MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Mon 29 June 2026
Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini
subtribe CRISTARIINA Lopes-Lima, Bogan & Froufe in Lopes-Lima et al., 2017: cladomics
Cladomics pages are experimental — even more so than the rest of this site!
The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.
The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.
Following the summary, the available published clades representing the taxon are listed.
For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!
Cladomics Summary
number of species = 38 (41 total)
number of subtaxa (rank: genus) = 10
analyzed in 205 cladograms in 122 publications
maximum number of terminals in an analysis = 33 (mean = 5.16, median = 4)
maximum number of subtaxa (genus) = 9 (mean = 2.78, median = 2)
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 149
maximum number of included terminals = 33, (mean = 6.72 median = 4)
maximum number of subtaxa (genus) = 9 (mean = 3.44, median = 3)
proportion of cladograms that support basic monophyly = 0.85
Rigorous Monophyly
number of cladograms that test rigorous monophyly = 91
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 33 (mean = 5.82, median = 4)
maximum number of subtaxa (genus) = 9 (mean = 3.67, median = 3)
proportion of cladograms that support rigorous monophyly = 0.82
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Cladograms
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14
** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.833 | 33 | 156 | 164 | 9 | 600 | 47 | 2 | ** |
+----- Amuranodonta **
+-----------------------------+
| | +-- Anemina **
| +--+
| +-- Buldowskia **
|
| +-- Beringiana **
| +-----+
+-----------------------------+ | +-- Pletholophus **
| | +-----------------------+
| | | | +----- Mimo **
| | | +--+
| | | | +-- Acudonta **
| | | +--+
| | | +-- Sinanodonta **
| | |
| +--+ +----------------------------- Cristaria **
| | |
| | | +----------------- Lepidodesma
| | | |
| | | | +----------- Lanceolaria
| | | | +--+
| | | +--------+ | | +-------- Pseudoacuticosta_retiaria__2
| | | | | | +--+
| +--+ | | | | +----- Acuticosta_chinensis__2
| | | | | +--+
| | | +--+ | +-- Acuticosta_chinensis__1
| | | | +--+
| | | | +-- Pseudoacuticosta_retiaria__1
| | | |
| | | | +----------- Globunio
| | | +--+
| | | | +-------- Aculamprotulina
| +--+ +--+
| | | +----- Middendorffinaiina
| | +--+
| | | +-- Unionina
| | +--+
| | +-- Oxynaiina
| |
--+ | +-------- Pseudodontoideus
| | +--------------+
| | | | +----- Lasmigona
| | | +--+
| | | | +-- Pterosyna
| +--+ +--+
| | +-- Arcidens
| |
| | +-------------------- Anodontina
| | |
| +--+ +-- Utterbackia
| | +--------------+
| | | +-- Pyganodon
| +--+
| | +-------------- Utterbackiana
| | |
| +--+ +----------- Simpsonaias
| | |
| | | +-- Alasminota
| +--+ +-----+
| | | +-- Alasmidonta
| | |
| +--+ +-- Prolasmidonta
| | +--+
| | | +-- Platynaias
| +--+
| | +-- Anodontoides
| +--+
| +-- Strophitus
|
| +-------------------------------------------------------------- Lamellidens
| |
| | +-- Balwantia
+--+ +--------------------------------------------------------+
| | +-- Trapezidens
| |
+--+ +-------------------------------------------------------- Leoparreysia
| |
| | +----------------------------------------------------- Pseudomulleria
| | |
+--+ | +-- Pseudoscabies
| | +--------------+
| | | +-- Indonaia
| | +--------------------------------+
| | | | +-------------- Indochinella
+--+ | | |
| | +--+ +----------- Radiatula
| | | |
| | +--+ +-- Scabiellus
| | | +-----+
| | | | +-- Unionetta_fabagina__2
| | +--+
| | | +----- Harmandia
+--+ +--+
| | +-- Unionetta_fabagina__1
| +--+
| +-- Scabies
|
| +-- Assamnaia
| +--------------------------------------------+
| | +-- Parreysia
| |
| | +----- Nitia_monceti__2
+--+ +--------------------------------------+
| | | +-- Nitia_teretiuscula
| | +--+
| | +-- Nitia_monceti__1
+--+
| +----------------------------------------- Grandidieria
| |
| | +----- Pseudospatha_tanganyicensis__2
| | +--------------------------------+
+--+ | | +-- Moncetia
| | +--+
| | +-- Pseudospatha_tanganyicensis__1
| |
| | +----- Coelatura_leopoldvillensis__2
+--+ +-----------------------------+
| | | +-- Coelatura_leopoldvillensis__1
| | +--+
| | +-- Coelatura_mesafricana
| |
| | +-------------------------------- Coelatura_aegyptiaca__3
| | |
+--+ | +-------- Coelatura_hauttecoeuri__2
| | |
| | +-------- Nitia_acuminata
| | |
| | | +----- Coelatura_aegyptiaca__2
| | +--------------------+ |
| | | | +----- Coelatura_alluaudi__2
+--+ | | |
| | | +----- Coelatura_hauttecoeuri__1
| | | |
| | +--+----- Coelatura_stuhlmanni__2
| | |
| | | +-- Coelatura_aegyptiaca__1
| | | |
| | | +-- Coelatura_stuhlmanni__1
| | | |
| | +--+-- Coelatura_alluaudi__1
+--+ |
| +-- Coelatura_bakeri__1
| |
| +-- Coelatura_bakeri__2
|
| +----------- Coelatura_horei__3
| |
| +--------------+ +-------- Coelatura_ratidota
| | | |
| | +--+ +----- Coelatura__2
| | | |
| | +--+ +-- Coelatura_horei__1
| | | |
+--+ +--+-- Coelatura__1
| |
| +-- Coelatura_horei__2
|
| +----------------------- Coelatura_gabonensis
| |
+--+ +-------------------- Coelatura_stanleyvillensis
| |
| | +----------------- Coelatura_kunenensis
+--+ |
| | +-------------- Nyassunio
| | |
+--+ | +----------- Coelatura_hypsiprymna__2
| | |
| | +----------- Coelatura_mossambicensis__1
| | |
+--+ | +-- Coelatura_hypsiprymna__3
| +--------+
| | +-- Coelatura_mossambicensis__2
| |
| | +----- Coelatura_luapulaensis__2
| | |
| | +----- Coelatura_choziensis__2
+--+-----+
| +----- Coelatura_hypsiprymna__1
| |
| | +-- Coelatura_choziensis__1
| +--+
| +-- Coelatura_luapulaensis__1
|
| +----- Coelatura_briarti
| +--+
| | | +-- Prisodontopsis_aviculaeformis__1
| | +--+
+--+ +-- Prisodontopsis_mweruensis__1
|
| +-- Prisodontopsis_aviculaeformis__2
| |
+-----+-- Prisodontopsis_aviculaeformis__3
|
+-- Prisodontopsis_mweruensis__2
Chen et al., 2026 [Publ 4805], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 33 | 33 | 31 | 9 | 1 | 0 | 0 | 153.0 |
+-------------------- Oxynaiina | --+ +----------------- Cristaria ** | | +--+ +-------------- Beringiana ** | | | | +----- Mimo ** +--+ +-----+ | | | +-- Acudonta ** | | +--+ +--+ +-- Sinanodonta ** | | +-------- Pletholophus ** +--+ | +----- Anemina ** +--+ | +-- Amuranodonta ** +--+ +-- Buldowskia **
Wang et al., 2025 [Publ 4488], fig. 3B
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.950 | 15 | 15 | 13 | 8 | 49 | 11 | 2 | 157.0 |
+----- Anodonta
+--------+
| | +-- Pseudanodonta
| +--+
| +-- Alasmidontina
--+
| +----------- Cristaria **
| |
| | +----- Amuranodonta **
+--+ +--+
| | | +-- Anemina **
| | +--+
| | +-- Buldowskia **
+--+
| +-- Beringiana **
| +--+
| | +-- Pletholophus **
+--+
| +-- Acudonta **
+--+
+-- Sinanodonta **
Wu et al., 2026 [Publ 4811], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.950 | 15 | 15 | 13 | 8 | 80 | 11 | 2 | 157.0 |
+----- Anodonta
+--------+
| | +-- Pseudanodonta
| +--+
| +-- Alasmidontina
--+
| +----------- Cristaria **
| |
| | +----- Amuranodonta **
+--+ +--+
| | | +-- Anemina **
| | +--+
| | +-- Buldowskia **
+--+
| +-- Beringiana **
| +--+
| | +-- Pletholophus **
+--+
| +-- Acudonta **
+--+
+-- Sinanodonta **
Wu et al., 2026 [Publ 4825], fig. 6B
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.950 | 14 | 14 | 13 | 8 | 51 | 11 | 2 | 159.0 |
+----- Anodonta
+--------+
| | +-- Pseudanodonta
| +--+
| +-- Alasmidontina
--+
| +----------- Cristaria **
| |
| | +----- Amuranodonta **
+--+ +--+
| | | +-- Anemina **
| | +--+
| | +-- Buldowskia **
+--+
| +-- Beringiana **
| +--+
| | +-- Pletholophus **
+--+
| +-- Acudonta **
+--+
+-- Sinanodonta **
Xiang et al., 2025 [Publ 4433], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 27 | 27 | 25 | 8 | 1 | 0 | 0 | 155.0 |
+----------------- Scabies | --+ +-------------- Pletholophus ** | | | | +----- Beringiana ** +--+ +-----+ | | | +-- Mimo ** | | +--+ +--+ +-- Sinanodonta ** | | +-------- Cristaria ** +--+ | +----- Anemina ** +--+ | +-- Amuranodonta ** +--+ +-- Buldowskia **
Wu et al., 2025 [Publ 4442], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.900 | 33 | 33 | 31 | 8 | 2 | 0 | 0 | 159.0 |
+-------------------- Oxynaiina | --+ +----------------- Cristaria ** | | | | +----- Beringiana ** +--+ +--------+ | | | +-- Acudonta ** | | +--+ | | +-- Sinanodonta ** +--+ | +----- Pletholophus_guangzhouensis ** | +-----+ | | | +-- Pletholophus_tenuis ** | | +--+ +--+ +-- Pletholophus_reinianus ** | | +-------- Pletholophus_honglinhensis ** +--+ | +----- Anemina ** +--+ | +-- Amuranodonta ** +--+ +-- Buldowskia **
Wu et al., 2025 [Publ 4442], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 15 | 15 | 13 | 8 | 2 | 0 | 0 | 157.0 |
+-------------- Unionina | --+ +----------- Cristaria ** | | | | +----- Amuranodonta ** +--+ +--+ | | | +-- Anemina ** | | +--+ | | +-- Buldowskia ** +--+ | +-- Beringiana ** | +--+ | | +-- Pletholophus ** +--+ | +-- Acudonta ** +--+ +-- Sinanodonta **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.895 | 24 | 24 | 19 | 7 | 98 | 9 | 2 | 151.0 |
+-------- Pseudanodonta
|
+--+-------- Alasmidontina
| |
| | +----- Anodonta_cygnea
| +--+
| | +-- Anodonta_anatina
| +--+
--+ +-- Brachyanodon
|
| +-------- Cristaria **
| |
| | +----- Anemina **
| +--+
+--+ | +-- Amuranodonta **
| +--+
| +-- Buldowskia **
|
| +----- Pletholophus **
+--+
| +-- Beringiana **
+--+
+-- Sinanodonta **
Bogan et al., 2023 [Publ 4295], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 27 | 27 | 24 | 7 | 4 | 1 | 1 | 148.0 |
+-------------- Anodontina | --+ +----------- Cristaria ** | | | | +----- Amuranodonta ** +--+ +--+ | | | +-- Anemina ** | | +--+ +--+ +-- Buldowskia ** | | +----- Pletholophus ** +--+ | +-- Beringiana ** +--+ +-- Sinanodonta **
Dai et al., 2024 [Publ 4385], fig. 3-BI
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 27 | 27 | 24 | 7 | 4 | 0 | 0 | 150.0 |
+-- Trapezidens
+-----------+
| +-- Scabies
|
--+ +----- Amuranodonta **
| +-----+
| | | +-- Anemina **
| | +--+
+--+ +-- Buldowskia **
|
| +-------- Cristaria **
+--+
| +----- Pletholophus **
+--+
| +-- Beringiana **
+--+
+-- Sinanodonta **
Dai et al., 2024 [Publ 4385], fig. 3-ML
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 27 | 27 | 25 | 7 | 4 | 0 | 0 | 151.0 |
+-- Trapezidens
+-----------+
| +-- Scabies
|
--+ +----- Amuranodonta **
| +-----+
| | | +-- Anemina **
| | +--+
+--+ +-- Buldowskia **
|
| +-------- Cristaria **
+--+
| +----- Pletholophus **
+--+
| +-- Beringiana **
+--+
+-- Sinanodonta **
Dai et al., 2026 [Publ 4757], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 30 | 30 | 29 | 7 | 4 | 0 | 0 | 151.0 |
+-- Scabies
+-----------+
| +-- Trapezidens
|
--+ +----- Amuranodonta **
| +-----+
| | | +-- Anemina **
| | +--+
+--+ +-- Buldowskia **
|
| +-------- Cristaria **
+--+
| +----- Pletholophus **
+--+
| +-- Beringiana **
+--+
+-- Sinanodonta **
Sano et al., 2020 [Publ 4138], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.875 | 11 | 36 | 36 | 6 | 58 | 8 | 2 | 145.0 |
+----------------- Lampsilis
|
+--------+ +-------------- Sinohyriopsis
| | |
| +--+ +----------- Inversidens
| | |
| +--+ +-- Ptychorhynchus
| | +-----+
| | | +-- Obovalis
| +--+
| | +----- Lamprotula
--+ +--+
| | +-- Pronodularia
| +--+
| +-- Potomida
|
| +-- Anemina **
| +--------------------+
| | +-- Buldowskia **
| |
+--+ +----- Pletholophus **
| +--------------+
| | | +-- Beringiana **
| | +--+
+--+ +-- Sinanodonta **
|
| +----------------- Cristaria **
| |
+--+ +-- Anodontina
| +-----------+
| | +-- Alasmidontina
+--+
| +----------- Lanceolaria
+--+
| +-------- Lepidodesma
+--+
| +----- Aculamprotulina
+--+
| +-- Unionina
+--+
+-- Oxynaiina
Sano et al., 2020 [Publ 4138], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.875 | 11 | 36 | 37 | 6 | 58 | 8 | 2 | 143.0 |
+-------------- Sinohyriopsis
|
+-----------+ +----------- Lampsilis
| | |
| | | +----- Lamprotula
| +--+ +--+
| | | | +-- Potomida
| | | +--+
| +--+ +-- Pronodularia
| |
| | +----- Inversidens
--+ +--+
| | +-- Obovalis
| +--+
| +-- Ptychorhynchus
|
| +-- Anemina **
| +--------------------+
| | +-- Buldowskia **
| |
+--+ +----- Sinanodonta **
| +--------------+
| | | +-- Pletholophus **
| | +--+
+--+ +-- Beringiana **
|
| +----------------- Cristaria **
| |
+--+ +-- Anodontina
| +-----------+
| | +-- Alasmidontina
+--+
| +----------- Lanceolaria
+--+
| +-------- Lepidodesma
+--+
| +----- Aculamprotulina
+--+
| +-- Unionina
+--+
+-- Oxynaiina
Sano et al., 2020 [Publ 4138], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.875 | 11 | 36 | 37 | 6 | 58 | 8 | 2 | 141.0 |
+----------------- Lampsilis
|
+--------+ +-------------- Sinohyriopsis
| | |
| +--+ +----------- Inversidens
| | |
| +--+ +-- Obovalis
| | +-----+
| | | +-- Ptychorhynchus
| +--+
| | +----- Lamprotula
--+ +--+
| | +-- Pronodularia
| +--+
| +-- Potomida
|
| +-- Anemina **
| +--------------------+
| | +-- Buldowskia **
| |
+--+ +----- Sinanodonta **
| +--------------+
| | | +-- Pletholophus **
| | +--+
+--+ +-- Beringiana **
|
| +----------------- Cristaria **
| |
+--+ +-- Anodontina
| +-----------+
| | +-- Alasmidontina
+--+
| +----------- Lepidodesma
+--+
| +-------- Lanceolaria
+--+
| +----- Aculamprotulina
+--+
| +-- Unionina
+--+
+-- Oxynaiina
Chen et al., 2023 [Publ 4302], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.857 | 6 | 32 | 24 | 6 | 35 | 4 | 2 | 162.0 |
+-- Coelatura
|
+--------------------------+-- Indonaia
| |
| +-- Parreysia
|
| +-- Anemina **
--+ +-----------------------+
| | +-- Amuranodonta **
| |
| | +----- Pletholophus **
| | +-----------------+
+--+ | | +-- Sinanodonta **
| | +--+
| | +-- Beringiana **
| |
| | +-------------------- Lanceolaria_gladiola
+--+ |
| +-------------------- Acuticosta
| |
| +-------------------- Lepidodesma
| |
| +-------------------- Lanceolaria_oxyrhyncha
| |
+--+-------------------- Cristaria **
|
| +-- Alasmidontina
+-----------------+
| +-- Anodontina
|
| +----------------- Aculamprotulina
| |
| | +-- Middendorffinaiina
+--+ +-----------+
| | +-- Nodularia
| |
+--+ +----------- Cuneopsis
| |
| | +-- Diaurora
+--+--------+
| +-- Schistodesmus
|
| +-------- Pseudocuneopsis
+--+
| +----- Inversiunio
+--+
| +-- Tchangsinaia
+--+
+-- Unionina
Chen et al., 2023 [Publ 4302], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 6 | 6 | 4 | 6 | 35 | 4 | 2 | 166.0 |
+-- Alasmidontina
+-----------+
| +-- Anodontina
--+
| +----------- Cristaria **
| |
+--+ +-- Anemina **
| +-----+
| | +-- Amuranodonta **
+--+
| +----- Pletholophus **
+--+
| +-- Sinanodonta **
+--+
+-- Beringiana **
Dai et al., 2025 [Publ 4440], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.857 | 6 | 38 | 31 | 6 | 38 | 4 | 2 | 159.0 |
+-- Lampsilis
+-----------------------------------------+
| +-- Quadrulina
|
| +-- Anemina **
| +--------------------------------------+
--+ | +-- Amuranodonta **
| |
| | +----- Parreysia
| +-----------------------------------+
| | | +-- Coelatura
| | +--+
+--+ +-- Indonaia
|
| +----- Pletholophus **
| +--------------------------------+
| | | +-- Sinanodonta **
| | +--+
| | +-- Beringiana **
+--+
| +----------------------------------- Cristaria **
| |
| | +-- Alasmidontina
| +--------------------------------+
+--+ +-- Anodontina
|
| +-------------------------------- Lanceolaria
| |
+--+ +----------------------------- Lepidodesma
| |
+--+ +-------------------------- Acuticosta
| |
+--+ +----------------------- Globunionina
| |
+--+ +-------------------- Aculamprotulina
| |
+--+ +----------------- Middendorffinaiina
| |
+--+ +-------------- Nodularia
| |
+--+ +----------- Cuneopsis
| |
| | +-- Diaurora
+--+--------+
| +-- Schistodesmus
|
| +-------- Pseudocuneopsis
+--+
| +----- Unionina
+--+
| +-- Tchangsinaia
+--+
+-- Inversiunio
Dai et al., 2025 [Publ 4440], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.857 | 6 | 35 | 29 | 6 | 38 | 4 | 2 | 163.0 |
+-- Coelatura
|
+-----------------------------------+-- Indonaia
| |
| +-- Parreysia
|
--+ +-- Anemina **
| +--------------------------------+
| | +-- Amuranodonta **
| |
| | +----- Pletholophus **
+--+ +--------------------------+
| | | +-- Sinanodonta **
| | +--+
| | +-- Beringiana **
+--+
| +-- Alasmidontina
| +--------------------------+
| | +-- Anodontina
| |
+--+ +-------------------------- Lanceolaria_gladiola
| |
| +-------------------------- Lanceolaria_oxyrhyncha
| |
+--+-------------------------- Cristaria **
|
| +----------------------- Acuticosta
| |
| +----------------------- Lepidodesma
+--+
| +-------------------- Globunionina
| |
+--+ +----------------- Aculamprotulina
| |
| | +-- Middendorffinaiina
+--+ +-----------+
| | +-- Nodularia
| |
+--+ +----------- Cuneopsis
| |
| | +-- Diaurora
+--+--------+
| +-- Schistodesmus
|
| +-------- Pseudocuneopsis
+--+
| +----- Inversiunio
+--+
| +-- Tchangsinaia
+--+
+-- Unionina
Chen et al., 2025 [Publ 4487], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 6 | 6 | 4 | 6 | 35 | 4 | 2 | 166.0 |
+-- Alasmidontina
+-----------+
| +-- Anodontina
--+
| +----------- Cristaria **
| |
+--+ +-- Anemina **
| +-----+
| | +-- Amuranodonta **
+--+
| +----- Pletholophus **
+--+
| +-- Sinanodonta **
+--+
+-- Beringiana **
Chen et al., 2025 [Publ 4487], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.857 | 6 | 9 | 7 | 6 | 35 | 4 | 2 | 165.0 |
+-- Alasmidontina
+-----------+
| +-- Anodontina
|
| +----- Lepidodesma
--+ +-----+
| | | +-- Cristaria **
| | +--+
| | +-- Lanceolaria
+--+
| +-- Anemina **
| +-----+
| | +-- Amuranodonta **
+--+
| +----- Pletholophus **
+--+
| +-- Sinanodonta **
+--+
+-- Beringiana **
Bolotov et al., 2020 [Publ 4084], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.909 | 9 | 9 | 7 | 6 | 27 | 3 | 1 | 159.0 |
+----------- Anodontina | | +----- Amuranodonta ** --+ +--+ | | | +-- Anemina ** | | +--+ +--+ +-- Buldowskia ** | | +----- Cristaria ** +--+ | +-- Beringiana ** +--+ +-- Sinanodonta **
Konopleva et al., 2023 [Publ 4269], fig. S1_4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.909 | 13 | 13 | 11 | 6 | 43 | 3 | 1 | 157.0 |
+----------- Anodontina | | +----- Anemina ** --+ +--+ | | | +-- Amuranodonta ** | | +--+ +--+ +-- Buldowskia ** | | +----- Cristaria ** +--+ | +-- Beringiana ** +--+ +-- Sinanodonta **
Sano et al., 2020 [Publ 4138], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.250 | 10 | 22 | 5 | 6 | 24 | 0 | 0 | 175.0 |
+----- Sinohyriopsis
+-----+
| | +-- Lamprotula
| +--+
+--+ +-- Pronodularia
| |
| | +-------- Sinosolenaia
| | |
| +--+ +----- Inversidens
| | |
| +--+ +-- Obovalis_omiensis__1
| | |
| +--+-- Obovalis_omiensis__2
| |
| +-- Ptychorhynchus
|
| +----- Lanceolaria_oxyrhyncha__1
| |
| +----- Lanceolaria_oxyrhyncha__2
| |
| +----- Cristaria_plicata__1 **
| |
--+ +----- Cristaria_plicata__2 **
| |
| +----- Sinanodonta_lauta **
| |
| +----- Beringiana_japonica **
| |
| +----- Beringiana_fukuharai__1 **
| |
| +----- Beringiana_beringiana **
| |
| +----- Beringiana_fukuharai__2 **
| |
| +----- Beringiana_fukuharai__3 **
| |
| +----- Buldowskia_iwakawai__1 **
| |
| +----- Buldowskia_iwakawai__2 **
| |
| +----- Sinanodonta_tumens__1 **
+--------+
+----- Sinanodonta_tumens__2 **
|
+----- Pletholophus **
|
+----- Anemina **
|
+----- Lanceolaria_lanceolata
|
+----- Cristaria_plicata__3 **
|
+----- Lanceolaria_gladiola
|
+----- Sinanodonta_angula **
|
+----- Lepidodesma
|
| +-- Aculamprotulina
+--+
| +-- Acuticosta
|
| +-- Cuneopsis
+--+
| +-- Nodularia
|
| +-- Schistodesmus
+--+
+-- Inversiunio
Xiang et al., 2025 [Publ 4433], fig. 7
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 7 | 7 | 5 | 5 | 6 | 2 | 1 | 163.0 |
+-------------- Anodontina --+ | +----------- Cristaria ** +--+ | +-------- Anemina ** +--+ | +----- Beringiana ** +--+ | +-- Mimo ** +--+ +-- Sinanodonta **
Sano et al., 2017 [Publ 3799], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 8 | 8 | 6 | 5 | 13 | 0 | 0 | 166.0 |
+-- Lanceolaria
+--------+
| +-- Oxynaiina
|
--+ +-- Cristaria **
| +-----+
| | +-- Buldowskia **
+--+
| +----- Pletholophus **
+--+
| +-- Sinanodonta **
+--+
+-- Beringiana **
Sano et al., 2017 [Publ 3799], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 9 | 9 | 7 | 5 | 24 | 0 | 0 | 167.0 |
+-- Cristaria **
+-----+
| +-- Buldowskia **
+-----+
| | +----- Beringiana **
| +--+
| | +-- Pletholophus **
| +--+
--+ +-- Sinanodonta **
|
| +-- Tchangsinaia
| +--------+
| | +-- Nodularia
| |
+--+ +-- Schistodesmus
| +-----+
| | +-- Acuticosta
+--+
| +----- Inversiunio
+--+
| +-- Lepidodesma
+--+
+-- Lanceolaria
Sano et al., 2022 [Publ 4311], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 12 | 12 | 12 | 5 | 0 | 0 | 0 | 170.0 |
+-- Beringiana **
+-----+
| +-- Sinanodonta **
--+
| +----- Pletholophus **
+--+
| +-- Cristaria **
+--+
+-- Buldowskia **
Sano et al., 2022 [Publ 4311], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 18 | 18 | 16 | 5 | 0 | 0 | 0 | 162.0 |
+-- Beringiana **
+-----+
| +-- Sinanodonta **
--+
| +----- Pletholophus **
+--+
| +-- Cristaria **
+--+
+-- Buldowskia **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.938 | 5 | 5 | 3 | 4 | 76 | 9 | 2 | 165.0 |
+-- Anodontina
+--------+
| +-- Alasmidontina
--+
| +-------- Cristaria **
+--+
| +----- Anemina **
+--+
| +-- Pletholophus **
+--+
+-- Sinanodonta **
Huang et al., 2019 [Publ 3843], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.824 | 9 | 9 | 7 | 4 | 192 | 11 | 2 | 159.0 |
+-------- Anemina **
+--+
| | +----- Cristaria **
| +--+
| | +-- Pletholophus **
| +--+
| +-- Sinanodonta **
|
--+ +-------- Alasmidonta
| |
| +-------- Lasmigona
| |
| +-------- Pseudanodonta
| |
| +-------- Simpsonaias
+--+
+-------- Strophitus
|
| +-- Pyganodon
+-----+
| +-- Utterbackia
|
| +----- Anodonta_anatina
+--+
| +-- Anodonta_cygnea
+--+
+-- Brachyanodon
Huang et al., 2019 [Publ 3843], fig. S3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.824 | 9 | 9 | 7 | 4 | 190 | 11 | 2 | 159.0 |
+-------- Anemina **
+-----+
| | +----- Cristaria **
| +--+
| | +-- Pletholophus **
| +--+
| +-- Sinanodonta **
|
--+ +-- Alasmidonta
| +--------+
| | +-- Lasmigona
| |
| | +----- Anodonta_anatina
| +-----+
| | | +-- Anodonta_cygnea
+--+ +--+
| +-- Brachyanodon
|
| +-- Pyganodon
| +-----+
| | +-- Utterbackia
+--+
| +----- Pseudanodonta
+--+
| +-- Simpsonaias
+--+
+-- Strophitus
Huang et al., 2019 [Publ 3843], fig. S4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.706 | 9 | 48 | 25 | 4 | 192 | 11 | 2 | 153.0 |
+----------------- Chamberlainia | +----------------- Megalonaias | +----------------- Uniomerus | +----------------- Quadrulina | +----------------- Sinohyriopsis | | +-- Indopseudodontina +--------------+ | +-- Pseudodontina | | +-- Rectidentina +--------------+ | +-- Contradentina | | +----- Lamprotula +-----------+ | | +-- Potomida | +--+ | +-- Pronodularia | | +-------- Sinosolenaia_carinata | | | | +----- Gonidea | +--+ | | | +-- Leguminaia | | +--+ | | +-- Microcondylaea +--------+ | | +----- Ptychorhynchus | +--+ | | | +-- Parvasolenaia | | +--+ | | +-- Koreosolenaia | | | | +----- Sinosolenaia_recognita | +--+ | | +-- Sinosolenaia_iridinea | +--+ | +-- Sinosolenaia_oleivora | | +-------------- Amblema | | | +-------------- Toxolasma | | | | +-- Obliquaria | +-----------+ | | +-- Plectomerus | | | | +----------- Elliptoideus --+ +--+ | | | +-------- Elliptio | | +--+ | | | +----- Eurynia +--+ +--+ | | | +-- Fusconaia | | +--+ | | +-- Pleurobema | | | | +----- Ellipsaria | | | | | +----- Potamilus_leptodon | | | | | +----- Medionidus | | | | | +----- Potamilus_alatus | | | | | +----- Ptychobranchus | +--------+ | +----- Truncilla | | | +----- Venustaconcha | | | +----- Cambarunio | | | +----- Leaunio | | | | +-- Ortmanniana | +--+ | +-- Lampsilis | | +-------- Acuticosta | | | +-------- Alasmidonta | | | +-------- Anodonta_anatina | | | +-------- Anemina ** | | | +-------- Anodonta_cygnea | | | +-------- Cristaria ** | | | +-------- Brachyanodon | | | +-------- Lasmigona | | | +-------- Lepidodesma | | | +-------- Pseudanodonta | | | +-------- Pyganodon | | +--------+-------- Simpsonaias | +-------- Strophitus | +-------- Utterbackia | +-------- Schistodesmus | +-------- Unionina | +-------- Aculamprotulina | +-------- Lanceolaria | | +-- Pletholophus ** +-----+ | +-- Sinanodonta ** | | +----- Cuneopsis_celtiformis | | | +----- Tchangsinaia | | +--+----- Cuneopsis_rufescens | +----- Nodularia | | +-- Cuneopsis_szechenyii +--+ +-- Cuneopsis_heudei
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 6 | 6 | 4 | 4 | 89 | 12 | 2 | 164.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
|
--+ +-- Anemina **
| +--+
| | +-- Cristaria **
+--+
| +-- Beringiana **
+--+
+-- Sinanodonta **
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.882 | 6 | 6 | 4 | 4 | 89 | 12 | 2 | 164.0 |
+-------- Cristaria **
+--+
| | +----- Anemina **
| +--+
| | +-- Beringiana **
| +--+
--+ +-- Sinanodonta **
|
| +-------- Brachyanodon
| |
| +-------- Pseudanodonta
+--+
| +----- Anodonta_cygnea
| |
+--+----- Alasmidontina
|
| +-- Anodonta_exulcerata
+--+
+-- Anodonta_anatina
Wu et al., 2024 [Publ 4350], fig. 8a_S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.941 | 6 | 6 | 4 | 4 | 85 | 12 | 2 | 164.0 |
+-------- Cristaria **
+--+
| | +----- Anemina **
| +--+
| | +-- Beringiana **
| +--+
--+ +-- Sinanodonta **
|
| +-- Pseudanodonta
| +--+
| | +-- Anodonta
+-----+
| +-- Brachyanodon
+--+
+-- Alasmidontina
Wu et al., 2024 [Publ 4350], fig. 8b_S2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.941 | 6 | 6 | 4 | 4 | 85 | 12 | 2 | 164.0 |
+-------- Cristaria **
+--+
| | +----- Anemina **
| +--+
| | +-- Beringiana **
--+ +--+
| +-- Sinanodonta **
|
| +-------- Brachyanodon
+--+
| +----- Alasmidontina
+--+
| +-- Pseudanodonta
+--+
+-- Anodonta
Wu et al., 2024 [Publ 4379], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.941 | 6 | 6 | 4 | 4 | 47 | 12 | 2 | 164.0 |
+-------- Cristaria **
+--+
| | +----- Anemina **
| +--+
| | +-- Beringiana **
--+ +--+
| +-- Sinanodonta **
|
| +-------- Brachyanodon
+--+
| +----- Alasmidontina
+--+
| +-- Pseudanodonta
+--+
+-- Anodonta
Wu et al., 2024 [Publ 4388], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 49 | 3 | 2 | 166.0 |
+-- Anodontina
+--------+
| +-- Alasmidontina
--+
| +-------- Cristaria **
+--+
| +----- Anemina **
+--+
| +-- Beringiana **
+--+
+-- Sinanodonta **
Hou et al., 2024 [Publ 4400], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.941 | 6 | 6 | 4 | 4 | 69 | 12 | 2 | 164.0 |
+-------- Cristaria **
+--+
| | +----- Anemina **
| +--+
| | +-- Beringiana **
| +--+
--+ +-- Sinanodonta **
|
| +-- Pseudanodonta
| +--+
| | +-- Anodonta
+-----+
| +-- Brachyanodon
+--+
+-- Alasmidontina
Hou et al., 2025 [Publ 4445], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.941 | 6 | 6 | 4 | 4 | 46 | 12 | 2 | 164.0 |
+-------- Cristaria **
+--+
| | +----- Anemina **
| +--+
| | +-- Beringiana **
--+ +--+
| +-- Sinanodonta **
|
| +-------- Brachyanodon
+--+
| +----- Alasmidontina
+--+
| +-- Pseudanodonta
+--+
+-- Anodonta
Froufe et al., 2017 [Publ 3802], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 10 | 10 | 7 | 4 | 2 | 2 | 1 | 159.0 |
+----------- Anodontina --+ | +-------- Anemina ** +--+ | +----- Cristaria ** +--+ | +-- Beringiana ** +--+ +-- Sinanodonta **
Lopes-Lima et al., 2020 [Publ 4080], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 9 | 9 | 7 | 4 | 2 | 0 | 0 | 163.0 |
+----------- Margaritifera --+ | +-------- Sinanodonta ** +--+ | +----- Amuranodonta ** +--+ | +-- Anemina ** +--+ +-- Buldowskia **
Wu et al., 2016 [Publ 3711], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.900 | 3 | 7 | 5 | 3 | 18 | 4 | 2 | 163.0 |
+----- Aculamprotulina
+--------+
| | +-- Unionina
| +--+
--+ +-- Oxynaiina
|
| +----------- Sinanodonta **
+--+
| +-------- Anemina **
+--+
| +----- Cristaria **
+--+
| +-- Anodontina
+--+
+-- Alasmidontina
Fonseca et al., 2016 [Publ 3775], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 3 | 27 | 5 | 2 | 166.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
--+
| +----- Cristaria **
+--+
| +-- Anemina **
+--+
+-- Sinanodonta **
Bolotov et al., 2017 [Publ 3761], fig. S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.867 | 6 | 6 | 4 | 3 | 133 | 9 | 2 | 162.0 |
+----- Cristaria **
+--------+
| | +-- Pletholophus **
| +--+
--+ +-- Sinanodonta **
|
| +----------- Anodonta_anatina
+--+
| +-------- Alasmidontina
+--+
| +----- Pseudanodonta
+--+
| +-- Anodonta_cygnea
+--+
+-- Brachyanodon
Bolotov et al., 2017 [Publ 3761], fig. S2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.800 | 6 | 6 | 4 | 3 | 133 | 9 | 2 | 162.0 |
+-------- Lanceolaria | | +----- Cristaria ** +--+ | | +-- Pletholophus ** --+ +--+ | +-- Sinanodonta ** | | +----- Anodonta_anatina | | | +----- Pseudanodonta +--+ +----- Alasmidontina | | +-- Anodonta_cygnea +--+ +-- Brachyanodon
Bolotov et al., 2017 [Publ 3764], fig. S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.533 | 6 | 18 | 0 | 3 | 119 | 9 | 2 | 167.0 |
+-- Unio_crassus
|
+-- Unio_pictorum
|
+-- Unio_tumidus
|
+-- Aculamprotulina
|
+-- Cuneopsis_szechenyii
+--+
| +-- Tchangsinaia
| |
| +-- Cuneopsis_rufescens
| |
| +-- Nodularia_douglasiae
| |
| +-- Nodularia_nuxpersicae
| |
| +-- Schistodesmus
|
| +-- Alasmidonta
| |
| +-- Anodonta_anatina
--+ |
| +-- Anodonta_cygnea
| |
| +-- Brachyanodon
| |
| +-- Lasmigona
| |
| +-- Pseudanodonta
| |
| +-- Pyganodon
| |
| +-- Simpsonaias
| |
| +-- Strophitus
+--+
+-- Sinanodonta_lauta **
|
+-- Cristaria_plicata **
|
+-- Cristaria_radiata **
|
+-- Pletholophus **
|
+-- Sinanodonta_lucida **
|
+-- Sinanodonta_jourdyi **
|
+-- Lanceolaria_gladiola
|
+-- Lanceolaria_grayii
|
+-- Lanceolaria_danzhouensis
Lopes-Lima et al., 2017 [Publ 3792], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 3 | 28 | 5 | 2 | 166.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
--+
| +----- Cristaria **
+--+
| +-- Anemina **
+--+
+-- Sinanodonta **
Soroka & Burzynski, 2017 [Publ 3837], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 3 | 19 | 6 | 2 | 166.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
--+
| +----- Cristaria **
+--+
| +-- Anemina **
+--+
+-- Sinanodonta **
Bolotov et al., 2018 [Publ 3826], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 3 | 78 | 6 | 2 | 164.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
--+
| +----- Cristaria **
+--+
| +-- Pletholophus **
+--+
+-- Sinanodonta **
Huang et al., 2018 [Publ 3828], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 3 | 38 | 5 | 2 | 166.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
--+
| +----- Cristaria **
+--+
| +-- Anemina **
+--+
+-- Sinanodonta **
Wu et al., 2018 [Publ 3836], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.833 | 6 | 6 | 3 | 3 | 47 | 7 | 2 | 163.0 |
+-- Anemina **
|
+-----+-- Cristaria **
| |
--+ +-- Sinanodonta **
|
| +----- Anodonta_cygnea
+--+
| +-- Anodonta_anatina
+--+
+-- Alasmidontina
Wang et al., 2018 [Publ 3846], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 3 | 27 | 3 | 2 | 166.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
--+
| +----- Cristaria **
+--+
| +-- Anemina **
+--+
+-- Sinanodonta **
Aunins et al., 2018 [Publ 3863], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 3 | 36 | 7 | 2 | 166.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
--+
| +----- Cristaria **
+--+
| +-- Anemina **
+--+
+-- Sinanodonta **
Wang et al., 2018 [Publ 3865], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 3 | 27 | 3 | 2 | 166.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
--+
| +----- Cristaria **
+--+
| +-- Anemina **
+--+
+-- Sinanodonta **
Cha et al., 2018 [Publ 3916], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 3 | 34 | 5 | 2 | 166.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
--+
| +----- Cristaria **
+--+
| +-- Anemina **
+--+
+-- Sinanodonta **
Wu et al., 2018 [Publ 4143], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.900 | 3 | 7 | 5 | 3 | 19 | 4 | 2 | 163.0 |
+----- Aculamprotulina
+--------+
| | +-- Unionina
| +--+
--+ +-- Oxynaiina
|
| +----------- Sinanodonta **
+--+
| +-------- Anemina **
+--+
| +----- Cristaria **
+--+
| +-- Anodontina
+--+
+-- Alasmidontina
Huang et al., 2019 [Publ 3843], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.818 | 4 | 4 | 2 | 3 | 52 | 6 | 2 | 166.0 |
+----- Cristaria **
+--+
| | +-- Anemina **
| +--+
--+ +-- Sinanodonta **
|
| +----- Anodonta_cygnea
+--+
| +-- Anodonta_anatina
+--+
+-- Alasmidontina
Wu et al., 2019 [Publ 3864], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 3 | 42 | 5 | 2 | 166.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
--+
| +----- Cristaria **
+--+
| +-- Anemina **
+--+
+-- Sinanodonta **
Froufe et al., 2019 [Publ 3936], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 3 | 40 | 5 | 2 | 166.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
--+
| +----- Cristaria **
+--+
| +-- Anemina **
+--+
+-- Sinanodonta **
Froufe et al., 2019 [Publ 3936], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 3 | 40 | 5 | 2 | 166.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
--+
| +----- Cristaria **
+--+
| +-- Anemina **
+--+
+-- Sinanodonta **
Riccardi et al., 2020 [Publ 4089], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.846 | 4 | 4 | 2 | 3 | 15 | 9 | 2 | 166.0 |
+----- Cristaria **
+--------+
| | +-- Anemina **
| +--+
--+ +-- Sinanodonta **
|
| +----------- Anodonta_anatina
| |
+--+ +-- Brachyanodon
| +-----+
| | +-- Alasmidontina
+--+
| +----- Pseudanodonta
+--+
| +-- Anodonta_exulcerata
+--+
+-- Anodonta_cygnea
Fukata & Iigo, 2020 [Publ 4106], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 3 | 50 | 6 | 2 | 166.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
--+
| +----- Cristaria **
+--+
| +-- Anemina **
+--+
+-- Sinanodonta **
Wu et al., 2020 [Publ 4121], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 3 | 44 | 5 | 2 | 166.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
--+
| +----- Cristaria **
+--+
| +-- Anemina **
+--+
+-- Sinanodonta **
Wu et al., 2020 [Publ 4147], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 27 | 4 | 2 | 167.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
--+
| +----- Cristaria **
+--+
| +-- Sinanodonta **
+--+
+-- Anemina **
Chen et al., 2020 [Publ 4170], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.692 | 3 | 34 | 32 | 3 | 31 | 8 | 2 | 191.0 |
+----- Prolasmidonta
+--+
| | +-- Alasmidonta
| +--+
+-----------------+ +-- Anemina **
| |
| | +----- Pseudodontina
| +--+
| | +-- Lamprotula_caveata
| +--+
| +-- Chamberlainia
|
| +----- Oxynaiina
| +-----+
| | | +-- Lepidodesma
--+ | +--+
| | +-- Aculamprotulina
| +-----------+
| | | +----- Platynaias
| | | +--+
| | | | | +-- Utterbackia_imbecillis
| | | | +--+
| | +--+ +-- Pyganodon
| | |
| | | +----- Sinanodonta **
| | +--+
| | | +-- Cristaria **
+--+ +--+
| +-- Anodontina
|
| +-- Utterbackia_peninsularis
| +--------+
| | +-- Potamilus_alatus
| +--------+
| | | +-------- Lamprotula_leaii
| | +--+
| | | +----- Sinohyriopsis
| | +--+
+--+ | +-- Parvasolenaia
| +--+
| +-- Sinosolenaia
|
| +----------------- Gibbosula
+--+
| +-------------- Quadrulina
+--+
| +----------- Toxolasma
+--+
| +-------- Lampsilis_ornata
+--+
| +----- Potamilus_leptodon
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Lampsilis_powellii
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 3 | 44 | 5 | 2 | 166.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
--+
| +----- Cristaria **
+--+
| +-- Anemina **
+--+
+-- Sinanodonta **
Wu et al., 2021 [Publ 4366], fig. 2_I
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.833 | 4 | 4 | 2 | 3 | 87 | 6 | 2 | 164.0 |
+----- Anemina **
+--------+
| | +-- Cristaria **
| +--+
--+ +-- Sinanodonta **
|
| +----------- Lasmigona
+--+
| +-------- Anodonta
+--+
| +----- Pseudanodonta
+--+
| +-- Pyganodon
+--+
+-- Strophitus
Wu et al., 2021 [Publ 4366], fig. 3-BI
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.846 | 4 | 4 | 2 | 3 | 69 | 9 | 2 | 166.0 |
+----- Cristaria **
+--+
| | +-- Anemina **
| +--+
--+ +-- Sinanodonta **
|
| +----- Anodonta_cygnea
+--+
| +-- Anodonta_anatina
+--+
+-- Alasmidontina
Wu et al., 2021 [Publ 4366], fig. 3-ML
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.846 | 4 | 4 | 2 | 3 | 69 | 9 | 2 | 166.0 |
+----- Cristaria **
+--+
| | +-- Anemina **
| +--+
--+ +-- Sinanodonta **
|
| +----- Anodonta_cygnea
+--+
| +-- Anodonta_anatina
+--+
+-- Alasmidontina
Wu et al., 2021 [Publ 4366], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.846 | 4 | 4 | 2 | 3 | 64 | 8 | 2 | 166.0 |
+----- Cristaria **
+--+
| | +-- Anemina **
| +--+
--+ +-- Sinanodonta **
|
| +----- Anodonta_cygnea
+--+
| +-- Anodonta_anatina
+--+
+-- Alasmidontina
Wu et al., 2021 [Publ 4366], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 4 | 4 | 2 | 3 | 87 | 6 | 2 | 164.0 |
+----- Anemina **
+--------+
| | +-- Cristaria **
| +--+
--+ +-- Sinanodonta **
|
| +----------- Anodonta_cygnea
| |
+--+ +-- Lasmigona
| +-----+
| | +-- Anodonta_anatina
+--+
| +----- Pseudanodonta
+--+
| +-- Pyganodon
+--+
+-- Strophitus
Wu et al., 2022 [Publ 4238], fig. 2d
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 3 | 34 | 6 | 2 | 166.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
--+
| +----- Cristaria **
+--+
| +-- Anemina **
+--+
+-- Sinanodonta **
Wu et al., 2022 [Publ 4238], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 3 | 69 | 6 | 2 | 166.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
--+
| +----- Cristaria **
+--+
| +-- Anemina **
+--+
+-- Sinanodonta **
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 3 | 71 | 9 | 2 | 163.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
--+
| +----- Cristaria **
+--+
| +-- Pletholophus **
+--+
+-- Sinanodonta **
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.933 | 5 | 5 | 3 | 3 | 71 | 9 | 2 | 163.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
--+
| +----- Cristaria **
+--+
| +-- Pletholophus **
+--+
+-- Sinanodonta **
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 34 | 2 | 2 | 167.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
--+
| +----- Cristaria **
+--+
| +-- Sinanodonta **
+--+
+-- Anemina **
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 3 | 35 | 2 | 2 | 166.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
--+
| +----- Cristaria **
+--+
| +-- Anemina **
+--+
+-- Sinanodonta **
Dai et al., 2023 [Publ 4285], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 50 | 4 | 2 | 167.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
--+
| +----- Cristaria **
+--+
| +-- Anemina **
+--+
+-- Sinanodonta **
Wu et al., 2023 [Publ 4339], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.900 | 4 | 4 | 2 | 3 | 15 | 5 | 2 | 164.0 |
+-- Alasmidontina
+-----+
| +-- Anodontina
--+
| +----- Anemina **
+--+
| +-- Cristaria **
+--+
+-- Sinanodonta **
Dai et al., 2024 [Publ 4358], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 50 | 4 | 2 | 167.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
--+
| +----- Cristaria **
+--+
| +-- Anemina **
+--+
+-- Sinanodonta **
Dai et al., 2025 [Publ 4440], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 3 | 83 | 6 | 2 | 166.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
--+
| +----- Cristaria **
+--+
| +-- Anemina **
+--+
+-- Sinanodonta **
Chen et al., 2025 [Publ 4448], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 3 | 80 | 6 | 2 | 165.0 |
+-- Anodontina
+-----+
| +-- Alasmidontina
--+
| +----- Cristaria **
+--+
| +-- Anemina **
+--+
+-- Sinanodonta **
Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.875 | 4 | 4 | 2 | 3 | 136 | 13 | 2 | 166.0 |
+----- Cristaria **
+--------+
| | +-- Anemina **
| +--+
| +-- Sinanodonta **
--+
| +-- Anodonta_cygnea
| +--------+
| | +-- Anodonta_exulcerata
+--+
| +-------- Anodonta_anatina
+--+
| +----- Alasmidontina
+--+
| +-- Brachyanodon
+--+
+-- Pseudanodonta
Xue et al., 2016 [Publ 3659], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.857 | 4 | 5 | 3 | 3 | 11 | 1 | 1 | 165.0 |
+-- Cristaria **
+--+
| +-- Alasmidontina
+--------+
| | +-- Anemina **
| +--+
--+ +-- Sinanodonta **
|
| +----------- Lanceolaria
+--+
| +-------- Aculamprotulina
+--+
| +----- Schistodesmus
+--+
| +-- Nodularia
+--+
+-- Unionina
Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 3 | 3 | 1 | 167.0 |
+-------- Anodontina --+ | +----- Cristaria ** +--+ | +-- Anemina ** +--+ +-- Sinanodonta **
Soroka, 2010 [Publ 3404], fig. 9a
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 3 | 0 | 0 | 0 | 163.0 |
+----- Buldowskia ** --+ | +-- Beringiana ** +--+ +-- Sinanodonta **
Klishko et al., 2014 [Publ 3666], fig. 8
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 3 | 0 | 0 | 0 | 164.0 |
+----- Cristaria ** --+ | +-- Beringiana ** +--+ +-- Sinanodonta **
Klishko et al., 2016 [Publ 3733], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 3 | 0 | 0 | 0 | 164.0 |
+----- Cristaria ** --+ | +-- Sinanodonta ** +--+ +-- Beringiana **
Huang et al., 2019 [Publ 3843], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.600 | 8 | 32 | 30 | 3 | 38 | 0 | 0 | 164.0 |
+-------------------- Sinohyriopsis | --+ +----------------- Anemina ** | | +--+ +-------------- Acuticosta | | +--+ +----------- Sinanodonta ** | | +--+ +-------- Aculamprotulina | | +--+ +-- Cristaria ** | +--+ | | +-- Lanceolaria +--+ | +-- Lepidodesma +--+ +-- Oxynaiina
Huang et al., 2019 [Publ 3843], fig. S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.600 | 8 | 33 | 36 | 3 | 39 | 0 | 0 | 160.0 |
+----------------- Anemina **
|
+--+ +-------------- Sinanodonta **
| | |
| +--+ +----------- Acuticosta
| | |
| +--+ +-- Lepidodesma
| | +-----+
| | | +-- Cristaria **
| +--+
--+ | +----- Lanceolaria
| +--+
| | +-- Aculamprotulina
| +--+
| +-- Oxynaiina
|
| +----------------- Sinohyriopsis
| |
| | +----- Parvasolenaia_rivularis
+--+ +--------+
| | | +-- Parvasolenaia_triangularis
| | +--+
| | +-- Ptychorhynchus
+--+
| +-- Sinosolenaia_carinata
| +--------+
| | +-- Lamprotula
+--+
| +-------- Sinosolenaia_recognita
| |
+--+ +----- Sinosolenaia_oleivora__2
| |
+--+ +-- Sinosolenaia_oleivora__1
| |
+--+-- Sinosolenaia_iridinea
|
+-- Sinosolenaia_oleivora__3
Kano et al., 2019 [Publ 4194], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 7 | 0 | 0 | 165.0 |
+----- Sinanodonta **
+-----+
| | +-- Pletholophus **
| +--+
| +-- Beringiana **
--+
| +-- Lanceolaria
| +-----+
| | +-- Oxynaiina
+--+
| +----- Inversidens
+--+
| +-- Obovalis
+--+
+-- Pronodularia
Lopes-Lima et al., 2020 [Publ 4080], fig. 9
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 3 | 2 | 0 | 0 | 164.0 |
+-------- Margaritifera --+ | +----- Cristaria ** +--+ | +-- Sinanodonta ** +--+ +-- Pletholophus **
Mabuchi & Nishida, 2020 [Publ 4120], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.600 | 5 | 5 | 5 | 3 | 7 | 0 | 0 | 170.0 |
+----- Nodularia
+-----------+
| | +-- Lanceolaria
| +--+
| +-- Inversiunio
|
--+ +-------------- Cristaria **
| |
| | +----------- Sinanodonta_tumens **
| | |
| | | +-- Sinanodonta_lauta__5 **
| | | |
+--+ | +-- Sinanodonta_lauta__6 **
| | |
| | +-- Beringiana_japonica__15 **
| | |
| | +-- Beringiana_japonica__16 **
| | +-----+
| | | +-- Beringiana_japonica__17 **
+--+ | |
| | +-- Beringiana_japonica__18 **
| | |
| | +-- Beringiana_japonica__19 **
| | |
| | +-- Beringiana_japonica__20 **
| |
| | +----- Beringiana_japonica__1 **
| | |
| | +----- Beringiana_japonica__2 **
| | |
+--+ +----- Beringiana_japonica__3 **
| |
| +----- Beringiana_japonica__4 **
| |
| +----- Beringiana_japonica__5 **
| |
| +----- Sinanodonta_lauta__1 **
| |
| +----- Beringiana_japonica__6 **
| |
| +----- Sinanodonta_lauta__2 **
| |
| +----- Sinanodonta_calipygos__1 **
| |
+--+----- Beringiana_japonica__7 **
|
+----- Sinanodonta_lauta__4 **
|
+----- Beringiana_japonica__8 **
|
+----- Beringiana_japonica__9 **
|
+----- Beringiana_japonica__10 **
|
+----- Beringiana_japonica__11 **
|
+----- Beringiana_japonica__12 **
|
+----- Beringiana_japonica__13 **
|
+----- Beringiana_japonica__14 **
|
| +-- Sinanodonta_lauta__3 **
+--+
+-- Sinanodonta_calipygos__2 **
Bogan et al., 2023 [Publ 4295], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 6 | 6 | 4 | 3 | 1 | 0 | 0 | 162.0 |
+-------- Gibbosula --+ | +----- Cristaria ** +--+ | +-- Sinanodonta ** +--+ +-- Pletholophus **
Bogan et al., 2023 [Publ 4295], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 3 | 1 | 0 | 0 | 163.0 |
+-------- Gibbosula --+ | +----- Cristaria ** +--+ | +-- Sinanodonta ** +--+ +-- Pletholophus **
Bogan et al., 2023 [Publ 4295], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 12 | 12 | 9 | 3 | 1 | 0 | 0 | 163.0 |
+-------- Gibbosula --+ | +----- Pletholophus ** +--+ | +-- Cristaria ** +--+ +-- Sinanodonta **
Dai et al., 2024 [Publ 4389], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 43 | 0 | 0 | 167.0 |
+----- Sinanodonta **
+-----------------------------+
| | +-- Anemina **
| +--+
| +-- Cristaria **
|
--+ +----- Ctenodesmina
| +--------------------------+
| | | +-- Rectidentina
| | +--+
| | +-- Contradentina
| |
+--+ +-- Chamberlainia
| +--------------------------+
| | +-- Sinohyriopsis
| |
| | +----- Lamprotula
+--+ +--------------------+
| | | +-- Pronodularia
| | +--+
| | +-- Potomida
+--+
| +-- Schepmania
| +--------------------+
| | +-- Pseudodontina
| |
+--+ +-------------------- Sinosolenaia
| |
| | +----- Gonidea
+--+ +-----------+
| | | +-- Leguminaia
| | +--+
+--+ +-- Microcondylaea
|
| +-------------- Pseudopostolata
| |
+--+ +-- Parvasolenaia
| +--------+
| | +-- Koreosolenaia
+--+
| +-------- Postolata
+--+
| +----- Ptychorhynchus
+--+
| +-- Cosmopseudodon
+--+
+-- Obovalis
Dai et al., 2025 [Publ 4441], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 44 | 0 | 0 | 167.0 |
+----- Sinanodonta **
+-----------------------------+
| | +-- Anemina **
| +--+
| +-- Cristaria **
|
--+ +----- Ctenodesmina
| +--------------------------+
| | | +-- Rectidentina
| | +--+
| | +-- Contradentina
| |
+--+ +-- Chamberlainia
| +--------------------------+
| | +-- Sinohyriopsis
| |
| | +----- Potomida
+--+ +--------------------+
| | | +-- Pronodularia
| | +--+
| | +-- Lamprotula
+--+
| +-- Schepmania
| +--------------------+
| | +-- Pseudodontina
| |
+--+ +-------------------- Sinosolenaia
| |
| | +----- Gonidea
+--+ +-----------+
| | | +-- Leguminaia
| | +--+
+--+ +-- Microcondylaea
|
| +-------------- Pseudopostolata
| |
+--+ +-- Koreosolenaia
| +--------+
| | +-- Parvasolenaia
+--+
| +-------- Postolata
| |
+--+ +-- Guiunio
| +--+
| | +-- Cosmopseudodon
+--+
| +-- Obovalis
+--+
+-- Ptychorhynchus
Chen et al., 2025 [Publ 4663], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 45 | 0 | 0 | 167.0 |
+----- Sinanodonta **
+--------------------------------+
| | +-- Anemina **
| +--+
| +-- Cristaria **
|
--+ +----- Ctenodesmina
| +-----------------------------+
| | | +-- Rectidentina
| | +--+
| | +-- Contradentina
| |
+--+ +-- Chamberlainia
| +-----------------------------+
| | +-- Sinohyriopsis
| |
| | +----- Potomida
+--+ +-----------------------+
| | | +-- Pronodularia
| | +--+
| | +-- Lamprotula
+--+
| +-- Schepmania
| +-----------------------+
| | +-- Pseudodontina
| |
+--+ +----------------------- Sinosolenaia
| |
| | +----- Gonidea
+--+ +--------------+
| | | +-- Leguminaia
| | +--+
+--+ +-- Microcondylaea
|
| +----------------- Pseudopostolata
| |
+--+ +-- Koreosolenaia
| +-----------+
| | +-- Parvasolenaia
+--+
| +----------- Postolata
| |
+--+ +-- Guiunio
| +-----+
| | +-- Cosmopseudodon
+--+
| +----- Obovalis
+--+
| +-- Tenuiunio
+--+
+-- Ptychorhynchus
Chong et al., 2008 [Publ 2668], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.706 | 2 | 2 | 0 | 2 | 16 | 15 | 2 | -- |
+----------------- Anodonta_cygnea | +----------------- Pyganodon | +----------------- Alasminota | +----------------- Anodonta_anatina | +----------------- Unionina | | +-- Brachyanodon_nuttalliana +--------------+ | +-- Brachyanodon_californiensis | --+ +-- Beringiana ** +--------------+ | +-- Sinanodonta ** | | +-- Pseudodontoideus | | +--------------+-- Alasmidonta | | | +-- Pterosyna | | +-------------- Pseudanodonta | | | | +----- Strophitus | +--------+ +--+ | +-- Prolasmidonta | +--+ | +-- Platynaias | | +----------- Brachyanodon_oregonensis__4 +--+ | +-------- Brachyanodon_oregonensis__3 +--+ | +----- Brachyanodon_oregonensis__2 +--+ | +-- Brachyanodon_oregonensis__1 +--+ +-- Brachyanodon_kennerlyi
Feng et al., 2016 [Publ 3800], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.778 | 3 | 3 | 1 | 2 | 11 | 3 | 2 | 165.0 |
+----- Alasmidontina --+ | +-- Anemina ** +--+ +-- Sinanodonta **
Guerra et al., 2017 [Publ 3772], fig. 2a
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.900 | 2 | 8 | 0 | 2 | 23 | 6 | 2 | 160.0 |
+-- Aculamprotulina
+--------+
| +-- Unionina
--+
| +-------- Sinanodonta **
+--+
| +----- Cristaria **
+--+
| +-- Anodontina
+--+
+-- Alasmidontina
Guerra et al., 2017 [Publ 3772], fig. 2b
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 23 | 6 | 2 | -- |
+-- Cristaria **
+--+
| +-- Sinanodonta **
--+
| +-- Anodontina
+--+
+-- Alasmidontina
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 55 | 2 | 2 | -- |
+-- Cristaria **
+--+
| +-- Sinanodonta **
--+
| +-- Anodontina
+--+
+-- Alasmidontina
Pfeiffer et al., 2019 [Publ 3937], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 55 | 2 | 2 | -- |
+-- Cristaria **
+--+
| +-- Sinanodonta **
--+
| +-- Anodontina
+--+
+-- Alasmidontina
Whelan et al., 2023 [Publ 4287], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.913 | 4 | 4 | 2 | 2 | 37 | 37 | 2 | 164.0 |
+-- Anemina **
+--------+
| +-- Sinanodonta **
--+
| +-------- Anodonta_cygnea
| |
+--+ +-- Brachyanodon
| +--+
| | +-- Pseudanodonta
+--+
| +-- Anodonta_anatina
+--+
+-- Alasmidontina
Galal-Khallaf et al., 2024 [Publ 4459], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.667 | 4 | 4 | 2 | 2 | 24 | 9 | 2 | 166.0 |
+-- Beringiana **
+-----------+
| +-- Sinanodonta **
|
--+ +-- Anodonta_exulcerata
| +--------+
| | +-- Anodonta_cygnea
| |
+--+ +-- Brachyanodon_californiensis
| +-----+
| | +-- Brachyanodon_nuttalliana
| |
+--+ +-- Brachyanodon_kennerlyi
| +--+
| | +-- Brachyanodon_oregonensis
+--+
| +-- Anodonta_anatina
+--+
+-- Alasmidontina
Huang et al., 2002 [Publ 2657], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 32 | 5 | 1 | -- |
+----- Alasmidontina --+ | +-- Sinanodonta ** +--+ +-- Cristaria **
Huang et al., 2002 [Publ 2657], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.800 | 2 | 2 | 0 | 2 | 32 | 5 | 1 | -- |
+-------- Strophitus | +-------- Schistodesmus | +-------- Lepidodesma | +-------- Acuticosta | +-------- Lanceolaria | --+ +-- Sinanodonta ** +-----+ | +-- Cristaria ** | | +-- Nodularia +-----+ | +-- Tchangsinaia | | +----- Utterbackia | | +--+----- Utterbackiana | | +-- Alasmidonta +--+ +-- Lasmigona
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 29 | 3 | 1 | -- |
+----- Alasmidontina --+ | +-- Cristaria ** +--+ +-- Sinanodonta **
Huang et al., 2013 [Publ 3629], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 14 | 4 | 1 | -- |
+----- Alasmidontina --+ | +-- Sinanodonta ** +--+ +-- Cristaria **
Zhou et al., 2016 [Publ 3710], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.875 | 2 | 6 | 0 | 2 | 18 | 4 | 1 | 162.0 |
+----- Lepidodesma
+--+
| | +-- Oxynaiina
| +--+
--+ +-- Unionina
|
| +----- Sinanodonta **
+--+
| +-- Cristaria **
+--+
+-- Alasmidontina
Wen et al., 2017 [Publ 3795], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 14 | 1 | 1 | -- |
+----- Anodontina --+ | +-- Sinanodonta ** +--+ +-- Cristaria **
Riccardi et al., 2020 [Publ 4089], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.857 | 2 | 2 | 0 | 2 | 4 | 4 | 1 | -- |
+----- Anodontina --+ | +-- Sinanodonta ** +--+ +-- Anemina **
Pfarrer et al., 2022 [Publ 4305], fig. 2-5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 10 | 3 | 1 | -- |
+----- Anodontina --+ | +-- Cristaria ** +--+ +-- Sinanodonta **
Zhang et al., 2004 [Publ 2735], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 12 | 0 | 0 | -- |
+-- Sinanodonta **
+-----+
| +-- Cristaria **
--+
| +----- Oxynaiina
+--+
| +-- Acuticosta
+--+
+-- Lanceolaria
Zhou et al., 2007 [Publ 2658], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 19 | 0 | 0 | -- |
+-- Sinanodonta **
+--------------+
| +-- Cristaria **
|
| +----------- Sinohyriopsis
| +--+
--+ | | +-------- Lamprotula_leaii
| | +--+
| | | +----- Lamprotula_caveata
| | +--+
| | | +-- Gibbosula
+--+ +--+
| +-- Lamprotula_cornuumlunae
|
| +----- Aculamprotulina
| +--+
| | | +-- Tchangsinaia
| | +--+
+-----+ +-- Nodularia
|
| +----- Schistodesmus
+--+
| +-- Lanceolaria
+--+
+-- Acuticosta
Zhou et al., 2007 [Publ 2658], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.500 | 2 | 18 | 0 | 2 | 16 | 0 | 0 | 172.0 |
+-------------- Sinanodonta ** | --+ +----------- Cristaria ** | | +--+ +-- Lamprotula | +-----+ | | +-- Quadrulina +--+ | +----- Aculamprotulina +--+ | +-- Acuticosta +--+ +-- Oxynaiina
Soroka, 2010 [Publ 3404], fig. 9b
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 2 | 0 | 0 | 0 | 164.0 |
+-- Buldowskia ** --+ +-- Sinanodonta **
Soroka, 2010 [Publ 3404], fig. 9c
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 3 | 2 | 0 | 0 | 0 | 167.0 |
+-- Buldowskia ** --+ +-- Sinanodonta **
Ouyang et al., 2011 [Publ 3408], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 2 | 10 | 0 | 0 | 165.0 |
+-- Anemina **
+--------+
| +-- Sinanodonta **
--+
| +-------- Oxynaiina
| |
+--+ +----- Lanceolaria_grayii
| |
+--+----- Acuticosta
|
| +-- Lanceolaria_gladiola
+--+
+-- Aculamprotulina
Zhang et al., 2013 [Publ 4214], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.500 | 2 | 4 | 0 | 2 | 2 | 0 | 0 | 168.0 |
+-------- Cristaria ** --+ | +----- Sinanodonta ** +--+ | +-- Margaritifera +--+ +-- Lamprotula
Ouyang et al., 2015 [Publ 4493], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 2 | 27 | 0 | 0 | 164.0 |
+----- Acuticosta --+ | +-- Anemina ** +--+ +-- Sinanodonta **
Ouyang et al., 2015 [Publ 4493], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 2 | 27 | 0 | 0 | 166.0 |
+----- Oxynaiina --+ | +-- Anemina ** +--+ +-- Sinanodonta **
Xia et al., 2016 [Publ 3712], fig. 7B
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.500 | 2 | 3 | 0 | 2 | 1 | 0 | 0 | 167.0 |
+----- Sinanodonta ** --+ | +-- Cristaria ** +--+ +-- Sinohyriopsis
Sayenko et al., 2017 [Publ 3760], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 2 | 0 | 0 | 0 | 164.0 |
+-- Beringiana ** --+ +-- Sinanodonta **
Vikhrev et al., 2017 [Publ 3810], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 8 | 8 | 6 | 2 | 2 | 0 | 0 | 160.0 |
+-- Gibbosula
+--+
| +-- Margaritifera
--+
| +-- Anemina **
+--+
+-- Sinanodonta **
Bespalaya et al., 2018 [Publ 3811], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 8 | 8 | 7 | 2 | 2 | 0 | 0 | 167.0 |
+-- Gibbosula
+--+
| +-- Margaritifera
--+
| +-- Anemina **
+--+
+-- Sinanodonta **
Kondakov et al., 2018 [Publ 3914], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 8 | 8 | 6 | 2 | 0 | 0 | 0 | 160.0 |
+-- Anemina ** --+ +-- Sinanodonta **
Lopes-Lima et al., 2020 [Publ 4080], fig. 10
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 12 | 12 | 8 | 2 | 2 | 0 | 0 | 160.0 |
+----- Margaritifera --+ | +-- Beringiana ** +--+ +-- Sinanodonta **
Lopes-Lima et al., 2020 [Publ 4080], fig. 7
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 6 | 6 | 4 | 2 | 2 | 0 | 0 | 166.0 |
+----- Margaritifera --+ | +-- Sinanodonta ** +--+ +-- Beringiana **
Lopes-Lima et al., 2020 [Publ 4080], fig. 8
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 2 | 2 | 0 | 0 | 165.0 |
+----- Margaritifera --+ | +-- Sinanodonta ** +--+ +-- Cristaria **
Bauer et al., 2021 [Publ 4365], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 8 | 8 | 5 | 2 | 0 | 0 | 0 | 161.0 |
+-- Anemina ** --+ +-- Sinanodonta **
Douda et al., 2024 [Publ 4364], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 13 | 13 | 12 | 2 | 1 | 0 | 0 | 159.0 |
+----- Margaritifera --+ | +-- Beringiana ** +--+ +-- Sinanodonta **
Wu et al., 2024 [Publ 4379], fig. 4_6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 2 | 20 | 0 | 0 | 165.0 |
+----- Oxynaiina --+ | +-- Cristaria ** +--+ +-- Sinanodonta **
Kashiri et al., 2024 [Publ 4822], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 8 | 8 | 6 | 2 | 2 | 0 | 0 | 160.0 |
+-- Gibbosula
+--+
| +-- Margaritifera
--+
| +-- Anemina **
+--+
+-- Sinanodonta **
Itoh et al., 2026 [Publ 4842], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 2 | 0 | 0 | 0 | 163.0 |
+-- Sinanodonta ** --+ +-- Beringiana **
Soroka, 2008 [Publ 3695], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 3 | 2 | 2 | -- |
+----- Sinanodonta ** --+ | +-- Anodontina +--+ +-- Alasmidontina
Soroka, 2008 [Publ 3695], fig. 1-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 3 | 2 | 2 | -- |
+----- Sinanodonta ** --+ | +-- Anodontina +--+ +-- Alasmidontina
Plazzi et al., 2011 [Publ 3676], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 1 | 1 | 0 | 1 | 17 | 4 | 2 | -- |
+----------------- Cristaria ** | | +-------------- Lanceolaria | | --+ +-------------- Anodonta | | | +-------------- Unionina | | | | +-- Alasmidontina +--+-----------+ | +-- Pseudanodonta | | +-- Alathyria | | | +--------+-- Pronodularia | | | | | +-- Sinohyriopsis +--+ | +-------- Quadrulina | | +--+-------- Parvaspina | | +----- Lampsilis +--+ | +-- Epioblasma +--+ +-- Venustaconcha
Huang et al., 2018 [Publ 3828], fig. 6-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 16 | 3 | 2 | -- |
+----- Sinanodonta ** --+ | +-- Anodontina +--+ +-- Alasmidontina
Chase et al., 2018 [Publ 3847], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 38 | 13 | 2 | -- |
+----- Sinanodonta ** --+ | +-- Anodontina +--+ +-- Alasmidontina
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.500 | 1 | 1 | 0 | 1 | 104 | 19 | 2 | -- |
+-- Alasmidonta_marginata | +-- Prolasmidonta | +-- Alasmidonta_undulata | +-- Beringiana ** | +-- Brachyanodon_kennerlyi | +-- Brachyanodon_nuttalliana | +-- Brachyanodon_oregonensis | +-- Arcidens_confragosus | +-- Arcidens_wheeleri | +-- Pterosyna | --+-- Platynaias_compressa | +-- Lasmigona | +-- Platynaias_decorata | +-- Platynaias_subviridis | +-- Pyganodon_grandis | +-- Pyganodon_lacustris | +-- Pseudodontoideus | +-- Strophitus | +-- Utterbackia_imbecillis | +-- Utterbackia_peninsularis__1 | +-- Utterbackia_peninsularis__2
Burzynski & Soroka, 2018 [Publ 3862], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 3 | 3 | 2 | -- |
+----- Sinanodonta ** --+ | +-- Anodontina +--+ +-- Alasmidontina
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 19 | 3 | 2 | -- |
+-- Sinanodonta ** | --+-- Anodontina | +-- Alasmidontina
Burzynski & Soroka, 2018 [Publ 3862], fig. 5b
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 9 | 3 | 2 | -- |
+----- Sinanodonta ** --+ | +-- Anodontina +--+ +-- Alasmidontina
Huang et al., 2019 [Publ 3843], fig. 3-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 22 | 3 | 2 | -- |
+----- Sinanodonta ** --+ | +-- Anodontina +--+ +-- Alasmidontina
Froufe et al., 2019 [Publ 3936], fig. 3-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 18 | 3 | 2 | -- |
+----- Sinanodonta ** --+ | +-- Anodontina +--+ +-- Alasmidontina
Froufe et al., 2019 [Publ 3936], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 18 | 3 | 2 | -- |
+----- Sinanodonta ** --+ | +-- Anodontina +--+ +-- Alasmidontina
Guerra et al., 2019 [Publ 4168], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 40 | 6 | 2 | -- |
+----- Sinanodonta ** --+ | +-- Anodontina +--+ +-- Alasmidontina
Guerra et al., 2019 [Publ 4168], fig. 1-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 21 | 3 | 2 | -- |
+----- Sinanodonta ** --+ | +-- Anodontina +--+ +-- Alasmidontina
Guerra et al., 2019 [Publ 4168], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 40 | 6 | 2 | -- |
+----- Sinanodonta ** --+ | +-- Anodontina +--+ +-- Alasmidontina
Guerra et al., 2019 [Publ 4168], fig. 2-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 21 | 3 | 2 | -- |
+----- Alasmidontina --+ | +-- Anodontina +--+ +-- Sinanodonta **
Zieritz et al., 2021 [Publ 4231], fig. 3-M_4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 22 | 3 | 2 | -- |
+----- Sinanodonta ** --+ | +-- Anodontina +--+ +-- Alasmidontina
Wu et al., 2022 [Publ 4238], fig. 2e
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 17 | 3 | 2 | -- |
+----- Sinanodonta ** --+ | +-- Anodontina +--+ +-- Alasmidontina
Klymus et al., 2025 [Publ 4814], fig. 3A
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 35 | 3 | 2 | -- |
+----- Sinanodonta ** --+ | +-- Anodontina +--+ +-- Alasmidontina
Klymus et al., 2025 [Publ 4814], fig. 3B
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 35 | 3 | 2 | -- |
+----- Sinanodonta ** --+ | +-- Anodontina +--+ +-- Alasmidontina
Campbell & Lydeard, 2026 [Publ 4810], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 46 | 3 | 2 | -- |
+----- Sinanodonta ** --+ | +-- Anodontina +--+ +-- Alasmidontina
Nagel et al., 1996 [Publ 3400], fig. 351
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.667 | 1 | 1 | 0 | 1 | 4 | 4 | 1 | -- |
+-------------- Pseudanodonta --+ | +----------- Anodonta_cygnea +--+ | +-------- Sinanodonta ** +--+ | +----- Anodonta__2 +--+ | +-- Anodonta_anatina +--+ +-- Anodonta__1
Nagel et al., 1998 [Publ 1946], fig. 7
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.667 | 1 | 1 | 0 | 1 | 4 | 4 | 1 | -- |
+-------------- Pseudanodonta --+ | +----------- Anodonta_cygnea +--+ | +-------- Sinanodonta ** +--+ | +----- Anodonta__2 +--+ | +-- Anodonta_anatina +--+ +-- Anodonta__1
Mock et al., 2004 [Publ 2686], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 1 | 1 | 1 | -- |
+-- Anodontina --+ +-- Sinanodonta **
Mock et al., 2004 [Publ 2686], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 1 | 1 | 1 | -- |
+-- Anodontina --+ +-- Sinanodonta **
Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 4 | 2 | 1 | -- |
+-- Beringiana ** --+ +-- Anodontina
Soroka, 2010 [Publ 3404], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.667 | 1 | 1 | 0 | 1 | 6 | 3 | 1 | -- |
+----- Sinanodonta ** --+ | +-- Unionina +--+ +-- Anodontina
Soroka, 2010 [Publ 3404], fig. 2-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.800 | 1 | 1 | 0 | 1 | 4 | 1 | 1 | -- |
+----- Sinanodonta ** --+ | +-- Anodontina +--+ +-- Unionina
Soroka, 2010 [Publ 3404], fig. 3a
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.667 | 2 | 2 | 0 | 1 | 7 | 3 | 1 | -- |
+----- Sinanodonta ** --+ | +-- Anodontina +--+ +-- Unionina
Soroka, 2010 [Publ 3404], fig. 3a-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.800 | 2 | 2 | 0 | 1 | 4 | 1 | 1 | -- |
+-- Sinanodonta ** --+ +-- Unionina
Soroka, 2010 [Publ 3404], fig. 3b
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.500 | 2 | 2 | 0 | 1 | 7 | 3 | 1 | -- |
+-- Sinanodonta ** --+ +-- Unionina
Soroka, 2010 [Publ 3404], fig. 3b-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.800 | 2 | 2 | 0 | 1 | 4 | 1 | 1 | -- |
+-- Sinanodonta ** --+ +-- Unionina
Soroka, 2010 [Publ 3404], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.500 | 1 | 1 | 0 | 1 | 6 | 3 | 1 | -- |
+-- Sinanodonta ** --+ +-- Anodontina
Soroka, 2010 [Publ 3404], fig. 4-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 1 | 1 | 1 | -- |
+-- Anodontina --+ +-- Sinanodonta **
Soroka, 2010 [Publ 3404], fig. 5a
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.667 | 2 | 2 | 0 | 1 | 6 | 3 | 1 | -- |
+-------- Anodonta_anatina --+ | +----- Sinanodonta ** +--+ | +-- Anodonta_cygnea +--+ +-- Pseudanodonta
Soroka, 2010 [Publ 3404], fig. 5a-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 1 | 1 | 1 | -- |
+-- Anodontina --+ +-- Sinanodonta **
Soroka, 2010 [Publ 3404], fig. 5b
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.500 | 2 | 2 | 0 | 1 | 6 | 3 | 1 | -- |
+-- Anodontina | --+-- Sinanodonta ** | +-- Unionina
Soroka, 2010 [Publ 3404], fig. 5b-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 1 | 1 | 1 | -- |
+-- Anodontina --+ +-- Sinanodonta **
Soroka, 2010 [Publ 3404], fig. 8a
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 5 | 2 | 1 | -- |
+-- Sinanodonta ** --+ +-- Anodontina
Soroka, 2010 [Publ 3404], fig. 8b
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.800 | 1 | 1 | 0 | 1 | 5 | 2 | 1 | -- |
+----- Sinanodonta ** --+ | +-- Anodontina +--+ +-- Unionina
Breton et al., 2011 [Publ 3376], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 37 | 11 | 1 | -- |
+-- Cristaria ** --+ +-- Alasmidontina
Chen et al., 2012 [Publ 3624], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 6 | 1 | 1 | -- |
+-- Alasmidontina --+ +-- Cristaria **
Zieritz et al., 2012 [Publ 3673], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.667 | 1 | 1 | 0 | 1 | 7 | 3 | 1 | -- |
+-- Sinanodonta ** | --+-- Margaritifera | +-- Unionina
Sheng et al., 2014 [Publ 3704], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.800 | 1 | 1 | 0 | 1 | 7 | 1 | 1 | -- |
+-------------- Cristaria ** | --+ +----------- Alasmidontina | | +--+ +-------- Lampsilis | | +--+ +-- Venustaconcha | +--+ | | +-- Quadrulina +--+ | +-- Lamprotula +--+ +-- Sinohyriopsis
Mezhzherin et al., 2014 [Publ 3705], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 3 | 3 | 1 | -- |
+-- Sinanodonta ** --+ +-- Anodontina
Mezhzherin et al., 2014 [Publ 3705], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.833 | 1 | 1 | 0 | 1 | 3 | 3 | 1 | -- |
+-- Sinanodonta ** --+ +-- Anodontina
Wei et al., 2016 [Publ 3665], fig. 0
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.800 | 1 | 1 | 0 | 1 | 5 | 1 | 1 | -- |
+-- Cristaria ** | +-- Lampsilis --+ +-- Alasmidontina | +-- Quadrulina
Amaro et al., 2017 [Publ 3791], fig. 2-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.857 | 1 | 1 | 0 | 1 | 9 | 3 | 1 | -- |
+----- Pyganodon | --+ +-- Lasmigona | | +--+-- Sinanodonta ** | +-- Utterbackia
Froufe et al., 2017 [Publ 3802], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 3 | 3 | 1 | -- |
+-- Cristaria ** --+ +-- Anodontina
Froufe et al., 2017 [Publ 3802], fig. 7
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 9 | 3 | 1 | -- |
+-- Sinanodonta ** --+ +-- Anodontina
Smith et al., 2018 [Publ 3790], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 17 | 17 | 1 | -- |
+-- Cristaria ** --+ +-- Alasmidontina
Tomilova et al., 2020 [Publ 4097], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.667 | 2 | 2 | 0 | 1 | 6 | 4 | 1 | -- |
+-- Unionina | --+-- Sinanodonta ** | +-- Anodontina
Smith, 2021 [Publ 4331], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 19 | 2 | 1 | -- |
+-- Cristaria ** --+ +-- Alasmidontina
Dai et al., 2024 [Publ 4386], fig. 5-BI
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.800 | 1 | 1 | 0 | 1 | 21 | 1 | 1 | -- |
+----------- Acuticosta | +----------- Anemina ** | --+----------- Anodontina | | +-------- Lanceolaria | | | | +-- Aculamprotulina +--+-----+ | +-- Lepidodesma | | +----- Unionina +--+ | +-- Middendorffinaiina +--+ +-- Oxynaiina
Dai et al., 2024 [Publ 4386], fig. 5-ML
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.800 | 1 | 1 | 0 | 1 | 21 | 1 | 1 | -- |
+-- Acuticosta --+ +-- Anemina **
Dai et al., 2024 [Publ 4386], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 40 | 2 | 1 | -- |
+-- Sinanodonta ** --+ +-- Alasmidontina
Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 3 | 0 | 0 | -- |
+----- Sinanodonta ** | --+----- Lampsilis | | +-- Sinohyriopsis +--+ +-- Pronodularia
Soroka, 2010 [Publ 3404], fig. 8a-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 2 | 0 | 0 | -- |
+-- Sinanodonta ** --+ +-- Unionina
Soroka, 2010 [Publ 3404], fig. 8b-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 2 | 0 | 0 | -- |
+-- Sinanodonta ** --+ +-- Unionina
Jiang et al., 2010 [Publ 3438], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 3 | 0 | 0 | -- |
+----- Lampsilis | --+----- Cristaria ** | | +-- Pronodularia +--+ +-- Sinohyriopsis
Peng et al., 2012 [Publ 3689], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 2 | 0 | 0 | -- |
+-- Cristaria ** --+ +-- Sinohyriopsis
Bolotov et al., 2016 [Publ 3732], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 6 | 6 | 5 | 1 | 2 | 0 | 0 | 167.0 |
+----- Sinanodonta ** --+ | +-- Gibbosula +--+ +-- Margaritifera
Zhou et al., 2017 [Publ 3788], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 15 | 0 | 0 | -- |
+----------------- Anemina ** | | +----- Lanceolaria --+ +--------+ | | | +-- Oxynaiina | | +--+ | | +-- Aculamprotulina +--+ | +----- Quadrulina | +-----+ | | | +-- Potamilus | | +--+ +--+ +-- Lampsilis | | +-------- Sinohyriopsis +--+ | +----- Lamprotula +--+ | +-- Ptychorhynchus +--+ +-- Sinosolenaia
Liu et al., 2017 [Publ 3939], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- |
+-- Anemina ** --+ +-- Sinosolenaia
Bulakhova et al., 2023 [Publ 4325], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 1 | 0 | 0 | 0 | 166.0 |
-- Beringiana **
