Bivalvia | Unionida | Hyrioidea | Hyriidae

subfamily HYRIINAE Swainson, 1840: cladomics

4 tribes, 1 genus

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 78 (88 total)
number of subtaxa (rank: tribe) = 5
analyzed in 101 cladograms in 53 publications
maximum number of terminals in an analysis = 19 (mean = 2.96, median = 2)
maximum number of subtaxa (tribe) = 5 (mean = 2.18, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 56
maximum number of included terminals = 19, (mean = 4.54 median = 3)
maximum number of subtaxa (tribe) = 5 (mean = 3.13, median = 3)
proportion of cladograms that support basic monophyly = 0.46

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 51
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (tribe) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 19 (mean = 4.59, median = 3)
maximum number of subtaxa (tribe) = 5 (mean = 3.22, median = 3)
proportion of cladograms that support rigorous monophyly = 0.41

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Hyriinae cladogram distances
PCoA Ordination

NJ Tree of Hyriinae cladogram distances
NJ Tree

Cladograms

Huang et al., 2019 [Publ 3843], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
0.882 7 7 5 5 194 7 1 **
Cladomics cladogram Westralunio Echyridella HYRIDELLINI RHIPIDODONTINI CASTALIINI HYRIINI Alathyria jacksoni Alathyria profuga Velesunio Alathyria pertexta Lortiella Microdontia

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

cst iT tT bi riS oT roT roS iD
0.682 14 20 20 5 7 6 1 19.0
Cladomics cladogram Neotrigonia Guyanhyrida HYRIINI CASTALIINI Westralunio Diplodon Hyridella drapeta Hyridella depressa Hyridella australis Cucumerunio Echyridella Velesunio ambiguus Microdontia Lortiella Alathyria Velesunio angasi

Santos-Neto et al., 2016 [Publ 3739], fig. 4

cst iT tT bi riS oT roT roS iD
0.682 14 20 21 5 7 6 1 20.0
Cladomics cladogram Neotrigonia Guyanhyrida HYRIINI CASTALIINI Westralunio Diplodon Hyridella drapeta Hyridella depressa Echyridella Cucumerunio Hyridella australis Velesunio ambiguus Alathyria Velesunio angasi Lortiella Microdontia

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
0.882 7 7 5 5 192 7 1 0
Cladomics cladogram Westralunio Echyridella HYRIDELLINI RHIPIDODONTINI CASTALIINI HYRIINI Velesunio Alathyria jacksoni Alathyria profuga Alathyria pertexta Lortiella Microdontia

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.824 7 14 9 5 194 7 1 8.0
Cladomics cladogram Echyridella Westralunio HYRIDELLINI RHIPIDODONTINI CASTALIINI HYRIINI Velesunio Alathyria jacksoni Alathyria profuga Alathyria pertexta Lortiella Microdontia Gibbosula rochechouartii Gibbosula laosensis MARGARITIFERINAE PARREYSIINAE Chamberlainia Megalonaias Uniomerus QUADRULINA Sinohyriopsis LAMPROTULINI GONIDEINI PSEUDODONTINI Rectidentini UNIONINAE Amblema Toxolasma PLEUROBEMINI Obliquaria Plectomerus Ellipsaria Potamilus leptodon Medionidus Potamilus alatus Ptychobranchus Truncilla Venustaconcha Cambarunio Leaunio Ortmanniana Lampsilis

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.636 19 27 32 5 614 10 1 31.0
Cladomics cladogram Westralunio Guyanhyrida HYRIINI CASTALIINI Diplodon Hyridella depressa Hyridella drapeta Cucumerunio Hyridella australis Echyridella Velesunio 1 Alathyria jacksoni 1 Velesunio ambiguus Alathyria profuga Velesunio 3 Microdontia Lortiella Velesunio angasi Alathyria pertexta 2 Velesunio 2 Alathyria jacksoni 2 Alathyria pertexta 1

Prié et al., 2026 [Publ 4844], fig. 2

cst iT tT bi riS oT roT roS iD
0.826 15 20 15 5 8 5 1 18.0
Cladomics cladogram MARGARITIFERINAE Echyridella Westralunio HYRIDELLINI Velesunio Alathyria Lortiella Microdontia Diplodon Guyanhyrida CASTALIINI Iridea HYRIINI

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

cst iT tT bi riS oT roT roS iD
0.818 5 5 4 4 6 1 1 8.5
Cladomics cladogram VELESUNIONINAE RHIPIDODONTINI CASTALIINI Hyridella australis 2 Hyridella australis 1 Echyridella Hyridella depressa

Baker et al., 2003 [Publ 2690], fig. 5

cst iT tT bi riS oT roT roS iD
0.727 5 7 5 4 3 2 1 12.0
Cladomics cladogram AMBLEMINAE Hyridella australis RHIPIDODONTINI Hyridella depressa VELESUNIONINAE Echyridella CASTALIINI

Baker et al., 2004 [Publ 2685], fig. 2-3

cst iT tT bi riS oT roT roS iD
0.667 5 8 9 4 4 3 1 16.0
Cladomics cladogram AMBLEMINAE Hyridella depressa Hyridella australis RHIPIDODONTINI VELESUNIONINAE Echyridella CASTALIINI

Graf & Cummings, 2006 [Publ 1968], fig. 2

cst iT tT bi riS oT roT roS iD
0.917 6 6 4 4 41 3 1 4.5
Cladomics cladogram VELESUNIONINAE Echyridella HYRIDELLINI CASTALIINI RHIPIDODONTINI

Walker et al., 2006 [Publ 2666], fig. 6

cst iT tT bi riS oT roT roS iD
0.692 5 10 7 4 31 5 1 15.0
Cladomics cladogram CASTALIINI RHIPIDODONTINI Hyridella depressa Hyridella australis Echyridella VELESUNIONINAE UNIONINAE MARGARITIFERINAE GONIDEINAE AMBLEMINAE IRIDININAE Etheria ASPATHARIINAE Mulleria ANODONTITINAE MONOCONDYLAEINAE

Hoeh et al., 2009 [Publ 2716], fig. 3

cst iT tT bi riS oT roT roS iD
0.917 6 6 4 4 41 3 1 4.5
Cladomics cladogram VELESUNIONINAE Echyridella HYRIDELLINI CASTALIINI RHIPIDODONTINI

Hoeh et al., 2009 [Publ 2716], fig. 4

cst iT tT bi riS oT roT roS iD
0.917 6 6 4 4 40 3 1 4.5
Cladomics cladogram VELESUNIONINAE Echyridella HYRIDELLINI CASTALIINI RHIPIDODONTINI

Hoeh et al., 2009 [Publ 2716], fig. 5

cst iT tT bi riS oT roT roS iD
0.917 6 6 4 4 40 3 1 4.5
Cladomics cladogram VELESUNIONINAE Echyridella HYRIDELLINI CASTALIINI RHIPIDODONTINI

Graf et al., 2015 [Publ 3619], fig. 1

cst iT tT bi riS oT roT roS iD
0.882 7 7 5 4 13 7 1 2.0
Cladomics cladogram VELESUNIONINAE HYRIDELLINI RHIPIDODONTINI HYRIINI CASTALIINI

Guerra et al., 2019 [Publ 4168], fig. 1

cst iT tT bi riS oT roT roS iD
0.909 4 5 3 4 37 1 1 6.0
Cladomics cladogram Echyridella VELESUNIONINAE RHIPIDODONTINI CASTALIINI HYRIINI GIBBOSULINAE MARGARITIFERINAE AMBLEMINAE GONIDEINAE UNIONINAE

Guerra et al., 2019 [Publ 4168], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 37 1 1 5.0
Cladomics cladogram VELESUNIONINAE Echyridella RHIPIDODONTINI CASTALIINI HYRIINI

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 136 1 1 5.0
Cladomics cladogram VELESUNIONINAE Echyridella RHIPIDODONTINI CASTALIINI HYRIINI

Graf, 2000 [Publ 2647], fig. 1

cst iT tT bi riS oT roT roS iD
0.900 3 4 2 3 14 1 1 9.0
Cladomics cladogram CASTALIINI RHIPIDODONTINI HYRIDELLINI VELESUNIONINAE IRIDININAE ANODONTITINAE MYCETOPODINAE Etheria Mulleria

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

cst iT tT bi riS oT roT roS iD
0.900 3 5 3 3 31 2 1 8.0
Cladomics cladogram CASTALIINI RHIPIDODONTINI Echyridella VELESUNIONINAE IRIDININAE Etheria Mulleria MONOCONDYLAEINAE ANODONTITINAE PARREYSIINAE Pseudomulleria ANODONTINI GONIDEINAE UNIONINI MARGARITIFERINAE AMBLEMINAE

Hoeh et al., 2001 [Publ 2652], fig. 14f2

cst iT tT bi riS oT roT roS iD
0.800 3 5 1 3 28 2 1 9.0
Cladomics cladogram IRIDININAE Anodontites guanarensis Anodontites trigona MONOCONDYLAEINAE RHIPIDODONTINI Echyridella Lortiella CASTALIINI Velesunio

Hoeh et al., 2001 [Publ 2652], fig. 14f4

cst iT tT bi riS oT roT roS iD
0.900 3 5 3 3 28 2 1 8.0
Cladomics cladogram CASTALIINI RHIPIDODONTINI Echyridella VELESUNIONINAE IRIDININAE MONOCONDYLAEINAE ANODONTITINAE PARREYSIINAE MARGARITIFERINAE GONIDEINAE UNIONINAE AMBLEMINAE

Graf, 2002 [Publ 2651], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 15 1 1 5.5
Cladomics cladogram VELESUNIONINAE HYRIDELLINI RHIPIDODONTINI CASTALIINI

Hoeh et al., 2002 [Publ 2653], fig. 2

cst iT tT bi riS oT roT roS iD
0.900 3 5 3 3 28 2 1 8.0
Cladomics cladogram CASTALIINI RHIPIDODONTINI Echyridella VELESUNIONINAE IRIDININAE MONOCONDYLAEINAE ANODONTITINAE PARREYSIINAE MARGARITIFERINAE AMBLEMINAE GONIDEINAE UNIONINAE

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

cst iT tT bi riS oT roT roS iD
0.800 3 5 2 3 28 2 1 10.0
Cladomics cladogram IRIDININAE MONOCONDYLAEINAE ANODONTITINAE CASTALIINI Velesunio Echyridella Lortiella RHIPIDODONTINI

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

cst iT tT bi riS oT roT roS iD
0.900 3 5 3 3 28 2 1 8.0
Cladomics cladogram IRIDININAE MONOCONDYLAEINAE ANODONTITINAE CASTALIINI RHIPIDODONTINI Echyridella VELESUNIONINAE

Whelan et al., 2011 [Publ 3380], fig. 1-ML

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 33 1 1 4.0
Cladomics cladogram VELESUNIONINAE HYRIDELLINI CASTALIINI HYRIINI

Whelan et al., 2011 [Publ 3380], fig. 1-MP

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 33 1 1 4.0
Cladomics cladogram VELESUNIONINAE HYRIDELLINI CASTALIINI HYRIINI

Marshall et al., 2014 [Publ 3593], fig. 1

cst iT tT bi riS oT roT roS iD
0.818 7 10 8 3 7 3 1 12.0
Cladomics cladogram AMBLEMINAE Echyridella VELESUNIONINAE HYRIDELLINI RHIPIDODONTINI

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 37 1 1 4.0
Cladomics cladogram VELESUNIONINAE HYRIDELLINI HYRIINI CASTALIINI

Bolotov et al., 2016 [Publ 3743], fig. 2

cst iT tT bi riS oT roT roS iD
0.923 3 3 1 3 27 6 1 4.0
Cladomics cladogram VELESUNIONINAE RHIPIDODONTINI HYRIINI CASTALIINI

Bolotov et al., 2016 [Publ 3743], fig. 4

cst iT tT bi riS oT roT roS iD
0.923 3 3 1 3 27 6 1 4.0
Cladomics cladogram VELESUNIONINAE RHIPIDODONTINI HYRIINI CASTALIINI

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
0.923 3 3 1 3 136 6 1 4.0
Cladomics cladogram VELESUNIONINAE RHIPIDODONTINI CASTALIINI HYRIINI

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
0.923 3 3 1 3 136 6 1 4.0
Cladomics cladogram VELESUNIONINAE RHIPIDODONTINI CASTALIINI HYRIINI

Lawrie et al., 2026 [Publ 4843], fig. 1

cst iT tT bi riS oT roT roS iD
0.733 6 17 21 3 12 11 1 21.0
Cladomics cladogram UNIONINAE HYRIDELLINI VELESUNIONINAE CASTALIINI HYRIINI

Santos-Neto et al., 2016 [Publ 3739], fig. 3_5

cst iT tT bi riS oT roT roS iD
0.909 7 7 6 3 0 0 0 10.0
Cladomics cladogram RHIPIDODONTINI HYRIINI CASTALIINI

Konopleva et al., 2017 [Publ 3807], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 28 0 0 4.0
Cladomics cladogram RHIPIDODONTINI HYRIINI CASTALIINI MONOCONDYLAEINAE Etheria ASPATHARIINAE MARGARITIFERINAE GONIDEINAE PARREYSIINAE

Combosch et al., 2017 [Publ 3806], fig. 1

cst iT tT bi riS oT roT roS iD
0.875 2 3 0 2 211 1 1 6.0
Cladomics cladogram HYRIINI VELESUNIONINAE HYRIDELLINI UNIONINAE AMBLEMINAE PARREYSIINAE MARGARITIFERINAE ASPATHARIINAE Etheria IRIDININAE MONOCONDYLAEINAE ANODONTITINAE

Combosch et al., 2017 [Publ 3806], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 212 1 1 --
Cladomics cladogram VELESUNIONINAE HYRIINI HYRIDELLINI

Huang et al., 2018 [Publ 3828], fig. 3_5

cst iT tT bi riS oT roT roS iD
0.875 2 3 0 2 27 1 1 6.0
Cladomics cladogram HYRIINI VELESUNIONINAE HYRIDELLINI IRIDININAE ASPATHARIINAE Etheria MONOCONDYLAEINAE ANODONTITINAE GIBBOSULINAE MARGARITIFERINAE GONIDEINAE UNIONINAE AMBLEMINAE PARREYSIINAE

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

cst iT tT bi riS oT roT roS iD
0.875 2 3 0 2 26 1 1 6.0
Cladomics cladogram HYRIINI VELESUNIONINAE HYRIDELLINI IRIDININAE ASPATHARIINAE Etheria ANODONTITINAE MONOCONDYLAEINAE GIBBOSULINAE MARGARITIFERINAE GONIDEINAE AMBLEMINAE UNIONINAE

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

cst iT tT bi riS oT roT roS iD
0.875 2 3 0 2 26 1 1 6.0
Cladomics cladogram HYRIINI VELESUNIONINAE HYRIDELLINI IRIDININAE ASPATHARIINAE Etheria MONOCONDYLAEINAE ANODONTITINAE GIBBOSULINAE MARGARITIFERINAE GONIDEINAE AMBLEMINAE UNIONINAE

Wu et al., 2020 [Publ 4121], fig. 2

cst iT tT bi riS oT roT roS iD
0.875 2 3 0 2 32 1 1 6.0
Cladomics cladogram HYRIINI HYRIDELLINI VELESUNIONINAE IRIDININAE ASPATHARIINAE Etheria MONOCONDYLAEINAE ANODONTITINAE Cumberlandia Margaritifera GIBBOSULINAE Pseudunio AMBLEMINAE PARREYSIINAE GONIDEINAE UNIONINAE

Wu et al., 2020 [Publ 4121], fig. S1

cst iT tT bi riS oT roT roS iD
0.875 2 3 0 2 32 1 1 6.0
Cladomics cladogram HYRIINI HYRIDELLINI VELESUNIONINAE IRIDININAE ASPATHARIINAE Etheria MONOCONDYLAEINAE ANODONTITINAE Gibbosula rochechouartii Gibbosula laosensis MARGARITIFERINAE AMBLEMINAE PARREYSIINAE GONIDEINAE UNIONINAE

Sano et al., 2020 [Publ 4138], fig. 2

cst iT tT bi riS oT roT roS iD
0.875 2 3 0 2 67 1 1 6.0
Cladomics cladogram HYRIINI HYRIDELLINI VELESUNIONINAE IRIDININAE ASPATHARIINAE Etheria MONOCONDYLAEINAE ANODONTITINAE Cumberlandia Margaritifera GIBBOSULINAE Pseudunio PARREYSIINAE UNIONINAE AMBLEMINAE GONIDEINAE

Sano et al., 2020 [Publ 4138], fig. 3

cst iT tT bi riS oT roT roS iD
0.875 2 3 0 2 67 1 1 6.0
Cladomics cladogram HYRIINI HYRIDELLINI VELESUNIONINAE IRIDININAE ASPATHARIINAE Etheria MONOCONDYLAEINAE ANODONTITINAE Cumberlandia Pseudunio GIBBOSULINAE Margaritifera PARREYSIINAE UNIONINAE CHAMBERLAINIINI AMBLEMINAE LAMPROTULINI GONIDEINI

Sano et al., 2020 [Publ 4138], fig. 4

cst iT tT bi riS oT roT roS iD
0.875 2 3 0 2 67 1 1 6.0
Cladomics cladogram HYRIINI HYRIDELLINI VELESUNIONINAE IRIDININAE ASPATHARIINAE Etheria MONOCONDYLAEINAE ANODONTITINAE Cumberlandia Pseudunio GIBBOSULINAE Margaritifera PARREYSIINAE UNIONINAE AMBLEMINAE GONIDEINAE

Sonowal et al., 2021 [Publ 4221], fig. 3a

cst iT tT bi riS oT roT roS iD
0.875 2 3 0 2 13 1 1 6.0
Cladomics cladogram Neotrigonia HYRIINI HYRIDELLINI VELESUNIONINAE

Sonowal et al., 2021 [Publ 4221], fig. 3b

cst iT tT bi riS oT roT roS iD
0.875 2 3 0 2 13 1 1 6.0
Cladomics cladogram HYRIINI VELESUNIONINAE HYRIDELLINI ACULAMPROTULINA Lamprotula LAMPSILINI Potomida Unio pictorum ANODONTINI Unio ravoisieri PARREYSIINAE PLEUROBEMINI

Sonowal et al., 2021 [Publ 4221], fig. 4

cst iT tT bi riS oT roT roS iD
0.875 2 3 0 2 14 1 1 6.0
Cladomics cladogram HYRIINI VELESUNIONINAE HYRIDELLINI Lamellidens phenchooganjensis Lamprotula Potomida LAMPSILINI PLEUROBEMINI ACULAMPROTULINA Unio pictorum ANODONTINI Unio ravoisieri Lamellidens corrianus

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 89 1 1 --
Cladomics cladogram VELESUNIONINAE CASTALIINI RHIPIDODONTINI

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 89 1 1 --
Cladomics cladogram VELESUNIONINAE RHIPIDODONTINI CASTALIINI

Santos-Neto et al., 2018 [Publ 3888], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 2 0 0 0 8.0
Cladomics cladogram HYRIINI CASTALIINI

Chen et al., 2025 [Publ 4455], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 24 0 0 --
Cladomics cladogram HYRIDELLINI HYRIINI ASPATHARIINAE Etheria MONOCONDYLAEINAE ANODONTITINAE AMBLEMINAE GONIDEINAE Pseudunio marocanus Pseudunio auricularius 1 Pseudunio auricularius 2 Pseudunio homsensis GIBBOSULINAE Cumberlandia Margaritifera

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 80 1 1 --
Cladomics cladogram HYRIDELLINI VELESUNIONINAE

Guerra et al., 2019 [Publ 4168], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 1 1 --
Cladomics cladogram Echyridella VELESUNIONINAE

Guerra et al., 2019 [Publ 4168], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 1 1 --
Cladomics cladogram Echyridella VELESUNIONINAE

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 121 1 1 --
Cladomics cladogram HYRIDELLINI VELESUNIONINAE

Klunzinger et al., 2021 [Publ 4086], fig. 2

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 5 3 1 --
Cladomics cladogram Westralunio Velesunio HYRIDELLINI

Klunzinger et al., 2022 [Publ 4259], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 3 1 --
Cladomics cladogram HYRIDELLINI VELESUNIONINAE

Gomes-dos-Santos et al., 2023 [Publ 4463], fig. 2a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 20 1 1 --
Cladomics cladogram Echyridella VELESUNIONINAE

Gomes-dos-Santos et al., 2023 [Publ 4463], fig. 2b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 1 1 --
Cladomics cladogram VELESUNIONINAE Echyridella

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 46 1 1 --
Cladomics cladogram Echyridella VELESUNIONINAE

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 0 0 --
Cladomics cladogram HYRIDELLINI MARGARITIFERINAE UNIONINI ANODONTINI AMBLEMINAE

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 0 0 --
Cladomics cladogram HYRIDELLINI MARGARITIFERINAE UNIONINI ANODONTINI AMBLEMINAE

Hoeh et al., 2002 [Publ 2673], fig. 1-F

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 0 0 --
Cladomics cladogram Echyridella UNIONINAE AMBLEMINAE

Hoeh et al., 2002 [Publ 2673], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 0 0 --
Cladomics cladogram Echyridella UNIONINAE AMBLEMINAE

Walker et al., 2006 [Publ 2666], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram HYRIDELLINI UNIONINAE LAMPROTULINI GONIDEINI PSEUDODONTINI LAMPSILINI PLEUROBEMINI MARGARITIFERINAE

Walker et al., 2006 [Publ 2666], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram HYRIDELLINI MARGARITIFERINAE PSEUDODONTINI Gonidea Inversidens UNIONINAE AMBLEMINAE

Walker et al., 2006 [Publ 2666], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram HYRIDELLINI MARGARITIFERINAE PSEUDODONTINI Gonidea Inversidens UNIONINAE AMBLEMINAE

Walker et al., 2006 [Publ 2666], fig. 6-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram Echyridella MARGARITIFERINAE GONIDEINAE UNIONINAE AMBLEMINAE

Breton et al., 2011 [Publ 3376], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 37 0 0 --
Cladomics cladogram Echyridella MARGARITIFERINAE QUADRULINA LAMPROTULINI CHAMBERLAINIINI UNIONINAE Megalonaias PLEUROBEMINI AMBLEMINI LAMPSILINI

Guerra et al., 2017 [Publ 3772], fig. 2a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 24 0 0 --
Cladomics cladogram Echyridella MARGARITIFERINAE UNIONINAE AMBLEMINAE GONIDEINAE

Guerra et al., 2017 [Publ 3772], fig. 2a-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 0 0 --
Cladomics cladogram Echyridella MARGARITIFERINAE GONIDEINAE UNIONINAE AMBLEMINAE

Guerra et al., 2017 [Publ 3772], fig. 2b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 24 0 0 --
Cladomics cladogram Echyridella MARGARITIFERINAE AMBLEMINAE GONIDEINAE UNIONINAE

Guerra et al., 2017 [Publ 3772], fig. 2b-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 0 0 --
Cladomics cladogram Echyridella MARGARITIFERINAE GONIDEINAE UNIONINAE AMBLEMINAE

Huang et al., 2018 [Publ 3828], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 41 0 0 --
Cladomics cladogram Echyridella GIBBOSULINAE MARGARITIFERINAE UNIONINAE AMBLEMINAE GONIDEINAE

Huang et al., 2018 [Publ 3828], fig. 6-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 16 0 0 --
Cladomics cladogram Echyridella MARGARITIFERINAE AMBLEMINAE GONIDEINAE UNIONINAE

Chase et al., 2018 [Publ 3847], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 38 0 0 --
Cladomics cladogram Echyridella AMBLEMINAE

Chase et al., 2018 [Publ 3847], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 0 0 --
Cladomics cladogram MARGARITIFERINAE Echyridella

Chase et al., 2018 [Publ 3861], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 0 0 --
Cladomics cladogram Echyridella MARGARITIFERINAE AMBLEMINAE GONIDEINAE UNIONINAE

Chase et al., 2018 [Publ 3861], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 0 0 --
Cladomics cladogram MARGARITIFERINAE Echyridella

Huang et al., 2019 [Publ 3843], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 55 0 0 --
Cladomics cladogram Echyridella GIBBOSULINAE MARGARITIFERINAE UNIONINAE GONIDEINAE AMBLEMINAE

Huang et al., 2019 [Publ 3843], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 0 0 --
Cladomics cladogram Echyridella MARGARITIFERINAE AMBLEMINAE GONIDEINAE UNIONINAE

Wu et al., 2019 [Publ 3864], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 45 0 0 --
Cladomics cladogram Echyridella IRIDININAE

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 0 0 --
Cladomics cladogram HYRIDELLINI MARGARITIFERINAE AMBLEMINAE GONIDEINAE UNIONINAE PARREYSIINAE

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 0 0 --
Cladomics cladogram HYRIDELLINI MARGARITIFERINAE

Wu et al., 2020 [Publ 4121], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 0 0 --
Cladomics cladogram Echyridella ANODONTITINAE IRIDININAE GIBBOSULINAE MARGARITIFERINAE AMBLEMINAE GONIDEINAE UNIONINAE

Olivera-Hyde et al., 2020 [Publ 4249], fig. 10

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 0 0 0 8.0
Cladomics cladogram CASTALIINI

Wu et al., 2021 [Publ 4366], fig. 3-BI

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 72 0 0 --
Cladomics cladogram Echyridella ANODONTITINAE IRIDININAE GIBBOSULINAE MARGARITIFERINAE AMBLEMINAE GONIDEINAE UNIONINAE

Wu et al., 2021 [Publ 4366], fig. 3-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 72 0 0 --
Cladomics cladogram Echyridella ANODONTITINAE IRIDININAE GIBBOSULINAE MARGARITIFERINAE AMBLEMINAE GONIDEINAE UNIONINAE

Wu et al., 2021 [Publ 4366], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 67 0 0 --
Cladomics cladogram Echyridella IRIDININAE ANODONTITINAE GIBBOSULINAE MARGARITIFERINAE AMBLEMINAE GONIDEINAE UNIONINAE

Wu et al., 2022 [Publ 4238], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 72 0 0 --
Cladomics cladogram Echyridella GIBBOSULINAE MARGARITIFERINAE AMBLEMINAE GONIDEINAE UNIONINAE

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 0 0 --
Cladomics cladogram Echyridella ANODONTITINAE IRIDININAE GIBBOSULINAE MARGARITIFERINAE AMBLEMINAE GONIDEINAE PARREYSIINAE UNIONINAE

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 0 0 --
Cladomics cladogram Echyridella ANODONTITINAE IRIDININAE GIBBOSULINAE MARGARITIFERINAE AMBLEMINAE GONIDEINAE PARREYSIINAE UNIONINAE

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 0 0 --
Cladomics cladogram Echyridella ANODONTITINAE IRIDININAE MARGARITIFERINAE AMBLEMINAE GONIDEINAE PARREYSIINAE UNIONINAE

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 0 0 --
Cladomics cladogram Echyridella ANODONTITINAE IRIDININAE MARGARITIFERINAE AMBLEMINAE GONIDEINAE PARREYSIINAE UNIONINAE

Dai et al., 2025 [Publ 4440], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 86 0 0 --
Cladomics cladogram Echyridella GIBBOSULINAE MARGARITIFERINAE AMBLEMINAE GONIDEINAE UNIONINAE

Chen et al., 2025 [Publ 4448], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 84 0 0 --
Cladomics cladogram Echyridella GIBBOSULINAE MARGARITIFERINAE

Prié et al., 2026 [Publ 4844], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram CASTALIINI MARGARITIFERINAE