|
Bivalvia
| Unionoida
| Unionoidea
| Unionidae
| Ambleminae
tribe POPENAIADINI Heard & Guckert, 1970: cladomics6 genera
Cladomics pages are experimental — even more so than the rest of this site!
The purpose of the cladomics pages for each genus and family-group level taxon is to
provide statistics and references indicating the extent to which published phylogenetic
work supports the classification used here. The Cladomics Summary provides an
overview of the number of studies that have included terminal species representing this
taxon as well as an indication of how frequently it has been recovered as monophyletic.
Following the summary, the available published clades representing the taxon are listed.
For more information, visit the
Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various
statistics are explained on the FARQ!
Cladomics Summary
number of species = 36 (36 total)
number of subtaxa (rank: genus) = 6
analyzed in 43 cladograms in 27 publications
maximum number of terminals in an analysis = 5 (mean = 1.47, median = 1)
maximum number of subtaxa (genus) = 3 (mean = 1.28, median = 1)
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 11
maximum number of included terminals = 5, (mean = 2.82 median = 2)
maximum number of subtaxa (genus) = 3 (mean = 2.09, median = 2)
proportion of cladograms that support basic monophyly = 1
Rigorous Monophyly
number of cladograms that test rigorous monophyly = 11
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 5 (mean = 2.82, median = 2)
maximum number of subtaxa (genus) = 3 (mean = 2.09, median = 2)
proportion of cladograms that support rigorous monophyly = 1
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome.
See the Cladmoics FARQ for more information.
Cladograms
** Top Monophyly Cladogram **
iT| tT | bi | riS | oT | roT | roS | iD |
| 4 |
4 |
3 |
3 |
2 |
2 |
1 |
0 |
+-------- Lampsilini
--+
| +----- Popenaias **
+--+
| +-- Nephronaias **
+--+
+-- Psoronaias **
** Top Sister-Group Cladogram **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
101 |
89 |
4 |
-- |
+-------------- Quadrulini
|
--+ +----------- Pleurobemini
| |
+--+ +-- Plectomerus
| +-----+
| | +-- Reginaia
+--+
| +----- Lampsilini
+--+
| +-- Popenaias **
+--+
+-- Amblema
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
7 |
2 |
17 |
17 |
3 |
7 |
+-------------- Quadrulini
--+
| +----------- Reginaia
+--+
| +-------- Amblema
+--+
| +----- Lampsilini
+--+
| +-- Popenaias **
+--+
+-- Psoronaias **
iT| tT | bi | riS | oT | roT | roS | iD |
| 5 |
5 |
3 |
2 |
18 |
18 |
3 |
8 |
+-------------- Quadrulini
--+
| +----------- Reginaia
+--+
| +-------- Amblema
+--+
| +----- Lampsilini
+--+
| +-- Popenaias **
+--+
+-- Psoronaias **
iT| tT | bi | riS | oT | roT | roS | iD |
| 3 |
3 |
2 |
2 |
28 |
27 |
4 |
3 |
+-------------- Quadrulini
--+
| +----------- Pleurobemini
+--+
| +-------- Amblemini
+--+
| +----- Lampsilini
+--+
| +-- Popenaias **
+--+
+-- Psoronaias **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
26 |
26 |
4 |
-- |
+-------------- Quadrulini
--+
| +----------- Pleurobemini
+--+
| +-------- Amblemini
+--+
| +----- Lampsilini
+--+
| +-- Popenaias **
+--+
+-- Psoronaias **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
9 |
9 |
4 |
-- |
+----- Lampsilini
+--+
| | +-- Nephronaias **
| +--+
--+ +-- Popenaias **
|
| +----- Amblemini
+--+
| +-- Pleurobemini
+--+
+-- Quadrulini
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
26 |
26 |
4 |
-- |
+-------------- Quadrulini
--+
| +----------- Pleurobemini
+--+
| +-------- Amblemini
+--+
| +----- Lampsilini
+--+
| +-- Popenaias **
+--+
+-- Psoronaias **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
10 |
10 |
4 |
-- |
+----- Lampsilini
+--+
| | +-- Nephronaias **
| +--+
--+ +-- Popenaias **
|
| +----- Amblemini
+--+
| +-- Pleurobemini
+--+
+-- Quadrulini
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
55 |
13 |
4 |
-- |
+-------------- Quadrulini
--+
| +----------- Pleurobemini
+--+
| +-------- Amblemini
+--+
| +----- Lampsilini
+--+
| +-- Psoronaias **
+--+
+-- Popenaias **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
10 |
10 |
4 |
-- |
+----- Amblemini
+--+
| | +-- Pleurobemini
| +--+
| +-- Quadrulini
--+
| +----- Obliquaria
| |
| +----- Toxolasma
| |
+--+ +-- Glebula
+--+
| +-- Cyrtonaias
|
| +-- Nephronaias **
+--+
+-- Popenaias **
iT| tT | bi | riS | oT | roT | roS | iD |
| 2 |
2 |
0 |
2 |
55 |
13 |
4 |
-- |
+----------- Quadrulini
|
--+ +-- Pleurobemini
| +-----+
| | +-- Amblemini
+--+
| +----- Lampsilini
+--+
| +-- Psoronaias **
+--+
+-- Popenaias **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
85 |
14 |
4 |
-- |
+----------- Quadrulini
--+
| +-------- Pleurobemini
+--+
| +----- Amblemini
+--+
| +-- Popenaias **
+--+
+-- Lampsilini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
91 |
16 |
4 |
-- |
+----------- Quadrulini
--+
| +-------- Pleurobemini
+--+
| +----- Amblemini
+--+
| +-- Popenaias **
+--+
+-- Lampsilini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
41 |
41 |
4 |
-- |
+-------------------- Quadrulini
|
--+ +----------------- Pleurobemini
| |
+--+ +-- Reginaia
| +-----------+
| | +-- Amblema
+--+
| +----------- Popenaias **
| |
+--+ +-------- Cyrtonaias
| |
+--+-------- Plectomerus
|
| +----- Lemiox
+--+
| +-- Lampsilis
+--+
+-- Obovaria
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
42 |
42 |
4 |
-- |
+-------------------------------- Quadrulini
|
| +-------------- Elliptoideus
| +--------------+
| | | +----------- Plectomerus
--+ | +--+
| | | +-------- Obovaria_unicolor
| | +--+
| | | +----- Obovaria_olivaria
| | +--+
| | | +-- Lampsilis
+--+ +--+
| +-- Lemiox
|
| +-------- Cyrtonaias
| +-----------------+
| | | +----- Reginaia
| | +--+
| | | +-- Popenaias **
+--+ +--+
| +-- Amblema
|
| +----------------------- Elliptio
| |
+--+ +-------------------- Eurynia
| |
| | +----------------- Parvaspina
+--+ |
| | +-- Hemistena
| +--------------+
+--+ +-- Plethobasus
|
| +-------------- Pleuronaia
| |
| | +-- Pleurobema_cordatum
+--+ +--------+
| | +-- Fusconaia
| |
+--+ +-- Pleurobema_stabile
| +-----+
| | +-- Pleurobema_sintoxia
+--+
| +----- Pleurobema_decisum
+--+
| +-- Pleurobema_clava
+--+
+-- Pleurobema_pyriforme
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
53 |
9 |
3 |
-- |
+-------- Quadrulini
--+
| +----- Pleurobemini
+--+
| +-- Popenaias **
+--+
+-- Lampsilini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
95 |
16 |
4 |
-- |
+----------- Quadrulini
--+
| +-------- Pleurobemini
+--+
| +----- Amblemini
+--+
| +-- Popenaias **
+--+
+-- Lampsilini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
20 |
19 |
4 |
-- |
+-------------- Quadrulini
--+
| +----------- Pleurobemini
+--+
| +-------- Amblema
+--+
| +----- Plectomerus
+--+
| +-- Popenaias **
+--+
+-- Lampsilini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
14 |
14 |
4 |
-- |
+----------- Toxolasma
+--+
| | +-------- Popenaias **
| +--+
| | +----- Amblemini
| +--+
--+ | +-- Quadrulini
| +--+
| +-- Pleurobemini
|
| +----------- Potamilus
| |
+--+ +-- Lampsilis_cardium
| +-----+
| | +-- Lampsilis_ornata
+--+
| +----- Venustaconcha
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Lampsilis_powellii
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
18 |
13 |
4 |
-- |
+----------- Quadrulini
--+
| +-------- Pleurobemini
+--+
| +----- Amblemini
+--+
| +-- Popenaias **
+--+
+-- Lampsilini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
55 |
53 |
3 |
-- |
+-------- Lampsilini
--+
| +----- Pleurobemini
+--+
| +-- Amblemini
+--+
+-- Popenaias **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
73 |
14 |
4 |
-- |
+----------- Quadrulini
--+
| +-------- Pleurobemini
+--+
| +----- Amblemini
+--+
| +-- Popenaias **
+--+
+-- Lampsilini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
52 |
11 |
4 |
-- |
+----------- Quadrulini
--+
| +-------- Pleurobemini
+--+
| +----- Amblemini
+--+
| +-- Popenaias **
+--+
+-- Lampsilini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
101 |
89 |
4 |
-- |
+-------------------------- Quadrulini
|
| +----------------------- Pleurobemini
--+ |
| | +----- Popenaias **
| | +--------------+
| | | | +-- Amblema
+--+ | +--+
| | +-- Reginaia
| |
| | +-- Plectomerus
| | +--------------+
+--+ | +-- Toxolasma
| |
| | +----- Obliquaria
| | +--------+
| | | | +-- Cyrtonaias
| | | +--+
| | | +-- Glebula
| | |
+--+ | +----------- Sagittunio
| | |
| | +----------- Leaunio
| | |
| | +----------- Potamilus
| | |
| | +----------- Hamiota
| | |
| | | +-- Ellipsaria
+--+ +--------+
| | +-- Truncilla
| |
| | +----- Lampsilis_teres
| +-----+
| | | +-- Ortmanniana_ligamentina
| | +--+
| | +-- Lampsilis_siliquoidea
| |
| | +----- Villosa
| +-----+
| | | +-- Ligumia
| | +--+
| | +-- Cambarunio
| |
+--+ +-------- Ortmanniana_pectorosa
+--+
| | +----- Lampsilis_ornata
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ovata
|
| +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha_ellipsiformis__2
+--+
| +-------- Epioblasma
| |
| | +----- Obovaria_unicolor
| +--+
| | +-- Obovaria_arkansasensis
| +--+
| +-- Obovaria_subrotunda
|
| +-- Ptychobranchus
| +-----+
| | +-- Venustaconcha_ellipsiformis__1
| |
+--+ +----- Medionidus_conradicus
| |
| | +-- Lemiox
+--+--+
| +-- Medionidus_acutissimus
|
| +-- Dromus
+--+
+-- Cyprogenia
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
41 |
41 |
4 |
-- |
+-------------------- Quadrulini
|
--+ +----------------- Pleurobemini
| |
| | +-- Popenaias **
+--+ +-----------+
| | +-- Amblema
| |
+--+ +-- Cyrtonaias
| +--------+
| | +-- Reginaia
+--+
| +-------- Plectomerus
+--+
| +----- Lemiox
+--+
| +-- Lampsilis
+--+
+-- Obovaria
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
73 |
14 |
4 |
-- |
+----------- Quadrulini
--+
| +-------- Pleurobemini
+--+
| +----- Amblemini
+--+
| +-- Popenaias **
+--+
+-- Lampsilini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
72 |
72 |
4 |
-- |
+----------------------------------- Quadrulini
|
--+ +-------------------------------- Pleurobemini
| |
| | +-- Amblema
+--+ +--------------------------+
| | +-- Reginaia
| |
+--+ +-------------------------- Popenaias **
| |
| | +-- Glebula
+--+ +--------------------+
| | +-- Paetulunio
| |
+--+ +-- Toxolasma
| +-----------------+
| | +-- Plectomerus
| |
+--+ +----------------- Obliquaria
| |
| | +----- Potamilus
+--+ +--------+
| | | +-- Dromus
| | +--+
+--+ +-- Lemiox
|
| +----------- Lampsilis_ovata
| |
+--+ +-------- Lampsilis_virescens
| |
+--+ +-- Epioblasma
| +--+
| | +-- Obovaria_arkansasensis
+--+
| +-- Obovaria_olivaria
+--+
+-- Venustaconcha
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
40 |
40 |
3 |
-- |
+----------- Amblema
|
--+ +-- Lampsilis
| +-----+
| | +-- Plectomerus
+--+
| +----- Popenaias **
+--+
| +-- Cyrtonaias
+--+
+-- Pleurobemini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
20 |
19 |
4 |
-- |
+-------------- Quadrulini
--+
| +----------- Pleurobemini
+--+
| +-------- Amblema
+--+
| +----- Plectomerus
+--+
| +-- Popenaias **
+--+
+-- Lampsilini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
87 |
15 |
4 |
-- |
+----------- Quadrulini
--+
| +-------- Pleurobemini
+--+
| +----- Amblemini
+--+
| +-- Popenaias **
+--+
+-- Lampsilini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
41 |
41 |
4 |
-- |
+----------------- Quadrulini
|
--+ +-------------- Pleurobemini
| |
| | +-- Popenaias **
+--+ +--------+
| | +-- Amblema
| |
+--+ +-------- Plectomerus
| |
| | +-- Cyrtonaias
+--+-----+
| +-- Reginaia
|
| +----- Lemiox
+--+
| +-- Lampsilis
+--+
+-- Obovaria
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
103 |
81 |
4 |
-- |
+-------- Reginaia
|
+-------- Amblema
|
| +-- Quadrulini
+-----+
| +-- Pleurobemini
--+
| +----- Popenaias **
| |
| | +-- Dromus
| | |
| | +-- Epioblasma
| | |
+--+ +-- Glebula
| |
| +-- Lampsilis_ovata
| |
| +-- Lampsilis_virescens
| |
| +-- Lemiox
+--+
+-- Obliquaria
|
+-- Obovaria_olivaria
|
+-- Obovaria_unicolor
|
+-- Plectomerus
|
+-- Potamilus_alatus
|
+-- Potamilus_capax
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
73 |
14 |
4 |
-- |
+----------- Quadrulini
--+
| +-------- Pleurobemini
+--+
| +----- Amblemini
+--+
| +-- Popenaias **
+--+
+-- Lampsilini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
91 |
16 |
4 |
-- |
+----------- Quadrulini
--+
| +-------- Pleurobemini
+--+
| +----- Amblemini
+--+
| +-- Popenaias **
+--+
+-- Lampsilini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
68 |
11 |
4 |
-- |
+----------- Quadrulini
--+
| +-------- Pleurobemini
+--+
| +----- Amblemini
+--+
| +-- Popenaias **
+--+
+-- Lampsilini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
32 |
32 |
4 |
-- |
+-------------------------- Quadrulini
|
| +----------------------- Plectomerus
--+ |
| | +-------------------- Popenaias **
| | |
| | +-------------------- Pleurobemini
+--+ |
| | +-- Cyrtonaias
| | +--+
| | | +-- Glebula
| +--------------+
| | | +-- Reginaia
+--+ +--+
| +-- Obliquaria
|
| +----------------- Toxolasma
| |
| | +-------------- Potamilus
| | |
| | | +-- Ellipsaria
| | +-----------+
+--+ | +-- Truncilla
| |
| | +----- Medionidus
| +--------+
| | | +-- Lemiox
| | +--+
+--+ +-- Dromus
|
| +----------- Cambarunio
| |
| +----------- Leaunio
| |
| +----------- Villosa
| |
| | +-- Sagittunio
| +--------+
+--+ +-- Lampsilis_ovata
|
| +----- Venustaconcha
+-----+
| | +-- Epioblasma
| +--+
| +-- Obovaria
|
| +-------- Lampsilis_teres
+--+
| +----- Lampsilis_siliquoidea
+--+
| +-- Lampsilis_hydiana
+--+
+-- Lampsilis_straminea
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
95 |
16 |
4 |
-- |
+----------- Quadrulini
--+
| +-------- Pleurobemini
+--+
| +----- Amblemini
+--+
| +-- Popenaias **
+--+
+-- Lampsilini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
52 |
14 |
4 |
-- |
+----------- Quadrulini
--+
| +-------- Pleurobemini
+--+
| +----- Amblemini
+--+
| +-- Popenaias **
+--+
+-- Lampsilini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
40 |
40 |
3 |
-- |
+-------------------------------------- Quadrulini
|
| +-- Amblema
--+ +--------------------------------+
| | +-- Reginaia
| |
+--+ +-------------------------------- Plectomerus
| |
| | +----------------------------- Toxolasma
+--+ |
| | +----- Popenaias **
| | +--------------------+
+--+ | | +-- Glebula
| | +--+
| | +-- Cyrtonaias
| |
+--+ +----------------------- Obliquaria
| |
| | +-- Medionidus
| | +--+
| | | +-- Lemiox
+--+ +--------------+
| | | +-- Cyprogenia
| | +--+
| | +-- Ptychobranchus
| |
+--+ +-------- Potamilus
| +--------+
| | | +----- Atlanticoncha
| | +--+
| | | +-- Ellipsaria
| | +--+
+--+ +-- Truncilla
|
| +-- Hamiota
| +-----------+
| | +-- Villosa
| |
+--+ +----------- Ligumia
| |
| | +----- Venustaconcha
+--+ +--+
| | | +-- Obovaria
| | +--+
+--+ +-- Epioblasma
|
| +----- Ortmanniana_pectorosa
+--+
| +-- Ortmanniana_ligamentina
+--+
+-- Lampsilis
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
26 |
25 |
4 |
-- |
+----------- Quadrulini
--+
| +-------- Pleurobemini
+--+
| +----- Amblemini
+--+
| +-- Lampsilini
+--+
+-- Popenaias **
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
56 |
54 |
3 |
-- |
+----- Pleurobemini
|
--+ +-- Amblemini
| |
+--+-- Popenaias **
|
+-- Lampsilini
iT| tT | bi | riS | oT | roT | roS | iD |
| 1 |
1 |
0 |
1 |
72 |
72 |
4 |
-- |
+-------------- Quadrulini
|
--+ +----------- Pleurobemini
| |
+--+ +-- Plectomerus
| +-----+
| | +-- Reginaia
+--+
| +----- Lampsilini
+--+
| +-- Popenaias **
+--+
+-- Amblema
|
