|
Data last updated
Mon 1 Dec 2025
|
Bivalvia
| Unionoida
| Unionoidea
| Unionidae
| Ambleminae
tribe LAMPSILINI Ihering, 1901: cladomics32 genera
Cladomics pages are experimental — even more so than the rest of this site!
The purpose of the cladomics pages for each genus and family-group level taxon is to
provide statistics and references indicating the extent to which published phylogenetic
work supports the classification used here. The Cladomics Summary provides an
overview of the number of studies that have included terminal species representing this
taxon as well as an indication of how frequently it has been recovered as monophyletic.
Following the summary, the available published clades representing the taxon are listed.
For more information, visit the
Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various
statistics are explained on the FARQ!
Cladomics Summary
number of species = 155 (180 total)
number of subtaxa (rank: genus) = 32
analyzed in 270 cladograms in 160 publications
maximum number of terminals in an analysis = 48 (mean = 5.68, median = 3)
maximum number of subtaxa (genus) = 23 (mean = 3.84, median = 3)
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 196
maximum number of included terminals = 48, (mean = 7.45 median = 5)
maximum number of subtaxa (genus) = 23 (mean = 4.91, median = 4)
proportion of cladograms that support basic monophyly = 0.78
Rigorous Monophyly
number of cladograms that test rigorous monophyly = 168
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 48 (mean = 7.7, median = 5)
maximum number of subtaxa (genus) = 23 (mean = 5.28, median = 4)
proportion of cladograms that support rigorous monophyly = 0.74
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome.
See the Cladmoics FARQ for more information.
Cladograms
** Top Monophyly Cladogram **
iT| tT | bi | riS | oT | roT | roS | iD |
| 43 |
43 |
46 |
23 |
59 |
47 |
4 |
0 |
+----------------------------------------- Quadrulini
|
| +-------------------------------------- Pleurobemini
--+ |
| | +-- Plectomerus
| | +--------------------------------+
+--+ | +-- Reginaia
| |
| | +-- Popenaiadini
| | +-----------------------------+
+--+ | +-- Amblema
| |
| | +-------- Toxolasma **
| | +--------------------+
| | | | +----- Obliquaria **
| | | +--+
+--+ | | +-- Cyrtonaias **
| | +--+
| | +-- Glebula **
| |
| | +-------------- Potamilus **
| | |
| | | +-------- Medionidus_acutissimus **
+--+ | +--+
| +-----------+ | | +----- Lemiox **
| | | | +--+
| | | | | +-- Dromus **
| | | | +--+
| | +--+ +-- Cyprogenia **
| | |
| | | +-- Sagittunio **
| | | +--+
| | | | +-- Medionidus_conradicus **
+--+ +-----+
| | +-- Ptychobranchus **
| +--+
| +-- Venustaconcha_ellipsiformis__2 **
|
| +-- Ellipsaria **
| +--------------------+
| | +-- Truncilla **
| |
| | +-------------------- Villosa **
+--+ |
| | +-- Obovaria_olivaria **
| | +--------+
| | | +-- Venustaconcha_ellipsiformis__1 **
| | +-----+
+--+ | | +-------- Epioblasma **
| | +--+
| | | +----- Obovaria_unicolor **
| | +--+
| | | +-- Obovaria_arkansasensis **
| | +--+
| | +-- Obovaria_subrotunda **
+--+
| +-------- Hamiota **
| |
| +-----+ +-- Lampsilis_cardium **
| | | +--+
| | | | +-- Lampsilis_ovata **
| | +--+
| | | +-- Ortmanniana_pectorosa **
+--+ +--+
| +-- Lampsilis_ornata **
|
| +-- Ligumia **
| +--------+
| | +-- Cambarunio **
+--+
| +-------- Lampsilis_teres **
+--+
| +----- Ortmanniana_ligamentina **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Leaunio **
** Top Sister-Group Cladogram **
iT| tT | bi | riS | oT | roT | roS | iD |
| 11 |
12 |
0 |
8 |
93 |
71 |
4 |
51 |
+-------- Reginaia
|
+-------- Amblema
|
| +-- Quadrulini
+-----+
| +-- Pleurobemini
--+
| +----- Popenaiadini
| |
| | +-- Dromus **
| | |
| | +-- Epioblasma **
| | |
+--+ +-- Glebula **
| |
| +-- Lampsilis_ovata **
| |
| +-- Lampsilis_virescens **
| |
| +-- Lemiox **
+--+
+-- Obliquaria **
|
+-- Obovaria_olivaria **
|
+-- Obovaria_unicolor **
|
+-- Plectomerus
|
+-- Potamilus_alatus **
|
+-- Potamilus_capax **
iT| tT | bi | riS | oT | roT | roS | iD |
| 48 |
48 |
42 |
23 |
9 |
7 |
3 |
45 |
+-------------------------------------------- Pleurobemini
|
| +----------------------------------------- Amblemini
--+ |
| +----------------------------------------- Popenaiadini
| |
| | +-- Paetulunio **
+--+ +-----------------------------------+
| | +-- Toxolasma **
| |
| | +----- Obliquaria **
| | +-----------------------------+
+--+ | | +-- Glebula **
| | +--+
| | +-- Cyrtonaias **
| |
| | +----- Ptychobranchus **
| | +--+
+--+ | | +-- Dromus **
| | +--+
| +-----------------------+ +-- Cyprogenia **
| | |
| | | +----- Medionidus_conradicus **
| | +--+
| | | +-- Medionidus_acutissimus **
| | +--+
+--+ +-- Lemiox **
|
| +----------------------------- Sagittunio **
| |
| +----------------------------- Potamilus **
| |
| | +-- Truncilla **
+--+ +-----------------------+
| | +-- Ellipsaria **
| |
| | +-- Ortmanniana_pectorosa **
| | +-----+
| | | +-- Lampsilis_fasciola **
+--+ +--------------+
| | | +----- Lampsilis_ornata **
| | +--+
| | | +-- Lampsilis_ovata **
| | +--+
| | +-- Lampsilis_cardium **
+--+
| +-- Ligumia **
| +--+
| | +-- Cambarunio **
| +--------------+
| | | +-- Lampsilis_teres **
| | +--+
+--+ +-- Hamiota **
|
| +----------------- Leaunio **
| |
+--+ +-------------- Ortmanniana_ligamentina **
| |
+--+ +----------- Lampsilis_siliquoidea **
| |
+--+ +-------- Obovaria_olivaria **
| |
| +-------- Venustaconcha **
+--+
+-------- Epioblasma **
|
| +----- Obovaria_unicolor **
+--+
| +-- Obovaria_subrotunda **
+--+
+-- Obovaria_arkansasensis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 48 |
48 |
45 |
23 |
8 |
6 |
3 |
47 |
+----- Pleurobemini
+--------------------------------+
| | +-- Amblemini
| +--+
| +-- Popenaiadini
--+
| +-- Cyrtonaias **
| +--------------------------------+
| | +-- Glebula **
| |
+--+ +-------------------------------- Paetulunio **
| |
| | +-- Obliquaria **
+--+ +--------------------------+
| | +-- Toxolasma **
| |
| | +-------------------------- Sagittunio **
+--+ |
| | +----------- Potamilus **
| | |
| | +-----------+ +-- Medionidus_conradicus **
| | | | +-----+
+--+ | | | +-- Ptychobranchus **
| | +--+
| | | +----- Medionidus_acutissimus **
| | | |
| | +--+----- Lemiox **
| | |
| | | +-- Dromus **
+--+ +--+
| +-- Cyprogenia **
|
| +----- Lampsilis_teres **
| +--------------+
| | | +-- Truncilla **
| | +--+
| | +-- Ellipsaria **
| |
+--+ +----------------- Leaunio **
| |
| | +----------- Hamiota **
| | |
| | +--+ +-- Lampsilis_fasciola **
+--+ | | +-----+
| | | | +-- Ortmanniana_pectorosa **
| | +--+
| | | +----- Lampsilis_ornata **
| | +--+
| | | +-- Lampsilis_ovata **
+--+ +--+
| +-- Lampsilis_cardium **
|
| +----- Venustaconcha **
| +-----+
| | | +-- Obovaria **
| | +--+
+--+ +-- Epioblasma **
|
| +-------- Lampsilis_siliquoidea **
+--+
| +----- Ortmanniana_ligamentina **
+--+
| +-- Cambarunio **
+--+
+-- Ligumia **
iT| tT | bi | riS | oT | roT | roS | iD |
| 43 |
44 |
37 |
23 |
59 |
47 |
4 |
38 |
+-------------------------- Quadrulini
|
| +----------------------- Pleurobemini
--+ |
| | +----- Popenaiadini
| | +--------------+
| | | | +-- Amblema
+--+ | +--+
| | +-- Reginaia
| |
| | +-- Plectomerus
| | +--------------+
+--+ | +-- Toxolasma **
| |
| | +----- Obliquaria **
| | +--------+
| | | | +-- Cyrtonaias **
| | | +--+
| | | +-- Glebula **
| | |
+--+ | +----------- Sagittunio **
| | |
| | +----------- Leaunio **
| | |
| | +----------- Potamilus **
| | |
| | +----------- Hamiota **
| | |
| | | +-- Ellipsaria **
+--+ +--------+
| | +-- Truncilla **
| |
| | +----- Lampsilis_teres **
| +-----+
| | | +-- Ortmanniana_ligamentina **
| | +--+
| | +-- Lampsilis_siliquoidea **
| |
| | +----- Villosa **
| +-----+
| | | +-- Ligumia **
| | +--+
| | +-- Cambarunio **
| |
+--+ +-------- Ortmanniana_pectorosa **
+--+
| | +----- Lampsilis_ornata **
| +--+
| | +-- Lampsilis_cardium **
| +--+
| +-- Lampsilis_ovata **
|
| +-------- Obovaria_olivaria **
| |
| +-------- Venustaconcha_ellipsiformis__2 **
+--+
| +-------- Epioblasma **
| |
| | +----- Obovaria_unicolor **
| +--+
| | +-- Obovaria_arkansasensis **
| +--+
| +-- Obovaria_subrotunda **
|
| +-- Ptychobranchus **
| +-----+
| | +-- Venustaconcha_ellipsiformis__1 **
| |
+--+ +----- Medionidus_conradicus **
| |
| | +-- Lemiox **
+--+--+
| +-- Medionidus_acutissimus **
|
| +-- Dromus **
+--+
+-- Cyprogenia **
iT| tT | bi | riS | oT | roT | roS | iD |
| 36 |
37 |
36 |
20 |
5 |
5 |
3 |
58 |
+-------------------------------------- Quadrulini
|
| +-- Amblema
--+ +--------------------------------+
| | +-- Reginaia
| |
+--+ +-------------------------------- Plectomerus
| |
| | +----------------------------- Toxolasma **
+--+ |
| | +----- Popenaiadini
| | +--------------------+
+--+ | | +-- Glebula **
| | +--+
| | +-- Cyrtonaias **
| |
+--+ +----------------------- Obliquaria **
| |
| | +-- Medionidus **
| | +--+
| | | +-- Lemiox **
+--+ +--------------+
| | | +-- Cyprogenia **
| | +--+
| | +-- Ptychobranchus **
| |
+--+ +-------- Potamilus **
| +--------+
| | | +----- Atlanticoncha **
| | +--+
| | | +-- Ellipsaria **
| | +--+
+--+ +-- Truncilla **
|
| +-- Hamiota **
| +-----------+
| | +-- Villosa **
| |
+--+ +----------- Ligumia **
| |
| | +----- Venustaconcha **
+--+ +--+
| | | +-- Obovaria **
| | +--+
+--+ +-- Epioblasma **
|
| +----- Ortmanniana_pectorosa **
+--+
| +-- Ortmanniana_ligamentina **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 27 |
27 |
27 |
7 |
15 |
14 |
3 |
57 |
+-------------------------------------------- Quadrulini
|
--+ +----------------------------------------- Pleurobemini
| |
+--+ +-------------------------------------- Amblemini
| |
+--+ +----------------------------------- Toxolasma **
| |
+--+ +-------------------------------- Potamilus **
| |
| | +-- Ellipsaria **
+--+ +--------------------------+
| | +-- Truncilla **
| |
| | +-------------------------- Leaunio **
+--+ |
| | +-------------------- Lampsilis_bracteata **
| | |
| | +--+ +----------------- Lampsilis_fasciola **
| | | | |
+--+ | +--+ +-------------- Lampsilis_ornata **
| | | |
| | +--+ +-- Lampsilis_ovata **
| | | +--------+
| | | | +-- Lampsilis_cariosa **
| | +--+
| | | +-------- Lampsilis_cardium__3 **
+--+ +--+
| | +----- Lampsilis_cardium__2 **
| +--+
| | +-- Lampsilis_satura **
| +--+
| +-- Lampsilis_cardium__1 **
|
| +-- Lampsilis_higginsii **
| +--------------+
| | +-- Ortmanniana **
+-----+
| +-------------- Lampsilis_teres **
| |
+--+ +-- Lampsilis_radiata **
| +--------+
| | +-- Lampsilis_siliquoidea **
+--+
| +-------- Lampsilis_bergmanni **
+--+
| +----- Lampsilis_hydiana **
+--+
| +-- Lampsilis_virescens **
+--+
+-- Lampsilis_straminea **
iT| tT | bi | riS | oT | roT | roS | iD |
| 26 |
30 |
18 |
18 |
7 |
7 |
4 |
56 |
+-------------------------- Quadrulini
|
| +----------------------- Plectomerus
--+ |
| | +-------------------- Popenaiadini
| | |
| | +-------------------- Pleurobemini
+--+ |
| | +-- Cyrtonaias **
| | +--+
| | | +-- Glebula **
| +--------------+
| | | +-- Reginaia
+--+ +--+
| +-- Obliquaria **
|
| +----------------- Toxolasma **
| |
| | +-------------- Potamilus **
| | |
| | | +-- Ellipsaria **
| | +-----------+
+--+ | +-- Truncilla **
| |
| | +----- Medionidus **
| +--------+
| | | +-- Lemiox **
| | +--+
+--+ +-- Dromus **
|
| +----------- Cambarunio **
| |
| +----------- Leaunio **
| |
| +----------- Villosa **
| |
| | +-- Sagittunio **
| +--------+
+--+ +-- Lampsilis_ovata **
|
| +----- Venustaconcha **
+-----+
| | +-- Epioblasma **
| +--+
| +-- Obovaria **
|
| +-------- Lampsilis_teres **
+--+
| +----- Lampsilis_siliquoidea **
+--+
| +-- Lampsilis_hydiana **
+--+
+-- Lampsilis_straminea **
iT| tT | bi | riS | oT | roT | roS | iD |
| 26 |
26 |
26 |
7 |
15 |
14 |
3 |
61 |
+----------------------------------------------- Quadrulini
|
--+ +-------------------------------------------- Reginaia
| |
+--+ +----------------------------------------- Pleurobemini
| |
+--+ +-------------------------------------- Amblema
| |
+--+ +----------------------------------- Toxolasma **
| |
+--+ +-------------------------------- Ellipsaria **
| |
+--+ +----------------------------- Truncilla **
| |
| | +-------------------------- Potamilus **
+--+ |
| | +----------------------- Leaunio **
| | |
+--+ | +----------------- Lampsilis_bracteata **
| | |
| | +--+ +-- Lampsilis_ornata **
| | | | +-----------+
+--+ | | | +-- Lampsilis_fasciola **
| | +--+
| | | +----------- Lampsilis_ovata **
| | +--+
| | | +-------- Lampsilis_cardium__3 **
| | +--+
+--+ | +----- Lampsilis_cardium__2 **
| +--+
| | +-- Lampsilis_satura **
| +--+
| +-- Lampsilis_cardium__1 **
|
| +-------------- Lampsilis_teres **
| |
| | +-- Lampsilis_higginsii **
+-----+ +--------+
| | +-- Ortmanniana **
| |
+--+ +-- Lampsilis_radiata **
| +-----+
| | +-- Lampsilis_siliquoidea **
| |
+--+ +-- Lampsilis_hydiana **
| +--+
| | +-- Lampsilis_bergmanni **
+--+
| +-- Lampsilis_virescens **
+--+
+-- Lampsilis_straminea **
iT| tT | bi | riS | oT | roT | roS | iD |
| 20 |
20 |
18 |
8 |
8 |
8 |
4 |
52 |
+-------------------------- Quadrulini
|
--+ +----------------------- Pleurobemini
| |
+--+ +-------------------- Amblemini
| |
+--+ +----------------- Popenaiadini
| |
+--+ +-------------- Obliquaria **
| |
+--+ +----------- Cyrtonaias **
| |
| | +-- Pachynaias **
+--+ +-----+
| | +-- Lampsilis **
| |
+--+ +-- Ellipsaria **
| +--+
| | +-- Truncilla **
+--+
| +-- Atlanticoncha **
+--+
+-- Potamilus **
iT| tT | bi | riS | oT | roT | roS | iD |
| 20 |
20 |
18 |
8 |
8 |
8 |
4 |
52 |
+-------------------------- Quadrulini
|
--+ +----------------------- Pleurobemini
| |
+--+ +-------------------- Amblemini
| |
+--+ +----------------- Popenaiadini
| |
+--+ +-------------- Obliquaria **
| |
+--+ +----------- Cyrtonaias **
| |
| | +-- Pachynaias **
+--+ +-----+
| | +-- Lampsilis **
| |
+--+ +-- Ellipsaria **
| +--+
| | +-- Truncilla **
+--+
| +-- Atlanticoncha **
+--+
+-- Potamilus **
iT| tT | bi | riS | oT | roT | roS | iD |
| 17 |
18 |
13 |
6 |
1 |
1 |
1 |
52 |
+-------------------- Obliquaria **
|
+-------------------- Amblemini
|
| +----------------- Lampsilis_teres **
| |
| +----------------- Obovaria **
--+ |
| | +-- Leaunio **
| +--------------+
| | +-- Cambarunio **
| |
| | +-------- Ortmanniana **
| | |
+--+--------+ +-- Lampsilis_hydiana **
| | +--+
| | | +-- Lampsilis_virescens **
| +--+
| | +-- Lampsilis_siliquoidea **
| +--+
| +-- Lampsilis_radiata **
|
| +-------------- Lampsilis_bracteata **
| |
+--+ +----------- Lampsilis_ornata **
| |
+--+ +-------- Lampsilis_ovata **
| |
+--+ +----- Lampsilis_cardium__3 **
| |
+--+ +-- Lampsilis_satura **
| |
+--+-- Lampsilis_cardium__1 **
|
+-- Lampsilis_cardium__2 **
iT| tT | bi | riS | oT | roT | roS | iD |
| 17 |
17 |
15 |
6 |
1 |
1 |
1 |
54 |
+-------------------------- Amblemini
|
| +----------------------- Obliquaria **
--+ |
| | +-- Cambarunio **
| | +-----------------+
| | | +-- Leaunio **
+--+ |
| | +----------- Lampsilis_teres **
| | |
| +--------+ +-------- Ortmanniana **
| | | |
| | +--+ +-- Lampsilis_hydiana **
+--+ | +--+
| | | +-- Lampsilis_virescens **
| +--+
| | +-- Lampsilis_siliquoidea **
| +--+
| +-- Lampsilis_radiata **
|
| +----------------- Obovaria **
| |
+--+ +-------------- Lampsilis_bracteata **
| |
+--+ +----------- Lampsilis_ornata **
| |
+--+ +-------- Lampsilis_ovata **
| |
+--+ +----- Lampsilis_cardium__3 **
| |
+--+ +-- Lampsilis_satura **
| |
+--+-- Lampsilis_cardium__1 **
|
+-- Lampsilis_cardium__2 **
iT| tT | bi | riS | oT | roT | roS | iD |
| 16 |
16 |
14 |
2 |
0 |
0 |
0 |
51 |
+-- Truncilla **
--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 16 |
22 |
18 |
10 |
22 |
13 |
3 |
49 |
+-------------------------------- Quadrulina
|
--+ +----------------------------- Megalonaias
| |
+--+ +-------------------------- Toxolasma **
| |
+--+ +----------------------- Amblema
| |
| | +-------------------- Pleurobemini
+--+ |
| | +-- Reginaia
| +-----------------+
+--+ +-- Obliquaria **
|
| +----------------- Potamilus **
| |
| | +-- Ellipsaria **
+--+ +-----------+
| | +-- Truncilla **
| |
+--+ +----------- Lampsilis_cardium **
| |
| | +----- Ligumia **
| +-----+
+--+ | +-- Obovaria **
| +--+
| +-- Epioblasma **
|
| +-------- Lampsilis_teres **
+--+
| +----- Lampsilis_siliquoidea **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis_higginsii **
iT| tT | bi | riS | oT | roT | roS | iD |
| 16 |
16 |
9 |
10 |
19 |
13 |
3 |
50 |
+-------------------- Reginaia
|
--+ +-- Pleurobemini
| +--------------+
| | +-- Quadrulini
+--+
| +-------------- Amblema
| |
+--+ +----------- Toxolasma **
| |
| | +-------- Obliquaria **
+--+ |
| | +----- Obovaria **
| | |
| | +----- Lampsilis_cardium **
+--+ |
| +----- Potamilus **
| |
| | +-- Epioblasma **
| +--+
| | +-- Lampsilis_siliquoidea **
+--+
| +-- Ortmanniana **
+--+
| +-- Lampsilis_higginsii **
|
| +-- Ligumia **
+--+
| +-- Lampsilis_teres **
|
| +-- Ellipsaria **
+--+
+-- Truncilla **
iT| tT | bi | riS | oT | roT | roS | iD |
| 16 |
16 |
12 |
10 |
1 |
1 |
1 |
45 |
+----------------- Amblemini
|
| +-------- Lemiox **
| +-----+
--+ | | +----- Medionidus **
| | +--+
| | | +-- Dromus **
| | +--+
+--+ +-- Cyprogenia **
|
| +----------- Potamilus **
| |
| | +----- Lampsilis_siliquoidea **
| | +--+
+--+ | | +-- Lampsilis_higginsii **
| | +--+
| | +-- Ortmanniana **
| |
| | +----- Lampsilis_fasciola **
+--+--+
| | +-- Lampsilis_cardium **
| +--+
| +-- Lampsilis_ovata **
|
| +-- Cambarunio **
| |
+-----+-- Ligumia **
|
+-- Epioblasma **
iT| tT | bi | riS | oT | roT | roS | iD |
| 15 |
16 |
14 |
11 |
58 |
58 |
4 |
48 |
+----------------------------------- Quadrulini
|
--+ +-------------------------------- Pleurobemini
| |
| | +-- Amblema
+--+ +--------------------------+
| | +-- Reginaia
| |
+--+ +-------------------------- Popenaiadini
| |
| | +-- Glebula **
+--+ +--------------------+
| | +-- Paetulunio **
| |
+--+ +-- Toxolasma **
| +-----------------+
| | +-- Plectomerus
| |
+--+ +----------------- Obliquaria **
| |
| | +----- Potamilus **
+--+ +--------+
| | | +-- Dromus **
| | +--+
+--+ +-- Lemiox **
|
| +----------- Lampsilis_ovata **
| |
+--+ +-------- Lampsilis_virescens **
| |
+--+ +-- Epioblasma **
| +--+
| | +-- Obovaria_arkansasensis **
+--+
| +-- Obovaria_olivaria **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 15 |
15 |
11 |
10 |
1 |
1 |
1 |
48 |
+-------------------------- Amblemini
|
--+ +----------------------- Ellipsaria **
| |
+--+ +-------------------- Truncilla **
| |
+--+ +----------------- Obliquaria **
| |
| | +-------------- Potamilus **
+--+ |
| | +----- Lampsilis_ornata **
| | +-----+
| | | | +-- Lampsilis_cardium **
+--+ | +--+
| | +-- Lampsilis_ovata **
| |
| | +-------- Lampsilis_teres **
| +--+
+--+ | +----- Lampsilis_siliquoidea **
| +--+
| | +-- Ortmanniana **
| +--+
| +-- Lampsilis_higginsii **
|
| +----- Hamiota **
| |
+-----+ +-- Epioblasma **
| |
+--+-- Cambarunio **
|
+-- Ligumia **
iT| tT | bi | riS | oT | roT | roS | iD |
| 15 |
15 |
9 |
11 |
58 |
58 |
4 |
51 |
+-------------------------- Quadrulini
|
--+ +----------------------- Pleurobemini
| |
| | +-- Plectomerus
+--+ +-----------------+
| | +-- Reginaia
| |
| | +-- Popenaiadini
+--+ +--------------+
| | +-- Amblema
| |
| | +-------------- Potamilus_alatus **
| | |
+--+ +-------------- Potamilus_capax **
| |
| +-------------- Glebula **
| |
| | +-- Toxolasma **
| +-----------+
+--+ +-- Paetulunio **
|
| +-- Dromus **
+-----------+
| +-- Lemiox **
|
| +-- Lampsilis_ovata **
| +--------+
| | +-- Obliquaria **
+--+
| +-------- Lampsilis_virescens **
| |
+--+ +-- Epioblasma **
| +--+
| | +-- Obovaria_arkansasensis **
+--+
| +-- Obovaria_olivaria **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 14 |
15 |
10 |
12 |
187 |
11 |
3 |
49 |
+-------------------------- Quadrulini
|
--+ +----------------------- Pleurobemini
| |
+--+ +-------------------- Amblema
| |
| | +----- Toxolasma **
+--+ +-----------+
| | | +-- Obliquaria **
| | +--+
| | +-- Plectomerus
+--+
| +-------------- Potamilus_leptodon **
| |
| | +----------- Potamilus_alatus **
+--+ |
| | +-------- Venustaconcha **
| | |
+--+ +-------- Leaunio **
| |
| | +-- Ellipsaria **
| +-----+
+--+ +-- Truncilla **
|
| +-- Medionidus **
+-----+
| +-- Ptychobranchus **
|
| +----- Cambarunio **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 14 |
21 |
8 |
12 |
187 |
11 |
3 |
54 |
+-------- Amblema
|
+-------- Toxolasma **
|
+-------- Pleurobemini
|
| +-- Obliquaria **
+-----+
| +-- Plectomerus
--+
| +----- Ellipsaria **
| |
| +----- Potamilus_leptodon **
| |
| +----- Medionidus **
| |
| +----- Potamilus_alatus **
| |
| +----- Ptychobranchus **
+--+
+----- Truncilla **
|
+----- Venustaconcha **
|
+----- Cambarunio **
|
+----- Leaunio **
|
| +-- Ortmanniana **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 14 |
15 |
13 |
12 |
185 |
11 |
3 |
50 |
+----------------------------------- Quadrulini
|
--+ +-------------------------------- Pleurobemini
| |
+--+ +----------------------------- Amblema
| |
+--+ +-------------------------- Toxolasma **
| |
+--+ +-- Obliquaria **
| +--------------------+
| | +-- Plectomerus
+--+
| +-------------------- Potamilus_leptodon **
| |
+--+ +----------------- Potamilus_alatus **
| |
| | +-- Medionidus **
+--+ +-----------+
| | +-- Ptychobranchus **
| |
+--+ +-- Ellipsaria **
| +--------+
| | +-- Truncilla **
+--+
| +-------- Leaunio **
| |
+--+ +-- Venustaconcha **
| +--+
| | +-- Cambarunio **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 13 |
13 |
11 |
11 |
8 |
7 |
4 |
47 |
+----------------------------------------- Quadrulini
|
--+ +-------------------------------------- Pleurobemini
| |
+--+ +----------------------------------- Amblema
| |
+--+ +-------------------------------- Plectomerus
| |
+--+ +----------------------------- Popenaiadini
| |
+--+ +-------------------------- Toxolasma **
| |
+--+ +----------------------- Obliquaria **
| |
| | +-- Cyrtonaias **
+--+ +-----------------+
| | +-- Glebula **
| |
+--+ +-- Ptychobranchus **
| +--------------+
| | +-- Lemiox **
| |
+--+ +-- Venustaconcha **
| +-----------+
| | +-- Obovaria **
+--+
| +----------- Hamiota **
+--+
| +-------- Lampsilis_ovata **
+--+
| +----- Ortmanniana **
+--+
| +-- Lampsilis_straminea **
+--+
+-- Lampsilis_hydiana **
iT| tT | bi | riS | oT | roT | roS | iD |
| 13 |
13 |
11 |
11 |
8 |
7 |
4 |
47 |
+----------------------------------------- Quadrulini
|
--+ +-------------------------------------- Pleurobemini
| |
+--+ +----------------------------------- Amblema
| |
+--+ +-------------------------------- Plectomerus
| |
+--+ +----------------------------- Popenaiadini
| |
+--+ +-------------------------- Toxolasma **
| |
+--+ +----------------------- Obliquaria **
| |
| | +-- Cyrtonaias **
+--+ +-----------------+
| | +-- Glebula **
| |
+--+ +-- Ptychobranchus **
| +--------------+
| | +-- Lemiox **
| |
+--+ +-- Venustaconcha **
| +-----------+
| | +-- Obovaria **
+--+
| +----------- Hamiota **
+--+
| +-------- Lampsilis_ovata **
+--+
| +----- Ortmanniana **
+--+
| +-- Lampsilis_straminea **
+--+
+-- Lampsilis_hydiana **
iT| tT | bi | riS | oT | roT | roS | iD |
| 12 |
12 |
6 |
9 |
18 |
5 |
3 |
49 |
+-- Quadrulini
+--------------+
| +-- Pleurobemini
|
--+ +-------------- Amblemini
| |
| | +----------- Ligumia **
| | |
+--+ +----------- Truncilla **
| |
| | +-- Ptychobranchus **
| +--------+
+--+ +-- Sagittunio **
|
| +-- Lampsilis_siliquoidea **
| +-----+
| | +-- Epioblasma **
| |
+--+ +----- Ortmanniana **
| |
| +----- Cambarunio **
+--+
+----- Leaunio **
|
| +-- Lampsilis_cardium **
+--+
+-- Lampsilis_fasciola **
iT| tT | bi | riS | oT | roT | roS | iD |
| 12 |
12 |
10 |
9 |
18 |
5 |
3 |
53 |
+-- Quadrulini
+-----------------------+
| +-- Pleurobemini
--+
| +----------------------- Amblemini
| |
+--+ +-------------------- Truncilla **
| |
+--+ +----------------- Ligumia **
| |
| | +-- Ptychobranchus **
+--+ +-----------+
| | +-- Sagittunio **
| |
+--+ +----------- Lampsilis_siliquoidea **
| |
| | +----- Ortmanniana **
+--+ +--+
| | | +-- Lampsilis_cardium **
| | +--+
+--+ +-- Lampsilis_fasciola **
|
| +----- Epioblasma **
+--+
| +-- Cambarunio **
+--+
+-- Leaunio **
iT| tT | bi | riS | oT | roT | roS | iD |
| 12 |
12 |
5 |
12 |
11 |
11 |
3 |
48 |
+----------------------- Quadrulini
|
--+ +-------------------- Reginaia
| |
+--+ +----------------- Amblema
| |
+--+ +-------------- Popenaiadini
| |
| | +-- Glebula **
+--+ +--------+
| | +-- Cyrtonaias **
| |
+--+ +-------- Potamilus **
| |
| | +-- Ptychobranchus **
+--+ +--+
| | +-- Truncilla **
| |
| | +-- Ortmanniana **
+--+ |
| +-- Epioblasma **
| |
| +-- Lampsilis **
| |
+--+-- Ligumia **
|
+-- Cambarunio **
|
+-- Pachynaias **
|
+-- Mesonaias **
iT| tT | bi | riS | oT | roT | roS | iD |
| 12 |
12 |
10 |
8 |
25 |
6 |
3 |
53 |
+-------------------------- Quadrulini
|
--+ +----------------------- Pleurobemini
| |
+--+ +-------------------- Amblemini
| |
+--+ +----------------- Toxolasma **
| |
+--+ +-------------- Ellipsaria **
| |
+--+ +----------- Truncilla **
| |
+--+ +-- Lemiox **
| +-----+
| | +-- Potamilus **
+--+
| +----- Lampsilis **
+--+
| +-- Cambarunio **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 12 |
12 |
10 |
8 |
26 |
5 |
3 |
53 |
+----------------- Toxolasma **
|
--+ +-------------- Venustaconcha **
| |
+--+ +-- Lemiox **
| +--------+
| | +-- Lampsilis **
+--+
| +-------- Potamilus **
+--+
| +----- Cambarunio **
+--+
| +-- Ellipsaria **
+--+
+-- Truncilla **
iT| tT | bi | riS | oT | roT | roS | iD |
| 12 |
12 |
10 |
12 |
19 |
18 |
4 |
51 |
+----------------------------------- Quadrulini
|
--+ +-------------------------------- Pleurobemini
| |
+--+ +----------------------------- Amblemini
| |
+--+ +-------------------------- Popenaiadini
| |
+--+ +-- Cyrtonaias **
| +--------------------+
| | +-- Glebula **
+--+
| +-------------------- Potamilus **
| |
+--+ +-- Truncilla **
| +--------------+
| | +-- Ptychobranchus **
+--+
| +-------------- Pachynaias **
| |
+--+ +----------- Mesonaias **
| |
+--+ +-- Lampsilis **
| +-----+
| | +-- Ortmanniana **
+--+
| +----- Cambarunio **
+--+
| +-- Epioblasma **
+--+
+-- Ligumia **
iT| tT | bi | riS | oT | roT | roS | iD |
| 11 |
11 |
8 |
5 |
81 |
6 |
4 |
48 |
+----------------------------- Quadrulini
|
--+ +-------------------------- Pleurobemini
| |
+--+ +----------------------- Amblemini
| |
+--+ +-------------------- Popenaiadini
| |
+--+ +----------------- Toxolasma **
| |
+--+ +-------------- Potamilus **
| |
+--+ +----------- Leaunio **
| |
+--+ +-------- Venustaconcha **
| |
| | +-- Lampsilis_powellii **
+--+-----+
| +-- Lampsilis_siliquoidea **
|
| +----- Lampsilis_ornata **
+--+
| +-- Lampsilis_cardium **
+--+
+-- Lampsilis_cariosa **
iT| tT | bi | riS | oT | roT | roS | iD |
| 11 |
11 |
9 |
11 |
11 |
11 |
3 |
54 |
+----------------------------------- Quadrulini
|
--+ +-------------------------------- Reginaia
| |
+--+ +----------------------------- Amblema
| |
+--+ +-------------------------- Popenaiadini
| |
+--+ +----------------------- Cyrtonaias **
| |
+--+ +-------------------- Ptychobranchus **
| |
+--+ +----------------- Ligumia **
| |
+--+ +-------------- Potamilus **
| |
+--+ +----------- Pachynaias **
| |
| | +-- Epioblasma **
+--+ +-----+
| | +-- Truncilla **
| |
+--+ +-- Mesonaias **
| +--+
| | +-- Cambarunio **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 11 |
11 |
9 |
5 |
85 |
6 |
4 |
49 |
+----------------------------- Quadrulini
|
--+ +-------------------------- Pleurobemini
| |
+--+ +----------------------- Amblemini
| |
+--+ +-------------------- Popenaiadini
| |
+--+ +----------------- Toxolasma **
| |
+--+ +-------------- Potamilus **
| |
+--+ +----------- Leaunio **
| |
| | +----- Lampsilis_ornata **
+--+ +--+
| | | +-- Lampsilis_cardium **
| | +--+
+--+ +-- Lampsilis_cariosa **
|
| +----- Venustaconcha **
+--+
| +-- Lampsilis_powellii **
+--+
+-- Lampsilis_siliquoidea **
iT| tT | bi | riS | oT | roT | roS | iD |
| 11 |
11 |
9 |
6 |
0 |
0 |
0 |
48 |
+----------- Obliquaria **
|
--+----------- Lampsilis **
|
| +-------- Medionidus **
+--+
| +----- Ptychobranchus **
+--+
| +-- Cyprogenia **
+--+
+-- Dromus **
iT| tT | bi | riS | oT | roT | roS | iD |
| 11 |
11 |
9 |
5 |
85 |
6 |
4 |
49 |
+----------------------------- Quadrulini
|
--+ +-------------------------- Pleurobemini
| |
+--+ +----------------------- Amblemini
| |
+--+ +-------------------- Popenaiadini
| |
+--+ +----------------- Toxolasma **
| |
+--+ +-------------- Potamilus **
| |
+--+ +----------- Leaunio **
| |
| | +----- Lampsilis_ornata **
+--+ +--+
| | | +-- Lampsilis_cardium **
| | +--+
+--+ +-- Lampsilis_cariosa **
|
| +----- Venustaconcha **
+--+
| +-- Lampsilis_powellii **
+--+
+-- Lampsilis_siliquoidea **
iT| tT | bi | riS | oT | roT | roS | iD |
| 11 |
12 |
10 |
8 |
22 |
10 |
3 |
53 |
+-- Megalonaias
+-----------------------+
| +-- Quadrulina
|
| +----------- Elliptoideus
| +-----------+
--+ | | +-------- Pleurobema
| | +--+
| | | +----- Uniomerus
| | +--+
| | | +-- Fusconaia
+--+ +--+
| +-- Eurynia
|
| +-------------------- Amblema
| |
| | +----- Toxolasma **
| | +-----------+
+--+ | | +-- Plectomerus
| | +--+
| | +-- Obliquaria **
| |
+--+ +----- Ptychobranchus **
| +--------+
| | | +-- Potamilus_fragilis **
| | +--+
| | +-- Medionidus **
+--+
| +-- Villosa **
| +--------+
| | +-- Ellipsaria **
+--+
| +-------- Lampsilis_teres **
+--+
| +----- Lampsilis_siliquoidea **
+--+
| +-- Potamilus_purpuratus **
+--+
+-- Potamilus_alatus **
iT| tT | bi | riS | oT | roT | roS | iD |
| 11 |
11 |
9 |
5 |
81 |
6 |
4 |
49 |
+----------------------------- Quadrulini
|
--+ +-------------------------- Pleurobemini
| |
+--+ +----------------------- Amblemini
| |
+--+ +-------------------- Popenaiadini
| |
+--+ +----------------- Toxolasma **
| |
+--+ +-------------- Potamilus **
| |
+--+ +----------- Leaunio **
| |
| | +----- Lampsilis_ornata **
+--+ +--+
| | | +-- Lampsilis_cardium **
| | +--+
+--+ +-- Lampsilis_cariosa **
|
| +----- Venustaconcha **
+--+
| +-- Lampsilis_powellii **
+--+
+-- Lampsilis_siliquoidea **
iT| tT | bi | riS | oT | roT | roS | iD |
| 10 |
13 |
10 |
6 |
7 |
5 |
3 |
52 |
+-- Pleurobemini
+-----------+
| +-- Quadrulini
|
--+ +-- Paetulunio **
| +--------+
| | +-- Amblemini
+--+
| +-------- Lampsilis **
| |
+--+ +----- Leaunio **
| |
+--+----- Obovaria **
|
| +-- Villosa **
+--+
+-- Cambarunio **
iT| tT | bi | riS | oT | roT | roS | iD |
| 10 |
10 |
8 |
5 |
78 |
6 |
4 |
48 |
+----------------------------- Quadrulini
|
--+ +-------------------------- Pleurobemini
| |
+--+ +----------------------- Amblemini
| |
+--+ +-------------------- Popenaiadini
| |
+--+ +----------------- Toxolasma **
| |
+--+ +-------------- Potamilus **
| |
+--+ +----------- Leaunio **
| |
+--+ +-- Lampsilis_cardium **
| +-----+
| | +-- Lampsilis_ornata **
+--+
| +----- Venustaconcha **
+--+
| +-- Lampsilis_powellii **
+--+
+-- Lampsilis_siliquoidea **
iT| tT | bi | riS | oT | roT | roS | iD |
| 10 |
10 |
8 |
5 |
0 |
0 |
0 |
44 |
+----- Ligumia **
|
+----- Obliquaria **
--+
+----- Lampsilis **
|
| +-- Villosa **
+--+
+-- Hamiota **
iT| tT | bi | riS | oT | roT | roS | iD |
| 10 |
10 |
7 |
2 |
0 |
0 |
0 |
54 |
+-- Ortmanniana **
|
+-----------+-- Lampsilis_abrupta **
| |
| +-- Lampsilis_higginsii **
--+
| +----------- Lampsilis_sietmani **
| |
| | +-- Lampsilis_siliquoidea **
+--+ +-----+
| | +-- Lampsilis_radiata **
| |
+--+ +----- Lampsilis_hydiana__2 **
| |
| +----- Lampsilis_straminea__2 **
| |
+--+ +-- Lampsilis_hydiana__1 **
+--+
| +-- Lampsilis_bergmanni **
|
| +-- Lampsilis_straminea__1 **
+--+
+-- Lampsilis_virescens **
iT| tT | bi | riS | oT | roT | roS | iD |
| 9 |
9 |
7 |
5 |
65 |
6 |
4 |
47 |
+----------------------------- Quadrulini
|
--+ +-------------------------- Pleurobemini
| |
+--+ +----------------------- Amblemini
| |
+--+ +-------------------- Popenaiadini
| |
+--+ +----------------- Toxolasma **
| |
+--+ +-------------- Potamilus **
| |
+--+ +----------- Leaunio **
| |
+--+ +-- Lampsilis_cardium **
| +-----+
| | +-- Lampsilis_ornata **
+--+
| +----- Venustaconcha **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Lampsilis_powellii **
iT| tT | bi | riS | oT | roT | roS | iD |
| 9 |
9 |
7 |
7 |
19 |
9 |
3 |
48 |
+-- Megalonaias
+-----------------------------+
| +-- Quadrulina
|
| +-------- Elliptoideus
--+ +--------------------+
| | | +----- Uniomerus
| | +--+
| | | +-- Eurynia
+--+ +--+
| +-- Pleurobema
|
| +-------------------------- Amblema
+--+
| +----------------------- Plectomerus
+--+
| +-------------------- Obliquaria **
+--+
| +----------------- Medionidus **
+--+
| +-------------- Ptychobranchus **
+--+
| +----------- Potamilus_fragilis **
+--+
| +-------- Potamilus_purpuratus **
+--+
| +----- Ellipsaria **
+--+
| +-- Villosa **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 9 |
10 |
7 |
7 |
18 |
9 |
3 |
52 |
+-- Megalonaias
+-----------------------+
| +-- Quadrulina
|
| +----- Elliptoideus
--+ |
| +-----------------+ +-- Eurynia
| | | |
| | +--+-- Pleurobema
| | |
+--+ +-- Uniomerus
|
| +-------------------- Amblema
| |
+--+ +-- Plectomerus
| +--------------+
| | +-- Obliquaria **
+--+
| +-------------- Potamilus_fragilis **
| |
+--+ +----------- Ptychobranchus **
| |
+--+ +-------- Medionidus **
| |
+--+ +----- Potamilus_purpuratus **
| |
+--+----- Lampsilis **
|
| +-- Ellipsaria **
+--+
+-- Villosa **
iT| tT | bi | riS | oT | roT | roS | iD |
| 9 |
9 |
7 |
5 |
65 |
6 |
4 |
47 |
+----------------------------- Quadrulini
|
--+ +-------------------------- Pleurobemini
| |
+--+ +----------------------- Amblemini
| |
+--+ +-------------------- Popenaiadini
| |
+--+ +----------------- Toxolasma **
| |
+--+ +-------------- Potamilus **
| |
+--+ +----------- Leaunio **
| |
+--+ +-- Lampsilis_ornata **
| +-----+
| | +-- Lampsilis_cardium **
+--+
| +----- Venustaconcha **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Lampsilis_powellii **
iT| tT | bi | riS | oT | roT | roS | iD |
| 9 |
15 |
13 |
4 |
6 |
6 |
4 |
51 |
+----------- Toxolasma **
+--+
| | +-------- Popenaiadini
| +--+
| | +----- Amblemini
| +--+
--+ | +-- Quadrulini
| +--+
| +-- Pleurobemini
|
| +----------- Potamilus **
| |
+--+ +-- Lampsilis_cardium **
| +-----+
| | +-- Lampsilis_ornata **
+--+
| +----- Venustaconcha **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Lampsilis_powellii **
iT| tT | bi | riS | oT | roT | roS | iD |
| 9 |
9 |
7 |
5 |
65 |
6 |
4 |
47 |
+----------------------------- Quadrulini
|
--+ +-------------------------- Pleurobemini
| |
+--+ +----------------------- Amblemini
| |
+--+ +-------------------- Popenaiadini
| |
+--+ +----------------- Toxolasma **
| |
+--+ +-------------- Potamilus **
| |
+--+ +----------- Leaunio **
| |
+--+ +-- Lampsilis_ornata **
| +-----+
| | +-- Lampsilis_cardium **
+--+
| +----- Venustaconcha **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Lampsilis_powellii **
iT| tT | bi | riS | oT | roT | roS | iD |
| 9 |
9 |
7 |
4 |
6 |
6 |
3 |
53 |
+-------------------- Quadrula_quadrula
|
+-------------------- Quadrula_verrucosa
--+
| +----------------- Pleurobemini
| |
| | +-------------- Amblemini
+--+ |
| | +-- Obovaria **
| | +--------+
+--+ | +-- Ortmanniana **
| |
| | +-- Obliquaria **
| +--------+
+--+ +-- Lampsilis_ornata **
|
| +----- Lampsilis_radiata__2 **
| +--+
| | | +-- Lampsilis_siliquoidea **
| | +--+
+--+ +-- Lampsilis_radiata__1 **
|
| +----- Lampsilis_virescens **
+--+
| +-- Lampsilis_hydiana **
+--+
+-- Lampsilis_straminea **
iT| tT | bi | riS | oT | roT | roS | iD |
| 9 |
10 |
5 |
3 |
1 |
1 |
1 |
49 |
+-------------- Pleurobemini
|
+-------------- Obliquaria **
--+
| +----------- Potamilus_capax **
| |
| +----------- Lampsilis **
+--+
| +-------- Potamilus_fragilis **
| |
| | +----- Potamilus_inflatus **
+--+ |
| | +-- Potamilus_ohiensis **
| +--+
+--+ +-- Potamilus_amphichaenus **
|
| +-- Potamilus_purpuratus__1 **
| |
+--+-- Potamilus_purpuratus__2 **
|
+-- Potamilus_alatus **
iT| tT | bi | riS | oT | roT | roS | iD |
| 9 |
9 |
7 |
5 |
77 |
6 |
4 |
47 |
+----------------------------- Quadrulini
|
--+ +-------------------------- Pleurobemini
| |
+--+ +----------------------- Amblemini
| |
+--+ +-------------------- Popenaiadini
| |
+--+ +----------------- Toxolasma **
| |
+--+ +-------------- Potamilus **
| |
+--+ +----------- Leaunio **
| |
+--+ +-- Lampsilis_cardium **
| +-----+
| | +-- Lampsilis_ornata **
+--+
| +----- Venustaconcha **
+--+
| +-- Lampsilis_powellii **
+--+
+-- Lampsilis_siliquoidea **
iT| tT | bi | riS | oT | roT | roS | iD |
| 9 |
9 |
7 |
3 |
5 |
3 |
3 |
52 |
+-- Amblemini
+--------------------+
| +-- Pleurobemini
--+
| +-------------------- Medionidus **
| |
+--+ +----------------- Lampsilis_teres **
| |
| +----------------- Villosa **
+--+
| +-- Lampsilis_satura **
| +-----------+
| | +-- Lampsilis_cardium **
+--+
| +----------- Lampsilis_reeveiana **
| |
+--+ +-------- Lampsilis_hydiana__1 **
| |
+--+-------- Lampsilis_hydiana__2 **
|
| +----- Lampsilis_siliquoidea__2 **
+--+
| +-- Lampsilis_siliquoidea__1 **
+--+
+-- Lampsilis_powellii **
iT| tT | bi | riS | oT | roT | roS | iD |
| 9 |
9 |
7 |
5 |
44 |
6 |
4 |
47 |
+----------------------------- Quadrulini
|
--+ +-------------------------- Pleurobemini
| |
+--+ +----------------------- Amblemini
| |
+--+ +-------------------- Popenaiadini
| |
+--+ +----------------- Toxolasma **
| |
+--+ +-------------- Potamilus **
| |
+--+ +----------- Leaunio **
| |
+--+ +-- Lampsilis_cardium **
| +-----+
| | +-- Lampsilis_ornata **
+--+
| +----- Venustaconcha **
+--+
| +-- Lampsilis_powellii **
+--+
+-- Lampsilis_siliquoidea **
iT| tT | bi | riS | oT | roT | roS | iD |
| 8 |
8 |
6 |
3 |
2 |
2 |
2 |
52 |
+-- Amblemini
+-----+
| +-- Pleurobemini
--+
| +----- Potamilus **
+--+
| +-- Leaunio **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 8 |
8 |
6 |
4 |
11 |
6 |
4 |
45 |
+-------------------------- Quadrulini
|
--+ +----------------------- Pleurobemini
| |
+--+ +-------------------- Amblemini
| |
+--+ +----------------- Popenaiadini
| |
+--+ +-------------- Toxolasma **
| |
+--+ +----------- Potamilus **
| |
+--+ +-- Lampsilis_cardium **
| +-----+
| | +-- Lampsilis_ornata **
+--+
| +----- Venustaconcha **
+--+
| +-- Lampsilis_powellii **
+--+
+-- Lampsilis_siliquoidea **
iT| tT | bi | riS | oT | roT | roS | iD |
| 8 |
13 |
10 |
8 |
38 |
5 |
2 |
51 |
+----- Pleurobemini
+--------------+
| | +-- Obliquaria **
| +--+
--+ +-- Quadrulini
|
| +----------------- Ptychobranchus **
+--+
| +-------------- Truncilla **
+--+
| +----------- Ortmanniana **
+--+
| +-------- Lampsilis **
+--+
| +----- Cambarunio **
+--+
| +-- Ligumia **
+--+
+-- Epioblasma **
iT| tT | bi | riS | oT | roT | roS | iD |
| 8 |
14 |
12 |
8 |
39 |
6 |
3 |
52 |
+-------- Amblemini
+-----------+
| | +----- Pleurobemini
| +--+
| | +-- Obliquaria **
--+ +--+
| +-- Quadrulini
|
| +----------------- Ptychobranchus **
+--+
| +-------------- Truncilla **
+--+
| +----------- Ortmanniana **
+--+
| +-------- Lampsilis **
+--+
| +----- Cambarunio **
+--+
| +-- Ligumia **
+--+
+-- Epioblasma **
iT| tT | bi | riS | oT | roT | roS | iD |
| 8 |
9 |
7 |
7 |
26 |
6 |
3 |
52 |
+----------------------------- Megalonaias
|
| +----- Elliptoideus
--+ +--------------------+
| | | +-- Uniomerus
| | +--+
+--+ +-- Pleurobema
|
| +----------------------- Amblema
| |
+--+ +-- Obliquaria **
| +-----------------+
| | +-- Plectomerus
+--+
| +----------------- Medionidus **
+--+
| +-------------- Potamilus_fragilis **
+--+
| +----------- Ptychobranchus **
+--+
| +-------- Potamilus_purpuratus **
+--+
| +----- Lampsilis **
+--+
| +-- Villosa **
+--+
+-- Ellipsaria **
iT| tT | bi | riS | oT | roT | roS | iD |
| 8 |
8 |
6 |
4 |
46 |
3 |
2 |
48 |
+-------------------- Quadrulini
|
--+ +----------------- Pleurobemini
| |
+--+ +-------------- Potamilus **
| |
+--+ +----------- Leaunio **
| |
+--+ +-- Lampsilis_cardium **
| +-----+
| | +-- Lampsilis_ornata **
+--+
| +----- Venustaconcha **
+--+
| +-- Lampsilis_powellii **
+--+
+-- Lampsilis_siliquoidea **
iT| tT | bi | riS | oT | roT | roS | iD |
| 8 |
11 |
9 |
8 |
17 |
5 |
3 |
55 |
+----- Eurynia
+-----------+
| | +-- Cambarunio **
| +--+
| +-- Ptychobranchus **
--+
| +-- Amblemini
| +-----------+
| | +-- Pleurobema
+--+
| +----------- Truncilla **
| |
+--+ +-- Lampsilis **
| +-----+
| | +-- Epioblasma **
+--+
| +----- Ligumia **
+--+
| +-- Obliquaria **
+--+
+-- Ortmanniana **
iT| tT | bi | riS | oT | roT | roS | iD |
| 8 |
8 |
6 |
4 |
61 |
4 |
4 |
45 |
+-------------------------- Quadrulini
|
--+ +----------------------- Pleurobemini
| |
+--+ +-------------------- Amblemini
| |
+--+ +----------------- Popenaiadini
| |
+--+ +-------------- Toxolasma **
| |
+--+ +----------- Potamilus **
| |
+--+ +-- Lampsilis_ornata **
| +-----+
| | +-- Lampsilis_cardium **
+--+
| +----- Venustaconcha **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Lampsilis_powellii **
iT| tT | bi | riS | oT | roT | roS | iD |
| 8 |
8 |
4 |
8 |
39 |
6 |
3 |
49 |
+----------------------- Quadrulini
|
--+ +-------------------- Amblemini
| |
+--+ +----------------- Pleurobemini
| |
+--+ +-------------- Obliquaria **
| |
| +-------------- Truncilla **
+--+
+-------------- Ptychobranchus **
|
| +----------- Ligumia **
+--+
| +-------- Cambarunio **
+--+
| +----- Epioblasma **
+--+
| +-- Lampsilis **
+--+
+-- Ortmanniana **
iT| tT | bi | riS | oT | roT | roS | iD |
| 8 |
10 |
2 |
7 |
26 |
6 |
3 |
52 |
+----------- Megalonaias
|
| +----- Elliptoideus
| +--+
--+ | | +-- Uniomerus
| | +--+
| | +-- Pleurobema
| |
| | +----- Potamilus_fragilis **
+--+ |
| +----- Medionidus **
| |
| +----- Amblema
| |
| | +-- Obliquaria **
+--+--+
| +-- Plectomerus
|
| +-- Villosa **
| |
| +-- Ellipsaria **
| |
+--+-- Lampsilis **
|
+-- Potamilus_purpuratus **
|
+-- Ptychobranchus **
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
7 |
3 |
4 |
0 |
0 |
0 |
48 |
+----- Lampsilis **
|
--+----- Epioblasma **
|
| +-- Cambarunio **
+--+
+-- Villosa **
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
13 |
7 |
7 |
39 |
6 |
3 |
50 |
+-- Obliquaria **
+--------+
| +-- Quadrulini
|
--+ +-------- Amblemini
| |
| +-------- Pleurobemini
+--+
| +----- Truncilla **
| |
| +----- Ptychobranchus **
+--+
| +-- Ortmanniana **
| |
| +-- Cambarunio **
+--+
+-- Ligumia **
|
+-- Epioblasma **
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
7 |
8 |
5 |
0 |
0 |
0 |
44 |
+-------------------- Obliquaria **
|
+-------------------- Lampsilis **
--+
| +----------------- Ptychobranchus **
| |
+--+ +-- Cyprogenia_aberti__5 **
| +-----------+
| | +-- Dromus **
+--+
| +----------- Cyprogenia_aberti__4 **
| |
+--+ +-------- Cyprogenia_aberti__3 **
| |
+--+ +----- Cyprogenia_stegaria__2 **
| |
+--+----- Cyprogenia_aberti__2 **
|
| +-- Cyprogenia_stegaria__1 **
+--+
+-- Cyprogenia_aberti__1 **
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
7 |
5 |
5 |
49 |
3 |
2 |
45 |
+----------------- Quadrulini
--+
| +-------------- Pleurobemini
+--+
| +----------- Toxolasma **
+--+
| +-------- Potamilus **
+--+
| +----- Leaunio **
+--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
7 |
3 |
3 |
4 |
4 |
3 |
50 |
+-------------- Quadrula
|
--+-------------- Pustulosa
|
| +----------- Pleurobemini
+--+
| +-------- Amblemini
+--+
| +----- Obovaria **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
7 |
5 |
4 |
0 |
0 |
0 |
47 |
+----- Lampsilis **
|
--+----- Epioblasma **
|
| +-- Cambarunio **
+--+
+-- Villosa **
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
7 |
8 |
5 |
0 |
0 |
0 |
44 |
+-------------------- Obliquaria **
|
+-------------------- Lampsilis **
--+
| +----------------- Ptychobranchus **
| |
+--+ +-- Cyprogenia_aberti__5 **
| +-----------+
| | +-- Dromus **
+--+
| +----------- Cyprogenia_aberti__4 **
| |
+--+ +-------- Cyprogenia_aberti__3 **
| |
+--+ +----- Cyprogenia_stegaria__2 **
| |
+--+----- Cyprogenia_aberti__2 **
|
| +-- Cyprogenia_stegaria__1 **
+--+
+-- Cyprogenia_aberti__1 **
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
7 |
5 |
1 |
0 |
0 |
0 |
50 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
7 |
9 |
5 |
0 |
0 |
0 |
44 |
+----------------------- Obliquaria **
|
+----------------------- Lampsilis **
--+
| +-------------------- Ptychobranchus **
| |
+--+ +-- Cyprogenia_aberti__6 **
| +--------------+
| | +-- Dromus **
+--+
| +-------------- Cyprogenia_aberti__5 **
| |
+--+ +----------- Cyprogenia_aberti__4 **
| |
+--+ +-------- Cyprogenia_aberti__3 **
| |
+--+ +----- Cyprogenia_stegaria__2 **
| |
+--+----- Cyprogenia_aberti__2 **
|
| +-- Cyprogenia_stegaria__1 **
+--+
+-- Cyprogenia_aberti__1 **
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
7 |
8 |
2 |
3 |
1 |
1 |
50 |
+----------------------- Pleurobemini
|
--+ +-------------------- Lampsilis_siliquoidea **
| |
| | +----------------- Lampsilis_fasciola **
+--+ |
| | +-- Epioblasma **
| +--------------+
+--+ +-- Lampsilis_radiata **
|
| +-------------- Lampsilis_ornata **
| |
+--+ +-- Lampsilis_cardium__1 **
| +--------+
| | +-- Lampsilis_ovata__1 **
+--+
| +-------- Lampsilis_cardium__6 **
| |
+--+ +----- Lampsilis_cardium__5 **
| |
+--+ +-- Lampsilis_cardium__2 **
| |
| +-- Lampsilis_ovata__2 **
+--+
+-- Lampsilis_cardium__3 **
|
+-- Lampsilis_cardium__4 **
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
29 |
27 |
6 |
22 |
22 |
3 |
63 |
+-------------------------- Lampsilis **
|
--+ +----------------------- Potamilus **
| |
| | +----- Medionidus **
+--+ +--------------+
| | | +-- Dromus **
| | +--+
+--+ +-- Cyprogenia **
|
| +----------------- Obliquaria **
| |
+--+ +-------------- Amblema
| |
| | +----- Reginaia
+--+ +-----+
| | | +-- Elliptoideus
| | +--+
+--+ +-- Plectomerus
|
| +-------- Quadrulini
+--+
| +----- Hemistena
+--+
| +-- Elliptio
+--+
+-- Fusconaia
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
7 |
10 |
5 |
0 |
0 |
0 |
45 |
+-------------------------- Lampsilis **
|
--+ +----------------------- Obliquaria **
| |
+--+ +-------------------- Ptychobranchus **
| |
+--+ +-- Cyprogenia_aberti__6 **
| +--------------+
| | +-- Dromus **
+--+
| +-------------- Cyprogenia_aberti__4 **
| |
+--+ +----------- Cyprogenia_aberti__3 **
| |
+--+ +-------- Cyprogenia_aberti__5 **
| |
+--+ +----- Cyprogenia_stegaria__2 **
| |
+--+----- Cyprogenia_aberti__2 **
|
| +-- Cyprogenia_stegaria__1 **
+--+
+-- Cyprogenia_aberti__1 **
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
7 |
1 |
1 |
0 |
0 |
0 |
49 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
7 |
7 |
5 |
0 |
0 |
0 |
46 |
+-------------- Obliquaria **
|
| +-- Ptychobranchus **
+-----------+
--+ +-- Lampsilis **
|
| +-- Cyprogenia_aberti__6 **
| +--------+
| | +-- Dromus **
+--+
| +-------- Cyprogenia_aberti__5 **
| |
+--+ +----- Cyprogenia_aberti__4 **
| |
| | +-- Cyprogenia_stegaria__1 **
+--+ |
| +-- Cyprogenia_stegaria__2 **
| |
+--+-- Cyprogenia_aberti__1 **
|
+-- Cyprogenia_aberti__2 **
|
+-- Cyprogenia_aberti__3 **
iT| tT | bi | riS | oT | roT | roS | iD |
| 7 |
29 |
20 |
6 |
22 |
22 |
3 |
62 |
+----------- Lampsilis **
|
| +-------- Obliquaria **
| |
| +-------- Amblema
| |
--+ +-------- Reginaia
| |
| +-------- Potamilus **
| |
| +-------- Quadrulini
| |
+--+ +-- Elliptoideus
+-----+
| +-- Plectomerus
|
| +----- Medionidus **
+--+
| | +-- Dromus **
| +--+
| +-- Cyprogenia **
|
| +----- Hemistena
+--+
| +-- Elliptio
+--+
+-- Fusconaia
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
4 |
33 |
1 |
1 |
45 |
+-------------- Quadrulini
|
--+ +----------- Toxolasma **
| |
+--+ +-------- Potamilus **
| |
+--+ +-- Lampsilis_cardium **
| +--+
| | +-- Lampsilis_ornata **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
11 |
5 |
6 |
28 |
5 |
3 |
49 |
+-------- Toxolasma **
|
+-------- Amblemini
|
| +-- Glebula **
+-----+
--+ +-- Cyrtonaias **
|
| +-- Quadrulini
+-----+
| +-- Pleurobemini
|
| +----- Potamilus **
+--+
| +-- Ortmanniana **
+--+
+-- Ligumia **
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
5 |
47 |
6 |
4 |
46 |
+----------------------- Quadrulini
--+
| +-------------------- Pleurobemini
+--+
| +----------------- Amblemini
+--+
| +-------------- Popenaiadini
+--+
| +----------- Toxolasma **
+--+
| +-------- Potamilus **
+--+
| +----- Leaunio **
+--+
| +-- Lampsilis **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
11 |
8 |
6 |
25 |
5 |
3 |
50 |
+-- Glebula **
+--------+
| +-- Cyrtonaias **
|
--+ +-------- Toxolasma **
| |
| | +----- Potamilus **
| +--+
+--+ | +-- Ligumia **
| +--+
| +-- Ortmanniana **
|
| +----- Amblemini
+--+
| +-- Quadrulini
+--+
+-- Pleurobemini
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
6 |
1 |
0 |
0 |
0 |
50 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
6 |
25 |
5 |
3 |
48 |
+----------------- Amblemini
|
| +-- Quadrulini
--+ +-----------+
| | +-- Pleurobemini
| |
+--+ +-- Cyrtonaias **
| +--------+
| | +-- Glebula **
+--+
| +-------- Toxolasma **
+--+
| +----- Ortmanniana **
+--+
| +-- Ligumia **
+--+
+-- Potamilus **
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
4 |
31 |
1 |
1 |
45 |
+-------------- Quadrulini
|
--+ +----------- Toxolasma **
| |
+--+ +-------- Potamilus **
| |
+--+ +-- Lampsilis_cardium **
| +--+
| | +-- Lampsilis_ornata **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
6 |
32 |
31 |
3 |
47 |
+----------------------------- Amblemini
|
| +-- Cyrtonaias **
| +-----+
| | +-- Glebula **
--+ |
| +-----------------+ +-- Lampsilis **
| | | +--+
| | | | +-- Leaunio **
| | +--+
| | | +-- Truncilla **
+--+ +--+
| +-- Potamilus **
|
| +----------------------- Elliptoideus
| |
+--+ +-------------------- Quadrulini
| |
+--+ +----------------- Elliptio
| |
| | +-------------- Parvaspina
+--+ |
| +-------------- Plethobasus
| |
| | +-- Hemistena
+--+-----------+
| +-- Eurynia
|
| +----------- Fusconaia
| |
| | +----- Pleurobema_riddellii
+--+ +--+
| | | +-- Pleurobema_sintoxia
| | +--+
+--+ +-- Pleurobema_cordatum
|
| +----- Pleuronaia
| |
+--+ +-- Pleurobema_pyriforme
| |
+--+-- Pleurobema_strodeanum
|
+-- Pleurobema_clava
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
12 |
9 |
6 |
25 |
5 |
3 |
57 |
+----------------------- Fusconaia
|
+----------------------- Amblemini
--+
| +-------------------- Quadrulini
| |
+--+ +----------------- Pleurobema
| |
| | +-------------- Eurynia
+--+ |
| | +----- Cyrtonaias **
| +--------+
+--+ | +-- Unionini
| +--+
| +-- Anodontini
|
| +----------- Ortmanniana **
+--+
| +-------- Glebula **
+--+
| +----- Potamilus **
+--+
| +-- Ligumia **
+--+
+-- Toxolasma **
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
16 |
14 |
3 |
28 |
1 |
1 |
58 |
+-- Anodontini
+-----+
| +-- Potamilus_alatus **
+-----------+
| | +----- Lamprotulini
| +--+
| | +-- Chamberlainiini
--+ +--+
| +-- Gonideini
|
| +----------------- Gibbosula
+--+
| +-------------- Quadrulini
+--+
| +----------- Toxolasma **
+--+
| +-------- Lampsilis_ornata **
+--+
| +----- Potamilus_leptodon **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Lampsilis_powellii **
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
28 |
16 |
6 |
25 |
5 |
3 |
69 |
+-------------------- Amblemini
|
+-------------------- Coelaturini
|
+-------------------- Eurynia
|
+-------------------- Fusconaia
|
+-------------------- Gonideini
|
+-------------------- Pleurobema
|
+-------------------- Quadrulini
--+
| +----------- Ortmanniana **
+--------+
| | +-------- Glebula **
| +--+
| | +----- Potamilus **
| +--+
| | +-- Ligumia **
| +--+
| +-- Toxolasma **
|
| +----------------- Cyrtonaias **
| |
+--+ +-------------- Unionini
| |
| | +----------- Anodontini
+--+ |
| | +----- Mutela
| | |
+--+ +--+ +-- Anodontites_guanarensis
| | | |
| | +--+-- Anodontites_trigona
| | |
+--+ +-- Monocondylaea
|
| +----- Rhipidodontini
| |
| +----- Echyridella
+--+
+----- Lortiella
|
| +-- Castaliini
+--+
+-- Velesunio
iT| tT | bi | riS | oT | roT | roS | iD |
| 6 |
6 |
4 |
6 |
25 |
5 |
3 |
46 |
+----------------- Amblemini
|
| +-- Quadrulini
--+ +-----------+
| | +-- Pleurobemini
| |
+--+ +-- Cyrtonaias **
| +--------+
| | +-- Glebula **
+--+
| +-------- Toxolasma **
+--+
| +----- Potamilus **
+--+
| +-- Ligumia **
+--+
+-- Ortmanniana **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
5 |
49 |
5 |
3 |
47 |
+-------------------- Quadrulini
--+
| +----------------- Pleurobemini
+--+
| +-------------- Popenaiadini
+--+
| +----------- Toxolasma **
+--+
| +-------- Potamilus **
+--+
| +----- Leaunio **
+--+
| +-- Lampsilis **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
4 |
7 |
4 |
2 |
49 |
+-------------- Quadrulini
|
--+ +----------- Plectomerus
| |
+--+ +-------- Amblema
| |
+--+ +-- Potamilus **
| +--+
| | +-- Glebula **
+--+
| +-- Leaunio **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
4 |
41 |
1 |
1 |
44 |
+----------- Quadrulini
--+
| +-------- Toxolasma **
+--+
| +----- Potamilus **
+--+
| +-- Lampsilis **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
4 |
26 |
1 |
1 |
44 |
+----------- Quadrulini
--+
| +-------- Toxolasma **
+--+
| +----- Potamilus **
+--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
9 |
7 |
4 |
6 |
4 |
2 |
52 |
+----------------- Glebula **
|
--+ +-- Amblema
| +-----------+
| | +-- Quadrulini
+--+
| +----------- Plectomerus
+--+
| +-------- Leaunio **
+--+
| +----- Lampsilis_straminea **
+--+
| +-- Lampsilis_teres **
+--+
+-- Potamilus **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
5 |
10 |
1 |
1 |
46 |
+----------- Mesonaias **
--+
| +-------- Potamilus **
+--+
| +----- Ligumia **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
3 |
1 |
1 |
1 |
48 |
+-------- Quadrulini
--+
| +----- Potamilus **
+--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
1 |
3 |
0 |
0 |
0 |
49 |
+-- Paetulunio **
|
--+-- Epioblasma **
|
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
5 |
9 |
1 |
1 |
46 |
+-------------- Pleurobemini
--+
| +----------- Potamilus **
+--+
| +-------- Cyrtonaias **
+--+
| +----- Ligumia **
+--+
| +-- Lampsilis **
+--+
+-- Ortmanniana **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
5 |
52 |
10 |
4 |
49 |
+-------------------- Quadrulini
|
--+ +-- Pleurobemini
| +--------------+
| | +-- Amblemini
+--+
| +-------------- Popenaiadini
+--+
| +----------- Obliquaria **
+--+
| +-------- Cyrtonaias **
+--+
| +----- Leaunio **
+--+
| +-- Lampsilis **
+--+
+-- Pachynaias **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
6 |
3 |
4 |
37 |
37 |
4 |
45 |
+-------------------- Quadrulini
|
--+ +----------------- Pleurobemini
| |
+--+ +-- Reginaia
| +-----------+
| | +-- Amblema
+--+
| +----------- Popenaiadini
| |
+--+ +-------- Cyrtonaias **
| |
+--+-------- Plectomerus
|
| +----- Lemiox **
+--+
| +-- Lampsilis **
+--+
+-- Obovaria **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
2 |
0 |
0 |
0 |
45 |
+-- Lampsilis_siliquoidea__1 **
|
--+-- Lampsilis_siliquoidea__2 **
|
+-- Epioblasma **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
6 |
3 |
5 |
31 |
1 |
1 |
47 |
+----------- Cyrtonaias **
|
--+----------- Pleurobemini
|
| +-------- Potamilus **
+--+
| +----- Lampsilis **
+--+
| +-- Ligumia **
+--+
+-- Ortmanniana **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
5 |
52 |
10 |
4 |
49 |
+----------------------- Quadrulini
--+
| +-------------------- Pleurobemini
+--+
| +----------------- Amblemini
+--+
| +-------------- Popenaiadini
+--+
| +----------- Obliquaria **
+--+
| +-------- Cyrtonaias **
+--+
| +----- Leaunio **
+--+
| +-- Lampsilis **
+--+
+-- Pachynaias **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
4 |
1 |
0 |
0 |
0 |
49 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
4 |
43 |
1 |
1 |
44 |
+----------- Quadrulini
--+
| +-------- Toxolasma **
+--+
| +----- Potamilus **
+--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
2 |
0 |
0 |
0 |
50 |
+-------- Lampsilis_radiata **
|
--+ +-- Lampsilis_ovata **
| +--+
| | +-- Ortmanniana **
+--+
| +-- Lampsilis_hydiana **
+--+
+-- Lampsilis_straminea **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
5 |
10 |
1 |
1 |
46 |
+-------------- Pleurobemini
--+
| +----------- Mesonaias **
+--+
| +-------- Potamilus **
+--+
| +----- Ligumia **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
8 |
6 |
4 |
37 |
37 |
4 |
46 |
+-------------------- Quadrulini
|
--+ +----------------- Pleurobemini
| |
| | +-- Popenaiadini
+--+ +-----------+
| | +-- Amblema
| |
+--+ +-- Cyrtonaias **
| +--------+
| | +-- Reginaia
+--+
| +-------- Plectomerus
+--+
| +----- Lemiox **
+--+
| +-- Lampsilis **
+--+
+-- Obovaria **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
4 |
39 |
1 |
1 |
44 |
+----------- Quadrulini
--+
| +-------- Toxolasma **
+--+
| +----- Potamilus **
+--+
| +-- Lampsilis **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
8 |
5 |
4 |
37 |
37 |
4 |
46 |
+----------------- Quadrulini
|
--+ +-------------- Pleurobemini
| |
| | +-- Popenaiadini
+--+ +--------+
| | +-- Amblema
| |
+--+ +-------- Plectomerus
| |
| | +-- Cyrtonaias **
+--+-----+
| +-- Reginaia
|
| +----- Lemiox **
+--+
| +-- Lampsilis **
+--+
+-- Obovaria **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
4 |
11 |
1 |
1 |
44 |
+----------- Quadrulini
--+
| +-------- Toxolasma **
+--+
| +----- Potamilus **
+--+
| +-- Lampsilis **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
5 |
10 |
1 |
1 |
46 |
+-------------- Pleurobemini
--+
| +----------- Potamilus **
+--+
| +-------- Mesonaias **
+--+
| +----- Ligumia **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
36 |
35 |
4 |
38 |
38 |
4 |
73 |
+-------------------------------- Quadrulini
|
| +-------------- Elliptoideus
| +--------------+
| | | +----------- Plectomerus
--+ | +--+
| | | +-------- Obovaria_unicolor **
| | +--+
| | | +----- Obovaria_olivaria **
| | +--+
| | | +-- Lampsilis **
+--+ +--+
| +-- Lemiox **
|
| +-------- Cyrtonaias **
| +-----------------+
| | | +----- Reginaia
| | +--+
| | | +-- Popenaiadini
+--+ +--+
| +-- Amblema
|
| +----------------------- Elliptio
| |
+--+ +-------------------- Eurynia
| |
| | +----------------- Parvaspina
+--+ |
| | +-- Hemistena
| +--------------+
+--+ +-- Plethobasus
|
| +-------------- Pleuronaia
| |
| | +-- Pleurobema_cordatum
+--+ +--------+
| | +-- Fusconaia
| |
+--+ +-- Pleurobema_stabile
| +-----+
| | +-- Pleurobema_sintoxia
+--+
| +----- Pleurobema_decisum
+--+
| +-- Pleurobema_clava
+--+
+-- Pleurobema_pyriforme
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
2 |
0 |
0 |
0 |
49 |
+-- Ellipsaria **
--+
+-- Truncilla **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
4 |
37 |
1 |
1 |
44 |
+----------- Quadrulini
--+
| +-------- Toxolasma **
+--+
| +----- Potamilus **
+--+
| +-- Lampsilis **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
5 |
4 |
1 |
1 |
46 |
+-------------- Pleurobemini
--+
| +----------- Potamilus **
+--+
| +-------- Cyrtonaias **
+--+
| +----- Ligumia **
+--+
| +-- Lampsilis **
+--+
+-- Ortmanniana **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
4 |
44 |
1 |
1 |
44 |
+----------- Quadrulini
--+
| +-------- Toxolasma **
+--+
| +----- Potamilus **
+--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
4 |
39 |
1 |
1 |
44 |
+----------- Quadrulini
--+
| +-------- Toxolasma **
+--+
| +----- Potamilus **
+--+
| +-- Lampsilis **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
5 |
4 |
1 |
1 |
48 |
+-------------- Pleurobemini
--+
| +----------- Cyrtonaias **
+--+
| +-------- Ortmanniana **
+--+
| +----- Lampsilis **
+--+
| +-- Potamilus **
+--+
+-- Ligumia **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
4 |
26 |
1 |
1 |
44 |
+----------- Quadrulini
--+
| +-------- Toxolasma **
+--+
| +----- Potamilus **
+--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
6 |
2 |
4 |
8 |
8 |
4 |
49 |
+----- Amblemini
+--+
| | +-- Pleurobemini
| +--+
| +-- Quadrulini
--+
| +----- Obliquaria **
| |
| +----- Toxolasma **
+--+
+----- Popenaiadini
|
| +-- Glebula **
+--+
+-- Cyrtonaias **
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
2 |
4 |
15 |
2 |
2 |
47 |
+-- Pleurobemini
+--------+
| +-- Quadrulini
--+
| +-------- Obliquaria **
+--+
| +----- Truncilla **
+--+
| +-- Lampsilis **
+--+
+-- Potamilus **
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
2 |
4 |
87 |
2 |
2 |
45 |
+-------------- Quadrulini
--+
| +----------- Pleurobemini
+--+
| +-------- Obliquaria **
+--+
| +----- Potamilus **
+--+
| +-- Lampsilis **
+--+
+-- Truncilla **
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
2 |
4 |
8 |
8 |
4 |
46 |
+----- Amblemini
+--------+
| | +-- Pleurobemini
| +--+
--+ +-- Quadrulini
|
| +----------- Popenaiadini
+--+
| +-------- Toxolasma **
+--+
| +----- Obliquaria **
+--+
| +-- Glebula **
+--+
+-- Cyrtonaias **
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
2 |
3 |
0 |
0 |
0 |
49 |
+----- Venustaconcha **
--+
| +-- Ligumia **
+--+
+-- Sagittunio **
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
2 |
3 |
1 |
0 |
0 |
48 |
+----- Lampsilis **
--+
| +-- Epioblasma **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
2 |
3 |
34 |
1 |
1 |
45 |
+-------- Quadrulini
--+
| +----- Toxolasma **
+--+
| +-- Lampsilis **
+--+
+-- Potamilus **
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
2 |
3 |
16 |
5 |
3 |
46 |
+-------------- Quadrulini
--+
| +----------- Pleurobemini
+--+
| +-------- Amblemini
+--+
| +----- Potamilus **
+--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
2 |
4 |
87 |
2 |
2 |
45 |
+-- Quadrulini
+--------+
| +-- Pleurobemini
--+
| +-------- Obliquaria **
+--+
| +----- Potamilus **
+--+
| +-- Lampsilis **
+--+
+-- Truncilla **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
1 |
13 |
0 |
0 |
50 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
17 |
0 |
2 |
19 |
8 |
3 |
59 |
+-- Gonideini
|
+-- Pyganodon_grandis
|
+-- Utterbackia
|
+-- Pyganodon_cataracta
|
+-- Lampsilis_straminea **
|
+-- Lampsilis_teres **
|
+-- Megalonaias
|
+-- Obliquaria **
|
--+-- Theliderma
|
+-- Quadrula
|
+-- Uniomerus
|
+-- Amblema
|
+-- Elliptio
|
+-- Fusconaia
|
+-- Plectomerus
|
+-- Aculamprotula
|
+-- Unio
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
1 |
0 |
0 |
0 |
48 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
2 |
1 |
3 |
0 |
0 |
51 |
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
3 |
22 |
1 |
1 |
46 |
+-------- Quadrulini
--+
| +----- Toxolasma **
+--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
2 |
11 |
9 |
3 |
46 |
+-------- Amblemini
|
--+ +-- Lampsilis **
| +--+
| | +-- Obovaria **
+--+
| +-- Quadrulini
+--+
+-- Pleurobemini
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
3 |
13 |
3 |
2 |
46 |
+-------- Quadrulini
--+
| +----- Toxolasma **
+--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
3 |
8 |
8 |
4 |
46 |
+----- Amblemini
+-----+
| | +-- Pleurobemini
| +--+
--+ +-- Quadrulini
|
| +-------- Popenaiadini
+--+
| +----- Obliquaria **
+--+
| +-- Glebula **
+--+
+-- Cyrtonaias **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
3 |
73 |
6 |
3 |
48 |
+-- Quadrulini
+--------+
| +-- Pleurobemini
--+
| +-------- Amblemini
+--+
| +----- Ortmanniana **
+--+
| +-- Cambarunio **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
3 |
28 |
1 |
1 |
46 |
+-------- Quadrulini
--+
| +----- Toxolasma **
+--+
| +-- Lampsilis **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
3 |
136 |
6 |
3 |
48 |
+-- Quadrulini
+--------+
| +-- Pleurobemini
--+
| +-------- Amblemini
+--+
| +----- Ortmanniana **
+--+
| +-- Lampsilis **
+--+
+-- Cambarunio **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
3 |
15 |
2 |
2 |
47 |
+-------- Quadrulini
|
--+-------- Pleurobemini
|
| +----- Lampsilis **
+--+
| +-- Epioblasma **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
3 |
17 |
1 |
1 |
46 |
+-------- Quadrulini
--+
| +----- Toxolasma **
+--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
7 |
0 |
3 |
33 |
4 |
3 |
50 |
+-- Amblemini
|
+-- Ortmanniana **
|
+-- Lampsilis **
|
--+-- Cambarunio **
|
+-- Elliptio
|
+-- Pleurobema
|
+-- Quadrulini
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
3 |
22 |
1 |
1 |
46 |
+-------- Quadrulini
--+
| +----- Toxolasma **
+--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
2 |
11 |
9 |
3 |
48 |
+----- Obovaria_unicolor **
--+
| +-- Obovaria_olivaria **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
3 |
37 |
1 |
1 |
46 |
+-------- Quadrulini
--+
| +----- Toxolasma **
+--+
| +-- Lampsilis **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
3 |
38 |
1 |
1 |
46 |
+-------- Quadrulini
--+
| +----- Toxolasma **
+--+
| +-- Venustaconcha **
+--+
+-- Potamilus **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
3 |
136 |
6 |
3 |
48 |
+-- Quadrulini
+--------+
| +-- Pleurobemini
--+
| +-------- Amblemini
+--+
| +----- Cambarunio **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
13 |
0 |
3 |
80 |
4 |
3 |
55 |
+-- Lamprotula
|
+-- Potomida
|
+-- Pronodularia
|
+-- Gonidea
|
+-- Leguminaia
|
+-- Microcondylaea
|
--+-- Amblemini
|
+-- Ortmanniana **
|
+-- Lampsilis **
|
+-- Cambarunio **
|
+-- Elliptio
|
+-- Pleurobema
|
+-- Quadrulini
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
2 |
7 |
0 |
0 |
48 |
+-- Potamilus **
--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
9 |
0 |
3 |
122 |
6 |
3 |
52 |
+-- Amblemini
|
+-- Ortmanniana **
|
+-- Lampsilis **
|
+-- Cambarunio **
|
--+-- Elliptio
|
+-- Eurynia
|
+-- Pleurobema
|
+-- Quadrula_quadrula
|
+-- Quadrula_verrucosa
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
3 |
77 |
6 |
3 |
48 |
+-- Quadrulini
+--------+
| +-- Pleurobemini
--+
| +-------- Amblemini
+--+
| +----- Cambarunio **
+--+
| +-- Lampsilis **
+--+
+-- Ortmanniana **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
3 |
29 |
1 |
1 |
46 |
+-------- Quadrulini
--+
| +----- Toxolasma **
+--+
| +-- Lampsilis **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
3 |
13 |
1 |
1 |
46 |
+-------- Quadrulini
--+
| +----- Toxolasma **
+--+
| +-- Lampsilis **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
3 |
37 |
2 |
2 |
46 |
+----------- Quadrulini
--+
| +-------- Amblemini
+--+
| +----- Potamilus **
+--+
| +-- Lampsilis **
+--+
+-- Truncilla **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
17 |
0 |
2 |
19 |
9 |
3 |
59 |
+-- Gonideini
|
+-- Pyganodon_grandis
|
+-- Utterbackia
|
+-- Pyganodon_cataracta
|
+-- Lampsilis_straminea **
|
+-- Lampsilis_teres **
|
+-- Megalonaias
|
+-- Obliquaria **
|
--+-- Theliderma
|
+-- Quadrula
|
+-- Uniomerus
|
+-- Amblema
|
+-- Elliptio
|
+-- Fusconaia
|
+-- Plectomerus
|
+-- Pleurobema
|
+-- Unionini
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
2 |
11 |
9 |
3 |
46 |
+-- Lampsilis **
--+
+-- Obovaria **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
10 |
0 |
2 |
16 |
8 |
3 |
54 |
+-- Pyganodon_cataracta
|
+-- Pyganodon_grandis
|
+-- Utterbackia
|
+-- Lampsilis_teres **
|
+-- Lampsilis_straminea **
--+
+-- Megalonaias
|
+-- Obliquaria **
|
+-- Theliderma
|
+-- Quadrula
|
+-- Uniomerus
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
3 |
38 |
1 |
1 |
46 |
+-------- Quadrulini
--+
| +----- Toxolasma **
+--+
| +-- Venustaconcha **
+--+
+-- Potamilus **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
2 |
11 |
9 |
3 |
46 |
+-- Lampsilis **
+-----+
| +-- Obovaria **
--+
| +----- Amblemini
+--+
| +-- Quadrulini
+--+
+-- Pleurobemini
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
3 |
73 |
6 |
3 |
48 |
+-- Quadrulini
+--------+
| +-- Pleurobemini
--+
| +-------- Amblemini
+--+
| +----- Ortmanniana **
+--+
| +-- Lampsilis **
+--+
+-- Cambarunio **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
1 |
2 |
38 |
1 |
1 |
48 |
+----- Quadrulini
--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
19 |
1 |
1 |
-- |
+----- Quadrulini
--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
8 |
2 |
2 |
-- |
+-- Pleurobemini
+--+
| +-- Quadrulini
--+
| +-- Lampsilis **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
39 |
0 |
2 |
39 |
39 |
3 |
78 |
+----------- Amblema
|
--+ +-- Lampsilis **
| +-----+
| | +-- Plectomerus
+--+
| +----- Popenaiadini
+--+
| +-- Cyrtonaias **
+--+
+-- Pleurobemini
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
26 |
22 |
3 |
-- |
+-------- Amblemini
|
--+ +-- Lampsilis **
| +--+
| | +-- Potamilus **
+--+
| +-- Pleurobemini
+--+
+-- Quadrulini
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
17 |
1 |
1 |
-- |
+----- Quadrulini
--+
| +-- Potamilus **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
11 |
3 |
2 |
-- |
+-- Lampsilis **
+--+
| +-- Sagittunio **
--+
| +-- Amblemini
+--+
+-- Pleurobemini
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
28 |
1 |
1 |
-- |
+----- Quadrulini
--+
| +-- Toxolasma **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
11 |
11 |
3 |
-- |
+----------- Quadrulini
--+
| +-------- Pleurobemini
+--+
| +----- Amblemini
+--+
| +-- Obovaria **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
6 |
0 |
2 |
6 |
1 |
1 |
50 |
+-------- Lampsilis **
|
--+ +-- Venustaconcha **
| +--+
| | +-- Quadrulini
+--+
| +-- Lamprotulini
+--+
+-- Chamberlainiini
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
16 |
2 |
2 |
-- |
+-- Lampsilis **
+--+
| +-- Obliquaria **
--+
| +-- Pleurobemini
+--+
+-- Amblemini
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
1 |
9 |
1 |
1 |
-- |
+-- Quadrulini
--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
10 |
1 |
1 |
-- |
+-- Toxolasma **
--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
10 |
1 |
1 |
-- |
+----- Quadrulini
--+
| +-- Toxolasma **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
21 |
1 |
1 |
-- |
+----- Quadrulini
--+
| +-- Venustaconcha **
+--+
+-- Potamilus **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
13 |
0 |
0 |
-- |
+-- Lampsilis **
--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
15 |
1 |
1 |
-- |
+----- Quadrulini
--+
| +-- Venustaconcha **
+--+
+-- Potamilus **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
1 |
9 |
1 |
1 |
-- |
+-- Quadrulini
--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
20 |
1 |
1 |
-- |
+----- Quadrulini
--+
| +-- Venustaconcha **
+--+
+-- Potamilus **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
17 |
1 |
1 |
-- |
+----- Quadrulini
--+
| +-- Potamilus **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
24 |
20 |
3 |
-- |
+----------- Quadrulini
--+
| +-------- Amblemini
+--+
| +----- Pleurobemini
+--+
| +-- Lampsilis **
+--+
+-- Potamilus **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
12 |
1 |
1 |
-- |
+----- Quadrulini
--+
| +-- Potamilus **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
27 |
23 |
3 |
-- |
+-------------- Megalonaias
|
--+ +-- Uniomerus
| +--------+
| | +-- Quadrulina
+--+
| +-------- Pleurobemini
+--+
| +----- Amblemini
+--+
| +-- Lampsilis **
+--+
+-- Potamilus **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
21 |
1 |
1 |
-- |
+----- Quadrulini
--+
| +-- Venustaconcha **
+--+
+-- Potamilus **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
18 |
1 |
1 |
-- |
+----- Quadrulini
--+
| +-- Potamilus **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
20 |
1 |
1 |
-- |
+----- Quadrulini
--+
| +-- Venustaconcha **
+--+
+-- Potamilus **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
4 |
4 |
1 |
-- |
+----- Popenaiadini
--+
| +-- Mesonaias **
+--+
+-- Pachynaias **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
20 |
1 |
1 |
-- |
+----- Quadrulini
--+
| +-- Venustaconcha **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
29 |
3 |
3 |
-- |
+----------- Quadrulini
--+
| +-------- Pleurobemini
+--+
| +----- Amblemini
+--+
| +-- Lampsilis **
+--+
+-- Truncilla **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
4 |
1 |
1 |
-- |
+----- Quadrulini
--+
| +-- Potamilus **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
34 |
3 |
3 |
-- |
+----------- Pleurobemini
--+
| +-------- Quadrulini
+--+
| +----- Amblemini
+--+
| +-- Truncilla **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
34 |
3 |
3 |
-- |
+-------- Quadrulini
|
--+-------- Pleurobemini
|
| +----- Amblemini
+--+
| +-- Lampsilis **
+--+
+-- Truncilla **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
14 |
1 |
1 |
-- |
+----- Quadrulini
--+
| +-- Potamilus **
+--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
5 |
1 |
1 |
-- |
+----- Quadrulini
--+
| +-- Lampsilis **
+--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
6 |
1 |
1 |
-- |
-- Venustaconcha_ellipsiformis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
24 |
24 |
3 |
-- |
+----------------------- Quadrulini
|
+----------------------- Amblemini
|
| +-------------------- Cyrtonaias **
| |
| +-------------------- Eurynia
| |
| +-------------------- Fusconaia_cerina
--+ |
| +-------------------- Fusconaia_flava
| |
| +-------------------- Hemistena
| |
| +-------------------- Plethobasus
| |
| +-------------------- Parvaspina
| |
+--+-------------------- Pleurobema_cordatum
|
+-------------------- Pleuronaia
|
+-------------------- Pleurobema_oviforme__2
|
+-------------------- Pleurobema_stabile
|
+-------------------- Elliptio
|
| +----------------- Pleurobema_decisum
| |
| +----------------- Pleurobema_sintoxia
| |
| +----------------- Pleurobema_clava
+--+
+----------------- Pleurobema_oviforme__1
|
| +-------------- Pleurobema_beadleianum
| |
+--+ +----------- Pleurobema_pyriforme
| |
+--+ +-------- Pleurobema_rubellum
| |
| +-------- Pleurobema_hanleyianum__2
+--+
+-------- Pleurobema_georgianum__2
|
| +----- Pleurobema_hanleyianum__1
+--+
| +-- Pleurobema_georgianum__1
+--+
+-- Pleurobema_athearni
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
24 |
24 |
3 |
-- |
+----------- Quadrulini
|
+----------- Amblemini
|
| +-- Elliptio_arctata
| +-----+
| | +-- Elliptio_crassidens
| |
| | +----- Pleurobema_decisum
--+ | |
| | +----- Pleurobema_rubellum
| | |
| | +----- Pleurobema_georgianum__2
| | |
| | +----- Pleurobema_stabile
| | |
| | +----- Pleurobema_pyriforme
| | |
+--+ +----- Pleurobema_oviforme__2
| |
| +----- Pleurobema_hanleyianum__2
| |
| +----- Pleuronaia
| |
| +----- Pleurobema_georgianum__3
| |
| +----- Pleurobema_cordatum
| |
| +----- Pleurobema_beadleianum
| |
| +----- Hemistena
+--+
+----- Fusconaia_flava
|
+----- Fusconaia_cerina
|
+----- Eurynia
|
+----- Elliptio_arca
|
+----- Cyrtonaias **
|
| +-- Plethobasus
+--+
| +-- Parvaspina
|
| +-- Pleurobema_sintoxia
| |
+--+-- Pleurobema_oviforme__1
| |
| +-- Pleurobema_clava
|
| +-- Pleurobema_athearni
| |
+--+-- Pleurobema_georgianum__1
|
+-- Pleurobema_hanleyianum__1
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
5 |
1 |
1 |
-- |
+-- Quadrulini
--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
13 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
3 |
2 |
1 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
7 |
0 |
0 |
-- |
-- Epioblasma_triquetra **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
44 |
1 |
1 |
-- |
+-- Lampsilis **
--+
+-- Quadrulini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
6 |
1 |
1 |
-- |
+-- Venustaconcha **
--+
+-- Quadrulini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
14 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
3 |
2 |
1 |
-- |
+-- Lampsilis **
--+
+-- Quadrulini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
52 |
8 |
3 |
-- |
+-- Amblemini
--+
+-- Obovaria **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
10 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
3 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
40 |
1 |
1 |
-- |
+-- Lampsilis **
--+
+-- Quadrulini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
31 |
1 |
1 |
-- |
+-- Amblemini
--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
1 |
1 |
1 |
-- |
+-- Quadrulini
--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
10 |
1 |
1 |
-- |
+-- Amblemini
--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
1 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
7 |
1 |
1 |
-- |
+-- Venustaconcha **
--+
+-- Quadrulini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
7 |
1 |
1 |
-- |
+-- Venustaconcha **
--+
+-- Quadrulini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
2 |
1 |
1 |
-- |
+-- Pleurobemini
--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
33 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
27 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
3 |
1 |
1 |
-- |
+-- Lampsilis **
--+
+-- Pleurobemini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
6 |
1 |
1 |
-- |
+-- Venustaconcha **
--+
+-- Quadrulini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
3 |
3 |
1 |
-- |
+-- Villosa **
--+
+-- Amblemini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
5 |
1 |
1 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
11 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
67 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
4 |
2 |
2 |
-- |
+-- Lampsilis **
--+
+-- Pleurobemini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
44 |
1 |
1 |
-- |
+-- Lampsilis **
--+
+-- Quadrulini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
4 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
14 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
3 |
2 |
1 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
6 |
1 |
1 |
-- |
-- Venustaconcha_ellipsiformis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
29 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
27 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
3 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
25 |
25 |
2 |
-- |
+-- Cyrtonaias **
|
--+-- Amblemini
|
+-- Pleurobemini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
45 |
1 |
1 |
-- |
+-- Amblemini
--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
40 |
1 |
1 |
-- |
+-- Lampsilis **
--+
+-- Quadrulini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
9 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
15 |
1 |
1 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
28 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
33 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
26 |
25 |
4 |
-- |
+----------- Quadrulini
--+
| +-------- Pleurobemini
+--+
| +----- Amblemini
+--+
| +-- Lampsilis **
+--+
+-- Popenaiadini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
9 |
1 |
1 |
-- |
+-- Venustaconcha **
--+
+-- Quadrulini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
5 |
1 |
1 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
12 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
13 |
1 |
1 |
-- |
+-- Amblemini
--+
+-- Lampsilis **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
14 |
1 |
1 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
52 |
1 |
1 |
-- |
+-- Lampsilis **
--+
+-- Quadrulini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
121 |
3 |
3 |
-- |
+-- Amblemini
|
+-- Pleurobemini
--+
+-- Lampsilis **
|
+-- Quadrulini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
7 |
1 |
1 |
-- |
+-- Venustaconcha **
--+
+-- Quadrulini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
25 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
8 |
0 |
0 |
-- |
-- Epioblasma_triquetra **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
25 |
25 |
2 |
-- |
+-- Cyrtonaias **
|
--+-- Amblemini
|
+-- Pleurobemini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
75 |
1 |
1 |
-- |
+-- Lampsilis **
--+
+-- Quadrulini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
67 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
12 |
1 |
1 |
-- |
+-- Quadrulini
--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
11 |
1 |
1 |
-- |
+-- Venustaconcha **
--+
+-- Quadrulini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
19 |
3 |
3 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
9 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
55 |
1 |
1 |
-- |
+-- Lampsilis **
--+
+-- Amblemini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
11 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
14 |
1 |
1 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
29 |
29 |
2 |
-- |
+-------------------------------- Lampsilis **
|
--+ +----------------------------- Parvaspina
| |
| | +-------------------------- Amblemini
+--+ |
| | +----- Eurynia
| | +-----------------+
+--+ | | +-- Plethobasus
| | +--+
| | +-- Elliptio
| |
| | +-------------------- Pleurobema_plenum
+--+ |
| +-------------------- Pleuronaia
| |
| | +-- Pleurobema_cordatum__1
| | +-----+
| | | +-- Fusconaia
+--+ +--------+
| | | +----- Pleurobema_sintoxia__1
| | | |
| | +--+ +-- Pleurobema_cordatum__2
| | | |
| | +--+-- Pleurobema_sintoxia__2
| | |
+--+ +-- Pleurobema_sintoxia__3
|
| +-------------- Pleurobema_stabile
| |
| | +-- Pleurobema_rubellum
| | +-----+
| | | +-- Pleurobema_strodeanum
+--+ +--+
| | | +----- Pleurobema_beadleianum
| | +--+
| | | +-- Pleurobema_perovatum
| | +--+
+--+ +-- Pleurobema_taitianum
|
| +-- Pleurobema_clava
| +-----+
| | +-- Pleurobema_oviforme
| |
+--+ +-- Pleurobema_decisum
| +--+
| | +-- Pleurobema_hanleyianum
+--+
| +-- Pleurobema_pyriforme
+--+
+-- Pleurobema_georgianum
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
29 |
0 |
0 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
8 |
1 |
1 |
-- |
+-- Venustaconcha **
--+
+-- Quadrulini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
16 |
1 |
1 |
-- |
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
12 |
1 |
1 |
-- |
+-- Quadrulini
--+
+-- Venustaconcha **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
67 |
0 |
0 |
-- |
|
