Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae

tribe LAMPSILINI Ihering, 1901: cladomics

32 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 158 (183 total)
number of subtaxa (rank: genus) = 32
analyzed in 285 cladograms in 168 publications
maximum number of terminals in an analysis = 93 (mean = 5.88, median = 3)
maximum number of subtaxa (genus) = 27 (mean = 3.86, median = 3)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 206
maximum number of included terminals = 93, (mean = 7.75 median = 5)
maximum number of subtaxa (genus) = 27 (mean = 4.96, median = 4)
proportion of cladograms that support basic monophyly = 0.78

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 177
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 93 (mean = 8.07, median = 5)
maximum number of subtaxa (genus) = 27 (mean = 5.33, median = 4)
proportion of cladograms that support rigorous monophyly = 0.74

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Lampsilini cladogram distances
PCoA Ordination

NJ Tree of Lampsilini cladogram distances
NJ Tree

Cladograms

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
0.636 93 97 113 27 540 98 4 **
Cladomics cladogram QUADRULINI PLEUROBEMINI Amblema Nephronaias Psoronaias Reginaia Popenaias Cyrtonaias Glebula Paetulunio Toxolasma parvum Toxolasma lividum 2 Toxolasma cylindrellus Toxolasma lividum 1 Toxolasma texasiense 2 Toxolasma glans Toxolasma paulum Toxolasma texasiense 1 Toxolasma 4 Toxolasma granulatum Toxolasma 2 Toxolasma 3 Toxolasma pullus Toxolasma germanum Toxolasma 1 Obliquaria Plectomerus Potamilus inflatus Potamilus ohiensis Potamilus amphichaenus Potamilus streckersoni Potamilus fragilis 2 Potamilus leptodon Potamilus fragilis 1 Potamilus metnecktayi Potamilus alatus 3 Potamilus alatus 2 Potamilus purpuratus 2 Potamilus alatus 1 Potamilus purpuratus 1 Villosa villosa Atlanticoncha Cambarunio nebulosus Ligumia Leaunio umbrans Cambarunio taeniatus Cambarunio iris Cambarunio simus Hamiota Pachynaias Sagittunio Ellipsaria Potamilus capax Truncilla Medionidus acutissimus Lemiox Medionidus penicillatus Ptychobranchus Cyprogenia aberti 2 Dromus Cyprogenia stegaria 2 Cyprogenia aberti 1 Cyprogenia stegaria 1 Villosa delumbis Villosa vibex Obovaria olivaria Venustaconcha Epioblasma Obovaria retusa Obovaria choctawensis Obovaria unicolor Obovaria subrotunda Mesonaias Obovaria arkansasensis Lampsilis splendida Lampsilis bracteata Lampsilis fasciola Ortmanniana pectorosa Lampsilis ornata Lampsilis ovata 2 Lampsilis dolabraeformis Lampsilis cardium 2 Lampsilis cardium 1 Lampsilis satura Lampsilis cariosa 2 Lampsilis ovata 1 Lampsilis cariosa 1 Ortmanniana ligamentina 2 Lampsilis abrupta Lampsilis higginsii 2 Ortmanniana ligamentina 1 Lampsilis higginsii 1 Medionidus conradicus Leaunio lienosus Leaunio vanuxemensis Lampsilis floridensis Lampsilis teres Lampsilis siliquoidea 1 Lampsilis siliquoidea 2 Lampsilis powellii 1 Lampsilis powellii 2 Lampsilis sietmani Lampsilis straminea 2 Lampsilis virescens Lampsilis hydiana 2 Lampsilis hydiana 1 Lampsilis bergmanni Lampsilis straminea 1

Campbell et al., 2005 [Publ 2660], fig. 1

cst iT tT bi riS oT roT roS iD
0.857 43 43 46 23 59 47 4 130.0
Cladomics cladogram QUADRULINI PLEUROBEMINI Plectomerus Reginaia POPENAIADINI Amblema Toxolasma Obliquaria Cyrtonaias Glebula Potamilus Medionidus acutissimus Lemiox Dromus Cyprogenia Sagittunio Medionidus conradicus Ptychobranchus Venustaconcha ellipsiformis 2 Ellipsaria Truncilla Villosa Obovaria olivaria Venustaconcha ellipsiformis 1 Epioblasma Obovaria unicolor Obovaria arkansasensis Obovaria subrotunda Hamiota Lampsilis cardium Lampsilis ovata Ortmanniana pectorosa Lampsilis ornata Ligumia Cambarunio Lampsilis teres Ortmanniana ligamentina Lampsilis siliquoidea Leaunio

Campbell et al., 2005 [Publ 2660], fig. 2

cst iT tT bi riS oT roT roS iD
0.796 43 44 37 23 59 47 4 115.0
Cladomics cladogram QUADRULINI PLEUROBEMINI POPENAIADINI Amblema Reginaia Plectomerus Toxolasma Obliquaria Cyrtonaias Glebula Sagittunio Leaunio Potamilus Hamiota Ellipsaria Truncilla Lampsilis teres Ortmanniana ligamentina Lampsilis siliquoidea Villosa Ligumia Cambarunio Ortmanniana pectorosa Lampsilis ornata Lampsilis cardium Lampsilis ovata Obovaria olivaria Venustaconcha ellipsiformis 2 Epioblasma Obovaria unicolor Obovaria arkansasensis Obovaria subrotunda Ptychobranchus Venustaconcha ellipsiformis 1 Medionidus conradicus Lemiox Medionidus acutissimus Dromus Cyprogenia

Zanatta & Murphy, 2006 [Publ 2672], fig. 1

cst iT tT bi riS oT roT roS iD
0.912 48 48 45 23 8 6 3 120.0
Cladomics cladogram PLEUROBEMINI AMBLEMINI POPENAIADINI Cyrtonaias Glebula Paetulunio Obliquaria Toxolasma Sagittunio Potamilus Medionidus conradicus Ptychobranchus Medionidus acutissimus Lemiox Dromus Cyprogenia Lampsilis teres Truncilla Ellipsaria Leaunio Hamiota Lampsilis fasciola Ortmanniana pectorosa Lampsilis ornata Lampsilis ovata Lampsilis cardium Venustaconcha Obovaria Epioblasma Lampsilis siliquoidea Ortmanniana ligamentina Cambarunio Ligumia

Zanatta & Murphy, 2006 [Publ 2672], fig. 2

cst iT tT bi riS oT roT roS iD
0.886 48 48 42 23 9 7 3 117.0
Cladomics cladogram PLEUROBEMINI AMBLEMINI POPENAIADINI Paetulunio Toxolasma Obliquaria Glebula Cyrtonaias Ptychobranchus Dromus Cyprogenia Medionidus conradicus Medionidus acutissimus Lemiox Sagittunio Potamilus Truncilla Ellipsaria Ortmanniana pectorosa Lampsilis fasciola Lampsilis ornata Lampsilis ovata Lampsilis cardium Ligumia Cambarunio Lampsilis teres Hamiota Leaunio Ortmanniana ligamentina Lampsilis siliquoidea Obovaria olivaria Venustaconcha Epioblasma Obovaria unicolor Obovaria subrotunda Obovaria arkansasensis

Smith et al., 2019 [Publ 3951], fig. 2

cst iT tT bi riS oT roT roS iD
0.903 36 37 36 20 5 5 3 122.0
Cladomics cladogram QUADRULINI Amblema Reginaia Plectomerus Toxolasma POPENAIADINI Glebula Cyrtonaias Obliquaria Medionidus Lemiox Cyprogenia Ptychobranchus Potamilus Atlanticoncha Ellipsaria Truncilla Hamiota Villosa Ligumia Venustaconcha Obovaria Epioblasma Ortmanniana pectorosa Ortmanniana ligamentina Lampsilis

Burlakova et al., 2019 [Publ 3899], fig. 1

cst iT tT bi riS oT roT roS iD
0.906 26 30 18 18 7 7 4 115.0
Cladomics cladogram QUADRULINI Plectomerus POPENAIADINI PLEUROBEMINI Cyrtonaias Glebula Reginaia Obliquaria Toxolasma Potamilus Ellipsaria Truncilla Medionidus Lemiox Dromus Cambarunio Leaunio Villosa Sagittunio Lampsilis ovata Venustaconcha Epioblasma Obovaria Lampsilis teres Lampsilis siliquoidea Lampsilis hydiana Lampsilis straminea

Pfeiffer et al., 2019 [Publ 3866], fig. 1

cst iT tT bi riS oT roT roS iD
0.938 12 12 10 12 19 18 4 119.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI POPENAIADINI Cyrtonaias Glebula Potamilus Truncilla Ptychobranchus Pachynaias Mesonaias Lampsilis Ortmanniana Cambarunio Epioblasma Ligumia

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
0.893 14 15 10 12 187 11 3 118.0
Cladomics cladogram QUADRULINI PLEUROBEMINI Amblema Toxolasma Obliquaria Plectomerus Potamilus leptodon Potamilus alatus Venustaconcha Leaunio Ellipsaria Truncilla Medionidus Ptychobranchus Cambarunio Ortmanniana Lampsilis

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
0.893 14 15 13 12 185 11 3 119.0
Cladomics cladogram QUADRULINI PLEUROBEMINI Amblema Toxolasma Obliquaria Plectomerus Potamilus leptodon Potamilus alatus Medionidus Ptychobranchus Ellipsaria Truncilla Leaunio Venustaconcha Cambarunio Ortmanniana Lampsilis

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.821 14 21 8 12 187 11 3 122.0
Cladomics cladogram Chamberlainia Megalonaias Uniomerus QUADRULINA Sinohyriopsis LAMPROTULINI GONIDEINI PSEUDODONTINI Rectidentini Amblema Toxolasma PLEUROBEMINI Obliquaria Plectomerus Ellipsaria Potamilus leptodon Medionidus Potamilus alatus Ptychobranchus Truncilla Venustaconcha Cambarunio Leaunio Ortmanniana Lampsilis ACUTICOSTINI Alasmidonta Anodonta anatina Anemina Anodonta cygnea Cristaria Brachyanodon Lasmigona LEPIDODESMINI Pseudanodonta Pyganodon Simpsonaias Strophitus Utterbackia Schistodesmus UNIONINA ACULAMPROTULINA LANCEOLARIINI Pletholophus Sinanodonta Cuneopsis celtiformis Tchangsinaia Cuneopsis rufescens Nodularia Cuneopsis szechenyii Cuneopsis heudei

Inoue et al., 2020 [Publ 4128], fig. 2

cst iT tT bi riS oT roT roS iD
0.957 12 12 5 12 11 11 3 114.0
Cladomics cladogram QUADRULINI Reginaia Amblema POPENAIADINI Glebula Cyrtonaias Potamilus Ptychobranchus Truncilla Ortmanniana Epioblasma Lampsilis Ligumia Cambarunio Pachynaias Mesonaias

Chapman et al., 2008 [Publ 2667], fig. 1

cst iT tT bi riS oT roT roS iD
0.920 13 13 11 11 8 7 4 114.0
Cladomics cladogram QUADRULINI PLEUROBEMINI Amblema Plectomerus POPENAIADINI Toxolasma Obliquaria Cyrtonaias Glebula Ptychobranchus Lemiox Venustaconcha Obovaria Hamiota Lampsilis ovata Ortmanniana Lampsilis straminea Lampsilis hydiana

Chapman et al., 2008 [Publ 2744], fig. 2

cst iT tT bi riS oT roT roS iD
0.920 13 13 11 11 8 7 4 114.0
Cladomics cladogram QUADRULINI PLEUROBEMINI Amblema Plectomerus POPENAIADINI Toxolasma Obliquaria Cyrtonaias Glebula Ptychobranchus Lemiox Venustaconcha Obovaria Hamiota Lampsilis ovata Ortmanniana Lampsilis straminea Lampsilis hydiana

Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2

cst iT tT bi riS oT roT roS iD
0.806 15 15 9 11 58 58 4 122.0
Cladomics cladogram QUADRULINI PLEUROBEMINI Plectomerus Reginaia POPENAIADINI Amblema Potamilus alatus Potamilus capax Glebula Toxolasma Paetulunio Dromus Lemiox Lampsilis ovata Obliquaria Lampsilis virescens Epioblasma Obovaria arkansasensis Obovaria olivaria Venustaconcha

Campbell & Lydeard, 2012 [Publ 3309], fig. 3

cst iT tT bi riS oT roT roS iD
0.833 15 16 14 11 58 58 4 121.0
Cladomics cladogram QUADRULINI PLEUROBEMINI Amblema Reginaia POPENAIADINI Glebula Paetulunio Toxolasma Plectomerus Obliquaria Potamilus Dromus Lemiox Lampsilis ovata Lampsilis virescens Epioblasma Obovaria arkansasensis Obovaria olivaria Venustaconcha

Inoue et al., 2020 [Publ 4128], fig. 3

cst iT tT bi riS oT roT roS iD
0.955 11 11 9 11 11 11 3 124.0
Cladomics cladogram QUADRULINI Reginaia Amblema POPENAIADINI Cyrtonaias Ptychobranchus Ligumia Potamilus Pachynaias Epioblasma Truncilla Mesonaias Cambarunio Ortmanniana Lampsilis

Boyer et al., 2011 [Publ 2718], fig. 3

cst iT tT bi riS oT roT roS iD
0.852 16 16 9 10 19 13 3 114.0
Cladomics cladogram Reginaia PLEUROBEMINI QUADRULINI Amblema Toxolasma Obliquaria Obovaria Lampsilis cardium Potamilus Epioblasma Lampsilis siliquoidea Ortmanniana Lampsilis higginsii Ligumia Lampsilis teres Ellipsaria Truncilla

Boyer et al., 2011 [Publ 2718], fig. 4

cst iT tT bi riS oT roT roS iD
0.815 16 22 18 10 22 13 3 117.0
Cladomics cladogram QUADRULINA Megalonaias Toxolasma Amblema PLEUROBEMINI Reginaia Obliquaria Potamilus Ellipsaria Truncilla Lampsilis cardium Ligumia Obovaria Epioblasma Lampsilis teres Lampsilis siliquoidea Ortmanniana Lampsilis higginsii

Grobler et al., 2011 [Publ 3359], fig. 2

cst iT tT bi riS oT roT roS iD
0.929 16 16 12 10 1 1 1 117.0
Cladomics cladogram AMBLEMINI Lemiox Medionidus Dromus Cyprogenia Potamilus Lampsilis siliquoidea Lampsilis higginsii Ortmanniana Lampsilis fasciola Lampsilis cardium Lampsilis ovata Cambarunio Ligumia Epioblasma

Szumowski et al., 2012 [Publ 3569], fig. 4

cst iT tT bi riS oT roT roS iD
0.929 15 15 11 10 1 1 1 120.0
Cladomics cladogram AMBLEMINI Ellipsaria Truncilla Obliquaria Potamilus Lampsilis ornata Lampsilis cardium Lampsilis ovata Lampsilis teres Lampsilis siliquoidea Ortmanniana Lampsilis higginsii Hamiota Epioblasma Cambarunio Ligumia

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

cst iT tT bi riS oT roT roS iD
0.950 12 12 6 9 18 5 3 119.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI Ligumia Truncilla Ptychobranchus Sagittunio Lampsilis siliquoidea Epioblasma Ortmanniana Cambarunio Leaunio Lampsilis cardium Lampsilis fasciola

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

cst iT tT bi riS oT roT roS iD
0.950 12 12 10 9 18 5 3 123.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI Truncilla Ligumia Ptychobranchus Sagittunio Lampsilis siliquoidea Ortmanniana Lampsilis cardium Lampsilis fasciola Epioblasma Cambarunio Leaunio

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

cst iT tT bi riS oT roT roS iD
0.758 11 12 0 8 94 72 4 117.0
Cladomics cladogram Reginaia Amblema QUADRULINI PLEUROBEMINI POPENAIADINI Dromus Epioblasma Glebula Lampsilis ovata Lampsilis virescens Lemiox Obliquaria Obovaria olivaria Obovaria unicolor Plectomerus Potamilus alatus Potamilus capax

Smith et al., 2020 [Publ 4129], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 20 20 18 8 8 8 4 119.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI POPENAIADINI Obliquaria Cyrtonaias Pachynaias Lampsilis Ellipsaria Truncilla Atlanticoncha Potamilus

Smith et al., 2020 [Publ 4129], fig. 2-3

cst iT tT bi riS oT roT roS iD
1.000 20 20 18 8 8 8 4 119.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI POPENAIADINI Obliquaria Cyrtonaias Pachynaias Lampsilis Ellipsaria Truncilla Atlanticoncha Potamilus

Krebs et al., 2003 [Publ 2691], fig. 3

cst iT tT bi riS oT roT roS iD
0.708 11 12 10 8 22 10 3 122.0
Cladomics cladogram Megalonaias QUADRULINA Elliptoideus Pleurobema Uniomerus Fusconaia Eurynia Amblema Toxolasma Plectomerus Obliquaria Ptychobranchus Potamilus fragilis Medionidus Villosa Ellipsaria Lampsilis teres Lampsilis siliquoidea Potamilus purpuratus Potamilus alatus

Graf & Cummings, 2006 [Publ 1968], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 8 8 4 8 39 6 3 117.0
Cladomics cladogram QUADRULINI AMBLEMINI PLEUROBEMINI Obliquaria Truncilla Ptychobranchus Ligumia Cambarunio Epioblasma Lampsilis Ortmanniana

Hoeh et al., 2009 [Publ 2716], fig. 3

cst iT tT bi riS oT roT roS iD
0.947 8 14 12 8 39 6 3 126.0
Cladomics cladogram GONIDEINI PSEUDODONTINI AMBLEMINI PLEUROBEMINI Obliquaria QUADRULINI Ptychobranchus Truncilla Ortmanniana Lampsilis Cambarunio Ligumia Epioblasma

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

cst iT tT bi riS oT roT roS iD
0.833 8 11 9 8 17 5 3 127.0
Cladomics cladogram GONIDEINI Eurynia Cambarunio Ptychobranchus AMBLEMINI Pleurobema Truncilla Lampsilis Epioblasma Ligumia Obliquaria Ortmanniana

Breton et al., 2011 [Publ 3376], fig. 3

cst iT tT bi riS oT roT roS iD
0.900 13 13 11 8 25 6 3 122.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI Toxolasma Ellipsaria Truncilla Lemiox Potamilus Lampsilis Cambarunio Venustaconcha

Chase et al., 2018 [Publ 3847], fig. 4

cst iT tT bi riS oT roT roS iD
0.947 13 13 11 8 26 5 3 116.0
Cladomics cladogram Echyridella AMBLEMINI PLEUROBEMINI QUADRULINI Toxolasma Venustaconcha Lemiox Lampsilis Potamilus Cambarunio Ellipsaria Truncilla

Hoeh et al., 2009 [Publ 2716], fig. 4

cst iT tT bi riS oT roT roS iD
0.941 8 13 10 8 38 5 2 126.0
Cladomics cladogram GONIDEINI PLEUROBEMINI Obliquaria QUADRULINI Ptychobranchus Truncilla Ortmanniana Lampsilis Cambarunio Ligumia Epioblasma

Lydeard et al., 1996 [Publ 2656], fig. 3

cst iT tT bi riS oT roT roS iD
0.773 9 10 7 7 18 9 3 119.0
Cladomics cladogram Megalonaias QUADRULINA Elliptoideus Eurynia Pleurobema Uniomerus Amblema Plectomerus Obliquaria Potamilus fragilis Ptychobranchus Medionidus Potamilus purpuratus Lampsilis Ellipsaria Villosa

Lydeard et al., 1996 [Publ 2656], fig. 4

cst iT tT bi riS oT roT roS iD
0.818 9 9 7 7 19 9 3 121.0
Cladomics cladogram Megalonaias QUADRULINA Elliptoideus Uniomerus Eurynia Pleurobema Amblema Plectomerus Obliquaria Medionidus Ptychobranchus Potamilus fragilis Potamilus purpuratus Ellipsaria Villosa Lampsilis

Huang et al., 2002 [Publ 2657], fig. 1

cst iT tT bi riS oT roT roS iD
0.722 8 9 7 7 26 6 3 121.0
Cladomics cladogram Megalonaias Elliptoideus Uniomerus Pleurobema Amblema Obliquaria Plectomerus Medionidus Potamilus fragilis Ptychobranchus Potamilus purpuratus Lampsilis Villosa Ellipsaria

Huang et al., 2002 [Publ 2657], fig. 2

cst iT tT bi riS oT roT roS iD
0.722 8 10 2 7 26 6 3 119.0
Cladomics cladogram Megalonaias Elliptoideus Uniomerus Pleurobema Potamilus fragilis Medionidus Amblema Obliquaria Plectomerus Villosa Ellipsaria Lampsilis Potamilus purpuratus Ptychobranchus

Hoeh et al., 2009 [Publ 2716], fig. 5

cst iT tT bi riS oT roT roS iD
0.944 7 13 7 7 39 6 3 124.0
Cladomics cladogram GONIDEINI Obliquaria QUADRULINI AMBLEMINI PLEUROBEMINI Truncilla Ptychobranchus Ortmanniana Cambarunio Ligumia Epioblasma

Inoue et al., 2020 [Publ 3941], fig. 3

cst iT tT bi riS oT roT roS iD
0.957 27 27 27 7 15 14 3 104.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI Toxolasma Potamilus Ellipsaria Truncilla Leaunio Lampsilis bracteata Lampsilis fasciola Lampsilis ornata Lampsilis ovata Lampsilis cariosa Lampsilis cardium 3 Lampsilis cardium 2 Lampsilis satura Lampsilis cardium 1 Lampsilis higginsii Ortmanniana Lampsilis teres Lampsilis radiata Lampsilis siliquoidea Lampsilis bergmanni Lampsilis hydiana Lampsilis virescens Lampsilis straminea

Inoue et al., 2020 [Publ 3941], fig. 4

cst iT tT bi riS oT roT roS iD
0.913 26 26 26 7 15 14 3 114.0
Cladomics cladogram QUADRULINI Reginaia PLEUROBEMINI Amblema Toxolasma Ellipsaria Truncilla Potamilus Leaunio Lampsilis bracteata Lampsilis ornata Lampsilis fasciola Lampsilis ovata Lampsilis cardium 3 Lampsilis cardium 2 Lampsilis satura Lampsilis cardium 1 Lampsilis teres Lampsilis higginsii Ortmanniana Lampsilis radiata Lampsilis siliquoidea Lampsilis hydiana Lampsilis bergmanni Lampsilis virescens Lampsilis straminea

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cst iT tT bi riS oT roT roS iD
0.947 12 12 10 6 128 6 4 115.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI POPENAIADINI Toxolasma Potamilus Leaunio Ortmanniana Lampsilis hydiana Lampsilis siliquoidea Venustaconcha Lampsilis ornata Lampsilis cardium Lampsilis cariosa

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

cst iT tT bi riS oT roT roS iD
0.941 6 11 5 6 28 5 3 119.0
Cladomics cladogram Cumberlandia Margaritifera Toxolasma AMBLEMINI Glebula Cyrtonaias QUADRULINI PLEUROBEMINI Potamilus Ortmanniana Ligumia

Hoeh et al., 2001 [Publ 2652], fig. 14f2

cst iT tT bi riS oT roT roS iD
0.824 6 28 16 6 25 5 3 138.0
Cladomics cladogram Cumberlandia Margaritifera AMBLEMINI COELATURINI Eurynia Fusconaia GONIDEINI Pleurobema QUADRULINI Ortmanniana Glebula Potamilus Ligumia Toxolasma Cyrtonaias UNIONINI ANODONTINI Mutela Anodontites guanarensis Anodontites trigona Monocondylaea RHIPIDODONTINI Echyridella Lortiella CASTALIINI Velesunio

Hoeh et al., 2001 [Publ 2652], fig. 14f4

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 6 25 5 3 113.0
Cladomics cladogram AMBLEMINI QUADRULINI PLEUROBEMINI Cyrtonaias Glebula Toxolasma Potamilus Ligumia Ortmanniana

Hoeh et al., 2002 [Publ 2653], fig. 2

cst iT tT bi riS oT roT roS iD
0.941 6 11 8 6 25 5 3 118.0
Cladomics cladogram GONIDEINI Unio pictorum Unio tumidus ANODONTINI Glebula Cyrtonaias Toxolasma Potamilus Ligumia Ortmanniana AMBLEMINI QUADRULINI PLEUROBEMINI

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

cst iT tT bi riS oT roT roS iD
0.824 6 12 9 6 25 5 3 127.0
Cladomics cladogram Fusconaia AMBLEMINI QUADRULINI Pleurobema Eurynia Cyrtonaias UNIONINI ANODONTINI Ortmanniana Glebula Potamilus Ligumia Toxolasma

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 6 25 5 3 115.0
Cladomics cladogram AMBLEMINI QUADRULINI PLEUROBEMINI Cyrtonaias Glebula Toxolasma Ortmanniana Ligumia Potamilus

Serb et al., 2003 [Publ 2659], fig. 2

cst iT tT bi riS oT roT roS iD
0.870 7 29 20 6 22 22 3 135.0
Cladomics cladogram Lampsilis Obliquaria Amblema Reginaia Potamilus QUADRULINI Elliptoideus Plectomerus Medionidus Dromus Cyprogenia Hemistena Elliptio Fusconaia

Serb et al., 2003 [Publ 2659], fig. 3

cst iT tT bi riS oT roT roS iD
0.826 7 29 27 6 22 22 3 135.0
Cladomics cladogram Lampsilis Potamilus Medionidus Dromus Cyprogenia Obliquaria Amblema Reginaia Elliptoideus Plectomerus QUADRULINI Hemistena Elliptio Fusconaia

Inoue et al., 2013 [Publ 3528], fig. 3

cst iT tT bi riS oT roT roS iD
0.938 10 13 10 6 7 5 3 122.0
Cladomics cladogram PLEUROBEMINI QUADRULINI Paetulunio AMBLEMINI Lampsilis Leaunio Obovaria Villosa Cambarunio

Pfeiffer et al., 2016 [Publ 3660], fig. 2

cst iT tT bi riS oT roT roS iD
0.852 6 6 4 6 32 31 3 115.0
Cladomics cladogram AMBLEMINI Cyrtonaias Glebula Lampsilis Leaunio Truncilla Potamilus Elliptoideus QUADRULINI Elliptio Parvaspina Plethobasus Hemistena Eurynia Fusconaia Pleurobema riddellii Pleurobema sintoxia Pleurobema cordatum Pleuronaia Pleurobema pyriforme Pleurobema strodeanum Pleurobema clava

Porto-Hannes et al., 2019 [Publ 3969], fig. 2

cst iT tT bi riS oT roT roS iD
0.900 17 17 15 6 1 1 1 115.0
Cladomics cladogram AMBLEMINI Obliquaria Cambarunio Leaunio Lampsilis teres Ortmanniana Lampsilis hydiana Lampsilis virescens Lampsilis siliquoidea Lampsilis radiata Obovaria Lampsilis bracteata Lampsilis ornata Lampsilis ovata Lampsilis cardium 3 Lampsilis satura Lampsilis cardium 1 Lampsilis cardium 2

Porto-Hannes et al., 2019 [Publ 3969], fig. 3

cst iT tT bi riS oT roT roS iD
0.800 17 18 13 6 1 1 1 114.0
Cladomics cladogram Obliquaria AMBLEMINI Lampsilis teres Obovaria Leaunio Cambarunio Ortmanniana Lampsilis hydiana Lampsilis virescens Lampsilis siliquoidea Lampsilis radiata Lampsilis bracteata Lampsilis ornata Lampsilis ovata Lampsilis cardium 3 Lampsilis satura Lampsilis cardium 1 Lampsilis cardium 2

Roe, 2013 [Publ 3571], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 11 11 9 6 0 0 0 119.0
Cladomics cladogram Obliquaria Lampsilis Medionidus Ptychobranchus Cyprogenia Dromus

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 52 10 4 118.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI POPENAIADINI Obliquaria Cyrtonaias Leaunio Lampsilis Pachynaias

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 52 10 4 118.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI POPENAIADINI Obliquaria Cyrtonaias Leaunio Lampsilis Pachynaias

Wu et al., 2021 [Publ 4366], fig. 3-BI

cst iT tT bi riS oT roT roS iD
0.944 9 9 7 5 64 6 4 114.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI POPENAIADINI Toxolasma Potamilus Leaunio Lampsilis ornata Lampsilis cardium Venustaconcha Lampsilis siliquoidea Lampsilis powellii

Wu et al., 2021 [Publ 4366], fig. 3-ML

cst iT tT bi riS oT roT roS iD
0.944 9 9 7 5 64 6 4 114.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI POPENAIADINI Toxolasma Potamilus Leaunio Lampsilis ornata Lampsilis cardium Venustaconcha Lampsilis siliquoidea Lampsilis powellii

Wu et al., 2022 [Publ 4238], fig. 3

cst iT tT bi riS oT roT roS iD
0.944 9 9 7 5 64 6 4 114.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI POPENAIADINI Toxolasma Potamilus Leaunio Lampsilis cardium Lampsilis ornata Venustaconcha Lampsilis siliquoidea Lampsilis powellii

Dai et al., 2023 [Publ 4285], fig. 5

cst iT tT bi riS oT roT roS iD
0.944 9 9 7 5 44 6 4 114.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI POPENAIADINI Toxolasma Potamilus Leaunio Lampsilis cardium Lampsilis ornata Venustaconcha Lampsilis powellii Lampsilis siliquoidea

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cst iT tT bi riS oT roT roS iD
0.944 11 11 9 5 84 6 4 114.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI POPENAIADINI Toxolasma Potamilus Leaunio Lampsilis ornata Lampsilis cardium Lampsilis cariosa Venustaconcha Lampsilis powellii Lampsilis siliquoidea

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cst iT tT bi riS oT roT roS iD
0.944 11 11 9 5 84 6 4 114.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI POPENAIADINI Toxolasma Potamilus Leaunio Lampsilis ornata Lampsilis cardium Lampsilis cariosa Venustaconcha Lampsilis powellii Lampsilis siliquoidea

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cst iT tT bi riS oT roT roS iD
0.944 11 11 9 5 80 6 4 114.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI POPENAIADINI Toxolasma Potamilus Leaunio Lampsilis ornata Lampsilis cardium Lampsilis cariosa Venustaconcha Lampsilis powellii Lampsilis siliquoidea

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cst iT tT bi riS oT roT roS iD
0.944 11 11 8 5 80 6 4 113.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI POPENAIADINI Toxolasma Potamilus Leaunio Venustaconcha Lampsilis powellii Lampsilis siliquoidea Lampsilis ornata Lampsilis cardium Lampsilis cariosa

Dai et al., 2024 [Publ 4358], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 5 47 6 4 115.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI POPENAIADINI Toxolasma Potamilus Leaunio Lampsilis Venustaconcha

Dai et al., 2025 [Publ 4440], fig. 3

cst iT tT bi riS oT roT roS iD
0.944 10 10 8 5 77 6 4 115.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI POPENAIADINI Toxolasma Potamilus Leaunio Lampsilis cardium Lampsilis ornata Venustaconcha Lampsilis powellii Lampsilis siliquoidea

Chen et al., 2025 [Publ 4448], fig. 3

cst iT tT bi riS oT roT roS iD
0.944 9 9 7 5 76 6 4 114.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI POPENAIADINI Toxolasma Potamilus Leaunio Lampsilis cardium Lampsilis ornata Venustaconcha Lampsilis powellii Lampsilis siliquoidea

Wu et al., 2024 [Publ 4388], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 48 5 3 116.0
Cladomics cladogram QUADRULINI PLEUROBEMINI POPENAIADINI Toxolasma Potamilus Leaunio Lampsilis Venustaconcha

Huang et al., 2019 [Publ 3843], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 5 49 3 2 114.0
Cladomics cladogram QUADRULINI PLEUROBEMINI Toxolasma Potamilus Leaunio Venustaconcha Lampsilis

Hoeh et al., 2002 [Publ 2673], fig. 1-F

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 4 1 1 118.0
Cladomics cladogram PLEUROBEMINI Cyrtonaias Ortmanniana Lampsilis Potamilus Ligumia

Hoeh et al., 2002 [Publ 2673], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 4 1 1 116.0
Cladomics cladogram PLEUROBEMINI Potamilus Cyrtonaias Ligumia Lampsilis Ortmanniana

Walker et al., 2006 [Publ 2666], fig. 2

cst iT tT bi riS oT roT roS iD
0.889 5 5 3 5 10 1 1 119.0
Cladomics cladogram PLEUROBEMINI Cumberlandia Mesonaias Potamilus Ligumia Ortmanniana Lampsilis

Walker et al., 2006 [Publ 2666], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 10 1 1 117.0
Cladomics cladogram PLEUROBEMINI Potamilus Mesonaias Ligumia Ortmanniana Lampsilis

Walker et al., 2006 [Publ 2666], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 10 1 1 119.0
Cladomics cladogram PLEUROBEMINI Mesonaias Potamilus Ligumia Ortmanniana Lampsilis

Walker et al., 2006 [Publ 2666], fig. 6

cst iT tT bi riS oT roT roS iD
0.889 5 6 3 5 31 1 1 117.0
Cladomics cladogram GONIDEINI PSEUDODONTINI Cyrtonaias PLEUROBEMINI Potamilus Lampsilis Ligumia Ortmanniana

Walker et al., 2006 [Publ 2666], fig. 6-M

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 9 1 1 116.0
Cladomics cladogram PLEUROBEMINI Potamilus Cyrtonaias Ligumia Lampsilis Ortmanniana

Roe et al., 2002 [Publ 2689], fig. 3a

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 5 0 0 0 115.0
Cladomics cladogram Ligumia Obliquaria Lampsilis Villosa Hamiota

Serb, 2006 [Publ 2678], fig. 2

cst iT tT bi riS oT roT roS iD
0.857 7 7 7 5 0 0 0 115.0
Cladomics cladogram Obliquaria Ptychobranchus Lampsilis Cyprogenia aberti 6 Dromus Cyprogenia aberti 5 Cyprogenia aberti 4 Cyprogenia stegaria 1 Cyprogenia stegaria 2 Cyprogenia aberti 1 Cyprogenia aberti 2 Cyprogenia aberti 3

Serb, 2006 [Publ 2678], fig. 3

cst iT tT bi riS oT roT roS iD
0.857 7 7 8 5 0 0 0 113.0
Cladomics cladogram Obliquaria Lampsilis Ptychobranchus Cyprogenia aberti 5 Dromus Cyprogenia aberti 4 Cyprogenia aberti 3 Cyprogenia stegaria 2 Cyprogenia aberti 2 Cyprogenia stegaria 1 Cyprogenia aberti 1

Serb, 2006 [Publ 2678], fig. 4

cst iT tT bi riS oT roT roS iD
0.857 7 7 8 5 0 0 0 113.0
Cladomics cladogram Obliquaria Lampsilis Ptychobranchus Cyprogenia aberti 5 Dromus Cyprogenia aberti 4 Cyprogenia aberti 3 Cyprogenia stegaria 2 Cyprogenia aberti 2 Cyprogenia stegaria 1 Cyprogenia aberti 1

Serb & Barnhart, 2008 [Publ 2674], fig. 3

cst iT tT bi riS oT roT roS iD
0.857 7 7 10 5 0 0 0 114.0
Cladomics cladogram Lampsilis Obliquaria Ptychobranchus Cyprogenia aberti 6 Dromus Cyprogenia aberti 4 Cyprogenia aberti 3 Cyprogenia aberti 5 Cyprogenia stegaria 2 Cyprogenia aberti 2 Cyprogenia stegaria 1 Cyprogenia aberti 1

Serb & Barnhart, 2008 [Publ 2674], fig. 4

cst iT tT bi riS oT roT roS iD
0.857 7 7 9 5 0 0 0 113.0
Cladomics cladogram Obliquaria Lampsilis Ptychobranchus Cyprogenia aberti 6 Dromus Cyprogenia aberti 5 Cyprogenia aberti 4 Cyprogenia aberti 3 Cyprogenia stegaria 2 Cyprogenia aberti 2 Cyprogenia stegaria 1 Cyprogenia aberti 1

Campbell & Lydeard, 2012 [Publ 3308], fig. 1

cst iT tT bi riS oT roT roS iD
0.852 5 8 6 4 37 37 4 115.0
Cladomics cladogram QUADRULINI PLEUROBEMINI POPENAIADINI Amblema Cyrtonaias Reginaia Plectomerus Lemiox Lampsilis Obovaria

Campbell & Lydeard, 2012 [Publ 3308], fig. 2

cst iT tT bi riS oT roT roS iD
0.889 5 6 3 4 37 37 4 114.0
Cladomics cladogram QUADRULINI PLEUROBEMINI Reginaia Amblema POPENAIADINI Cyrtonaias Plectomerus Lemiox Lampsilis Obovaria

Campbell & Lydeard, 2012 [Publ 3308], fig. 3

cst iT tT bi riS oT roT roS iD
0.786 5 36 35 4 38 38 4 136.0
Cladomics cladogram QUADRULINI Elliptoideus Plectomerus Obovaria unicolor Obovaria olivaria Lampsilis Lemiox Cyrtonaias Reginaia POPENAIADINI Amblema Elliptio Eurynia Parvaspina Hemistena Plethobasus Pleuronaia Pleurobema cordatum Fusconaia Pleurobema stabile Pleurobema sintoxia Pleurobema decisum Pleurobema clava Pleurobema pyriforme

Campbell & Lydeard, 2012 [Publ 3308], fig. 4

cst iT tT bi riS oT roT roS iD
0.852 5 8 5 4 37 37 4 116.0
Cladomics cladogram QUADRULINI PLEUROBEMINI POPENAIADINI Amblema Plectomerus Cyrtonaias Reginaia Lemiox Lampsilis Obovaria

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

cst iT tT bi riS oT roT roS iD
0.941 8 8 6 4 11 6 4 113.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI POPENAIADINI Toxolasma Potamilus Lampsilis cardium Lampsilis ornata Venustaconcha Lampsilis powellii Lampsilis siliquoidea

Wu et al., 2021 [Publ 4366], fig. 4

cst iT tT bi riS oT roT roS iD
0.933 8 8 6 4 60 4 4 113.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI POPENAIADINI Toxolasma Potamilus Lampsilis ornata Lampsilis cardium Venustaconcha Lampsilis siliquoidea Lampsilis powellii

Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F

cst iT tT bi riS oT roT roS iD
0.882 9 15 13 4 6 6 4 121.0
Cladomics cladogram Toxolasma POPENAIADINI AMBLEMINI QUADRULINI PLEUROBEMINI Potamilus Lampsilis cardium Lampsilis ornata Venustaconcha Lampsilis siliquoidea Lampsilis powellii

Freeman et al., 2025 [Publ 4443], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 8 8 4 117.0
Cladomics cladogram AMBLEMINI PLEUROBEMINI QUADRULINI POPENAIADINI Toxolasma Obliquaria Glebula Cyrtonaias

Freeman et al., 2025 [Publ 4443], fig. 4

cst iT tT bi riS oT roT roS iD
0.947 4 6 2 4 8 8 4 117.0
Cladomics cladogram AMBLEMINI PLEUROBEMINI QUADRULINI Obliquaria Toxolasma POPENAIADINI Glebula Cyrtonaias

Porto-Hannes et al., 2021 [Publ 4227], fig. 3

cst iT tT bi riS oT roT roS iD
0.714 9 9 7 4 6 6 3 117.0
Cladomics cladogram Quadrula quadrula Quadrula verrucosa PLEUROBEMINI AMBLEMINI Obovaria Ortmanniana Obliquaria Lampsilis ornata Lampsilis radiata 2 Lampsilis siliquoidea Lampsilis radiata 1 Lampsilis virescens Lampsilis hydiana Lampsilis straminea

Curole & Kocher, 2005 [Publ 2682], fig. 1

cst iT tT bi riS oT roT roS iD
0.769 5 9 7 4 6 4 2 120.0
Cladomics cladogram ANODONTINI Margaritifera Glebula Amblema QUADRULINI Plectomerus Leaunio Lampsilis straminea Lampsilis teres Potamilus

Curole & Kocher, 2005 [Publ 2682], fig. 1-M

cst iT tT bi riS oT roT roS iD
0.923 5 5 3 4 7 4 2 116.0
Cladomics cladogram QUADRULINI Plectomerus Amblema Potamilus Glebula Leaunio Lampsilis

Fukata & Iigo, 2020 [Publ 4106], fig. 1

cst iT tT bi riS oT roT roS iD
0.917 8 8 6 4 46 3 2 115.0
Cladomics cladogram QUADRULINI PLEUROBEMINI Potamilus Leaunio Lampsilis cardium Lampsilis ornata Venustaconcha Lampsilis powellii Lampsilis siliquoidea

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 87 2 2 115.0
Cladomics cladogram QUADRULINI PLEUROBEMINI Obliquaria Potamilus Lampsilis Truncilla

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 87 2 2 115.0
Cladomics cladogram QUADRULINI PLEUROBEMINI Obliquaria Potamilus Lampsilis Truncilla

Wu et al., 2023 [Publ 4339], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 15 2 2 117.0
Cladomics cladogram PLEUROBEMINI QUADRULINI Obliquaria Truncilla Lampsilis Potamilus

Wen et al., 2017 [Publ 3795], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 4 11 1 1 114.0
Cladomics cladogram QUADRULINI Toxolasma Potamilus Lampsilis Venustaconcha

Huang et al., 2018 [Publ 3828], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 4 37 1 1 114.0
Cladomics cladogram QUADRULINI Toxolasma Potamilus Lampsilis Venustaconcha

Wang et al., 2018 [Publ 3846], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 4 26 1 1 114.0
Cladomics cladogram QUADRULINI Toxolasma Potamilus Venustaconcha Lampsilis

Wang et al., 2018 [Publ 3865], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 4 26 1 1 114.0
Cladomics cladogram QUADRULINI Toxolasma Potamilus Venustaconcha Lampsilis

Wu et al., 2019 [Publ 3864], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 4 41 1 1 114.0
Cladomics cladogram QUADRULINI Toxolasma Potamilus Lampsilis Venustaconcha

Froufe et al., 2019 [Publ 3936], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 4 39 1 1 114.0
Cladomics cladogram QUADRULINI Toxolasma Potamilus Lampsilis Venustaconcha

Froufe et al., 2019 [Publ 3936], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 4 39 1 1 114.0
Cladomics cladogram QUADRULINI Toxolasma Potamilus Lampsilis Venustaconcha

Wu et al., 2020 [Publ 4121], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 4 43 1 1 114.0
Cladomics cladogram QUADRULINI Toxolasma Potamilus Venustaconcha Lampsilis

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 4 43 1 1 114.0
Cladomics cladogram QUADRULINI Toxolasma Potamilus Venustaconcha Lampsilis

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

cst iT tT bi riS oT roT roS iD
0.889 6 6 4 4 31 1 1 115.0
Cladomics cladogram QUADRULINI Toxolasma Potamilus Lampsilis cardium Lampsilis ornata Lampsilis siliquoidea Venustaconcha

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

cst iT tT bi riS oT roT roS iD
0.889 6 6 4 4 33 1 1 115.0
Cladomics cladogram QUADRULINI Toxolasma Potamilus Lampsilis cardium Lampsilis ornata Lampsilis siliquoidea Venustaconcha

Klymus et al., 2025 [Publ 4814], fig. 3A

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 4 31 1 1 114.0
Cladomics cladogram QUADRULINI Potamilus Venustaconcha Ortmanniana Lampsilis

Klymus et al., 2025 [Publ 4814], fig. 3B

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 4 31 1 1 114.0
Cladomics cladogram QUADRULINI Potamilus Venustaconcha Ortmanniana Lampsilis

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 4 41 1 1 113.0
Cladomics cladogram QUADRULINI Potamilus Venustaconcha Ortmanniana Lampsilis

Buhay et al., 2002 [Publ 2661], fig. 2A

cst iT tT bi riS oT roT roS iD
1.000 7 7 3 4 0 0 0 117.0
Cladomics cladogram Lampsilis Epioblasma Cambarunio Villosa

Buhay et al., 2002 [Publ 2661], fig. 2B

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 4 0 0 0 118.0
Cladomics cladogram Lampsilis Epioblasma Cambarunio Villosa

Freeman et al., 2025 [Publ 4443], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 8 8 4 116.0
Cladomics cladogram AMBLEMINI PLEUROBEMINI QUADRULINI POPENAIADINI Obliquaria Glebula Cyrtonaias

Barraza et al., 2026 [Publ 4851], fig. 3

cst iT tT bi riS oT roT roS iD
0.824 3 5 3 3 16 9 4 114.0
Cladomics cladogram Amblema Psoronaias Nephronaias Reginaia QUADRULINI PLEUROBEMINI Mesonaias Pachynaias Cyrtonaias Popenaias

Turner et al., 2000 [Publ 2722], fig. 2

cst iT tT bi riS oT roT roS iD
0.833 9 9 7 3 5 3 3 117.0
Cladomics cladogram AMBLEMINI PLEUROBEMINI Medionidus Lampsilis teres Villosa Lampsilis satura Lampsilis cardium Lampsilis reeveiana Lampsilis hydiana 1 Lampsilis hydiana 2 Lampsilis siliquoidea 2 Lampsilis siliquoidea 1 Lampsilis powellii

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 78 6 3 116.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI Cambarunio Lampsilis Ortmanniana

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 136 6 3 118.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI Ortmanniana Lampsilis Cambarunio

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 136 6 3 116.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI Cambarunio Ortmanniana Lampsilis

Bolotov et al., 2017 [Publ 3764], fig. S1

cst iT tT bi riS oT roT roS iD
0.643 3 9 0 3 122 6 3 123.0
Cladomics cladogram AMBLEMINI Ortmanniana Lampsilis Cambarunio Elliptio Eurynia Pleurobema Quadrula quadrula Quadrula verrucosa Chamberlainia Sinohyriopsis Lamprotula caveata Lamprotula leaii Potomida Pronodularia Gonidea Leguminaia wheatleyi Leguminaia Microcondylaea Sinosolenaia carinata Sinosolenaia oleivora Sinosolenaia recognita

Bolotov et al., 2018 [Publ 3826], fig. 2

cst iT tT bi riS oT roT roS iD
0.769 3 13 0 3 79 4 3 125.0
Cladomics cladogram UNIONINI LANCEOLARIINI ANODONTINI Lamprotula Potomida Pronodularia Gonidea Leguminaia Microcondylaea AMBLEMINI Ortmanniana Lampsilis Cambarunio Elliptio Pleurobema QUADRULINI

Bolotov et al., 2020 [Publ 4084], fig. 1

cst iT tT bi riS oT roT roS iD
0.846 3 7 0 3 33 4 3 121.0
Cladomics cladogram Margaritifera AMBLEMINI Ortmanniana Lampsilis Cambarunio Elliptio Pleurobema QUADRULINI UNIONINI LANCEOLARIINI ANODONTINI Lamprotula Potomida Pronodularia Gonidea Leguminaia Microcondylaea

Porto-Hannes et al., 2021 [Publ 4227], fig. 4

cst iT tT bi riS oT roT roS iD
0.846 7 7 3 3 4 4 3 117.0
Cladomics cladogram Quadrula Pustulosa PLEUROBEMINI AMBLEMINI Obovaria Ortmanniana Lampsilis

Smith, 2021 [Publ 4331], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 16 5 3 115.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI Potamilus Venustaconcha Lampsilis

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 73 6 3 118.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI Ortmanniana Cambarunio Lampsilis

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 73 6 3 118.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI Ortmanniana Lampsilis Cambarunio

Plazzi et al., 2011 [Publ 3676], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 15 2 2 116.0
Cladomics cladogram QUADRULINI PLEUROBEMINI Lampsilis Epioblasma Venustaconcha

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 37 2 2 116.0
Cladomics cladogram QUADRULINI AMBLEMINI Potamilus Lampsilis Truncilla

Amaro et al., 2017 [Publ 3791], fig. 2

cst iT tT bi riS oT roT roS iD
0.900 3 3 1 3 13 3 2 116.0
Cladomics cladogram QUADRULINI Toxolasma Venustaconcha Lampsilis

Keogh & Simons, 2019 [Publ 3935], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 3 2 2 2 113.0
Cladomics cladogram AMBLEMINI PLEUROBEMINI Potamilus Leaunio Lampsilis

Roe & Lydeard, 1998 [Publ 2693], fig. 3

cst iT tT bi riS oT roT roS iD
0.714 9 10 5 3 1 1 1 115.0
Cladomics cladogram PLEUROBEMINI Obliquaria Potamilus capax Lampsilis Potamilus fragilis Potamilus inflatus Potamilus ohiensis Potamilus amphichaenus Potamilus purpuratus 1 Potamilus purpuratus 2 Potamilus alatus

Huang et al., 2013 [Publ 3629], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 13 1 1 116.0
Cladomics cladogram QUADRULINI Toxolasma Lampsilis Venustaconcha

Ouyang et al., 2015 [Publ 4493], fig. 1

cst iT tT bi riS oT roT roS iD
0.900 3 9 7 3 28 6 1 122.0
Cladomics cladogram LAMPROTULINI CHAMBERLAINIINI GONIDEINI Epioblasma Lampsilis Cyrtonaias PLEUROBEMINI

Ouyang et al., 2015 [Publ 4493], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 28 6 1 116.0
Cladomics cladogram PLEUROBEMINI Cyrtonaias Lampsilis Epioblasma

Zhou et al., 2016 [Publ 3710], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 17 1 1 116.0
Cladomics cladogram QUADRULINI Toxolasma Venustaconcha Lampsilis

Fonseca et al., 2016 [Publ 3775], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 28 1 1 116.0
Cladomics cladogram QUADRULINI Toxolasma Lampsilis Venustaconcha

Guerra et al., 2017 [Publ 3772], fig. 2a

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 22 1 1 116.0
Cladomics cladogram QUADRULINI Toxolasma Venustaconcha Lampsilis

Guerra et al., 2017 [Publ 3772], fig. 2b

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 22 1 1 116.0
Cladomics cladogram QUADRULINI Toxolasma Venustaconcha Lampsilis

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 29 1 1 116.0
Cladomics cladogram QUADRULINI Toxolasma Lampsilis Venustaconcha

Aunins et al., 2018 [Publ 3863], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 37 1 1 116.0
Cladomics cladogram QUADRULINI Toxolasma Lampsilis Venustaconcha

Cha et al., 2018 [Publ 3916], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 34 1 1 115.0
Cladomics cladogram QUADRULINI Toxolasma Lampsilis Potamilus

Guerra et al., 2019 [Publ 4168], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 38 1 1 116.0
Cladomics cladogram QUADRULINI Toxolasma Venustaconcha Potamilus

Guerra et al., 2019 [Publ 4168], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 38 1 1 116.0
Cladomics cladogram QUADRULINI Toxolasma Venustaconcha Potamilus

Chen et al., 2020 [Publ 4170], fig. 1

cst iT tT bi riS oT roT roS iD
0.500 6 16 14 3 28 1 1 126.0
Cladomics cladogram OXYNAIINA LEPIDODESMINI ACULAMPROTULINA Platynaias Utterbackia imbecillis Pyganodon Sinanodonta Cristaria ANODONTINA Utterbackia peninsularis Potamilus alatus LAMPROTULINI CHAMBERLAINIINI GONIDEINI Gibbosula QUADRULINI Toxolasma Lampsilis ornata Potamilus leptodon Lampsilis siliquoidea Lampsilis powellii

Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 3 1 1 1 114.0
Cladomics cladogram QUADRULINI Potamilus Venustaconcha Lampsilis

Martínez et al., 2026 [Publ 4820], fig. 2

cst iT tT bi riS oT roT roS iD
0.778 3 3 1 3 13 6 1 116.0
Cladomics cladogram Nephronaias hjalmarsoni Psoronaias Nephronaias goascoranensis Nephronaias tempisquensis Popenaias Cyrtonaias Mesonaias Pachynaias

Zanatta & Harris, 2013 [Publ 3530], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 1 0 0 115.0
Cladomics cladogram ANODONTINI Lampsilis Epioblasma Venustaconcha

Gangloff et al., 2013 [Publ 3551], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 0 0 0 117.0
Cladomics cladogram Venustaconcha Ligumia Sagittunio

Lane et al., 2016 [Publ 3735], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 5 5 1 3 0 0 0 117.0
Cladomics cladogram Paetulunio Epioblasma Venustaconcha

Rosenberg et al., 1994 [Publ 2719], fig. 3

cst iT tT bi riS oT roT roS iD
0.625 3 10 0 2 16 8 3 122.0
Cladomics cladogram Elliptio Fusconaia AMBLEMINI Pleurobema UNIONINI Pyganodon cataracta Pyganodon grandis Utterbackia Lampsilis teres Lampsilis straminea Megalonaias Obliquaria Theliderma Quadrula Uniomerus

Rosenberg et al., 1997 [Publ 2720], fig. 2A

cst iT tT bi riS oT roT roS iD
0.588 3 17 0 2 19 9 3 127.0
Cladomics cladogram ANODONTINA GONIDEINI Pyganodon grandis Utterbackia Pyganodon cataracta Lampsilis straminea Lampsilis teres Megalonaias Obliquaria Theliderma Quadrula Uniomerus Amblema Elliptio Fusconaia Plectomerus Pleurobema UNIONINI

Rosenberg et al., 1997 [Publ 2720], fig. 2B

cst iT tT bi riS oT roT roS iD
0.562 3 17 0 2 19 8 3 127.0
Cladomics cladogram ANODONTINA GONIDEINI Pyganodon grandis Utterbackia Pyganodon cataracta Lampsilis straminea Lampsilis teres Megalonaias Obliquaria Theliderma Quadrula Uniomerus Amblema Elliptio Fusconaia Plectomerus ACULAMPROTULINA UNIONINA

Lydeard et al., 2000 [Publ 2654], fig. 2a

cst iT tT bi riS oT roT roS iD
0.923 3 3 1 2 11 9 3 116.0
Cladomics cladogram ANODONTINI Cumberlandia AMBLEMINI Lampsilis Obovaria QUADRULINI PLEUROBEMINI

Lydeard et al., 2000 [Publ 2654], fig. 2b

cst iT tT bi riS oT roT roS iD
0.923 3 3 1 2 11 9 3 116.0
Cladomics cladogram Lampsilis Obovaria AMBLEMINI QUADRULINI PLEUROBEMINI

Lydeard et al., 2000 [Publ 2654], fig. 2c

cst iT tT bi riS oT roT roS iD
0.846 3 3 1 2 11 9 3 118.0
Cladomics cladogram ANODONTINI Cumberlandia QUADRULINI AMBLEMINI PLEUROBEMINI Obovaria unicolor Obovaria olivaria Lampsilis

Lydeard et al., 2000 [Publ 2654], fig. 2d

cst iT tT bi riS oT roT roS iD
0.923 3 3 1 2 11 9 3 116.0
Cladomics cladogram AMBLEMINI Lampsilis Obovaria QUADRULINI PLEUROBEMINI

Whelan et al., 2011 [Publ 3380], fig. 1-ML

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 34 3 3 --
Cladomics cladogram PLEUROBEMINI QUADRULINI AMBLEMINI Truncilla Lampsilis

Whelan et al., 2011 [Publ 3380], fig. 1-MP

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 34 3 3 --
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI Lampsilis Truncilla

Burlakova et al., 2012 [Publ 3486], fig. 1

cst iT tT bi riS oT roT roS iD
0.833 2 39 0 2 39 39 3 141.0
Cladomics cladogram Amblema Lampsilis Plectomerus POPENAIADINI Cyrtonaias PLEUROBEMINI

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 29 3 3 --
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI Lampsilis Truncilla

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

cst iT tT bi riS oT roT roS iD
0.882 2 2 0 2 26 22 3 --
Cladomics cladogram AMBLEMINI Lampsilis Potamilus PLEUROBEMINI QUADRULINI

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

cst iT tT bi riS oT roT roS iD
0.875 2 2 0 2 27 23 3 --
Cladomics cladogram Megalonaias Uniomerus QUADRULINA PLEUROBEMINI AMBLEMINI Lampsilis Potamilus

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

cst iT tT bi riS oT roT roS iD
0.875 2 2 0 2 24 20 3 --
Cladomics cladogram QUADRULINI AMBLEMINI PLEUROBEMINI Lampsilis Potamilus

Hauck et al., 2023 [Publ 4333], fig. 9

cst iT tT bi riS oT roT roS iD
0.917 2 2 0 2 11 11 3 --
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI Obovaria Lampsilis

Graf, 2002 [Publ 2651], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 16 2 2 --
Cladomics cladogram Lampsilis Obliquaria PLEUROBEMINI AMBLEMINI

Manendo et al., 2008 [Publ 2669], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 11 3 2 --
Cladomics cladogram Lampsilis Sagittunio AMBLEMINI PLEUROBEMINI

Amaro et al., 2017 [Publ 3791], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 8 2 2 --
Cladomics cladogram PLEUROBEMINI QUADRULINI Lampsilis Venustaconcha

Chen et al., 2012 [Publ 3624], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 5 1 1 --
Cladomics cladogram QUADRULINI Lampsilis Venustaconcha

Sheng et al., 2014 [Publ 3704], fig. 2

cst iT tT bi riS oT roT roS iD
0.714 2 6 0 2 6 1 1 121.0
Cladomics cladogram ANODONTINI Lampsilis Venustaconcha QUADRULINI LAMPROTULINI CHAMBERLAINIINI

Wu et al., 2016 [Publ 3711], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 19 1 1 --
Cladomics cladogram QUADRULINI Venustaconcha Lampsilis

Feng et al., 2016 [Publ 3800], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 12 1 1 --
Cladomics cladogram QUADRULINI Potamilus Lampsilis

Zhou et al., 2017 [Publ 3788], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 14 1 1 --
Cladomics cladogram QUADRULINI Potamilus Lampsilis

Amaro et al., 2017 [Publ 3791], fig. 3A

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 10 1 1 --
Cladomics cladogram QUADRULINI Toxolasma Venustaconcha

Amaro et al., 2017 [Publ 3791], fig. 3B

cst iT tT bi riS oT roT roS iD
0.857 2 2 0 2 10 1 1 --
Cladomics cladogram Toxolasma Venustaconcha QUADRULINI LAMPROTULINI GONIDEINI

Wen et al., 2017 [Publ 3795], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 4 1 1 --
Cladomics cladogram QUADRULINI Potamilus Venustaconcha

Huang et al., 2018 [Publ 3828], fig. 6-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 15 1 1 --
Cladomics cladogram QUADRULINI Venustaconcha Potamilus

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 18 1 1 --
Cladomics cladogram QUADRULINI Potamilus Venustaconcha

Wu et al., 2018 [Publ 4143], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 20 1 1 --
Cladomics cladogram QUADRULINI Venustaconcha Lampsilis

Huang et al., 2019 [Publ 3843], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 21 1 1 --
Cladomics cladogram QUADRULINI Venustaconcha Potamilus

Froufe et al., 2019 [Publ 3936], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 17 1 1 --
Cladomics cladogram QUADRULINI Potamilus Venustaconcha

Froufe et al., 2019 [Publ 3936], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 17 1 1 --
Cladomics cladogram QUADRULINI Potamilus Venustaconcha

Hewitt et al., 2019 [Publ 3962], fig. 2

cst iT tT bi riS oT roT roS iD
0.833 7 7 8 2 3 1 1 116.0
Cladomics cladogram PLEUROBEMINI Lampsilis siliquoidea Lampsilis fasciola Epioblasma Lampsilis radiata Lampsilis ornata Lampsilis cardium 1 Lampsilis ovata 1 Lampsilis cardium 6 Lampsilis cardium 5 Lampsilis cardium 2 Lampsilis ovata 2 Lampsilis cardium 3 Lampsilis cardium 4

Guerra et al., 2019 [Publ 4168], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 20 1 1 --
Cladomics cladogram QUADRULINI Venustaconcha Potamilus

Guerra et al., 2019 [Publ 4168], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 20 1 1 --
Cladomics cladogram QUADRULINI Venustaconcha Potamilus

Wu et al., 2020 [Publ 4147], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 28 1 1 --
Cladomics cladogram QUADRULINI Toxolasma Venustaconcha

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 21 1 1 --
Cladomics cladogram QUADRULINI Venustaconcha Potamilus

Dai et al., 2024 [Publ 4386], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 38 1 1 116.0
Cladomics cladogram QUADRULINI Venustaconcha Lampsilis

Freeman et al., 2025 [Publ 4443], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 4 4 1 --
Cladomics cladogram POPENAIADINI Mesonaias Pachynaias

Buhay et al., 2002 [Publ 2661], fig. 1

cst iT tT bi riS oT roT roS iD
0.750 5 5 3 2 0 0 0 117.0
Cladomics cladogram Lampsilis siliquoidea 1 Lampsilis siliquoidea 2 Epioblasma

Krebs et al., 2013 [Publ 3515], fig. 2A

cst iT tT bi riS oT roT roS iD
0.750 5 5 3 2 0 0 0 118.0
Cladomics cladogram Lampsilis radiata Lampsilis ovata Ortmanniana Lampsilis hydiana Lampsilis straminea

Xue et al., 2016 [Publ 3659], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 13 0 0 --
Cladomics cladogram CHAMBERLAINIINI LAMPROTULINI Lampsilis Venustaconcha

Chase et al., 2018 [Publ 3847], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 7 0 0 116.0
Cladomics cladogram Potamilus Lampsilis ANODONTINI UNIONINI

Inoue et al., 2020 [Publ 3941], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 16 16 14 2 0 0 0 113.0
Cladomics cladogram Truncilla Lampsilis

Stodola et al., 2021 [Publ 4266], fig. 4

cst iT tT bi riS oT roT roS iD
0.750 10 10 7 2 0 0 0 118.0
Cladomics cladogram Ortmanniana Lampsilis abrupta Lampsilis higginsii Lampsilis sietmani Lampsilis siliquoidea Lampsilis radiata Lampsilis hydiana 2 Lampsilis straminea 2 Lampsilis hydiana 1 Lampsilis bergmanni Lampsilis straminea 1 Lampsilis virescens

Saxon et al., 2023 [Publ 4338], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 2 0 0 0 116.0
Cladomics cladogram Ellipsaria Truncilla

Neemuchwala et al., 2023 [Publ 4292], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 26 25 4 --
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI Lampsilis POPENAIADINI

Campbell et al., 2008 [Publ 2675], fig. 4

cst iT tT bi riS oT roT roS iD
0.812 1 1 0 1 24 24 3 --
Cladomics cladogram QUADRULINI AMBLEMINI Cyrtonaias Eurynia Fusconaia cerina Fusconaia flava Hemistena Plethobasus Parvaspina Pleurobema cordatum Pleuronaia Pleurobema oviforme 2 Pleurobema stabile Elliptio Pleurobema decisum Pleurobema sintoxia Pleurobema clava Pleurobema oviforme 1 Pleurobema beadleianum Pleurobema pyriforme Pleurobema rubellum Pleurobema hanleyianum 2 Pleurobema georgianum 2 Pleurobema hanleyianum 1 Pleurobema georgianum 1 Pleurobema athearni

Campbell et al., 2008 [Publ 2675], fig. 5

cst iT tT bi riS oT roT roS iD
0.750 1 1 0 1 24 24 3 --
Cladomics cladogram QUADRULINI AMBLEMINI Elliptio arctata Elliptio crassidens Pleurobema decisum Pleurobema rubellum Pleurobema georgianum 2 Pleurobema stabile Pleurobema pyriforme Pleurobema oviforme 2 Pleurobema hanleyianum 2 Pleuronaia Pleurobema georgianum 3 Pleurobema cordatum Pleurobema beadleianum Hemistena Fusconaia flava Fusconaia cerina Eurynia Elliptio arca Cyrtonaias Plethobasus Parvaspina Pleurobema sintoxia Pleurobema oviforme 1 Pleurobema clava Pleurobema athearni Pleurobema georgianum 1 Pleurobema hanleyianum 1

Inoue et al., 2014 [Publ 3635], fig. 3A

cst iT tT bi riS oT roT roS iD
0.900 1 1 0 1 19 3 3 --
Cladomics cladogram AMBLEMINI PLEUROBEMINI Lampsilis QUADRULINI GONIDEINI ANODONTINI

Wu et al., 2018 [Publ 3836], fig. 2

cst iT tT bi riS oT roT roS iD
0.923 1 1 0 1 52 8 3 --
Cladomics cladogram AMBLEMINI Obovaria

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 121 3 3 --
Cladomics cladogram AMBLEMINI PLEUROBEMINI Lampsilis QUADRULINI

Krebs, 2004 [Publ 2692], fig. 2

cst iT tT bi riS oT roT roS iD
0.875 1 1 0 1 4 2 2 --
Cladomics cladogram Lampsilis PLEUROBEMINI

Burdick & White, 2007 [Publ 2679], fig. 3

cst iT tT bi riS oT roT roS iD
0.846 1 1 0 1 29 29 2 --
Cladomics cladogram Lampsilis Parvaspina AMBLEMINI Eurynia Plethobasus Elliptio Pleurobema plenum Pleuronaia Pleurobema cordatum 1 Fusconaia Pleurobema sintoxia 1 Pleurobema cordatum 2 Pleurobema sintoxia 2 Pleurobema sintoxia 3 Pleurobema stabile Pleurobema rubellum Pleurobema strodeanum Pleurobema beadleianum Pleurobema perovatum Pleurobema taitianum Pleurobema clava Pleurobema oviforme Pleurobema decisum Pleurobema hanleyianum Pleurobema pyriforme Pleurobema georgianum

Campbell et al., 2008 [Publ 2675], fig. 2

cst iT tT bi riS oT roT roS iD
0.929 1 1 0 1 25 25 2 --
Cladomics cladogram Cyrtonaias AMBLEMINI PLEUROBEMINI

Campbell et al., 2008 [Publ 2675], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 25 25 2 --
Cladomics cladogram Cyrtonaias AMBLEMINI PLEUROBEMINI

Graf, 2000 [Publ 2647], fig. 1

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 16 1 1 --
Cladomics cladogram ANODONTINI Lampsilis

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 1 1 --
Cladomics cladogram AMBLEMINI Lampsilis

Curole & Kocher, 2002 [Publ 2712], fig. 1-F

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 2 1 --
Cladomics cladogram Lampsilis QUADRULINI

Curole & Kocher, 2002 [Publ 2712], fig. 1a-M

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 3 2 1 --
Cladomics cladogram ANODONTINI Lampsilis

Curole & Kocher, 2002 [Publ 2712], fig. 1b-M

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 3 2 1 --
Cladomics cladogram Lampsilis ANODONTINI QUADRULINI

Giribet & Distel, 2003 [Publ 3670], fig. 3f5

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 5 1 1 --
Cladomics cladogram PLEUROBEMINI Lampsilis UNIONINI

Krebs, 2004 [Publ 2692], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 1 1 --
Cladomics cladogram Lampsilis PLEUROBEMINI

Huang et al., 2013 [Publ 3629], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 1 1 --
Cladomics cladogram QUADRULINI Venustaconcha

Froufe et al., 2015 [Publ 3664], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 1 1 --
Cladomics cladogram Venustaconcha QUADRULINI

Froufe et al., 2015 [Publ 3664], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 1 1 --
Cladomics cladogram Venustaconcha QUADRULINI

Xue et al., 2016 [Publ 3659], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 1 1 1 --
Cladomics cladogram QUADRULINI Venustaconcha

Wei et al., 2016 [Publ 3665], fig. 0

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 5 1 1 --
Cladomics cladogram CRISTARIINA Lampsilis ALASMIDONTINA QUADRULINI

Fonseca et al., 2016 [Publ 3775], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 1 1 --
Cladomics cladogram Venustaconcha QUADRULINI

Guerra et al., 2017 [Publ 3772], fig. 2a-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 1 1 --
Cladomics cladogram Venustaconcha QUADRULINI

Guerra et al., 2017 [Publ 3772], fig. 2b-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 1 1 --
Cladomics cladogram Venustaconcha QUADRULINI

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 12 1 1 --
Cladomics cladogram QUADRULINI Venustaconcha

Burzynski et al., 2017 [Publ 3774], fig. 1-top

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 12 1 1 --
Cladomics cladogram QUADRULINI Venustaconcha

Amaro et al., 2017 [Publ 3791], fig. 3A-M

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 6 1 1 --
Cladomics cladogram Venustaconcha ANODONTINI

Amaro et al., 2017 [Publ 3791], fig. 3B-M

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 6 1 1 --
Cladomics cladogram Venustaconcha ANODONTINI LAMPROTULINI GONIDEINI

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 1 1 --
Cladomics cladogram Venustaconcha QUADRULINI

Araujo et al., 2017 [Publ 3801], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 1 1 --
Cladomics cladogram AMBLEMINI Lampsilis

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 6 1 1 --
Cladomics cladogram Venustaconcha QUADRULINI

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 6 1 1 --
Cladomics cladogram Venustaconcha QUADRULINI

Chase et al., 2018 [Publ 3861], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 9 1 1 --
Cladomics cladogram QUADRULINI Lampsilis

Chase et al., 2018 [Publ 3861], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 9 1 1 --
Cladomics cladogram QUADRULINI Lampsilis

Metzger et al., 2018 [Publ 3952], fig. 1C

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 2 1 1 --
Cladomics cladogram PLEUROBEMINI Lampsilis

Zieritz et al., 2021 [Publ 4218], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 1 1 --
Cladomics cladogram AMBLEMINI Lampsilis

Sonowal et al., 2021 [Publ 4221], fig. 3a

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 14 1 1 --
Cladomics cladogram PLEUROBEMINI Lampsilis ANODONTINI Unio ravoisieri Unio pictorum ACULAMPROTULINA

Sonowal et al., 2021 [Publ 4221], fig. 3b

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 14 1 1 --
Cladomics cladogram ACULAMPROTULINA Lamprotula Lampsilis Potomida Unio pictorum ANODONTINI Unio ravoisieri LAMELLIDENTINI PLEUROBEMINI

Sonowal et al., 2021 [Publ 4221], fig. 4

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 15 1 1 --
Cladomics cladogram Lampsilis PLEUROBEMINI ACULAMPROTULINA Unio pictorum ANODONTINI Unio ravoisieri LAMELLIDENTINI

Patterson et al., 2021 [Publ 4222], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 3 1 --
Cladomics cladogram Villosa AMBLEMINI

Konopleva et al., 2023 [Publ 4269], fig. S1_4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 55 1 1 --
Cladomics cladogram Lampsilis AMBLEMINI

Chen et al., 2023 [Publ 4302], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 1 1 --
Cladomics cladogram Lampsilis QUADRULINI

Chen et al., 2023 [Publ 4302], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 1 1 --
Cladomics cladogram Lampsilis QUADRULINI

Zieritz et al., 2024 [Publ 4381], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 44 1 1 --
Cladomics cladogram AMBLEMINI Lampsilis

Hou et al., 2024 [Publ 4400], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 74 1 1 --
Cladomics cladogram Lampsilis QUADRULINI

Dai et al., 2025 [Publ 4440], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 43 1 1 --
Cladomics cladogram Lampsilis QUADRULINI

Dai et al., 2025 [Publ 4440], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 43 1 1 --
Cladomics cladogram Lampsilis QUADRULINI

Hou et al., 2025 [Publ 4445], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 51 1 1 --
Cladomics cladogram Lampsilis QUADRULINI

Chen et al., 2025 [Publ 4487], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 1 1 --
Cladomics cladogram Lampsilis QUADRULINI

Chen et al., 2025 [Publ 4487], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 1 1 --
Cladomics cladogram QUADRULINI Lampsilis

Schilling et al., 2025 [Publ 4737], fig. 3

cst iT tT bi riS oT roT roS iD
0.857 2 2 0 1 9 9 1 --
Cladomics cladogram Lampsilis PLEUROBEMINI

Schilling et al., 2025 [Publ 4737], fig. 4

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 9 9 1 --
Cladomics cladogram Lampsilis PLEUROBEMINI

Baker et al., 2003 [Publ 2690], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 0 0 --
Cladomics cladogram Epioblasma Hyridella australis RHIPIDODONTINI Hyridella depressa Velesunio Echyridella CASTALIINI

Baker et al., 2004 [Publ 2685], fig. 2-3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 0 0 --
Cladomics cladogram Epioblasma Hyridella depressa Hyridella australis RHIPIDODONTINI Echyridella CASTALIINI Velesunio 3 Velesunio 4 Velesunio 2 Velesunio angasi Velesunio 1 Alathyria

Huff et al., 2004 [Publ 2697], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 0 0 --
Cladomics cladogram Lampsilis UNIONINI ANODONTINI

Huff et al., 2004 [Publ 2697], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 0 0 --
Cladomics cladogram Lampsilis UNIONINI ANODONTINI

Huff et al., 2004 [Publ 2697], fig. 5-6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 0 0 --
Cladomics cladogram UNIONINI ANODONTINI Lampsilis

Jones et al., 2006 [Publ 2681], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 5 5 4 1 0 0 0 117.0
Cladomics cladogram Epioblasma

Cyr et al., 2007 [Publ 2745], fig. 2A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 4 0 0 --
Cladomics cladogram Lampsilis ANODONTINI

Jones & Neves, 2010 [Publ 2646], fig. 16

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 1 0 0 0 116.0
Cladomics cladogram Epioblasma

Jones & Neves, 2010 [Publ 2646], fig. 17

cst iT tT bi riS oT roT roS iD
1.000 7 7 1 1 0 0 0 118.0
Cladomics cladogram Epioblasma

Plazzi & Passamonti, 2010 [Publ 3354], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 1 0 0 --
Cladomics cladogram LAMPROTULINI Lampsilis

Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 0 0 --
Cladomics cladogram ANODONTINI Lampsilis CHAMBERLAINIINI LAMPROTULINI

Jiang et al., 2010 [Publ 3438], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 0 0 --
Cladomics cladogram Lampsilis ANODONTINI LAMPROTULINI CHAMBERLAINIINI

Moyer & Díaz-Ferguson, 2012 [Publ 3375], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 3 3 2 1 3 0 0 118.0
Cladomics cladogram ANODONTINI Lampsilis

Takeuchi et al., 2015 [Publ 3661], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 0 0 --
Cladomics cladogram Lampsilis Pseudunio marocanus Pseudunio auricularius Cumberlandia Margaritifera

Takeuchi et al., 2015 [Publ 3661], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 0 0 --
Cladomics cladogram Lampsilis Pseudunio auricularius 1 Pseudunio auricularius 2 Cumberlandia Margaritifera

Bolotov et al., 2016 [Publ 3743], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 0 0 --
Cladomics cladogram Lampsilis ANODONTINI UNIONINI

Bolotov et al., 2016 [Publ 3743], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 0 0 --
Cladomics cladogram Lampsilis ANODONTINI UNIONINI

Combosch et al., 2017 [Publ 3806], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 212 0 0 --
Cladomics cladogram Lampsilis INDOCHINELLINI

Combosch et al., 2017 [Publ 3806], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 213 0 0 --
Cladomics cladogram INDOCHINELLINI Lampsilis

Vikhrev et al., 2018 [Publ 3812], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 12 0 0 --
Cladomics cladogram UNIONINI Lampsilis

Huang et al., 2018 [Publ 3828], fig. 3_5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 28 0 0 --
Cladomics cladogram Lampsilis INDOCHINELLINI

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 27 0 0 --
Cladomics cladogram Lampsilis UNIONINI

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 27 0 0 --
Cladomics cladogram Lampsilis UNIONINI

Keogh & Simons, 2019 [Publ 3935], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 6 6 6 1 0 0 0 117.0
Cladomics cladogram Lampsilis

Lemer et al., 2019 [Publ 4093], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram Margaritifera Lampsilis

Wu et al., 2020 [Publ 4121], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 0 0 --
Cladomics cladogram Lampsilis INDOCHINELLINI

Wu et al., 2020 [Publ 4121], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 0 0 --
Cladomics cladogram Lampsilis INDOCHINELLINI

Sano et al., 2020 [Publ 4138], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 68 0 0 --
Cladomics cladogram Lampsilis CHAMBERLAINIINI Inversidens LAMPROTULINI Ptychorhynchus Obovalis

Sano et al., 2020 [Publ 4138], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 68 0 0 --
Cladomics cladogram Lampsilis LAMPROTULINI GONIDEINI

Sano et al., 2020 [Publ 4138], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 68 0 0 --
Cladomics cladogram Lampsilis CHAMBERLAINIINI Inversidens LAMPROTULINI Obovalis Ptychorhynchus

Wu et al., 2020 [Publ 4147], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram Lampsilis UNIONINI GONIDEINI LAMPROTULINI

Smith & Johnson, 2020 [Publ 4229], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 0 0 0 116.0
Cladomics cladogram Potamilus

Stewart et al., 2021 [Publ 4451], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 13 0 0 118.0
Cladomics cladogram Toxolasma ANODONTINI

Gomes-dos-Santos et al., 2023 [Publ 4463], fig. 2a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 20 0 0 --
Cladomics cladogram Lampsilis LAMPROTULINI ANODONTINI UNIONINI

Gomes-dos-Santos et al., 2023 [Publ 4463], fig. 2b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 0 0 --
Cladomics cladogram Lampsilis LAMPROTULINI ANODONTINI UNIONINI

Chen et al., 2025 [Publ 4455], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 25 0 0 --
Cladomics cladogram Lampsilis LAMPROTULINI