Bivalvia | Unionida | Hyrioidea

family HYRIIDAE Swainson, 1840: cladomics

2 genera, 2 subfamilies

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 94 (136 total)
number of subtaxa (rank: subfamily) = 4
analyzed in 115 cladograms in 58 publications
maximum number of terminals in an analysis = 29 (mean = 4.22, median = 3)
maximum number of subtaxa (subfamily) = 2 (mean = 1.52, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 66
maximum number of included terminals = 29, (mean = 6.61 median = 5)
maximum number of subtaxa (subfamily) = 2 (mean = 1.91, median = 2)
proportion of cladograms that support basic monophyly = 1

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Hyriidae cladogram distances
PCoA Ordination

NJ Tree of Hyriidae cladogram distances
NJ Tree

Cladograms

Graf, 2002 [Publ 2651], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 2 14 0 0 **
Cladomics cladogram VELESUNIONINAE HYRIINAE MARGARITIFERIDAE UNIONIDAE

Graf, 2000 [Publ 2647], fig. 1

cst iT tT bi riS oT roT roS iD
0.909 4 4 2 2 13 0 0 5.5
Cladomics cladogram CASTALIINI RHIPIDODONTINI HYRIDELLINI VELESUNIONINAE IRIDINIDAE ANODONTITINAE MYCETOPODINAE ETHERIIDAE

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

cst iT tT bi riS oT roT roS iD
0.833 6 6 5 2 5 0 0 3.0
Cladomics cladogram MARGARITIFERIDAE UNIONIDAE VELESUNIONINAE HYRIINAE

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

cst iT tT bi riS oT roT roS iD
0.909 5 5 3 2 29 0 0 5.5
Cladomics cladogram CASTALIINI RHIPIDODONTINI Echyridella VELESUNIONINAE IRIDINIDAE Etheria Mulleria MYCETOPODIDAE PARREYSIINAE Pseudomulleria ANODONTINI GONIDEINAE UNIONINI MARGARITIFERIDAE AMBLEMINAE

Hoeh et al., 2001 [Publ 2652], fig. 14f2

cst iT tT bi riS oT roT roS iD
0.818 5 5 1 2 26 0 0 7.5
Cladomics cladogram IRIDINIDAE MYCETOPODIDAE RHIPIDODONTINI Echyridella Lortiella CASTALIINI Velesunio

Hoeh et al., 2001 [Publ 2652], fig. 14f4

cst iT tT bi riS oT roT roS iD
0.909 5 5 3 2 26 0 0 5.5
Cladomics cladogram CASTALIINI RHIPIDODONTINI Echyridella VELESUNIONINAE IRIDINIDAE MYCETOPODIDAE PARREYSIINAE MARGARITIFERIDAE GONIDEINAE UNIONINAE AMBLEMINAE

Hoeh et al., 2002 [Publ 2653], fig. 2

cst iT tT bi riS oT roT roS iD
0.909 5 5 3 2 26 0 0 5.5
Cladomics cladogram CASTALIINI RHIPIDODONTINI Echyridella VELESUNIONINAE IRIDINIDAE MYCETOPODIDAE PARREYSIINAE MARGARITIFERIDAE AMBLEMINAE GONIDEINAE UNIONINAE

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

cst iT tT bi riS oT roT roS iD
0.818 5 5 2 2 26 0 0 7.5
Cladomics cladogram IRIDINIDAE MYCETOPODIDAE CASTALIINI Velesunio Echyridella Lortiella RHIPIDODONTINI

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

cst iT tT bi riS oT roT roS iD
0.909 5 5 3 2 26 0 0 5.5
Cladomics cladogram IRIDINIDAE MYCETOPODIDAE CASTALIINI RHIPIDODONTINI Echyridella VELESUNIONINAE

Baker et al., 2003 [Publ 2690], fig. 5

cst iT tT bi riS oT roT roS iD
0.750 7 7 5 2 1 0 0 8.5
Cladomics cladogram UNIONIDAE Hyridella australis RHIPIDODONTINI Hyridella depressa VELESUNIONINAE Echyridella CASTALIINI

Baker et al., 2004 [Publ 2685], fig. 2-3

cst iT tT bi riS oT roT roS iD
0.692 8 8 9 2 1 0 0 13.0
Cladomics cladogram UNIONIDAE Hyridella depressa Hyridella australis RHIPIDODONTINI VELESUNIONINAE Echyridella CASTALIINI

Graf & Cummings, 2006 [Publ 1968], fig. 2

cst iT tT bi riS oT roT roS iD
0.923 9 9 7 2 38 0 0 5.0
Cladomics cladogram VELESUNIONINAE HYRIINAE ETHERIIDAE MYCETOPODIDAE IRIDINIDAE

Walker et al., 2006 [Publ 2666], fig. 6

cst iT tT bi riS oT roT roS iD
0.714 10 10 7 2 26 0 0 12.5
Cladomics cladogram CASTALIINI RHIPIDODONTINI Hyridella depressa Hyridella australis Echyridella VELESUNIONINAE UNIONINAE MARGARITIFERIDAE GONIDEINAE AMBLEMINAE IRIDININAE Etheria ASPATHARIINAE Mulleria MYCETOPODIDAE

Hoeh et al., 2009 [Publ 2716], fig. 3

cst iT tT bi riS oT roT roS iD
0.923 9 9 7 2 38 0 0 5.0
Cladomics cladogram VELESUNIONINAE HYRIINAE IRIDINIDAE Etheria Mulleria MYCETOPODIDAE MARGARITIFERIDAE PARREYSIINAE Pseudomulleria UNIONINAE Rectidentini AMBLEMINAE GONIDEINI PSEUDODONTINI

Hoeh et al., 2009 [Publ 2716], fig. 4

cst iT tT bi riS oT roT roS iD
0.923 9 9 7 2 37 0 0 5.0
Cladomics cladogram VELESUNIONINAE HYRIINAE Etheria Mulleria MYCETOPODIDAE IRIDINIDAE PARREYSIINAE Pseudomulleria MARGARITIFERIDAE PSEUDODONTINI Rectidentini UNIONINAE GONIDEINI AMBLEMINAE

Hoeh et al., 2009 [Publ 2716], fig. 5

cst iT tT bi riS oT roT roS iD
0.923 9 9 7 2 37 0 0 5.0
Cladomics cladogram VELESUNIONINAE HYRIINAE Pseudomulleria Mulleria Etheria MYCETOPODIDAE IRIDINIDAE

Whelan et al., 2011 [Publ 3380], fig. 1-ML

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 2 32 0 0 3.5
Cladomics cladogram VELESUNIONINAE HYRIINAE MARGARITIFERIDAE UNIONIDAE ASPATHARIINAE IRIDININAE MYCETOPODIDAE

Whelan et al., 2011 [Publ 3380], fig. 1-MP

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 2 32 0 0 3.5
Cladomics cladogram MARGARITIFERIDAE VELESUNIONINAE HYRIINAE

Marshall et al., 2014 [Publ 3593], fig. 1

cst iT tT bi riS oT roT roS iD
0.833 10 10 8 2 4 0 0 9.0
Cladomics cladogram UNIONIDAE Echyridella VELESUNIONINAE HYRIDELLINI RHIPIDODONTINI

Graf et al., 2015 [Publ 3619], fig. 1

cst iT tT bi riS oT roT roS iD
0.889 14 14 12 2 6 0 0 11.5
Cladomics cladogram VELESUNIONINAE HYRIINAE UNIONIDAE MARGARITIFERIDAE IRIDINIDAE MYCETOPODIDAE ETHERIIDAE

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 2 36 0 0 3.5
Cladomics cladogram VELESUNIONINAE HYRIINAE MARGARITIFERIDAE UNIONIDAE ETHERIIDAE MYCETOPODIDAE IRIDINIDAE

Bolotov et al., 2016 [Publ 3743], fig. 2

cst iT tT bi riS oT roT roS iD
0.929 9 9 7 2 21 0 0 8.5
Cladomics cladogram HYRIINAE VELESUNIONINAE UNIONIDAE MARGARITIFERIDAE MYCETOPODIDAE ETHERIIDAE IRIDINIDAE

Bolotov et al., 2016 [Publ 3743], fig. 4

cst iT tT bi riS oT roT roS iD
0.929 9 9 7 2 21 0 0 8.5
Cladomics cladogram HYRIINAE VELESUNIONINAE UNIONIDAE MARGARITIFERIDAE MYCETOPODIDAE ETHERIIDAE IRIDINIDAE

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

cst iT tT bi riS oT roT roS iD
0.696 20 20 20 2 1 0 0 23.5
Cladomics cladogram TRIGONIIDAE Guyanhyrida HYRIINI CASTALIINI Westralunio Diplodon Hyridella drapeta Hyridella depressa Hyridella australis Cucumerunio Echyridella Velesunio ambiguus Microdontia Lortiella Alathyria Velesunio angasi

Santos-Neto et al., 2016 [Publ 3739], fig. 4

cst iT tT bi riS oT roT roS iD
0.696 20 20 21 2 1 0 0 24.5
Cladomics cladogram TRIGONIIDAE Guyanhyrida HYRIINI CASTALIINI Westralunio Diplodon Hyridella drapeta Hyridella depressa Echyridella Cucumerunio Hyridella australis Velesunio ambiguus Alathyria Velesunio angasi Lortiella Microdontia

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 79 0 0 --
Cladomics cladogram HYRIINAE VELESUNIONINAE MARGARITIFERIDAE UNIONIDAE IRIDININAE ASPATHARIINAE ETHERIIDAE MYCETOPODIDAE

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
0.929 9 9 7 2 130 0 0 8.5
Cladomics cladogram HYRIINAE VELESUNIONINAE MYCETOPODIDAE ETHERIIDAE IRIDINIDAE

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
0.929 9 9 7 2 130 0 0 8.5
Cladomics cladogram HYRIINAE VELESUNIONINAE IRIDINIDAE MYCETOPODIDAE ETHERIIDAE

Combosch et al., 2017 [Publ 3806], fig. 1

cst iT tT bi riS oT roT roS iD
0.889 3 3 1 2 210 0 0 4.0
Cladomics cladogram HYRIINI VELESUNIONINAE HYRIDELLINI UNIONIDAE MARGARITIFERIDAE ASPATHARIINAE ETHERIIDAE IRIDININAE MYCETOPODIDAE

Combosch et al., 2017 [Publ 3806], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 211 0 0 4.0
Cladomics cladogram VELESUNIONINAE HYRIINAE MARGARITIFERIDAE UNIONIDAE

Huang et al., 2018 [Publ 3828], fig. 3_5

cst iT tT bi riS oT roT roS iD
0.889 3 3 1 2 26 0 0 4.0
Cladomics cladogram HYRIINI VELESUNIONINAE HYRIDELLINI UNIONIDAE MARGARITIFERIDAE IRIDININAE ASPATHARIINAE ETHERIIDAE MYCETOPODIDAE

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

cst iT tT bi riS oT roT roS iD
0.889 3 3 1 2 25 0 0 4.0
Cladomics cladogram HYRIINI VELESUNIONINAE HYRIDELLINI UNIONIDAE MARGARITIFERIDAE IRIDININAE ASPATHARIINAE ETHERIIDAE MYCETOPODIDAE

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

cst iT tT bi riS oT roT roS iD
0.889 3 3 1 2 25 0 0 4.0
Cladomics cladogram HYRIINI VELESUNIONINAE HYRIDELLINI UNIONIDAE MARGARITIFERIDAE IRIDININAE ASPATHARIINAE ETHERIIDAE MYCETOPODIDAE

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
0.889 14 14 12 2 187 0 0 11.5
Cladomics cladogram Westralunio HYRIINAE Alathyria jacksoni Alathyria profuga Velesunio Alathyria pertexta Lortiella Microdontia MARGARITIFERIDAE UNIONIDAE Anodontites IRIDININAE ETHERIIDAE ASPATHARIINAE MYCETOPODINAE MONOCONDYLAEINAE Lamproscapha

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
0.889 14 14 12 2 185 0 0 11.5
Cladomics cladogram Westralunio HYRIINAE Velesunio Alathyria jacksoni Alathyria profuga Alathyria pertexta Lortiella Microdontia MARGARITIFERIDAE UNIONIDAE Anodontites IRIDININAE ETHERIIDAE ASPATHARIINAE MYCETOPODINAE MONOCONDYLAEINAE Lamproscapha

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.833 14 14 9 2 187 0 0 12.5
Cladomics cladogram MARGARITIFERIDAE UNIONIDAE Echyridella Westralunio HYRIDELLINI RHIPIDODONTINI CASTALIINI HYRIINI Velesunio Alathyria jacksoni Alathyria profuga Alathyria pertexta Lortiella Microdontia

Guerra et al., 2019 [Publ 4168], fig. 1

cst iT tT bi riS oT roT roS iD
0.917 5 5 3 2 36 0 0 5.5
Cladomics cladogram MARGARITIFERIDAE UNIONIDAE Echyridella VELESUNIONINAE RHIPIDODONTINI CASTALIINI HYRIINI

Guerra et al., 2019 [Publ 4168], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 20 0 0 --
Cladomics cladogram HYRIINAE VELESUNIONINAE MARGARITIFERIDAE UNIONIDAE

Guerra et al., 2019 [Publ 4168], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 2 36 0 0 5.5
Cladomics cladogram VELESUNIONINAE HYRIINAE MARGARITIFERIDAE UNIONIDAE

Guerra et al., 2019 [Publ 4168], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 20 0 0 --
Cladomics cladogram HYRIINAE VELESUNIONINAE MARGARITIFERIDAE UNIONIDAE

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 120 0 0 --
Cladomics cladogram HYRIINAE VELESUNIONINAE MARGARITIFERIDAE UNIONIDAE MYCETOPODIDAE ETHERIIDAE IRIDINIDAE

Wu et al., 2020 [Publ 4121], fig. 2

cst iT tT bi riS oT roT roS iD
0.889 3 3 1 2 31 0 0 4.0
Cladomics cladogram HYRIINI HYRIDELLINI VELESUNIONINAE UNIONIDAE MARGARITIFERIDAE IRIDININAE ASPATHARIINAE ETHERIIDAE MYCETOPODIDAE

Wu et al., 2020 [Publ 4121], fig. S1

cst iT tT bi riS oT roT roS iD
0.889 3 3 1 2 31 0 0 4.0
Cladomics cladogram HYRIINI HYRIDELLINI VELESUNIONINAE UNIONIDAE MARGARITIFERIDAE IRIDININAE ASPATHARIINAE ETHERIIDAE MYCETOPODIDAE

Sano et al., 2020 [Publ 4138], fig. 2

cst iT tT bi riS oT roT roS iD
0.889 3 3 1 2 66 0 0 4.0
Cladomics cladogram HYRIINI HYRIDELLINI VELESUNIONINAE MARGARITIFERIDAE UNIONIDAE IRIDININAE ASPATHARIINAE ETHERIIDAE MYCETOPODIDAE

Sano et al., 2020 [Publ 4138], fig. 3

cst iT tT bi riS oT roT roS iD
0.889 3 3 1 2 66 0 0 4.0
Cladomics cladogram HYRIINI HYRIDELLINI VELESUNIONINAE MARGARITIFERIDAE UNIONIDAE IRIDININAE ASPATHARIINAE ETHERIIDAE MYCETOPODIDAE

Sano et al., 2020 [Publ 4138], fig. 4

cst iT tT bi riS oT roT roS iD
0.889 3 3 1 2 66 0 0 4.0
Cladomics cladogram HYRIINI HYRIDELLINI VELESUNIONINAE MARGARITIFERIDAE UNIONIDAE IRIDININAE ASPATHARIINAE ETHERIIDAE MYCETOPODIDAE

Klunzinger et al., 2021 [Publ 4086], fig. 2

cst iT tT bi riS oT roT roS iD
0.875 4 4 2 2 2 0 0 5.0
Cladomics cladogram MARGARITIFERIDAE UNIONIDAE Westralunio Velesunio HYRIINAE

Sonowal et al., 2021 [Publ 4221], fig. 3a

cst iT tT bi riS oT roT roS iD
0.889 3 3 0 2 12 0 0 4.0
Cladomics cladogram TRIGONIIDAE HYRIINI HYRIDELLINI VELESUNIONINAE

Sonowal et al., 2021 [Publ 4221], fig. 3b

cst iT tT bi riS oT roT roS iD
0.889 3 3 0 2 12 0 0 4.0
Cladomics cladogram UNIONIDAE HYRIINI VELESUNIONINAE HYRIDELLINI

Sonowal et al., 2021 [Publ 4221], fig. 4

cst iT tT bi riS oT roT roS iD
0.889 3 3 1 2 13 0 0 4.0
Cladomics cladogram UNIONIDAE HYRIINI VELESUNIONINAE HYRIDELLINI

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 88 0 0 3.5
Cladomics cladogram VELESUNIONINAE HYRIINAE MARGARITIFERIDAE UNIONIDAE IRIDINIDAE ETHERIIDAE MYCETOPODIDAE

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 88 0 0 3.5
Cladomics cladogram VELESUNIONINAE HYRIINAE MARGARITIFERIDAE UNIONIDAE

Klunzinger et al., 2022 [Publ 4259], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 2 0 0 0 5.0
Cladomics cladogram HYRIINAE VELESUNIONINAE

Gomes-dos-Santos et al., 2023 [Publ 4463], fig. 2a

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 19 0 0 --
Cladomics cladogram HYRIINAE VELESUNIONINAE UNIONIDAE MARGARITIFERIDAE

Gomes-dos-Santos et al., 2023 [Publ 4463], fig. 2b

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 20 0 0 --
Cladomics cladogram VELESUNIONINAE HYRIINAE UNIONIDAE MARGARITIFERIDAE

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 2 135 0 0 5.5
Cladomics cladogram VELESUNIONINAE HYRIINAE MARGARITIFERIDAE UNIONIDAE

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 45 0 0 --
Cladomics cladogram HYRIINAE VELESUNIONINAE MARGARITIFERIDAE UNIONIDAE

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.652 29 29 34 2 604 0 0 31.5
Cladomics cladogram Westralunio Guyanhyrida HYRIINI CASTALIINI Diplodon Hyridella depressa Hyridella drapeta Cucumerunio Hyridella australis Echyridella Velesunio 1 Alathyria jacksoni 1 Velesunio ambiguus Alathyria profuga Velesunio 3 Microdontia Lortiella Velesunio angasi Alathyria pertexta 2 Velesunio 2 Alathyria jacksoni 2 Alathyria pertexta 1 UNIONINAE Lamellidens Balwantia Trapezidens LEOPARREYSIINI ETHERIIDAE INDOCHINELLINI PARREYSIINI COELATURINI Rectidentini MARGARITIFERIDAE AMBLEMINAE INDOPSEUDODONTINA Lannanaia Pilsbryoconcha Songkhlanaia Nyeinchanconcha Sundadontina Isannaia Namkongnaia Thaiconcha Bineurus Pseudodon walpolei Pseudodon vagulus Pseudodon vondembuschianus Pseudodon mekongi SCHEPMANIINI Pseudodon lenyanensis 2 Pseudodon inoscularis Pseudodon lenyanensis 1 CHAMBERLAINIINI Parvasolenaia Pseudopostolata Koreosolenaia Cosmopseudodon Guiunio Ptychorhynchus Obovalis Postolata Caudiculatus LAMPROTULINI Sinosolenaia carinata Gonidea Microcondylaea Leguminaia Inversidens Sinosolenaia recognita Sinosolenaia iridinea Sinosolenaia oleivora

Lawrie et al., 2026 [Publ 4843], fig. 1

cst iT tT bi riS oT roT roS iD
0.750 17 17 21 2 1 0 0 23.5
Cladomics cladogram UNIONIDAE HYRIDELLINI VELESUNIONINAE CASTALIINI HYRIINI

Prié et al., 2026 [Publ 4844], fig. 2

cst iT tT bi riS oT roT roS iD
0.833 20 20 15 2 3 0 0 18.5
Cladomics cladogram MARGARITIFERIDAE Echyridella Westralunio HYRIDELLINI Velesunio Alathyria Lortiella Microdontia Diplodon Guyanhyrida CASTALIINI Iridea HYRIINI

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 0 0 --
Cladomics cladogram HYRIINAE MARGARITIFERIDAE UNIONIDAE

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 0 0 --
Cladomics cladogram HYRIINAE MARGARITIFERIDAE UNIONIDAE

Hoeh et al., 2002 [Publ 2673], fig. 1-F

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 0 0 --
Cladomics cladogram HYRIINAE UNIONIDAE

Hoeh et al., 2002 [Publ 2673], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 0 0 --
Cladomics cladogram HYRIINAE UNIONIDAE

Walker et al., 2006 [Publ 2666], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram HYRIINAE UNIONINAE LAMPROTULINI GONIDEINI PSEUDODONTINI LAMPSILINI PLEUROBEMINI MARGARITIFERIDAE

Walker et al., 2006 [Publ 2666], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram HYRIINAE MARGARITIFERIDAE UNIONIDAE

Walker et al., 2006 [Publ 2666], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram HYRIINAE MARGARITIFERIDAE UNIONIDAE

Walker et al., 2006 [Publ 2666], fig. 6-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram HYRIINAE MARGARITIFERIDAE UNIONIDAE

Breton et al., 2011 [Publ 3376], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 37 0 0 --
Cladomics cladogram HYRIINAE MARGARITIFERIDAE UNIONIDAE

Plazzi et al., 2011 [Publ 3676], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 0 0 --
Cladomics cladogram VELESUNIONINAE LAMPROTULINI CHAMBERLAINIINI

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 --
Cladomics cladogram MARGARITIFERIDAE VELESUNIONINAE UNIONIDAE

Bieler et al., 2014 [Publ 3622], fig. 26

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram VELESUNIONINAE UNIONIDAE IRIDINIDAE MARGARITIFERIDAE

Bieler et al., 2014 [Publ 3622], fig. 27

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram VELESUNIONINAE UNIONIDAE IRIDINIDAE MARGARITIFERIDAE

Bieler et al., 2014 [Publ 3622], fig. 28

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram UNIONIDAE VELESUNIONINAE

Bieler et al., 2014 [Publ 3622], fig. 29

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram VELESUNIONINAE UNIONIDAE IRIDINIDAE MARGARITIFERIDAE

Bieler et al., 2014 [Publ 3622], fig. 30

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram VELESUNIONINAE UNIONIDAE IRIDINIDAE MARGARITIFERIDAE

Bieler et al., 2014 [Publ 3622], fig. 31

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram VELESUNIONINAE UNIONIDAE MARGARITIFERIDAE IRIDINIDAE

González & Giribet, 2015 [Publ 3632], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram UNIONIDAE VELESUNIONINAE

González & Giribet, 2015 [Publ 3632], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram UNIONIDAE VELESUNIONINAE IRIDINIDAE MARGARITIFERIDAE

González & Giribet, 2015 [Publ 3632], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram VELESUNIONINAE UNIONIDAE MARGARITIFERIDAE

González & Giribet, 2015 [Publ 3632], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram VELESUNIONINAE UNIONIDAE IRIDINIDAE MARGARITIFERIDAE

Elderkin et al., 2016 [Publ 3693], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 2 0 0 --
Cladomics cladogram ETHERIIDAE IRIDINIDAE VELESUNIONINAE

Santos-Neto et al., 2016 [Publ 3739], fig. 3_5

cst iT tT bi riS oT roT roS iD
0.917 7 7 6 1 0 0 0 9.5
Cladomics cladogram HYRIINAE

Guerra et al., 2017 [Publ 3772], fig. 2a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 24 0 0 --
Cladomics cladogram HYRIINAE MARGARITIFERIDAE UNIONIDAE

Guerra et al., 2017 [Publ 3772], fig. 2a-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 0 0 --
Cladomics cladogram HYRIINAE MARGARITIFERIDAE UNIONIDAE

Guerra et al., 2017 [Publ 3772], fig. 2b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 24 0 0 --
Cladomics cladogram HYRIINAE MARGARITIFERIDAE UNIONIDAE

Guerra et al., 2017 [Publ 3772], fig. 2b-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 0 0 --
Cladomics cladogram HYRIINAE MARGARITIFERIDAE UNIONIDAE

Konopleva et al., 2017 [Publ 3807], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 28 0 0 4.5
Cladomics cladogram HYRIINAE MARGARITIFERIDAE UNIONIDAE MYCETOPODIDAE ETHERIIDAE IRIDINIDAE

Huang et al., 2018 [Publ 3828], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 41 0 0 --
Cladomics cladogram HYRIINAE MARGARITIFERIDAE UNIONIDAE

Huang et al., 2018 [Publ 3828], fig. 6-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 16 0 0 --
Cladomics cladogram HYRIINAE MARGARITIFERIDAE UNIONIDAE

Chase et al., 2018 [Publ 3847], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 38 0 0 --
Cladomics cladogram HYRIINAE UNIONIDAE

Chase et al., 2018 [Publ 3847], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 0 0 --
Cladomics cladogram MARGARITIFERIDAE HYRIINAE

Chase et al., 2018 [Publ 3861], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 0 0 --
Cladomics cladogram HYRIINAE MARGARITIFERIDAE UNIONIDAE

Chase et al., 2018 [Publ 3861], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 0 0 --
Cladomics cladogram MARGARITIFERIDAE HYRIINAE

Santos-Neto et al., 2018 [Publ 3888], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 1 0 0 0 7.5
Cladomics cladogram HYRIINAE

Huang et al., 2019 [Publ 3843], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 55 0 0 --
Cladomics cladogram HYRIINAE MARGARITIFERIDAE UNIONIDAE

Huang et al., 2019 [Publ 3843], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 0 0 --
Cladomics cladogram HYRIINAE MARGARITIFERIDAE UNIONIDAE

Wu et al., 2019 [Publ 3864], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 45 0 0 --
Cladomics cladogram HYRIINAE IRIDINIDAE

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 0 0 --
Cladomics cladogram HYRIINAE MARGARITIFERIDAE UNIONIDAE

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 0 0 --
Cladomics cladogram HYRIINAE MARGARITIFERIDAE

Wu et al., 2020 [Publ 4121], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 0 0 --
Cladomics cladogram HYRIINAE MYCETOPODIDAE IRIDINIDAE MARGARITIFERIDAE UNIONIDAE

Olivera-Hyde et al., 2020 [Publ 4249], fig. 10

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 0 0 0 5.5
Cladomics cladogram CASTALIINI

Wu et al., 2021 [Publ 4366], fig. 3-BI

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 72 0 0 --
Cladomics cladogram HYRIINAE MYCETOPODIDAE IRIDINIDAE MARGARITIFERIDAE UNIONIDAE

Wu et al., 2021 [Publ 4366], fig. 3-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 72 0 0 --
Cladomics cladogram HYRIINAE MYCETOPODIDAE IRIDINIDAE MARGARITIFERIDAE UNIONIDAE

Wu et al., 2021 [Publ 4366], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 67 0 0 --
Cladomics cladogram HYRIINAE IRIDINIDAE MYCETOPODIDAE MARGARITIFERIDAE UNIONIDAE

Wu et al., 2022 [Publ 4238], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 72 0 0 --
Cladomics cladogram HYRIINAE MARGARITIFERIDAE UNIONIDAE

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 0 0 --
Cladomics cladogram HYRIINAE MYCETOPODIDAE IRIDINIDAE MARGARITIFERIDAE UNIONIDAE

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 0 0 --
Cladomics cladogram HYRIINAE MYCETOPODIDAE IRIDINIDAE MARGARITIFERIDAE UNIONIDAE

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 0 0 --
Cladomics cladogram HYRIINAE MYCETOPODIDAE IRIDINIDAE MARGARITIFERIDAE UNIONIDAE

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 0 0 --
Cladomics cladogram HYRIINAE MYCETOPODIDAE IRIDINIDAE MARGARITIFERIDAE UNIONIDAE

Dai et al., 2025 [Publ 4440], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 86 0 0 --
Cladomics cladogram HYRIINAE MARGARITIFERIDAE UNIONIDAE

Chen et al., 2025 [Publ 4448], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 84 0 0 --
Cladomics cladogram HYRIINAE MARGARITIFERIDAE

Chen et al., 2025 [Publ 4455], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 24 0 0 --
Cladomics cladogram HYRIINAE UNIONIDAE MARGARITIFERIDAE IRIDINIDAE ETHERIIDAE MYCETOPODIDAE

Lawrie et al., 2026 [Publ 4843], fig. 2

cst iT tT bi riS oT roT roS iD
0.714 9 9 13 1 0 0 0 22.0
Cladomics cladogram VELESUNIONINAE

Prié et al., 2026 [Publ 4844], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram HYRIINAE MARGARITIFERIDAE