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Data last updated
Thu 18 Dec 2025

Bivalvia | Unionoida | Unionoidea | Unionidae | Parreysiinae

tribe LAMELLIDENTINI Modell, 1942: cladomics

4 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 21 (23 total)
number of subtaxa (rank: genus) = 4
analyzed in 52 cladograms in 32 publications
maximum number of terminals in an analysis = 17 (mean = 5.25, median = 2)
maximum number of subtaxa (genus) = 3 (mean = 1.63, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 32
maximum number of included terminals = 17, (mean = 7.91 median = 8)
maximum number of subtaxa (genus) = 3 (mean = 2.03, median = 2)
proportion of cladograms that support basic monophyly = 0.94

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 29
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 17 (mean = 8.17, median = 8)
maximum number of subtaxa (genus) = 3 (mean = 2.07, median = 2)
proportion of cladograms that support rigorous monophyly = 0.97

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Lamellidentini cladogram distances
PCoA Ordination

NJ Tree of Lamellidentini cladogram distances
NJ Tree

Cladograms

Bolotov et al., 2024 [Publ 4369], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
17 17 15 3 45 45 4 0 
     +----- Lamellidens **   
  +--+                    
  |  |  +-- Balwantia **     
  |  +--+                 
  |     +-- Trapezidens **   
--+                       
  |     +-- Leoparreysiini
  |  +--+                 
  |  |  +-- Parreysiini   
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2024 [Publ 4369], fig. S1

iTtTbiriSoTroTroSiD
17 17 15 3 46 45 4 6 
  +-------------- Parreysia     
  |                             
  |        +----- Assamnaia     
--+  +-----+                    
  |  |     |  +-- Coelaturini   
  |  |     +--+                 
  +--+        +-- Indochinellini
     |                          
     |  +-------- Leoparreysiini
     +--+                       
        |  +----- Lamellidens **   
        +--+                    
           |  +-- Balwantia **     
           +--+                 
              +-- Trapezidens **

Bolotov et al., 2022 [Publ 4252], fig. 2

iTtTbiriSoTroTroSiD
16 16 12 2 45 41 4 11 
           +-- Trapezidens **   
  +--------+                 
  |        +-- Lamellidens **   
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2022 [Publ 4252], fig. S1

iTtTbiriSoTroTroSiD
16 16 14 2 45 41 4 9 
  +----------- Parreysiini   
  |                          
--+        +-- Trapezidens **   
  |  +-----+                 
  |  |     +-- Lamellidens **   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2022 [Publ 4239], fig. S2

iTtTbiriSoTroTroSiD
14 14 12 2 44 40 4 13 
  +----------- Parreysiini   
  |                          
--+        +-- Trapezidens **   
  |  +-----+                 
  |  |     +-- Lamellidens **   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2022 [Publ 4239], fig. 2

iTtTbiriSoTroTroSiD
14 14 12 2 44 40 4 13 
        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
  |                       
--+     +-- Parreysiini   
  |  +--+                 
  |  |  +-- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2022 [Publ 4239], fig. S1

iTtTbiriSoTroTroSiD
14 14 11 2 44 40 4 12 
           +-- Trapezidens **   
  +--------+                 
  |        +-- Lamellidens **   
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
11 11 7 2 188 23 4 15 
           +-- Trapezidens **   
  +--------+                 
  |        +-- Lamellidens **   
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
11 11 6 2 190 23 4 10 
  +-------- Parreysiini   
  |                       
  |     +-- Trapezidens **   
--+-----+                 
  |     +-- Lamellidens **   
  |                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
11 11 10 2 190 23 4 13 
           +-- Trapezidens **   
  +--------+                 
  |        +-- Lamellidens **   
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
11 38 28 2 114 27 4 46 
           +-------------- Trapezidens **             
           |                                       
  +--------+  +----------- Lamellidens_lamellatus **  
  |        |  |                                    
  |        +--+  +-------- Lamellidens_jenkinsianus **
  |           |  |                                 
  |           |  |  +----- Lamellidens_generosus **   
  |           +--+  |                              
  |              |  +----- Lamellidens_mainwaringi ** 
  |              |  |                              
  |              +--+  +-- Lamellidens_ferrugineus ** 
--+                 +--+                           
  |                 |  +-- Lamellidens_marginalis **  
  |                 |                              
  |                 |  +-- Lamellidens_savadiensis ** 
  |                 +--+                           
  |                    +-- Lamellidens_corrianus__1 **
  |                                                
  |  +-------------------- Parreysiini             
  |  |                                             
  +--+  +----------------- Leoparreysiini          
     |  |                                          
     +--+  +-------------- Coelaturini             
        |  |                                       
        |  |        +----- Indochinella            
        +--+  +-----+                              
           |  |     |  +-- Scabies                 
           |  |     +--+                           
           +--+        +-- Radiatula               
              |                                    
              |  +-------- Indonaia_shurtleffiana  
              |  |                                 
              +--+  +----- Indonaia_rugosa         
                 |  |                              
                 |  |  +-- Indonaia_andersoniana   
                 +--+--+                           
                    |  +-- Indonaia_caerulea       
                    |                              
                    |  +-- Indonaia_bonneaudii     
                    +--+                           
                       +-- Lamellidens_corrianus__2 **

Bolotov et al., 2022 [Publ 4250], fig. 8

iTtTbiriSoTroTroSiD
9 9 7 2 40 15 3 8 
        +-- Lamellidens **   
  +-----+                 
  |     +-- Trapezidens **   
--+                       
  |  +----- Parreysiini   
  +--+                    
     |  +-- Leoparreysiini
     +--+                 
        +-- Indochinellini

Bolotov et al., 2022 [Publ 4250], fig. S4

iTtTbiriSoTroTroSiD
9 9 6 2 0 0 0 11 
  +-- Lamellidens **
--+              
  +-- Trapezidens **

Konopleva et al., 2020 [Publ 4072], fig. 2

iTtTbiriSoTroTroSiD
9 9 7 2 3 3 1 12 
  +----- Leoparreysiini
--+                    
  |  +-- Lamellidens **   
  +--+                 
     +-- Trapezidens **

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
8 8 6 2 110 33 4 11 
        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
  |                       
--+     +-- Parreysiini   
  |  +--+                 
  |  |  +-- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
8 8 5 2 131 21 4 12 
     +----- Parreysiini   
  +--+                    
  |  |  +-- Trapezidens **   
  |  +--+                 
--+     +-- Lamellidens **   
  |                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
8 8 6 2 104 32 4 11 
        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
  |                       
--+     +-- Parreysiini   
  |  +--+                 
  |  |  +-- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
8 8 6 2 131 21 4 11 
           +-- Trapezidens **   
  +--------+                 
  |        +-- Lamellidens **   
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
6 6 4 2 77 18 4 11 
           +-- Trapezidens **   
  +--------+                 
  |        +-- Lamellidens **   
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
5 5 3 2 26 6 3 12 
        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Konopleva et al., 2017 [Publ 3807], fig. 5

iTtTbiriSoTroTroSiD
5 5 3 2 0 0 0 12 
  +-- Trapezidens **
--+              
  +-- Lamellidens **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
4 4 2 2 87 9 3 13 
        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
4 4 2 2 87 9 3 13 
        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 2 55 8 3 -- 
        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
2 2 0 2 54 5 3 -- 
        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
--+                       
  |  +----- Parreysiini   
  +--+                    
     |  +-- Leoparreysiini
     +--+                 
        +-- Indochinellini

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
2 10 0 1 14 0 0 23 
  +-------------------------- Lamellidens_mainwaringi **
--+                                                  
  |  +----------------------- Lamprotula             
  +--+                                               
     |  +-------------------- Potomida               
     +--+                                            
        |  +----------------- Lampsilini             
        +--+                                         
           |  +-------------- Pleurobemini           
           +--+                                      
              |  +----------- Aculamprotula          
              +--+                                   
                 |  +-------- Unio_pictorum          
                 +--+                                
                    |  +----- Anodontini             
                    +--+                             
                       |  +-- Unio_ravoisieri        
                       +--+                          
                          +-- Lamellidens_corrianus **

Pfeiffer et al., 2018 [Publ 3827], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 26 25 4 -- 
           +-- Trapezidens **   
  +--------+                 
  |        +-- Lamellidens **   
--+                          
  |  +-------- Parreysiini   
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
2 2 0 2 55 8 3 -- 
        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 38 6 3 -- 
        +-- Trapezidens **   
  +-----+                 
  |     +-- Lamellidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
2 2 0 2 34 7 3 -- 
        +-- Lamellidens **   
  +-----+                 
  |     +-- Trapezidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Indochinellini
     +--+                 
        +-- Coelaturini

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
2 2 0 2 34 7 3 -- 
        +-- Lamellidens **   
  +-----+                 
  |     +-- Trapezidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Indochinellini
     +--+                 
        +-- Coelaturini

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 21 21 4 -- 
           +-- Lamellidens **   
  +--------+                 
  |        +-- Trapezidens **   
--+                          
  |  +-------- Leoparreysiini
  +--+                       
     |  +----- Parreysiini   
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iTtTbiriSoTroTroSiD
1 1 0 1 24 0 0 -- 
-- Lamellidens_corrianus **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
1 1 0 1 91 0 0 -- 
-- Lamellidens_jenkinsianus **

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 45 4 3 -- 
  +-------- Lamellidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Indochinellini
     +--+                 
        +-- Coelaturini

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
1 1 0 1 95 0 0 -- 
-- Lamellidens_jenkinsianus **

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Lamellidens_mainwaringi **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 1 1 -- 
  +-- Lamellidens **
--+              
  +-- Coelaturini

Dai et al., 2024 [Publ 4385], fig. 3-BI

iTtTbiriSoTroTroSiD
1 1 0 1 30 1 1 -- 
  +-- Trapezidens **   
--+                 
  +-- Indochinellini

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
1 1 0 1 79 3 3 -- 
  +-------- Trapezidens **   
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Lamellidens_corrianus **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 53 0 0 -- 
-- Lamellidens_marginalis **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 36 0 0 -- 
-- Lamellidens_jenkinsianus **

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 13 1 1 -- 
  +-- Lamellidens **
--+              
  +-- Coelaturini

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
1 1 0 1 95 0 0 -- 
-- Lamellidens_jenkinsianus **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
1 1 0 1 38 0 0 -- 
-- Lamellidens_jenkinsianus **

Dai et al., 2024 [Publ 4385], fig. 3-ML

iTtTbiriSoTroTroSiD
1 1 0 1 30 1 1 -- 
  +-- Trapezidens **   
--+                 
  +-- Indochinellini

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
1 1 0 1 20 2 1 -- 
  +-- Trapezidens **   
--+                 
  +-- Indochinellini

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
1 1 0 1 20 2 1 -- 
  +-- Trapezidens **   
--+                 
  +-- Indochinellini

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
1 1 0 1 91 0 0 -- 
-- Lamellidens_jenkinsianus **

Choi et al., 2020 [Publ 4169], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 9 2 2 -- 
  +----- Indochinellini
--+                    
  |  +-- Lamellidens **   
  +--+                 
     +-- Leoparreysiini

Imsong & Murali, 2023 [Publ 4370], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 2 2 2 -- 
  +----- Lamellidens **   
--+                    
  |  +-- Indochinellini
  +--+                 
     +-- Parreysiini
 
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