Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae

tribe LANCEOLARIINI Froufe, Lopes-Lima, & Bogan in Lopes-Lima et al., 2017: cladomics

1 genus

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 15 (17 total)
number of subtaxa (rank: genus) = 1
analyzed in 106 cladograms in 64 publications
maximum number of terminals in an analysis = 8 (mean = 2.27, median = 2)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 77
maximum number of included terminals = 8, (mean = 2.75 median = 2)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)
proportion of cladograms that support basic monophyly = 0.91

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Lanceolariini cladogram distances
PCoA Ordination

NJ Tree of Lanceolariini cladogram distances
NJ Tree

Cladograms

Sano et al., 2020 [Publ 4138], fig. 1

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
3 22 5 1 31 19 4 0 
  +----- Lanceolaria_grayii__1 **  
  |                             
  +----- Lanceolaria_grayii__2 **  
  |                             
  +----- Cristaria_plicata__1   
  |                             
  +----- Cristaria_plicata__2   
  |                             
  +----- Sinanodonta_lauta      
  |                             
  +----- Beringiana_japonica    
  |                             
  +----- Beringiana_fukuharai__1
  |                             
  +----- Beringiana_beringiana  
  |                             
  +----- Beringiana_fukuharai__2
  |                             
  +----- Beringiana_fukuharai__3
  |                             
  +----- Buldowskia_iwakawai__1 
  |                             
  +----- Buldowskia_iwakawai__2 
  |                             
  +----- Sinanodonta_tumens__1  
--+                             
  +----- Sinanodonta_tumens__2  
  |                             
  +----- Pletholophus           
  |                             
  +----- Anemina                
  |                             
  +----- Lanceolaria_lanceolata ** 
  |                             
  +----- Cristaria_plicata__3   
  |                             
  +----- Lanceolaria_gladiola **   
  |                             
  +----- Sinanodonta_angula     
  |                             
  +----- Lepidodesmini          
  |                             
  |  +-- Aculamprotulina        
  +--+                          
  |  +-- Acuticostini           
  |                             
  |  +-- Cuneopsis              
  +--+                          
  |  +-- Nodularia              
  |                             
  |  +-- Schistodesmus          
  +--+                          
     +-- Inversiunio

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
8 8 7 1 624 149 4 35 
  +----------------- Lepidodesmini               
  |                                              
  |     +----------- Lanceolaria **                 
  |  +--+                                        
--+  |  |  +-------- Pseudoacuticosta_retiaria__2
  |  |  +--+                                     
  |  |     |  +----- Acuticosta_chinensis__2     
  |  |     +--+                                  
  +--+        |  +-- Acuticosta_chinensis__1     
     |        +--+                               
     |           +-- Pseudoacuticosta_retiaria__1
     |                                           
     |  +----------- Globunio                    
     +--+                                        
        |  +-------- Aculamprotulina             
        +--+                                     
           |  +----- Middendorffinaiina          
           +--+                                  
              |  +-- Unionina                    
              +--+                               
                 +-- Oxynaiina

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
8 8 6 1 114 63 4 36 
  +----- Acuticostini 
  |                   
  +----- Lepidodesmini
--+                   
  +----- Unionini     
  |                   
  |  +-- Lanceolaria **  
  +--+                
     +-- Anodontini

Lopes-Lima et al., 2020 [Publ 4080], fig. 11

iTtTbiriSoTroTroSiD
8 8 6 1 4 2 1 36 
  +-- Unionini   
--+              
  +-- Lanceolaria **

Huang et al., 2019 [Publ 3843], fig. S1

iTtTbiriSoTroTroSiD
5 5 4 1 42 28 4 30 
  +-------------- Anemina      
  |                            
--+  +----------- Sinanodonta  
  |  |                         
  +--+  +-------- Acuticostini 
     |  |                      
     +--+     +-- Lepidodesmini
        |  +--+                
        |  |  +-- Cristaria    
        +--+                   
           |  +-- Lanceolaria **  
           +--+                
              +-- Unionini

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
4 4 2 1 41 27 4 33 
  +----------------- Anemina        
  |                                 
--+  +-------------- Acuticostini   
  |  |                              
  +--+  +----------- Sinanodonta    
     |  |                           
     +--+  +-------- Aculamprotulina
        |  |                        
        +--+     +-- Cristaria      
           |  +--+                  
           |  |  +-- Lanceolaria **    
           +--+                     
              |  +-- Lepidodesmini  
              +--+                  
                 +-- Oxynaiina

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 1 196 43 4 33 
  +-------- Acuticostini 
  |                      
--+  +----- Lepidodesmini
  |  |                   
  +--+----- Unionini     
     |                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
4 4 2 1 194 42 4 33 
  +----------- Acuticostini 
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lepidodesmini
     +--+                   
        |  +-- Lanceolaria **  
        +--+                
           +-- Anodontini

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
4 4 2 1 196 43 4 33 
  +-------- Acuticostini         
  |                              
  +-------- Alasmidonta          
  |                              
  +-------- Anodonta_anatina     
  |                              
  +-------- Anemina              
  |                              
  +-------- Anodonta_cygnea      
  |                              
  +-------- Cristaria            
  |                              
  +-------- Brachyanodon         
  |                              
  +-------- Lasmigona            
  |                              
  +-------- Lepidodesmini        
  |                              
  +-------- Pseudanodonta        
  |                              
  +-------- Pyganodon            
  |                              
--+-------- Simpsonaias          
  |                              
  +-------- Strophitus           
  |                              
  +-------- Utterbackia          
  |                              
  +-------- Lanceolaria **          
  |                              
  +-------- Schistodesmus        
  |                              
  +-------- Unionina             
  |                              
  +-------- Aculamprotulina      
  |                              
  |     +-- Pletholophus         
  +-----+                        
  |     +-- Sinanodonta          
  |                              
  |  +----- Cuneopsis_celtiformis
  |  |                           
  |  +----- Tchangsinaia         
  |  |                           
  +--+----- Cuneopsis_rufescens  
     |                           
     +----- Nodularia            
     |                           
     |  +-- Cuneopsis_szechenyii 
     +--+                        
        +-- Cuneopsis_heudei

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 1 49 29 4 33 
  +-------- Lepidodesmini
  |                      
--+-------- Unionini     
  |                      
  |  +----- Anodontini   
  +--+                   
     |  +-- Acuticostini 
     +--+                
        +-- Lanceolaria **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 1 71 33 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 1 53 23 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
3 3 2 1 77 25 2 32 
  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 1 73 69 3 32 
  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 1 20 18 3 32 
  +-------- Acuticostini
--+                     
  |  +----- Lanceolaria ** 
  +--+                  
     |  +-- Anodontini  
     +--+               
        +-- Unionini

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 1 66 30 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
3 3 1 1 89 44 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
3 3 1 1 93 44 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Xiang et al., 2025 [Publ 4433], fig. 7

iTtTbiriSoTroTroSiD
3 3 1 1 10 9 1 32 
  +-- Lanceolaria **
--+              
  +-- Anodontini

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
3 3 1 1 53 30 4 33 
  +-------- Unionini     
  |                      
--+     +-- Lepidodesmini
  |  +--+                
  |  |  +-- Lanceolaria **  
  +--+                   
     |  +-- Acuticostini 
     +--+                
        +-- Anodontini

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 1 51 25 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Lanceolaria **  
  +--+                      
     |  +----- Unionini     
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 1 93 63 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
3 3 1 1 71 31 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
3 3 1 1 89 44 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 1 85 40 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Kim et al., 2021 [Publ 4197], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 1 15 14 3 32 
  +-------- Aculamprotulina
  |                        
--+     +-- Acuticostini   
  |  +--+                  
  |  |  +-- Lanceolaria **    
  +--+                     
     |  +-- Unionina       
     +--+                  
        +-- Oxynaiina

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
3 3 1 1 36 36 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
3 3 1 1 88 23 4 32 
  +-------- Acuticostini 
  |                      
--+     +-- Lepidodesmini
  |  +--+                
  |  |  +-- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 1 14 14 2 32 
        +-- Unionina       
  +-----+                  
  |     +-- Oxynaiina      
--+                        
  |  +----- Aculamprotulina
  +--+                     
     |  +-- Acuticostini   
     +--+                  
        +-- Lanceolaria **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 1 83 41 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
3 21 14 1 21 19 3 43 
  +-------- Acuticostini          
  |                               
--+  +----- Lanceolaria_grayii **    
  |  |                            
  |  +----- Lanceolaria_gladiola **  
  +--+                            
     +----- Lanceolaria_lanceolata **
     |                            
     |  +-- Anodontini            
     +--+                         
        +-- Unionini

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
3 3 1 1 93 44 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
3 3 1 1 71 31 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
3 3 1 1 51 49 3 32 
  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
3 3 1 1 88 23 4 32 
  +----------- Acuticostini 
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Lepidodesmini
        +--+                
           +-- Anodontini

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
3 21 19 1 21 19 3 46 
  +----------- Acuticostini          
  |                                  
--+        +-- Lanceolaria_lanceolata **
  |  +-----+                         
  |  |     +-- Lanceolaria_grayii **    
  +--+                               
     |  +----- Lanceolaria_gladiola **  
     +--+                            
        |  +-- Anodontini            
        +--+                         
           +-- Unionini

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 39 30 4 -- 
  +-------------------- Unionini     
  |                                  
--+  +----------------- Acuticostini 
  |  |                               
  |  |              +-- Alasmidontina
  +--+  +-----------+                
     |  |           +-- Anodontina   
     |  |                            
     |  |        +----- Lepidodesmini
     +--+  +-----+                   
        |  |     |  +-- Cristaria    
        |  |     +--+                
        |  |        +-- Lanceolaria **  
        +--+                         
           |        +-- Anemina      
           |  +-----+                
           |  |     +-- Amuranodonta 
           +--+                      
              |  +----- Pletholophus 
              +--+                   
                 |  +-- Sinanodonta  
                 +--+                
                    +-- Beringiana

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 44 24 4 -- 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 80 15 2 -- 
  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 30 16 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 1 66 33 3 -- 
                 +-- Anemina      
  +--------------+                
  |              +-- Buldowskia   
  |                               
--+           +----- Sinanodonta  
  |  +--------+                   
  |  |        |  +-- Pletholophus 
  |  |        +--+                
  +--+           +-- Beringiana   
     |                            
     |  +----------- Cristaria    
     |  |                         
     +--+        +-- Anodontina   
        |  +-----+                
        |  |     +-- Alasmidontina
        +--+                      
           |  +----- Lepidodesmini
           +--+                   
              |  +-- Lanceolaria **  
              +--+                
                 +-- Unionini

Zhang et al., 2004 [Publ 2735], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 12 10 3 -- 
  +-------- Anodontini  
--+                     
  |  +----- Unionini    
  +--+                  
     |  +-- Acuticostini
     +--+               
        +-- Lanceolaria **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
2 2 0 1 37 15 4 -- 
  +----------- Anodontini     
  |                           
  |        +-- Acuticostini   
--+  +-----+                  
  |  |     +-- Lanceolaria **    
  |  |                        
  +--+     +-- Lepidodesmini  
     |  +--+                  
     |  |  +-- Aculamprotulina
     +--+                     
        |  +-- Unionina       
        +--+                  
           +-- Oxynaiina

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 21 21 3 -- 
  +-------- Anodontini   
--+                      
  |  +----- Lanceolaria **  
  +--+                   
     |  +-- Lepidodesmini
     +--+                
        +-- Unionini

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
2 2 0 1 139 67 4 -- 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 32 12 4 -- 
  +----------- Anodontini   
  |                         
  |     +----- Acuticostini 
--+  +--+                   
  |  |  |  +-- Nodularia    
  |  |  +--+                
  +--+     +-- Tchangsinaia 
     |                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Schistodesmus
        +--+                
           +-- Lepidodesmini

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
2 2 0 1 136 25 2 -- 
  +----------- Unionini        
  |                            
--+  +-------- Lanceolaria **     
  |  |                         
  |  +-------- Cristariina     
  +--+                         
     |  +----- Anodonta_anatina
     |  |                      
     |  +----- Pseudanodonta   
     +--+                      
        +----- Alasmidontina   
        |                      
        |  +-- Anodonta_cygnea 
        +--+                   
           +-- Brachyanodon

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 32 12 4 -- 
  +-------- Strophitus   
  |                      
  +-------- Schistodesmus
  |                      
  +-------- Lepidodesmini
  |                      
  +-------- Acuticostini 
  |                      
  +-------- Cristariina  
--+                      
  +-------- Lanceolaria **  
  |                      
  |     +-- Nodularia    
  +-----+                
  |     +-- Tchangsinaia 
  |                      
  |  +----- Utterbackia  
  |  |                   
  +--+----- Utterbackiana
     |                   
     |  +-- Alasmidonta  
     +--+                
        +-- Lasmigona

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 29 16 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Sano et al., 2017 [Publ 3799], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 31 19 4 -- 
  +-------------- Anodontini   
  |                            
  |           +-- Tchangsinaia 
--+  +--------+                
  |  |        +-- Nodularia    
  |  |                         
  +--+        +-- Schistodesmus
     |  +-----+                
     |  |     +-- Acuticostini 
     +--+                      
        |  +----- Inversiunio  
        +--+                   
           |  +-- Lepidodesmini
           +--+                
              +-- Lanceolaria **

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 29 15 3 -- 
  +-------- Unionini    
--+                     
  |  +----- Anodontini  
  +--+                  
     |  +-- Acuticostini
     +--+               
        +-- Lanceolaria **

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 34 17 2 -- 
  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Xue et al., 2016 [Publ 3659], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 13 9 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Wu et al., 2026 [Publ 4811], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 15 8 3 -- 
  +----------- Unionina    
--+                        
  |  +-------- Lanceolaria ** 
  +--+                     
     |  +----- Globunionina
     +--+                  
        |  +-- Acuticostini
        +--+               
           +-- Anodontini

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 43 34 4 -- 
           +----- Pletholophus 
  +--------+                   
  |        |  +-- Sinanodonta  
  |        +--+                
  |           +-- Beringiana   
--+                            
  |  +----------- Cristaria    
  |  |                         
  |  |        +-- Alasmidontina
  +--+--------+                
     |        +-- Anodontina   
     |                         
     |  +-------- Lanceolaria **  
     +--+                      
        |  +----- Lepidodesmini
        +--+                   
           |  +-- Acuticostini 
           +--+                
              +-- Unionini

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 29 15 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
2 27 0 1 43 34 4 42 
              +-- Anemina               
  +-----------+                         
  |           +-- Amuranodonta          
  |                                     
  |        +----- Pletholophus          
--+  +-----+                            
  |  |     |  +-- Sinanodonta           
  |  |     +--+                         
  |  |        +-- Beringiana            
  +--+                                  
     |        +-- Alasmidontina         
     |  +-----+                         
     |  |     +-- Anodontina            
     |  |                               
     +--+  +----- Lanceolaria_gladiola **  
        |  |                            
        |  +----- Lanceolaria_oxyrhyncha **
        |  |                            
        +--+----- Cristaria             
           |                            
           |  +-- Acuticostini          
           |  |                         
           +--+-- Lepidodesmini         
              |                         
              +-- Unionini

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
2 2 0 1 136 25 2 -- 
  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 38 24 4 -- 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Riccardi et al., 2020 [Publ 4089], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 17 15 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
2 27 0 1 39 30 4 42 
           +-- Anemina               
  +--------+                         
  |        +-- Amuranodonta          
  |                                  
  |     +----- Pletholophus          
--+  +--+                            
  |  |  |  +-- Sinanodonta           
  |  |  +--+                         
  |  |     +-- Beringiana            
  |  |                               
  +--+  +----- Lanceolaria_gladiola **  
     |  |                            
     |  +----- Acuticostini          
     |  |                            
     |  +----- Lepidodesmini         
     |  |                            
     +--+----- Lanceolaria_oxyrhyncha **
        |                            
        +----- Cristaria             
        |                            
        +----- Unionini              
        |                            
        |  +-- Alasmidontina         
        +--+                         
           +-- Anodontina

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
2 17 0 1 122 25 2 37 
  +----- Unionini             
  |                           
  |  +-- Alasmidonta          
  |  |                        
  |  +-- Anodonta_anatina     
  |  |                        
  |  +-- Anodonta_cygnea      
  |  |                        
  |  +-- Brachyanodon         
--+  |                        
  |  +-- Lasmigona            
  |  |                        
  |  +-- Pseudanodonta        
  |  |                        
  |  +-- Pyganodon            
  |  |                        
  |  +-- Simpsonaias          
  |  |                        
  |  +-- Strophitus           
  +--+                        
     +-- Sinanodonta_lauta    
     |                        
     +-- Cristaria_plicata    
     |                        
     +-- Cristaria_radiata    
     |                        
     +-- Pletholophus         
     |                        
     +-- Sinanodonta_lucida   
     |                        
     +-- Sinanodonta_jourdyi  
     |                        
     +-- Lanceolaria_gladiola ** 
     |                        
     +-- Lanceolaria_grayii__1 **
     |                        
     +-- Lanceolaria_grayii__2 **

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 39 30 4 -- 
  +-------------- Globunionina   
  |                              
  |        +----- Aculamprotulina
--+  +-----+                     
  |  |     |  +-- Unionina       
  |  |     +--+                  
  +--+        +-- Oxynaiina      
     |                           
     |  +-------- Acuticostini   
     +--+                        
        |  +----- Lepidodesmini  
        +--+                     
           |  +-- Lanceolaria **    
           +--+                  
              +-- Anodontini

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 39 30 4 -- 
  +-------- Acuticostini 
  |                      
--+  +----- Lepidodesmini
  |  |                   
  +--+----- Unionini     
     |                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 1 35 14 4 -- 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Ouyang et al., 2015 [Publ 4493], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 29 14 3 -- 
  +-------- Unionini    
--+                     
  |  +----- Lanceolaria ** 
  +--+                  
     |  +-- Acuticostini
     +--+               
        +-- Anodontini

Ouyang et al., 2011 [Publ 3408], fig. 1

iTtTbiriSoTroTroSiD
2 4 0 1 11 7 3 32 
  +----------- Anodontini          
  |                                
--+  +-------- Oxynaiina           
  |  |                             
  +--+  +----- Lanceolaria_grayii **  
     |  |                          
     +--+----- Acuticostini        
        |                          
        |  +-- Lanceolaria_gladiola **
        +--+                       
           +-- Aculamprotulina

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 1 42 20 3 -- 
  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Ouyang et al., 2015 [Publ 4493], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 29 14 3 -- 
  +-------- Lanceolaria **    
  |                        
--+     +-- Aculamprotulina
  |  +--+                  
  |  |  +-- Acuticostini   
  +--+                     
     |  +-- Anodontini     
     +--+                  
        +-- Oxynaiina

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 47 25 4 -- 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 42 20 3 -- 
  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 66 33 3 -- 
                 +-- Anemina      
  +--------------+                
  |              +-- Buldowskia   
  |                               
--+           +----- Pletholophus 
  |  +--------+                   
  |  |        |  +-- Beringiana   
  |  |        +--+                
  +--+           +-- Sinanodonta  
     |                            
     |  +----------- Cristaria    
     |  |                         
     +--+        +-- Anodontina   
        |  +-----+                
        |  |     +-- Alasmidontina
        +--+                      
           |  +----- Lanceolaria **  
           +--+                   
              |  +-- Lepidodesmini
              +--+                
                 +-- Unionini

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
2 2 0 1 74 24 2 -- 
  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
2 2 0 1 46 20 3 -- 
  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 19 11 3 -- 
     +----- Aculamprotulina
  +--+                     
  |  |  +-- Tchangsinaia   
  |  +--+                  
--+     +-- Nodularia      
  |                        
  |  +----- Schistodesmus  
  +--+                     
     |  +-- Lanceolaria **    
     +--+                  
        +-- Acuticostini

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 66 33 3 -- 
                 +-- Anemina      
  +--------------+                
  |              +-- Buldowskia   
  |                               
--+           +----- Sinanodonta  
  |  +--------+                   
  |  |        |  +-- Pletholophus 
  |  |        +--+                
  +--+           +-- Beringiana   
     |                            
     |  +----------- Cristaria    
     |  |                         
     +--+        +-- Anodontina   
        |  +-----+                
        |  |     +-- Alasmidontina
        +--+                      
           |  +----- Lanceolaria **  
           +--+                   
              |  +-- Lepidodesmini
              +--+                
                 +-- Unionini

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
2 2 0 1 74 24 2 -- 
  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 12 2 -- 
  +-------- Aculamprotulina
  |                        
--+     +-- Oxynaiina      
  |  +--+                  
  |  |  +-- Unionina       
  +--+                     
     |  +-- Lanceolaria **    
     +--+                  
        +-- Anodontini

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
1 1 0 1 21 16 4 -- 
  +----------- Anodontina   
  |                         
--+        +-- Acuticostini 
  |  +-----+                
  |  |     +-- Cristariina  
  +--+                      
     |  +----- Lepidodesmini
     +--+                   
        |  +-- Lanceolaria **  
        +--+                
           +-- Unionini

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 16 4 -- 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 18 0 0 -- 
-- Lanceolaria_grayii **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 7 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 41 27 4 -- 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Klymus et al., 2025 [Publ 4814], fig. 3B

iTtTbiriSoTroTroSiD
1 1 0 1 35 14 2 -- 
  +----------- Anodontini     
--+                           
  |  +-------- Oxynaiina      
  +--+                        
     |  +----- Unionina       
     +--+                     
        |  +-- Aculamprotulina
        +--+                  
           +-- Lanceolaria **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 52 17 3 -- 
  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Klymus et al., 2025 [Publ 4814], fig. 3A

iTtTbiriSoTroTroSiD
1 1 0 1 35 14 2 -- 
  +----- Lanceolaria **
--+                 
  |  +-- Anodontini 
  +--+              
     +-- Unionini

Sano et al., 2017 [Publ 3799], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 20 12 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 11 7 2 -- 
  +-------- Anodontini 
--+                    
  |  +----- Nodularia  
  +--+                 
     |  +-- Lanceolaria **
     +--+              
        +-- Inversiunio

Zhou et al., 2017 [Publ 3788], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 15 5 2 -- 
  +-- Lanceolaria **
--+              
  +-- Unionini

Wu et al., 2022 [Publ 4238], fig. 2e

iTtTbiriSoTroTroSiD
1 1 0 1 17 17 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

iTtTbiriSoTroTroSiD
1 1 0 1 21 21 2 -- 
  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 52 27 4 -- 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 9 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Plazzi et al., 2011 [Publ 3676], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 17 8 2 -- 
-- Lanceolaria_grayii **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iTtTbiriSoTroTroSiD
1 1 0 1 9 9 2 -- 
  +----- Lanceolaria **
--+                 
  |  +-- Anodontini 
  +--+              
     +-- Unionini

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 37 20 3 -- 
  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
1 1 0 1 19 9 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 9 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 7 2 -- 
  +----- Lanceolaria **
--+                 
  |  +-- Unionini   
  +--+              
     +-- Anodontini

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

iTtTbiriSoTroTroSiD
1 1 0 1 21 21 2 -- 
  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 46 18 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 22 9 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
1 1 0 1 21 16 4 -- 
  +----------- Acuticostini      
  |                              
  +----------- Cristariina       
  |                              
--+----------- Anodontina        
  |                              
  |  +-------- Lanceolaria **       
  |  |                           
  |  |     +-- Aculamprotulina   
  +--+-----+                     
     |     +-- Lepidodesmini     
     |                           
     |  +----- Unionina          
     +--+                        
        |  +-- Middendorffinaiina
        +--+                     
           +-- Oxynaiina

Liu et al., 2017 [Publ 3939], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 13 3 -- 
  +-- Lanceolaria **
--+              
  +-- Unionini

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 12 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Kano et al., 2019 [Publ 4194], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 9 5 2 -- 
  +-- Lanceolaria **
--+              
  +-- Unionini