemina Beringiana Sinanodonta Brachyanodon ALASMIDONTINA Pseudanodonta Anodonta

Wu et al., 2024 [Publ 4388], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 49 3 2 166.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Anemina Beringiana Sinanodonta

Hou et al., 2024 [Publ 4400], fig. 4

cst iT tT bi riS oT roT roS iD
0.941 6 6 4 4 69 12 2 164.0
Cladomics cladogram Cristaria Anemina Beringiana Sinanodonta Pseudanodonta Anodonta Brachyanodon ALASMIDONTINA

Hou et al., 2025 [Publ 4445], fig. 5

cst iT tT bi riS oT roT roS iD
0.941 6 6 4 4 46 12 2 164.0
Cladomics cladogram Cristaria Anemina Beringiana Sinanodonta Brachyanodon ALASMIDONTINA Pseudanodonta Anodonta

Froufe et al., 2017 [Publ 3802], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 10 10 7 4 2 2 1 159.0
Cladomics cladogram ANODONTINA Anemina Cristaria Beringiana Sinanodonta

Altaba et al., 2025 [Publ 4849], fig. 11

cst iT tT bi riS oT roT roS iD
0.778 4 4 2 4 10 6 1 166.0
Cladomics cladogram ANODONTINA Cristaria Amuranodonta Sinanodonta Beringiana

Altaba et al., 2025 [Publ 4849], fig. 12

cst iT tT bi riS oT roT roS iD
0.750 7 9 8 4 6 2 1 172.0
Cladomics cladogram Unio mancus Unio pictorum Unio tumidus OXYNAIINA Amuranodonta Cristaria Beringiana Sinanodonta woodiana 2 Sinanodonta woodiana 1 Sinanodonta lauta Sinanodonta tumens ANODONTINA

Lopes-Lima et al., 2020 [Publ 4080], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 9 9 7 4 2 0 0 163.0
Cladomics cladogram Margaritifera Sinanodonta Amuranodonta Anemina Buldowskia

Tumpeesuwan et al., 2026 [Publ 4845], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 4 0 0 0 162.0
Cladomics cladogram Beringiana Sinanodonta Buldowskia Cristaria

Wu et al., 2016 [Publ 3711], fig. 1

cst iT tT bi riS oT roT roS iD
0.900 3 7 5 3 18 4 2 163.0
Cladomics cladogram ACULAMPROTULINA UNIONINA OXYNAIINA Sinanodonta Anemina Cristaria ANODONTINA ALASMIDONTINA

Fonseca et al., 2016 [Publ 3775], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 27 5 2 166.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Anemina Sinanodonta

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
0.867 6 6 4 3 133 9 2 162.0
Cladomics cladogram Cristaria Pletholophus Sinanodonta Anodonta anatina ALASMIDONTINA Pseudanodonta Anodonta cygnea Brachyanodon

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
0.800 6 6 4 3 133 9 2 162.0
Cladomics cladogram Lanceolaria Cristaria Pletholophus Sinanodonta Anodonta anatina Pseudanodonta ALASMIDONTINA Anodonta cygnea Brachyanodon

Bolotov et al., 2017 [Publ 3764], fig. S1

cst iT tT bi riS oT roT roS iD
0.533 6 18 0 3 119 9 2 167.0
Cladomics cladogram Unio crassus Unio pictorum Unio tumidus ACULAMPROTULINA Cuneopsis szechenyii Tchangsinaia Cuneopsis rufescens Nodularia douglasiae Nodularia nuxpersicae Schistodesmus Alasmidonta Anodonta anatina Anodonta cygnea Brachyanodon Lasmigona Pseudanodonta Pyganodon Simpsonaias Strophitus Sinanodonta lauta Cristaria plicata Cristaria radiata Pletholophus Sinanodonta lucida Sinanodonta jourdyi Lanceolaria gladiola Lanceolaria grayii Lanceolaria danzhouensis

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 28 5 2 166.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Anemina Sinanodonta

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 19 6 2 166.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Anemina Sinanodonta

Bolotov et al., 2018 [Publ 3826], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 78 6 2 164.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Pletholophus Sinanodonta

Huang et al., 2018 [Publ 3828], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 38 5 2 166.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Anemina Sinanodonta

Wu et al., 2018 [Publ 3836], fig. 2

cst iT tT bi riS oT roT roS iD
0.833 6 6 3 3 47 7 2 163.0
Cladomics cladogram Anemina Cristaria Sinanodonta Anodonta cygnea Anodonta anatina ALASMIDONTINA

Wang et al., 2018 [Publ 3846], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 27 3 2 166.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Anemina Sinanodonta

Aunins et al., 2018 [Publ 3863], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 36 7 2 166.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Anemina Sinanodonta

Wang et al., 2018 [Publ 3865], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 27 3 2 166.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Anemina Sinanodonta

Cha et al., 2018 [Publ 3916], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 34 5 2 166.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Anemina Sinanodonta

Wu et al., 2018 [Publ 4143], fig. 2

cst iT tT bi riS oT roT roS iD
0.900 3 7 5 3 19 4 2 163.0
Cladomics cladogram ACULAMPROTULINA UNIONINA OXYNAIINA Sinanodonta Anemina Cristaria ANODONTINA ALASMIDONTINA

Huang et al., 2019 [Publ 3843], fig. 3

cst iT tT bi riS oT roT roS iD
0.818 4 4 2 3 52 6 2 166.0
Cladomics cladogram Cristaria Anemina Sinanodonta Anodonta cygnea Anodonta anatina ALASMIDONTINA

Wu et al., 2019 [Publ 3864], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 42 5 2 166.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Anemina Sinanodonta

Froufe et al., 2019 [Publ 3936], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 40 5 2 166.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Anemina Sinanodonta

Froufe et al., 2019 [Publ 3936], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 40 5 2 166.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Anemina Sinanodonta

Riccardi et al., 2020 [Publ 4089], fig. 2

cst iT tT bi riS oT roT roS iD
0.846 4 4 2 3 15 9 2 166.0
Cladomics cladogram Cristaria Anemina Sinanodonta Anodonta anatina Brachyanodon ALASMIDONTINA Pseudanodonta Anodonta exulcerata Anodonta cygnea

Fukata & Iigo, 2020 [Publ 4106], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 50 6 2 166.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Anemina Sinanodonta

Wu et al., 2020 [Publ 4121], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 44 5 2 166.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Anemina Sinanodonta

Wu et al., 2020 [Publ 4147], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 27 4 2 167.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Sinanodonta Anemina

Chen et al., 2020 [Publ 4170], fig. 1

cst iT tT bi riS oT roT roS iD
0.692 3 34 32 3 31 8 2 191.0
Cladomics cladogram Prolasmidonta Alasmidonta Anemina PSEUDODONTINA Lamprotula caveata Chamberlainia OXYNAIINA Lepidodesma ACULAMPROTULINA Platynaias Utterbackia imbecillis Pyganodon Sinanodonta Cristaria ANODONTINA Utterbackia peninsularis Potamilus alatus Lamprotula leaii Sinohyriopsis Parvasolenaia Sinosolenaia Gibbosula QUADRULINA Toxolasma Lampsilis ornata Potamilus leptodon Lampsilis siliquoidea Lampsilis powellii

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 44 5 2 166.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Anemina Sinanodonta

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
0.833 4 4 2 3 87 6 2 164.0
Cladomics cladogram Anemina Cristaria Sinanodonta Lasmigona Anodonta Pseudanodonta Pyganodon Strophitus

Wu et al., 2021 [Publ 4366], fig. 3-BI

cst iT tT bi riS oT roT roS iD
0.846 4 4 2 3 69 9 2 166.0
Cladomics cladogram Cristaria Anemina Sinanodonta Anodonta cygnea Anodonta anatina ALASMIDONTINA

Wu et al., 2021 [Publ 4366], fig. 3-ML

cst iT tT bi riS oT roT roS iD
0.846 4 4 2 3 69 9 2 166.0
Cladomics cladogram Cristaria Anemina Sinanodonta Anodonta cygnea Anodonta anatina ALASMIDONTINA

Wu et al., 2021 [Publ 4366], fig. 4

cst iT tT bi riS oT roT roS iD
0.846 4 4 2 3 64 8 2 166.0
Cladomics cladogram Cristaria Anemina Sinanodonta Anodonta cygnea Anodonta anatina ALASMIDONTINA

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
0.750 4 4 2 3 87 6 2 164.0
Cladomics cladogram Anemina Cristaria Sinanodonta Anodonta cygnea Lasmigona Anodonta anatina Pseudanodonta Pyganodon Strophitus

Wu et al., 2022 [Publ 4238], fig. 2d

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 34 6 2 166.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Anemina Sinanodonta

Wu et al., 2022 [Publ 4238], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 69 6 2 166.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Anemina Sinanodonta

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 3 71 9 2 163.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Pletholophus Sinanodonta

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

cst iT tT bi riS oT roT roS iD
0.933 5 5 3 3 71 9 2 163.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Pletholophus Sinanodonta

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 34 2 2 167.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Sinanodonta Anemina

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 35 2 2 166.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Anemina Sinanodonta

Dai et al., 2023 [Publ 4285], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 50 4 2 167.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Anemina Sinanodonta

Wu et al., 2023 [Publ 4339], fig. 5

cst iT tT bi riS oT roT roS iD
0.900 4 4 2 3 15 5 2 164.0
Cladomics cladogram ALASMIDONTINA ANODONTINA Anemina Cristaria Sinanodonta

Dai et al., 2024 [Publ 4358], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 50 4 2 167.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Anemina Sinanodonta

Dai et al., 2025 [Publ 4440], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 83 6 2 166.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Anemina Sinanodonta

Chen et al., 2025 [Publ 4448], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 3 80 6 2 165.0
Cladomics cladogram ANODONTINA ALASMIDONTINA Cristaria Anemina Sinanodonta

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cst iT tT bi riS oT roT roS iD
0.875 4 4 2 3 136 13 2 166.0
Cladomics cladogram Cristaria Anemina Sinanodonta Anodonta cygnea Anodonta exulcerata Anodonta anatina ALASMIDONTINA Brachyanodon Pseudanodonta

Xue et al., 2016 [Publ 3659], fig. 1

cst iT tT bi riS oT roT roS iD
0.857 4 5 3 3 11 1 1 165.0
Cladomics cladogram Cristaria ALASMIDONTINA Anemina Sinanodonta Lanceolaria ACULAMPROTULINA Schistodesmus Nodularia UNIONINA

Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 3 3 1 167.0
Cladomics cladogram ANODONTINA Cristaria Anemina Sinanodonta

Soroka, 2010 [Publ 3404], fig. 9a

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 3 0 0 0 163.0
Cladomics cladogram Buldowskia Beringiana Sinanodonta

Klishko et al., 2014 [Publ 3666], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 0 0 0 164.0
Cladomics cladogram Cristaria Beringiana Sinanodonta

Klishko et al., 2016 [Publ 3733], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 0 0 0 164.0
Cladomics cladogram Cristaria Sinanodonta Beringiana

Huang et al., 2019 [Publ 3843], fig. 1

cst iT tT bi riS oT roT roS iD
0.600 8 32 30 3 38 0 0 164.0
Cladomics cladogram Sinohyriopsis Anemina Acuticosta Sinanodonta ACULAMPROTULINA Cristaria Lanceolaria Lepidodesma OXYNAIINA

Huang et al., 2019 [Publ 3843], fig. S1

cst iT tT bi riS oT roT roS iD
0.600 8 33 36 3 39 0 0 160.0
Cladomics cladogram Anemina Sinanodonta Acuticosta Lepidodesma Cristaria Lanceolaria ACULAMPROTULINA OXYNAIINA Sinohyriopsis Parvasolenaia rivularis Parvasolenaia triangularis Ptychorhynchus Sinosolenaia carinata Lamprotula Sinosolenaia recognita Sinosolenaia oleivora 2 Sinosolenaia oleivora 1 Sinosolenaia iridinea Sinosolenaia oleivora 3

Kano et al., 2019 [Publ 4194], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 7 0 0 165.0
Cladomics cladogram Sinanodonta Pletholophus Beringiana Lanceolaria OXYNAIINA Inversidens Obovalis Pronodularia

Lopes-Lima et al., 2020 [Publ 4080], fig. 9

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 2 0 0 164.0
Cladomics cladogram Margaritifera Cristaria Sinanodonta Pletholophus

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

cst iT tT bi riS oT roT roS iD
0.600 5 5 5 3 7 0 0 170.0
Cladomics cladogram Nodularia Lanceolaria Inversiunio Cristaria Sinanodonta tumens Sinanodonta lauta 5 Sinanodonta lauta 6 Beringiana japonica 15 Beringiana japonica 16 Beringiana japonica 17 Beringiana japonica 18 Beringiana japonica 19 Beringiana japonica 20 Beringiana japonica 1 Beringiana japonica 2 Beringiana japonica 3 Beringiana japonica 4 Beringiana japonica 5 Sinanodonta lauta 1 Beringiana japonica 6 Sinanodonta lauta 2 Sinanodonta calipygos 1 Beringiana japonica 7 Sinanodonta lauta 4 Beringiana japonica 8 Beringiana japonica 9 Beringiana japonica 10 Beringiana japonica 11 Beringiana japonica 12 Beringiana japonica 13 Beringiana japonica 14 Sinanodonta lauta 3 Sinanodonta calipygos 2

Bogan et al., 2023 [Publ 4295], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 3 1 0 0 162.0
Cladomics cladogram Gibbosula Cristaria Sinanodonta Pletholophus

Bogan et al., 2023 [Publ 4295], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 3 1 0 0 163.0
Cladomics cladogram Gibbosula Cristaria Sinanodonta Pletholophus

Bogan et al., 2023 [Publ 4295], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 12 12 9 3 1 0 0 163.0
Cladomics cladogram Gibbosula Pletholophus Cristaria Sinanodonta

Dai et al., 2024 [Publ 4389], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 43 0 0 167.0
Cladomics cladogram Sinanodonta Anemina Cristaria CTENODESMINA LIMNOSCAPHINA CONTRADENTINA Chamberlainia Sinohyriopsis Lamprotula Pronodularia Potomida Schepmania PSEUDODONTINA Sinosolenaia Gonidea Leguminaia Microcondylaea Pseudopostolata Parvasolenaia Koreosolenaia Postolata Ptychorhynchus Cosmopseudodon Obovalis

Dai et al., 2025 [Publ 4441], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 44 0 0 167.0
Cladomics cladogram Sinanodonta Anemina Cristaria CTENODESMINA LIMNOSCAPHINA CONTRADENTINA Chamberlainia Sinohyriopsis Potomida Pronodularia Lamprotula Schepmania PSEUDODONTINA Sinosolenaia Gonidea Leguminaia Microcondylaea Pseudopostolata Koreosolenaia Parvasolenaia Postolata Guiunio Cosmopseudodon Obovalis Ptychorhynchus

Chen et al., 2025 [Publ 4663], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 45 0 0 167.0
Cladomics cladogram Sinanodonta Anemina Cristaria CTENODESMINA LIMNOSCAPHINA CONTRADENTINA Chamberlainia Sinohyriopsis Potomida Pronodularia Lamprotula Schepmania PSEUDODONTINA Sinosolenaia Gonidea Leguminaia Microcondylaea Pseudopostolata Koreosolenaia Parvasolenaia Postolata Guiunio Cosmopseudodon Obovalis Tenuiunio Ptychorhynchus

Chong et al., 2008 [Publ 2668], fig. 3

cst iT tT bi riS oT roT roS iD
0.706 2 2 0 2 16 15 2 --
Cladomics cladogram Anodonta cygnea Pyganodon Alasminota Anodonta anatina UNIONINA Brachyanodon nuttalliana Brachyanodon californiensis Beringiana Sinanodonta Pseudodontoideus Alasmidonta Pterosyna Pseudanodonta Strophitus Prolasmidonta Platynaias Brachyanodon oregonensis 4 Brachyanodon oregonensis 3 Brachyanodon oregonensis 2 Brachyanodon oregonensis 1 Brachyanodon kennerlyi

Feng et al., 2016 [Publ 3800], fig. 2

cst iT tT bi riS oT roT roS iD
0.778 3 3 1 2 11 3 2 165.0
Cladomics cladogram ALASMIDONTINA Anemina Sinanodonta

Guerra et al., 2017 [Publ 3772], fig. 2a

cst iT tT bi riS oT roT roS iD
0.900 2 8 0 2 23 6 2 160.0
Cladomics cladogram ACULAMPROTULINA UNIONINA Sinanodonta Cristaria ANODONTINA ALASMIDONTINA

Guerra et al., 2017 [Publ 3772], fig. 2b

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 23 6 2 --
Cladomics cladogram Cristaria Sinanodonta ANODONTINA ALASMIDONTINA

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 55 2 2 --
Cladomics cladogram Cristaria Sinanodonta ANODONTINA ALASMIDONTINA

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 55 2 2 --
Cladomics cladogram Cristaria Sinanodonta ANODONTINA ALASMIDONTINA

Whelan et al., 2023 [Publ 4287], fig. 2

cst iT tT bi riS oT roT roS iD
0.913 4 4 2 2 37 37 2 164.0
Cladomics cladogram Anemina Sinanodonta Anodonta cygnea Brachyanodon Pseudanodonta Anodonta anatina ALASMIDONTINA

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

cst iT tT bi riS oT roT roS iD
0.667 4 4 2 2 24 9 2 166.0
Cladomics cladogram Beringiana Sinanodonta Anodonta exulcerata Anodonta cygnea Brachyanodon californiensis Brachyanodon nuttalliana Brachyanodon kennerlyi Brachyanodon oregonensis Anodonta anatina ALASMIDONTINA

Huang et al., 2002 [Publ 2657], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 32 5 1 --
Cladomics cladogram ALASMIDONTINA Sinanodonta Cristaria

Huang et al., 2002 [Publ 2657], fig. 2

cst iT tT bi riS oT roT roS iD
0.800 2 2 0 2 32 5 1 --
Cladomics cladogram Strophitus Schistodesmus Lepidodesma Acuticosta Lanceolaria Sinanodonta Cristaria Nodularia Tchangsinaia Utterbackia Utterbackiana Alasmidonta Lasmigona

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 29 3 1 --
Cladomics cladogram ALASMIDONTINA Cristaria Sinanodonta

Huang et al., 2013 [Publ 3629], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 14 4 1 --
Cladomics cladogram ALASMIDONTINA Sinanodonta Cristaria

Zhou et al., 2016 [Publ 3710], fig. 1

cst iT tT bi riS oT roT roS iD
0.875 2 6 0 2 18 4 1 162.0
Cladomics cladogram Lepidodesma OXYNAIINA UNIONINA Sinanodonta Cristaria ALASMIDONTINA

Wen et al., 2017 [Publ 3795], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 14 1 1 --
Cladomics cladogram ANODONTINA Sinanodonta Cristaria

Riccardi et al., 2020 [Publ 4089], fig. 1

cst iT tT bi riS oT roT roS iD
0.857 2 2 0 2 4 4 1 --
Cladomics cladogram ANODONTINA Sinanodonta Anemina

Pfarrer et al., 2022 [Publ 4305], fig. 2-5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 10 3 1 --
Cladomics cladogram ANODONTINA Cristaria Sinanodonta

Zhang et al., 2004 [Publ 2735], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 12 0 0 --
Cladomics cladogram Sinanodonta Cristaria OXYNAIINA Acuticosta Lanceolaria

Zhou et al., 2007 [Publ 2658], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 19 0 0 --
Cladomics cladogram Sinanodonta Cristaria Sinohyriopsis Lamprotula leaii Lamprotula caveata Gibbosula Lamprotula cornuumlunae ACULAMPROTULINA Tchangsinaia Nodularia Schistodesmus Lanceolaria Acuticosta

Zhou et al., 2007 [Publ 2658], fig. 2

cst iT tT bi riS oT roT roS iD
0.500 2 18 0 2 16 0 0 172.0
Cladomics cladogram Sinanodonta Cristaria Lamprotula QUADRULINA ACULAMPROTULINA Acuticosta OXYNAIINA

Soroka, 2010 [Publ 3404], fig. 9b

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 2 0 0 0 164.0
Cladomics cladogram Buldowskia Sinanodonta

Soroka, 2010 [Publ 3404], fig. 9c

cst iT tT bi riS oT roT roS iD
1.000 4 4 3 2 0 0 0 167.0
Cladomics cladogram Buldowskia Sinanodonta

Ouyang et al., 2011 [Publ 3408], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 10 0 0 165.0
Cladomics cladogram Anemina Sinanodonta OXYNAIINA Lanceolaria grayii Acuticosta Lanceolaria gladiola ACULAMPROTULINA

Zhang et al., 2013 [Publ 4214], fig. 5

cst iT tT bi riS oT roT roS iD
0.500 2 4 0 2 2 0 0 168.0
Cladomics cladogram Cristaria Sinanodonta Margaritifera Lamprotula

Ouyang et al., 2015 [Publ 4493], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 2 27 0 0 164.0
Cladomics cladogram Acuticosta Anemina Sinanodonta

Ouyang et al., 2015 [Publ 4493], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 2 27 0 0 166.0
Cladomics cladogram OXYNAIINA Anemina Sinanodonta

Xia et al., 2016 [Publ 3712], fig. 7B

cst iT tT bi riS oT roT roS iD
0.500 2 3 0 2 1 0 0 167.0
Cladomics cladogram Sinanodonta Cristaria Sinohyriopsis

Sayenko et al., 2017 [Publ 3760], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 2 0 0 0 164.0
Cladomics cladogram Beringiana Sinanodonta

Vikhrev et al., 2017 [Publ 3810], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 2 2 0 0 160.0
Cladomics cladogram Gibbosula Margaritifera Anemina Sinanodonta

Bespalaya et al., 2018 [Publ 3811], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 8 8 7 2 2 0 0 167.0
Cladomics cladogram Gibbosula Margaritifera Anemina Sinanodonta

Kondakov et al., 2018 [Publ 3914], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 2 0 0 0 160.0
Cladomics cladogram Anemina Sinanodonta

Lopes-Lima et al., 2020 [Publ 4080], fig. 10

cst iT tT bi riS oT roT roS iD
1.000 12 12 8 2 2 0 0 160.0
Cladomics cladogram Margaritifera Beringiana Sinanodonta

Lopes-Lima et al., 2020 [Publ 4080], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 2 2 0 0 166.0
Cladomics cladogram Margaritifera Sinanodonta Beringiana

Lopes-Lima et al., 2020 [Publ 4080], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 2 2 0 0 165.0
Cladomics cladogram Margaritifera Sinanodonta Cristaria

Bauer et al., 2021 [Publ 4365], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 8 8 5 2 0 0 0 161.0
Cladomics cladogram Anemina Sinanodonta

Douda et al., 2024 [Publ 4364], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 13 13 12 2 1 0 0 159.0
Cladomics cladogram Margaritifera Beringiana Sinanodonta

Wu et al., 2024 [Publ 4379], fig. 4_6

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 20 0 0 165.0
Cladomics cladogram OXYNAIINA Cristaria Sinanodonta

Kashiri et al., 2024 [Publ 4822], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 2 2 0 0 160.0
Cladomics cladogram Gibbosula Margaritifera Anemina Sinanodonta

Itoh et al., 2026 [Publ 4842], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 2 0 0 0 163.0
Cladomics cladogram Sinanodonta Beringiana

Soroka, 2008 [Publ 3695], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 2 2 --
Cladomics cladogram Sinanodonta ANODONTINA ALASMIDONTINA

Soroka, 2008 [Publ 3695], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 2 2 --
Cladomics cladogram Sinanodonta ANODONTINA ALASMIDONTINA

Plazzi et al., 2011 [Publ 3676], fig. 5

cst iT tT bi riS oT roT roS iD
0.750 1 1 0 1 17 4 2 --
Cladomics cladogram Cristaria Lanceolaria Anodonta UNIONINA ALASMIDONTINA Pseudanodonta Alathyria Pronodularia Sinohyriopsis QUADRULINA Parvaspina Lampsilis Epioblasma Venustaconcha

Huang et al., 2018 [Publ 3828], fig. 6-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 16 3 2 --
Cladomics cladogram Sinanodonta ANODONTINA ALASMIDONTINA

Chase et al., 2018 [Publ 3847], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 38 13 2 --
Cladomics cladogram Sinanodonta ANODONTINA ALASMIDONTINA

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

cst iT tT bi riS oT roT roS iD
0.500 1 1 0 1 104 19 2 --
Cladomics cladogram Alasmidonta marginata Prolasmidonta Alasmidonta undulata Beringiana Brachyanodon kennerlyi Brachyanodon nuttalliana Brachyanodon oregonensis Arcidens confragosus Arcidens wheeleri Pterosyna Platynaias compressa Lasmigona Platynaias decorata Platynaias subviridis Pyganodon grandis Pyganodon lacustris Pseudodontoideus Strophitus Utterbackia imbecillis Utterbackia peninsularis 1 Utterbackia peninsularis 2

Burzynski & Soroka, 2018 [Publ 3862], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 3 2 --
Cladomics cladogram Sinanodonta ANODONTINA ALASMIDONTINA

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 3 2 --
Cladomics cladogram Sinanodonta ANODONTINA ALASMIDONTINA

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 3 2 --
Cladomics cladogram Sinanodonta ANODONTINA ALASMIDONTINA

Huang et al., 2019 [Publ 3843], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 3 2 --
Cladomics cladogram Sinanodonta ANODONTINA ALASMIDONTINA

Froufe et al., 2019 [Publ 3936], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 18 3 2 --
Cladomics cladogram Sinanodonta ANODONTINA ALASMIDONTINA

Froufe et al., 2019 [Publ 3936], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 18 3 2 --
Cladomics cladogram Sinanodonta ANODONTINA ALASMIDONTINA

Guerra et al., 2019 [Publ 4168], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 6 2 --
Cladomics cladogram Sinanodonta ANODONTINA ALASMIDONTINA

Guerra et al., 2019 [Publ 4168], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 3 2 --
Cladomics cladogram Sinanodonta ANODONTINA ALASMIDONTINA

Guerra et al., 2019 [Publ 4168], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 6 2 --
Cladomics cladogram Sinanodonta ANODONTINA ALASMIDONTINA

Guerra et al., 2019 [Publ 4168], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 3 2 --
Cladomics cladogram ALASMIDONTINA ANODONTINA Sinanodonta

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 3 2 --
Cladomics cladogram Sinanodonta ANODONTINA ALASMIDONTINA

Wu et al., 2022 [Publ 4238], fig. 2e

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 3 2 --
Cladomics cladogram Sinanodonta ANODONTINA ALASMIDONTINA

Klymus et al., 2025 [Publ 4814], fig. 3A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 3 2 --
Cladomics cladogram Sinanodonta ANODONTINA ALASMIDONTINA

Klymus et al., 2025 [Publ 4814], fig. 3B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 3 2 --
Cladomics cladogram Sinanodonta ANODONTINA ALASMIDONTINA

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 46 3 2 --
Cladomics cladogram Sinanodonta ANODONTINA ALASMIDONTINA

Nagel et al., 1996 [Publ 3400], fig. 351

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 4 4 1 --
Cladomics cladogram Pseudanodonta Anodonta cygnea Sinanodonta Anodonta 2 Anodonta anatina Anodonta 1

Nagel et al., 1998 [Publ 1946], fig. 7

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 4 4 1 --
Cladomics cladogram Pseudanodonta Anodonta cygnea Sinanodonta Anodonta 2 Anodonta anatina Anodonta 1

Mock et al., 2004 [Publ 2686], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 1 1 1 --
Cladomics cladogram ANODONTINA Sinanodonta

Mock et al., 2004 [Publ 2686], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 1 1 1 --
Cladomics cladogram ANODONTINA Sinanodonta

Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 4 2 1 --
Cladomics cladogram Beringiana ANODONTINA

Soroka, 2010 [Publ 3404], fig. 2

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 6 3 1 --
Cladomics cladogram Sinanodonta UNIONINA ANODONTINA

Soroka, 2010 [Publ 3404], fig. 2-M

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 4 1 1 --
Cladomics cladogram Sinanodonta ANODONTINA UNIONINA

Soroka, 2010 [Publ 3404], fig. 3a

cst iT tT bi riS oT roT roS iD
0.667 2 2 0 1 7 3 1 --
Cladomics cladogram Sinanodonta ANODONTINA UNIONINA

Soroka, 2010 [Publ 3404], fig. 3a-M

cst iT tT bi riS oT roT roS iD
0.800 2 2 0 1 4 1 1 --
Cladomics cladogram Sinanodonta UNIONINA

Soroka, 2010 [Publ 3404], fig. 3b

cst iT tT bi riS oT roT roS iD
0.500 2 2 0 1 7 3 1 --
Cladomics cladogram Sinanodonta UNIONINA

Soroka, 2010 [Publ 3404], fig. 3b-M

cst iT tT bi riS oT roT roS iD
0.800 2 2 0 1 4 1 1 --
Cladomics cladogram Sinanodonta UNIONINA

Soroka, 2010 [Publ 3404], fig. 4

cst iT tT bi riS oT roT roS iD
0.500 1 1 0 1 6 3 1 --
Cladomics cladogram Sinanodonta ANODONTINA

Soroka, 2010 [Publ 3404], fig. 4-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 1 1 1 --
Cladomics cladogram ANODONTINA Sinanodonta

Soroka, 2010 [Publ 3404], fig. 5a

cst iT tT bi riS oT roT roS iD
0.667 2 2 0 1 6 3 1 --
Cladomics cladogram Anodonta anatina Sinanodonta Anodonta cygnea Pseudanodonta

Soroka, 2010 [Publ 3404], fig. 5a-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 1 1 1 --
Cladomics cladogram ANODONTINA Sinanodonta

Soroka, 2010 [Publ 3404], fig. 5b

cst iT tT bi riS oT roT roS iD
0.500 2 2 0 1 6 3 1 --
Cladomics cladogram ANODONTINA Sinanodonta UNIONINA

Soroka, 2010 [Publ 3404], fig. 5b-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 1 1 1 --
Cladomics cladogram ANODONTINA Sinanodonta

Soroka, 2010 [Publ 3404], fig. 8a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 2 1 --
Cladomics cladogram Sinanodonta ANODONTINA

Soroka, 2010 [Publ 3404], fig. 8b

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 5 2 1 --
Cladomics cladogram Sinanodonta ANODONTINA UNIONINA

Breton et al., 2011 [Publ 3376], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 37 11 1 --
Cladomics cladogram Cristaria ALASMIDONTINA

Chen et al., 2012 [Publ 3624], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 6 1 1 --
Cladomics cladogram ALASMIDONTINA Cristaria

Zieritz et al., 2012 [Publ 3673], fig. 4

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 7 3 1 --
Cladomics cladogram Sinanodonta Margaritifera UNIONINA

Sheng et al., 2014 [Publ 3704], fig. 2

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 7 1 1 --
Cladomics cladogram Cristaria ALASMIDONTINA Lampsilis Venustaconcha QUADRULINA Lamprotula Sinohyriopsis

Mezhzherin et al., 2014 [Publ 3705], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 3 1 --
Cladomics cladogram Sinanodonta ANODONTINA

Mezhzherin et al., 2014 [Publ 3705], fig. 4

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 3 3 1 --
Cladomics cladogram Sinanodonta ANODONTINA

Wei et al., 2016 [Publ 3665], fig. 0

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 5 1 1 --
Cladomics cladogram Cristaria Lampsilis ALASMIDONTINA QUADRULINA

Amaro et al., 2017 [Publ 3791], fig. 2-M

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 9 3 1 --
Cladomics cladogram Pyganodon Lasmigona Sinanodonta Utterbackia

Froufe et al., 2017 [Publ 3802], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 3 1 --
Cladomics cladogram Cristaria ANODONTINA

Froufe et al., 2017 [Publ 3802], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 3 1 --
Cladomics cladogram Sinanodonta ANODONTINA

Smith et al., 2018 [Publ 3790], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 17 1 --
Cladomics cladogram Cristaria ALASMIDONTINA

Tomilova et al., 2020 [Publ 4097], fig. 3

cst iT tT bi riS oT roT roS iD
0.667 2 2 0 1 6 4 1 --
Cladomics cladogram UNIONINA Sinanodonta ANODONTINA

Smith, 2021 [Publ 4331], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 2 1 --
Cladomics cladogram Cristaria ALASMIDONTINA

Dai et al., 2024 [Publ 4386], fig. 5-BI

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 21 1 1 --
Cladomics cladogram Acuticosta Anemina ANODONTINA Lanceolaria ACULAMPROTULINA Lepidodesma UNIONINA MIDDENDORFFINAIINA OXYNAIINA

Dai et al., 2024 [Publ 4386], fig. 5-ML

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 21 1 1 --
Cladomics cladogram Acuticosta Anemina

Dai et al., 2024 [Publ 4386], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 2 1 --
Cladomics cladogram Sinanodonta ALASMIDONTINA

Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 0 0 --
Cladomics cladogram Sinanodonta Lampsilis Sinohyriopsis Pronodularia

Soroka, 2010 [Publ 3404], fig. 8a-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 2 0 0 --
Cladomics cladogram Sinanodonta UNIONINA

Soroka, 2010 [Publ 3404], fig. 8b-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 2 0 0 --
Cladomics cladogram Sinanodonta UNIONINA

Jiang et al., 2010 [Publ 3438], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 0 0 --
Cladomics cladogram Lampsilis Cristaria Pronodularia Sinohyriopsis

Peng et al., 2012 [Publ 3689], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 2 0 0 --
Cladomics cladogram Cristaria Sinohyriopsis

Bolotov et al., 2016 [Publ 3732], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 6 6 5 1 2 0 0 167.0
Cladomics cladogram Sinanodonta Gibbosula Margaritifera

Zhou et al., 2017 [Publ 3788], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 15 0 0 --
Cladomics cladogram Anemina Lanceolaria OXYNAIINA ACULAMPROTULINA QUADRULINA Potamilus Lampsilis Sinohyriopsis Lamprotula Ptychorhynchus Sinosolenaia

Liu et al., 2017 [Publ 3939], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram Anemina Sinosolenaia

Bulakhova et al., 2023 [Publ 4325], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 0 0 0 166.0
Cladomics cladogram Beringiana

Barraza et al., 2026 [Publ 4851], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 18 0 0 --
Cladomics cladogram Sinanodonta Amblema Psoronaias Nephronaias Reginaia QUADRULINA Eurynia Mesonaias Pachynaias Cyrtonaias Popenaias