Bivalvia | Unionida | Unionoidea | Unionidae

subfamily PARREYSIINAE Henderson, 1935: cladomics

5 tribes

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 108 (114 total)
number of subtaxa (rank: tribe) = 5
analyzed in 97 cladograms in 60 publications
maximum number of terminals in an analysis = 84 (mean = 12.6, median = 3)
maximum number of subtaxa (tribe) = 5 (mean = 2.53, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 63
maximum number of included terminals = 84, (mean = 18.86 median = 10)
maximum number of subtaxa (tribe) = 5 (mean = 3.35, median = 3)
proportion of cladograms that support basic monophyly = 0.95

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 46
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (tribe) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 84 (mean = 19.83, median = 9.5)
maximum number of subtaxa (tribe) = 5 (mean = 3.76, median = 4)
proportion of cladograms that support rigorous monophyly = 0.96

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Parreysiinae cladogram distances
PCoA Ordination

NJ Tree of Parreysiinae cladogram distances
NJ Tree

Cladograms

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
84 86 91 5 547 483 3 0 
        +-------------------------------- Lamellidens **                 
        |                                                             
  +-----+                             +-- Balwantia **                   
  |     |  +--------------------------+                               
  |     |  |                          +-- Trapezidens **                 
  |     +--+                                                          
  |        |  +-------------------------- Leoparreysiini **              
  |        |  |                                                       
  |        +--+  +----------------------- Pseudomulleria              
  |           |  |                                                    
  |           |  |                    +-- Parreysiini **                 
  |           +--+  +-----------------+                               
  |              |  |                 +-- Coelaturini **                 
  |              |  |                                                 
  |              +--+                 +-- Pseudoscabies **               
  |                 |  +--------------+                               
  |                 |  |              +-- Indonaia **                    
  |                 +--+                                              
  |                    |  +-------------- Indochinella **                
  |                    |  |                                           
  |                    +--+  +----------- Radiatula **                   
--+                       |  |                                        
  |                       +--+        +-- Gonideinae                  
  |                          |  +-----+                               
  |                          |  |     +-- Unionetta_fabagina__2 **       
  |                          +--+                                     
  |                             |  +----- Harmandia **                   
  |                             +--+                                  
  |                                |  +-- Unionetta_fabagina__1 **       
  |                                +--+                               
  |                                   +-- Scabies **                     
  |                                                                   
  |                                +----- Amuranodonta                
  |  +-----------------------------+                                  
  |  |                             |  +-- Anemina                     
  |  |                             +--+                               
  |  |                                +-- Buldowskia                  
  |  |                                                                
  |  |                                +-- Beringiana                  
  |  |                          +-----+                               
  |  |                          |     +-- Pletholophus                
  +--+  +-----------------------+                                     
     |  |                       |  +----- Mimo                        
     |  |                       +--+                                  
     |  |                          |  +-- Acudonta                    
     |  |                          +--+                               
     |  |                             +-- Sinanodonta                 
     |  |                                                             
     |  |  +----------------------------- Cristaria                   
     +--+  |                                                          
        |  |           +----------------- Lepidodesmini               
        |  |           |                                              
        |  |           |     +----------- Lanceolariini               
        |  |           |  +--+                                        
        |  |  +--------+  |  |  +-------- Pseudoacuticosta_retiaria__2
        |  |  |        |  |  +--+                                     
        |  |  |        |  |     |  +----- Acuticosta_chinensis__2     
        +--+  |        |  |     +--+                                  
           |  |        +--+        |  +-- Acuticosta_chinensis__1     
           |  |           |        +--+                               
           |  |           |           +-- Pseudoacuticosta_retiaria__1
           |  |           |                                           
           |  |           |  +----------- Globunio                    
           |  |           +--+                                        
           |  |              |  +-------- Aculamprotulina             
           |  |              +--+                                     
           +--+                 |  +----- Middendorffinaiina          
              |                 +--+                                  
              |                    |  +-- Unionina                    
              |                    +--+                               
              |                       +-- Oxynaiina                   
              |                                                       
              |                 +-------- Pseudodontoideus            
              |  +--------------+                                     
              |  |              |  +----- Lasmigona                   
              |  |              +--+                                  
              |  |                 |  +-- Pterosyna                   
              |  |                 +--+                               
              +--+                    +-- Arcidens                    
                 |                                                    
                 |  +-------------------- Anodontina                  
                 |  |                                                 
                 |  |     +-------------- Utterbackia                 
                 |  |  +--+                                           
                 +--+  |  |  +----------- Pyganodon_grandis           
                    |  |  +--+                                        
                    |  |     |  +-------- Pyganodon_lacustris         
                    |  |     +--+                                     
                    |  |        |  +----- Pyganodon_cataracta__2      
                    +--+        +--+                                  
                       |           |  +-- Pyganodon_cataracta__1      
                       |           +--+                               
                       |              +-- Ambleminae                  
                       |                                              
                       |  +-------------- Utterbackiana               
                       |  |                                           
                       +--+  +----------- Simpsonaias                 
                          |  |                                        
                          |  |        +-- Alasminota                  
                          +--+  +-----+                               
                             |  |     +-- Alasmidonta                 
                             |  |                                     
                             +--+     +-- Prolasmidonta               
                                |  +--+                               
                                |  |  +-- Platynaias                  
                                +--+                                  
                                   |  +-- Anodontoides                
                                   +--+                               
                                      +-- Strophitus

Bolotov et al., 2024 [Publ 4369], fig. 2

iTtTbiriSoTroTroSiD
62 62 60 5 0 0 0 91 
  +-------- Lamellidentini **
  |                       
--+     +-- Leoparreysiini **
  |  +--+                 
  |  |  +-- Parreysiini **   
  +--+                    
     |  +-- Coelaturini **   
     +--+                 
        +-- Indochinellini **

Bolotov et al., 2024 [Publ 4369], fig. S1

iTtTbiriSoTroTroSiD
62 62 60 5 1 1 1 77 
  +-------------- Gonideinae    
  |                             
--+  +----------- Parreysia **     
  |  |                          
  +--+        +-- Leoparreysiini **
     |  +-----+                 
     |  |     +-- Lamellidentini **
     +--+                       
        |  +----- Assamnaia **     
        +--+                    
           |  +-- Coelaturini **   
           +--+                 
              +-- Indochinellini **

Bolotov et al., 2022 [Publ 4252], fig. 2

iTtTbiriSoTroTroSiD
57 57 53 5 4 4 1 96 
  +-------------- Gonideinae    
--+                             
  |  +----------- Lamellidentini **
  +--+                          
     |  +-------- Parreysiini **   
     +--+                       
        |  +----- Leoparreysiini **
        +--+                    
           |  +-- Coelaturini **   
           +--+                 
              +-- Indochinellini **

Bolotov et al., 2022 [Publ 4252], fig. S1

iTtTbiriSoTroTroSiD
57 57 47 5 4 4 1 86 
  +-------------- Gonideinae    
--+                             
  |  +----------- Parreysiini **   
  +--+                          
     |  +-------- Lamellidentini **
     +--+                       
        |  +----- Leoparreysiini **
        +--+                    
           |  +-- Coelaturini **   
           +--+                 
              +-- Indochinellini **

Bolotov et al., 2022 [Publ 4239], fig. 2

iTtTbiriSoTroTroSiD
54 54 52 5 4 4 1 83 
  +----------- Gonideinae    
  |                          
--+  +-------- Lamellidentini **
  |  |                       
  +--+     +-- Parreysiini **   
     |  +--+                 
     |  |  +-- Leoparreysiini **
     +--+                    
        |  +-- Coelaturini **   
        +--+                 
           +-- Indochinellini **

Bolotov et al., 2022 [Publ 4239], fig. S1

iTtTbiriSoTroTroSiD
54 54 50 5 4 4 1 85 
  +-------------- Gonideinae    
--+                             
  |  +----------- Lamellidentini **
  +--+                          
     |  +-------- Parreysiini **   
     +--+                       
        |  +----- Leoparreysiini **
        +--+                    
           |  +-- Coelaturini **   
           +--+                 
              +-- Indochinellini **

Bolotov et al., 2022 [Publ 4239], fig. S2

iTtTbiriSoTroTroSiD
54 54 52 5 4 4 1 79 
  +-------------- Gonideinae    
--+                             
  |  +----------- Parreysiini **   
  +--+                          
     |  +-------- Lamellidentini **
     +--+                       
        |  +----- Leoparreysiini **
        +--+                    
           |  +-- Coelaturini **   
           +--+                 
              +-- Indochinellini **

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
41 41 39 5 76 71 1 98 
  +----------- Gonideinae    
  |                          
--+  +-------- Lamellidentini **
  |  |                       
  +--+     +-- Parreysiini **   
     |  +--+                 
     |  |  +-- Leoparreysiini **
     +--+                    
        |  +-- Coelaturini **   
        +--+                 
           +-- Indochinellini **

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
40 40 37 5 71 71 1 98 
  +----------- Gonideinae    
  |                          
--+  +-------- Lamellidentini **
  |  |                       
  +--+     +-- Parreysiini **   
     |  +--+                 
     |  |  +-- Leoparreysiini **
     +--+                    
        |  +-- Coelaturini **   
        +--+                 
           +-- Indochinellini **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
38 38 28 5 86 86 3 105 
              +-------------- Pseudodontini           
              |                                       
  +-----------+  +----------- Rectidentini            
  |           |  |                                    
  |           |  |  +-------- Unioninae               
  |           +--+  |                                 
  |              |  |  +----- Ambleminae              
  |              |  |  |                              
  |              +--+  |  +-- Chamberlainia           
  |                 |  |  |                           
  |                 |  |  +-- Sinohyriopsis           
  |                 |  |  |                           
  |                 |  |  +-- Lamprotula_caveata      
  |                 +--+  |                           
  |                    |  +-- Lamprotula_leaii        
  |                    |  |                           
  |                    |  +-- Potomida                
  |                    |  |                           
  |                    |  +-- Pronodularia            
  |                    |  |                           
  |                    +--+-- Gonidea                 
  |                       |                           
  |                       +-- Leguminaia_wheatleyi    
--+                       |                           
  |                       +-- Leguminaia              
  |                       |                           
  |                       +-- Microcondylaea          
  |                       |                           
  |                       +-- Sinosolenaia_carinata   
  |                       |                           
  |                       +-- Sinosolenaia_oleivora   
  |                       |                           
  |                       +-- Sinosolenaia_recognita  
  |                                                   
  |           +-------------- Trapezidens **             
  |           |                                       
  |  +--------+  +----------- Lamellidens_lamellatus **  
  |  |        |  |                                    
  |  |        +--+  +-------- Lamellidens_jenkinsianus **
  |  |           |  |                                 
  |  |           |  |  +----- Lamellidens_generosus **   
  |  |           +--+  |                              
  |  |              |  +----- Lamellidens_mainwaringi ** 
  |  |              |  |                              
  |  |              +--+  +-- Lamellidens_ferrugineus ** 
  +--+                 +--+                           
     |                 |  +-- Lamellidens_marginalis **  
     |                 |                              
     |                 |  +-- Lamellidens_savadiensis ** 
     |                 +--+                           
     |                    +-- Lamellidens_corrianus__1 **
     |                                                
     |  +-------------------- Parreysiini **             
     |  |                                             
     +--+  +----------------- Leoparreysiini **          
        |  |                                          
        +--+  +-------------- Coelaturini **             
           |  |                                       
           |  |        +----- Indochinella **            
           +--+  +-----+                              
              |  |     |  +-- Scabies **                 
              |  |     +--+                           
              +--+        +-- Radiatula **               
                 |                                    
                 |  +-------- Indonaia_shurtleffiana **  
                 |  |                                 
                 +--+  +----- Indonaia_rugosa **         
                    |  |                              
                    |  |  +-- Indonaia_andersoniana **   
                    +--+--+                           
                       |  +-- Indonaia_caerulea **       
                       |                              
                       |  +-- Indonaia_bonneaudii **     
                       +--+                           
                          +-- Lamellidens_corrianus__2 **

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 1

iTtTbiriSoTroTroSiD
36 36 22 3 0 0 0 95 
  +----- Leoparreysiini **
--+                    
  |  +-- Indochinellini **
  +--+                 
     +-- Coelaturini **

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 2

iTtTbiriSoTroTroSiD
36 36 20 3 0 0 0 97 
  +----- Leoparreysiini **
--+                    
  |  +-- Indochinellini **
  +--+                 
     +-- Coelaturini **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
34 34 30 5 166 127 3 100 
           +----------- Lamellidentini ** 
  +--------+                           
  |        |  +-------- Parreysiini **    
  |        +--+                        
  |           |  +----- Leoparreysiini ** 
  |           +--+                     
--+              |  +-- Coelaturini **    
  |              +--+                  
  |                 +-- Indochinellini ** 
  |                                    
  |  +----------------- Unioninae      
  +--+                                 
     |  +-------------- Rectidentini   
     +--+                              
        |  +----------- Ambleminae     
        +--+                           
           |  +-------- Chamberlainiini
           +--+                        
              |  +----- Lamprotulini   
              +--+                     
                 |  +-- Gonideini      
                 +--+                  
                    +-- Pseudodontini

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
34 34 30 5 164 125 3 95 
           +----------- Lamellidentini ** 
  +--------+                           
  |        |  +-------- Parreysiini **    
  |        +--+                        
  |           |  +----- Leoparreysiini ** 
  |           +--+                     
--+              |  +-- Coelaturini **    
  |              +--+                  
  |                 +-- Indochinellini ** 
  |                                    
  |  +----------------- Pseudodontini  
  +--+                                 
     |  +-------------- Rectidentini   
     +--+                              
        |  +----------- Unioninae      
        +--+                           
           |  +-------- Chamberlainiini
           +--+                        
              |  +----- Ambleminae     
              +--+                     
                 |  +-- Lamprotulini   
                 +--+                  
                    +-- Gonideini

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
34 34 25 5 166 127 3 95 
        +-------- Parreysiini **       
        |                           
  +-----+-------- Lamellidentini **    
  |     |                           
  |     |  +----- Leoparreysiini **    
  |     +--+                        
  |        |  +-- Coelaturini **       
  |        +--+                     
  |           +-- Indochinellini **    
  |                                 
  |  +----------- Chamberlainia     
  |  |                              
  |  +----------- Megalonaias       
--+  |                              
  |  +----------- Uniomerus         
  |  |                              
  |  +----------- Quadrulina        
  |  |                              
  |  +----------- Sinohyriopsis     
  |  |                              
  |  +----------- Lamprotulini      
  |  |                              
  |  +----------- Gonideini         
  |  |                              
  +--+----------- Pseudodontini     
     |                              
     +----------- Rectidentini      
     |                              
     +----------- Unioninae         
     |                              
     |  +-------- Amblema           
     |  |                           
     |  +-------- Toxolasma         
     |  |                           
     |  +-------- Pleurobemini      
     |  |                           
     |  |     +-- Obliquaria        
     |  +-----+                     
     |  |     +-- Plectomerus       
     +--+                           
        |  +----- Ellipsaria        
        |  |                        
        |  +----- Potamilus_leptodon
        |  |                        
        |  +----- Medionidus        
        |  |                        
        |  +----- Potamilus_alatus  
        |  |                        
        |  +----- Ptychobranchus    
        +--+                        
           +----- Truncilla         
           |                        
           +----- Venustaconcha     
           |                        
           +----- Cambarunio        
           |                        
           +----- Leaunio           
           |                        
           |  +-- Ortmanniana       
           +--+                     
              +-- Lampsilis

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
29 29 26 5 109 84 3 94 
                    +-- Parreysiini **    
              +-----+                  
              |     +-- Lamellidentini ** 
  +-----------+                        
  |           |  +----- Leoparreysiini ** 
  |           +--+                     
  |              |  +-- Coelaturini **    
--+              +--+                  
  |                 +-- Indochinellini ** 
  |                                    
  |  +----------------- Unioninae      
  +--+                                 
     |  +-------------- Rectidentini   
     +--+                              
        |  +----------- Ambleminae     
        +--+                           
           |  +-------- Pseudodontini  
           +--+                        
              |  +----- Gonideini      
              +--+                     
                 |  +-- Chamberlainiini
                 +--+                  
                    +-- Lamprotulini

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
29 29 27 5 109 84 3 95 
           +----------- Lamellidentini ** 
  +--------+                           
  |        |  +-------- Parreysiini **    
  |        +--+                        
  |           |  +----- Leoparreysiini ** 
  |           +--+                     
--+              |  +-- Coelaturini **    
  |              +--+                  
  |                 +-- Indochinellini ** 
  |                                    
  |  +----------------- Rectidentini   
  +--+                                 
     |  +-------------- Pseudodontini  
     +--+                              
        |  +----------- Unioninae      
        +--+                           
           |  +-------- Ambleminae     
           +--+                        
              |  +----- Gonideini      
              +--+                     
                 |  +-- Chamberlainiini
                 +--+                  
                    +-- Lamprotulini

Pfeiffer et al., 2018 [Publ 3827], fig. 2

iTtTbiriSoTroTroSiD
27 27 25 5 1 0 0 99 
  +----------- Lamellidentini **
--+                          
  |  +-------- Parreysiini **   
  +--+                       
     |  +----- Leoparreysiini **
     +--+                    
        |  +-- Coelaturini **   
        +--+                 
           +-- Indochinellini **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
24 24 22 5 58 58 3 95 
     +----------- Lamellidentini **
  +--+                          
  |  |  +-------- Parreysiini **   
  |  +--+                       
  |     |  +----- Leoparreysiini **
  |     +--+                    
  |        |  +-- Coelaturini **   
--+        +--+                 
  |           +-- Indochinellini **
  |                             
  |  +----------- Pseudodontini 
  |  |                          
  |  |  +-------- Rectidentini  
  +--+  |                       
     |  |  +----- Unioninae     
     |  |  |                    
     +--+  |  +-- Lamprotula    
        |  |  |                 
        |  |  +-- Potomida      
        |  |  |                 
        |  |  +-- Pronodularia  
        +--+  |                 
           |  +-- Gonidea       
           |  |                 
           |  +-- Leguminaia    
           |  |                 
           |  +-- Microcondylaea
           |  |                 
           +--+-- Amblemini     
              |                 
              +-- Ortmanniana   
              |                 
              +-- Lampsilis     
              |                 
              +-- Cambarunio    
              |                 
              +-- Elliptio      
              |                 
              +-- Pleurobema    
              |                 
              +-- Quadrulini

Bolotov et al., 2022 [Publ 4250], fig. 8

iTtTbiriSoTroTroSiD
24 24 22 4 25 23 1 91 
  +----------- Gonideinae    
--+                          
  |  +-------- Lamellidentini **
  +--+                       
     |  +----- Parreysiini **   
     +--+                    
        |  +-- Leoparreysiini **
        +--+                 
           +-- Indochinellini **

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

iTtTbiriSoTroTroSiD
23 23 21 5 0 0 0 98 
  +----------- Lamellidentini **
--+                          
  |  +-------- Leoparreysiini **
  +--+                       
     |  +----- Parreysiini **   
     +--+                    
        |  +-- Coelaturini **   
        +--+                 
           +-- Indochinellini **

Bolotov et al., 2022 [Publ 4250], fig. S2

iTtTbiriSoTroTroSiD
15 15 13 3 0 0 0 102 
  +----- Indochinellini **
--+                    
  |  +-- Parreysiini **   
  +--+                 
     +-- Leoparreysiini **

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
14 14 10 1 68 68 1 105 
  +-- Indochinellini **
--+                 
  +-- Gonideinae

Bolotov et al., 2019 [Publ 3974], fig. 1

iTtTbiriSoTroTroSiD
13 13 11 3 5 5 1 98 
  +-------- Gonideinae    
--+                       
  |  +----- Leoparreysiini **
  +--+                    
     |  +-- Coelaturini **   
     +--+                 
        +-- Indochinellini **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
13 13 11 4 78 61 3 100 
              +-------- Lamellidentini ** 
  +-----------+                        
  |           |  +----- Leoparreysiini ** 
  |           +--+                     
  |              |  +-- Coelaturini **    
--+              +--+                  
  |                 +-- Indochinellini ** 
  |                                    
  |  +----------------- Unioninae      
  +--+                                 
     |  +-------------- Rectidentini   
     +--+                              
        |  +----------- Ambleminae     
        +--+                           
           |  +-------- Chamberlainiini
           +--+                        
              |  +----- Lamprotulini   
              +--+                     
                 |  +-- Pseudodontini  
                 +--+                  
                    +-- Gonideini

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
13 13 11 4 78 61 3 98 
              +-------- Lamellidentini ** 
  +-----------+                        
  |           |  +----- Leoparreysiini ** 
  |           +--+                     
  |              |  +-- Coelaturini **    
--+              +--+                  
  |                 +-- Indochinellini ** 
  |                                    
  |  +----------------- Unioninae      
  +--+                                 
     |  +-------------- Rectidentini   
     +--+                              
        |  +----------- Ambleminae     
        +--+                           
           |  +-------- Chamberlainiini
           +--+                        
              |  +----- Lamprotulini   
              +--+                     
                 |  +-- Pseudodontini  
                 +--+                  
                    +-- Gonideini

Konopleva et al., 2020 [Publ 4072], fig. 2

iTtTbiriSoTroTroSiD
12 12 9 2 0 0 0 102 
  +-- Leoparreysiini **
--+                 
  +-- Lamellidentini **

Konopleva et al., 2019 [Publ 3900], fig. 1

iTtTbiriSoTroTroSiD
12 12 9 1 39 37 1 100 
  +----- Pseudodontini 
--+                    
  |  +-- Indochinellini **
  +--+                 
     +-- Rectidentini

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

iTtTbiriSoTroTroSiD
11 11 12 1 17 13 1 100 
1

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
11 11 9 4 20 10 1 98 
  +----------- Gonideinae    
--+                          
  |  +-------- Lamellidentini **
  +--+                       
     |  +----- Leoparreysiini **
     +--+                    
        |  +-- Coelaturini **   
        +--+                 
           +-- Indochinellini **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
10 10 8 4 47 40 3 99 
  +-------------- Ambleminae    
  |                             
--+-------------- Gonideinae    
  |                             
  |  +----------- Unioninae     
  +--+                          
     |  +-------- Lamellidentini **
     +--+                       
        |  +----- Leoparreysiini **
        +--+                    
           |  +-- Coelaturini **   
           +--+                 
              +-- Indochinellini **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
10 10 8 4 47 40 3 99 
  +----------------- Ambleminae    
--+                                
  |  +-------------- Gonideinae    
  +--+                             
     |  +----------- Unioninae     
     +--+                          
        |  +-------- Lamellidentini **
        +--+                       
           |  +----- Leoparreysiini **
           +--+                    
              |  +-- Coelaturini **   
              +--+                 
                 +-- Indochinellini **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
9 9 8 4 27 15 3 100 
        +-------- Lamellidentini **
  +-----+                       
  |     |  +----- Leoparreysiini **
  |     +--+                    
  |        |  +-- Indochinellini **
  |        +--+                 
--+           +-- Coelaturini **   
  |                             
  |           +-- Pseudodontini 
  |  +--------+                 
  |  |        +-- Rectidentini  
  +--+                          
     |  +-------- Unioninae     
     +--+                       
        |  +----- Gonideini     
        +--+                    
           |  +-- Lamprotulini  
           +--+                 
              +-- Ambleminae

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
9 9 7 4 27 15 3 100 
           +-------- Lamellidentini **
  +--------+                       
  |        |  +----- Leoparreysiini **
  |        +--+                    
  |           |  +-- Indochinellini **
--+           +--+                 
  |              +-- Coelaturini **   
  |                                
  |  +-------------- Rectidentini  
  +--+                             
     |  +----------- Pseudodontini 
     +--+                          
        |  +-------- Unioninae     
        +--+                       
           |  +----- Gonideini     
           +--+                    
              |  +-- Lamprotulini  
              +--+                 
                 +-- Ambleminae

Bolotov et al., 2022 [Publ 4250], fig. S4

iTtTbiriSoTroTroSiD
9 9 6 1 0 0 0 101 
1

Graf et al., 2014 [Publ 3591], fig. 3

iTtTbiriSoTroTroSiD
9 9 12 2 2 1 1 93 
  +-- Leoparreysiini **
--+                 
  +-- Coelaturini **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
8 8 6 4 32 22 3 101 
        +----- Unioninae     
  +-----+                    
  |     |  +-- Ambleminae    
  |     +--+                 
--+        +-- Gonideinae    
  |                          
  |  +-------- Lamellidentini **
  +--+                       
     |  +----- Leoparreysiini **
     +--+                    
        |  +-- Coelaturini **   
        +--+                 
           +-- Indochinellini **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
7 7 5 4 49 48 3 104 
        +----- Unioninae     
  +-----+                    
  |     |  +-- Ambleminae    
  |     +--+                 
--+        +-- Gonideinae    
  |                          
  |  +-------- Lamellidentini **
  +--+                       
     |  +----- Parreysiini **   
     +--+                    
        |  +-- Leoparreysiini **
        +--+                 
           +-- Indochinellini **

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
5 5 3 4 40 38 3 102 
  +----------------- Ambleminae     
  |                                 
  |     +----------- Rectidentini   
  |     |                           
  |  +--+        +-- Schepmaniini   
--+  |  |  +-----+                  
  |  |  |  |     +-- Pseudodontini  
  |  |  +--+                        
  |  |     |  +----- Caudiculatus   
  |  |     +--+                     
  |  |        |  +-- Potomida       
  +--+        +--+                  
     |           +-- Gonideini      
     |                              
     |        +----- Unioninae      
     |  +-----+                     
     |  |     |  +-- Chamberlainiini
     |  |     +--+                  
     +--+        +-- Discomya       
        |                           
        |  +-------- Lamellidentini ** 
        +--+                        
           |  +----- Leoparreysiini ** 
           +--+                     
              |  +-- Indochinellini ** 
              +--+                  
                 +-- Coelaturini **

Bolotov et al., 2022 [Publ 4250], fig. S3

iTtTbiriSoTroTroSiD
5 5 3 1 0 0 0 102 
1

Konopleva et al., 2017 [Publ 3807], fig. 5

iTtTbiriSoTroTroSiD
5 5 3 1 0 0 0 102 
1

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
4 4 2 4 76 65 3 103 
                    +-------- Lamellidentini ** 
  +-----------------+                        
  |                 |  +----- Leoparreysiini ** 
  |                 +--+                     
  |                    |  +-- Coelaturini **    
--+                    +--+                  
  |                       +-- Indochinellini ** 
  |                                          
  |  +----------------------- Unioninae      
  |  |                                       
  +--+  +-------------------- Rectidentini   
     |  |                                    
     +--+  +----------------- Ambleminae     
        |  |                                 
        |  |              +-- Chamberlainiini
        +--+  +-----------+                  
           |  |           +-- Lamprotulini   
           |  |                              
           +--+           +-- Parvasolenaia  
              |  +--------+                  
              |  |        +-- Pseudodontini  
              +--+                           
                 |  +-------- Sinosolenaia   
                 +--+                        
                    |  +----- Gonidea        
                    +--+                     
                       |  +-- Microcondylaea 
                       +--+                  
                          +-- Leguminaia

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
3 3 1 2 19 17 1 104 
  +----- Unioninae     
--+                    
  |  +-- Lamellidentini **
  +--+                 
     +-- Indochinellini **

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 38 36 3 106 
           +-- Gonideinae    
  +--------+                 
  |        +-- Ambleminae    
--+                          
  |  +-------- Unioninae     
  +--+                       
     |  +----- Parreysiini **   
     +--+                    
        |  +-- Coelaturini **   
        +--+                 
           +-- Indochinellini **

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 38 36 3 106 
           +-- Gonideinae    
  +--------+                 
  |        +-- Ambleminae    
--+                          
  |  +-------- Unioninae     
  +--+                       
     |  +----- Parreysiini **   
     +--+                    
        |  +-- Coelaturini **   
        +--+                 
           +-- Indochinellini **

Imsong & Murali, 2023 [Publ 4370], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 0 0 0 104 
  +----- Lamellidentini **
--+                    
  |  +-- Indochinellini **
  +--+                 
     +-- Parreysiini **

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 38 36 3 106 
           +-- Ambleminae    
  +--------+                 
  |        +-- Gonideinae    
--+                          
  |  +-------- Unioninae     
  +--+                       
     |  +----- Parreysiini **   
     +--+                    
        |  +-- Coelaturini **   
        +--+                 
           +-- Indochinellini **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
3 3 0 3 119 108 3 105 
                 +-- Coelaturini **    
                 |                  
  +--------------+-- Leoparreysiini ** 
  |              |                  
  |              +-- Indochinellini ** 
--+                                 
  |  +-------------- Unioninae      
  |  |                              
  +--+  +----------- Rectidentini   
     |  |                           
     +--+  +-------- Ambleminae     
        |  |                        
        +--+     +-- Chamberlainiini
           |  +--+                  
           |  |  +-- Lamprotulini   
           +--+                     
              |  +-- Pseudodontini  
              +--+                  
                 +-- Gonideini

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 42 40 3 106 
  +----------------------- Gonideinae    
  |                                      
  |  +-------------------- Ambleminae    
--+  |                                   
  |  |                 +-- Anemina       
  |  |  +--------------+                 
  |  |  |              +-- Amuranodonta  
  +--+  |                                
     |  |           +----- Parreysiini **   
     |  +-----------+                    
     |  |           |  +-- Coelaturini **   
     |  |           +--+                 
     +--+              +-- Indochinellini **
        |                                
        |           +----- Pletholophus  
        |  +--------+                    
        |  |        |  +-- Sinanodonta   
        |  |        +--+                 
        |  |           +-- Beringiana    
        +--+                             
           |  +----------- Cristaria     
           |  |                          
           |  |        +-- Alasmidontina 
           +--+--------+                 
              |        +-- Anodontina    
              |                          
              |  +-------- Lanceolariini 
              +--+                       
                 |  +----- Lepidodesmini 
                 +--+                    
                    |  +-- Acuticostini  
                    +--+                 
                       +-- Unionini

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
3 3 1 2 19 17 1 104 
  +----- Unioninae     
--+                    
  |  +-- Lamellidentini **
  +--+                 
     +-- Indochinellini **

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
3 3 0 3 38 36 3 105 
        +-- Gonideinae    
  +-----+                 
  |     +-- Ambleminae    
--+                       
  |  +----- Unioninae     
  |  |                    
  +--+  +-- Coelaturini **   
     |  |                 
     +--+-- Indochinellini **
        |                 
        +-- Parreysiini **

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
3 3 0 3 42 40 3 105 
        +-- Gonideinae    
  +-----+                 
  |     +-- Ambleminae    
--+                       
  |  +----- Unioninae     
  |  |                    
  +--+  +-- Coelaturini **   
     |  |                 
     +--+-- Indochinellini **
        |                 
        +-- Parreysiini **

Choi et al., 2020 [Publ 4169], fig. 5

iTtTbiriSoTroTroSiD
3 3 1 3 7 7 1 106 
  +-------- Unioninae     
--+                       
  |  +----- Indochinellini **
  +--+                    
     |  +-- Lamellidentini **
     +--+                 
        +-- Leoparreysiini **

Lyubas et al., 2022 [Publ 4255], fig. S1

iTtTbiriSoTroTroSiD
2 2 0 1 29 29 2 -- 
  +----- Parreysiini **
--+                 
  |  +-- Gonideinae 
  +--+              
     +-- Unioninae

Dai et al., 2026 [Publ 4757], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 32 30 1 -- 
  +----- Unioninae     
--+                    
  |  +-- Indochinellini **
  +--+                 
     +-- Lamellidentini **

Lyubas et al., 2022 [Publ 4255], fig. 7

iTtTbiriSoTroTroSiD
2 2 0 1 29 29 2 -- 
  +----- Parreysiini **
--+                 
  |  +-- Gonideinae 
  +--+              
     +-- Unioninae

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 12 11 3 -- 
        +-- Lamellidentini **
  +-----+                 
  |     +-- Coelaturini **   
--+                       
  |  +----- Ambleminae    
  +--+                    
     |  +-- Unioninae     
     +--+                 
        +-- Gonideinae

Dai et al., 2024 [Publ 4385], fig. 3-BI

iTtTbiriSoTroTroSiD
2 2 0 2 29 27 1 -- 
  +----- Unioninae     
--+                    
  |  +-- Lamellidentini **
  +--+                 
     +-- Indochinellini **

Graf, 2000 [Publ 2647], fig. 1

iTtTbiriSoTroTroSiD
2 11 0 2 15 4 2 113 
              +-------- Unionini      
  +-----------+                       
  |           |  +----- Pleurobemini  
  |           +--+                    
  |              |  +-- Anodontini    
--+              +--+                 
  |                 +-- Lampsilini    
  |                                   
  |  +----------------- Parreysiini **   
  |  |                                
  +--+  +-------------- Coelaturini **   
     |  |                             
     |  |     +-------- Castaliini    
     +--+  +--+                       
        |  |  |  +----- Rhipidodontini
        |  |  +--+                    
        |  |     |  +-- Hyridellini   
        +--+     +--+                 
           |        +-- Velesunioninae
           |                          
           |  +-------- Iridininae    
           |  |                       
           +--+  +----- Anodontitinae 
              |  |                    
              +--+----- Mycetopodinae 
                 |                    
                 |  +-- Etheria       
                 +--+                 
                    +-- Mulleria

Dai et al., 2024 [Publ 4385], fig. 3-ML

iTtTbiriSoTroTroSiD
2 2 0 2 29 27 1 -- 
  +----- Unioninae     
--+                    
  |  +-- Lamellidentini **
  +--+                 
     +-- Indochinellini **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 29 27 3 -- 
        +-- Lamellidentini **
  +-----+                 
  |     +-- Coelaturini **   
--+                       
  |  +----- Unioninae     
  +--+                    
     |  +-- Ambleminae    
     +--+                 
        +-- Gonideinae

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
2 10 0 1 14 8 3 112 
  +-------------------------- Lamellidens_mainwaringi **
--+                                                  
  |  +----------------------- Lamprotula             
  +--+                                               
     |  +-------------------- Potomida               
     +--+                                            
        |  +----------------- Lampsilini             
        +--+                                         
           |  +-------------- Pleurobemini           
           +--+                                      
              |  +----------- Aculamprotulina        
              +--+                                   
                 |  +-------- Unio_pictorum          
                 +--+                                
                    |  +----- Anodontini             
                    +--+                             
                       |  +-- Unio_ravoisieri        
                       +--+                          
                          +-- Lamellidens_corrianus **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
1 1 0 1 140 113 3 -- 
  +-------- Ambleminae    
--+                       
  |  +----- Gonideinae    
  +--+                    
     |  +-- Lamellidentini **
     +--+                 
        +-- Unioninae

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
1 1 0 1 95 84 3 -- 
  +-------- Ambleminae    
--+                       
  |  +----- Gonideinae    
  +--+                    
     |  +-- Lamellidentini **
     +--+                 
        +-- Unioninae

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 67 44 3 -- 
  +-------------- Indochinellini ** 
--+                              
  |  +----------- Unioninae      
  +--+                           
     |  +-------- Chamberlainiini
     +--+                        
        |  +----- Ambleminae     
        +--+                     
           |  +-- Lamprotulini   
           +--+                  
              +-- Gonideini

Wu et al., 2021 [Publ 4215], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 23 23 2 -- 
  +----- Indochinellini **
--+                    
  |  +-- Unioninae     
  +--+                 
     +-- Gonideinae

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
1 1 0 1 14 8 3 -- 
-- Lamellidens_corrianus **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
1 1 0 1 91 85 3 -- 
  +-------- Ambleminae    
--+                       
  |  +----- Gonideinae    
  +--+                    
     |  +-- Lamellidentini **
     +--+                 
        +-- Unioninae

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 33 10 3 -- 
     +-- Ambleminae    
  +--+                 
  |  +-- Indochinellini **
--+                    
  |  +-- Gonideinae    
  +--+                 
     +-- Unioninae

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 33 10 3 -- 
     +-- Ambleminae    
  +--+                 
  |  +-- Indochinellini **
--+                    
  |  +-- Gonideinae    
  +--+                 
     +-- Unioninae

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 67 44 3 -- 
  +-------- Indochinellini **
--+                       
  |  +----- Unioninae     
  +--+                    
     |  +-- Ambleminae    
     +--+                 
        +-- Gonideinae

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
1 1 0 1 14 8 3 -- 
  +-- Aculamprotulina
  |                  
  +-- Lamprotula     
  |                  
  +-- Lampsilini     
  |                  
  +-- Potomida       
  |                  
--+-- Unio_pictorum  
  |                  
  +-- Anodontini     
  |                  
  +-- Unio_ravoisieri
  |                  
  +-- Lamellidentini ** 
  |                  
  +-- Pleurobemini

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
1 1 0 1 38 36 3 -- 
     +-- Lamellidentini **
  +--+                 
  |  +-- Unioninae     
--+                    
  |  +-- Ambleminae    
  +--+                 
     +-- Gonideinae

Smith, 2021 [Publ 4331], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 19 15 3 -- 
  +-------- Ambleminae    
--+                       
  |  +----- Gonideinae    
  +--+                    
     |  +-- Indochinellini **
     +--+                 
        +-- Unioninae

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 46 24 3 -- 
-- Coelatura_aegyptiaca **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
1 1 0 1 95 84 3 -- 
  +-------- Ambleminae    
--+                       
  |  +----- Gonideinae    
  +--+                    
     |  +-- Lamellidentini **
     +--+                 
        +-- Unioninae

Dai et al., 2024 [Publ 4386], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 13 13 1 -- 
  +-- Indochinellini **
--+                 
  +-- Unioninae

Combosch et al., 2017 [Publ 3806], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 14 2 2 -- 
  +----- Unioninae     
--+                    
  |  +-- Ambleminae    
  +--+                 
     +-- Indochinellini **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 36 34 3 -- 
  +-------- Ambleminae    
--+                       
  |  +----- Gonideinae    
  +--+                    
     |  +-- Lamellidentini **
     +--+                 
        +-- Unioninae

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iTtTbiriSoTroTroSiD
1 1 0 1 24 24 3 -- 
  +-------------------------------- Lamellidentini **
  |                                               
--+  +----------------------------- Unioninae     
  |  |                                            
  +--+  +-------------------------- Lamprotulini  
     |  |                                         
     +--+  +----------------------- Quadrulini    
        |  |                                      
        +--+  +-------------------- Gonideini     
           |  |                                   
           |  |              +----- Eurynia       
           +--+  +-----------+                    
              |  |           |  +-- Cambarunio    
              |  |           +--+                 
              |  |              +-- Ptychobranchus
              +--+                                
                 |              +-- Amblemini     
                 |  +-----------+                 
                 |  |           +-- Pleurobema    
                 +--+                             
                    |  +----------- Truncilla     
                    |  |                          
                    +--+        +-- Lampsilis     
                       |  +-----+                 
                       |  |     +-- Epioblasma    
                       +--+                       
                          |  +----- Ligumia       
                          +--+                    
                             |  +-- Obliquaria    
                             +--+                 
                                +-- Ortmanniana

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
1 1 0 1 30 17 3 -- 
  +----------------------------- Gonideinae 
  |                                         
--+  +-------------------------- Coelaturini **
  |  |                                      
  +--+  +----------------------- Fusconaia  
     |  |                                   
     |  +----------------------- Amblemini  
     +--+                                   
        |  +-------------------- Quadrulini 
        |  |                                
        +--+  +----------------- Pleurobema 
           |  |                             
           +--+  +-------------- Eurynia    
              |  |                          
              |  |           +-- Cyrtonaias 
              +--+-----------+              
                 |           +-- Unioninae  
                 |                          
                 |  +----------- Ortmanniana
                 +--+                       
                    |  +-------- Glebula    
                    +--+                    
                       |  +----- Potamilus  
                       +--+                 
                          |  +-- Ligumia    
                          +--+              
                             +-- Toxolasma

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
1 1 0 1 30 17 3 -- 
  +-------- Coelaturini **
--+                    
  |  +----- Unioninae  
  +--+                 
     |  +-- Gonideinae 
     +--+              
        +-- Ambleminae

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
1 1 0 1 30 17 3 -- 
-- Coelatura_aegyptiaca **

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
1 1 0 1 28 5 3 -- 
  +-------- Gonideinae    
--+                       
  |  +----- Unioninae     
  +--+                    
     |  +-- Ambleminae    
     +--+                 
        +-- Indochinellini **

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 45 23 3 -- 
-- Coelatura_aegyptiaca **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
1 1 0 1 30 17 3 -- 
-- Coelatura_aegyptiaca **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
1 1 0 1 91 85 3 -- 
  +-------- Ambleminae    
--+                       
  |  +----- Gonideinae    
  +--+                    
     |  +-- Lamellidentini **
     +--+                 
        +-- Unioninae

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 67 44 3 -- 
  +-------- Indochinellini **
--+                       
  |  +----- Unioninae     
  +--+                    
     |  +-- Ambleminae    
     +--+                 
        +-- Gonideinae

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 52 50 3 -- 
  +-------- Ambleminae    
--+                       
  |  +----- Gonideinae    
  +--+                    
     |  +-- Lamellidentini **
     +--+                 
        +-- Unioninae

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 46 24 3 -- 
  +-------- Pseudodontini
  |                      
--+  +----- Coelaturini **  
  |  |                   
  |  +----- Rectidentini 
  +--+                   
     +----- Unioninae    
     |                   
     |  +-- Gonideini    
     +--+                
        +-- Ambleminae

Xiang et al., 2025 [Publ 4433], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 27 27 1 -- 
  +-- Indochinellini **
--+                 
  +-- Unioninae

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 17 3 -- 
-- Coelatura_aegyptiaca **

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 45 23 3 -- 
-- Coelatura_aegyptiaca **

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 8 8 1 -- 
  +----- Rectidentini  
--+                    
  |  +-- Pseudodontini 
  +--+                 
     +-- Indochinellini **

Combosch et al., 2017 [Publ 3806], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 2 2 -- 
  +----- Unioninae     
--+                    
  |  +-- Indochinellini **
  +--+                 
     +-- Ambleminae

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 17 3 -- 
-- Coelatura_aegyptiaca **