Bivalvia | Unionida | Hyrioidea | Hyriidae

subfamily HYRIINAE Swainson, 1840: cladomics

4 tribes, 1 genus

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 76 (82 total)
number of subtaxa (rank: tribe) = 5
analyzed in 96 cladograms in 50 publications
maximum number of terminals in an analysis = 18 (mean = 2.83, median = 2)
maximum number of subtaxa (tribe) = 5 (mean = 2.18, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 54
maximum number of included terminals = 18, (mean = 4.26 median = 3)
maximum number of subtaxa (tribe) = 5 (mean = 3.09, median = 3)
proportion of cladograms that support basic monophyly = 0.48

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 49
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (tribe) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 18 (mean = 4.29, median = 3)
maximum number of subtaxa (tribe) = 5 (mean = 3.18, median = 3)
proportion of cladograms that support rigorous monophyly = 0.43

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Hyriinae cladogram distances
PCoA Ordination

NJ Tree of Hyriinae cladogram distances
NJ Tree

Cladograms

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
18 26 32 5 613 10 1 0 
  +----------------------------------- Westralunio           
  |                                                          
  |                             +----- Diplodon_suavidicus__2 **
  |  +--------------------------+                            
--+  |                          |  +-- Hyriini **               
  |  |                          +--+                         
  |  |                             +-- Castaliini **            
  |  |                                                       
  |  |                             +-- Hyridella_depressa **    
  +--+                       +-----+                         
     |                       |     +-- Hyridella_drapeta **     
     |  +--------------------+                               
     |  |                    |  +----- Diplodon_suavidicus__1 **
     |  |                    +--+                            
     |  |                       |  +-- Diplodon_chilensis **    
     |  |                       +--+                         
     +--+                          +-- Diplodon_gratus **       
        |                                                    
        |                          +-- Cucumerunio **           
        |  +-----------------------+                         
        |  |                       +-- Hyridella_australis **   
        |  |                                                 
        +--+  +----------------------- Echyridella **           
           |  |                                              
           |  |              +-------- Velesunio__1          
           +--+  +-----------+                               
              |  |           |  +----- Alathyria_jacksoni__1 
              |  |           +--+                            
              |  |              |  +-- Velesunio_ambiguus    
              +--+              +--+                         
                 |                 +-- Alathyria_profuga     
                 |                                           
                 |  +----------------- Velesunio__3          
                 |  |                                        
                 +--+  +-------------- Microdontia           
                    |  |                                     
                    +--+           +-- Lortiella             
                       |  +--------+                         
                       |  |        +-- Velesunio_angasi      
                       +--+                                  
                          |  +-------- Alathyria_pertexta__2 
                          +--+                               
                             |  +----- Velesunio__2          
                             +--+                            
                                |  +-- Alathyria_jacksoni__2 
                                +--+                         
                                   +-- Alathyria_pertexta__1

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

iTtTbiriSoTroTroSiD
13 19 20 5 7 6 1 20.5 
                 +----- Diplodon_suavidicus__1 **
  +--------------+                            
  |              |  +-- Hyriini **               
  |              +--+                         
  |                 +-- Castaliini **            
  |                                           
  |           +-------- Westralunio           
  |           |                               
  +-----------+     +-- Hyridella_drapeta **     
  |           |  +--+                         
--+           |  |  +-- Hyridella_depressa **    
  |           +--+                            
  |              |  +-- Diplodon_gratus **       
  |              +--+                         
  |                 +-- Diplodon_suavidicus__2 **
  |                                           
  |                 +-- Hyridella_australis **   
  |  +--------------+                         
  |  |              +-- Cucumerunio **           
  +--+                                        
     |  +-------------- Echyridella **           
     +--+                                     
        |  +----------- Velesunio_ambiguus    
        +--+                                  
           |  +-------- Microdontia           
           +--+                               
              |  +----- Lortiella             
              +--+                            
                 |  +-- Alathyria             
                 +--+                         
                    +-- Velesunio_angasi

Santos-Neto et al., 2016 [Publ 3739], fig. 4

iTtTbiriSoTroTroSiD
13 19 21 5 7 6 1 21.5 
                    +----- Diplodon_suavidicus__2 **
  +-----------------+                            
  |                 |  +-- Hyriini **               
  |                 +--+                         
  |                    +-- Castaliini **            
  |                                              
  |              +-------- Westralunio           
--+              |                               
  |  +-----------+     +-- Hyridella_drapeta **     
  |  |           |  +--+                         
  |  |           |  |  +-- Hyridella_depressa **    
  |  |           +--+                            
  |  |              |  +-- Diplodon_suavidicus__1 **
  +--+              +--+                         
     |                 +-- Diplodon_gratus **       
     |                                           
     |  +----------------- Echyridella **           
     |  |                                        
     +--+              +-- Cucumerunio **           
        |  +-----------+                         
        |  |           +-- Hyridella_australis **   
        +--+                                     
           |  +----------- Velesunio_ambiguus    
           +--+                                  
              |  +-------- Alathyria             
              +--+                               
                 |  +----- Velesunio_angasi      
                 +--+                            
                    |  +-- Lortiella             
                    +--+                         
                       +-- Microdontia

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
7 7 5 5 193 7 1 33 
     +-------------- Westralunio       
  +--+                                 
  |  |  +----------- Echyridella **       
  |  +--+                              
  |     |  +-------- Hyridellini **       
  |     +--+                           
  |        |  +----- Rhipidodontini **    
  |        +--+                        
--+           |  +-- Castaliini **        
  |           +--+                     
  |              +-- Hyriini **           
  |                                    
  |           +----- Alathyria_jacksoni
  |        +--+                        
  |        |  |  +-- Alathyria_profuga 
  |        |  +--+                     
  +--------+     +-- Velesunio         
           |                           
           |  +----- Alathyria_pertexta
           +--+                        
              |  +-- Lortiella         
              +--+                     
                 +-- Microdontia

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
7 7 5 5 191 7 1 33 
     +-------------- Westralunio       
  +--+                                 
  |  |  +----------- Echyridella **       
  |  +--+                              
  |     |  +-------- Hyridellini **       
  |     +--+                           
  |        |  +----- Rhipidodontini **    
  |        +--+                        
--+           |  +-- Castaliini **        
  |           +--+                     
  |              +-- Hyriini **           
  |                                    
  |           +----- Velesunio         
  |        +--+                        
  |        |  |  +-- Alathyria_jacksoni
  |        |  +--+                     
  +--------+     +-- Alathyria_profuga 
           |                           
           |  +----- Alathyria_pertexta
           +--+                        
              |  +-- Lortiella         
              +--+                     
                 +-- Microdontia

Marshall et al., 2014 [Publ 3593], fig. 1

iTtTbiriSoTroTroSiD
7 10 8 3 7 3 1 29.5 
  +-------- Echyridella **   
--+                       
  |  +----- Velesunioninae
  +--+                    
     |  +-- Hyridellini **   
     +--+                 
        +-- Rhipidodontini **

Santos-Neto et al., 2016 [Publ 3739], fig. 3_5

iTtTbiriSoTroTroSiD
7 7 6 3 0 0 0 27 
  +----- Rhipidodontini **
--+                    
  |  +-- Hyriini **       
  +--+                 
     +-- Castaliini **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
7 14 9 5 193 7 1 37 
  +----------- Echyridella **       
  |                              
  +----------- Westralunio       
  |                              
  +----------- Hyridellini **       
  |                              
  |     +----- Rhipidodontini **    
  +-----+                        
--+     |  +-- Castaliini **        
  |     +--+                     
  |        +-- Hyriini **           
  |                              
  |     +----- Velesunio         
  |  +--+                        
  |  |  |  +-- Alathyria_jacksoni
  |  |  +--+                     
  +--+     +-- Alathyria_profuga 
     |                           
     |  +----- Alathyria_pertexta
     +--+                        
        |  +-- Lortiella         
        +--+                     
           +-- Microdontia

Graf et al., 2015 [Publ 3619], fig. 1

iTtTbiriSoTroTroSiD
7 7 5 4 13 7 1 31 
  +----------- Velesunioninae
--+                          
  |  +-------- Hyridellini **   
  +--+                       
     |  +----- Rhipidodontini **
     +--+                    
        |  +-- Hyriini **       
        +--+                 
           +-- Castaliini **

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
6 6 4 4 41 3 1 31.5 
  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Hyridellini **   
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Rhipidodontini **

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
6 6 4 4 40 3 1 31.5 
  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Hyridellini **   
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Rhipidodontini **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
6 6 4 4 41 3 1 31.5 
  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Hyridellini **   
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Rhipidodontini **

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
6 6 4 4 40 3 1 31.5 
  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Hyridellini **   
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Rhipidodontini **

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

iTtTbiriSoTroTroSiD
5 5 4 4 6 1 1 33.5 
  +-------------- Velesunioninae        
  |                                     
--+           +-- Rhipidodontini **        
  |  +--------+                         
  |  |        +-- Castaliini **            
  +--+                                  
     |  +-------- Hyridella_australis__2 **
     +--+                               
        |  +----- Hyridella_australis__1 **
        +--+                            
           |  +-- Echyridella **           
           +--+                         
              +-- Hyridella_depressa **

Walker et al., 2006 [Publ 2666], fig. 6

iTtTbiriSoTroTroSiD
5 10 7 4 31 5 1 27 
  +-------- Castaliini **         
  |                            
  +-------- Rhipidodontini **     
--+                            
  |  +----- Hyridella_depressa ** 
  |  |                         
  +--+----- Hyridella_australis **
     |                         
     |  +-- Echyridella **        
     +--+                      
        +-- Velesunioninae

Baker et al., 2004 [Publ 2685], fig. 2-3

iTtTbiriSoTroTroSiD
5 8 9 4 4 3 1 33 
     +----- Hyridella_depressa ** 
  +--+                         
  |  |  +-- Hyridella_australis **
  |  +--+                      
--+     +-- Rhipidodontini **     
  |                            
  |  +----- Velesunioninae     
  +--+                         
     |  +-- Echyridella **        
     +--+                      
        +-- Castaliini **

Baker et al., 2003 [Publ 2690], fig. 5

iTtTbiriSoTroTroSiD
5 7 5 4 3 2 1 35 
           +-- Hyridella_australis **
  +--------+                      
  |        +-- Rhipidodontini **     
--+                               
  |  +-------- Hyridella_depressa ** 
  +--+                            
     |  +----- Velesunioninae     
     +--+                         
        |  +-- Echyridella **        
        +--+                      
           +-- Castaliini **

Santos-Neto et al., 2018 [Publ 3888], fig. 5

iTtTbiriSoTroTroSiD
5 5 3 2 0 0 0 29 
  +-- Hyriini **   
--+             
  +-- Castaliini **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
4 4 2 4 137 1 1 32 
  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Rhipidodontini **
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Hyriini **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 4 37 1 1 32 
  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Rhipidodontini **
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Hyriini **

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
4 5 3 4 37 1 1 32 
        +-- Echyridella **   
  +-----+                 
  |     +-- Velesunioninae
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Castaliini **    
     +--+                 
        +-- Hyriini **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
3 3 1 3 33 1 1 31 
  +-------- Velesunioninae
--+                       
  |  +----- Hyridellini **   
  +--+                    
     |  +-- Castaliini **    
     +--+                 
        +-- Hyriini **

Graf, 2002 [Publ 2651], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 15 1 1 30.5 
  +-------- Velesunioninae
--+                       
  |  +----- Hyridellini **   
  +--+                    
     |  +-- Rhipidodontini **
     +--+                 
        +-- Castaliini **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
3 3 1 3 33 1 1 31 
  +-------- Velesunioninae
--+                       
  |  +----- Hyridellini **   
  +--+                    
     |  +-- Castaliini **    
     +--+                 
        +-- Hyriini **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
3 3 1 3 135 6 1 31 
  +-------- Velesunioninae
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Castaliini **    
     +--+                 
        +-- Hyriini **

Bolotov et al., 2016 [Publ 3743], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 27 6 1 31 
  +-------- Velesunioninae
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Hyriini **       
     +--+                 
        +-- Castaliini **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
3 5 2 3 28 2 1 34 
     +-- Castaliini **    
  +--+                 
  |  +-- Velesunio     
--+                    
  |  +-- Echyridella **   
  |  |                 
  +--+-- Lortiella     
     |                 
     +-- Rhipidodontini **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 37 1 1 31 
  +-------- Velesunioninae
--+                       
  |  +----- Hyridellini **   
  +--+                    
     |  +-- Hyriini **       
     +--+                 
        +-- Castaliini **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
3 5 3 3 28 2 1 31 
     +-- Castaliini **    
  +--+                 
  |  +-- Rhipidodontini **
--+                    
  |  +-- Echyridella **   
  +--+                 
     +-- Velesunioninae

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
3 5 3 3 28 2 1 31 
     +-- Castaliini **    
  +--+                 
  |  +-- Rhipidodontini **
--+                    
  |  +-- Echyridella **   
  +--+                 
     +-- Velesunioninae

Graf, 2000 [Publ 2647], fig. 1

iTtTbiriSoTroTroSiD
3 4 2 3 14 1 1 32.5 
  +-------- Castaliini **    
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Hyridellini **   
     +--+                 
        +-- Velesunioninae

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
3 5 1 3 28 2 1 33 
  +----- Rhipidodontini **
  |                    
  +----- Echyridella **   
--+                    
  +----- Lortiella     
  |                    
  |  +-- Castaliini **    
  +--+                 
     +-- Velesunio

Olivera-Hyde et al., 2020 [Publ 4249], fig. 10

iTtTbiriSoTroTroSiD
3 3 1 1 0 0 0 31 
1

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
3 3 1 3 28 0 0 31 
  +----- Rhipidodontini **
--+                    
  |  +-- Hyriini **       
  +--+                 
     +-- Castaliini **

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
3 5 3 3 31 2 1 31 
     +-- Castaliini **    
  +--+                 
  |  +-- Rhipidodontini **
--+                    
  |  +-- Echyridella **   
  +--+                 
     +-- Velesunioninae

Bolotov et al., 2016 [Publ 3743], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 27 6 1 31 
  +-------- Velesunioninae
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Hyriini **       
     +--+                 
        +-- Castaliini **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
3 3 1 3 135 6 1 31 
  +-------- Velesunioninae
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Castaliini **    
     +--+                 
        +-- Hyriini **

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
3 5 3 3 28 2 1 31.5 
     +-- Castaliini **    
  +--+                 
  |  +-- Rhipidodontini **
--+                    
  |  +-- Echyridella **   
  +--+                 
     +-- Velesunioninae

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
2 3 0 2 66 1 1 31 
  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
2 3 0 2 13 1 1 32 
  +-- Hyriini **       
  |                 
--+-- Hyridellini **   
  |                 
  +-- Velesunioninae

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
2 3 0 2 32 1 1 31 
  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
2 3 0 2 32 1 1 31 
  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
2 3 0 2 66 1 1 31 
  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
2 3 0 2 13 1 1 32 
  +-- Hyriini **       
  |                 
--+-- Velesunioninae
  |                 
  +-- Hyridellini **

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iTtTbiriSoTroTroSiD
2 3 0 2 26 1 1 31 
  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae
  +--+                 
     +-- Hyridellini **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 2 89 1 1 -- 
  +----- Velesunioninae
--+                    
  |  +-- Rhipidodontini **
  +--+                 
     +-- Castaliini **

Combosch et al., 2017 [Publ 3806], fig. 1

iTtTbiriSoTroTroSiD
2 3 0 2 13 1 1 31 
  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae
  +--+                 
     +-- Hyridellini **

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
2 3 0 2 27 1 1 31 
  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae
  +--+                 
     +-- Hyridellini **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
2 3 0 2 66 1 1 31 
  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iTtTbiriSoTroTroSiD
2 3 0 2 26 1 1 31 
  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae
  +--+                 
     +-- Hyridellini **

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
2 3 0 2 14 1 1 31 
  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae
  +--+                 
     +-- Hyridellini **

Chen et al., 2025 [Publ 4455], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 24 0 0 -- 
  +-- Hyridellini **
--+              
  +-- Hyriini **

Combosch et al., 2017 [Publ 3806], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 13 1 1 -- 
  +----- Velesunioninae
--+                    
  |  +-- Hyriini **       
  +--+                 
     +-- Hyridellini **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
2 2 0 2 89 1 1 -- 
  +----- Velesunioninae
--+                    
  |  +-- Castaliini **    
  +--+                 
     +-- Rhipidodontini **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 48 0 0 -- 
-- Echyridella_menziesii **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
1 1 0 1 95 0 0 -- 
-- Echyridella_menziesii **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Hyridella_depressa **

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 1 1 -- 
  +-- Echyridella **   
--+                 
  +-- Velesunioninae

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
1 1 0 1 73 0 0 -- 
-- Echyridella_menziesii **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
1 1 0 1 91 0 0 -- 
-- Echyridella_menziesii **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 45 0 0 -- 
-- Echyridella_menziesii **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 73 0 0 -- 
-- Echyridella_menziesii **

Guerra et al., 2017 [Publ 3772], fig. 2b-M

iTtTbiriSoTroTroSiD
1 1 0 1 7 0 0 -- 
-- Echyridella_menziesii **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 55 0 0 -- 
-- Echyridella_menziesii **

Guerra et al., 2017 [Publ 3772], fig. 2a-M

iTtTbiriSoTroTroSiD
1 1 0 1 7 0 0 -- 
-- Echyridella_menziesii **

Walker et al., 2006 [Publ 2666], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Hyridella **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
1 1 0 1 121 1 1 -- 
  +-- Hyridellini **   
--+                 
  +-- Velesunioninae

Chase et al., 2018 [Publ 3847], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Echyridella_menziesii **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
1 1 0 1 95 0 0 -- 
-- Echyridella_menziesii **

Chase et al., 2018 [Publ 3861], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Echyridella_menziesii **

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
1 1 0 1 24 0 0 -- 
-- Echyridella_menziesii **

Hoeh et al., 2002 [Publ 2673], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 8 0 0 -- 
-- Echyridella_menziesii **

Klunzinger et al., 2022 [Publ 4259], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 3 3 1 -- 
  +-- Hyridellini **   
--+                 
  +-- Velesunioninae

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 1 1 -- 
  +-- Echyridella **   
--+                 
  +-- Velesunioninae

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 41 0 0 -- 
-- Echyridella_menziesii **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
1 1 0 1 73 0 0 -- 
-- Echyridella_menziesii **

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 46 1 1 -- 
  +-- Echyridella **   
--+                 
  +-- Velesunioninae

Klunzinger et al., 2021 [Publ 4086], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 5 3 1 -- 
  +----- Westralunio
--+                 
  |  +-- Velesunio  
  +--+              
     +-- Hyridellini **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
1 1 0 1 91 0 0 -- 
-- Echyridella_menziesii **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Hyridella_australis **

Chase et al., 2018 [Publ 3847], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 38 0 0 -- 
-- Echyridella_menziesii **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
1 1 0 1 79 1 1 -- 
  +-- Hyridellini **   
--+                 
  +-- Velesunioninae

Walker et al., 2006 [Publ 2666], fig. 6-M

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Echyridella_menziesii **

Chase et al., 2018 [Publ 3861], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Echyridella_menziesii **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 68 0 0 -- 
-- Echyridella_menziesii **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 87 0 0 -- 
-- Echyridella_menziesii **

Walker et al., 2006 [Publ 2666], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Hyridella **

Breton et al., 2011 [Publ 3376], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 37 0 0 -- 
-- Echyridella_menziesii **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 85 0 0 -- 
-- Echyridella_menziesii **

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
1 1 0 1 24 0 0 -- 
-- Echyridella_menziesii **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Hyridella_depressa **

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 22 0 0 -- 
-- Echyridella_menziesii **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Hyridella_australis **

Walker et al., 2006 [Publ 2666], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Hyridella **

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
1 1 0 1 16 0 0 -- 
-- Echyridella_menziesii **

Hoeh et al., 2002 [Publ 2673], fig. 1-F

iTtTbiriSoTroTroSiD
1 1 0 1 8 0 0 -- 
-- Echyridella_menziesii **