Bivalvia | Unionida | Unionoidea | Unionidae | Parreysiinae

tribe LAMELLIDENTINI Modell, 1942: cladomics

4 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 21 (23 total)
number of subtaxa (rank: genus) = 4
analyzed in 56 cladograms in 35 publications
maximum number of terminals in an analysis = 17 (mean = 5.25, median = 2)
maximum number of subtaxa (genus) = 3 (mean = 1.64, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 34
maximum number of included terminals = 17, (mean = 8 median = 8)
maximum number of subtaxa (genus) = 3 (mean = 2.06, median = 2)
proportion of cladograms that support basic monophyly = 0.88

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 31
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 17 (mean = 8.26, median = 8)
maximum number of subtaxa (genus) = 3 (mean = 2.1, median = 2)
proportion of cladograms that support rigorous monophyly = 0.9

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Lamellidentini cladogram distances
PCoA Ordination

NJ Tree of Lamellidentini cladogram distances
NJ Tree

Cladograms

Bolotov et al., 2024 [Publ 4369], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cstiTtTbiriSoTroTroSiD
0.957 17 17 15 3 45 45 4 ** 
     +----- Lamellidens **   
  +--+                    
  |  |  +-- Balwantia **     
  |  +--+                 
  |     +-- Trapezidens **   
--+                       
  |     +-- Leoparreysiini
  |  +--+                 
  |  |  +-- Parreysiini   
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2024 [Publ 4369], fig. S1

cstiTtTbiriSoTroTroSiD
0.913 17 17 15 3 46 45 4 6.0 
  +-------------- Parreysia     
  |                             
  |        +----- Assamnaia     
--+  +-----+                    
  |  |     |  +-- Coelaturini   
  |  |     +--+                 
  +--+        +-- Indochinellini
     |                          
     |  +-------- Leoparreysiini
     +--+                       
        |  +----- Lamellidens **   
        +--+                    
           |  +-- Balwantia **     
           +--+                 
              +-- Trapezidens **

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cstiTtTbiriSoTroTroSiD
0.741 17 86 91 3 616 68 4 102.0 
                       +----------------- Lamellidens **                 
                       |                                              
  +--------------------+              +-- Balwantia **                   
  |                    |  +-----------+                               
  |                    |  |           +-- Trapezidens **                 
  |                    +--+                                           
  |                       |  +----------- Leoparreysiini              
  |                       +--+                                        
  |                          |  +-------- Pseudomulleria              
  |                          +--+                                     
  |                             |  +----- Indochinellini              
  |                             +--+                                  
  |                                |  +-- Parreysiini                 
--+                                +--+                               
  |                                   +-- Coelaturini                 
  |                                                                   
  |                                +----- Amuranodonta                
  |  +-----------------------------+                                  
  |  |                             |  +-- Anemina                     
  |  |                             +--+                               
  |  |                                +-- Buldowskia                  
  |  |                                                                
  |  |                                +-- Beringiana                  
  |  |                          +-----+                               
  +--+                          |     +-- Pletholophus                
     |  +-----------------------+                                     
     |  |                       |  +----- Mimo                        
     |  |                       +--+                                  
     |  |                          |  +-- Acudonta                    
     |  |                          +--+                               
     |  |                             +-- Sinanodonta                 
     |  |                                                             
     +--+  +----------------------------- Cristaria                   
        |  |                                                          
        |  |           +----------------- Lepidodesmini               
        |  |           |                                              
        |  |           |     +----------- Lanceolariini               
        |  |           |  +--+                                        
        |  |  +--------+  |  |  +-------- Pseudoacuticosta_retiaria__2
        |  |  |        |  |  +--+                                     
        +--+  |        |  |     |  +----- Acuticosta_chinensis__2     
           |  |        |  |     +--+                                  
           |  |        +--+        |  +-- Acuticosta_chinensis__1     
           |  |           |        +--+                               
           |  |           |           +-- Pseudoacuticosta_retiaria__1
           |  |           |                                           
           |  |           |  +----------- Globunio                    
           |  |           +--+                                        
           |  |              |  +-------- Aculamprotulina             
           +--+              +--+                                     
              |                 |  +----- Middendorffinaiina          
              |                 +--+                                  
              |                    |  +-- Unionina                    
              |                    +--+                               
              |                       +-- Oxynaiina                   
              |                                                       
              |                 +-------- Pseudodontoideus            
              |  +--------------+                                     
              |  |              |  +----- Lasmigona                   
              |  |              +--+                                  
              |  |                 |  +-- Pterosyna                   
              +--+                 +--+                               
                 |                    +-- Arcidens                    
                 |                                                    
                 |  +-------------------- Anodontina                  
                 |  |                                                 
                 +--+                 +-- Utterbackia                 
                    |  +--------------+                               
                    |  |              +-- Pyganodon                   
                    +--+                                              
                       |  +-------------- Utterbackiana               
                       |  |                                           
                       +--+  +----------- Simpsonaias                 
                          |  |                                        
                          |  |        +-- Alasminota                  
                          +--+  +-----+                               
                             |  |     +-- Alasmidonta                 
                             |  |                                     
                             +--+     +-- Prolasmidonta               
                                |  +--+                               
                                |  |  +-- Platynaias                  
                                +--+                                  
                                   |  +-- Anodontoides                
                                   +--+                               
                                      +-- Strophitus

Bolotov et al., 2017 [Publ 3761], fig. S1

cstiTtTbiriSoTroTroSiD
0.882 8 8 6 2 131 21 4 11.0 
           +-- Trapezidens **   
  +--------+                 
  |        +-- Lamellidens **   
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2017 [Publ 3761], fig. S2

cstiTtTbiriSoTroTroSiD
0.882 8 8 5 2 131 21 4 12.0 
     +----- Parreysiini   
  +--+                    
  |  |  +-- Trapezidens **   
  |  +--+                 
--+     +-- Lamellidens **   
  |                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2017 [Publ 3764], fig. S1

cstiTtTbiriSoTroTroSiD
0.647 11 38 28 2 114 27 4 46.0 
              +-------------- Trapezidens **             
              |                                       
     +--------+  +----------- Lamellidens_lamellatus **  
     |        |  |                                    
     |        +--+  +-------- Lamellidens_jenkinsianus **
     |           |  |                                 
     |           |  |  +----- Lamellidens_generosus **   
     |           +--+  |                              
     |              |  +----- Lamellidens_mainwaringi ** 
     |              |  |                              
     |              +--+  +-- Lamellidens_ferrugineus ** 
  +--+                 +--+                           
  |  |                 |  +-- Lamellidens_marginalis **  
  |  |                 |                              
  |  |                 |  +-- Lamellidens_savadiensis ** 
  |  |                 +--+                           
  |  |                    +-- Lamellidens_corrianus__1 **
  |  |                                                
  |  |  +-------------------- Parreysiini             
  |  |  |                                             
  |  +--+  +----------------- Leoparreysiini          
  |     |  |                                          
  |     +--+  +-------------- Coelaturini             
  |        |  |                                       
  |        |  |        +----- Indochinella            
  |        +--+  +-----+                              
  |           |  |     |  +-- Scabies                 
  |           |  |     +--+                           
  |           +--+        +-- Radiatula               
  |              |                                    
  |              |  +-------- Indonaia_shurtleffiana  
--+              |  |                                 
  |              +--+  +----- Indonaia_rugosa         
  |                 |  |                              
  |                 |  |  +-- Indonaia_andersoniana   
  |                 +--+--+                           
  |                    |  +-- Indonaia_caerulea       
  |                    |                              
  |                    |  +-- Indonaia_bonneaudii     
  |                    +--+                           
  |                       +-- Lamellidens_corrianus__2 **
  |                                                   
  |           +-------------- Pseudodontini           
  |           |                                       
  |           |  +----------- Rectidentini            
  |           |  |                                    
  |           |  |     +----- Unionini                
  |           |  |     |                              
  |           |  |     |  +-- Alasmidonta             
  |           |  |     |  |                           
  |           |  |     |  +-- Anodonta_anatina        
  +-----------+  |     |  |                           
              |  |     |  +-- Anodonta_cygnea         
              |  |     |  |                           
              |  |     |  +-- Brachyanodon            
              |  |  +--+  |                           
              |  |  |  |  +-- Lasmigona               
              |  |  |  |  |                           
              |  |  |  |  +-- Pseudanodonta           
              |  |  |  |  |                           
              |  |  |  |  +-- Pyganodon               
              +--+  |  |  |                           
                 |  |  |  +-- Simpsonaias             
                 |  |  |  |                           
                 |  |  |  +-- Strophitus              
                 |  |  +--+                           
                 |  |     +-- Sinanodonta_lauta       
                 |  |     |                           
                 |  |     +-- Cristaria_plicata       
                 |  |     |                           
                 |  |     +-- Cristaria_radiata       
                 |  |     |                           
                 |  |     +-- Pletholophus            
                 |  |     |                           
                 |  |     +-- Sinanodonta_lucida      
                 |  |     |                           
                 |  |     +-- Sinanodonta_jourdyi     
                 |  |     |                           
                 |  |     +-- Lanceolaria_gladiola    
                 +--+     |                           
                    |     +-- Lanceolaria_grayii      
                    |     |                           
                    |     +-- Lanceolaria_danzhouensis
                    |                                 
                    |     +-- Amblemini               
                    |     |                           
                    |     +-- Ortmanniana             
                    |     |                           
                    |     +-- Lampsilis               
                    |     |                           
                    |     +-- Cambarunio              
                    |     |                           
                    |  +--+-- Elliptio                
                    |  |  |                           
                    |  |  +-- Eurynia                 
                    |  |  |                           
                    |  |  +-- Pleurobema              
                    |  |  |                           
                    |  |  +-- Quadrula_quadrula       
                    |  |  |                           
                    |  |  +-- Quadrula_verrucosa      
                    |  |                              
                    |  |  +-- Chamberlainia           
                    +--+  |                           
                       |  +-- Sinohyriopsis           
                       |  |                           
                       |  +-- Lamprotula_caveata      
                       |  |                           
                       |  +-- Lamprotula_leaii        
                       |  |                           
                       |  +-- Potomida                
                       |  |                           
                       |  +-- Pronodularia            
                       |  |                           
                       +--+-- Gonidea                 
                          |                           
                          +-- Leguminaia_wheatleyi    
                          |                           
                          +-- Leguminaia              
                          |                           
                          +-- Microcondylaea          
                          |                           
                          +-- Sinosolenaia_carinata   
                          |                           
                          +-- Sinosolenaia_oleivora   
                          |                           
                          +-- Sinosolenaia_recognita

Bolotov et al., 2018 [Publ 3826], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 6 6 4 2 76 18 4 11.0 
           +-- Trapezidens **   
  +--------+                 
  |        +-- Lamellidens **   
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Pfeiffer et al., 2018 [Publ 3827], fig. 2

cstiTtTbiriSoTroTroSiD
0.950 2 2 0 2 26 25 4 -- 
           +-- Trapezidens **   
  +--------+                 
  |        +-- Lamellidens **   
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Huang et al., 2019 [Publ 3843], fig. 2

cstiTtTbiriSoTroTroSiD
0.941 11 11 10 2 190 23 4 13.0 
           +-- Trapezidens **   
  +--------+                 
  |        +-- Lamellidens **   
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Huang et al., 2019 [Publ 3843], fig. S3

cstiTtTbiriSoTroTroSiD
0.941 11 11 7 2 188 23 4 15.0 
           +-- Trapezidens **   
  +--------+                 
  |        +-- Lamellidens **   
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Huang et al., 2019 [Publ 3843], fig. S4

cstiTtTbiriSoTroTroSiD
0.941 11 11 6 2 190 23 4 10.0 
  +-------- Parreysiini   
  |                       
  |     +-- Trapezidens **   
--+-----+                 
  |     +-- Lamellidens **   
  |                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 21 21 4 -- 
           +-- Lamellidens **   
  +--------+                 
  |        +-- Trapezidens **   
--+                          
  |  +-------- Leoparreysiini
  +--+                       
     |  +----- Parreysiini   
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2020 [Publ 4100], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 8 8 6 2 109 33 4 11.0 
        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
  |                       
--+     +-- Parreysiini   
  |  +--+                 
  |  |  +-- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2020 [Publ 4100], fig. S1

cstiTtTbiriSoTroTroSiD
0.947 8 8 6 2 103 32 4 11.0 
        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
  |                       
--+     +-- Parreysiini   
  |  +--+                 
  |  |  +-- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2022 [Publ 4239], fig. 2

cstiTtTbiriSoTroTroSiD
0.950 14 14 12 2 44 40 4 13.0 
        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
  |                       
--+     +-- Parreysiini   
  |  +--+                 
  |  |  +-- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2022 [Publ 4239], fig. S1

cstiTtTbiriSoTroTroSiD
0.950 14 14 11 2 44 40 4 12.0 
           +-- Trapezidens **   
  +--------+                 
  |        +-- Lamellidens **   
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2022 [Publ 4239], fig. S2

cstiTtTbiriSoTroTroSiD
0.950 14 14 12 2 44 40 4 13.0 
  +----------- Parreysiini   
  |                          
--+        +-- Trapezidens **   
  |  +-----+                 
  |  |     +-- Lamellidens **   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2022 [Publ 4252], fig. 2

cstiTtTbiriSoTroTroSiD
0.950 16 16 12 2 45 41 4 11.0 
           +-- Trapezidens **   
  +--------+                 
  |        +-- Lamellidens **   
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2022 [Publ 4252], fig. S1

cstiTtTbiriSoTroTroSiD
0.950 16 16 14 2 45 41 4 9.0 
  +----------- Parreysiini   
  |                          
--+        +-- Trapezidens **   
  |  +-----+                 
  |  |     +-- Lamellidens **   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Chen et al., 2026 [Publ 4833], fig. 1

cstiTtTbiriSoTroTroSiD
0.789 2 25 0 2 25 22 4 38.0 
  +----------------------------------- Gibbosula                   
  |                                                                
--+  +-------------------------------- Lamellidens **                 
  |  |                                                             
  +--+  +----------------------------- Trapezidens **                 
     |  |                                                          
     +--+  +-------------------------- Parreysiini                 
        |  |                                                       
        +--+  +----------------------- Leoparreysiini              
           |  |                                                    
           +--+  +-------------------- Coelaturini                 
              |  |                                                 
              +--+                 +-- Indochinella                
                 |  +--------------+                               
                 |  |              +-- Indonaia                    
                 +--+                                              
                    |  +-------------- Radiatula                   
                    |  |                                           
                    +--+  +----------- Pseudodontini               
                       |  |                                        
                       +--+  +-------- Harmandia                   
                          |  |                                     
                          +--+     +-- Unionetta_xishuangbannaensis
                             |  +--+                               
                             |  |  +-- Unionetta_fabagina__2       
                             +--+                                  
                                |  +-- Unionetta_fabagina__1       
                                +--+                               
                                   +-- Scabies

Whelan et al., 2011 [Publ 3380], fig. 1-ML

cstiTtTbiriSoTroTroSiD
0.923 2 2 0 2 34 7 3 -- 
        +-- Lamellidens **   
  +-----+                 
  |     +-- Trapezidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Indochinellini
     +--+                 
        +-- Coelaturini

Whelan et al., 2011 [Publ 3380], fig. 1-MP

cstiTtTbiriSoTroTroSiD
0.923 2 2 0 2 34 7 3 -- 
        +-- Lamellidens **   
  +-----+                 
  |     +-- Trapezidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Indochinellini
     +--+                 
        +-- Coelaturini

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 38 6 3 -- 
        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Konopleva et al., 2017 [Publ 3807], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 5 5 3 2 26 6 3 12.0 
        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 55 8 3 -- 
        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 55 8 3 -- 
        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Wu et al., 2021 [Publ 4366], fig. 2_I

cstiTtTbiriSoTroTroSiD
1.000 4 4 2 2 87 9 3 13.0 
        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Wu et al., 2021 [Publ 4366], fig. 6

cstiTtTbiriSoTroTroSiD
0.917 4 4 2 2 87 9 3 13.0 
        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2022 [Publ 4250], fig. 8

cstiTtTbiriSoTroTroSiD
1.000 9 9 7 2 40 15 3 8.0 
        +-- Lamellidens **   
  +-----+                 
  |     +-- Trapezidens **   
--+                       
  |  +----- Parreysiini   
  +--+                    
     |  +-- Leoparreysiini
     +--+                 
        +-- Indochinellini

Konopleva et al., 2023 [Publ 4269], fig. S1_4

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 54 5 3 -- 
        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
--+                       
  |  +----- Parreysiini   
  +--+                    
     |  +-- Leoparreysiini
     +--+                 
        +-- Indochinellini

Konopleva et al., 2020 [Publ 4072], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 9 9 7 2 3 3 1 12.0 
  +----- Leoparreysiini
--+                    
  |  +-- Lamellidens **   
  +--+                 
     +-- Trapezidens **

Konopleva et al., 2017 [Publ 3807], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 5 5 3 2 0 0 0 12.0 
  +-- Trapezidens **
--+              
  +-- Lamellidens **

Bolotov et al., 2022 [Publ 4250], fig. S4

cstiTtTbiriSoTroTroSiD
1.000 9 9 6 2 0 0 0 11.0 
  +-- Lamellidens **
--+              
  +-- Trapezidens **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 80 3 3 -- 
  +-------- Trapezidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Zieritz et al., 2024 [Publ 4381], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 44 4 3 -- 
  +-------- Lamellidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Indochinellini
     +--+                 
        +-- Coelaturini

Choi et al., 2020 [Publ 4169], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 9 2 2 -- 
  +----- Indochinellini
--+                    
  |  +-- Lamellidens **   
  +--+                 
     +-- Leoparreysiini

Imsong & Murali, 2023 [Publ 4370], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 2 2 2 -- 
  +----- Lamellidens **   
--+                    
  |  +-- Indochinellini
  +--+                 
     +-- Parreysiini

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 1 1 -- 
  +-- Lamellidens **
--+              
  +-- Coelaturini

Araujo et al., 2017 [Publ 3801], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 13 1 1 -- 
  +-- Lamellidens **
--+              
  +-- Coelaturini

Dai et al., 2024 [Publ 4385], fig. 3-BI

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 1 1 -- 
  +-- Trapezidens **   
--+                 
  +-- Indochinellini

Dai et al., 2024 [Publ 4385], fig. 3-ML

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 1 1 -- 
  +-- Trapezidens **   
--+                 
  +-- Indochinellini

Dai et al., 2024 [Publ 4386], fig. 5-BI

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 21 2 1 -- 
  +-- Trapezidens **   
--+                 
  +-- Indochinellini

Dai et al., 2024 [Publ 4386], fig. 5-ML

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 21 2 1 -- 
  +-- Trapezidens **   
--+                 
  +-- Indochinellini

Dai et al., 2026 [Publ 4757], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 33 1 1 -- 
  +-- Indochinellini
--+                 
  +-- Trapezidens **

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 24 0 0 -- 
  +-------------------------------- Lamellidens **     
  |                                                 
  |                 +-------------- Oxynaiina       
  |                 |                               
  |  +--------------+  +----------- Anodonta_cygnea 
--+  |              |  |                            
  |  |              +--+  +-------- Unio_pictorum__2
  |  |                 |  |                         
  |  |                 +--+     +-- Unio_pictorum__1
  |  |                    |  +--+                   
  |  |                    |  |  +-- Unio_caffer     
  +--+                    +--+                      
     |                       |  +-- Anodonta        
     |                       +--+                   
     |                          +-- Alasmidontina   
     |                                              
     |  +-------------------------- Lamprotulini    
     |  |                                           
     +--+  +----------------------- Quadrulini      
        |  |                                        
        +--+  +-------------------- Gonideini       
           |  |                                     
           |  |              +----- Eurynia         
           +--+  +-----------+                      
              |  |           |  +-- Cambarunio      
              |  |           +--+                   
              |  |              +-- Ptychobranchus  
              +--+                                  
                 |              +-- Amblemini       
                 |  +-----------+                   
                 |  |           +-- Pleurobema      
                 +--+                               
                    |  +----------- Truncilla       
                    |  |                            
                    +--+        +-- Lampsilis       
                       |  +-----+                   
                       |  |     +-- Epioblasma      
                       +--+                         
                          |  +----- Ligumia         
                          +--+                      
                             |  +-- Obliquaria      
                             +--+                   
                                +-- Ortmanniana

Sonowal et al., 2021 [Publ 4221], fig. 3a

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 14 0 0 -- 
  +----------- Lamellidens **    
  |                           
  +----------- Lamprotulini   
  |                           
  |     +----- Neotrigonia    
--+     |                     
  |  +--+  +-- Hyriini        
  |  |  |  |                  
  |  |  +--+-- Hyridellini    
  |  |     |                  
  +--+     +-- Velesunio      
     |                        
     |  +----- Pleurobemini   
     |  |                     
     |  +----- Lampsilini     
     +--+                     
        |  +-- Anodontini     
        |  |                  
        |  +-- Unio_ravoisieri
        +--+                  
           +-- Unio_pictorum  
           |                  
           +-- Aculamprotulina

Sonowal et al., 2021 [Publ 4221], fig. 3b

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 14 0 0 -- 
  +-- Aculamprotulina
  |                  
  +-- Lamprotula     
  |                  
  +-- Lampsilini     
  |                  
  +-- Potomida       
  |                  
--+-- Unio_pictorum  
  |                  
  +-- Anodontini     
  |                  
  +-- Unio_ravoisieri
  |                  
  +-- Lamellidens **    
  |                  
  +-- Pleurobemini

Sonowal et al., 2021 [Publ 4221], fig. 4

cstiTtTbiriSoTroTroSiD
0.000 2 10 0 1 14 0 0 23.0 
                          +----- Hyriini                
  +-----------------------+                             
  |                       |  +-- Velesunio              
  |                       +--+                          
--+                          +-- Hyridellini            
  |                                                     
  |  +-------------------------- Lamellidens_mainwaringi **
  +--+                                                  
     |  +----------------------- Lamprotula             
     +--+                                               
        |  +-------------------- Potomida               
        +--+                                            
           |  +----------------- Lampsilini             
           +--+                                         
              |  +-------------- Pleurobemini           
              +--+                                      
                 |  +----------- Aculamprotulina        
                 +--+                                   
                    |  +-------- Unio_pictorum          
                    +--+                                
                       |  +----- Anodontini             
                       +--+                             
                          |  +-- Unio_ravoisieri        
                          +--+                          
                             +-- Lamellidens_corrianus **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 36 0 0 -- 
  +-------------- Lamellidens **  
--+                            
  |  +----------- Lepidodesmini
  +--+                         
     |  +-------- Unionini     
     +--+                      
        |  +----- Lanceolariini
        +--+                   
           |  +-- Acuticostini 
           +--+                
              +-- Anodontini

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 38 0 0 -- 
  +-------------- Lamellidens **    
  |                              
--+  +----------- Anodontini     
  |  |                           
  |  |        +-- Acuticostini   
  +--+  +-----+                  
     |  |     +-- Lanceolariini  
     |  |                        
     +--+     +-- Lepidodesmini  
        |  +--+                  
        |  |  +-- Aculamprotulina
        +--+                     
           |  +-- Unionina       
           +--+                  
              +-- Oxynaiina

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 94 0 0 -- 
  +-------------- Lamellidens **  
--+                            
  |  +----------- Lepidodesmini
  +--+                         
     |  +-------- Unionini     
     +--+                      
        |  +----- Anodontini   
        +--+                   
           |  +-- Acuticostini 
           +--+                
              +-- Lanceolariini

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 94 0 0 -- 
  +-------------- Lamellidens **  
--+                            
  |  +----------- Lepidodesmini
  +--+                         
     |  +-------- Unionini     
     +--+                      
        |  +----- Anodontini   
        +--+                   
           |  +-- Acuticostini 
           +--+                
              +-- Lanceolariini

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 90 0 0 -- 
  +-------------- Lamellidens **  
--+                            
  |  +----------- Lepidodesmini
  +--+                         
     |  +-------- Unionini     
     +--+                      
        |  +----- Anodontini   
        +--+                   
           |  +-- Acuticostini 
           +--+                
              +-- Lanceolariini

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 90 0 0 -- 
  +-------------- Lamellidens **  
--+                            
  |  +----------- Lepidodesmini
  +--+                         
     |  +-------- Unionini     
     +--+                      
        |  +----- Anodontini   
        +--+                   
           |  +-- Acuticostini 
           +--+                
              +-- Lanceolariini

Wu et al., 2024 [Publ 4388], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 52 0 0 -- 
  +----------- Lamellidens **  
--+                         
  |  +-------- Lepidodesmini
  +--+                      
     |  +----- Unionini     
     +--+                   
        |  +-- Lanceolariini
        +--+                
           +-- Anodontini

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 139 0 0 -- 
  +-------------- Lamellidens **  
--+                            
  |  +----------- Lepidodesmini
  +--+                         
     |  +-------- Unionini     
     +--+                      
        |  +----- Lanceolariini
        +--+                   
           |  +-- Acuticostini 
           +--+                
              +-- Anodontini