Bivalvia | Unionida | Unionoidea | Unionidae | Parreysiinae

tribe LAMELLIDENTINI Modell, 1942: cladomics

4 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 22 (24 total)
number of subtaxa (rank: genus) = 4
analyzed in 61 cladograms in 37 publications
maximum number of terminals in an analysis = 18 (mean = 6.02, median = 2)
maximum number of subtaxa (genus) = 3 (mean = 1.72, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 38
maximum number of included terminals = 18, (mean = 9.05 median = 8.5)
maximum number of subtaxa (genus) = 3 (mean = 2.16, median = 2)
proportion of cladograms that support basic monophyly = 0.89

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 35
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 18 (mean = 9.37, median = 9)
maximum number of subtaxa (genus) = 3 (mean = 2.2, median = 2)
proportion of cladograms that support rigorous monophyly = 0.91

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Lamellidentini cladogram distances
PCoA Ordination

NJ Tree of Lamellidentini cladogram distances
NJ Tree

Cladograms

Bolotov et al., 2024 [Publ 4369], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
0.957 17 17 15 3 45 45 4 **
Cladomics cladogram Lamellidens Balwantia Trapezidens LEOPARREYSIINI PARREYSIINI COELATURINI INDOCHINELLINI

Bolotov et al., 2024 [Publ 4369], fig. S1

cst iT tT bi riS oT roT roS iD
0.913 17 17 15 3 46 45 4 6.0
Cladomics cladogram Parreysia Assamnaia COELATURINI INDOCHINELLINI LEOPARREYSIINI Lamellidens Balwantia Trapezidens

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.741 17 86 91 3 616 68 4 102.0
Cladomics cladogram Lamellidens Balwantia Trapezidens LEOPARREYSIINI Pseudomulleria INDOCHINELLINI PARREYSIINI COELATURINI Amuranodonta Anemina Buldowskia Beringiana Pletholophus Mimo Acudonta Sinanodonta Cristaria LEPIDODESMINI LANCEOLARIINI Pseudoacuticosta retiaria 2 Acuticosta chinensis 2 Acuticosta chinensis 1 Pseudoacuticosta retiaria 1 Globunio ACULAMPROTULINA MIDDENDORFFINAIINA UNIONINA OXYNAIINA Pseudodontoideus Lasmigona Pterosyna Arcidens ANODONTINA Utterbackia Pyganodon Utterbackiana Simpsonaias Alasminota Alasmidonta Prolasmidonta Platynaias Anodontoides Strophitus

Konopleva et al., 2026 [Publ 4847], fig. 2

cst iT tT bi riS oT roT roS iD
0.957 18 18 16 3 48 48 4 5.0
Cladomics cladogram Lamellidens Balwantia Trapezidens LEOPARREYSIINI PARREYSIINI COELATURINI INDOCHINELLINI

Konopleva et al., 2026 [Publ 4847], fig. S1

cst iT tT bi riS oT roT roS iD
0.957 18 18 15 3 48 48 4 6.0
Cladomics cladogram Lamellidens Balwantia Trapezidens LEOPARREYSIINI PARREYSIINI COELATURINI INDOCHINELLINI

Konopleva et al., 2026 [Publ 4847], fig. S2

cst iT tT bi riS oT roT roS iD
0.913 18 18 16 3 49 48 4 5.0
Cladomics cladogram Parreysia Assamnaia COELATURINI INDOCHINELLINI LEOPARREYSIINI Lamellidens Balwantia Trapezidens

Konopleva et al., 2026 [Publ 4847], fig. S3

cst iT tT bi riS oT roT roS iD
0.913 18 18 12 3 49 48 4 6.0
Cladomics cladogram Parreysia Assamnaia COELATURINI INDOCHINELLINI LEOPARREYSIINI Lamellidens Balwantia Trapezidens

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
0.882 8 8 6 2 131 21 4 11.0
Cladomics cladogram Trapezidens Lamellidens PARREYSIINI LEOPARREYSIINI COELATURINI INDOCHINELLINI

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
0.882 8 8 5 2 131 21 4 12.0
Cladomics cladogram PARREYSIINI Trapezidens Lamellidens LEOPARREYSIINI COELATURINI INDOCHINELLINI

Bolotov et al., 2017 [Publ 3764], fig. S1

cst iT tT bi riS oT roT roS iD
0.647 11 38 28 2 114 27 4 46.0
Cladomics cladogram Trapezidens Lamellidens lamellatus Lamellidens jenkinsianus Lamellidens generosus Lamellidens mainwaringi Lamellidens ferrugineus Lamellidens marginalis Lamellidens savadiensis Lamellidens corrianus 1 PARREYSIINI LEOPARREYSIINI COELATURINI Indochinella Scabies Radiatula Indonaia shurtleffiana Indonaia rugosa Indonaia andersoniana Indonaia caerulea Indonaia bonneaudii Lamellidens corrianus 2 PSEUDODONTINI LIMNOSCAPHINI UNIONINI Alasmidonta Anodonta anatina Anodonta cygnea Brachyanodon Lasmigona Pseudanodonta Pyganodon Simpsonaias Strophitus Sinanodonta lauta Cristaria plicata Cristaria radiata Pletholophus Sinanodonta lucida Sinanodonta jourdyi Lanceolaria gladiola Lanceolaria grayii Lanceolaria danzhouensis AMBLEMINI Ortmanniana Lampsilis Cambarunio Elliptio Eurynia Pleurobema Quadrula quadrula Quadrula verrucosa Chamberlainia Sinohyriopsis Lamprotula caveata Lamprotula leaii Potomida Pronodularia Gonidea Leguminaia wheatleyi Leguminaia Microcondylaea Sinosolenaia carinata Sinosolenaia oleivora Sinosolenaia recognita

Bolotov et al., 2018 [Publ 3826], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 2 76 18 4 11.0
Cladomics cladogram Trapezidens Lamellidens PARREYSIINI LEOPARREYSIINI COELATURINI INDOCHINELLINI

Pfeiffer et al., 2018 [Publ 3827], fig. 2

cst iT tT bi riS oT roT roS iD
0.950 2 2 0 2 26 25 4 --
Cladomics cladogram Trapezidens Lamellidens PARREYSIINI LEOPARREYSIINI COELATURINI INDOCHINELLINI

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
0.941 11 11 10 2 190 23 4 13.0
Cladomics cladogram Trapezidens Lamellidens PARREYSIINI LEOPARREYSIINI COELATURINI INDOCHINELLINI

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
0.941 11 11 7 2 188 23 4 15.0
Cladomics cladogram Trapezidens Lamellidens PARREYSIINI LEOPARREYSIINI COELATURINI INDOCHINELLINI

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.941 11 11 6 2 190 23 4 10.0
Cladomics cladogram PARREYSIINI Trapezidens Lamellidens LEOPARREYSIINI COELATURINI INDOCHINELLINI

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 21 21 4 --
Cladomics cladogram Lamellidens Trapezidens LEOPARREYSIINI PARREYSIINI COELATURINI INDOCHINELLINI

Bolotov et al., 2020 [Publ 4100], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 2 109 33 4 11.0
Cladomics cladogram Trapezidens Lamellidens PARREYSIINI LEOPARREYSIINI COELATURINI INDOCHINELLINI

Bolotov et al., 2020 [Publ 4100], fig. S1

cst iT tT bi riS oT roT roS iD
0.947 8 8 6 2 103 32 4 11.0
Cladomics cladogram Trapezidens Lamellidens PARREYSIINI LEOPARREYSIINI COELATURINI INDOCHINELLINI

Bolotov et al., 2022 [Publ 4239], fig. 2

cst iT tT bi riS oT roT roS iD
0.950 14 14 12 2 44 40 4 13.0
Cladomics cladogram Trapezidens Lamellidens PARREYSIINI LEOPARREYSIINI COELATURINI INDOCHINELLINI

Bolotov et al., 2022 [Publ 4239], fig. S1

cst iT tT bi riS oT roT roS iD
0.950 14 14 11 2 44 40 4 12.0
Cladomics cladogram Trapezidens Lamellidens PARREYSIINI LEOPARREYSIINI COELATURINI INDOCHINELLINI

Bolotov et al., 2022 [Publ 4239], fig. S2

cst iT tT bi riS oT roT roS iD
0.950 14 14 12 2 44 40 4 13.0
Cladomics cladogram PARREYSIINI Trapezidens Lamellidens LEOPARREYSIINI COELATURINI INDOCHINELLINI

Bolotov et al., 2022 [Publ 4252], fig. 2

cst iT tT bi riS oT roT roS iD
0.950 16 16 12 2 45 41 4 11.0
Cladomics cladogram Trapezidens Lamellidens PARREYSIINI LEOPARREYSIINI COELATURINI INDOCHINELLINI

Bolotov et al., 2022 [Publ 4252], fig. S1

cst iT tT bi riS oT roT roS iD
0.950 16 16 14 2 45 41 4 9.0
Cladomics cladogram PARREYSIINI Trapezidens Lamellidens LEOPARREYSIINI COELATURINI INDOCHINELLINI

Chen et al., 2026 [Publ 4833], fig. 1

cst iT tT bi riS oT roT roS iD
0.800 2 25 0 2 25 22 4 38.0
Cladomics cladogram Gibbosula Lamellidens Trapezidens PARREYSIINI LEOPARREYSIINI COELATURINI Indochinella Indonaia Radiatula PSEUDODONTINI Harmandia Chenunionetta Unionetta fabagina 2 Unionetta fabagina 1 Scabies

Whelan et al., 2011 [Publ 3380], fig. 1-ML

cst iT tT bi riS oT roT roS iD
0.923 2 2 0 2 34 7 3 --
Cladomics cladogram Lamellidens Trapezidens LEOPARREYSIINI INDOCHINELLINI COELATURINI

Whelan et al., 2011 [Publ 3380], fig. 1-MP

cst iT tT bi riS oT roT roS iD
0.923 2 2 0 2 34 7 3 --
Cladomics cladogram Lamellidens Trapezidens LEOPARREYSIINI INDOCHINELLINI COELATURINI

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 38 6 3 --
Cladomics cladogram Trapezidens Lamellidens LEOPARREYSIINI COELATURINI INDOCHINELLINI

Konopleva et al., 2017 [Publ 3807], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 2 26 6 3 12.0
Cladomics cladogram Trapezidens Lamellidens LEOPARREYSIINI COELATURINI INDOCHINELLINI

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 55 8 3 --
Cladomics cladogram Trapezidens Lamellidens LEOPARREYSIINI COELATURINI INDOCHINELLINI

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 55 8 3 --
Cladomics cladogram Trapezidens Lamellidens LEOPARREYSIINI COELATURINI INDOCHINELLINI

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 2 87 9 3 13.0
Cladomics cladogram Trapezidens Lamellidens LEOPARREYSIINI COELATURINI INDOCHINELLINI

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
0.917 4 4 2 2 87 9 3 13.0
Cladomics cladogram Trapezidens Lamellidens LEOPARREYSIINI COELATURINI INDOCHINELLINI

Bolotov et al., 2022 [Publ 4250], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 9 9 7 2 40 15 3 8.0
Cladomics cladogram Lamellidens Trapezidens PARREYSIINI LEOPARREYSIINI INDOCHINELLINI

Konopleva et al., 2023 [Publ 4269], fig. S1_4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 54 5 3 --
Cladomics cladogram Trapezidens Lamellidens PARREYSIINI LEOPARREYSIINI INDOCHINELLINI

Konopleva et al., 2020 [Publ 4072], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 9 9 7 2 3 3 1 12.0
Cladomics cladogram LEOPARREYSIINI Lamellidens Trapezidens

Konopleva et al., 2017 [Publ 3807], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 2 0 0 0 12.0
Cladomics cladogram Trapezidens Lamellidens

Bolotov et al., 2022 [Publ 4250], fig. S4

cst iT tT bi riS oT roT roS iD
1.000 9 9 6 2 0 0 0 11.0
Cladomics cladogram Lamellidens Trapezidens

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 80 3 3 --
Cladomics cladogram Trapezidens LEOPARREYSIINI COELATURINI INDOCHINELLINI

Zieritz et al., 2024 [Publ 4381], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 44 4 3 --
Cladomics cladogram Lamellidens LEOPARREYSIINI INDOCHINELLINI COELATURINI

Choi et al., 2020 [Publ 4169], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 2 2 --
Cladomics cladogram INDOCHINELLINI Lamellidens LEOPARREYSIINI

Imsong & Murali, 2023 [Publ 4370], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 2 2 2 --
Cladomics cladogram Lamellidens INDOCHINELLINI PARREYSIINI

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 1 1 --
Cladomics cladogram Lamellidens COELATURINI

Araujo et al., 2017 [Publ 3801], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 1 1 --
Cladomics cladogram Lamellidens COELATURINI

Dai et al., 2024 [Publ 4385], fig. 3-BI

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 1 1 --
Cladomics cladogram Trapezidens INDOCHINELLINI

Dai et al., 2024 [Publ 4385], fig. 3-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 1 1 --
Cladomics cladogram Trapezidens INDOCHINELLINI

Dai et al., 2024 [Publ 4386], fig. 5-BI

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 2 1 --
Cladomics cladogram Trapezidens INDOCHINELLINI

Dai et al., 2024 [Publ 4386], fig. 5-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 2 1 --
Cladomics cladogram Trapezidens INDOCHINELLINI

Dai et al., 2026 [Publ 4757], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 1 1 --
Cladomics cladogram INDOCHINELLINI Trapezidens

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 24 0 0 --
Cladomics cladogram Lamellidens OXYNAIINA Anodonta cygnea Unio pictorum 2 Unio pictorum 1 Unio caffer Anodonta ALASMIDONTINA LAMPROTULINI QUADRULINI GONIDEINI Eurynia Cambarunio Ptychobranchus AMBLEMINI Pleurobema Truncilla Lampsilis Epioblasma Ligumia Obliquaria Ortmanniana

Sonowal et al., 2021 [Publ 4221], fig. 3a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram Lamellidens LAMPROTULINI Neotrigonia HYRIINI HYRIDELLINI Velesunio PLEUROBEMINI LAMPSILINI ANODONTINI Unio ravoisieri Unio pictorum ACULAMPROTULINA

Sonowal et al., 2021 [Publ 4221], fig. 3b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram ACULAMPROTULINA Lamprotula LAMPSILINI Potomida Unio pictorum ANODONTINI Unio ravoisieri Lamellidens PLEUROBEMINI

Sonowal et al., 2021 [Publ 4221], fig. 4

cst iT tT bi riS oT roT roS iD
0.000 2 10 0 1 14 0 0 25.0
Cladomics cladogram HYRIINI Velesunio HYRIDELLINI Lamellidens phenchooganjensis Lamprotula Potomida LAMPSILINI PLEUROBEMINI ACULAMPROTULINA Unio pictorum ANODONTINI Unio ravoisieri Lamellidens corrianus

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 36 0 0 --
Cladomics cladogram Lamellidens LEPIDODESMINI UNIONINI LANCEOLARIINI ACUTICOSTINI ANODONTINI

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 38 0 0 --
Cladomics cladogram Lamellidens ANODONTINI ACUTICOSTINI LANCEOLARIINI LEPIDODESMINI ACULAMPROTULINA UNIONINA OXYNAIINA

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 0 0 --
Cladomics cladogram Lamellidens LEPIDODESMINI UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 0 0 --
Cladomics cladogram Lamellidens LEPIDODESMINI UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 0 0 --
Cladomics cladogram Lamellidens LEPIDODESMINI UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 0 0 --
Cladomics cladogram Lamellidens LEPIDODESMINI UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Wu et al., 2024 [Publ 4388], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 0 0 --
Cladomics cladogram Lamellidens LEPIDODESMINI UNIONINI LANCEOLARIINI ANODONTINI

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 139 0 0 --
Cladomics cladogram Lamellidens LEPIDODESMINI UNIONINI LANCEOLARIINI ACUTICOSTINI ANODONTINI

Wu et al., 2026 [Publ 4852], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 0 0 --
Cladomics cladogram Lamellidens UNIONINI GONIDEINI LAMPROTULINI