Bivalvia | Unionida | Hyrioidea | Hyriidae

subfamily HYRIINAE Swainson, 1840: cladomics

4 tribes, 1 genus

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 76 (82 total)
number of subtaxa (rank: tribe) = 5
analyzed in 96 cladograms in 50 publications
maximum number of terminals in an analysis = 18 (mean = 2.83, median = 2)
maximum number of subtaxa (tribe) = 5 (mean = 2.18, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 54
maximum number of included terminals = 18, (mean = 4.26 median = 3)
maximum number of subtaxa (tribe) = 5 (mean = 3.09, median = 3)
proportion of cladograms that support basic monophyly = 0.48

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 49
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (tribe) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 18 (mean = 4.29, median = 3)
maximum number of subtaxa (tribe) = 5 (mean = 3.18, median = 3)
proportion of cladograms that support rigorous monophyly = 0.43

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Hyriinae cladogram distances
PCoA Ordination

NJ Tree of Hyriinae cladogram distances
NJ Tree

Cladograms

Huang et al., 2019 [Publ 3843], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cstiTtTbiriSoTroTroSiD
0.882 7 7 5 5 194 7 1 ** 
     +-------------- Westralunio       
  +--+                                 
  |  |  +----------- Echyridella **       
  |  +--+                              
  |     |  +-------- Hyridellini **       
  |     +--+                           
  |        |  +----- Rhipidodontini **    
  |        +--+                        
--+           |  +-- Castaliini **        
  |           +--+                     
  |              +-- Hyriini **           
  |                                    
  |           +----- Alathyria_jacksoni
  |        +--+                        
  |        |  |  +-- Alathyria_profuga 
  |        |  +--+                     
  +--------+     +-- Velesunio         
           |                           
           |  +----- Alathyria_pertexta
           +--+                        
              |  +-- Lortiella         
              +--+                     
                 +-- Microdontia

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

cstiTtTbiriSoTroTroSiD
0.619 13 19 20 5 7 6 1 21.0 
  +----------------------- Neotrigonia           
  |                                              
  |                 +----- Diplodon_suavidicus__1 **
  |  +--------------+                            
  |  |              |  +-- Hyriini **               
  |  |              +--+                         
--+  |                 +-- Castaliini **            
  |  |                                           
  |  |           +-------- Westralunio           
  |  |           |                               
  |  +-----------+     +-- Hyridella_drapeta **     
  |  |           |  +--+                         
  +--+           |  |  +-- Hyridella_depressa **    
     |           +--+                            
     |              |  +-- Diplodon_gratus **       
     |              +--+                         
     |                 +-- Diplodon_suavidicus__2 **
     |                                           
     |                 +-- Hyridella_australis **   
     |  +--------------+                         
     |  |              +-- Cucumerunio **           
     +--+                                        
        |  +-------------- Echyridella **           
        +--+                                     
           |  +----------- Velesunio_ambiguus    
           +--+                                  
              |  +-------- Microdontia           
              +--+                               
                 |  +----- Lortiella             
                 +--+                            
                    |  +-- Alathyria             
                    +--+                         
                       +-- Velesunio_angasi

Santos-Neto et al., 2016 [Publ 3739], fig. 4

cstiTtTbiriSoTroTroSiD
0.619 13 19 21 5 7 6 1 22.0 
  +-------------------------- Neotrigonia           
  |                                                 
  |                    +----- Diplodon_suavidicus__2 **
  |  +-----------------+                            
--+  |                 |  +-- Hyriini **               
  |  |                 +--+                         
  |  |                    +-- Castaliini **            
  |  |                                              
  |  |              +-------- Westralunio           
  +--+              |                               
     |  +-----------+     +-- Hyridella_drapeta **     
     |  |           |  +--+                         
     |  |           |  |  +-- Hyridella_depressa **    
     |  |           +--+                            
     |  |              |  +-- Diplodon_suavidicus__1 **
     +--+              +--+                         
        |                 +-- Diplodon_gratus **       
        |                                           
        |  +----------------- Echyridella **           
        |  |                                        
        +--+              +-- Cucumerunio **           
           |  +-----------+                         
           |  |           +-- Hyridella_australis **   
           +--+                                     
              |  +----------- Velesunio_ambiguus    
              +--+                                  
                 |  +-------- Alathyria             
                 +--+                               
                    |  +----- Velesunio_angasi      
                    +--+                            
                       |  +-- Lortiella             
                       +--+                         
                          +-- Microdontia

Huang et al., 2019 [Publ 3843], fig. S3

cstiTtTbiriSoTroTroSiD
0.882 7 7 5 5 192 7 1 0 
     +-------------- Westralunio       
  +--+                                 
  |  |  +----------- Echyridella **       
  |  +--+                              
  |     |  +-------- Hyridellini **       
  |     +--+                           
  |        |  +----- Rhipidodontini **    
  |        +--+                        
--+           |  +-- Castaliini **        
  |           +--+                     
  |              +-- Hyriini **           
  |                                    
  |           +----- Velesunio         
  |        +--+                        
  |        |  |  +-- Alathyria_jacksoni
  |        |  +--+                     
  +--------+     +-- Alathyria_profuga 
           |                           
           |  +----- Alathyria_pertexta
           +--+                        
              |  +-- Lortiella         
              +--+                     
                 +-- Microdontia

Huang et al., 2019 [Publ 3843], fig. S4

cstiTtTbiriSoTroTroSiD
0.824 7 14 9 5 194 7 1 8.0 
           +----------- Echyridella **             
           |                                    
           +----------- Westralunio             
           |                                    
           +----------- Hyridellini **             
           |                                    
           |     +----- Rhipidodontini **          
           +-----+                              
  +--------+     |  +-- Castaliini **              
  |        |     +--+                           
  |        |        +-- Hyriini **                 
  |        |                                    
  |        |     +----- Velesunio               
  |        |  +--+                              
  |        |  |  |  +-- Alathyria_jacksoni      
  |        |  |  +--+                           
  |        +--+     +-- Alathyria_profuga       
  |           |                                 
  |           |  +----- Alathyria_pertexta      
--+           +--+                              
  |              |  +-- Lortiella               
  |              +--+                           
  |                 +-- Microdontia             
  |                                             
  |              +----- Gibbosula_rochechouartii
  |  +-----------+                              
  |  |           |  +-- Gibbosula_laosensis     
  |  |           +--+                           
  |  |              +-- Margaritiferinae        
  |  |                                          
  |  |  +-------------- Parreysiinae            
  +--+  |                                       
     |  |  +----------- Chamberlainia           
     |  |  |                                    
     |  |  +----------- Megalonaias             
     |  |  |                                    
     |  |  +----------- Uniomerus               
     |  |  |                                    
     +--+  +----------- Quadrulina              
        |  |                                    
        |  +----------- Sinohyriopsis           
        |  |                                    
        |  +----------- Lamprotulini            
        |  |                                    
        |  +----------- Gonideini               
        |  |                                    
        +--+----------- Pseudodontini           
           |                                    
           +----------- Rectidentini            
           |                                    
           +----------- Unioninae               
           |                                    
           |  +-------- Amblema                 
           |  |                                 
           |  +-------- Toxolasma               
           |  |                                 
           |  +-------- Pleurobemini            
           |  |                                 
           |  |     +-- Obliquaria              
           |  +-----+                           
           |  |     +-- Plectomerus             
           +--+                                 
              |  +----- Ellipsaria              
              |  |                              
              |  +----- Potamilus_leptodon      
              |  |                              
              |  +----- Medionidus              
              |  |                              
              |  +----- Potamilus_alatus        
              |  |                              
              |  +----- Ptychobranchus          
              +--+                              
                 +----- Truncilla               
                 |                              
                 +----- Venustaconcha           
                 |                              
                 +----- Cambarunio              
                 |                              
                 +----- Leaunio                 
                 |                              
                 |  +-- Ortmanniana             
                 +--+                           
                    +-- Lampsilis

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cstiTtTbiriSoTroTroSiD
0.571 18 26 32 5 614 10 1 33.0 
  +----------------------------------- Westralunio           
  |                                                          
  |                             +----- Diplodon_suavidicus__2 **
  |  +--------------------------+                            
--+  |                          |  +-- Hyriini **               
  |  |                          +--+                         
  |  |                             +-- Castaliini **            
  |  |                                                       
  |  |                             +-- Hyridella_depressa **    
  +--+                       +-----+                         
     |                       |     +-- Hyridella_drapeta **     
     |  +--------------------+                               
     |  |                    |  +----- Diplodon_suavidicus__1 **
     |  |                    +--+                            
     |  |                       |  +-- Diplodon_chilensis **    
     |  |                       +--+                         
     +--+                          +-- Diplodon_gratus **       
        |                                                    
        |                          +-- Cucumerunio **           
        |  +-----------------------+                         
        |  |                       +-- Hyridella_australis **   
        |  |                                                 
        +--+  +----------------------- Echyridella **           
           |  |                                              
           |  |              +-------- Velesunio__1          
           +--+  +-----------+                               
              |  |           |  +----- Alathyria_jacksoni__1 
              |  |           +--+                            
              |  |              |  +-- Velesunio_ambiguus    
              +--+              +--+                         
                 |                 +-- Alathyria_profuga     
                 |                                           
                 |  +----------------- Velesunio__3          
                 |  |                                        
                 +--+  +-------------- Microdontia           
                    |  |                                     
                    +--+           +-- Lortiella             
                       |  +--------+                         
                       |  |        +-- Velesunio_angasi      
                       +--+                                  
                          |  +-------- Alathyria_pertexta__2 
                          +--+                               
                             |  +----- Velesunio__2          
                             +--+                            
                                |  +-- Alathyria_jacksoni__2 
                                +--+                         
                                   +-- Alathyria_pertexta__1

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

cstiTtTbiriSoTroTroSiD
0.818 5 5 4 4 6 1 1 8.5 
  +-------------- Velesunioninae        
  |                                     
--+           +-- Rhipidodontini **        
  |  +--------+                         
  |  |        +-- Castaliini **            
  +--+                                  
     |  +-------- Hyridella_australis__2 **
     +--+                               
        |  +----- Hyridella_australis__1 **
        +--+                            
           |  +-- Echyridella **           
           +--+                         
              +-- Hyridella_depressa **

Baker et al., 2003 [Publ 2690], fig. 5

cstiTtTbiriSoTroTroSiD
0.727 5 7 5 4 3 2 1 12.0 
  +-------------- Ambleminae         
  |                                  
--+           +-- Hyridella_australis **
  |  +--------+                      
  |  |        +-- Rhipidodontini **     
  +--+                               
     |  +-------- Hyridella_depressa ** 
     +--+                            
        |  +----- Velesunioninae     
        +--+                         
           |  +-- Echyridella **        
           +--+                      
              +-- Castaliini **

Baker et al., 2004 [Publ 2685], fig. 2-3

cstiTtTbiriSoTroTroSiD
0.667 5 8 9 4 4 3 1 16.0 
  +----------- Ambleminae         
  |                               
  |     +----- Hyridella_depressa ** 
--+  +--+                         
  |  |  |  +-- Hyridella_australis **
  |  |  +--+                      
  +--+     +-- Rhipidodontini **     
     |                            
     |  +----- Velesunioninae     
     +--+                         
        |  +-- Echyridella **        
        +--+                      
           +-- Castaliini **

Graf & Cummings, 2006 [Publ 1968], fig. 2

cstiTtTbiriSoTroTroSiD
0.917 6 6 4 4 41 3 1 4.5 
  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Hyridellini **   
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Rhipidodontini **

Walker et al., 2006 [Publ 2666], fig. 6

cstiTtTbiriSoTroTroSiD
0.692 5 10 7 4 31 5 1 15.0 
              +-------- Castaliini **         
              |                            
              +-------- Rhipidodontini **     
  +-----------+                            
  |           |  +----- Hyridella_depressa ** 
  |           |  |                         
  |           +--+----- Hyridella_australis **
  |              |                         
  |              |  +-- Echyridella **        
  |              +--+                      
--+                 +-- Velesunioninae     
  |                                        
  |           +-------- Unioninae          
  |  +--------+                            
  |  |        |  +----- Margaritiferinae   
  |  |        +--+                         
  |  |           |  +-- Gonideinae         
  +--+           +--+                      
     |              +-- Ambleminae         
     |                                     
     |  +-------------- Iridininae         
     +--+                                  
        |  +----------- Etheria            
        +--+                               
           |  +-------- Aspathariinae      
           +--+                            
              |  +----- Mulleria           
              +--+                         
                 |  +-- Anodontitinae      
                 +--+                      
                    +-- Monocondylaeinae

Hoeh et al., 2009 [Publ 2716], fig. 3

cstiTtTbiriSoTroTroSiD
0.917 6 6 4 4 41 3 1 4.5 
  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Hyridellini **   
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Rhipidodontini **

Hoeh et al., 2009 [Publ 2716], fig. 4

cstiTtTbiriSoTroTroSiD
0.917 6 6 4 4 40 3 1 4.5 
  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Hyridellini **   
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Rhipidodontini **

Hoeh et al., 2009 [Publ 2716], fig. 5

cstiTtTbiriSoTroTroSiD
0.917 6 6 4 4 40 3 1 4.5 
  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Hyridellini **   
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Rhipidodontini **

Graf et al., 2015 [Publ 3619], fig. 1

cstiTtTbiriSoTroTroSiD
0.882 7 7 5 4 13 7 1 2.0 
  +----------- Velesunioninae
--+                          
  |  +-------- Hyridellini **   
  +--+                       
     |  +----- Rhipidodontini **
     +--+                    
        |  +-- Hyriini **       
        +--+                 
           +-- Castaliini **

Guerra et al., 2019 [Publ 4168], fig. 1

cstiTtTbiriSoTroTroSiD
0.909 4 5 3 4 37 1 1 5.0 
           +-- Echyridella **     
     +-----+                   
     |     +-- Velesunioninae  
  +--+                         
  |  |  +----- Rhipidodontini **  
  |  +--+                      
  |     |  +-- Castaliini **      
  |     +--+                   
--+        +-- Hyriini **         
  |                            
  |        +-- Gibbosulinae    
  |  +-----+                   
  |  |     +-- Margaritiferinae
  +--+                         
     |  +----- Ambleminae      
     +--+                      
        |  +-- Gonideinae      
        +--+                   
           +-- Unioninae

Guerra et al., 2019 [Publ 4168], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 4 4 2 4 37 1 1 4.0 
  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Rhipidodontini **
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Hyriini **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cstiTtTbiriSoTroTroSiD
1.000 4 4 2 4 136 1 1 4.0 
  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Rhipidodontini **
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Hyriini **

Graf, 2000 [Publ 2647], fig. 1

cstiTtTbiriSoTroTroSiD
0.900 3 4 2 3 14 1 1 8.5 
     +-------- Castaliini **    
  +--+                       
  |  |  +----- Rhipidodontini **
  |  +--+                    
  |     |  +-- Hyridellini **   
--+     +--+                 
  |        +-- Velesunioninae
  |                          
  |  +-------- Iridininae    
  |  |                       
  +--+  +----- Anodontitinae 
     |  |                    
     +--+----- Mycetopodinae 
        |                    
        |  +-- Etheria       
        +--+                 
           +-- Mulleria

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

cstiTtTbiriSoTroTroSiD
0.900 3 5 3 3 31 2 1 8.0 
                    +-- Castaliini **      
                 +--+                   
                 |  +-- Rhipidodontini **  
  +--------------+                      
  |              |  +-- Echyridella **     
  |              +--+                   
  |                 +-- Velesunioninae  
  |                                     
--+        +----------- Iridininae      
  |  +-----+                            
  |  |     |  +-------- Etheria         
  |  |     +--+                         
  |  |        |  +----- Mulleria        
  |  |        +--+                      
  +--+           |  +-- Monocondylaeinae
     |           +--+                   
     |              +-- Anodontitinae   
     |                                  
     |  +-------------- Parreysiinae    
     |  |                               
     +--+  +----------- Pseudomulleria  
        |  |                            
        +--+  +-------- Anodontini      
           |  |                         
           +--+  +----- Gonideinae      
              |  |                      
              +--+----- Unionini        
                 |                      
                 |  +-- Margaritiferinae
                 +--+                   
                    +-- Ambleminae

Hoeh et al., 2001 [Publ 2652], fig. 14f2

cstiTtTbiriSoTroTroSiD
0.800 3 5 1 3 28 2 1 9.0 
     +----- Iridininae             
     |                             
  +--+  +-- Anodontites_guanarensis
  |  |  |                          
  |  +--+-- Anodontites_trigona    
  |     |                          
--+     +-- Monocondylaeinae       
  |                                
  |  +----- Rhipidodontini **         
  |  |                             
  |  +----- Echyridella **            
  +--+                             
     +----- Lortiella              
     |                             
     |  +-- Castaliini **             
     +--+                          
        +-- Velesunio

Hoeh et al., 2001 [Publ 2652], fig. 14f4

cstiTtTbiriSoTroTroSiD
0.900 3 5 3 3 28 2 1 8.0 
                 +-- Castaliini **      
              +--+                   
              |  +-- Rhipidodontini **  
  +-----------+                      
  |           |  +-- Echyridella **     
  |           +--+                   
  |              +-- Velesunioninae  
--+                                  
  |           +----- Iridininae      
  |  +--------+                      
  |  |        |  +-- Monocondylaeinae
  |  |        +--+                   
  +--+           +-- Anodontitinae   
     |                               
     |  +----------- Parreysiinae    
     +--+                            
        |  +-------- Margaritiferinae
        +--+                         
           |  +----- Gonideinae      
           +--+                      
              |  +-- Unioninae       
              +--+                   
                 +-- Ambleminae

Graf, 2002 [Publ 2651], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 15 1 1 5.5 
  +-------- Velesunioninae
--+                       
  |  +----- Hyridellini **   
  +--+                    
     |  +-- Rhipidodontini **
     +--+                 
        +-- Castaliini **

Hoeh et al., 2002 [Publ 2653], fig. 2

cstiTtTbiriSoTroTroSiD
0.900 3 5 3 3 28 2 1 8.0 
                 +-- Castaliini **      
              +--+                   
              |  +-- Rhipidodontini **  
  +-----------+                      
  |           |  +-- Echyridella **     
  |           +--+                   
  |              +-- Velesunioninae  
--+                                  
  |           +----- Iridininae      
  |  +--------+                      
  |  |        |  +-- Monocondylaeinae
  |  |        +--+                   
  +--+           +-- Anodontitinae   
     |                               
     |  +----------- Parreysiinae    
     +--+                            
        |  +-------- Margaritiferinae
        +--+                         
           |  +----- Ambleminae      
           +--+                      
              |  +-- Gonideinae      
              +--+                   
                 +-- Unioninae

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

cstiTtTbiriSoTroTroSiD
0.800 3 5 2 3 28 2 1 10.0 
     +----- Iridininae      
  +--+                      
  |  |  +-- Monocondylaeinae
  |  +--+                   
  |     +-- Anodontitinae   
--+                         
  |     +-- Castaliini **      
  |  +--+                   
  |  |  +-- Velesunio       
  +--+                      
     |  +-- Echyridella **     
     |  |                   
     +--+-- Lortiella       
        |                   
        +-- Rhipidodontini **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

cstiTtTbiriSoTroTroSiD
0.900 3 5 3 3 28 2 1 8.0 
     +----- Iridininae      
  +--+                      
  |  |  +-- Monocondylaeinae
  |  +--+                   
  |     +-- Anodontitinae   
--+                         
  |     +-- Castaliini **      
  |  +--+                   
  |  |  +-- Rhipidodontini **  
  +--+                      
     |  +-- Echyridella **     
     +--+                   
        +-- Velesunioninae

Whelan et al., 2011 [Publ 3380], fig. 1-ML

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 33 1 1 4.0 
  +-------- Velesunioninae
--+                       
  |  +----- Hyridellini **   
  +--+                    
     |  +-- Castaliini **    
     +--+                 
        +-- Hyriini **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 33 1 1 4.0 
  +-------- Velesunioninae
--+                       
  |  +----- Hyridellini **   
  +--+                    
     |  +-- Castaliini **    
     +--+                 
        +-- Hyriini **

Marshall et al., 2014 [Publ 3593], fig. 1

cstiTtTbiriSoTroTroSiD
0.818 7 10 8 3 7 3 1 12.0 
  +----------- Ambleminae    
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Velesunioninae
     +--+                    
        |  +-- Hyridellini **   
        +--+                 
           +-- Rhipidodontini **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 37 1 1 4.0 
  +-------- Velesunioninae
--+                       
  |  +----- Hyridellini **   
  +--+                    
     |  +-- Hyriini **       
     +--+                 
        +-- Castaliini **

Bolotov et al., 2016 [Publ 3743], fig. 2

cstiTtTbiriSoTroTroSiD
0.923 3 3 1 3 27 6 1 4.0 
  +-------- Velesunioninae
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Hyriini **       
     +--+                 
        +-- Castaliini **

Bolotov et al., 2016 [Publ 3743], fig. 4

cstiTtTbiriSoTroTroSiD
0.923 3 3 1 3 27 6 1 4.0 
  +-------- Velesunioninae
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Hyriini **       
     +--+                 
        +-- Castaliini **

Bolotov et al., 2017 [Publ 3761], fig. S1

cstiTtTbiriSoTroTroSiD
0.923 3 3 1 3 136 6 1 4.0 
  +-------- Velesunioninae
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Castaliini **    
     +--+                 
        +-- Hyriini **

Bolotov et al., 2017 [Publ 3761], fig. S2

cstiTtTbiriSoTroTroSiD
0.923 3 3 1 3 136 6 1 4.0 
  +-------- Velesunioninae
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Castaliini **    
     +--+                 
        +-- Hyriini **

Santos-Neto et al., 2016 [Publ 3739], fig. 3_5

cstiTtTbiriSoTroTroSiD
0.909 7 7 6 3 0 0 0 9.0 
  +----- Rhipidodontini **
--+                    
  |  +-- Hyriini **       
  +--+                 
     +-- Castaliini **

Konopleva et al., 2017 [Publ 3807], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 28 0 0 4.0 
        +----- Rhipidodontini **  
  +-----+                      
  |     |  +-- Hyriini **         
  |     +--+                   
  |        +-- Castaliini **      
--+                            
  |     +----- Monocondylaeinae
  |  +--+                      
  |  |  |  +-- Etheria         
  |  |  +--+                   
  +--+     +-- Aspathariinae   
     |                         
     |  +----- Margaritiferinae
     +--+                      
        |  +-- Gonideinae      
        +--+                   
           +-- Parreysiinae

Combosch et al., 2017 [Publ 3806], fig. 1

cstiTtTbiriSoTroTroSiD
0.875 2 3 0 2 13 1 1 6.0 
                 +----- Hyriini **         
  +--------------+                      
  |              |  +-- Velesunioninae  
  |              +--+                   
  |                 +-- Hyridellini **     
--+                                     
  |              +----- Unioninae       
  |  +-----------+                      
  |  |           |  +-- Ambleminae      
  |  |           +--+                   
  +--+              +-- Parreysiinae    
     |                                  
     |  +-------------- Margaritiferinae
     +--+                               
        |  +----------- Aspathariinae   
        +--+                            
           |  +-------- Etheria         
           +--+                         
              |  +----- Iridininae      
              +--+                      
                 |  +-- Monocondylaeinae
                 +--+                   
                    +-- Anodontitinae

Combosch et al., 2017 [Publ 3806], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 13 1 1 -- 
  +----- Velesunioninae
--+                    
  |  +-- Hyriini **       
  +--+                 
     +-- Hyridellini **

Huang et al., 2018 [Publ 3828], fig. 3_5

cstiTtTbiriSoTroTroSiD
0.875 2 3 0 2 27 1 1 6.0 
              +----- Hyriini **         
  +-----------+                      
  |           |  +-- Velesunioninae  
  |           +--+                   
  |              +-- Hyridellini **     
  |                                  
  |     +----------- Iridininae      
--+  +--+                            
  |  |  |  +-------- Aspathariinae   
  |  |  +--+                         
  |  |     |  +----- Etheria         
  |  |     +--+                      
  |  |        |  +-- Monocondylaeinae
  +--+        +--+                   
     |           +-- Anodontitinae   
     |                               
     |           +-- Gibbosulinae    
     |  +--------+                   
     |  |        +-- Margaritiferinae
     +--+                            
        |  +-------- Gonideinae      
        +--+                         
           |  +----- Unioninae       
           +--+                      
              |  +-- Ambleminae      
              +--+                   
                 +-- Parreysiinae

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

cstiTtTbiriSoTroTroSiD
0.875 2 3 0 2 26 1 1 6.0 
              +----- Hyriini **         
  +-----------+                      
  |           |  +-- Velesunioninae  
  |           +--+                   
  |              +-- Hyridellini **     
  |                                  
  |     +----------- Iridininae      
--+  +--+                            
  |  |  |  +-------- Aspathariinae   
  |  |  +--+                         
  |  |     |  +----- Etheria         
  |  |     +--+                      
  |  |        |  +-- Anodontitinae   
  +--+        +--+                   
     |           +-- Monocondylaeinae
     |                               
     |           +-- Gibbosulinae    
     |     +-----+                   
     |     |     +-- Margaritiferinae
     +-----+                         
           |  +----- Gonideinae      
           +--+                      
              |  +-- Ambleminae      
              +--+                   
                 +-- Unioninae

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

cstiTtTbiriSoTroTroSiD
0.875 2 3 0 2 26 1 1 6.0 
              +----- Hyriini **         
  +-----------+                      
  |           |  +-- Velesunioninae  
  |           +--+                   
  |              +-- Hyridellini **     
  |                                  
  |     +----------- Iridininae      
--+  +--+                            
  |  |  |  +-------- Aspathariinae   
  |  |  +--+                         
  |  |     |  +----- Etheria         
  |  |     +--+                      
  |  |        |  +-- Monocondylaeinae
  +--+        +--+                   
     |           +-- Anodontitinae   
     |                               
     |           +-- Gibbosulinae    
     |     +-----+                   
     |     |     +-- Margaritiferinae
     +-----+                         
           |  +----- Gonideinae      
           +--+                      
              |  +-- Ambleminae      
              +--+                   
                 +-- Unioninae

Wu et al., 2020 [Publ 4121], fig. 2

cstiTtTbiriSoTroTroSiD
0.875 2 3 0 2 32 1 1 6.0 
              +----- Hyriini **         
  +-----------+                      
  |           |  +-- Hyridellini **     
  |           +--+                   
  |              +-- Velesunioninae  
  |                                  
  |     +----------- Iridininae      
--+  +--+                            
  |  |  |  +-------- Aspathariinae   
  |  |  +--+                         
  |  |     |  +----- Etheria         
  |  |     +--+                      
  |  |        |  +-- Monocondylaeinae
  |  |        +--+                   
  +--+           +-- Anodontitinae   
     |                               
     |     +-------- Cumberlandia    
     |  +--+                         
     |  |  |  +----- Margaritifera   
     |  |  +--+                      
     |  |     |  +-- Gibbosulinae    
     |  |     +--+                   
     +--+        +-- Pseudunio       
        |                            
        |        +-- Ambleminae      
        |     +--+                   
        |     |  +-- Parreysiinae    
        +-----+                      
              |  +-- Gonideinae      
              +--+                   
                 +-- Unioninae

Wu et al., 2020 [Publ 4121], fig. S1

cstiTtTbiriSoTroTroSiD
0.875 2 3 0 2 32 1 1 6.0 
              +----- Hyriini **                 
  +-----------+                              
  |           |  +-- Hyridellini **             
  |           +--+                           
  |              +-- Velesunioninae          
  |                                          
  |     +----------- Iridininae              
--+  +--+                                    
  |  |  |  +-------- Aspathariinae           
  |  |  +--+                                 
  |  |     |  +----- Etheria                 
  |  |     +--+                              
  |  |        |  +-- Monocondylaeinae        
  |  |        +--+                           
  +--+           +-- Anodontitinae           
     |                                       
     |        +----- Gibbosula_rochechouartii
     |     +--+                              
     |     |  |  +-- Gibbosula_laosensis     
     |     |  +--+                           
     |     |     +-- Margaritiferinae        
     +-----+                                 
           |     +-- Ambleminae              
           |  +--+                           
           |  |  +-- Parreysiinae            
           +--+                              
              |  +-- Gonideinae              
              +--+                           
                 +-- Unioninae

Sano et al., 2020 [Publ 4138], fig. 2

cstiTtTbiriSoTroTroSiD
0.875 2 3 0 2 67 1 1 6.0 
              +----- Hyriini **         
  +-----------+                      
  |           |  +-- Hyridellini **     
  |           +--+                   
  |              +-- Velesunioninae  
  |                                  
  |     +----------- Iridininae      
--+  +--+                            
  |  |  |  +-------- Aspathariinae   
  |  |  +--+                         
  |  |     |  +----- Etheria         
  |  |     +--+                      
  |  |        |  +-- Monocondylaeinae
  |  |        +--+                   
  +--+           +-- Anodontitinae   
     |                               
     |     +-------- Cumberlandia    
     |  +--+                         
     |  |  |  +----- Margaritifera   
     |  |  +--+                      
     |  |     |  +-- Gibbosulinae    
     +--+     +--+                   
        |        +-- Pseudunio       
        |                            
        |  +-------- Parreysiinae    
        +--+                         
           |  +----- Unioninae       
           +--+                      
              |  +-- Ambleminae      
              +--+                   
                 +-- Gonideinae

Sano et al., 2020 [Publ 4138], fig. 3

cstiTtTbiriSoTroTroSiD
0.875 2 3 0 2 67 1 1 6.0 
                    +----- Hyriini **         
  +-----------------+                      
  |                 |  +-- Hyridellini **     
  |                 +--+                   
  |                    +-- Velesunioninae  
  |                                        
  |           +----------- Iridininae      
--+  +--------+                            
  |  |        |  +-------- Aspathariinae   
  |  |        +--+                         
  |  |           |  +----- Etheria         
  |  |           +--+                      
  |  |              |  +-- Monocondylaeinae
  |  |              +--+                   
  +--+                 +-- Anodontitinae   
     |                                     
     |           +-------- Cumberlandia    
     |  +--------+                         
     |  |        |  +----- Pseudunio       
     |  |        +--+                      
     |  |           |  +-- Gibbosulinae    
     +--+           +--+                   
        |              +-- Margaritifera   
        |                                  
        |  +-------------- Parreysiinae    
        +--+                               
           |  +----------- Unioninae       
           +--+                            
              |  +-------- Chamberlainiini 
              +--+                         
                 |  +----- Ambleminae      
                 +--+                      
                    |  +-- Lamprotulini    
                    +--+                   
                       +-- Gonideini

Sano et al., 2020 [Publ 4138], fig. 4

cstiTtTbiriSoTroTroSiD
0.875 2 3 0 2 67 1 1 6.0 
              +----- Hyriini **         
  +-----------+                      
  |           |  +-- Hyridellini **     
  |           +--+                   
  |              +-- Velesunioninae  
  |                                  
  |     +----------- Iridininae      
--+  +--+                            
  |  |  |  +-------- Aspathariinae   
  |  |  +--+                         
  |  |     |  +----- Etheria         
  |  |     +--+                      
  |  |        |  +-- Monocondylaeinae
  |  |        +--+                   
  +--+           +-- Anodontitinae   
     |                               
     |     +-------- Cumberlandia    
     |  +--+                         
     |  |  |  +----- Pseudunio       
     |  |  +--+                      
     |  |     |  +-- Gibbosulinae    
     +--+     +--+                   
        |        +-- Margaritifera   
        |                            
        |  +-------- Parreysiinae    
        +--+                         
           |  +----- Unioninae       
           +--+                      
              |  +-- Ambleminae      
              +--+                   
                 +-- Gonideinae

Sonowal et al., 2021 [Publ 4221], fig. 3a

cstiTtTbiriSoTroTroSiD
0.875 2 3 0 2 13 1 1 6.0 
  +----- Neotrigonia   
  |                    
--+  +-- Hyriini **       
  |  |                 
  +--+-- Hyridellini **   
     |                 
     +-- Velesunioninae

Sonowal et al., 2021 [Publ 4221], fig. 3b

cstiTtTbiriSoTroTroSiD
0.875 2 3 0 2 13 1 1 6.0 
     +-- Hyriini **        
     |                  
  +--+-- Velesunioninae 
  |  |                  
  |  +-- Hyridellini **    
  |                     
  |  +-- Aculamprotulina
  |  |                  
--+  +-- Lamprotula     
  |  |                  
  |  +-- Lampsilini     
  |  |                  
  |  +-- Potomida       
  |  |                  
  +--+-- Unio_pictorum  
     |                  
     +-- Anodontini     
     |                  
     +-- Unio_ravoisieri
     |                  
     +-- Parreysiinae   
     |                  
     +-- Pleurobemini

Sonowal et al., 2021 [Publ 4221], fig. 4

cstiTtTbiriSoTroTroSiD
0.875 2 3 0 2 14 1 1 6.0 
                          +----- Hyriini **                
  +-----------------------+                             
  |                       |  +-- Velesunioninae         
  |                       +--+                          
--+                          +-- Hyridellini **            
  |                                                     
  |  +-------------------------- Lamellidens_mainwaringi
  +--+                                                  
     |  +----------------------- Lamprotula             
     +--+                                               
        |  +-------------------- Potomida               
        +--+                                            
           |  +----------------- Lampsilini             
           +--+                                         
              |  +-------------- Pleurobemini           
              +--+                                      
                 |  +----------- Aculamprotulina        
                 +--+                                   
                    |  +-------- Unio_pictorum          
                    +--+                                
                       |  +----- Anodontini             
                       +--+                             
                          |  +-- Unio_ravoisieri        
                          +--+                          
                             +-- Lamellidens_corrianus

Wu et al., 2021 [Publ 4366], fig. 2_I

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 89 1 1 -- 
  +----- Velesunioninae
--+                    
  |  +-- Castaliini **    
  +--+                 
     +-- Rhipidodontini **

Wu et al., 2021 [Publ 4366], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 89 1 1 -- 
  +----- Velesunioninae
--+                    
  |  +-- Rhipidodontini **
  +--+                 
     +-- Castaliini **

Santos-Neto et al., 2018 [Publ 3888], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 5 5 3 2 0 0 0 8.0 
  +-- Hyriini **   
--+             
  +-- Castaliini **

Chen et al., 2025 [Publ 4455], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 24 0 0 -- 
                    +-- Hyridellini **              
  +-----------------+                            
  |                 +-- Hyriini **                  
  |                                              
  |           +-------- Aspathariinae            
--+  +--------+                                  
  |  |        |  +----- Etheria                  
  |  |        +--+                               
  |  |           |  +-- Monocondylaeinae         
  |  |           +--+                            
  +--+              +-- Anodontitinae            
     |                                           
     |              +-- Ambleminae               
     |  +-----------+                            
     |  |           +-- Gonideinae               
     |  |                                        
     +--+  +----------- Pseudunio_marocanus      
        |  |                                     
        |  |        +-- Pseudunio_auricularius__1
        +--+        |                            
           |  +-----+-- Pseudunio_auricularius__2
           |  |     |                            
           +--+     +-- Pseudunio_homsensis      
              |                                  
              |  +----- Gibbosulinae             
              +--+                               
                 |  +-- Cumberlandia             
                 +--+                            
                    +-- Margaritifera

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 80 1 1 -- 
  +-- Hyridellini **   
--+                 
  +-- Velesunioninae

Guerra et al., 2019 [Publ 4168], fig. 1-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 21 1 1 -- 
  +-- Echyridella **   
--+                 
  +-- Velesunioninae

Guerra et al., 2019 [Publ 4168], fig. 2-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 21 1 1 -- 
  +-- Echyridella **   
--+                 
  +-- Velesunioninae

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 121 1 1 -- 
  +-- Hyridellini **   
--+                 
  +-- Velesunioninae

Klunzinger et al., 2021 [Publ 4086], fig. 2

cstiTtTbiriSoTroTroSiD
0.857 1 1 0 1 5 3 1 -- 
  +----- Westralunio
--+                 
  |  +-- Velesunio  
  +--+              
     +-- Hyridellini **

Klunzinger et al., 2022 [Publ 4259], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 3 3 1 -- 
  +-- Hyridellini **   
--+                 
  +-- Velesunioninae

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 46 1 1 -- 
  +-- Echyridella **   
--+                 
  +-- Velesunioninae

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 29 0 0 -- 
  +----------- Hyridellini **     
--+                            
  |  +-------- Margaritiferinae
  +--+                         
     |  +----- Unionini        
     +--+                      
        |  +-- Anodontini      
        +--+                   
           +-- Ambleminae

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 29 0 0 -- 
  +----------- Hyridellini **     
--+                            
  |  +-------- Margaritiferinae
  +--+                         
     |  +----- Unionini        
     +--+                      
        |  +-- Anodontini      
        +--+                   
           +-- Ambleminae

Hoeh et al., 2002 [Publ 2673], fig. 1-F

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 8 0 0 -- 
  +----- Echyridella **
--+                 
  |  +-- Unioninae  
  +--+              
     +-- Ambleminae

Hoeh et al., 2002 [Publ 2673], fig. 1-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 8 0 0 -- 
  +----- Echyridella **
--+                 
  |  +-- Unioninae  
  +--+              
     +-- Ambleminae

Walker et al., 2006 [Publ 2666], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 14 0 0 -- 
  +----------------- Hyridellini **     
  |                                  
--+  +-------------- Unioninae       
  |  |                               
  +--+  +----------- Lamprotulini    
     |  |                            
     +--+        +-- Gonideini       
        |  +-----+                   
        |  |     +-- Pseudodontini   
        +--+                         
           |  +----- Lampsilini      
           +--+                      
              |  +-- Pleurobemini    
              +--+                   
                 +-- Margaritiferinae

Walker et al., 2006 [Publ 2666], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 14 0 0 -- 
  +----------------- Hyridellini **     
--+                                  
  |  +-------------- Margaritiferinae
  +--+                               
     |  +----------- Pseudodontini   
     +--+                            
        |  +-------- Gonidea         
        +--+                         
           |  +----- Inversidens     
           +--+                      
              |  +-- Unioninae       
              +--+                   
                 +-- Ambleminae

Walker et al., 2006 [Publ 2666], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 14 0 0 -- 
  +----------------- Hyridellini **     
--+                                  
  |  +-------------- Margaritiferinae
  +--+                               
     |  +----------- Pseudodontini   
     +--+                            
        |  +-------- Gonidea         
        +--+                         
           |  +----- Inversidens     
           +--+                      
              |  +-- Unioninae       
              +--+                   
                 +-- Ambleminae

Walker et al., 2006 [Publ 2666], fig. 6-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 13 0 0 -- 
  +----------- Echyridella **     
--+                            
  |  +-------- Margaritiferinae
  +--+                         
     |  +----- Gonideinae      
     +--+                      
        |  +-- Unioninae       
        +--+                   
           +-- Ambleminae

Breton et al., 2011 [Publ 3376], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 37 0 0 -- 
  +-------------------- Echyridella **     
  |                                     
--+  +----------------- Margaritiferinae
  |  |                                  
  +--+  +-------------- Quadrulina      
     |  |                               
     |  |        +----- Lamprotulini    
     +--+  +-----+                      
        |  |     |  +-- Chamberlainiini 
        |  |     +--+                   
        +--+        +-- Unioninae       
           |                            
           |  +-------- Megalonaias     
           +--+                         
              |  +----- Pleurobemini    
              +--+                      
                 |  +-- Amblemini       
                 +--+                   
                    +-- Lampsilini

Guerra et al., 2017 [Publ 3772], fig. 2a

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 24 0 0 -- 
  +----------- Echyridella **     
--+                            
  |  +-------- Margaritiferinae
  +--+                         
     |  +----- Unioninae       
     +--+                      
        |  +-- Ambleminae      
        +--+                   
           +-- Gonideinae

Guerra et al., 2017 [Publ 3772], fig. 2a-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 7 0 0 -- 
  +-------- Echyridella **     
  |                         
--+  +----- Margaritiferinae
  |  |                      
  +--+  +-- Gonideinae      
     |  |                   
     +--+-- Unioninae       
        |                   
        +-- Ambleminae

Guerra et al., 2017 [Publ 3772], fig. 2b

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 24 0 0 -- 
  +----------- Echyridella **     
--+                            
  |  +-------- Margaritiferinae
  +--+                         
     |  +----- Ambleminae      
     +--+                      
        |  +-- Gonideinae      
        +--+                   
           +-- Unioninae

Guerra et al., 2017 [Publ 3772], fig. 2b-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 7 0 0 -- 
  +-------- Echyridella **     
  |                         
--+  +----- Margaritiferinae
  |  |                      
  +--+  +-- Gonideinae      
     |  |                   
     +--+-- Unioninae       
        |                   
        +-- Ambleminae

Huang et al., 2018 [Publ 3828], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 41 0 0 -- 
  +----------- Echyridella **     
  |                            
--+        +-- Gibbosulinae    
  |  +-----+                   
  |  |     +-- Margaritiferinae
  +--+                         
     |  +----- Unioninae       
     +--+                      
        |  +-- Ambleminae      
        +--+                   
           +-- Gonideinae

Huang et al., 2018 [Publ 3828], fig. 6-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 16 0 0 -- 
  +----------- Echyridella **     
--+                            
  |  +-------- Margaritiferinae
  +--+                         
     |  +----- Ambleminae      
     +--+                      
        |  +-- Gonideinae      
        +--+                   
           +-- Unioninae

Chase et al., 2018 [Publ 3847], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 38 0 0 -- 
  +-- Echyridella **
--+              
  +-- Ambleminae

Chase et al., 2018 [Publ 3847], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 9 0 0 -- 
  +-- Margaritiferinae
--+                   
  +-- Echyridella **

Chase et al., 2018 [Publ 3861], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 10 0 0 -- 
  +----------- Echyridella **     
--+                            
  |  +-------- Margaritiferinae
  +--+                         
     |  +----- Ambleminae      
     +--+                      
        |  +-- Gonideinae      
        +--+                   
           +-- Unioninae

Chase et al., 2018 [Publ 3861], fig. 1-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 10 0 0 -- 
  +-- Margaritiferinae
--+                   
  +-- Echyridella **

Huang et al., 2019 [Publ 3843], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 55 0 0 -- 
  +----------- Echyridella **     
  |                            
--+        +-- Gibbosulinae    
  |  +-----+                   
  |  |     +-- Margaritiferinae
  +--+                         
     |  +----- Unioninae       
     +--+                      
        |  +-- Gonideinae      
        +--+                   
           +-- Ambleminae

Huang et al., 2019 [Publ 3843], fig. 3-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 22 0 0 -- 
  +----------- Echyridella **     
--+                            
  |  +-------- Margaritiferinae
  +--+                         
     |  +----- Ambleminae      
     +--+                      
        |  +-- Gonideinae      
        +--+                   
           +-- Unioninae

Wu et al., 2019 [Publ 3864], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 45 0 0 -- 
  +-- Echyridella **
--+              
  +-- Iridininae

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 56 0 0 -- 
  +-------------- Hyridellini **     
--+                               
  |  +----------- Margaritiferinae
  +--+                            
     |  +-------- Ambleminae      
     +--+                         
        |  +----- Gonideinae      
        +--+                      
           |  +-- Unioninae       
           +--+                   
              +-- Parreysiinae

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 56 0 0 -- 
  +-- Hyridellini **     
--+                   
  +-- Margaritiferinae

Wu et al., 2020 [Publ 4121], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 47 0 0 -- 
  +-------------- Echyridella **     
  |                               
  |           +-- Anodontitinae   
--+  +--------+                   
  |  |        +-- Iridininae      
  |  |                            
  +--+        +-- Gibbosulinae    
     |  +-----+                   
     |  |     +-- Margaritiferinae
     +--+                         
        |  +----- Ambleminae      
        +--+                      
           |  +-- Gonideinae      
           +--+                   
              +-- Unioninae

Olivera-Hyde et al., 2020 [Publ 4249], fig. 10

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 1 0 0 0 8.0 
-- Castaliini **

Wu et al., 2021 [Publ 4366], fig. 3-BI

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 72 0 0 -- 
  +-------------- Echyridella **     
  |                               
  |           +-- Anodontitinae   
--+  +--------+                   
  |  |        +-- Iridininae      
  |  |                            
  +--+        +-- Gibbosulinae    
     |  +-----+                   
     |  |     +-- Margaritiferinae
     +--+                         
        |  +----- Ambleminae      
        +--+                      
           |  +-- Gonideinae      
           +--+                   
              +-- Unioninae

Wu et al., 2021 [Publ 4366], fig. 3-ML

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 72 0 0 -- 
  +-------------- Echyridella **     
  |                               
  |           +-- Anodontitinae   
--+  +--------+                   
  |  |        +-- Iridininae      
  |  |                            
  +--+        +-- Gibbosulinae    
     |  +-----+                   
     |  |     +-- Margaritiferinae
     +--+                         
        |  +----- Ambleminae      
        +--+                      
           |  +-- Gonideinae      
           +--+                   
              +-- Unioninae

Wu et al., 2021 [Publ 4366], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 67 0 0 -- 
  +-------------- Echyridella **     
  |                               
  |           +-- Iridininae      
--+  +--------+                   
  |  |        +-- Anodontitinae   
  |  |                            
  +--+        +-- Gibbosulinae    
     |  +-----+                   
     |  |     +-- Margaritiferinae
     +--+                         
        |  +----- Ambleminae      
        +--+                      
           |  +-- Gonideinae      
           +--+                   
              +-- Unioninae

Wu et al., 2022 [Publ 4238], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 72 0 0 -- 
  +----------- Echyridella **     
  |                            
--+        +-- Gibbosulinae    
  |  +-----+                   
  |  |     +-- Margaritiferinae
  +--+                         
     |  +----- Ambleminae      
     +--+                      
        |  +-- Gonideinae      
        +--+                   
           +-- Unioninae

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 94 0 0 -- 
  +----------------- Echyridella **     
  |                                  
  |              +-- Anodontitinae   
--+  +-----------+                   
  |  |           +-- Iridininae      
  |  |                               
  +--+           +-- Gibbosulinae    
     |  +--------+                   
     |  |        +-- Margaritiferinae
     +--+                            
        |  +-------- Ambleminae      
        +--+                         
           |  +----- Gonideinae      
           +--+                      
              |  +-- Parreysiinae    
              +--+                   
                 +-- Unioninae

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 94 0 0 -- 
  +----------------- Echyridella **     
  |                                  
  |              +-- Anodontitinae   
--+  +-----------+                   
  |  |           +-- Iridininae      
  |  |                               
  +--+           +-- Gibbosulinae    
     |  +--------+                   
     |  |        +-- Margaritiferinae
     +--+                            
        |  +-------- Ambleminae      
        +--+                         
           |  +----- Gonideinae      
           +--+                      
              |  +-- Parreysiinae    
              +--+                   
                 +-- Unioninae

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 90 0 0 -- 
  +----------------- Echyridella **     
  |                                  
--+              +-- Anodontitinae   
  |  +-----------+                   
  |  |           +-- Iridininae      
  +--+                               
     |  +----------- Margaritiferinae
     +--+                            
        |  +-------- Ambleminae      
        +--+                         
           |  +----- Gonideinae      
           +--+                      
              |  +-- Parreysiinae    
              +--+                   
                 +-- Unioninae

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 90 0 0 -- 
  +----------------- Echyridella **     
  |                                  
--+              +-- Anodontitinae   
  |  +-----------+                   
  |  |           +-- Iridininae      
  +--+                               
     |  +----------- Margaritiferinae
     +--+                            
        |  +-------- Ambleminae      
        +--+                         
           |  +----- Gonideinae      
           +--+                      
              |  +-- Parreysiinae    
              +--+                   
                 +-- Unioninae

Dai et al., 2025 [Publ 4440], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 86 0 0 -- 
  +----------- Echyridella **     
  |                            
--+        +-- Gibbosulinae    
  |  +-----+                   
  |  |     +-- Margaritiferinae
  +--+                         
     |  +----- Ambleminae      
     +--+                      
        |  +-- Gonideinae      
        +--+                   
           +-- Unioninae

Chen et al., 2025 [Publ 4448], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 84 0 0 -- 
  +----- Echyridella **     
--+                      
  |  +-- Gibbosulinae    
  +--+                   
     +-- Margaritiferinae