Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae

tribe LAMPROTULINI Modell, 1942: cladomics

4 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 22 (41 total)
number of subtaxa (rank: genus) = 4
analyzed in 202 cladograms in 123 publications
maximum number of terminals in an analysis = 5 (mean = 2.44, median = 2)
maximum number of subtaxa (genus) = 3 (mean = 1.96, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 133
maximum number of included terminals = 5, (mean = 3.19 median = 3)
maximum number of subtaxa (genus) = 3 (mean = 2.45, median = 3)
proportion of cladograms that support basic monophyly = 0.89

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 109
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 5 (mean = 3.39, median = 3)
maximum number of subtaxa (genus) = 3 (mean = 2.67, median = 3)
proportion of cladograms that support rigorous monophyly = 0.91

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Lamprotulini cladogram distances
PCoA Ordination

NJ Tree of Lamprotulini cladogram distances
NJ Tree

Cladograms

Chen et al., 2025 [Publ 4663], fig. 1

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
4 4 2 3 45 40 5 0 
  +-------------- Rectidentini   
  |                              
--+  +----------- Chamberlainiini
  |  |                           
  |  |     +----- Potomida **       
  +--+  +--+                     
     |  |  |  +-- Pronodularia **   
     |  |  +--+                  
     +--+     +-- Lamprotula **     
        |                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini

Zieritz et al., 2024 [Publ 4381], fig. 2

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
2 42 0 2 44 33 6 45 
     +----------- Rectidentini   
     |                           
  +--+        +-- Schepmaniini   
  |  |  +-----+                  
  |  |  |     +-- Pseudodontini  
  |  +--+                        
  |     |  +----- Caudiculatus   
  |     +--+                     
  |        |  +-- Potomida **       
  |        +--+                  
--+           +-- Gonideini      
  |                              
  |     +-------- Lamellidentini 
  |  +--+                        
  |  |  |  +----- Leoparreysiini 
  |  |  +--+                     
  |  |     |  +-- Indochinellini 
  |  |     +--+                  
  +--+        +-- Coelaturini    
     |                           
     |        +-- Anodontini     
     |     +--+                  
     |     |  +-- Unionini       
     +-----+                     
           |  +-- Chamberlainiini
           +--+                  
              +-- Discomya **

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
5 16 0 3 113 67 4 15 
  +-------- Rectidentini             
  |                                  
  |  +----- Pseudodontini            
  |  |                               
  |  |  +-- Sinosolenaia_iridinea    
--+  |  |                            
  |  |  +-- Sinosolenaia_recognita__1
  |  |  |                            
  |  |  +-- Sinosolenaia_carinata    
  |  |  |                            
  |  |  +-- Leguminaia               
  +--+  |                            
     |  +-- Microcondylaea           
     |  |                            
     |  +-- Gonidea                  
     |  |                            
     |  +-- Ptychorhynchus           
     |  |                            
     |  +-- Parvasolenaia            
     |  |                            
     +--+-- Lamprotula_leaii **         
        |                            
        +-- Lamprotula **               
        |                            
        +-- Lamprotula_caveata **       
        |                            
        +-- Potomida **                 
        |                            
        +-- Pronodularia **             
        |                            
        +-- Sinohyriopsis            
        |                            
        +-- Chamberlainia            
        |                            
        +-- Sinosolenaia_recognita__2
        |                            
        +-- Sinosolenaia_oleivora

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
5 5 4 3 26 6 4 6 
  +-------------- Chamberlainiini
  |                              
--+           +-- Gonideini      
  |  +--------+                  
  |  |        +-- Pseudodontini  
  +--+                           
     |  +-------- Rectidentini   
     +--+                        
        |  +----- Pronodularia **   
        +--+                     
           |  +-- Potomida **       
           +--+                  
              +-- Lamprotula **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
5 5 3 3 91 15 4 2 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **

Matos et al., 2024 [Publ 4384], fig. 3

iTtTbiriSoTroTroSiD
5 5 3 3 15 13 4 2 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
5 5 3 3 48 17 4 2 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
5 5 3 3 87 16 4 2 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
5 5 3 3 194 50 4 4 
  +-------- Gonideini   
--+                     
  |  +----- Lamprotula **  
  +--+                  
     |  +-- Potomida **    
     +--+               
        +-- Pronodularia **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
5 5 3 3 87 16 4 2 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
5 5 3 3 196 51 4 4 
  +----------- Chamberlainiini
  |                           
--+        +-- Gonideini      
  |  +-----+                  
  |  |     +-- Pseudodontini  
  +--+                        
     |  +----- Lamprotula **     
     +--+                     
        |  +-- Potomida **       
        +--+                  
           +-- Pronodularia **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
5 5 3 3 91 15 4 2 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **

Froufe et al., 2016 [Publ 3727], fig. 3_4

iTtTbiriSoTroTroSiD
5 7 3 3 2 2 1 5 
  +----- Potomida **      
  |                    
--+  +-- Pronodularia **  
  |  |                 
  |  +-- Lamprotula **    
  +--+                 
     +-- Leguminaia    
     |                 
     +-- Microcondylaea

Zhang et al., 2024 [Publ 4397], fig. 5

iTtTbiriSoTroTroSiD
5 5 3 3 22 20 4 2 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
5 5 4 3 117 28 4 6 
           +-- Pseudodontini  
  +--------+                  
  |        +-- Gonideini      
--+                           
  |  +-------- Chamberlainiini
  +--+                        
     |  +----- Pronodularia **   
     +--+                     
        |  +-- Potomida **       
        +--+                  
           +-- Lamprotula **

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
5 5 0 3 107 67 4 3 
  +-------------- Rectidentini      
  |                                 
--+  +----------- Pseudodontini     
  |  |                              
  +--+  +-------- Gonideini         
     |  |                           
     +--+  +----- Chamberlainiini   
        |  |                        
        |  |  +-- Lamprotula_leaii **  
        +--+  |                     
           |  +-- Lamprotula **        
           |  |                     
           +--+-- Lamprotula_caveata **
              |                     
              +-- Potomida **          
              |                     
              +-- Pronodularia **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
5 5 3 3 196 51 4 4 
  +----- Lamprotula **  
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Matos et al., 2024 [Publ 4383], fig. 3

iTtTbiriSoTroTroSiD
5 5 3 3 15 13 4 1 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
4 4 2 3 40 9 3 1 
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
4 4 2 3 87 25 4 1 
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 3 52 6 2 1 
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
4 4 2 3 76 24 4 3 
        +-------- Chamberlainiini
  +-----+                        
  |     |  +----- Lamprotula **     
  |     +--+                     
  |        |  +-- Potomida **       
  |        +--+                  
--+           +-- Pronodularia **   
  |                              
  |           +-- Parvasolenaia  
  |  +--------+                  
  |  |        +-- Pseudodontini  
  +--+                           
     |  +-------- Sinosolenaia   
     +--+                        
        |  +----- Gonidea        
        +--+                     
           |  +-- Microcondylaea 
           +--+                  
              +-- Leguminaia

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
4 4 3 2 26 4 2 3 
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Pronodularia **   
     +--+                  
        +-- Lamprotula **

Lopes-Lima et al., 2020 [Publ 4080], fig. 16

iTtTbiriSoTroTroSiD
4 4 3 2 4 2 1 3 
  +-------- Chamberlainia
--+                      
  |  +----- Sinohyriopsis
  +--+                   
     |  +-- Pronodularia ** 
     +--+                
        +-- Lamprotula **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
4 4 2 3 87 25 4 3 
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Pronodularia **   
     +--+                     
        |  +-- Potomida **       
        +--+                  
           +-- Lamprotula **

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
4 4 2 3 38 4 2 1 
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
4 4 2 3 44 39 5 0 
  +-------------- Rectidentini   
  |                              
--+  +----------- Chamberlainiini
  |  |                           
  |  |     +----- Potomida **       
  +--+  +--+                     
     |  |  |  +-- Pronodularia **   
     |  |  +--+                  
     +--+     +-- Lamprotula **     
        |                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
4 4 2 3 70 10 3 1 
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
4 4 2 3 49 15 4 1 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
4 4 2 3 38 6 3 1 
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
4 4 2 3 34 5 2 1 
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
4 4 2 3 41 39 4 0 
  +----------------------- Pseudodontini  
  |                                       
  |  +-------------------- Rectidentini   
--+  |                                    
  |  |  +----------------- Inversidens    
  |  |  |                                 
  +--+  |        +-------- Chamberlainiini
     |  +--------+                        
     |  |        |  +----- Potomida **       
     |  |        +--+                     
     +--+           |  +-- Pronodularia **   
        |           +--+                  
        |              +-- Lamprotula **     
        |                                 
        |  +-------------- Sinosolenaia   
        |  |                              
        |  |        +----- Gonidea        
        +--+  +-----+                     
           |  |     |  +-- Leguminaia     
           |  |     +--+                  
           |  |        +-- Microcondylaea 
           +--+                           
              |        +-- Koreosolenaia  
              |  +-----+                  
              |  |     +-- Parvasolenaia  
              +--+                        
                 |  +----- Postolata      
                 +--+                     
                    |  +-- Ptychorhynchus 
                    +--+                  
                       +-- Obovalis

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
4 4 2 3 70 10 3 1 
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 3 84 16 4 1 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
4 4 2 3 41 39 4 0 
  +----------------------- Pseudodontini  
  |                                       
  |  +-------------------- Rectidentini   
--+  |                                    
  |  |        +----------- Inversidens    
  |  |  +-----+                           
  +--+  |     |  +-------- Chamberlainiini
     |  |     +--+                        
     |  |        |  +----- Potomida **       
     |  |        +--+                     
     |  |           |  +-- Pronodularia **   
     +--+           +--+                  
        |              +-- Lamprotula **     
        |                                 
        |  +-------------- Sinosolenaia   
        |  |                              
        |  |        +----- Gonidea        
        +--+  +-----+                     
           |  |     |  +-- Leguminaia     
           |  |     +--+                  
           |  |        +-- Microcondylaea 
           +--+                           
              |        +-- Koreosolenaia  
              |  +-----+                  
              |  |     +-- Parvasolenaia  
              +--+                        
                 |  +----- Postolata      
                 +--+                     
                    |  +-- Ptychorhynchus 
                    +--+                  
                       +-- Obovalis

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
4 4 2 3 135 45 4 3 
  +----------- Gonideini      
--+                           
  |  +-------- Chamberlainiini
  +--+                        
     |  +----- Lamprotula **     
     +--+                     
        |  +-- Potomida **       
        +--+                  
           +-- Pronodularia **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
4 4 2 3 50 8 3 3 
  +-------------- Chamberlainiini
--+                              
  |  +----------- Pseudodontini  
  +--+                           
     |  +-------- Gonideini      
     +--+                        
        |  +----- Lamprotula **     
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Potomida **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 3 82 14 4 1 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
4 4 2 3 135 45 4 3 
  +-------------- Pseudodontini  
--+                              
  |  +----------- Gonideini      
  +--+                           
     |  +-------- Chamberlainiini
     +--+                        
        |  +----- Lamprotula **     
        +--+                     
           |  +-- Potomida **       
           +--+                  
              +-- Pronodularia **

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 3 40 9 3 1 
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
4 4 2 3 52 9 2 0 
  +----------- Gonideini      
--+                           
  |  +-------- Chamberlainiini
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 3 43 38 5 2 
  +-------------- Rectidentini   
  |                              
--+  +----------- Chamberlainiini
  |  |                           
  |  |     +----- Lamprotula **     
  +--+  +--+                     
     |  |  |  +-- Pronodularia **   
     |  |  +--+                  
     +--+     +-- Potomida **       
        |                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini

Wu et al., 2021 [Publ 4215], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 3 20 15 3 3 
  +-------------------- Chamberlainiini
  |                                    
  |  +----------------- Pseudodontini  
--+  |                                 
  |  |        +-------- Sinosolenaia   
  |  |  +-----+                        
  +--+  |     |  +----- Lamprotula **     
     |  |     +--+                     
     |  |        |  +-- Potomida **       
     |  |        +--+                  
     +--+           +-- Pronodularia **   
        |                              
        |           +-- Ptychorhynchus 
        |  +--------+                  
        |  |        +-- Parvasolenaia  
        +--+                           
           |  +-------- Inversidens    
           +--+                        
              |  +----- Gonidea        
              +--+                     
                 |  +-- Microcondylaea 
                 +--+                  
                    +-- Leguminaia

Wu et al., 2020 [Publ 4147], fig. 4

iTtTbiriSoTroTroSiD
4 4 2 2 10 1 1 3 
  +----- Gonideini 
--+                
  |  +-- Potomida **  
  +--+             
     +-- Lamprotula **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
4 13 0 3 121 47 4 12 
  +-- Chamberlainia         
  |                         
  +-- Sinohyriopsis         
  |                         
  +-- Lamprotula_caveata **    
  |                         
  +-- Lamprotula_leaii **      
  |                         
  +-- Potomida **              
  |                         
  +-- Pronodularia **          
  |                         
--+-- Gonidea               
  |                         
  +-- Leguminaia_wheatleyi  
  |                         
  +-- Leguminaia            
  |                         
  +-- Microcondylaea        
  |                         
  +-- Sinosolenaia_carinata 
  |                         
  +-- Sinosolenaia_oleivora 
  |                         
  +-- Sinosolenaia_recognita

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
4 4 2 3 44 13 4 1 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Pseudodontini  
     |  +-----+                  
     |  |     +-- Gonideini      
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Lamprotula **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 3 70 10 3 1 
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
3 4 2 1 18 2 1 4 
  +----------- Chamberlainiini        
--+                                   
  |  +-------- Lamprotula_leaii **       
  +--+                                
     |  +----- Lamprotula_caveata **     
     +--+                             
        |  +-- Gibbosula              
        +--+                          
           +-- Lamprotula_cornuumlunae **

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 38 1 1 4 
  +-------- Gonideini   
--+                     
  |  +----- Pronodularia **
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Potomida **

Jeratthitikul et al., 2025 [Publ 4423], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 48 48 3 1 
           +-- Schepmaniini 
  +--------+                
  |        +-- Pseudodontini
--+                         
  |  +-------- Gonideini    
  +--+                      
     |  +----- Potomida **     
     +--+                   
        |  +-- Lamprotula **   
        +--+                
           +-- Pronodularia **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
3 3 1 2 33 3 3 3 
  +-- Pronodularia **
--+               
  +-- Potomida **

Kongim et al., 2023 [Publ 4290], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 12 12 3 3 
  +----------- Chamberlainiini
  |                           
--+        +-- Gonideini      
  |  +-----+                  
  |  |     +-- Pseudodontini  
  +--+                        
     |  +----- Lamprotula **     
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Potomida **

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 1 42 9 2 4 
  +-- Lamprotula **
--+             
  +-- Gonideini

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 28 4 2 2 
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
3 3 1 3 17 2 2 2 
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
3 3 1 2 34 32 5 2 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Potomida **       
     |  +-----+                  
     |  |     +-- Lamprotula **     
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 2 28 5 2 2 
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Potomida **       
     +--+                  
        +-- Lamprotula **

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
3 3 1 3 19 1 1 2 
  +-------- Gonideini   
--+                     
  |  +----- Potomida **    
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Pronodularia **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
3 3 1 3 73 28 4 1 
  +-------- Chamberlainiini
--+                        
  |  +----- Potomida **       
  +--+                     
     |  +-- Lamprotula **     
     +--+                  
        +-- Pronodularia **

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 33 33 4 1 
  +----------------- Rectidentini   
--+                                 
  |  +-------------- Pseudodontini  
  +--+                              
     |  +----------- Gonideini      
     +--+                           
        |  +-------- Chamberlainiini
        +--+                        
           |  +----- Potomida **       
           +--+                     
              |  +-- Lamprotula **     
              +--+                  
                 +-- Pronodularia **

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 2 50 6 2 2 
  +----- Gonideini 
--+                
  |  +-- Potomida **  
  +--+             
     +-- Lamprotula **

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
3 3 1 2 34 32 5 2 
  +-------------------------- Chamberlainiini
  |                                          
--+  +----------------------- Rectidentini   
  |  |                                       
  |  |                    +-- Schepmaniini   
  +--+  +-----------------+                  
     |  |                 +-- Pseudodontini  
     |  |                                    
     +--+              +----- Inversidens    
        |  +-----------+                     
        |  |           |  +-- Potomida **       
        |  |           +--+                  
        +--+              +-- Lamprotula **     
           |                                 
           |  +-------------- Sinosolenaia   
           |  |                              
           +--+           +-- Gonidea        
              |  +--------+                  
              |  |        +-- Microcondylaea 
              +--+                           
                 |  +-------- Parvasolenaia  
                 +--+                        
                    |  +----- Koreosolenaia  
                    +--+                     
                       |  +-- Postolata      
                       +--+                  
                          +-- Ptychorhynchus

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
3 13 0 3 80 32 3 13 
  +-- Lamprotula **    
  |                 
  +-- Potomida **      
  |                 
  +-- Pronodularia **  
  |                 
  +-- Gonidea       
  |                 
  +-- Leguminaia    
  |                 
  +-- Microcondylaea
  |                 
--+-- Amblemini     
  |                 
  +-- Ortmanniana   
  |                 
  +-- Lampsilis     
  |                 
  +-- Cambarunio    
  |                 
  +-- Elliptio      
  |                 
  +-- Pleurobema    
  |                 
  +-- Quadrulini

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
3 6 0 3 33 3 1 6 
  +-- Lamprotula **    
  |                 
  +-- Potomida **      
  |                 
  +-- Pronodularia **  
--+                 
  +-- Gonidea       
  |                 
  +-- Leguminaia    
  |                 
  +-- Microcondylaea

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 2 31 1 1 2 
  +----- Gonideini 
--+                
  |  +-- Potomida **  
  +--+             
     +-- Lamprotula **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 65 5 2 4 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Inversidens    
  |  |                           
  +--+        +-- Obovalis       
     |  +-----+                  
     |  |     +-- Ptychorhynchus 
     +--+                        
        |  +----- Lamprotula **     
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Potomida **

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
3 4 2 2 34 32 5 3 
  +----------------------------- Schepmaniini   
  |                                             
--+  +-------------------------- Pseudodontini  
  |  |                                          
  +--+  +----------------------- Rectidentini   
     |  |                                       
     +--+  +-------------------- Inversidens    
        |  |                                    
        |  |              +----- Lamprotula **     
        +--+  +-----------+                     
           |  |           |  +-- Chamberlainiini
           |  |           +--+                  
           +--+              +-- Potomida **       
              |                                 
              |  +-------------- Sinosolenaia   
              |  |                              
              +--+           +-- Gonidea        
                 |  +--------+                  
                 |  |        +-- Microcondylaea 
                 +--+                           
                    |  +-------- Koreosolenaia  
                    +--+                        
                       |  +----- Parvasolenaia  
                       +--+                     
                          |  +-- Postolata      
                          +--+                  
                             +-- Ptychorhynchus

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iTtTbiriSoTroTroSiD
3 3 1 3 10 2 2 2 
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 65 5 2 4 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Inversidens    
  |  |                           
  +--+        +-- Ptychorhynchus 
     |  +-----+                  
     |  |     +-- Obovalis       
     +--+                        
        |  +----- Lamprotula **     
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Potomida **

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
3 3 1 3 14 1 1 2 
  +-------- Gonideini   
--+                     
  |  +----- Potomida **    
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Pronodularia **

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
3 3 1 2 31 1 1 2 
  +----- Gonideini 
--+                
  |  +-- Potomida **  
  +--+             
     +-- Lamprotula **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
3 3 1 3 73 28 4 3 
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotula **     
  +--+                     
     |  +-- Potomida **       
     +--+                  
        +-- Pronodularia **

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
3 3 1 2 34 32 5 2 
              +-- Schepmaniini   
  +-----------+                  
  |           +-- Pseudodontini  
--+                              
  |  +----------- Gonideini      
  +--+                           
     |  +-------- Rectidentini   
     +--+                        
        |  +----- Chamberlainiini
        +--+                     
           |  +-- Potomida **       
           +--+                  
              +-- Lamprotula **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
3 3 1 2 33 3 3 3 
  +-- Pronodularia **
--+               
  +-- Potomida **

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
3 3 1 3 19 1 1 2 
  +-------- Gonideini   
--+                     
  |  +----- Potomida **    
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Pronodularia **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 2 66 10 3 2 
  +-------- Chamberlainiini
  |                        
--+     +-- Potomida **       
  |  +--+                  
  |  |  +-- Lamprotula **     
  +--+                     
     |  +-- Pseudodontini  
     +--+                  
        +-- Gonideini

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 32 32 4 1 
  +----------------- Rectidentini   
--+                                 
  |  +-------------- Pseudodontini  
  +--+                              
     |  +----------- Gonideini      
     +--+                           
        |  +-------- Chamberlainiini
        +--+                        
           |  +----- Potomida **       
           +--+                     
              |  +-- Pronodularia **   
              +--+                  
                 +-- Lamprotula **

Zhou et al., 2007 [Publ 2658], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 1 15 0 0 3 
1

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 38 1 1 4 
  +-------- Gonideini   
--+                     
  |  +----- Pronodularia **
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Potomida **

Froufe et al., 2015 [Publ 3664], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 9 4 2 2 
  +-------- Gonideini   
--+                     
  |  +----- Potomida **    
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Pronodularia **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 46 5 2 2 
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
3 3 1 3 20 4 2 2 
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
3 3 1 3 36 9 3 2 
  +-------------- Chamberlainiini
--+                              
  |  +----------- Gonideini      
  +--+                           
     |  +-------- Pseudodontini  
     +--+                        
        |  +----- Potomida **       
        +--+                     
           |  +-- Lamprotula **     
           +--+                  
              +-- Pronodularia **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 65 5 2 4 
  +-------- Gonideini   
--+                     
  |  +----- Lamprotula **  
  +--+                  
     |  +-- Potomida **    
     +--+               
        +-- Pronodularia **

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
3 4 2 2 34 32 5 3 
  +----------------------------- Schepmania_nieuwenhuisi__1
  |                                                        
  +----------------------------- Schepmania_nieuwenhuisi__2
--+                                                        
  |  +-------------------------- Pseudodontini             
  |  |                                                     
  +--+  +----------------------- Rectidentini              
     |  |                                                  
     +--+  +-------------------- Inversidens               
        |  |                                               
        |  |              +----- Lamprotula **                
        +--+  +-----------+                                
           |  |           |  +-- Chamberlainiini           
           |  |           +--+                             
           +--+              +-- Potomida **                  
              |                                            
              |  +-------------- Sinosolenaia              
              |  |                                         
              +--+  +----------- Gonidea                   
                 |  |                                      
                 +--+----------- Microcondylaea            
                    |                                      
                    |  +-------- Koreosolenaia             
                    +--+                                   
                       |  +----- Parvasolenaia             
                       +--+                                
                          |  +-- Postolata                 
                          +--+                             
                             +-- Ptychorhynchus

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 50 8 3 2 
  +----------- Chamberlainiini
  |                           
--+        +-- Pseudodontini  
  |  +-----+                  
  |  |     +-- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
3 3 1 2 34 32 5 2 
  +-------------- Chamberlainiini
  |                              
--+  +----------- Rectidentini   
  |  |                           
  +--+        +-- Schepmaniini   
     |  +-----+                  
     |  |     +-- Pseudodontini  
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Potomida **       
           +--+                  
              +-- Lamprotula **

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 2 28 5 2 2 
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Potomida **       
     +--+                  
        +-- Lamprotula **

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 57 57 5 3 
           +-- Chamberlainiini
  +--------+                  
  |        +-- Rectidentini   
  |                           
--+     +----- Pronodularia **   
  |  +--+                     
  |  |  |  +-- Lamprotula **     
  |  |  +--+                  
  +--+     +-- Potomida **       
     |                        
     |  +----- Gonideini      
     +--+                     
        |  +-- Schepmaniini   
        +--+                  
           +-- Pseudodontini

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 11 2 1 2 
  +-------- Gonideini   
--+                     
  |  +----- Potomida **    
  +--+                  
     |  +-- Lamprotula **  
     +--+               
        +-- Pronodularia **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 43 5 2 2 
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Lamprotula **     
        +--+                  
           +-- Pronodularia **

Huang et al., 2019 [Publ 3843], fig. S1

iTtTbiriSoTroTroSiD
3 3 2 1 44 9 2 5 
  +----------------- Chamberlainiini           
  |                                            
  |           +----- Parvasolenaia_rivularis   
--+  +--------+                                
  |  |        |  +-- Parvasolenaia_triangularis
  |  |        +--+                             
  |  |           +-- Ptychorhynchus            
  +--+                                         
     |           +-- Sinosolenaia_carinata     
     |  +--------+                             
     |  |        +-- Lamprotula **                
     +--+                                      
        |  +-------- Sinosolenaia_recognita    
        |  |                                   
        +--+  +----- Sinosolenaia_oleivora__2  
           |  |                                
           +--+  +-- Sinosolenaia_oleivora__1  
              |  |                             
              +--+-- Sinosolenaia_iridinea     
                 |                             
                 +-- Sinosolenaia_oleivora__3

Wu et al., 2024 [Publ 4379], fig. 4_6

iTtTbiriSoTroTroSiD
3 3 1 2 20 0 0 2 
  +-- Potomida **  
--+             
  +-- Lamprotula **

Lopes-Lima et al., 2021 [Publ 4217], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 1 22 3 1 5 
  +-- Potomida ** 
--+            
  +-- Gonideini

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 29 4 2 2 
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Potomida **       
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iTtTbiriSoTroTroSiD
3 3 1 3 10 2 2 4 
  +----------- Chamberlainiini
--+                           
  |  +-------- Gonideini      
  +--+                        
     |  +----- Lamprotula **     
     +--+                     
        |  +-- Potomida **       
        +--+                  
           +-- Pronodularia **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 37 11 3 2 
  +-------------- Rectidentini 
--+                            
  |  +----------- Pseudodontini
  +--+                         
     |  +-------- Gonideini    
     +--+                      
        |  +----- Potomida **     
        +--+                   
           |  +-- Pronodularia ** 
           +--+                
              +-- Lamprotula **

Froufe et al., 2015 [Publ 3664], fig. 1-M

iTtTbiriSoTroTroSiD
2 2 0 2 6 1 1 -- 
  +----- Gonideini   
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 43 0 0 -- 
  +-- Pronodularia **
--+               
  +-- Lamprotula **

Campbell et al., 2005 [Publ 2660], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 100 2 2 -- 
  +----- Gonideini   
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 20 4 2 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Pronodularia **   
     +--+                  
        +-- Lamprotula **

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
2 2 0 2 81 67 3 -- 
  +----------- Pseudodontini
--+                         
  |  +-------- Rectidentini 
  +--+                      
     |  +----- Gonideini    
     +--+                   
        |  +-- Lamprotula **   
        +--+                
           +-- Potomida **

Sano et al., 2017 [Publ 3799], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 31 6 2 -- 
           +-- Obovalis       
  +--------+                  
  |        +-- Ptychorhynchus 
  |                           
--+        +-- Inversidens    
  |  +-----+                  
  |  |     +-- Sinosolenaia   
  +--+                        
     |  +----- Chamberlainiini
     +--+                     
        |  +-- Pronodularia **   
        +--+                  
           +-- Lamprotula **

Zhou et al., 2017 [Publ 3788], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 14 5 2 -- 
  +----- Chamberlainiini
--+                     
  |  +-- Lamprotula **     
  +--+                  
     +-- Gonideini

Konopleva et al., 2021 [Publ 4213], fig. 1

iTtTbiriSoTroTroSiD
2 27 0 2 25 25 2 28 
  +-- Lamprotula **   
  |                
  +-- Leguminaia   
  |                
--+-- Potomida **     
  |                
  +-- Gonidea      
  |                
  +-- Pseudodontini

Ouyang et al., 2015 [Publ 4493], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 29 4 2 -- 
1

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
2 2 0 2 21 10 4 -- 
  +-------------- Chamberlainiini
--+                              
  |  +----------- Rectidentini   
  +--+                           
     |  +-------- Pseudodontini  
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Pronodularia **   
           +--+                  
              +-- Potomida **

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
2 9 0 2 13 0 0 10 
  +-- Aculamprotulina
  |                  
  +-- Lamprotula **     
  |                  
  +-- Lampsilini     
  |                  
  +-- Potomida **       
  |                  
--+-- Unio_pictorum  
  |                  
  +-- Anodontini     
  |                  
  +-- Unio_ravoisieri
  |                  
  +-- Lamellidentini 
  |                  
  +-- Pleurobemini

Liu et al., 2024 [Publ 4380], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 1 10 10 1 -- 
  +-- Lamprotula **
--+             
  +-- Gonideini

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
2 2 0 2 27 0 0 -- 
  +-- Potomida **  
--+             
  +-- Lamprotula **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 35 9 3 -- 
  +-------- Chamberlainiini
  |                        
--+     +-- Lamprotula **     
  |  +--+                  
  |  |  +-- Potomida **       
  +--+                     
     |  +-- Pseudodontini  
     +--+                  
        +-- Gonideini

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 39 0 0 -- 
  +-- Pronodularia **
--+               
  +-- Lamprotula **

Fonseca et al., 2016 [Publ 3775], fig. 1-M

iTtTbiriSoTroTroSiD
2 2 0 2 7 1 1 -- 
  +----- Gonideini   
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Zhang et al., 2024 [Publ 4397], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 6 6 1 -- 
  +-- Lamprotula **
--+             
  +-- Gonideini

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 39 0 0 -- 
  +-- Pronodularia **
--+               
  +-- Lamprotula **

Froufe et al., 2019 [Publ 3936], fig. 3-M

iTtTbiriSoTroTroSiD
2 2 0 2 17 6 3 -- 
  +-------- Chamberlainiini
  |                        
--+     +-- Potomida **       
  |  +--+                  
  |  |  +-- Pronodularia **   
  +--+                     
     |  +-- Gonideini      
     +--+                  
        +-- Pseudodontini

Bolotov et al., 2022 [Publ 4250], fig. 8

iTtTbiriSoTroTroSiD
2 7 0 2 47 21 3 7 
  +-- Gibbosula    
  |                
  +-- Margaritifera
  |                
  +-- Sinosolenaia 
  |                
--+-- Leguminaia   
  |                
  +-- Gonidea      
  |                
  +-- Lamprotula **   
  |                
  +-- Potomida **

Zhou et al., 2016 [Publ 3710], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 18 4 2 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Lamprotula **     
     +--+                  
        +-- Pronodularia **

Froufe et al., 2019 [Publ 3936], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 17 6 3 -- 
  +-------- Chamberlainiini
  |                        
--+     +-- Potomida **       
  |  +--+                  
  |  |  +-- Pronodularia **   
  +--+                     
     |  +-- Gonideini      
     +--+                  
        +-- Pseudodontini

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 43 0 0 -- 
  +-- Pronodularia **
--+               
  +-- Lamprotula **

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 38 5 2 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Lamprotula **     
     +--+                  
        +-- Potomida **

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
2 9 0 2 14 0 0 10 
  +----------------------- Lamprotula **     
--+                                       
  |  +-------------------- Potomida **       
  +--+                                    
     |  +----------------- Lampsilini     
     +--+                                 
        |  +-------------- Pleurobemini   
        +--+                              
           |  +----------- Aculamprotulina
           +--+                           
              |  +-------- Unio_pictorum  
              +--+                        
                 |  +----- Anodontini     
                 +--+                     
                    |  +-- Unio_ravoisieri
                    +--+                  
                       +-- Lamellidentini

Lopes-Lima et al., 2020 [Publ 4080], fig. 14

iTtTbiriSoTroTroSiD
2 2 0 1 4 2 1 -- 
  +-- Gonideini 
--+             
  +-- Lamprotula **

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
2 2 0 2 13 0 0 -- 
  +-- Lamprotula **
--+             
  +-- Potomida **

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 39 0 0 -- 
  +-- Pronodularia **
--+               
  +-- Lamprotula **

Campbell et al., 2005 [Publ 2660], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 100 2 2 -- 
  +----- Gonideini   
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
2 34 0 1 32 9 3 34 
                       +----- Prolasmidonta           
                    +--+                              
                    |  |  +-- Alasmidonta             
                    |  +--+                           
  +-----------------+     +-- Anemina                 
  |                 |                                 
  |                 |  +----- Pseudodontini           
  |                 +--+                              
  |                    |  +-- Lamprotula_caveata **      
  |                    +--+                           
  |                       +-- Chamberlainia           
  |                                                   
  |                    +----- Oxynaiina               
  |              +-----+                              
  |              |     |  +-- Lepidodesmini           
--+              |     +--+                           
  |              |        +-- Aculamprotulina         
  |  +-----------+                                    
  |  |           |     +----- Platynaias              
  |  |           |  +--+                              
  |  |           |  |  |  +-- Utterbackia_imbecillis  
  |  |           |  |  +--+                           
  |  |           +--+     +-- Pyganodon               
  |  |              |                                 
  |  |              |  +----- Sinanodonta             
  |  |              +--+                              
  |  |                 |  +-- Cristaria               
  +--+                 +--+                           
     |                    +-- Anodontina              
     |                                                
     |                    +-- Utterbackia_peninsularis
     |              +-----+                           
     |              |     +-- Potamilus_alatus        
     |  +-----------+                                 
     |  |           |  +----- Lamprotula_leaii **        
     |  |           +--+                              
     |  |              |  +-- Sinohyriopsis           
     +--+              +--+                           
        |                 +-- Gonideini               
        |                                             
        |  +----------------- Gibbosula               
        +--+                                          
           |  +-------------- Quadrulini              
           +--+                                       
              |  +----------- Toxolasma               
              +--+                                    
                 |  +-------- Lampsilis_ornata        
                 +--+                                 
                    |  +----- Potamilus_leptodon      
                    +--+                              
                       |  +-- Lampsilis_siliquoidea   
                       +--+                           
                          +-- Lampsilis_powellii

Ouyang et al., 2015 [Publ 4493], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 29 4 2 -- 
  +----- Lamprotula **     
--+                     
  |  +-- Chamberlainiini
  +--+                  
     +-- Gonideini

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

iTtTbiriSoTroTroSiD
2 2 0 2 8 1 1 -- 
  +----- Gonideini   
--+                  
  |  +-- Potomida **    
  +--+               
     +-- Pronodularia **

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 39 0 0 -- 
  +-- Pronodularia **
--+               
  +-- Lamprotula **

Hou et al., 2024 [Publ 4400], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 13 0 0 -- 
1

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 19 4 2 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Lamprotula **     
     +--+                  
        +-- Pronodularia **

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 32 6 2 -- 
  +-------- Gonideini      
--+                        
  |  +----- Chamberlainiini
  +--+                     
     |  +-- Lamprotula **     
     +--+                  
        +-- Pronodularia **

Amaro et al., 2017 [Publ 3791], fig. 3A-M

iTtTbiriSoTroTroSiD
1 1 0 1 6 1 1 -- 
  +-- Pronodularia **
--+               
  +-- Gonideini

Chen et al., 2012 [Publ 3624], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 6 1 1 -- 
  +-- Chamberlainiini
--+                  
  +-- Lamprotula **

Khalloufi et al., 2011 [Publ 2748], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Potomida_littoralis **

Machordom et al., 2003 [Publ 2688], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 3 0 0 -- 
-- Potomida_littoralis **

Välilä et al., 2017 [Publ 3793], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 2 0 0 -- 
-- Potomida_littoralis **

Guerra et al., 2017 [Publ 3772], fig. 2a-M

iTtTbiriSoTroTroSiD
1 1 0 1 7 1 1 -- 
  +-- Gonideini   
--+               
  +-- Pronodularia **

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
1 1 0 1 24 4 2 -- 
  +----- Chamberlainiini
--+                     
  |  +-- Pronodularia **   
  +--+                  
     +-- Gonideini

Lopes-Lima et al., 2017 [Publ 3734], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 16 1 1 -- 
  +-- Potomida ** 
--+            
  +-- Gonideini

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Potomida_littoralis **

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 33 3 2 -- 
-- Lamprotula_leaii **

Kano et al., 2019 [Publ 4194], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 9 2 1 -- 
  +----- Inversidens 
--+                  
  |  +-- Obovalis    
  +--+               
     +-- Pronodularia **

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 18 1 1 -- 
  +-- Chamberlainiini
--+                  
  +-- Lamprotula **

Pfeiffer et al., 2021 [Publ 4126], fig. 9

iTtTbiriSoTroTroSiD
1 1 0 1 28 28 1 -- 
  +-- Lamprotula **  
--+               
  +-- Rectidentini

Chapman et al., 2008 [Publ 2744], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 20 0 0 -- 
-- Pronodularia_japanensis **

Reis et al., 2013 [Publ 3580], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Potomida_littoralis **

Araujo et al., 2009 [Publ 2589], fig. 7C

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Potomida_littoralis **

Amaro et al., 2017 [Publ 3791], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 9 1 1 -- 
  +-- Pronodularia **
--+               
  +-- Gonideini

Reis et al., 2013 [Publ 3580], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Potomida_littoralis **

Amaro et al., 2017 [Publ 3791], fig. 3A

iTtTbiriSoTroTroSiD
1 1 0 1 11 1 1 -- 
  +-- Pronodularia **
--+               
  +-- Gonideini

Chapman et al., 2008 [Publ 2667], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 20 0 0 -- 
-- Pronodularia_japanensis **

Amaro et al., 2017 [Publ 3791], fig. 3B

iTtTbiriSoTroTroSiD
1 1 0 1 11 1 1 -- 
  +-- Pronodularia **
--+               
  +-- Gonideini

Amaro et al., 2017 [Publ 3791], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 15 1 1 -- 
  +-- Pronodularia **
--+               
  +-- Gonideini

Zhang et al., 2004 [Publ 2735], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 13 1 1 -- 
  +-- Lamprotula **     
--+                  
  +-- Chamberlainiini

Xue et al., 2016 [Publ 3659], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 14 1 1 -- 
  +-- Chamberlainiini
--+                  
  +-- Lamprotula **

Goncalves et al., 2022 [Publ 4253], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 6 6 1 -- 
  +-- Lamprotula **     
--+                  
  +-- Chamberlainiini

Prié & Puillandre, 2014 [Publ 3529], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 16 0 0 -- 
-- Potomida_littoralis **

Walker et al., 2006 [Publ 2666], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 2 2 -- 
-- Pronodularia_japanensis **

Breton et al., 2011 [Publ 3376], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 36 1 1 -- 
-- Pronodularia_japanensis **

Nagel et al., 1998 [Publ 1946], fig. 9

iTtTbiriSoTroTroSiD
1 1 0 1 5 1 1 -- 
-- Potomida **

Pfarrer et al., 2022 [Publ 4305], fig. 2-5

iTtTbiriSoTroTroSiD
1 1 0 1 11 1 1 -- 
  +-- Potomida ** 
--+            
  +-- Gonideini

Plazzi et al., 2011 [Publ 3676], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 17 2 1 -- 
-- Pronodularia_japanensis **

Huang et al., 2013 [Publ 3629], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 5 1 1 -- 
  +-- Gonideini   
--+               
  +-- Pronodularia **

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 3 2 -- 
-- Lamprotula_leaii **

Araujo et al., 2009 [Publ 2589], fig. 7B

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Potomida_littoralis **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 27 0 0 -- 
-- Potomida_littoralis **

Sano et al., 2017 [Publ 3799], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 20 3 2 -- 
  +-------- Obovalis       
--+                        
  |  +----- Inversidens    
  +--+                     
     |  +-- Pronodularia **   
     +--+                  
        +-- Chamberlainiini

Sheng et al., 2014 [Publ 3704], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 7 2 1 -- 
  +-- Lamprotula **     
--+                  
  +-- Chamberlainiini

Zieritz et al., 2020 [Publ 4102], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 30 30 4 -- 
  +----------- Chamberlainiini
--+                           
  |  +-------- Lamprotula **     
  +--+                        
     |  +----- Gonideini      
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Rectidentini

Nagel et al., 1998 [Publ 1946], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 8 1 1 -- 
-- Potomida_littoralis **

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 11 3 2 -- 
  +-------- Obovalis       
--+                        
  |  +----- Inversidens    
  +--+                     
     |  +-- Chamberlainiini
     +--+                  
        +-- Pronodularia **

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
1 1 0 1 24 4 2 -- 
  +----- Chamberlainiini
--+                     
  |  +-- Pronodularia **   
  +--+                  
     +-- Gonideini

Plazzi & Passamonti, 2010 [Publ 3354], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 1 0 0 -- 
-- Pronodularia_japanensis **

Wen et al., 2017 [Publ 3795], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 15 0 0 -- 
-- Lamprotula_leaii **

Zhang et al., 2013 [Publ 4214], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 3 0 0 -- 
-- Lamprotula_leaii **

Smith, 2021 [Publ 4331], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 19 2 2 -- 
  +----- Chamberlainiini
--+                     
  |  +-- Rectidentini   
  +--+                  
     +-- Lamprotula **

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 27 0 0 -- 
-- Potomida_littoralis **

Shirai et al., 2010 [Publ 3353], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 2 2 1 -- 
  +-- Pronodularia **   
--+                  
  +-- Chamberlainiini

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 31 27 5 -- 
  +-------------- Chamberlainiini
--+                              
  |  +----------- Potomida **       
  +--+                           
     |  +-------- Rectidentini   
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini

Amaro et al., 2017 [Publ 3791], fig. 3B-M

iTtTbiriSoTroTroSiD
1 1 0 1 6 1 1 -- 
  +-- Pronodularia **
--+               
  +-- Gonideini

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 16 4 -- 
  +----------- Chamberlainiini
--+                           
  |  +-------- Rectidentini   
  +--+                        
     |  +----- Lamprotula **     
     +--+                     
        |  +-- Gonideini      
        +--+                  
           +-- Pseudodontini

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 23 1 1 -- 
  +-- Gonideini 
--+             
  +-- Lamprotula **

Ouyang et al., 2011 [Publ 3408], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 12 3 1 -- 
  +-- Lamprotula **
--+             
  +-- Gonideini

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 16 4 -- 
  +----------- Chamberlainiini
--+                           
  |  +-------- Rectidentini   
  +--+                        
     |  +----- Pseudodontini  
     +--+                     
        |  +-- Gonideini      
        +--+                  
           +-- Lamprotula **

Jiang et al., 2010 [Publ 3438], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 3 1 1 -- 
  +-- Pronodularia **   
--+                  
  +-- Chamberlainiini

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 23 1 1 -- 
  +-- Gonideini 
--+             
  +-- Lamprotula **

Araujo et al., 2016 [Publ 3746], fig. 1_2b

iTtTbiriSoTroTroSiD
1 1 0 1 12 0 0 -- 
-- Potomida_littoralis **

Chase et al., 2018 [Publ 3861], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Lamprotula_leaii **

Huang et al., 2013 [Publ 3629], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 15 3 2 -- 
  +----- Chamberlainiini
--+                     
  |  +-- Gonideini      
  +--+                  
     +-- Pronodularia **

Guerra et al., 2017 [Publ 3772], fig. 2b-M

iTtTbiriSoTroTroSiD
1 1 0 1 7 1 1 -- 
  +-- Gonideini   
--+               
  +-- Pronodularia **

Froufe et al., 2017 [Publ 3802], fig. 7

iTtTbiriSoTroTroSiD
1 1 0 1 9 1 1 -- 
  +-- Gonideini
--+            
  +-- Potomida **

Araujo et al., 2009 [Publ 2589], fig. 7A

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Potomida_littoralis **

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Potomida_littoralis **

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iTtTbiriSoTroTroSiD
1 1 0 1 24 1 1 -- 
-- Pronodularia_japanensis **

Machordom et al., 2003 [Publ 2688], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 2 0 0 -- 
-- Potomida_littoralis **

Chase et al., 2018 [Publ 3861], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Lamprotula_leaii **

Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3

iTtTbiriSoTroTroSiD
1 1 0 1 3 1 1 -- 
  +-- Chamberlainiini
--+                  
  +-- Pronodularia **

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 22 1 1 -- 
  +-- Gonideini 
--+             
  +-- Lamprotula **

Chen et al., 2025 [Publ 4455], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 25 0 0 -- 
-- Potomida_littoralis **

Soroka, 2005 [Publ 2726], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Pronodularia_japanensis **