Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Unionini

subtribe OXYNAIINA Starobogatov, 1970: cladomics

12 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 51 (57 total)
number of subtaxa (rank: genus) = 12
analyzed in 151 cladograms in 89 publications
maximum number of terminals in an analysis = 32 (mean = 7.12, median = 6)
maximum number of subtaxa (genus) = 7 (mean = 3.29, median = 3)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 128
maximum number of included terminals = 32, (mean = 8.22 median = 7)
maximum number of subtaxa (genus) = 7 (mean = 3.7, median = 4)
proportion of cladograms that support basic monophyly = 0.84

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 101
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 32 (mean = 9.15, median = 7)
maximum number of subtaxa (genus) = 7 (mean = 4.19, median = 4)
proportion of cladograms that support rigorous monophyly = 0.84

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Oxynaiina cladogram distances
PCoA Ordination

NJ Tree of Oxynaiina cladogram distances
NJ Tree

Cladograms

Dai et al., 2025 [Publ 4440], fig. 1

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cstiTtTbiriSoTroTroSiD
0.941 13 15 11 7 31 8 4 **
GLOBUNIONINA ACULAMPROTULINA MIDDENDORFFINAIINA Nodularia Cuneopsis Diaurora Schistodesmus Pseudocuneopsis UNIONINA Tchangsinaia Inversiunio

Dai et al., 2025 [Publ 4440], fig. 2

cstiTtTbiriSoTroTroSiD
0.941 13 16 13 7 31 8 4 4.0
GLOBUNIONINA ACULAMPROTULINA MIDDENDORFFINAIINA Nodularia Cuneopsis Diaurora Schistodesmus Pseudocuneopsis Inversiunio Tchangsinaia UNIONINA

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cstiTtTbiriSoTroTroSiD
0.833 32 32 33 7 601 34 4 35.0
GLOBUNIONINA ACULAMPROTULINA MIDDENDORFFINAIINA UNIONINA Schistodesmus Nodularia Tchangsinaia Inversiunio Pseudocuneopsis Diaurora Cuneopsis

Chen et al., 2023 [Publ 4302], fig. 1

cstiTtTbiriSoTroTroSiD
0.933 14 17 14 7 27 4 3 5.0
ACULAMPROTULINA MIDDENDORFFINAIINA Nodularia Cuneopsis Diaurora Schistodesmus Pseudocuneopsis Inversiunio Tchangsinaia UNIONINA

Chen et al., 2023 [Publ 4302], fig. 2

cstiTtTbiriSoTroTroSiD
0.933 14 17 14 7 27 4 3 5.0
ACULAMPROTULINA MIDDENDORFFINAIINA Nodularia Cuneopsis Diaurora Schistodesmus Pseudocuneopsis Inversiunio Tchangsinaia UNIONINA

Chen et al., 2025 [Publ 4487], fig. 2

cstiTtTbiriSoTroTroSiD
0.938 13 13 11 7 28 5 3 8.0
ACULAMPROTULINA UNIONINA Tchangsinaia Inversiunio Cuneopsis Pseudocuneopsis Nodularia Diaurora Schistodesmus

Chen et al., 2025 [Publ 4487], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 13 13 11 7 28 5 3 8.0
GLOBUNIONINA ACULAMPROTULINA UNIONINA Tchangsinaia Inversiunio Pseudocuneopsis Cuneopsis Nodularia Diaurora Schistodesmus

Wu et al., 2022 [Publ 4238], fig. 2d

cstiTtTbiriSoTroTroSiD
1.000 15 15 13 6 23 8 2 15.0
ACULAMPROTULINA UNIONINA Pseudocuneopsis Nodularia Schistodesmus Cuneopsis Tchangsinaia Inversiunio

Wu et al., 2023 [Publ 4296], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 10 10 8 6 13 3 2 14.0
UNIONINA ACULAMPROTULINA Cuneopsis Nodularia Schistodesmus Tchangsinaia Inversiunio Pseudocuneopsis

Liu et al., 2023 [Publ 4310], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 11 11 9 6 13 3 2 15.0
UNIONINA ACULAMPROTULINA Cuneopsis Nodularia Schistodesmus Tchangsinaia Inversiunio Pseudocuneopsis

Liu et al., 2023 [Publ 4310], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 11 11 7 6 13 3 2 13.0
UNIONINA ACULAMPROTULINA Cuneopsis Nodularia Schistodesmus Tchangsinaia Inversiunio Pseudocuneopsis

Chen et al., 2025 [Publ 4448], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 19 19 17 6 66 8 2 23.0
ACULAMPROTULINA UNIONINA Pseudocuneopsis Nodularia Tongchuania Schistodesmus Tchangsinaia Cuneopsis

Wu et al., 2022 [Publ 4238], fig. 2a_S1

cstiTtTbiriSoTroTroSiD
1.000 11 11 9 6 1 1 1 14.5
ACULAMPROTULINA Nodularia Pseudocuneopsis Schistodesmus Cuneopsis Tchangsinaia Inversiunio

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cstiTtTbiriSoTroTroSiD
1.000 21 21 19 5 119 25 4 23.0
ACULAMPROTULINA GLOBUNIONINA MIDDENDORFFINAIINA UNIONINA Tchangsinaia Cuneopsis Pseudocuneopsis Schistodesmus Nodularia

Wu et al., 2026 [Publ 4811], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 20 20 18 5 75 12 4 23.0
GLOBUNIONINA ACULAMPROTULINA UNIONINA MIDDENDORFFINAIINA Pseudocuneopsis Nodularia Schistodesmus Tchangsinaia Cuneopsis

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cstiTtTbiriSoTroTroSiD
1.000 18 18 15 5 104 10 3 14.0
ACULAMPROTULINA MIDDENDORFFINAIINA UNIONINA Nodularia Schistodesmus Cuneopsis Tchangsinaia Inversiunio

Hou et al., 2024 [Publ 4400], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 24 24 22 5 51 24 3 26.0
ACULAMPROTULINA UNIONINA MIDDENDORFFINAIINA Pseudocuneopsis Nodularia Schistodesmus Tchangsinaia Cuneopsis

Dai et al., 2025 [Publ 4440], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 16 16 14 5 71 11 3 19.0
GLOBUNIONINA ACULAMPROTULINA UNIONINA Nodularia Pseudocuneopsis Schistodesmus Tchangsinaia Cuneopsis

Wang et al., 2025 [Publ 4488], fig. 3B

cstiTtTbiriSoTroTroSiD
1.000 23 23 21 5 41 9 3 26.0
ACULAMPROTULINA UNIONINA MIDDENDORFFINAIINA Pseudocuneopsis Nodularia Schistodesmus Tchangsinaia Cuneopsis

Wu et al., 2026 [Publ 4825], fig. 6B

cstiTtTbiriSoTroTroSiD
1.000 20 20 18 5 45 9 3 23.0
ACULAMPROTULINA UNIONINA MIDDENDORFFINAIINA Pseudocuneopsis Nodularia Schistodesmus Tchangsinaia Cuneopsis

Wu et al., 2019 [Publ 3864], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 5 5 3 5 41 8 2 16.0
ACULAMPROTULINA UNIONINA Pseudocuneopsis Nodularia Tchangsinaia Cuneopsis Schistodesmus

Fukata & Iigo, 2019 [Publ 4108], fig. 1

cstiTtTbiriSoTroTroSiD
0.909 8 8 6 5 9 5 2 19.0
UNIONINA Pseudocuneopsis Cuneopsis Schistodesmus Tchangsinaia Nodularia

Fukata & Iigo, 2020 [Publ 4106], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 6 6 4 5 48 7 2 17.0
ACULAMPROTULINA UNIONINA Cuneopsis Schistodesmus Pseudocuneopsis Nodularia Tchangsinaia

Wu et al., 2020 [Publ 4121], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 5 5 3 5 43 8 2 16.0
ACULAMPROTULINA UNIONINA Pseudocuneopsis Nodularia Tchangsinaia Cuneopsis Schistodesmus

Wu et al., 2020 [Publ 4147], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 6 6 4 5 24 5 2 17.0
ACULAMPROTULINA UNIONINA Pseudocuneopsis Nodularia Schistodesmus Cuneopsis Tchangsinaia

Wu et al., 2021 [Publ 4366], fig. 3-BI

cstiTtTbiriSoTroTroSiD
1.000 7 7 5 5 66 8 2 16.0
ACULAMPROTULINA UNIONINA Pseudocuneopsis Nodularia Schistodesmus Cuneopsis Tchangsinaia

Wu et al., 2021 [Publ 4366], fig. 3-ML

cstiTtTbiriSoTroTroSiD
1.000 7 7 5 5 66 8 2 16.0
ACULAMPROTULINA UNIONINA Pseudocuneopsis Nodularia Tchangsinaia Cuneopsis Schistodesmus

Wu et al., 2021 [Publ 4366], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 7 7 5 5 61 8 2 16.0
ACULAMPROTULINA UNIONINA Pseudocuneopsis Nodularia Schistodesmus Tchangsinaia Cuneopsis

Wu et al., 2022 [Publ 4238], fig. 2e

cstiTtTbiriSoTroTroSiD
1.000 9 9 7 5 9 4 2 15.0
ACULAMPROTULINA UNIONINA Pseudocuneopsis Nodularia Tchangsinaia Cuneopsis Schistodesmus

Wu et al., 2022 [Publ 4238], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 12 12 10 5 61 8 2 16.0
ACULAMPROTULINA UNIONINA Pseudocuneopsis Nodularia Schistodesmus Tchangsinaia Cuneopsis

Dai et al., 2023 [Publ 4285], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 5 5 3 5 48 2 2 15.0
ACULAMPROTULINA UNIONINA Pseudocuneopsis Nodularia Schistodesmus Cuneopsis Tchangsinaia

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cstiTtTbiriSoTroTroSiD
1.000 12 12 10 5 83 10 2 16.0
ACULAMPROTULINA UNIONINA Nodularia Pseudocuneopsis Tchangsinaia Schistodesmus Cuneopsis

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cstiTtTbiriSoTroTroSiD
0.909 12 12 10 5 83 10 2 16.0
ACULAMPROTULINA UNIONINA Pseudocuneopsis Nodularia Schistodesmus lampreyanus Cuneopsis Tchangsinaia Schistodesmus xinyuensis Schistodesmus spinosus

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cstiTtTbiriSoTroTroSiD
1.000 12 12 9 5 79 10 2 15.0
ACULAMPROTULINA UNIONINA Nodularia Pseudocuneopsis Schistodesmus Tchangsinaia Cuneopsis

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cstiTtTbiriSoTroTroSiD
1.000 12 12 9 5 79 10 2 15.0
ACULAMPROTULINA UNIONINA Nodularia Pseudocuneopsis Schistodesmus Tchangsinaia Cuneopsis

Dai et al., 2024 [Publ 4358], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 14 14 12 5 39 4 2 18.0
ACULAMPROTULINA UNIONINA Pseudocuneopsis Nodularia Schistodesmus Tchangsinaia Cuneopsis

Wu et al., 2024 [Publ 4379], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 19 19 17 5 34 10 2 23.0
ACULAMPROTULINA UNIONINA Pseudocuneopsis Nodularia Schistodesmus Tchangsinaia Cuneopsis

Dai et al., 2024 [Publ 4386], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 11 11 9 5 30 10 2 19.0
ACULAMPROTULINA UNIONINA Pseudocuneopsis Nodularia Schistodesmus Tchangsinaia Cuneopsis

Wu et al., 2024 [Publ 4388], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 7 7 5 5 46 2 2 16.0
ACULAMPROTULINA UNIONINA Pseudocuneopsis Nodularia Tchangsinaia Cuneopsis Schistodesmus

Hou et al., 2025 [Publ 4445], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 22 22 20 5 30 10 2 25.0
ACULAMPROTULINA UNIONINA Pseudocuneopsis Nodularia Schistodesmus Tchangsinaia Cuneopsis

Klymus et al., 2025 [Publ 4814], fig. 3A

cstiTtTbiriSoTroTroSiD
1.000 5 5 3 5 31 5 2 16.0
ACULAMPROTULINA UNIONINA Pseudocuneopsis Schistodesmus Nodularia Tchangsinaia Cuneopsis

Klymus et al., 2025 [Publ 4814], fig. 3B

cstiTtTbiriSoTroTroSiD
0.909 5 5 3 5 31 5 2 16.0
UNIONINA ACULAMPROTULINA Lanceolaria Nodularia Pseudocuneopsis Schistodesmus Tchangsinaia Cuneopsis

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 9 9 7 5 38 5 2 15.0
ACULAMPROTULINA UNIONINA Pseudocuneopsis Nodularia Tchangsinaia Cuneopsis Schistodesmus

Hou et al., 2025 [Publ 4401], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 25 25 23 5 3 1 1 27.0
MIDDENDORFFINAIINA Pseudocuneopsis Nodularia Schistodesmus Tchangsinaia Cuneopsis

Hou et al., 2025 [Publ 4432], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 25 25 23 5 3 1 1 27.0
MIDDENDORFFINAIINA Pseudocuneopsis Nodularia Schistodesmus Tchangsinaia Cuneopsis

Zhang et al., 2004 [Publ 2735], fig. 1

cstiTtTbiriSoTroTroSiD
0.857 7 7 5 5 7 0 0 16.0
Acuticosta Lanceolaria Tchangsinaia Cuneopsis heudei Nodularia Pseudocuneopsis Cuneopsis rufescens Schistodesmus

Dai et al., 2024 [Publ 4386], fig. 5-BI

cstiTtTbiriSoTroTroSiD
0.917 9 9 6 4 13 3 3 17.0
UNIONINA MIDDENDORFFINAIINA Cuneopsis Nodularia Inversiunio Schistodesmus

Dai et al., 2024 [Publ 4386], fig. 5-ML

cstiTtTbiriSoTroTroSiD
1.000 9 9 6 4 13 3 3 16.0
ACULAMPROTULINA UNIONINA MIDDENDORFFINAIINA Nodularia Cuneopsis Inversiunio Schistodesmus

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cstiTtTbiriSoTroTroSiD
1.000 6 6 4 4 75 5 2 14.0
UNIONINA ACULAMPROTULINA Nodularia Schistodesmus Tchangsinaia Cuneopsis

Bolotov et al., 2017 [Publ 3761], fig. S1

cstiTtTbiriSoTroTroSiD
1.000 6 6 4 4 133 4 2 14.0
UNIONINA ACULAMPROTULINA Nodularia Cuneopsis Tchangsinaia Schistodesmus

Bolotov et al., 2017 [Publ 3761], fig. S2

cstiTtTbiriSoTroTroSiD
0.800 6 7 3 4 133 4 2 16.0
UNIONINA ACULAMPROTULINA Schistodesmus Nodularia Cuneopsis szechenyii Tchangsinaia Cuneopsis rufescens

Bolotov et al., 2017 [Publ 3764], fig. S1

cstiTtTbiriSoTroTroSiD
0.500 6 10 0 4 119 4 2 17.0
Unio crassus Unio pictorum Unio tumidus ACULAMPROTULINA Cuneopsis szechenyii Tchangsinaia Cuneopsis rufescens Nodularia douglasiae Nodularia nuxpersicae Schistodesmus Alasmidonta Anodonta anatina Anodonta cygnea Brachyanodon Lasmigona Pseudanodonta Pyganodon Simpsonaias Strophitus Sinanodonta lauta Cristaria plicata Cristaria radiata Pletholophus Sinanodonta lucida Sinanodonta jourdyi Lanceolaria gladiola Lanceolaria grayii Lanceolaria danzhouensis

Wu et al., 2018 [Publ 3836], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 7 7 5 4 46 7 2 13.0
ACULAMPROTULINA UNIONINA Nodularia Schistodesmus Tchangsinaia Cuneopsis

Aunins et al., 2018 [Publ 3863], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 4 4 2 4 36 7 2 14.0
ACULAMPROTULINA UNIONINA Nodularia Tchangsinaia Cuneopsis Schistodesmus

Huang et al., 2019 [Publ 3843], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 12 12 9 4 189 9 2 17.0
ACULAMPROTULINA UNIONINA Schistodesmus Nodularia Tchangsinaia Cuneopsis

Huang et al., 2019 [Publ 3843], fig. S3

cstiTtTbiriSoTroTroSiD
1.000 11 11 9 4 188 9 2 15.0
ACULAMPROTULINA UNIONINA Nodularia Schistodesmus Tchangsinaia Cuneopsis

Huang et al., 2019 [Publ 3843], fig. S4

cstiTtTbiriSoTroTroSiD
0.700 12 48 25 4 189 9 2 52.0
Chamberlainia Megalonaias Uniomerus QUADRULINA Sinohyriopsis INDOPSEUDODONTINA PSEUDODONTINA RECTIDENTINA CONTRADENTINA Lamprotula Potomida Pronodularia Sinosolenaia carinata Gonidea Leguminaia Microcondylaea Ptychorhynchus Parvasolenaia Koreosolenaia Sinosolenaia recognita Sinosolenaia iridinea Sinosolenaia oleivora Amblema Toxolasma Obliquaria Plectomerus Elliptoideus Elliptio Eurynia Fusconaia Pleurobema Ellipsaria Potamilus leptodon Medionidus Potamilus alatus Ptychobranchus Truncilla Venustaconcha Cambarunio Leaunio Ortmanniana Lampsilis Acuticosta Alasmidonta Anodonta anatina Anemina Anodonta cygnea Cristaria Brachyanodon Lasmigona Lepidodesma Pseudanodonta Pyganodon Simpsonaias Strophitus Utterbackia Schistodesmus UNIONINA ACULAMPROTULINA Lanceolaria Pletholophus Sinanodonta Cuneopsis celtiformis Tchangsinaia Cuneopsis rufescens Nodularia Cuneopsis szechenyii Cuneopsis heudei

Sano et al., 2020 [Publ 4138], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 7 7 7 4 62 6 2 18.0
ACULAMPROTULINA UNIONINA Inversiunio Nodularia Tchangsinaia Schistodesmus

Sano et al., 2020 [Publ 4138], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 7 7 8 4 62 6 2 18.0
ACULAMPROTULINA UNIONINA Inversiunio Nodularia Schistodesmus Tchangsinaia

Sano et al., 2020 [Publ 4138], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 7 7 8 4 62 6 2 16.0
ACULAMPROTULINA UNIONINA Nodularia Inversiunio Schistodesmus Tchangsinaia

Wu et al., 2021 [Publ 4366], fig. 2_I

cstiTtTbiriSoTroTroSiD
1.000 6 6 4 4 85 5 2 12.0
ACULAMPROTULINA UNIONINA Nodularia Cuneopsis Tchangsinaia Schistodesmus

Wu et al., 2021 [Publ 4366], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 6 6 4 4 85 5 2 12.0
ACULAMPROTULINA UNIONINA Nodularia Cuneopsis Tchangsinaia Schistodesmus

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

cstiTtTbiriSoTroTroSiD
0.900 6 6 4 4 70 4 2 14.0
UNIONINA ACULAMPROTULINA Nodularia Schistodesmus Cuneopsis szechenyii Tchangsinaia Cuneopsis rufescens

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

cstiTtTbiriSoTroTroSiD
0.900 6 6 4 4 70 4 2 14.0
UNIONINA ACULAMPROTULINA Nodularia Schistodesmus Cuneopsis szechenyii Tchangsinaia Cuneopsis rufescens

Ouyang et al., 2015 [Publ 4493], fig. 1

cstiTtTbiriSoTroTroSiD
0.750 7 8 6 4 24 1 1 16.0
Lanceolaria Acuticosta CRISTARIINA Schistodesmus ACULAMPROTULINA Tchangsinaia Cuneopsis rufescens Nodularia Cuneopsis szechenyii Cuneopsis celtiformis

Ouyang et al., 2015 [Publ 4493], fig. 2

cstiTtTbiriSoTroTroSiD
0.750 7 7 8 4 24 1 1 17.0
CRISTARIINA Nodularia Cuneopsis rufescens 2 Schistodesmus Cuneopsis celtiformis 2 Tchangsinaia Cuneopsis celtiformis 1 Cuneopsis szechenyii 2 Cuneopsis szechenyii 1 Cuneopsis rufescens 1

Huang et al., 2019 [Publ 3843], fig. 1

cstiTtTbiriSoTroTroSiD
0.750 12 12 10 4 34 5 1 17.0
Lepidodesma Schistodesmus Nodularia Cuneopsis rufescens Tchangsinaia Cuneopsis celtiformis Cuneopsis szechenyii Cuneopsis heudei

Huang et al., 2019 [Publ 3843], fig. S1

cstiTtTbiriSoTroTroSiD
1.000 12 12 10 4 35 5 1 15.0
ACULAMPROTULINA Nodularia Tchangsinaia Cuneopsis Schistodesmus

Sano et al., 2020 [Publ 4138], fig. 1

cstiTtTbiriSoTroTroSiD
0.750 6 22 5 4 28 1 1 39.0
Sinohyriopsis Lamprotula Pronodularia Sinosolenaia Inversidens Obovalis omiensis 1 Obovalis omiensis 2 Ptychorhynchus Lanceolaria oxyrhyncha 1 Lanceolaria oxyrhyncha 2 Cristaria plicata 1 Cristaria plicata 2 Sinanodonta lauta Beringiana japonica Beringiana fukuharai 1 Beringiana beringiana Beringiana fukuharai 2 Beringiana fukuharai 3 Buldowskia iwakawai 1 Buldowskia iwakawai 2 Sinanodonta tumens 1 Sinanodonta tumens 2 Pletholophus Anemina Lanceolaria lanceolata Cristaria plicata 3 Lanceolaria gladiola Sinanodonta angula Lepidodesma ACULAMPROTULINA Acuticosta Cuneopsis Nodularia Schistodesmus Inversiunio

Choi et al., 2020 [Publ 4169], fig. 2a

cstiTtTbiriSoTroTroSiD
0.875 11 12 11 4 1 1 1 17.0
Nodularia Schistodesmus Cuneopsis UNIONINA Tchangsinaia

Sano et al., 2017 [Publ 3799], fig. 2

cstiTtTbiriSoTroTroSiD
0.667 8 12 13 4 25 0 0 20.0
CRISTARIINA Tchangsinaia Nodularia Schistodesmus Acuticosta Inversiunio Lepidodesma Lanceolaria

Wu et al., 2024 [Publ 4379], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 13 13 10 4 0 0 0 19.0
Nodularia Cuneopsis Tchangsinaia Pseudocuneopsis

Konopleva et al., 2023 [Publ 4269], fig. S1_4

cstiTtTbiriSoTroTroSiD
1.000 4 4 2 3 52 8 3 13.0
MIDDENDORFFINAIINA ACULAMPROTULINA UNIONINA Nodularia Tchangsinaia Cuneopsis

Dai et al., 2024 [Publ 4386], fig. 3

cstiTtTbiriSoTroTroSiD
0.909 8 8 6 3 6 3 3 18.0
MIDDENDORFFINAIINA Inversiunio Cuneopsis Nodularia

Fonseca et al., 2016 [Publ 3775], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 28 4 2 15.0
ACULAMPROTULINA UNIONINA Schistodesmus Nodularia Tchangsinaia

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 29 4 2 15.0
ACULAMPROTULINA UNIONINA Schistodesmus Tchangsinaia Nodularia

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 20 7 2 15.0
ACULAMPROTULINA UNIONINA Schistodesmus Tchangsinaia Nodularia

Wang et al., 2018 [Publ 3846], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 28 5 2 13.0
ACULAMPROTULINA UNIONINA Nodularia Tchangsinaia Cuneopsis

Cha et al., 2018 [Publ 3916], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 35 7 2 15.5
ACULAMPROTULINA UNIONINA Schistodesmus Nodularia Tchangsinaia

Huang et al., 2019 [Publ 3843], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 53 8 2 13.0
ACULAMPROTULINA UNIONINA Nodularia Tchangsinaia Schistodesmus

Froufe et al., 2019 [Publ 3936], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 41 7 2 13.0
ACULAMPROTULINA UNIONINA Nodularia Tchangsinaia Schistodesmus

Froufe et al., 2019 [Publ 3936], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 41 7 2 13.0
ACULAMPROTULINA UNIONINA Nodularia Tchangsinaia Schistodesmus

Kim et al., 2021 [Publ 4197], fig. 1

cstiTtTbiriSoTroTroSiD
0.889 7 7 5 3 11 5 2 15.0
UNIONINA Nodularia Cuneopsis Schistodesmus

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 45 7 2 13.0
ACULAMPROTULINA UNIONINA Nodularia Schistodesmus Tchangsinaia

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 4 4 2 3 33 3 2 16.0
ACULAMPROTULINA UNIONINA Schistodesmus Nodularia Tchangsinaia

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

cstiTtTbiriSoTroTroSiD
0.889 4 4 2 3 35 3 2 16.0
UNIONINA Schistodesmus Nodularia Tchangsinaia

Zhou et al., 2007 [Publ 2658], fig. 1

cstiTtTbiriSoTroTroSiD
0.714 4 11 9 3 17 3 1 20.0
Sinohyriopsis Lamprotula leaii Lamprotula caveata Gibbosula Lamprotula cornuumlunae ACULAMPROTULINA Tchangsinaia Nodularia Schistodesmus Lanceolaria Acuticosta

Liu et al., 2017 [Publ 3939], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 7 7 5 3 8 4 1 16.0
UNIONINA Schistodesmus Cuneopsis Nodularia

Hou et al., 2024 [Publ 4400], fig. 1

cstiTtTbiriSoTroTroSiD
0.714 12 13 11 3 3 1 1 20.0
Lamprotula Cuneopsis rufescens MIDDENDORFFINAIINA Pseudocuneopsis Tchangsinaia Cuneopsis kiangsiensis Cuneopsis celtiformis Cuneopsis szechenyii Cuneopsis heudei Cuneopsis demangei

Hou et al., 2025 [Publ 4432], fig. 3

cstiTtTbiriSoTroTroSiD
0.857 9 10 8 3 2 1 1 19.0
Lepidodesma Tchangsinaia Pseudocuneopsis MIDDENDORFFINAIINA Cuneopsis

Huang et al., 2002 [Publ 2657], fig. 1

cstiTtTbiriSoTroTroSiD
0.600 3 7 5 3 31 0 0 18.0
CRISTARIINA ALASMIDONTINA Acuticosta Nodularia Tchangsinaia Lanceolaria Schistodesmus Lepidodesma

Huang et al., 2002 [Publ 2657], fig. 2

cstiTtTbiriSoTroTroSiD
0.600 3 14 5 3 31 0 0 25.0
Strophitus Schistodesmus Lepidodesma Acuticosta CRISTARIINA Lanceolaria Nodularia Tchangsinaia Utterbackia Utterbackiana Alasmidonta Lasmigona Ptychorhynchus Sinohyriopsis Lamprotula Sinosolenaia Megalonaias Elliptoideus Uniomerus Pleurobema Potamilus fragilis Medionidus Amblema Obliquaria Plectomerus Villosa Ellipsaria Lampsilis Potamilus purpuratus Ptychobranchus

Dai et al., 2024 [Publ 4358], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 6 6 4 3 0 0 0 15.0
Pseudocuneopsis Schistodesmus Nodularia

Xue et al., 2016 [Publ 3659], fig. 1

cstiTtTbiriSoTroTroSiD
0.875 2 3 0 2 13 2 2 15.0
ACULAMPROTULINA Schistodesmus Nodularia UNIONINA

Huang et al., 2018 [Publ 3828], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 40 7 2 --
ACULAMPROTULINA UNIONINA Tchangsinaia Nodularia

Wang et al., 2018 [Publ 3865], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 29 5 2 --
ACULAMPROTULINA UNIONINA Tchangsinaia Nodularia

Sayenko et al., 2021 [Publ 4189], fig. 6

cstiTtTbiriSoTroTroSiD
0.875 10 11 9 2 3 3 2 19.0
UNIONINA Inversiunio MIDDENDORFFINAIINA Nodularia

Zhou et al., 2007 [Publ 2658], fig. 2

cstiTtTbiriSoTroTroSiD
0.833 2 2 0 2 16 3 1 --
Acuticosta Nodularia Cuneopsis

Ouyang et al., 2011 [Publ 3408], fig. 1

cstiTtTbiriSoTroTroSiD
0.833 2 2 0 2 11 1 1 --
Nodularia Cuneopsis Lanceolaria grayii Acuticosta Lanceolaria gladiola ACULAMPROTULINA

Zhou et al., 2017 [Publ 3788], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 14 2 1 --
ACULAMPROTULINA Nodularia Tchangsinaia

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 55 1 1 --
UNIONINA Nodularia Tchangsinaia

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 55 1 1 --
UNIONINA Tchangsinaia Nodularia

Choi et al., 2020 [Publ 4169], fig. 3a

cstiTtTbiriSoTroTroSiD
0.833 5 6 5 2 1 1 1 15.0
Nodularia Cuneopsis UNIONINA

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 20 17 1 --
UNIONINA Tchangsinaia Nodularia

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 20 17 1 --
UNIONINA Tchangsinaia Nodularia

Wu et al., 2024 [Publ 4379], fig. 4_6

cstiTtTbiriSoTroTroSiD
0.833 12 12 10 2 11 3 1 19.0
CRISTARIINA Schistodesmus Nodularia

Wu et al., 2026 [Publ 4825], fig. 5

cstiTtTbiriSoTroTroSiD
0.833 5 5 3 2 14 2 1 14.0
Schistodesmus Nodularia ACULAMPROTULINA Lanceolaria

Sano et al., 2017 [Publ 3799], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 4 4 5 2 17 0 0 15.0
Lanceolaria Nodularia Inversiunio

Kano et al., 2019 [Publ 4194], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 8 0 0 --
Lanceolaria Inversiunio Nodularia

Lopes-Lima et al., 2020 [Publ 4080], fig. 12

cstiTtTbiriSoTroTroSiD
1.000 6 6 4 2 2 0 0 13.0
Margaritifera Nodularia Inversiunio

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

cstiTtTbiriSoTroTroSiD
0.500 2 3 0 2 10 0 0 14.0
CRISTARIINA Nodularia Lanceolaria Inversiunio

Wu et al., 2021 [Publ 4215], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 2 21 0 0 14.0
Acuticosta Nodularia Inversiunio

Mabuchi et al., 2021 [Publ 4233], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 5 5 3 2 0 0 0 16.0
Tchangsinaia Nodularia

Hou et al., 2025 [Publ 4445], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 7 7 5 2 0 0 0 15.0
Cuneopsis Schistodesmus

Chen et al., 2025 [Publ 4448], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 8 8 6 2 1 0 0 17.0
Sinosolenaia Nodularia Pseudocuneopsis

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 4 2 --
ACULAMPROTULINA UNIONINA Nodularia

Wu et al., 2016 [Publ 3711], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 20 4 2 --
ACULAMPROTULINA UNIONINA Nodularia

Araujo et al., 2017 [Publ 3801], fig. 2

cstiTtTbiriSoTroTroSiD
0.857 1 1 0 1 13 2 2 --
Schistodesmus ACULAMPROTULINA

Wu et al., 2018 [Publ 4143], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 21 5 2 --
ACULAMPROTULINA UNIONINA Nodularia

Huang et al., 2019 [Publ 3843], fig. 3-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 22 4 2 --
Schistodesmus ACULAMPROTULINA UNIONINA

Klishko et al., 2019 [Publ 3886], fig. 9

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 1 8 5 2 14.0
UNIONINA MIDDENDORFFINAIINA Nodularia

Guerra et al., 2019 [Publ 4168], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 40 4 2 --
ACULAMPROTULINA Schistodesmus UNIONINA

Guerra et al., 2019 [Publ 4168], fig. 1-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 21 4 2 --
Schistodesmus ACULAMPROTULINA UNIONINA

Guerra et al., 2019 [Publ 4168], fig. 2

cstiTtTbiriSoTroTroSiD
0.857 1 1 0 1 40 4 2 --
Schistodesmus UNIONINA

Guerra et al., 2019 [Publ 4168], fig. 2-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 21 4 2 --
ACULAMPROTULINA Schistodesmus UNIONINA

Bolotov et al., 2020 [Publ 4084], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 35 4 2 --
MIDDENDORFFINAIINA Nodularia UNIONINA

Araujo et al., 2009 [Publ 2589], fig. 7A

cstiTtTbiriSoTroTroSiD
0.400 1 1 0 1 10 6 1 --
Nodularia Unio gibbus Unio tumidus ANODONTINA Unio pictorum 1 Unio pictorum 2 Unio mancus Unio tumidiformis Unio crassus

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

cstiTtTbiriSoTroTroSiD
0.400 1 1 0 1 24 2 1 --
Nodularia Anodonta cygnea Unio pictorum 2 Unio pictorum 1 Unio caffer Anodonta ALASMIDONTINA

Zhou et al., 2016 [Publ 3710], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 19 1 1 --
Schistodesmus UNIONINA

Froufe et al., 2016 [Publ 3738], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 1 12 12 1 --
Nodularia UNIONINA

Klishko et al., 2017 [Publ 3946], fig. 8

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 1 4 4 1 --
Nodularia UNIONINA

Araujo et al., 2018 [Publ 3779], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 17 13 1 --
Nodularia UNIONINA

Bolotov et al., 2018 [Publ 3826], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 2 2 1 1 80 3 1 --
UNIONINA Nodularia

Klishko et al., 2018 [Publ 3945], fig. 8

cstiTtTbiriSoTroTroSiD
1.000 4 4 2 1 4 4 1 15.0
Nodularia UNIONINA

Lopes-Lima et al., 2020 [Publ 4080], fig. 13

cstiTtTbiriSoTroTroSiD
1.000 7 7 5 1 3 1 1 18.0
MIDDENDORFFINAIINA Nodularia

Lopes-Lima et al., 2020 [Publ 4080], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 1 7 5 1 --
Nodularia ACULAMPROTULINA

Chen et al., 2020 [Publ 4170], fig. 1

cstiTtTbiriSoTroTroSiD
0.800 1 1 0 1 33 2 1 --
Schistodesmus Lepidodesma ACULAMPROTULINA

Altaba et al., 2025 [Publ 4849], fig. 10

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 11 3 1 --
Nodularia UNIONINA

Altaba et al., 2025 [Publ 4849], fig. 11

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 13 3 1 --
Nodularia UNIONINA

Altaba et al., 2025 [Publ 4849], fig. 12

cstiTtTbiriSoTroTroSiD
0.600 1 1 0 1 12 3 1 --
Unio mancus Unio pictorum Unio tumidus Nodularia

Feng et al., 2016 [Publ 3800], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 13 0 0 --
Tchangsinaia ALASMIDONTINA CRISTARIINA

Lopes-Lima et al., 2020 [Publ 4080], fig. 11

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 1 10 0 0 --
Nodularia Lanceolaria

Choi et al., 2020 [Publ 4169], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 7 7 5 1 3 0 0 18.0
Nodularia Radiatula Lamellidens Leoparreysia

Bogan et al., 2023 [Publ 4295], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 0 0 --
Schistodesmus ANODONTINA CRISTARIINA

Gomes-dos-Santos et al., 2023 [Publ 4463], fig. 2a

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 20 0 0 --
ANODONTINA Nodularia

Gomes-dos-Santos et al., 2023 [Publ 4463], fig. 2b

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 21 0 0 --
ANODONTINA Nodularia

Wu et al., 2024 [Publ 4388], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 5 5 3 1 4 0 0 16.0
Nodularia Cosmopseudodon Ptychorhynchus Postolata Obovalis

Hou et al., 2025 [Publ 4401], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 11 11 9 1 0 0 0 22.0
Nodularia

Wu et al., 2025 [Publ 4442], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 1 33 0 0 --
Cuneopsis CRISTARIINA

Chen et al., 2026 [Publ 4805], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 33 0 0 --
Cuneopsis CRISTARIINA

Wu et al., 2026 [Publ 4825], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 1 10 0 0 --
Schistodesmus Lanceolaria