Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae

tribe LAMPROTULINI Modell, 1942: cladomics

4 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 22 (41 total)
number of subtaxa (rank: genus) = 4
analyzed in 202 cladograms in 123 publications
maximum number of terminals in an analysis = 5 (mean = 2.44, median = 2)
maximum number of subtaxa (genus) = 3 (mean = 1.96, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 133
maximum number of included terminals = 5, (mean = 3.19 median = 3)
maximum number of subtaxa (genus) = 3 (mean = 2.45, median = 3)
proportion of cladograms that support basic monophyly = 0.89

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 109
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 5 (mean = 3.39, median = 3)
maximum number of subtaxa (genus) = 3 (mean = 2.67, median = 3)
proportion of cladograms that support rigorous monophyly = 0.91

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Lamprotulini cladogram distances
PCoA Ordination

NJ Tree of Lamprotulini cladogram distances
NJ Tree

Cladograms

Chen et al., 2025 [Publ 4663], fig. 1

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
4 4 2 3 45 40 5 0 
  +-------------- Rectidentini   
  |                              
--+  +----------- Chamberlainiini
  |  |                           
  |  |     +----- Potomida **       
  +--+  +--+                     
     |  |  |  +-- Pronodularia **   
     |  |  +--+                  
     +--+     +-- Lamprotula **     
        |                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini

Zieritz et al., 2024 [Publ 4381], fig. 2

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
2 42 0 2 44 33 6 45 
     +----------- Rectidentini   
     |                           
  +--+        +-- Schepmaniini   
  |  |  +-----+                  
  |  |  |     +-- Pseudodontini  
  |  +--+                        
  |     |  +----- Caudiculatus   
  |     +--+                     
  |        |  +-- Potomida **       
  |        +--+                  
--+           +-- Gonideini      
  |                              
  |     +-------- Lamellidentini 
  |  +--+                        
  |  |  |  +----- Leoparreysiini 
  |  |  +--+                     
  |  |     |  +-- Indochinellini 
  |  |     +--+                  
  +--+        +-- Coelaturini    
     |                           
     |        +-- Anodontini     
     |     +--+                  
     |     |  +-- Unionini       
     +-----+                     
           |  +-- Chamberlainiini
           +--+                  
              +-- Discomya **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
5 5 3 3 91 15 4 2 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **

Froufe et al., 2016 [Publ 3727], fig. 3_4

iTtTbiriSoTroTroSiD
5 7 3 3 2 2 1 5 
  +----- Potomida **      
  |                    
--+  +-- Pronodularia **  
  |  |                 
  |  +-- Lamprotula **    
  +--+                 
     +-- Leguminaia    
     |                 
     +-- Microcondylaea

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
5 5 0 3 107 67 4 3 
  +-------------- Rectidentini      
  |                                 
--+  +----------- Pseudodontini     
  |  |                              
  +--+  +-------- Gonideini         
     |  |                           
     +--+  +----- Chamberlainiini   
        |  |                        
        |  |  +-- Lamprotula_leaii **  
        +--+  |                     
           |  +-- Lamprotula **        
           |  |                     
           +--+-- Lamprotula_caveata **
              |                     
              +-- Potomida **          
              |                     
              +-- Pronodularia **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
5 5 3 3 87 16 4 2 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
5 5 3 3 87 16 4 2 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
5 5 3 3 194 50 4 4 
  +-------- Gonideini   
--+                     
  |  +----- Lamprotula **  
  +--+                  
     |  +-- Potomida **    
     +--+               
        +-- Pronodularia **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
5 5 3 3 48 17 4 2 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
5 5 3 3 196 51 4 4 
  +----------- Chamberlainiini
  |                           
--+        +-- Gonideini      
  |  +-----+                  
  |  |     +-- Pseudodontini  
  +--+                        
     |  +----- Lamprotula **     
     +--+                     
        |  +-- Potomida **       
        +--+                  
           +-- Pronodularia **

Matos et al., 2024 [Publ 4383], fig. 3

iTtTbiriSoTroTroSiD
5 5 3 3 15 13 4 1 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **

Zhang et al., 2024 [Publ 4397], fig. 5

iTtTbiriSoTroTroSiD
5 5 3 3 22 20 4 2 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
5 5 3 3 91 15 4 2 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
5 5 3 3 196 51 4 4 
  +----- Lamprotula **  
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
5 5 4 3 117 28 4 6 
           +-- Pseudodontini  
  +--------+                  
  |        +-- Gonideini      
--+                           
  |  +-------- Chamberlainiini
  +--+                        
     |  +----- Pronodularia **   
     +--+                     
        |  +-- Potomida **       
        +--+                  
           +-- Lamprotula **

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
5 16 0 3 113 67 4 15 
  +-------- Rectidentini             
  |                                  
  |  +----- Pseudodontini            
  |  |                               
  |  |  +-- Sinosolenaia_iridinea    
--+  |  |                            
  |  |  +-- Sinosolenaia_recognita__1
  |  |  |                            
  |  |  +-- Sinosolenaia_carinata    
  |  |  |                            
  |  |  +-- Leguminaia               
  +--+  |                            
     |  +-- Microcondylaea           
     |  |                            
     |  +-- Gonidea                  
     |  |                            
     |  +-- Ptychorhynchus           
     |  |                            
     |  +-- Parvasolenaia            
     |  |                            
     +--+-- Lamprotula_leaii **         
        |                            
        +-- Lamprotula **               
        |                            
        +-- Lamprotula_caveata **       
        |                            
        +-- Potomida **                 
        |                            
        +-- Pronodularia **             
        |                            
        +-- Sinohyriopsis            
        |                            
        +-- Chamberlainia            
        |                            
        +-- Sinosolenaia_recognita__2
        |                            
        +-- Sinosolenaia_oleivora

Matos et al., 2024 [Publ 4384], fig. 3

iTtTbiriSoTroTroSiD
5 5 3 3 15 13 4 2 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
5 5 4 3 26 6 4 6 
  +-------------- Chamberlainiini
  |                              
--+           +-- Gonideini      
  |  +--------+                  
  |  |        +-- Pseudodontini  
  +--+                           
     |  +-------- Rectidentini   
     +--+                        
        |  +----- Pronodularia **   
        +--+                     
           |  +-- Potomida **       
           +--+                  
              +-- Lamprotula **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
4 13 0 3 121 47 4 12 
  +-- Chamberlainia         
  |                         
  +-- Sinohyriopsis         
  |                         
  +-- Lamprotula_caveata **    
  |                         
  +-- Lamprotula_leaii **      
  |                         
  +-- Potomida **              
  |                         
  +-- Pronodularia **          
  |                         
--+-- Gonidea               
  |                         
  +-- Leguminaia_wheatleyi  
  |                         
  +-- Leguminaia            
  |                         
  +-- Microcondylaea        
  |                         
  +-- Sinosolenaia_carinata 
  |                         
  +-- Sinosolenaia_oleivora 
  |                         
  +-- Sinosolenaia_recognita

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 3 40 9 3 1 
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 3 84 16 4 1 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
4 4 2 3 49 15 4 1 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **

Wu et al., 2021 [Publ 4215], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 3 20 15 3 3 
  +-------------------- Chamberlainiini
  |                                    
  |  +----------------- Pseudodontini  
--+  |                                 
  |  |        +-------- Sinosolenaia   
  |  |  +-----+                        
  +--+  |     |  +----- Lamprotula **     
     |  |     +--+                     
     |  |        |  +-- Potomida **       
     |  |        +--+                  
     +--+           +-- Pronodularia **   
        |                              
        |           +-- Ptychorhynchus 
        |  +--------+                  
        |  |        +-- Parvasolenaia  
        +--+                           
           |  +-------- Inversidens    
           +--+                        
              |  +----- Gonidea        
              +--+                     
                 |  +-- Microcondylaea 
                 +--+                  
                    +-- Leguminaia

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
4 4 2 3 87 25 4 3 
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Pronodularia **   
     +--+                     
        |  +-- Potomida **       
        +--+                  
           +-- Lamprotula **

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
4 4 2 3 44 39 5 0 
  +-------------- Rectidentini   
  |                              
--+  +----------- Chamberlainiini
  |  |                           
  |  |     +----- Potomida **       
  +--+  +--+                     
     |  |  |  +-- Pronodularia **   
     |  |  +--+                  
     +--+     +-- Lamprotula **     
        |                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 3 82 14 4 1 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **

Lopes-Lima et al., 2020 [Publ 4080], fig. 16

iTtTbiriSoTroTroSiD
4 4 3 2 4 2 1 3 
  +-------- Chamberlainia
--+                      
  |  +----- Sinohyriopsis
  +--+                   
     |  +-- Pronodularia ** 
     +--+                
        +-- Lamprotula **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
4 4 2 3 87 25 4 1 
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 3 70 10 3 1 
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
4 4 2 3 38 4 2 1 
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 3 52 6 2 1 
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
4 4 2 3 52 9 2 0 
  +----------- Gonideini      
--+                           
  |  +-------- Chamberlainiini
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
4 4 2 3 38 6 3 1 
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 3 43 38 5 2 
  +-------------- Rectidentini   
  |                              
--+  +----------- Chamberlainiini
  |  |                           
  |  |     +----- Lamprotula **     
  +--+  +--+                     
     |  |  |  +-- Pronodularia **   
     |  |  +--+                  
     +--+     +-- Potomida **       
        |                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini

Wu et al., 2020 [Publ 4147], fig. 4

iTtTbiriSoTroTroSiD
4 4 2 2 10 1 1 3 
  +----- Gonideini 
--+                
  |  +-- Potomida **  
  +--+             
     +-- Lamprotula **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
4 4 2 3 50 8 3 3 
  +-------------- Chamberlainiini
--+                              
  |  +----------- Pseudodontini  
  +--+                           
     |  +-------- Gonideini      
     +--+                        
        |  +----- Lamprotula **     
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Potomida **

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
4 4 2 3 41 39 4 0 
  +----------------------- Pseudodontini  
  |                                       
  |  +-------------------- Rectidentini   
--+  |                                    
  |  |        +----------- Inversidens    
  |  |  +-----+                           
  +--+  |     |  +-------- Chamberlainiini
     |  |     +--+                        
     |  |        |  +----- Potomida **       
     |  |        +--+                     
     |  |           |  +-- Pronodularia **   
     +--+           +--+                  
        |              +-- Lamprotula **     
        |                                 
        |  +-------------- Sinosolenaia   
        |  |                              
        |  |        +----- Gonidea        
        +--+  +-----+                     
           |  |     |  +-- Leguminaia     
           |  |     +--+                  
           |  |        +-- Microcondylaea 
           +--+                           
              |        +-- Koreosolenaia  
              |  +-----+                  
              |  |     +-- Parvasolenaia  
              +--+                        
                 |  +----- Postolata      
                 +--+                     
                    |  +-- Ptychorhynchus 
                    +--+                  
                       +-- Obovalis

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
4 4 2 3 44 13 4 1 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
4 4 2 3 135 45 4 3 
  +-------------- Pseudodontini  
--+                              
  |  +----------- Gonideini      
  +--+                           
     |  +-------- Chamberlainiini
     +--+                        
        |  +----- Lamprotula **     
        +--+                     
           |  +-- Potomida **       
           +--+                  
              +-- Pronodularia **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
4 4 2 3 70 10 3 1 
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
4 4 2 3 34 5 2 1 
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
4 4 2 3 76 24 4 3 
        +-------- Chamberlainiini
  +-----+                        
  |     |  +----- Lamprotula **     
  |     +--+                     
  |        |  +-- Potomida **       
  |        +--+                  
--+           +-- Pronodularia **   
  |                              
  |           +-- Parvasolenaia  
  |  +--------+                  
  |  |        +-- Pseudodontini  
  +--+                           
     |  +-------- Sinosolenaia   
     +--+                        
        |  +----- Gonidea        
        +--+                     
           |  +-- Microcondylaea 
           +--+                  
              +-- Leguminaia

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
4 4 3 2 26 4 2 3 
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Pronodularia **   
     +--+                  
        +-- Lamprotula **

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
4 4 2 3 41 39 4 0 
  +----------------------- Pseudodontini  
  |                                       
  |  +-------------------- Rectidentini   
--+  |                                    
  |  |  +----------------- Inversidens    
  |  |  |                                 
  +--+  |        +-------- Chamberlainiini
     |  +--------+                        
     |  |        |  +----- Potomida **       
     |  |        +--+                     
     +--+           |  +-- Pronodularia **   
        |           +--+                  
        |              +-- Lamprotula **     
        |                                 
        |  +-------------- Sinosolenaia   
        |  |                              
        |  |        +----- Gonidea        
        +--+  +-----+                     
           |  |     |  +-- Leguminaia     
           |  |     +--+                  
           |  |        +-- Microcondylaea 
           +--+                           
              |        +-- Koreosolenaia  
              |  +-----+                  
              |  |     +-- Parvasolenaia  
              +--+                        
                 |  +----- Postolata      
                 +--+                     
                    |  +-- Ptychorhynchus 
                    +--+                  
                       +-- Obovalis

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
4 4 2 3 70 10 3 1 
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
4 4 2 3 40 9 3 1 
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
4 4 2 3 135 45 4 3 
  +----------- Gonideini      
--+                           
  |  +-------- Chamberlainiini
  +--+                        
     |  +----- Lamprotula **     
     +--+                     
        |  +-- Potomida **       
        +--+                  
           +-- Pronodularia **

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
3 3 1 2 31 1 1 2 
  +----- Gonideini 
--+                
  |  +-- Potomida **  
  +--+             
     +-- Lamprotula **

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 38 1 1 4 
  +-------- Gonideini   
--+                     
  |  +----- Pronodularia **
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Potomida **

Jeratthitikul et al., 2025 [Publ 4423], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 48 48 3 1 
           +-- Schepmaniini 
  +--------+                
  |        +-- Pseudodontini
--+                         
  |  +-------- Gonideini    
  +--+                      
     |  +----- Potomida **     
     +--+                   
        |  +-- Lamprotula **   
        +--+                
           +-- Pronodularia **

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iTtTbiriSoTroTroSiD
3 3 1 3 10 2 2 4 
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Lamprotula **     
     +--+                     
        |  +-- Potomida **       
        +--+                  
           +-- Pronodularia **

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 2 28 5 2 2 
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Potomida **       
     +--+                  
        +-- Lamprotula **

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
3 3 1 2 34 32 5 2 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Schepmaniini   
     |  +-----+                  
     |  |     +-- Pseudodontini  
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Potomida **       
           +--+                  
              +-- Lamprotula **

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
3 3 1 3 20 4 2 2 
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
3 3 1 3 14 1 1 2 
  +-------- Gonideini   
--+                     
  |  +----- Potomida **    
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Pronodularia **

Kongim et al., 2023 [Publ 4290], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 12 12 3 3 
  +----------- Chamberlainiini
  |                           
--+        +-- Gonideini      
  |  +-----+                  
  |  |     +-- Pseudodontini  
  +--+                        
     |  +----- Lamprotula **     
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Potomida **

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 1 42 9 2 4 
  +-- Lamprotula **
--+             
  +-- Gonideini

Lopes-Lima et al., 2021 [Publ 4217], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 1 22 3 1 5 
  +-- Potomida ** 
--+            
  +-- Gonideini

Wu et al., 2024 [Publ 4379], fig. 4_6

iTtTbiriSoTroTroSiD
3 3 1 2 20 0 0 2 
  +-- Potomida **  
--+             
  +-- Lamprotula **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 65 5 2 4 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Inversidens    
  |  |                           
  +--+        +-- Obovalis       
     |  +-----+                  
     |  |     +-- Ptychorhynchus 
     +--+                        
        |  +----- Lamprotula **     
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Potomida **

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
3 3 1 2 34 32 5 2 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Potomida **       
     |  +-----+                  
     |  |     +-- Lamprotula **     
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
3 3 1 2 33 3 3 3 
  +-- Pronodularia **
--+               
  +-- Potomida **

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 2 28 5 2 2 
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Potomida **       
     +--+                  
        +-- Lamprotula **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 43 5 2 2 
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
3 3 1 3 73 28 4 3 
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotula **     
  +--+                     
     |  +-- Potomida **       
     +--+                  
        +-- Pronodularia **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 2 66 10 3 2 
  +-------- Chamberlainiini
  |                        
--+     +-- Potomida **       
  |  +--+                  
  |  |  +-- Lamprotula **     
  +--+                     
     |  +-- Pseudodontini  
     +--+                  
        +-- Gonideini

Huang et al., 2019 [Publ 3843], fig. S1

iTtTbiriSoTroTroSiD
3 3 2 1 44 9 2 5 
  +----------------- Chamberlainiini           
  |                                            
  |           +----- Parvasolenaia_rivularis   
--+  +--------+                                
  |  |        |  +-- Parvasolenaia_triangularis
  |  |        +--+                             
  |  |           +-- Ptychorhynchus            
  +--+                                         
     |           +-- Sinosolenaia_carinata     
     |  +--------+                             
     |  |        +-- Lamprotula **                
     +--+                                      
        |  +-------- Sinosolenaia_recognita    
        |  |                                   
        +--+  +----- Sinosolenaia_oleivora__2  
           |  |                                
           +--+  +-- Sinosolenaia_oleivora__1  
              |  |                             
              +--+-- Sinosolenaia_iridinea     
                 |                             
                 +-- Sinosolenaia_oleivora__3

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
3 4 2 2 34 32 5 3 
  +----------------------------- Schepmaniini   
  |                                             
--+  +-------------------------- Pseudodontini  
  |  |                                          
  +--+  +----------------------- Rectidentini   
     |  |                                       
     +--+  +-------------------- Inversidens    
        |  |                                    
        |  |              +----- Lamprotula **     
        +--+  +-----------+                     
           |  |           |  +-- Chamberlainiini
           |  |           +--+                  
           +--+              +-- Potomida **       
              |                                 
              |  +-------------- Sinosolenaia   
              |  |                              
              +--+           +-- Gonidea        
                 |  +--------+                  
                 |  |        +-- Microcondylaea 
                 +--+                           
                    |  +-------- Koreosolenaia  
                    +--+                        
                       |  +----- Parvasolenaia  
                       +--+                     
                          |  +-- Postolata      
                          +--+                  
                             +-- Ptychorhynchus

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 33 33 4 1 
  +----------------- Rectidentini   
--+                                 
  |  +-------------- Pseudodontini  
  +--+                              
     |  +----------- Gonideini      
     +--+                           
        |  +-------- Chamberlainiini
        +--+                        
           |  +----- Potomida **       
           +--+                     
              |  +-- Lamprotula **     
              +--+                  
                 +-- Pronodularia **

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iTtTbiriSoTroTroSiD
3 3 1 3 10 2 2 2 
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 2 50 6 2 2 
  +----- Gonideini 
--+                
  |  +-- Potomida **  
  +--+             
     +-- Lamprotula **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 46 5 2 2 
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
3 4 2 2 34 32 5 3 
  +----------------------------- Schepmania_nieuwenhuisi__1
  |                                                        
  +----------------------------- Schepmania_nieuwenhuisi__2
--+                                                        
  |  +-------------------------- Pseudodontini             
  |  |                                                     
  +--+  +----------------------- Rectidentini              
     |  |                                                  
     +--+  +-------------------- Inversidens               
        |  |                                               
        |  |              +----- Lamprotula **                
        +--+  +-----------+                                
           |  |           |  +-- Chamberlainiini           
           |  |           +--+                             
           +--+              +-- Potomida **                  
              |                                            
              |  +-------------- Sinosolenaia              
              |  |                                         
              +--+  +----------- Gonidea                   
                 |  |                                      
                 +--+----------- Microcondylaea            
                    |                                      
                    |  +-------- Koreosolenaia             
                    +--+                                   
                       |  +----- Parvasolenaia             
                       +--+                                
                          |  +-- Postolata                 
                          +--+                             
                             +-- Ptychorhynchus

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 28 4 2 2 
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
3 3 1 3 17 2 2 2 
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 65 5 2 4 
  +-------- Gonideini   
--+                     
  |  +----- Lamprotula **  
  +--+                  
     |  +-- Potomida **    
     +--+               
        +-- Pronodularia **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 50 8 3 2 
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 2 31 1 1 2 
  +----- Gonideini 
--+                
  |  +-- Potomida **  
  +--+             
     +-- Lamprotula **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
3 3 1 3 36 9 3 2 
  +-------------- Chamberlainiini
--+                              
  |  +----------- Gonideini      
  +--+                           
     |  +-------- Pseudodontini  
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Lamprotula **     
           +--+                  
              +-- Pronodularia **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
3 3 1 2 33 3 3 3 
  +-- Pronodularia **
--+               
  +-- Potomida **

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
3 6 0 3 33 3 1 6 
  +-- Lamprotula **    
  |                 
  +-- Potomida **      
  |                 
  +-- Pronodularia **  
--+                 
  +-- Gonidea       
  |                 
  +-- Leguminaia    
  |                 
  +-- Microcondylaea

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
3 3 1 3 19 1 1 2 
  +-------- Gonideini   
--+                     
  |  +----- Potomida **    
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Pronodularia **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 65 5 2 4 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Inversidens    
  |  |                           
  +--+        +-- Ptychorhynchus 
     |  +-----+                  
     |  |     +-- Obovalis       
     +--+                        
        |  +----- Lamprotula **     
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Potomida **

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
3 3 1 2 34 32 5 2 
              +-- Schepmaniini   
  +-----------+                  
  |           +-- Pseudodontini  
--+                              
  |  +----------- Gonideini      
  +--+                           
     |  +-------- Rectidentini   
     +--+                        
        |  +----- Chamberlainiini
        +--+                     
           |  +-- Potomida **       
           +--+                  
              +-- Lamprotula **

Zhou et al., 2007 [Publ 2658], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 1 15 0 0 3 
1

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 38 1 1 4 
  +-------- Gonideini   
--+                     
  |  +----- Pronodularia **
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Potomida **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
3 3 1 3 73 28 4 1 
  +-------- Chamberlainiini
--+                        
  |  +----- Potomida **       
  +--+                     
     |  +-- Lamprotula **     
     +--+                  
        +-- Pronodularia **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
3 13 0 3 80 32 3 13 
  +-- Lamprotula **    
  |                 
  +-- Potomida **      
  |                 
  +-- Pronodularia **  
  |                 
  +-- Gonidea       
  |                 
  +-- Leguminaia    
  |                 
  +-- Microcondylaea
  |                 
--+-- Amblemini     
  |                 
  +-- Ortmanniana   
  |                 
  +-- Lampsilis     
  |                 
  +-- Cambarunio    
  |                 
  +-- Elliptio      
  |                 
  +-- Pleurobema    
  |                 
  +-- Quadrulini

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
3 3 1 2 34 32 5 2 
  +-------------------------- Chamberlainiini
  |                                          
--+  +----------------------- Rectidentini   
  |  |                                       
  |  |                    +-- Schepmaniini   
  +--+  +-----------------+                  
     |  |                 +-- Pseudodontini  
     |  |                                    
     +--+              +----- Inversidens    
        |  +-----------+                     
        |  |           |  +-- Potomida **       
        |  |           +--+                  
        +--+              +-- Lamprotula **     
           |                                 
           |  +-------------- Sinosolenaia   
           |  |                              
           +--+           +-- Gonidea        
              |  +--------+                  
              |  |        +-- Microcondylaea 
              +--+                           
                 |  +-------- Parvasolenaia  
                 +--+                        
                    |  +----- Koreosolenaia  
                    +--+                     
                       |  +-- Postolata      
                       +--+                  
                          +-- Ptychorhynchus

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
3 3 1 3 19 1 1 2 
  +-------- Gonideini   
--+                     
  |  +----- Potomida **    
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Pronodularia **

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 11 2 1 2 
  +-------- Gonideini   
--+                     
  |  +----- Potomida **    
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Pronodularia **

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 57 57 5 3 
           +-- Chamberlainiini
  +--------+                  
  |        +-- Rectidentini   
  |                           
--+     +----- Pronodularia **   
  |  +--+                     
  |  |  |  +-- Lamprotula **     
  |  |  +--+                  
  +--+     +-- Potomida **       
     |                        
     |  +----- Gonideini      
     +--+                     
        |  +-- Schepmaniini   
        +--+                  
           +-- Pseudodontini

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 29 4 2 2 
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 37 11 3 2 
  +-------------- Rectidentini 
--+                            
  |  +----------- Pseudodontini
  +--+                         
     |  +-------- Gonideini    
     +--+                      
        |  +----- Potomida **     
        +--+                   
           |  +-- Pronodularia ** 
           +--+                
              +-- Lamprotula **

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 32 32 4 1 
  +----------------- Rectidentini   
--+                                 
  |  +-------------- Pseudodontini  
  +--+                              
     |  +----------- Gonideini      
     +--+                           
        |  +-------- Chamberlainiini
        +--+                        
           |  +----- Potomida **       
           +--+                     
              |  +-- Pronodularia **   
              +--+                  
                 +-- Lamprotula **

Froufe et al., 2015 [Publ 3664], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 9 4 2 2 
  +-------- Gonideini   
--+                     
  |  +----- Potomida **    
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Pronodularia **

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
3 4 2 1 18 2 1 4 
  +----------- Chamberlainiini        
--+                                   
  |  +-------- Lamprotula_leaii **       
  +--+                                
     |  +----- Lamprotula_caveata **     
     +--+                             
        |  +-- Gibbosula              
        +--+                          
           +-- Lamprotula_cornuumlunae **

Campbell et al., 2005 [Publ 2660], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 100 2 2 -- 
  +----- Gonideini   
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Zhou et al., 2017 [Publ 3788], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 14 5 2 -- 
  +----- Chamberlainiini
--+                     
  |  +-- Lamprotula **     
  +--+                  
     +-- Gonideini

Sano et al., 2017 [Publ 3799], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 31 6 2 -- 
           +-- Obovalis       
  +--------+                  
  |        +-- Ptychorhynchus 
  |                           
--+        +-- Inversidens    
  |  +-----+                  
  |  |     +-- Sinosolenaia   
  +--+                        
     |  +----- Chamberlainiini
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Froufe et al., 2019 [Publ 3936], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 17 6 3 -- 
  +-------- Chamberlainiini
  |                        
--+     +-- Potomida **       
  |  +--+                  
  |  |  +-- Pronodularia **   
  +--+                     
     |  +-- Gonideini      
     +--+                  
        +-- Pseudodontini

Konopleva et al., 2021 [Publ 4213], fig. 1

iTtTbiriSoTroTroSiD
2 27 0 2 25 25 2 28 
  +-- Lamprotula **   
  |                
  +-- Leguminaia   
  |                
--+-- Potomida **     
  |                
  +-- Gonidea      
  |                
  +-- Pseudodontini

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 19 4 2 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Lamprotula **     
     +--+                  
        +-- Pronodularia **

Ouyang et al., 2015 [Publ 4493], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 29 4 2 -- 
1

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

iTtTbiriSoTroTroSiD
2 2 0 2 8 1 1 -- 
  +----- Gonideini   
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 43 0 0 -- 
  +-- Pronodularia **
--+               
  +-- Lamprotula **

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 39 0 0 -- 
  +-- Pronodularia **
--+               
  +-- Lamprotula **

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
2 9 0 2 14 0 0 10 
  +----------------------- Lamprotula **     
--+                                       
  |  +-------------------- Potomida **       
  +--+                                    
     |  +----------------- Lampsilini     
     +--+                                 
        |  +-------------- Pleurobemini   
        +--+                              
           |  +----------- Aculamprotulina
           +--+                           
              |  +-------- Unio_pictorum  
              +--+                        
                 |  +----- Anodontini     
                 +--+                     
                    |  +-- Unio_ravoisieri
                    +--+                  
                       +-- Lamellidentini

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
2 2 0 2 13 0 0 -- 
  +-- Lamprotula **
--+             
  +-- Potomida **

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
2 2 0 2 81 67 3 -- 
  +----------- Pseudodontini
--+                         
  |  +-------- Rectidentini 
  +--+                      
     |  +----- Gonideini    
     +--+                   
        |  +-- Lamprotula **   
        +--+                
           +-- Potomida **

Zhang et al., 2024 [Publ 4397], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 6 6 1 -- 
  +-- Lamprotula **
--+             
  +-- Gonideini

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 39 0 0 -- 
  +-- Pronodularia **
--+               
  +-- Lamprotula **

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 39 0 0 -- 
  +-- Pronodularia **
--+               
  +-- Lamprotula **

Zhou et al., 2016 [Publ 3710], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 18 4 2 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Lamprotula **     
     +--+                  
        +-- Pronodularia **

Froufe et al., 2015 [Publ 3664], fig. 1-M

iTtTbiriSoTroTroSiD
2 2 0 2 6 1 1 -- 
  +----- Gonideini   
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
2 2 0 2 21 10 4 -- 
  +-------------- Chamberlainiini
--+                              
  |  +----------- Rectidentini   
  +--+                           
     |  +-------- Pseudodontini  
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Potomida **

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 43 0 0 -- 
  +-- Pronodularia **
--+               
  +-- Lamprotula **

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 38 5 2 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Lamprotula **     
     +--+                  
        +-- Potomida **

Lopes-Lima et al., 2020 [Publ 4080], fig. 14

iTtTbiriSoTroTroSiD
2 2 0 1 4 2 1 -- 
  +-- Gonideini 
--+             
  +-- Lamprotula **

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
2 2 0 2 27 0 0 -- 
  +-- Potomida **  
--+             
  +-- Lamprotula **

Fonseca et al., 2016 [Publ 3775], fig. 1-M

iTtTbiriSoTroTroSiD
2 2 0 2 7 1 1 -- 
  +----- Gonideini   
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Campbell et al., 2005 [Publ 2660], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 100 2 2 -- 
  +----- Gonideini   
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 20 4 2 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Pronodularia **   
     +--+                  
        +-- Lamprotula **

Bolotov et al., 2022 [Publ 4250], fig. 8

iTtTbiriSoTroTroSiD
2 7 0 2 47 21 3 7 
  +-- Gibbosula    
  |                
  +-- Margaritifera
  |                
  +-- Sinosolenaia 
  |                
--+-- Leguminaia   
  |                
  +-- Gonidea      
  |                
  +-- Lamprotula **   
  |                
  +-- Potomida **

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 39 0 0 -- 
  +-- Pronodularia **
--+               
  +-- Lamprotula **

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
2 34 0 1 32 9 3 34 
                       +----- Prolasmidonta           
                    +--+                              
                    |  |  +-- Alasmidonta             
                    |  +--+                           
  +-----------------+     +-- Anemina                 
  |                 |                                 
  |                 |  +----- Pseudodontini           
  |                 +--+                              
  |                    |  +-- Lamprotula_caveata **      
  |                    +--+                           
  |                       +-- Chamberlainia           
  |                                                   
  |                    +----- Oxynaiina               
  |              +-----+                              
  |              |     |  +-- Lepidodesmini           
--+              |     +--+                           
  |              |        +-- Aculamprotulina         
  |  +-----------+                                    
  |  |           |     +----- Platynaias              
  |  |           |  +--+                              
  |  |           |  |  |  +-- Utterbackia_imbecillis  
  |  |           |  |  +--+                           
  |  |           +--+     +-- Pyganodon               
  |  |              |                                 
  |  |              |  +----- Sinanodonta             
  |  |              +--+                              
  |  |                 |  +-- Cristaria               
  +--+                 +--+                           
     |                    +-- Anodontina              
     |                                                
     |                    +-- Utterbackia_peninsularis
     |              +-----+                           
     |              |     +-- Potamilus_alatus        
     |  +-----------+                                 
     |  |           |  +----- Lamprotula_leaii **        
     |  |           +--+                              
     |  |              |  +-- Sinohyriopsis           
     +--+              +--+                           
        |                 +-- Gonideini               
        |                                             
        |  +----------------- Gibbosula               
        +--+                                          
           |  +-------------- Quadrulini              
           +--+                                       
              |  +----------- Toxolasma               
              +--+                                    
                 |  +-------- Lampsilis_ornata        
                 +--+                                 
                    |  +----- Potamilus_leptodon      
                    +--+                              
                       |  +-- Lampsilis_siliquoidea   
                       +--+                           
                          +-- Lampsilis_powellii

Froufe et al., 2019 [Publ 3936], fig. 3-M

iTtTbiriSoTroTroSiD
2 2 0 2 17 6 3 -- 
  +-------- Chamberlainiini
  |                        
--+     +-- Potomida **       
  |  +--+                  
  |  |  +-- Pronodularia **   
  +--+                     
     |  +-- Gonideini      
     +--+                  
        +-- Pseudodontini

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
2 9 0 2 13 0 0 10 
  +-- Aculamprotulina
  |                  
  +-- Lamprotula **     
  |                  
  +-- Lampsilini     
  |                  
  +-- Potomida **       
  |                  
--+-- Unio_pictorum  
  |                  
  +-- Anodontini     
  |                  
  +-- Unio_ravoisieri
  |                  
  +-- Lamellidentini 
  |                  
  +-- Pleurobemini

Ouyang et al., 2015 [Publ 4493], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 29 4 2 -- 
  +----- Lamprotula **     
--+                     
  |  +-- Chamberlainiini
  +--+                  
     +-- Gonideini

Hou et al., 2024 [Publ 4400], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 13 0 0 -- 
1

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 32 6 2 -- 
  +-------- Gonideini      
--+                        
  |  +----- Chamberlainiini
  +--+                     
     |  +-- Lamprotula **     
     +--+                  
        +-- Pronodularia **

Liu et al., 2024 [Publ 4380], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 1 10 10 1 -- 
  +-- Lamprotula **
--+             
  +-- Gonideini

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 35 9 3 -- 
  +-------- Chamberlainiini
  |                        
--+     +-- Lamprotula **     
  |  +--+                  
  |  |  +-- Potomida **       
  +--+                     
     |  +-- Pseudodontini  
     +--+                  
        +-- Gonideini

Zieritz et al., 2020 [Publ 4102], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 30 30 4 -- 
  +----------- Chamberlainiini
--+                           
  |  +-------- Lamprotula **     
  +--+                        
     |  +----- Gonideini      
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Rectidentini

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 33 3 2 -- 
-- Lamprotula_leaii **

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
1 1 0 1 24 4 2 -- 
  +----- Chamberlainiini
--+                     
  |  +-- Pronodularia **   
  +--+                  
     +-- Gonideini

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Potomida_littoralis **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 27 0 0 -- 
-- Potomida_littoralis **

Pfeiffer et al., 2021 [Publ 4126], fig. 9

iTtTbiriSoTroTroSiD
1 1 0 1 28 28 1 -- 
  +-- Lamprotula **  
--+               
  +-- Rectidentini

Amaro et al., 2017 [Publ 3791], fig. 3B-M

iTtTbiriSoTroTroSiD
1 1 0 1 6 1 1 -- 
  +-- Pronodularia **
--+               
  +-- Gonideini

Shirai et al., 2010 [Publ 3353], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 2 2 1 -- 
  +-- Pronodularia **   
--+                  
  +-- Chamberlainiini

Pfarrer et al., 2022 [Publ 4305], fig. 2-5

iTtTbiriSoTroTroSiD
1 1 0 1 11 1 1 -- 
  +-- Potomida ** 
--+            
  +-- Gonideini

Wen et al., 2017 [Publ 3795], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 15 0 0 -- 
-- Lamprotula_leaii **

Chase et al., 2018 [Publ 3861], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Lamprotula_leaii **

Guerra et al., 2017 [Publ 3772], fig. 2b-M

iTtTbiriSoTroTroSiD
1 1 0 1 7 1 1 -- 
  +-- Gonideini   
--+               
  +-- Pronodularia **

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 18 1 1 -- 
  +-- Chamberlainiini
--+                  
  +-- Lamprotula **

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
1 1 0 1 24 4 2 -- 
  +----- Chamberlainiini
--+                     
  |  +-- Pronodularia **   
  +--+                  
     +-- Gonideini

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 23 1 1 -- 
  +-- Gonideini 
--+             
  +-- Lamprotula **

Nagel et al., 1998 [Publ 1946], fig. 9

iTtTbiriSoTroTroSiD
1 1 0 1 5 1 1 -- 
-- Potomida **

Khalloufi et al., 2011 [Publ 2748], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Potomida_littoralis **

Välilä et al., 2017 [Publ 3793], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 2 0 0 -- 
-- Potomida_littoralis **

Amaro et al., 2017 [Publ 3791], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 15 1 1 -- 
  +-- Pronodularia **
--+               
  +-- Gonideini

Goncalves et al., 2022 [Publ 4253], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 6 6 1 -- 
  +-- Lamprotula **     
--+                  
  +-- Chamberlainiini

Prié & Puillandre, 2014 [Publ 3529], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 16 0 0 -- 
-- Potomida_littoralis **

Chapman et al., 2008 [Publ 2667], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 20 0 0 -- 
-- Pronodularia_japanensis **

Huang et al., 2013 [Publ 3629], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 15 3 2 -- 
  +----- Chamberlainiini
--+                     
  |  +-- Gonideini      
  +--+                  
     +-- Pronodularia **

Walker et al., 2006 [Publ 2666], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 2 2 -- 
-- Pronodularia_japanensis **

Zhang et al., 2004 [Publ 2735], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 13 1 1 -- 
  +-- Lamprotula **     
--+                  
  +-- Chamberlainiini

Lopes-Lima et al., 2017 [Publ 3734], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 16 1 1 -- 
  +-- Potomida ** 
--+            
  +-- Gonideini

Sheng et al., 2014 [Publ 3704], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 7 2 1 -- 
  +-- Lamprotula **     
--+                  
  +-- Chamberlainiini

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 3 2 -- 
-- Lamprotula_leaii **

Amaro et al., 2017 [Publ 3791], fig. 3A

iTtTbiriSoTroTroSiD
1 1 0 1 11 1 1 -- 
  +-- Pronodularia **
--+               
  +-- Gonideini

Chapman et al., 2008 [Publ 2744], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 20 0 0 -- 
-- Pronodularia_japanensis **

Sano et al., 2017 [Publ 3799], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 20 3 2 -- 
  +-------- Obovalis       
--+                        
  |  +----- Inversidens    
  +--+                     
     |  +-- Pronodularia **   
     +--+                  
        +-- Chamberlainiini

Nagel et al., 1998 [Publ 1946], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 8 1 1 -- 
-- Potomida_littoralis **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 16 4 -- 
  +----------- Chamberlainiini
--+                           
  |  +-------- Rectidentini   
  +--+                        
     |  +----- Pseudodontini  
     +--+                     
        |  +-- Gonideini      
        +--+                  
           +-- Lamprotula **

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 11 3 2 -- 
  +-------- Obovalis       
--+                        
  |  +----- Inversidens    
  +--+                     
     |  +-- Chamberlainiini
     +--+                  
        +-- Pronodularia **

Reis et al., 2013 [Publ 3580], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Potomida_littoralis **

Amaro et al., 2017 [Publ 3791], fig. 3A-M

iTtTbiriSoTroTroSiD
1 1 0 1 6 1 1 -- 
  +-- Pronodularia **
--+               
  +-- Gonideini

Plazzi & Passamonti, 2010 [Publ 3354], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 1 0 0 -- 
-- Pronodularia_japanensis **

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 22 1 1 -- 
  +-- Gonideini 
--+             
  +-- Lamprotula **

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 31 27 5 -- 
  +-------------- Chamberlainiini
--+                              
  |  +----------- Potomida **       
  +--+                           
     |  +-------- Rectidentini   
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 27 0 0 -- 
-- Potomida_littoralis **

Plazzi et al., 2011 [Publ 3676], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 17 2 1 -- 
-- Pronodularia_japanensis **

Xue et al., 2016 [Publ 3659], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 14 1 1 -- 
  +-- Chamberlainiini
--+                  
  +-- Lamprotula **

Jiang et al., 2010 [Publ 3438], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 3 1 1 -- 
  +-- Pronodularia **   
--+                  
  +-- Chamberlainiini

Soroka, 2005 [Publ 2726], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Pronodularia_japanensis **

Chen et al., 2012 [Publ 3624], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 6 1 1 -- 
  +-- Chamberlainiini
--+                  
  +-- Lamprotula **

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Potomida_littoralis **

Chase et al., 2018 [Publ 3861], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Lamprotula_leaii **

Amaro et al., 2017 [Publ 3791], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 9 1 1 -- 
  +-- Pronodularia **
--+               
  +-- Gonideini

Machordom et al., 2003 [Publ 2688], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 3 0 0 -- 
-- Potomida_littoralis **

Huang et al., 2013 [Publ 3629], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 5 1 1 -- 
  +-- Gonideini   
--+               
  +-- Pronodularia **

Araujo et al., 2009 [Publ 2589], fig. 7C

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Potomida_littoralis **

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iTtTbiriSoTroTroSiD
1 1 0 1 24 1 1 -- 
-- Pronodularia_japanensis **

Froufe et al., 2017 [Publ 3802], fig. 7

iTtTbiriSoTroTroSiD
1 1 0 1 9 1 1 -- 
  +-- Gonideini
--+            
  +-- Potomida **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 16 4 -- 
  +----------- Chamberlainiini
--+                           
  |  +-------- Rectidentini   
  +--+                        
     |  +----- Lamprotula **     
     +--+                     
        |  +-- Gonideini      
        +--+                  
           +-- Pseudodontini

Zhang et al., 2013 [Publ 4214], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 3 0 0 -- 
-- Lamprotula_leaii **

Machordom et al., 2003 [Publ 2688], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 2 0 0 -- 
-- Potomida_littoralis **

Smith, 2021 [Publ 4331], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 19 2 2 -- 
  +----- Chamberlainiini
--+                     
  |  +-- Rectidentini   
  +--+                  
     +-- Lamprotula **

Amaro et al., 2017 [Publ 3791], fig. 3B

iTtTbiriSoTroTroSiD
1 1 0 1 11 1 1 -- 
  +-- Pronodularia **
--+               
  +-- Gonideini

Reis et al., 2013 [Publ 3580], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Potomida_littoralis **

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 23 1 1 -- 
  +-- Gonideini 
--+             
  +-- Lamprotula **

Araujo et al., 2009 [Publ 2589], fig. 7A

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Potomida_littoralis **

Guerra et al., 2017 [Publ 3772], fig. 2a-M

iTtTbiriSoTroTroSiD
1 1 0 1 7 1 1 -- 
  +-- Gonideini   
--+               
  +-- Pronodularia **

Ouyang et al., 2011 [Publ 3408], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 12 3 1 -- 
  +-- Lamprotula **
--+             
  +-- Gonideini

Chen et al., 2025 [Publ 4455], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 25 0 0 -- 
-- Potomida_littoralis **

Araujo et al., 2016 [Publ 3746], fig. 1_2b

iTtTbiriSoTroTroSiD
1 1 0 1 12 0 0 -- 
-- Potomida_littoralis **

Breton et al., 2011 [Publ 3376], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 36 1 1 -- 
-- Pronodularia_japanensis **

Kano et al., 2019 [Publ 4194], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 9 2 1 -- 
  +----- Inversidens 
--+                  
  |  +-- Obovalis    
  +--+               
     +-- Pronodularia **

Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3

iTtTbiriSoTroTroSiD
1 1 0 1 3 1 1 -- 
  +-- Chamberlainiini
--+                  
  +-- Pronodularia **

Araujo et al., 2009 [Publ 2589], fig. 7B

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Potomida_littoralis **