Bivalvia | Unionida | Hyrioidea | Hyriidae

subfamily HYRIINAE Swainson, 1840: cladomics

4 tribes, 1 genus

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 76 (82 total)
number of subtaxa (rank: tribe) = 5
analyzed in 96 cladograms in 50 publications
maximum number of terminals in an analysis = 18 (mean = 2.83, median = 2)
maximum number of subtaxa (tribe) = 5 (mean = 2.18, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 54
maximum number of included terminals = 18, (mean = 4.26 median = 3)
maximum number of subtaxa (tribe) = 5 (mean = 3.09, median = 3)
proportion of cladograms that support basic monophyly = 0.48

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 49
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (tribe) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 18 (mean = 4.29, median = 3)
maximum number of subtaxa (tribe) = 5 (mean = 3.18, median = 3)
proportion of cladograms that support rigorous monophyly = 0.43

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Hyriinae cladogram distances
PCoA Ordination

NJ Tree of Hyriinae cladogram distances
NJ Tree

Cladograms

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
18 26 32 5 614 10 1 0 
  +----------------------------------- Westralunio           
  |                                                          
  |                             +----- Diplodon_suavidicus__2 **
  |  +--------------------------+                            
--+  |                          |  +-- Hyriini **               
  |  |                          +--+                         
  |  |                             +-- Castaliini **            
  |  |                                                       
  |  |                             +-- Hyridella_depressa **    
  +--+                       +-----+                         
     |                       |     +-- Hyridella_drapeta **     
     |  +--------------------+                               
     |  |                    |  +----- Diplodon_suavidicus__1 **
     |  |                    +--+                            
     |  |                       |  +-- Diplodon_chilensis **    
     |  |                       +--+                         
     +--+                          +-- Diplodon_gratus **       
        |                                                    
        |                          +-- Cucumerunio **           
        |  +-----------------------+                         
        |  |                       +-- Hyridella_australis **   
        |  |                                                 
        +--+  +----------------------- Echyridella **           
           |  |                                              
           |  |              +-------- Velesunio__1          
           +--+  +-----------+                               
              |  |           |  +----- Alathyria_jacksoni__1 
              |  |           +--+                            
              |  |              |  +-- Velesunio_ambiguus    
              +--+              +--+                         
                 |                 +-- Alathyria_profuga     
                 |                                           
                 |  +----------------- Velesunio__3          
                 |  |                                        
                 +--+  +-------------- Microdontia           
                    |  |                                     
                    +--+           +-- Lortiella             
                       |  +--------+                         
                       |  |        +-- Velesunio_angasi      
                       +--+                                  
                          |  +-------- Alathyria_pertexta__2 
                          +--+                               
                             |  +----- Velesunio__2          
                             +--+                            
                                |  +-- Alathyria_jacksoni__2 
                                +--+                         
                                   +-- Alathyria_pertexta__1

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

iTtTbiriSoTroTroSiD
13 19 20 5 7 6 1 20.5 
                 +----- Diplodon_suavidicus__1 **
  +--------------+                            
  |              |  +-- Hyriini **               
  |              +--+                         
  |                 +-- Castaliini **            
  |                                           
  |           +-------- Westralunio           
  |           |                               
  +-----------+     +-- Hyridella_drapeta **     
  |           |  +--+                         
--+           |  |  +-- Hyridella_depressa **    
  |           +--+                            
  |              |  +-- Diplodon_gratus **       
  |              +--+                         
  |                 +-- Diplodon_suavidicus__2 **
  |                                           
  |                 +-- Hyridella_australis **   
  |  +--------------+                         
  |  |              +-- Cucumerunio **           
  +--+                                        
     |  +-------------- Echyridella **           
     +--+                                     
        |  +----------- Velesunio_ambiguus    
        +--+                                  
           |  +-------- Microdontia           
           +--+                               
              |  +----- Lortiella             
              +--+                            
                 |  +-- Alathyria             
                 +--+                         
                    +-- Velesunio_angasi

Santos-Neto et al., 2016 [Publ 3739], fig. 4

iTtTbiriSoTroTroSiD
13 19 21 5 7 6 1 21.5 
                    +----- Diplodon_suavidicus__2 **
  +-----------------+                            
  |                 |  +-- Hyriini **               
  |                 +--+                         
  |                    +-- Castaliini **            
  |                                              
  |              +-------- Westralunio           
--+              |                               
  |  +-----------+     +-- Hyridella_drapeta **     
  |  |           |  +--+                         
  |  |           |  |  +-- Hyridella_depressa **    
  |  |           +--+                            
  |  |              |  +-- Diplodon_suavidicus__1 **
  +--+              +--+                         
     |                 +-- Diplodon_gratus **       
     |                                           
     |  +----------------- Echyridella **           
     |  |                                        
     +--+              +-- Cucumerunio **           
        |  +-----------+                         
        |  |           +-- Hyridella_australis **   
        +--+                                     
           |  +----------- Velesunio_ambiguus    
           +--+                                  
              |  +-------- Alathyria             
              +--+                               
                 |  +----- Velesunio_angasi      
                 +--+                            
                    |  +-- Lortiella             
                    +--+                         
                       +-- Microdontia

Santos-Neto et al., 2016 [Publ 3739], fig. 3_5

iTtTbiriSoTroTroSiD
7 7 6 3 0 0 0 27 
  +----- Rhipidodontini **
--+                    
  |  +-- Hyriini **       
  +--+                 
     +-- Castaliini **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
7 7 5 5 191 7 1 33 
     +-------------- Westralunio       
  +--+                                 
  |  |  +----------- Echyridella **       
  |  +--+                              
  |     |  +-------- Hyridellini **       
  |     +--+                           
  |        |  +----- Rhipidodontini **    
  |        +--+                        
--+           |  +-- Castaliini **        
  |           +--+                     
  |              +-- Hyriini **           
  |                                    
  |           +----- Velesunio         
  |        +--+                        
  |        |  |  +-- Alathyria_jacksoni
  |        |  +--+                     
  +--------+     +-- Alathyria_profuga 
           |                           
           |  +----- Alathyria_pertexta
           +--+                        
              |  +-- Lortiella         
              +--+                     
                 +-- Microdontia

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
7 14 9 5 193 7 1 37 
  +----------- Echyridella **       
  |                              
  +----------- Westralunio       
  |                              
  +----------- Hyridellini **       
  |                              
  |     +----- Rhipidodontini **    
  +-----+                        
--+     |  +-- Castaliini **        
  |     +--+                     
  |        +-- Hyriini **           
  |                              
  |     +----- Velesunio         
  |  +--+                        
  |  |  |  +-- Alathyria_jacksoni
  |  |  +--+                     
  +--+     +-- Alathyria_profuga 
     |                           
     |  +----- Alathyria_pertexta
     +--+                        
        |  +-- Lortiella         
        +--+                     
           +-- Microdontia

Marshall et al., 2014 [Publ 3593], fig. 1

iTtTbiriSoTroTroSiD
7 10 8 3 7 3 1 29.5 
  +-------- Echyridella **   
--+                       
  |  +----- Velesunioninae
  +--+                    
     |  +-- Hyridellini **   
     +--+                 
        +-- Rhipidodontini **

Graf et al., 2015 [Publ 3619], fig. 1

iTtTbiriSoTroTroSiD
7 7 5 4 13 7 1 31 
  +----------- Velesunioninae
--+                          
  |  +-------- Hyridellini **   
  +--+                       
     |  +----- Rhipidodontini **
     +--+                    
        |  +-- Hyriini **       
        +--+                 
           +-- Castaliini **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
7 7 5 5 193 7 1 33 
     +-------------- Westralunio       
  +--+                                 
  |  |  +----------- Echyridella **       
  |  +--+                              
  |     |  +-------- Hyridellini **       
  |     +--+                           
  |        |  +----- Rhipidodontini **    
  |        +--+                        
--+           |  +-- Castaliini **        
  |           +--+                     
  |              +-- Hyriini **           
  |                                    
  |           +----- Alathyria_jacksoni
  |        +--+                        
  |        |  |  +-- Alathyria_profuga 
  |        |  +--+                     
  +--------+     +-- Velesunio         
           |                           
           |  +----- Alathyria_pertexta
           +--+                        
              |  +-- Lortiella         
              +--+                     
                 +-- Microdontia

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
6 6 4 4 40 3 1 31.5 
  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Hyridellini **   
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Rhipidodontini **

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
6 6 4 4 40 3 1 31.5 
  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Hyridellini **   
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Rhipidodontini **

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
6 6 4 4 41 3 1 31.5 
  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Hyridellini **   
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Rhipidodontini **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
6 6 4 4 41 3 1 31.5 
  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Hyridellini **   
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Rhipidodontini **

Walker et al., 2006 [Publ 2666], fig. 6

iTtTbiriSoTroTroSiD
5 10 7 4 31 5 1 27 
  +-------- Castaliini **         
  |                            
  +-------- Rhipidodontini **     
--+                            
  |  +----- Hyridella_depressa ** 
  |  |                         
  +--+----- Hyridella_australis **
     |                         
     |  +-- Echyridella **        
     +--+                      
        +-- Velesunioninae

Baker et al., 2003 [Publ 2690], fig. 5

iTtTbiriSoTroTroSiD
5 7 5 4 3 2 1 35 
           +-- Hyridella_australis **
  +--------+                      
  |        +-- Rhipidodontini **     
--+                               
  |  +-------- Hyridella_depressa ** 
  +--+                            
     |  +----- Velesunioninae     
     +--+                         
        |  +-- Echyridella **        
        +--+                      
           +-- Castaliini **

Baker et al., 2004 [Publ 2685], fig. 2-3

iTtTbiriSoTroTroSiD
5 8 9 4 4 3 1 33 
     +----- Hyridella_depressa ** 
  +--+                         
  |  |  +-- Hyridella_australis **
  |  +--+                      
--+     +-- Rhipidodontini **     
  |                            
  |  +----- Velesunioninae     
  +--+                         
     |  +-- Echyridella **        
     +--+                      
        +-- Castaliini **

Santos-Neto et al., 2018 [Publ 3888], fig. 5

iTtTbiriSoTroTroSiD
5 5 3 2 0 0 0 29 
  +-- Hyriini **   
--+             
  +-- Castaliini **

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

iTtTbiriSoTroTroSiD
5 5 4 4 6 1 1 33.5 
  +-------------- Velesunioninae        
  |                                     
--+           +-- Rhipidodontini **        
  |  +--------+                         
  |  |        +-- Castaliini **            
  +--+                                  
     |  +-------- Hyridella_australis__2 **
     +--+                               
        |  +----- Hyridella_australis__1 **
        +--+                            
           |  +-- Echyridella **           
           +--+                         
              +-- Hyridella_depressa **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 4 37 1 1 32 
  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Rhipidodontini **
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Hyriini **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
4 4 2 4 137 1 1 32 
  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Rhipidodontini **
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Hyriini **

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
4 5 3 4 37 1 1 32 
        +-- Echyridella **   
  +-----+                 
  |     +-- Velesunioninae
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Castaliini **    
     +--+                 
        +-- Hyriini **

Bolotov et al., 2016 [Publ 3743], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 27 6 1 31 
  +-------- Velesunioninae
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Hyriini **       
     +--+                 
        +-- Castaliini **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
3 3 1 3 135 6 1 31 
  +-------- Velesunioninae
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Castaliini **    
     +--+                 
        +-- Hyriini **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
3 5 1 3 28 2 1 33 
  +----- Rhipidodontini **
  |                    
  +----- Echyridella **   
--+                    
  +----- Lortiella     
  |                    
  |  +-- Castaliini **    
  +--+                 
     +-- Velesunio

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
3 5 3 3 31 2 1 31 
     +-- Castaliini **    
  +--+                 
  |  +-- Rhipidodontini **
--+                    
  |  +-- Echyridella **   
  +--+                 
     +-- Velesunioninae

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
3 5 3 3 28 2 1 31 
     +-- Castaliini **    
  +--+                 
  |  +-- Rhipidodontini **
--+                    
  |  +-- Echyridella **   
  +--+                 
     +-- Velesunioninae

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 37 1 1 31 
  +-------- Velesunioninae
--+                       
  |  +----- Hyridellini **   
  +--+                    
     |  +-- Hyriini **       
     +--+                 
        +-- Castaliini **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
3 3 1 3 135 6 1 31 
  +-------- Velesunioninae
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Castaliini **    
     +--+                 
        +-- Hyriini **

Olivera-Hyde et al., 2020 [Publ 4249], fig. 10

iTtTbiriSoTroTroSiD
3 3 1 1 0 0 0 31 
1

Bolotov et al., 2016 [Publ 3743], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 27 6 1 31 
  +-------- Velesunioninae
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Hyriini **       
     +--+                 
        +-- Castaliini **

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
3 5 3 3 28 2 1 31.5 
     +-- Castaliini **    
  +--+                 
  |  +-- Rhipidodontini **
--+                    
  |  +-- Echyridella **   
  +--+                 
     +-- Velesunioninae

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
3 5 3 3 28 2 1 31 
     +-- Castaliini **    
  +--+                 
  |  +-- Rhipidodontini **
--+                    
  |  +-- Echyridella **   
  +--+                 
     +-- Velesunioninae

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
3 5 2 3 28 2 1 34 
     +-- Castaliini **    
  +--+                 
  |  +-- Velesunio     
--+                    
  |  +-- Echyridella **   
  |  |                 
  +--+-- Lortiella     
     |                 
     +-- Rhipidodontini **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
3 3 1 3 33 1 1 31 
  +-------- Velesunioninae
--+                       
  |  +----- Hyridellini **   
  +--+                    
     |  +-- Castaliini **    
     +--+                 
        +-- Hyriini **

Graf, 2000 [Publ 2647], fig. 1

iTtTbiriSoTroTroSiD
3 4 2 3 14 1 1 32.5 
  +-------- Castaliini **    
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Hyridellini **   
     +--+                 
        +-- Velesunioninae

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
3 3 1 3 28 0 0 31 
  +----- Rhipidodontini **
--+                    
  |  +-- Hyriini **       
  +--+                 
     +-- Castaliini **

Graf, 2002 [Publ 2651], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 15 1 1 30.5 
  +-------- Velesunioninae
--+                       
  |  +----- Hyridellini **   
  +--+                    
     |  +-- Rhipidodontini **
     +--+                 
        +-- Castaliini **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
3 3 1 3 33 1 1 31 
  +-------- Velesunioninae
--+                       
  |  +----- Hyridellini **   
  +--+                    
     |  +-- Castaliini **    
     +--+                 
        +-- Hyriini **

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
2 3 0 2 14 1 1 31 
  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae
  +--+                 
     +-- Hyridellini **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
2 3 0 2 66 1 1 31 
  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
2 3 0 2 27 1 1 31 
  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae
  +--+                 
     +-- Hyridellini **

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
2 3 0 2 13 1 1 32 
  +-- Hyriini **       
  |                 
--+-- Hyridellini **   
  |                 
  +-- Velesunioninae

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
2 3 0 2 32 1 1 31 
  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iTtTbiriSoTroTroSiD
2 3 0 2 26 1 1 31 
  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae
  +--+                 
     +-- Hyridellini **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 2 89 1 1 -- 
  +----- Velesunioninae
--+                    
  |  +-- Rhipidodontini **
  +--+                 
     +-- Castaliini **

Combosch et al., 2017 [Publ 3806], fig. 1

iTtTbiriSoTroTroSiD
2 3 0 2 13 1 1 31 
  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae
  +--+                 
     +-- Hyridellini **

Combosch et al., 2017 [Publ 3806], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 13 1 1 -- 
  +----- Velesunioninae
--+                    
  |  +-- Hyriini **       
  +--+                 
     +-- Hyridellini **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
2 3 0 2 66 1 1 31 
  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
2 2 0 2 89 1 1 -- 
  +----- Velesunioninae
--+                    
  |  +-- Castaliini **    
  +--+                 
     +-- Rhipidodontini **

Chen et al., 2025 [Publ 4455], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 24 0 0 -- 
  +-- Hyridellini **
--+              
  +-- Hyriini **

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
2 3 0 2 32 1 1 31 
  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
2 3 0 2 13 1 1 32 
  +-- Hyriini **       
  |                 
--+-- Velesunioninae
  |                 
  +-- Hyridellini **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
2 3 0 2 66 1 1 31 
  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iTtTbiriSoTroTroSiD
2 3 0 2 26 1 1 31 
  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae
  +--+                 
     +-- Hyridellini **

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 22 0 0 -- 
-- Echyridella_menziesii **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 68 0 0 -- 
-- Echyridella_menziesii **

Walker et al., 2006 [Publ 2666], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Hyridella **

Chase et al., 2018 [Publ 3861], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Echyridella_menziesii **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
1 1 0 1 95 0 0 -- 
-- Echyridella_menziesii **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
1 1 0 1 73 0 0 -- 
-- Echyridella_menziesii **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Hyridella_australis **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
1 1 0 1 121 1 1 -- 
  +-- Hyridellini **   
--+                 
  +-- Velesunioninae

Hoeh et al., 2002 [Publ 2673], fig. 1-F

iTtTbiriSoTroTroSiD
1 1 0 1 8 0 0 -- 
-- Echyridella_menziesii **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 45 0 0 -- 
-- Echyridella_menziesii **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 73 0 0 -- 
-- Echyridella_menziesii **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
1 1 0 1 79 1 1 -- 
  +-- Hyridellini **   
--+                 
  +-- Velesunioninae

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 48 0 0 -- 
-- Echyridella_menziesii **

Klunzinger et al., 2021 [Publ 4086], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 5 3 1 -- 
  +----- Westralunio
--+                 
  |  +-- Velesunio  
  +--+              
     +-- Hyridellini **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 55 0 0 -- 
-- Echyridella_menziesii **

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 1 1 -- 
  +-- Echyridella **   
--+                 
  +-- Velesunioninae

Guerra et al., 2017 [Publ 3772], fig. 2a-M

iTtTbiriSoTroTroSiD
1 1 0 1 7 0 0 -- 
-- Echyridella_menziesii **

Walker et al., 2006 [Publ 2666], fig. 6-M

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Echyridella_menziesii **

Guerra et al., 2017 [Publ 3772], fig. 2b-M

iTtTbiriSoTroTroSiD
1 1 0 1 7 0 0 -- 
-- Echyridella_menziesii **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Hyridella_depressa **

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 1 1 -- 
  +-- Echyridella **   
--+                 
  +-- Velesunioninae

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
1 1 0 1 95 0 0 -- 
-- Echyridella_menziesii **

Hoeh et al., 2002 [Publ 2673], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 8 0 0 -- 
-- Echyridella_menziesii **

Chase et al., 2018 [Publ 3861], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Echyridella_menziesii **

Klunzinger et al., 2022 [Publ 4259], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 3 3 1 -- 
  +-- Hyridellini **   
--+                 
  +-- Velesunioninae

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
1 1 0 1 91 0 0 -- 
-- Echyridella_menziesii **

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 41 0 0 -- 
-- Echyridella_menziesii **

Walker et al., 2006 [Publ 2666], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Hyridella **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
1 1 0 1 73 0 0 -- 
-- Echyridella_menziesii **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Hyridella_depressa **

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
1 1 0 1 24 0 0 -- 
-- Echyridella_menziesii **

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
1 1 0 1 24 0 0 -- 
-- Echyridella_menziesii **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Hyridella_australis **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
1 1 0 1 91 0 0 -- 
-- Echyridella_menziesii **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 87 0 0 -- 
-- Echyridella_menziesii **

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 46 1 1 -- 
  +-- Echyridella **   
--+                 
  +-- Velesunioninae

Breton et al., 2011 [Publ 3376], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 37 0 0 -- 
-- Echyridella_menziesii **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 85 0 0 -- 
-- Echyridella_menziesii **

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
1 1 0 1 16 0 0 -- 
-- Echyridella_menziesii **

Walker et al., 2006 [Publ 2666], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Hyridella **

Chase et al., 2018 [Publ 3847], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Echyridella_menziesii **

Chase et al., 2018 [Publ 3847], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 38 0 0 -- 
-- Echyridella_menziesii **