Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae

tribe LEPIDODESMINI Huang & Wu in Huang et al. 2019: cladomics

1 genus

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 2 (3 total)
number of subtaxa (rank: genus) = 1
analyzed in 68 cladograms in 40 publications
maximum number of terminals in an analysis = 2 (mean = 1.19, median = 1)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 13
maximum number of included terminals = 2, (mean = 2 median = 2)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)
proportion of cladograms that support basic monophyly = 1

Cladograms

Aunins et al., 2018 [Publ 3863], fig. 1

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 39 25 4 **
Cladomics cladogram Lepidodesma UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Huang et al., 2002 [Publ 2657], fig. 1

cst iT tT bi riS oT roT roS iD
0.909 1 1 0 1 33 13 4 0.0
Cladomics cladogram ANODONTINI ACUTICOSTINI Nodularia Tchangsinaia LANCEOLARIINI Schistodesmus Lepidodesma

Huang et al., 2002 [Publ 2657], fig. 2

cst iT tT bi riS oT roT roS iD
0.818 1 1 0 1 33 13 4 0.0
Cladomics cladogram Strophitus Schistodesmus Lepidodesma ACUTICOSTINI CRISTARIINA LANCEOLARIINI Nodularia Tchangsinaia Utterbackia Utterbackiana Alasmidonta Lasmigona

Sano et al., 2017 [Publ 3799], fig. 2

cst iT tT bi riS oT roT roS iD
0.900 1 1 0 1 32 20 4 0.0
Cladomics cladogram ANODONTINI Tchangsinaia Nodularia Schistodesmus ACUTICOSTINI Inversiunio Lepidodesma LANCEOLARIINI

Wu et al., 2018 [Publ 3836], fig. 2

cst iT tT bi riS oT roT roS iD
0.931 1 1 0 1 52 32 4 0.0
Cladomics cladogram Lepidodesma UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Huang et al., 2019 [Publ 3843], fig. 1

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 45 31 4 0.0
Cladomics cladogram Anemina ACUTICOSTINI Sinanodonta ACULAMPROTULINA Cristaria LANCEOLARIINI Lepidodesma OXYNAIINA

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
0.912 1 1 0 1 200 47 4 0.0
Cladomics cladogram ACUTICOSTINI Lepidodesma UNIONINI LANCEOLARIINI ANODONTINI

Huang et al., 2019 [Publ 3843], fig. 3

cst iT tT bi riS oT roT roS iD
0.926 1 1 0 1 55 25 4 0.0
Cladomics cladogram Lepidodesma UNIONINI LANCEOLARIINI ACUTICOSTINI ANODONTINI

Huang et al., 2019 [Publ 3843], fig. S1

cst iT tT bi riS oT roT roS iD
0.900 1 1 0 1 46 32 4 0.0
Cladomics cladogram Anemina Sinanodonta ACUTICOSTINI Lepidodesma Cristaria LANCEOLARIINI UNIONINI

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
0.912 1 1 0 1 198 46 4 0.0
Cladomics cladogram ACUTICOSTINI UNIONINI Lepidodesma LANCEOLARIINI ANODONTINI

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.765 1 1 0 1 200 47 4 0.0
Cladomics cladogram ACUTICOSTINI Alasmidonta Anodonta anatina Anemina Anodonta cygnea Cristaria Brachyanodon Lasmigona Lepidodesma Pseudanodonta Pyganodon Simpsonaias Strophitus Utterbackia Schistodesmus UNIONINA ACULAMPROTULINA LANCEOLARIINI Pletholophus Sinanodonta Cuneopsis celtiformis Tchangsinaia Cuneopsis rufescens Nodularia Cuneopsis szechenyii Cuneopsis heudei

Wu et al., 2019 [Publ 3864], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 45 25 4 0.0
Cladomics cladogram Lepidodesma UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
0.949 1 1 0 1 121 70 4 0.0
Cladomics cladogram ACUTICOSTINI Lepidodesma UNIONINI LANCEOLARIINI ANODONTINI

Fukata & Iigo, 2020 [Publ 4106], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 53 27 4 0.0
Cladomics cladogram Lepidodesma LANCEOLARIINI UNIONINI ACUTICOSTINI ANODONTINI

Wu et al., 2020 [Publ 4121], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 25 4 0.0
Cladomics cladogram Lepidodesma UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Sano et al., 2020 [Publ 4138], fig. 1

cst iT tT bi riS oT roT roS iD
0.565 1 1 0 1 33 21 4 0.0
Cladomics cladogram Lanceolaria oxyrhyncha 1 Lanceolaria oxyrhyncha 2 Cristaria plicata 1 Cristaria plicata 2 Sinanodonta lauta Beringiana japonica Beringiana fukuharai 1 Beringiana beringiana Beringiana fukuharai 2 Beringiana fukuharai 3 Buldowskia iwakawai 1 Buldowskia iwakawai 2 Sinanodonta tumens 1 Sinanodonta tumens 2 Pletholophus Anemina Lanceolaria lanceolata Cristaria plicata 3 Lanceolaria gladiola Sinanodonta angula Lepidodesma ACULAMPROTULINA ACUTICOSTINI Cuneopsis Nodularia Schistodesmus Inversiunio

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
0.931 1 1 0 1 90 25 4 0.0
Cladomics cladogram ACUTICOSTINI Lepidodesma UNIONINI LANCEOLARIINI ANODONTINI

Wu et al., 2021 [Publ 4366], fig. 3-BI

cst iT tT bi riS oT roT roS iD
0.935 1 1 0 1 72 32 4 0.0
Cladomics cladogram Lepidodesma UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Wu et al., 2021 [Publ 4366], fig. 3-ML

cst iT tT bi riS oT roT roS iD
0.935 1 1 0 1 72 32 4 0.0
Cladomics cladogram Lepidodesma UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Wu et al., 2021 [Publ 4366], fig. 4

cst iT tT bi riS oT roT roS iD
0.935 1 1 0 1 67 31 4 0.0
Cladomics cladogram Lepidodesma UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
0.897 1 1 0 1 90 25 4 0.0
Cladomics cladogram ACUTICOSTINI UNIONINI LANCEOLARIINI Lepidodesma ANODONTINI

Wu et al., 2022 [Publ 4238], fig. 2d

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 37 37 4 0.0
Cladomics cladogram Lepidodesma UNIONINI LANCEOLARIINI ACUTICOSTINI ANODONTINI

Wu et al., 2022 [Publ 4238], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 72 34 4 0.0
Cladomics cladogram Lepidodesma UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 36 15 4 0.0
Cladomics cladogram Lepidodesma UNIONINI LANCEOLARIINI ACUTICOSTINI ANODONTINI

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

cst iT tT bi riS oT roT roS iD
0.960 1 1 0 1 38 16 4 0.0
Cladomics cladogram ANODONTINI ACUTICOSTINI LANCEOLARIINI Lepidodesma ACULAMPROTULINA UNIONINA OXYNAIINA

Konopleva et al., 2023 [Publ 4269], fig. S1_4

cst iT tT bi riS oT roT roS iD
0.966 1 1 0 1 55 32 4 0.0
Cladomics cladogram UNIONINI Lepidodesma LANCEOLARIINI ACUTICOSTINI ANODONTINI

Dai et al., 2023 [Publ 4285], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 16 4 0.0
Cladomics cladogram Lepidodesma UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Chen et al., 2023 [Publ 4302], fig. 1

cst iT tT bi riS oT roT roS iD
0.861 1 1 0 1 40 31 4 0.0
Cladomics cladogram Anemina Amuranodonta Pletholophus Sinanodonta Beringiana Lanceolaria gladiola ACUTICOSTINI Lepidodesma Lanceolaria oxyrhyncha Cristaria UNIONINI ALASMIDONTINA ANODONTINA

Chen et al., 2023 [Publ 4302], fig. 2

cst iT tT bi riS oT roT roS iD
0.972 1 1 0 1 40 31 4 0.0
Cladomics cladogram ACUTICOSTINI Lepidodesma UNIONINI LANCEOLARIINI ANODONTINI

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 93 44 4 0.0
Cladomics cladogram Lepidodesma UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cst iT tT bi riS oT roT roS iD
0.914 2 2 0 1 93 44 4 0.0
Cladomics cladogram Lepidodesma UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cst iT tT bi riS oT roT roS iD
0.971 2 2 0 1 89 44 4 0.0
Cladomics cladogram Lepidodesma UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cst iT tT bi riS oT roT roS iD
0.971 2 2 0 1 89 44 4 0.0
Cladomics cladogram Lepidodesma UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Dai et al., 2024 [Publ 4358], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 27 4 0.0
Cladomics cladogram Lepidodesma UNIONINI LANCEOLARIINI ACUTICOSTINI ANODONTINI

Dai et al., 2024 [Publ 4386], fig. 5-BI

cst iT tT bi riS oT roT roS iD
0.917 1 1 0 1 21 16 4 0.0
Cladomics cladogram ACUTICOSTINI CRISTARIINA ANODONTINA LANCEOLARIINI ACULAMPROTULINA Lepidodesma UNIONINA MIDDENDORFFINAIINA OXYNAIINA

Dai et al., 2024 [Publ 4386], fig. 5-ML

cst iT tT bi riS oT roT roS iD
0.958 1 1 0 1 21 16 4 0.0
Cladomics cladogram ANODONTINA ACUTICOSTINI CRISTARIINA Lepidodesma LANCEOLARIINI UNIONINI

Dai et al., 2024 [Publ 4386], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 26 4 0.0
Cladomics cladogram Lepidodesma UNIONINI LANCEOLARIINI ACUTICOSTINI ANODONTINI

Dai et al., 2025 [Publ 4440], fig. 1

cst iT tT bi riS oT roT roS iD
0.895 1 1 0 1 43 34 4 0.0
Cladomics cladogram Pletholophus Sinanodonta Beringiana Cristaria ALASMIDONTINA ANODONTINA LANCEOLARIINI Lepidodesma ACUTICOSTINI UNIONINI

Dai et al., 2025 [Publ 4440], fig. 2

cst iT tT bi riS oT roT roS iD
0.868 1 1 0 1 43 34 4 0.0
Cladomics cladogram Anemina Amuranodonta Pletholophus Sinanodonta Beringiana ALASMIDONTINA ANODONTINA Lanceolaria gladiola Lanceolaria oxyrhyncha Cristaria ACUTICOSTINI Lepidodesma UNIONINI

Dai et al., 2025 [Publ 4440], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 86 41 4 0.0
Cladomics cladogram Lepidodesma UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Chen et al., 2025 [Publ 4448], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 84 42 4 0.0
Cladomics cladogram Lepidodesma UNIONINI LANCEOLARIINI ACUTICOSTINI ANODONTINI

Chen et al., 2025 [Publ 4487], fig. 2

cst iT tT bi riS oT roT roS iD
0.973 1 1 0 1 40 31 4 0.0
Cladomics cladogram GLOBUNIONINA ACULAMPROTULINA UNIONINA OXYNAIINA ACUTICOSTINI Lepidodesma LANCEOLARIINI ANODONTINI

Chen et al., 2025 [Publ 4487], fig. 3

cst iT tT bi riS oT roT roS iD
0.946 1 1 0 1 40 31 4 0.0
Cladomics cladogram UNIONINI ACUTICOSTINI ALASMIDONTINA ANODONTINA Lepidodesma Cristaria LANCEOLARIINI Anemina Amuranodonta Pletholophus Sinanodonta Beringiana

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cst iT tT bi riS oT roT roS iD
0.947 2 2 0 1 138 66 4 0.0
Cladomics cladogram Lepidodesma UNIONINI LANCEOLARIINI ACUTICOSTINI ANODONTINI

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.818 2 2 0 1 631 154 4 0.0
Cladomics cladogram Amuranodonta Anemina Buldowskia Beringiana Pletholophus Mimo Acudonta Sinanodonta Cristaria Lepidodesma LANCEOLARIINI Pseudoacuticosta retiaria 2 Acuticosta chinensis 2 Acuticosta chinensis 1 Pseudoacuticosta retiaria 1 Globunio ACULAMPROTULINA MIDDENDORFFINAIINA UNIONINA OXYNAIINA Pseudodontoideus Lasmigona Pterosyna Arcidens ANODONTINA Utterbackia Pyganodon Utterbackiana Simpsonaias Alasminota Alasmidonta Prolasmidonta Platynaias Anodontoides Strophitus

Wu et al., 2026 [Publ 4811], fig. 4

cst iT tT bi riS oT roT roS iD
0.977 2 2 0 1 93 63 4 0.0
Cladomics cladogram Lepidodesma UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 22 3 0.0
Cladomics cladogram ANODONTINI LANCEOLARIINI Lepidodesma UNIONINI

Cha et al., 2018 [Publ 3916], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 37 20 3 0.0
Cladomics cladogram Lepidodesma UNIONINI LANCEOLARIINI ANODONTINI

Froufe et al., 2019 [Publ 3936], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 43 21 3 0.0
Cladomics cladogram Lepidodesma UNIONINI LANCEOLARIINI ANODONTINI

Froufe et al., 2019 [Publ 3936], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 43 21 3 0.0
Cladomics cladogram Lepidodesma UNIONINI LANCEOLARIINI ANODONTINI

Sano et al., 2020 [Publ 4138], fig. 2

cst iT tT bi riS oT roT roS iD
0.935 1 1 0 1 68 35 3 0.0
Cladomics cladogram Anemina Buldowskia Pletholophus Beringiana Sinanodonta Cristaria ANODONTINA ALASMIDONTINA LANCEOLARIINI Lepidodesma UNIONINI

Sano et al., 2020 [Publ 4138], fig. 3

cst iT tT bi riS oT roT roS iD
0.935 1 1 0 1 68 35 3 0.0
Cladomics cladogram Anemina Buldowskia Sinanodonta Pletholophus Beringiana Cristaria ANODONTINA ALASMIDONTINA LANCEOLARIINI Lepidodesma UNIONINI

Sano et al., 2020 [Publ 4138], fig. 4

cst iT tT bi riS oT roT roS iD
0.935 1 1 0 1 68 35 3 0.0
Cladomics cladogram Anemina Buldowskia Sinanodonta Pletholophus Beringiana Cristaria ANODONTINA ALASMIDONTINA Lepidodesma LANCEOLARIINI UNIONINI

Kim et al., 2021 [Publ 4197], fig. 1

cst iT tT bi riS oT roT roS iD
0.875 1 1 0 1 17 16 3 0.0
Cladomics cladogram CHAMBERLAINIINI Lepidodesma

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 21 3 0.0
Cladomics cladogram Lepidodesma UNIONINI LANCEOLARIINI ANODONTINI

Wu et al., 2024 [Publ 4379], fig. 5

cst iT tT bi riS oT roT roS iD
0.970 1 1 0 1 52 50 3 0.0
Cladomics cladogram Lepidodesma UNIONINI LANCEOLARIINI ANODONTINI

Wu et al., 2024 [Publ 4388], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 17 3 0.0
Cladomics cladogram Lepidodesma UNIONINI LANCEOLARIINI ANODONTINI

Hou et al., 2024 [Publ 4400], fig. 4

cst iT tT bi riS oT roT roS iD
0.971 2 2 0 1 73 69 3 0.0
Cladomics cladogram Lepidodesma UNIONINI LANCEOLARIINI ANODONTINI

Wang et al., 2025 [Publ 4488], fig. 3B

cst iT tT bi riS oT roT roS iD
0.974 2 2 0 1 62 60 3 0.0
Cladomics cladogram Lepidodesma UNIONINI ACUTICOSTINI ANODONTINI

Wu et al., 2026 [Publ 4825], fig. 6B

cst iT tT bi riS oT roT roS iD
0.974 2 2 0 1 63 61 3 0.0
Cladomics cladogram Lepidodesma UNIONINI LANCEOLARIINI ANODONTINI

Zhou et al., 2016 [Publ 3710], fig. 1

cst iT tT bi riS oT roT roS iD
0.938 1 1 0 1 19 8 2 0.0
Cladomics cladogram ANODONTINI Lepidodesma UNIONINI

Wen et al., 2017 [Publ 3795], fig. 3

cst iT tT bi riS oT roT roS iD
0.929 1 1 0 1 15 6 2 0.0
Cladomics cladogram ANODONTINI ACULAMPROTULINA Lepidodesma UNIONINA

Chen et al., 2020 [Publ 4170], fig. 1

cst iT tT bi riS oT roT roS iD
0.714 1 1 0 1 33 14 2 0.0
Cladomics cladogram ANODONTINI OXYNAIINA Lepidodesma ACULAMPROTULINA

Dai et al., 2024 [Publ 4386], fig. 3

cst iT tT bi riS oT roT roS iD
0.941 1 1 0 1 13 12 2 0.0
Cladomics cladogram UNIONINA MIDDENDORFFINAIINA OXYNAIINA ACULAMPROTULINA ANODONTINI Lepidodesma

Wu et al., 2023 [Publ 4339], fig. 5

cst iT tT bi riS oT roT roS iD
0.923 2 2 0 1 17 9 1 0.0
Cladomics cladogram Lepidodesma ANODONTINI

Hou et al., 2025 [Publ 4401], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 26 26 1 0.0
Cladomics cladogram Lepidodesma UNIONINI

Hou et al., 2025 [Publ 4432], fig. 3

cst iT tT bi riS oT roT roS iD
0.900 1 1 0 1 10 10 1 0.0
Cladomics cladogram Lepidodesma UNIONINI

Hou et al., 2025 [Publ 4432], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 26 26 1 0.0
Cladomics cladogram Lepidodesma UNIONINI