Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae

tribe LANCEOLARIINI Froufe, Lopes-Lima, & Bogan in Lopes-Lima et al., 2017: cladomics

1 genus

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 15 (17 total)
number of subtaxa (rank: genus) = 1
analyzed in 99 cladograms in 61 publications
maximum number of terminals in an analysis = 8 (mean = 2.25, median = 2)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 73
maximum number of included terminals = 8, (mean = 2.7 median = 2)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)
proportion of cladograms that support basic monophyly = 0.9

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Lanceolariini cladogram distances
PCoA Ordination

NJ Tree of Lanceolariini cladogram distances
NJ Tree

Cladograms

Sano et al., 2020 [Publ 4138], fig. 1

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
3 22 5 1 31 19 4 0 
  +----- Lanceolaria_grayii__1 **  
  |                             
  +----- Lanceolaria_grayii__2 **  
  |                             
  +----- Cristaria_plicata__1   
  |                             
  +----- Cristaria_plicata__2   
  |                             
  +----- Sinanodonta_lauta      
  |                             
  +----- Beringiana_japonica    
  |                             
  +----- Beringiana_fukuharai__1
  |                             
  +----- Beringiana_beringiana  
  |                             
  +----- Beringiana_fukuharai__2
  |                             
  +----- Beringiana_fukuharai__3
  |                             
  +----- Buldowskia_iwakawai__1 
  |                             
  +----- Buldowskia_iwakawai__2 
  |                             
  +----- Sinanodonta_tumens__1  
--+                             
  +----- Sinanodonta_tumens__2  
  |                             
  +----- Pletholophus           
  |                             
  +----- Anemina                
  |                             
  +----- Lanceolaria_lanceolata ** 
  |                             
  +----- Cristaria_plicata__3   
  |                             
  +----- Lanceolaria_gladiola **   
  |                             
  +----- Sinanodonta_angula     
  |                             
  +----- Lepidodesmini          
  |                             
  |  +-- Aculamprotulina        
  +--+                          
  |  +-- Acuticostini           
  |                             
  |  +-- Cuneopsis              
  +--+                          
  |  +-- Nodularia              
  |                             
  |  +-- Schistodesmus          
  +--+                          
     +-- Inversiunio

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
8 8 6 1 114 63 4 36 
  +----- Acuticostini 
  |                   
  +----- Lepidodesmini
--+                   
  +----- Unionini     
  |                   
  |  +-- Lanceolaria **  
  +--+                
     +-- Anodontini

Lopes-Lima et al., 2020 [Publ 4080], fig. 11

iTtTbiriSoTroTroSiD
8 8 6 1 4 2 1 36 
  +-- Unionini   
--+              
  +-- Lanceolaria **

Huang et al., 2019 [Publ 3843], fig. S1

iTtTbiriSoTroTroSiD
5 5 4 1 42 28 4 30 
  +-------------- Anemina      
  |                            
--+  +----------- Sinanodonta  
  |  |                         
  +--+  +-------- Acuticostini 
     |  |                      
     +--+     +-- Lepidodesmini
        |  +--+                
        |  |  +-- Cristaria    
        +--+                   
           |  +-- Lanceolaria **  
           +--+                
              +-- Unionini

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 1 49 29 4 33 
  +-------- Lepidodesmini
  |                      
--+-------- Unionini     
  |                      
  |  +----- Anodontini   
  +--+                   
     |  +-- Acuticostini 
     +--+                
        +-- Lanceolaria **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 1 197 43 4 33 
  +-------- Acuticostini 
  |                      
--+  +----- Lepidodesmini
  |  |                   
  +--+----- Unionini     
     |                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
4 4 2 1 195 42 4 33 
  +----------- Acuticostini 
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lepidodesmini
     +--+                   
        |  +-- Lanceolaria **  
        +--+                
           +-- Anodontini

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
4 4 2 1 197 43 4 33 
  +-------- Acuticostini         
  |                              
  +-------- Alasmidonta          
  |                              
  +-------- Anodonta_anatina     
  |                              
  +-------- Anemina              
  |                              
  +-------- Anodonta_cygnea      
  |                              
  +-------- Cristaria            
  |                              
  +-------- Anodonta_nuttalliana 
  |                              
  +-------- Lasmigona            
  |                              
  +-------- Lepidodesmini        
  |                              
  +-------- Pseudanodonta        
  |                              
  +-------- Pyganodon            
  |                              
--+-------- Simpsonaias          
  |                              
  +-------- Strophitus           
  |                              
  +-------- Utterbackia          
  |                              
  +-------- Lanceolaria **          
  |                              
  +-------- Schistodesmus        
  |                              
  +-------- Unionina             
  |                              
  +-------- Aculamprotulina      
  |                              
  |     +-- Pletholophus         
  +-----+                        
  |     +-- Sinanodonta          
  |                              
  |  +----- Cuneopsis_celtiformis
  |  |                           
  |  +----- Tchangsinaia         
  |  |                           
  +--+----- Cuneopsis_rufescens  
     |                           
     +----- Nodularia            
     |                           
     |  +-- Cuneopsis_szechenyii 
     +--+                        
        +-- Cuneopsis_heudei

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
4 4 2 1 41 27 4 33 
  +----------------- Anemina        
  |                                 
--+  +-------------- Acuticostini   
  |  |                              
  +--+  +----------- Sinanodonta    
     |  |                           
     +--+  +-------- Aculamprotulina
        |  |                        
        +--+     +-- Cristaria      
           |  +--+                  
           |  |  +-- Lanceolaria **    
           +--+                     
              |  +-- Lepidodesmini  
              +--+                  
                 +-- Oxynaiina

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
3 3 1 1 71 31 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
3 3 1 1 51 49 3 32 
  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
3 3 2 1 77 25 2 32 
  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 1 71 33 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 1 20 18 3 32 
  +-------- Acuticostini
--+                     
  |  +----- Lanceolaria ** 
  +--+                  
     |  +-- Anodontini  
     +--+               
        +-- Unionini

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
3 3 1 1 53 30 4 33 
  +-------- Unionini     
  |                      
--+     +-- Lepidodesmini
  |  +--+                
  |  |  +-- Lanceolaria **  
  +--+                   
     |  +-- Acuticostini 
     +--+                
        +-- Anodontini

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 1 66 30 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
3 3 1 1 89 44 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
3 3 1 1 36 36 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
3 3 1 1 93 44 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
3 3 1 1 88 23 4 32 
  +----------- Acuticostini 
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Lepidodesmini
        +--+                
           +-- Anodontini

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
3 3 1 1 93 44 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 1 53 23 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
3 3 1 1 89 44 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
3 3 1 1 88 23 4 32 
  +-------- Acuticostini 
  |                      
--+     +-- Lepidodesmini
  |  +--+                
  |  |  +-- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
3 21 14 1 21 19 3 43 
  +-------- Acuticostini          
  |                               
--+  +----- Lanceolaria_grayii **    
  |  |                            
  |  +----- Lanceolaria_gladiola **  
  +--+                            
     +----- Lanceolaria_lanceolata **
     |                            
     |  +-- Anodontini            
     +--+                         
        +-- Unionini

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 1 51 25 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Lanceolaria **  
  +--+                      
     |  +----- Unionini     
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Xiang et al., 2025 [Publ 4433], fig. 7

iTtTbiriSoTroTroSiD
3 3 1 1 10 9 1 32 
  +-- Lanceolaria **
--+              
  +-- Anodontini

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
3 21 19 1 21 19 3 46 
  +----------- Acuticostini          
  |                                  
--+        +-- Lanceolaria_lanceolata **
  |  +-----+                         
  |  |     +-- Lanceolaria_grayii **    
  +--+                               
     |  +----- Lanceolaria_gladiola **  
     +--+                            
        |  +-- Anodontini            
        +--+                         
           +-- Unionini

Kim et al., 2021 [Publ 4197], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 1 15 14 3 32 
  +-------- Aculamprotulina
  |                        
--+     +-- Acuticostini   
  |  +--+                  
  |  |  +-- Lanceolaria **    
  +--+                     
     |  +-- Unionina       
     +--+                  
        +-- Oxynaiina

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 1 85 40 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
3 3 1 1 71 31 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 1 14 14 2 32 
        +-- Unionina       
  +-----+                  
  |     +-- Oxynaiina      
--+                        
  |  +----- Aculamprotulina
  +--+                     
     |  +-- Acuticostini   
     +--+                  
        +-- Lanceolaria **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 1 83 41 4 32 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 1 73 69 3 32 
  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 19 11 3 -- 
     +----- Aculamprotulina
  +--+                     
  |  |  +-- Tchangsinaia   
  |  +--+                  
--+     +-- Nodularia      
  |                        
  |  +----- Schistodesmus  
  +--+                     
     |  +-- Lanceolaria **    
     +--+                  
        +-- Acuticostini

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 47 25 4 -- 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 29 15 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
2 2 0 1 74 24 2 -- 
  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
2 27 0 1 43 34 4 42 
              +-- Anemina               
  +-----------+                         
  |           +-- Amuranodonta          
  |                                     
  |        +----- Pletholophus          
--+  +-----+                            
  |  |     |  +-- Sinanodonta           
  |  |     +--+                         
  |  |        +-- Beringiana            
  +--+                                  
     |        +-- Alasmidontina         
     |  +-----+                         
     |  |     +-- Anodontina            
     |  |                               
     +--+  +----- Lanceolaria_gladiola **  
        |  |                            
        |  +----- Lanceolaria_oxyrhyncha **
        |  |                            
        +--+----- Cristaria             
           |                            
           |  +-- Acuticostini          
           |  |                         
           +--+-- Lepidodesmini         
              |                         
              +-- Unionini

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 42 20 3 -- 
  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 1 42 20 3 -- 
  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 66 33 3 -- 
                 +-- Anemina      
  +--------------+                
  |              +-- Buldowskia   
  |                               
--+           +----- Sinanodonta  
  |  +--------+                   
  |  |        |  +-- Pletholophus 
  |  |        +--+                
  +--+           +-- Beringiana   
     |                            
     |  +----------- Cristaria    
     |  |                         
     +--+        +-- Anodontina   
        |  +-----+                
        |  |     +-- Alasmidontina
        +--+                      
           |  +----- Lanceolaria **  
           +--+                   
              |  +-- Lepidodesmini
              +--+                
                 +-- Unionini

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
2 17 0 1 123 25 2 37 
  +----- Unionini             
  |                           
  |  +-- Alasmidonta          
  |  |                        
  |  +-- Anodonta_anatina     
  |  |                        
  |  +-- Anodonta_cygnea      
  |  |                        
  |  +-- Anodonta_nuttalliana 
--+  |                        
  |  +-- Lasmigona            
  |  |                        
  |  +-- Pseudanodonta        
  |  |                        
  |  +-- Pyganodon            
  |  |                        
  |  +-- Simpsonaias          
  |  |                        
  |  +-- Strophitus           
  +--+                        
     +-- Sinanodonta_lauta    
     |                        
     +-- Cristaria_plicata    
     |                        
     +-- Cristaria_radiata    
     |                        
     +-- Pletholophus         
     |                        
     +-- Sinanodonta_lucida   
     |                        
     +-- Sinanodonta_jourdyi  
     |                        
     +-- Lanceolaria_gladiola ** 
     |                        
     +-- Lanceolaria_grayii__1 **
     |                        
     +-- Lanceolaria_grayii__2 **

Riccardi et al., 2020 [Publ 4089], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 17 15 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 39 30 4 -- 
  +-------------- Globunionina   
  |                              
  |        +----- Aculamprotulina
--+  +-----+                     
  |  |     |  +-- Unionina       
  |  |     +--+                  
  +--+        +-- Oxynaiina      
     |                           
     |  +-------- Acuticostini   
     +--+                        
        |  +----- Lepidodesmini  
        +--+                     
           |  +-- Lanceolaria **    
           +--+                  
              +-- Anodontini

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 29 16 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 39 30 4 -- 
  +-------- Acuticostini 
  |                      
--+  +----- Lepidodesmini
  |  |                   
  +--+----- Unionini     
     |                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 44 24 4 -- 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Ouyang et al., 2015 [Publ 4493], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 29 14 3 -- 
  +-------- Lanceolaria **    
  |                        
--+     +-- Aculamprotulina
  |  +--+                  
  |  |  +-- Acuticostini   
  +--+                     
     |  +-- Anodontini     
     +--+                  
        +-- Oxynaiina

Zhang et al., 2004 [Publ 2735], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 12 10 3 -- 
  +-------- Anodontini  
--+                     
  |  +----- Unionini    
  +--+                  
     |  +-- Acuticostini
     +--+               
        +-- Lanceolaria **

Xue et al., 2016 [Publ 3659], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 13 9 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 66 33 3 -- 
                 +-- Anemina      
  +--------------+                
  |              +-- Buldowskia   
  |                               
--+           +----- Pletholophus 
  |  +--------+                   
  |  |        |  +-- Beringiana   
  |  |        +--+                
  +--+           +-- Sinanodonta  
     |                            
     |  +----------- Cristaria    
     |  |                         
     +--+        +-- Anodontina   
        |  +-----+                
        |  |     +-- Alasmidontina
        +--+                      
           |  +----- Lanceolaria **  
           +--+                   
              |  +-- Lepidodesmini
              +--+                
                 +-- Unionini

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 21 21 3 -- 
  +-------- Anodontini   
--+                      
  |  +----- Lanceolaria **  
  +--+                   
     |  +-- Lepidodesmini
     +--+                
        +-- Unionini

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 32 12 4 -- 
  +----------- Anodontini   
  |                         
  |     +----- Acuticostini 
--+  +--+                   
  |  |  |  +-- Nodularia    
  |  |  +--+                
  +--+     +-- Tchangsinaia 
     |                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Schistodesmus
        +--+                
           +-- Lepidodesmini

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 1 35 14 4 -- 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
2 2 0 1 46 20 3 -- 
  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 81 15 2 -- 
  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
2 2 0 1 74 24 2 -- 
  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 34 17 2 -- 
  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
2 2 0 1 37 15 4 -- 
  +----------- Anodontini     
  |                           
  |        +-- Acuticostini   
--+  +-----+                  
  |  |     +-- Lanceolaria **    
  |  |                        
  +--+     +-- Lepidodesmini  
     |  +--+                  
     |  |  +-- Aculamprotulina
     +--+                     
        |  +-- Unionina       
        +--+                  
           +-- Oxynaiina

Ouyang et al., 2015 [Publ 4493], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 29 14 3 -- 
  +-------- Unionini    
--+                     
  |  +----- Lanceolaria ** 
  +--+                  
     |  +-- Acuticostini
     +--+               
        +-- Anodontini

Ouyang et al., 2011 [Publ 3408], fig. 1

iTtTbiriSoTroTroSiD
2 4 0 1 11 7 3 32 
  +----------- Anodontini          
  |                                
--+  +-------- Oxynaiina           
  |  |                             
  +--+  +----- Lanceolaria_grayii **  
     |  |                          
     +--+----- Acuticostini        
        |                          
        |  +-- Lanceolaria_gladiola **
        +--+                       
           +-- Aculamprotulina

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 43 34 4 -- 
           +----- Pletholophus 
  +--------+                   
  |        |  +-- Sinanodonta  
  |        +--+                
  |           +-- Beringiana   
--+                            
  |  +----------- Cristaria    
  |  |                         
  |  |        +-- Alasmidontina
  +--+--------+                
     |        +-- Anodontina   
     |                         
     |  +-------- Lanceolaria **  
     +--+                      
        |  +----- Lepidodesmini
        +--+                   
           |  +-- Acuticostini 
           +--+                
              +-- Unionini

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 30 16 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 38 24 4 -- 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
2 2 0 1 137 25 2 -- 
  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
2 2 0 1 137 25 2 -- 
  +----------- Unionini            
  |                                
--+  +-------- Lanceolaria **         
  |  |                             
  |  +-------- Cristariina         
  +--+                             
     |  +----- Anodonta_anatina    
     |  |                          
     |  +----- Pseudanodonta       
     +--+                          
        +----- Alasmidontina       
        |                          
        |  +-- Anodonta_cygnea     
        +--+                       
           +-- Anodonta_nuttalliana

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
2 27 0 1 39 30 4 42 
           +-- Anemina               
  +--------+                         
  |        +-- Amuranodonta          
  |                                  
  |     +----- Pletholophus          
--+  +--+                            
  |  |  |  +-- Sinanodonta           
  |  |  +--+                         
  |  |     +-- Beringiana            
  |  |                               
  +--+  +----- Lanceolaria_gladiola **  
     |  |                            
     |  +----- Acuticostini          
     |  |                            
     |  +----- Lepidodesmini         
     |  |                            
     +--+----- Lanceolaria_oxyrhyncha **
        |                            
        +----- Cristaria             
        |                            
        +----- Unionini              
        |                            
        |  +-- Alasmidontina         
        +--+                         
           +-- Anodontina

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 32 12 4 -- 
  +-------- Strophitus   
  |                      
  +-------- Schistodesmus
  |                      
  +-------- Lepidodesmini
  |                      
  +-------- Acuticostini 
  |                      
  +-------- Cristariina  
--+                      
  +-------- Lanceolaria **  
  |                      
  |     +-- Nodularia    
  +-----+                
  |     +-- Tchangsinaia 
  |                      
  |  +----- Utterbackia  
  |  |                   
  +--+----- Utterbackiana
     |                   
     |  +-- Alasmidonta  
     +--+                
        +-- Lasmigona

Sano et al., 2017 [Publ 3799], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 31 19 4 -- 
  +-------------- Anodontini   
  |                            
  |           +-- Tchangsinaia 
--+  +--------+                
  |  |        +-- Nodularia    
  |  |                         
  +--+        +-- Schistodesmus
     |  +-----+                
     |  |     +-- Acuticostini 
     +--+                      
        |  +----- Inversiunio  
        +--+                   
           |  +-- Lepidodesmini
           +--+                
              +-- Lanceolaria **

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 29 15 3 -- 
  +-------- Unionini    
--+                     
  |  +----- Anodontini  
  +--+                  
     |  +-- Acuticostini
     +--+               
        +-- Lanceolaria **

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 39 30 4 -- 
  +-------------------- Unionini     
  |                                  
--+  +----------------- Acuticostini 
  |  |                               
  |  |              +-- Alasmidontina
  +--+  +-----------+                
     |  |           +-- Anodontina   
     |  |                            
     |  |        +----- Lepidodesmini
     +--+  +-----+                   
        |  |     |  +-- Cristaria    
        |  |     +--+                
        |  |        +-- Lanceolaria **  
        +--+                         
           |        +-- Anemina      
           |  +-----+                
           |  |     +-- Amuranodonta 
           +--+                      
              |  +----- Pletholophus 
              +--+                   
                 |  +-- Sinanodonta  
                 +--+                
                    +-- Beringiana

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 1 66 33 3 -- 
                 +-- Anemina      
  +--------------+                
  |              +-- Buldowskia   
  |                               
--+           +----- Sinanodonta  
  |  +--------+                   
  |  |        |  +-- Pletholophus 
  |  |        +--+                
  +--+           +-- Beringiana   
     |                            
     |  +----------- Cristaria    
     |  |                         
     +--+        +-- Anodontina   
        |  +-----+                
        |  |     +-- Alasmidontina
        +--+                      
           |  +----- Lepidodesmini
           +--+                   
              |  +-- Lanceolaria **  
              +--+                
                 +-- Unionini

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 18 0 0 -- 
-- Lanceolaria_grayii **

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 22 9 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iTtTbiriSoTroTroSiD
1 1 0 1 9 9 2 -- 
  +----- Lanceolaria **
--+                 
  |  +-- Anodontini 
  +--+              
     +-- Unionini

Wu et al., 2022 [Publ 4238], fig. 2e

iTtTbiriSoTroTroSiD
1 1 0 1 17 17 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 12 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 52 27 4 -- 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 9 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Kano et al., 2019 [Publ 4194], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 9 5 2 -- 
  +-- Lanceolaria **
--+              
  +-- Unionini

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 7 2 -- 
  +----- Lanceolaria **
--+                 
  |  +-- Unionini   
  +--+              
     +-- Anodontini

Plazzi et al., 2011 [Publ 3676], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 17 8 2 -- 
-- Lanceolaria_grayii **

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

iTtTbiriSoTroTroSiD
1 1 0 1 21 21 2 -- 
  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 37 20 3 -- 
  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 9 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Liu et al., 2017 [Publ 3939], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 13 3 -- 
  +-- Lanceolaria **
--+              
  +-- Unionini

Sano et al., 2017 [Publ 3799], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 20 12 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 41 27 4 -- 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 11 7 2 -- 
  +-------- Anodontini 
--+                    
  |  +----- Nodularia  
  +--+                 
     |  +-- Lanceolaria **
     +--+              
        +-- Inversiunio

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 12 2 -- 
  +-------- Aculamprotulina
  |                        
--+     +-- Oxynaiina      
  |  +--+                  
  |  |  +-- Unionina       
  +--+                     
     |  +-- Lanceolaria **    
     +--+                  
        +-- Anodontini

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
1 1 0 1 19 9 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 52 17 3 -- 
  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
1 1 0 1 21 16 4 -- 
  +----------- Anodontina   
  |                         
--+        +-- Acuticostini 
  |  +-----+                
  |  |     +-- Cristariina  
  +--+                      
     |  +----- Lepidodesmini
     +--+                   
        |  +-- Lanceolaria **  
        +--+                
           +-- Unionini

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
1 1 0 1 21 16 4 -- 
  +----------- Acuticostini      
  |                              
  +----------- Cristariina       
  |                              
--+----------- Anodontina        
  |                              
  |  +-------- Lanceolaria **       
  |  |                           
  |  |     +-- Aculamprotulina   
  +--+-----+                     
     |     +-- Lepidodesmini     
     |                           
     |  +----- Unionina          
     +--+                        
        |  +-- Middendorffinaiina
        +--+                     
           +-- Oxynaiina

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 7 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

iTtTbiriSoTroTroSiD
1 1 0 1 21 21 2 -- 
  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Zhou et al., 2017 [Publ 3788], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 15 5 2 -- 
  +-- Lanceolaria **
--+              
  +-- Unionini

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 16 4 -- 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **