MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Sun 12 April 2026
Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae
tribe PSEUDODONTINI Frierson, 1927: cladomics
Cladomics pages are experimental — even more so than the rest of this site!
The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.
The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.
Following the summary, the available published clades representing the taxon are listed.
For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!
Cladomics Summary
number of species = 51 (52 total)
number of subtaxa (rank: subtribe) = 2
analyzed in 106 cladograms in 64 publications
maximum number of terminals in an analysis = 45 (mean = 8.52, median = 4)
maximum number of subtaxa (subtribe) = 2 (mean = 1.45, median = 1)
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 96
maximum number of included terminals = 45, (mean = 9.3 median = 5)
maximum number of subtaxa (subtribe) = 2 (mean = 1.5, median = 1.5)
proportion of cladograms that support basic monophyly = 0.99
Rigorous Monophyly
number of cladograms that test rigorous monophyly = 38
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (subtribe) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 45 (mean = 14.05, median = 11)
maximum number of subtaxa (subtribe) = 2 (mean = 2, median = 2)
proportion of cladograms that support rigorous monophyly = 0.97
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Cladograms
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14
** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **
| tT | bi | riS | oT | roT | roS | iD | |
| 45 | 582 | 651 | 2 | 587 | 90 | 6 | 0 |
+-------------------------------- Lamellidens
|
+-----+ +-- Balwantia
| | +--------------------------+
| | | +-- Trapezidens
| +--+
| | +-------------------------- Leoparreysiini
| | |
| +--+ +----------------------- Pseudomulleria
| | |
| | | +-- Parreysiini
| +--+ +-----------------+
| | | +-- Coelaturini
| | |
| +--+ +-- Pseudoscabies
| | +--------------+
| | | +-- Indonaia
| +--+
| | +-------------- Indochinella
| | |
| +--+ +----------- Radiatula
+--------------------------------------------------+ | |
| | +--+ +-- Songkhlanaia_songkhramensis__2 **
| | | +-----+
| | | | +-- Unionetta_fabagina__2
| | +--+
| | | +----- Harmandia
| | +--+
| | | +-- Unionetta_fabagina__1
| | +--+
| | +-- Scabies
| |
| | +----- Amuranodonta
| | +-----------------------------+
| | | | +-- Anemina
| | | +--+
| | | +-- Buldowskia
| | |
| | | +-- Beringiana
| | | +-----+
| | | | +-- Pletholophus
| +--+ +-----------------------+
| | | | +----- Mimo
| | | +--+
| | | | +-- Acudonta
| | | +--+
| | | +-- Sinanodonta
| | |
| | | +----------------------------- Cristaria
| +--+ |
| | | +----------------- Lepidodesmini
| | | |
| | | | +----------- Lanceolariini
| | | | +--+
| | | +--------+ | | +-------- Pseudoacuticosta_retiaria__2
| | | | | | +--+
| | | | | | | +----- Acuticosta_chinensis__2
| +--+ | | | +--+
| | | +--+ | +-- Acuticosta_chinensis__1
| | | | +--+
| | | | +-- Pseudoacuticosta_retiaria__1
| | | |
| | | | +----------- Globunio
| | | +--+
| | | | +-------- Aculamprotulina
| | | +--+
| +--+ | +----- Middendorffinaiina
| | +--+
| | | +-- Unionina
| | +--+
| | +-- Oxynaiina
| |
| | +-------- Pseudodontoideus
--+ | +--------------+
| | | | +----- Lasmigona
| | | +--+
| | | | +-- Pterosyna
| | | +--+
| +--+ +-- Arcidens
| |
| | +-------------------- Anodontina
| | |
| | | +-------------- Utterbackia
| | | +--+
| +--+ | | +----------- Pyganodon_grandis
| | | +--+
| | | | +-------- Pyganodon_lacustris
| | | +--+
| | | | +----- Pyganodon_cataracta__2
| +--+ +--+
| | | +-- Pyganodon_cataracta__1
| | +--+
| | +-- Potamilus_fragilis__3
| |
| | +-------------- Utterbackiana
| | |
| +--+ +----------- Simpsonaias
| | |
| | | +-- Alasminota
| +--+ +-----+
| | | +-- Alasmidonta
| | |
| +--+ +-- Prolasmidonta
| | +--+
| | | +-- Platynaias
| +--+
| | +-- Anodontoides
| +--+
| +-- Strophitus
|
| +-------------------------------------------------------------------------------------- Rectidentini
| |
| | +-------- Pseudunio
| | +--------------------------------------------------------------------------+
| | | | +----- Gibbosula
| | | +--+
| | | | +-- Cumberlandia
| | | +--+
| | | +-- Margaritifera
| | |
| | | +-------------------- Chamberlainiini
| | | |
| | | +--+ +----------------- Parvasolenaia
| | | | | |
| | | | +--+ +-------------- Pseudopostolata
| | | | | |
+--+ | | +--+ +----------- Koreosolenaia
| | | | |
| | | +--+ +-- Cosmopseudodon
| | | | +-----+
| | | | | +-- Guiunio
| | | +--+
| | +--------+ | +----- Ptychorhynchus
| | | | +--+
| | | | | +-- Obovalis
| | | | +--+
| | | | +-- Postolata
| | | |
| | | | +-- Caudiculatus
| | | | +--------------+
| | | | | +-- Lamprotulini
| | | | |
| | | +-----+ +-------------- Sinosolenaia_carinata
+--+ | | |
| | | | +----- Gonidea
| | +--+ +-----+
| +-----------------------------------------------+ | | | +-- Microcondylaea
| | | | | +--+
| | | +--+ +-- Leguminaia
| | | |
| | | | +-------- Inversidens
| | | +--+
| | | | +----- Sinosolenaia_recognita
| | | +--+
| | | | +-- Sinosolenaia_iridinea
| | | +--+
| | | +-- Sinosolenaia_oleivora
| | |
| | | +----------------------------- Indopseudodontina **
| | | |
| | | | +-- Lannanaia **
| | +--+ +-----------------------+
| | | | +-- Pilsbryoconcha **
| | | |
| | | | +----- Songkhlanaia_tamodienica **
| | +--+ +-----------------+
| | | | | +-- Songkhlanaia_moreleti **
| | | | +--+
| | | | +-- Songkhlanaia_songkhramensis__1 **
| | | |
| | +--+ +-- Nyeinchanconcha **
| | | +-----------------+
| | | | +-- Sundadontina **
| | | |
+--+ | | +-- Isannaia **
| +--+ +--+
| | | +-- Namkongnaia **
| | +-----------+
| | | | +-- Thaiconcha **
| | | +--+
| +--+ +-- Bineurus **
| |
| | +-------------- Pseudodon_walpolei **
| | |
| +--+ +----------- Pseudodon_vagulus **
| | |
| | | +----- Pseudodon_vondembuschianus **
| +--+ +--+
| | | | +-- Pseudodon_mekongi **
| | | +--+
| +--+ +-- Schepmaniini
| |
| | +----- Pseudodon_lenyanensis__2 **
| +--+
| | +-- Pseudodon_inoscularis **
| +--+
| +-- Pseudodon_lenyanensis__1 **
|
| +-- Quadrulini
| +--------------------------------------------------------------------------+
| | +-- Pleurobemini
| |
| | +----- Amblema
+--+ +--------------------------------------------------------------------+
| | | +-- Nephronaias
| | +--+
| | +-- Psoronaias
| |
+--+ +----------------------------------------------------------------------- Reginaia
| |
| | +----- Popenaias
| | +--------------------------------------------------------------+
| | | | +-- Cyrtonaias
+--+ | +--+
| | +-- Glebula
| |
| | +----------------------------------------------------------------- Paetulunio
| | |
+--+ | +----------------- Toxolasma_parvum
| | |
| | | +----- Toxolasma_lividum__2
| | | +--------+
| | +--------------------------------------------+ | | +-- Toxolasma_cylindrellus
| | | | | +--+
| | | | | +-- Toxolasma_lividum__1
| | | | |
+--+ | +--+ +-------- Toxolasma_texasiense__2
| | | +--+
| | | | | +----- Toxolasma_glans
| | | | +--+
| | | | | +-- Toxolasma_paulum
| | +--+ +--+
| | | +-- Toxolasma_texasiense__1
| | |
| | | +-------- Toxolasma__4
| | +--+
+--+ | +----- Toxolasma_granulatum
| +--+
| | +-- Toxolasma__2
| +--+
| +-- Toxolasma__3
|
| +----- Toxolasma_pullus
| +-----------------------------------------------------+
| | | +-- Toxolasma_germanum
| | +--+
| | +-- Toxolasma__1
| |
| | +-- Obliquaria
| | +-----------------------------------------------------+
| | | +-- Plectomerus
+--+ |
| | +-------- Potamilus_inflatus
| | +--------+
| | | | +----- Potamilus_ohiensis
| | | +--+
| | | | +-- Potamilus_amphichaenus
| | | +--+
| | +-----------------------------------+ +-- Potamilus_streckersoni
| | | |
+--+ | | +----- Potamilus_fragilis__2
| | | +--------+
| | | | | +-- Potamilus_leptodon
| | | | +--+
| | +--+ +-- Potamilus_fragilis__1
| | |
| | | +----------- Potamilus_metnecktayi
| | | |
| | +--+ +-------- Potamilus_alatus__3
| | | |
| | +--+ +----- Potamilus_alatus__2
+--+ | |
| +--+----- Potamilus_purpuratus__2
| |
| | +-- Potamilus_alatus__1
| +--+
| +-- Potamilus_purpuratus__1
|
| +-- Villosa_villosa
| +-----------------------------------------------+
| | +-- Atlanticoncha
| |
| | +-- Cambarunio_nebulosus
| | +--------------------------------------------+
+--+ | +-- Ligumia
| |
| | +-------- Leaunio_umbrans
| | +-----------------------------------+
| | | | +----- Cambarunio_taeniatus
+--+ | +--+
| | | +-- Cambarunio_iris
| | +--+
| | +-- Cambarunio_simus
| |
| | +----------------------------------------- Hamiota
| | |
+--+ | +----------------------- Pachynaias
| | |
| | +--------------+ +-------------------- Sagittunio
| | | | |
| | | | | +----- Ellipsaria
| | | +--+ +-----------+
| | | | | | +-- Potamilus_capax
| | | | | +--+
| | | | | +-- Truncilla
+--+ | +--+
| | | +----- Medionidus_acutissimus
| | | +--------+
| | | | | +-- Lemiox
| | | | +--+
| | +--+ +-- Medionidus_penicillatus
| | |
| | | +----------- Ptychobranchus
| | | |
| | +--+ +-- Cyprogenia_aberti__2
| | | +-----+
+--+ | | +-- Dromus
| +--+
| | +----- Cyprogenia_stegaria__2
| +--+
| | +-- Cyprogenia_aberti__1
| +--+
| +-- Cyprogenia_stegaria__1
|
| +-- Villosa_delumbis
| +--------------------------------+
| | +-- Villosa_vibex
| |
| | +-------------- Obovaria_olivaria
| | |
| | +-----------------+ +-- Venustaconcha
| | | | +--------+
| | | | | +-- Epioblasma
| | | +--+
| | | | +-------- Obovaria_retusa
+--+ | +--+
| | | +----- Obovaria_choctawensis
| | +--+
| | | +-- Obovaria_unicolor
| | +--+
| | +-- Obovaria_subrotunda
| |
| | +-- Mesonaias
| | +-----------------------+
| | | +-- Obovaria_arkansasensis
| | |
+--+ +--+ +-- Lampsilis_splendida
| | | +--------------------+
| | | | +-- Lampsilis_bracteata
| | | |
| | +--+ +-- Lampsilis_fasciola
| | | +-----------------+
| | | | +-- Ortmanniana_pectorosa
| | +--+
| | | +----------------- Lampsilis_ornata
| | | |
| | +--+ +-------------- Lampsilis_ovata__2
| | | |
| | +--+ +----------- Lampsilis_dolabraeformis
| | | |
| | | | +----- Lampsilis_cardium__2
| | +--+ +--+
| | | | | +-- Lampsilis_cardium__1
+--+ | | +--+
| +--+ +-- Lampsilis_satura
| |
| | +----- Lampsilis_cariosa__2
| +--+
| | +-- Lampsilis_ovata__1
| +--+
| +-- Lampsilis_cariosa__1
|
| +----------- Ortmanniana_ligamentina__2
| +--------------+
| | | +-------- Lampsilis_abrupta
| | +--+
| | | +----- Lampsilis_higginsii__2
| | +--+
| | | +-- Ortmanniana_ligamentina__1
| | +--+
+--+ +-- Lampsilis_higginsii__1
|
| +----- Medionidus_conradicus
| +-----------------+
| | | +-- Leaunio_lienosus
| | +--+
| | +-- Leaunio_vanuxemensis
| |
+--+ +-- Lampsilis_floridensis
| +-----------------+
| | +-- Lampsilis_teres
| |
| | +-- Lampsilis_siliquoidea__1
+--+ |
| +-- Lampsilis_siliquoidea__2
| +--------------+
| | +-- Lampsilis_powellii__1
| | |
+--+ +-- Lampsilis_powellii__2
|
| +-------------- Lampsilis_sietmani
| |
+--+ +-- Lampsilis_straminea__2
| +--------+
| | +-- Lampsilis_virescens
+--+
| +-------- Lampsilis_hydiana__2
+--+
| +----- Lampsilis_hydiana__1
+--+
| +-- Lampsilis_bergmanni
+--+
+-- Lampsilis_straminea__1
Konopleva et al., 2025 [Publ 4446], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 44 | 44 | 42 | 2 | 0 | 0 | 0 | 684 |
+-- Indopseudodontina ** --+ +-- Pseudodontina **
Jeratthitikul et al., 2025 [Publ 4423], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 43 | 43 | 41 | 2 | 7 | 7 | 3 | 680 |
+-- Lamprotulini
+-----+
| +-- Gonideini
--+
| +----- Schepmaniini
+--+
| +-- Indopseudodontina **
+--+
+-- Pseudodontina **
Jeratthitikul et al., 2024 [Publ 4387], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 41 | 41 | 39 | 2 | 18 | 18 | 5 | 680 |
+-- Chamberlainiini
+-----------+
| +-- Rectidentini
--+
| +----------- Lamprotulini
+--+
| +-------- Gonideini
+--+
| +----- Schepmaniini
+--+
| +-- Indopseudodontina **
+--+
+-- Pseudodontina **
Bolotov et al., 2023 [Publ 4312], fig. S2
| tT | bi | riS | oT | roT | roS | iD | |
| 40 | 40 | 38 | 2 | 0 | 0 | 0 | 677 |
+-- Indopseudodontina ** --+ +-- Pseudodontina **
Bolotov et al., 2023 [Publ 4312], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 38 | 38 | 36 | 2 | 0 | 0 | 0 | 680 |
+-- Indopseudodontina ** --+ +-- Pseudodontina **
Bolotov et al., 2023 [Publ 4312], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 38 | 38 | 36 | 2 | 0 | 0 | 0 | 676 |
+-- Indopseudodontina ** --+ +-- Pseudodontina **
Konopleva et al., 2023 [Publ 4262], fig. 8
| tT | bi | riS | oT | roT | roS | iD | |
| 29 | 29 | 28 | 2 | 53 | 39 | 3 | 680 |
+-- Indopseudodontina **
+-----+
| +-- Pseudodontina **
--+
| +----- Rectidentini
+--+
| +-- Gonideini
+--+
+-- Lamprotulini
Konopleva et al., 2021 [Publ 4213], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 22 | 22 | 20 | 2 | 4 | 4 | 2 | 682 |
+----- Lamprotula | +----- Leguminaia | --+----- Potomida | +----- Gonidea | | +-- Indopseudodontina ** +--+ +-- Pseudodontina **
Jeratthitikul et al., 2021 [Publ 4219], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 21 | 21 | 19 | 2 | 14 | 14 | 4 | 682 |
+----------- Rectidentini | --+ +-- Indopseudodontina ** | +-----+ | | +-- Pseudodontina ** +--+ | +----- Gonideini +--+ | +-- Chamberlainiini +--+ +-- Lamprotulini
Bolotov et al., 2020 [Publ 4100], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 19 | 2 | 91 | 51 | 4 | 681 |
+----------- Rectidentini | --+ +-- Indopseudodontina ** | +-----+ | | +-- Pseudodontina ** +--+ | +----- Gonideini +--+ | +-- Chamberlainiini +--+ +-- Lamprotulini
Jeratthitikul et al., 2021 [Publ 4219], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 18 | 2 | 14 | 14 | 4 | 681 |
+----------- Rectidentini | --+ +-- Indopseudodontina ** | +-----+ | | +-- Pseudodontina ** +--+ | +----- Gonideini +--+ | +-- Chamberlainiini +--+ +-- Lamprotulini
Bolotov et al., 2020 [Publ 4100], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 19 | 2 | 97 | 51 | 4 | 683 |
+-------- Rectidentini | | +-- Indopseudodontina ** | +--+ | | +-- Pseudodontina ** | | --+ | +-- Sinosolenaia_iridinea | | | | | +-- Sinosolenaia_recognita__1 | | | | | +-- Sinosolenaia_carinata | | | +--+ +-- Leguminaia | | | +-- Microcondylaea | | | +-- Gonidea | | | +-- Ptychorhynchus | | | +-- Parvasolenaia | | +--+-- Lamprotula_leaii | +-- Lamprotula | +-- Lamprotula_caveata | +-- Potomida | +-- Pronodularia | +-- Sinohyriopsis | +-- Chamberlainia | +-- Sinosolenaia_recognita__2 | +-- Sinosolenaia_oleivora
Dai et al., 2023 [Publ 4285], fig. 3-BI
| tT | bi | riS | oT | roT | roS | iD | |
| 17 | 17 | 14 | 2 | 28 | 26 | 4 | 679 |
+-- Indopseudodontina **
+--------------------+
| +-- Pseudodontina **
|
--+ +-------------------- Rectidentini
| |
| | +----------------- Inversidens
+--+ |
| | +-- Chamberlainiini
| +--------------+
+--+ +-- Lamprotulini
|
| +-------------- Sinosolenaia
| |
| | +----- Gonidea
+--+ +-----+
| | | +-- Leguminaia
| | +--+
| | +-- Microcondylaea
+--+
| +-- Koreosolenaia
| +-----+
| | +-- Parvasolenaia
+--+
| +----- Postolata
+--+
| +-- Ptychorhynchus
+--+
+-- Obovalis
Dai et al., 2023 [Publ 4285], fig. 3-ML
| tT | bi | riS | oT | roT | roS | iD | |
| 17 | 17 | 15 | 2 | 28 | 26 | 4 | 680 |
+-- Indopseudodontina **
+--------------------+
| +-- Pseudodontina **
|
--+ +-------------------- Rectidentini
| |
| | +----- Inversidens
+--+ +-----------+
| | | +-- Chamberlainiini
| | +--+
| | +-- Lamprotulini
+--+
| +-------------- Sinosolenaia
| |
| | +----- Gonidea
+--+ +-----+
| | | +-- Leguminaia
| | +--+
| | +-- Microcondylaea
+--+
| +-- Koreosolenaia
| +-----+
| | +-- Parvasolenaia
+--+
| +----- Postolata
+--+
| +-- Ptychorhynchus
+--+
+-- Obovalis
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1
| tT | bi | riS | oT | roT | roS | iD | |
| 17 | 17 | 15 | 2 | 59 | 14 | 4 | 678 |
+-- Indopseudodontina ** --+ +-- Pseudodontina **
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2
| tT | bi | riS | oT | roT | roS | iD | |
| 17 | 17 | 15 | 2 | 59 | 14 | 4 | 678 |
+-- Indopseudodontina ** --+ +-- Pseudodontina **
Jeratthitikul et al., 2025 [Publ 4476], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 13 | 1 | 0 | 0 | 0 | 684 |
1
Huang et al., 2019 [Publ 3843], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 14 | 2 | 186 | 41 | 4 | 681 |
+----------- Chamberlainiini --+ | +-------- Lamprotulini +--+ | +----- Gonideini +--+ | +-- Indopseudodontina ** +--+ +-- Pseudodontina **
Huang et al., 2019 [Publ 3843], fig. S3
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 14 | 2 | 184 | 40 | 4 | 681 |
+-- Indopseudodontina ** --+ +-- Pseudodontina **
Huang et al., 2019 [Publ 3843], fig. S4
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 8 | 2 | 186 | 41 | 4 | 682 |
+-- Indopseudodontina ** --+ +-- Pseudodontina **
Bolotov et al., 2017 [Publ 3764], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 2 | 110 | 36 | 4 | 685 |
+-- Indopseudodontina ** --+ +-- Pseudodontina **
Bolotov et al., 2017 [Publ 3761], fig. S2
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 12 | 2 | 125 | 35 | 4 | 689 |
+----- Gonideini
+-----------+
| | +-- Chamberlainiini
| +--+
| +-- Lamprotulini
--+
| +-------------- Pilsbryoconcha **
| |
| | +----------- Indopseudodontina **
+--+ |
| | +-- Thaiconcha **
| +--------+
| | +-- Bineurus **
+--+
| +-- Pseudodon_mekongi **
| +-----+
| | +-- Pseudodon_vondembuschianus **
| |
+--+ +-- Pseudodon_inoscularis **
| +--+
| | +-- Sundadontina_tumida **
+--+
| +-- Nyeinchanconcha **
+--+
+-- Sundadontina_cumingii **
Jeratthitikul & Sutcharit, 2023 [Publ 4289], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 2 | 0 | 0 | 0 | 690 |
+-- Indopseudodontina ** --+ +-- Pseudodontina **
Bolotov et al., 2017 [Publ 3761], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 12 | 2 | 126 | 36 | 4 | 685 |
+-- Indopseudodontina ** --+ +-- Pseudodontina **
Dai et al., 2025 [Publ 4441], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 10 | 1 | 36 | 31 | 5 | 683 |
+-------------- Rectidentini --+ | +----------- Chamberlainiini +--+ | +-------- Lamprotulini +--+ | +----- Gonideini +--+ | +-- Schepmaniini +--+ +-- Pseudodontina **
Chen et al., 2025 [Publ 4663], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 10 | 1 | 37 | 32 | 5 | 683 |
+-------------- Rectidentini --+ | +----------- Chamberlainiini +--+ | +-------- Lamprotulini +--+ | +----- Gonideini +--+ | +-- Schepmaniini +--+ +-- Pseudodontina **
Dai et al., 2024 [Publ 4389], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 9 | 1 | 35 | 30 | 5 | 682 |
+-------------- Rectidentini --+ | +----------- Chamberlainiini +--+ | +-------- Lamprotulini +--+ | +----- Gonideini +--+ | +-- Schepmaniini +--+ +-- Pseudodontina **
Bolotov et al., 2018 [Publ 3826], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 2 | 2 | 72 | 24 | 3 | 686 |
+-- Indopseudodontina ** --+ +-- Pseudodontina **
Konopleva et al., 2019 [Publ 3900], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 2 | 2 | 41 | 27 | 1 | 686 |
+-- Indopseudodontina ** --+ +-- Pseudodontina **
Zieritz et al., 2024 [Publ 4381], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 2 | 36 | 25 | 6 | 684 |
+----------- Rectidentini | | +----- Caudiculatus --+ +--+ | | | +-- Lamprotulini | | +--+ +--+ +-- Gonideini | | +----- Schepmaniini +--+ | +-- Indopseudodontina ** +--+ +-- Pseudodontina **
Zieritz et al., 2026 [Publ 4812], fig. 4a
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 1 | 0 | 0 | 0 | 684 |
1
Zieritz et al., 2021 [Publ 4218], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 6 | 2 | 23 | 19 | 5 | 683 |
+----------------- Chamberlainiini --+ | +-------------- Lamprotulini +--+ | +----------- Rectidentini +--+ | +-------- Gonideini +--+ | +----- Schepmaniini +--+ | +-- Indopseudodontina ** +--+ +-- Pseudodontina **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 5 | 2 | 114 | 25 | 4 | 684 |
+-- Chamberlainiini
+-----+
| +-- Lamprotulini
--+
| +----- Gonideini
+--+
| +-- Indopseudodontina **
+--+
+-- Pseudodontina **
Zieritz et al., 2026 [Publ 4812], fig. 5a
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 8 | 1 | 0 | 0 | 0 | 689 |
1
Wu et al., 2024 [Publ 4350], fig. 5b
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 2 | 29 | 27 | 5 | 684 |
+----------------------------- Schepmaniini | --+ +-- Indopseudodontina ** | +-----------------------+ | | +-- Pseudodontina ** +--+ | +----------------------- Rectidentini | | +--+ +-------------------- Inversidens | | | | +----- Lamprotula +--+ +-----------+ | | | +-- Chamberlainiini | | +--+ +--+ +-- Potomida | | +-------------- Sinosolenaia | | +--+ +-- Gonidea | +--------+ | | +-- Microcondylaea +--+ | +-------- Koreosolenaia +--+ | +----- Parvasolenaia +--+ | +-- Postolata +--+ +-- Ptychorhynchus
Wu et al., 2024 [Publ 4350], fig. 5c
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 4 | 2 | 29 | 27 | 5 | 685 |
+----------------------------- Schepmania_nieuwenhuisi__1 | +----------------------------- Schepmania_nieuwenhuisi__2 --+ | +-- Indopseudodontina ** | +-----------------------+ | | +-- Pseudodontina ** +--+ | +----------------------- Rectidentini | | +--+ +-------------------- Inversidens | | | | +----- Lamprotula +--+ +-----------+ | | | +-- Chamberlainiini | | +--+ +--+ +-- Potomida | | +-------------- Sinosolenaia | | +--+ +----------- Gonidea | | +--+----------- Microcondylaea | | +-------- Koreosolenaia +--+ | +----- Parvasolenaia +--+ | +-- Postolata +--+ +-- Ptychorhynchus
Wu et al., 2024 [Publ 4388], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 2 | 29 | 27 | 5 | 682 |
+----------------- Chamberlainiini --+ | +-------------- Rectidentini +--+ | +----------- Lamprotulini +--+ | +-------- Gonideini +--+ | +----- Schepmaniini +--+ | +-- Indopseudodontina ** +--+ +-- Pseudodontina **
Zieritz et al., 2024 [Publ 4381], fig. 8A
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 1 | 0 | 0 | 0 | 688 |
1
Wu et al., 2024 [Publ 4350], fig. 6-7
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 2 | 29 | 27 | 5 | 682 |
+-------------- Chamberlainiini | --+ +----------- Rectidentini | | +--+ +-- Lamprotulini | +-----+ | | +-- Gonideini +--+ | +----- Schepmaniini +--+ | +-- Indopseudodontina ** +--+ +-- Pseudodontina **
Liu et al., 2024 [Publ 4380], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 2 | 29 | 27 | 5 | 682 |
+----- Schepmaniini
+-----+
| | +-- Indopseudodontina **
| +--+
--+ +-- Pseudodontina **
|
| +-------- Gonideini
+--+
| +----- Rectidentini
+--+
| +-- Chamberlainiini
+--+
+-- Lamprotulini
Wu et al., 2024 [Publ 4350], fig. 5a
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 2 | 29 | 27 | 5 | 682 |
+-------------------------- Chamberlainiini | --+ +----------------------- Rectidentini | | | | +----- Schepmaniini +--+ +--------------+ | | | +-- Indopseudodontina ** | | +--+ | | +-- Pseudodontina ** +--+ | +-- Inversidens | +--------------+ | | +-- Lamprotulini +--+ | +-------------- Sinosolenaia | | +--+ +-- Gonidea | +--------+ | | +-- Microcondylaea +--+ | +-------- Parvasolenaia +--+ | +----- Koreosolenaia +--+ | +-- Postolata +--+ +-- Ptychorhynchus
Konopleva et al., 2017 [Publ 3807], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 1 | 25 | 4 | 1 | 689 |
+-- Rectidentini --+ +-- Pseudodontina **
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 1 | 34 | 8 | 3 | 687 |
+-------- Rectidentini --+ | +----- Pseudodontina ** +--+ | +-- Gonideini +--+ +-- Lamprotulini
Bolotov et al., 2022 [Publ 4250], fig. 8
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 2 | 43 | 17 | 3 | 683 |
+----- Rectidentini --+ | +-- Pseudodontina ** +--+ +-- Indopseudodontina **
Wu et al., 2021 [Publ 4366], fig. 2_I
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 2 | 85 | 23 | 4 | 685 |
+----------- Chamberlainiini --+ | +-------- Lamprotulini +--+ | +----- Gonideini +--+ | +-- Indopseudodontina ** +--+ +-- Pseudodontina **
Wu et al., 2021 [Publ 4366], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 2 | 85 | 23 | 4 | 685 |
+----------- Chamberlainiini --+ | +-------- Lamprotulini +--+ | +----- Gonideini +--+ | +-- Indopseudodontina ** +--+ +-- Pseudodontina **
Pfeiffer et al., 2019 [Publ 3937], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 2 | 52 | 12 | 4 | 686 |
+----------- Chamberlainiini | --+ +-------- Rectidentini | | +--+ +-- Gonideini | +--+ | | +-- Lamprotulini +--+ | +-- Indopseudodontina ** +--+ +-- Pseudodontina **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 75 | 23 | 4 | 686 |
+-- Chamberlainiini
+-----------+
| +-- Lamprotulini
|
--+ +-- Parvasolenaia
| +--------+
| | +-- Pseudodontina **
+--+
| +-------- Sinosolenaia
+--+
| +----- Gonidea
+--+
| +-- Microcondylaea
+--+
+-- Leguminaia
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 2 | 52 | 12 | 4 | 688 |
+-------------- Chamberlainiini --+ | +----------- Rectidentini +--+ | +-------- Lamprotulini +--+ | +----- Gonideini +--+ | +-- Indopseudodontina ** +--+ +-- Pseudodontina **
Bolotov et al., 2022 [Publ 4239], fig. S2
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 0 | 2 | 54 | 0 | 0 | 687 |
+-- Pseudodon ** | +-- Nyeinchanconcha ** --+ +-- Indopseudodontina ** | +-- Sundadontina **
Bolotov et al., 2022 [Publ 4252], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 0 | 2 | 57 | 0 | 0 | 687 |
+-- Pseudodon ** | +-- Nyeinchanconcha ** --+ +-- Indopseudodontina ** | +-- Sundadontina **
Bolotov et al., 2022 [Publ 4252], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 0 | 2 | 57 | 0 | 0 | 687 |
+-- Pseudodon ** | +-- Nyeinchanconcha ** --+ +-- Indopseudodontina ** | +-- Sundadontina **
Bolotov et al., 2022 [Publ 4239], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 0 | 2 | 54 | 0 | 0 | 687 |
+-- Pseudodon ** | +-- Nyeinchanconcha ** --+ +-- Indopseudodontina ** | +-- Sundadontina **
Bolotov et al., 2022 [Publ 4239], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 0 | 2 | 54 | 0 | 0 | 687 |
+-- Pseudodon ** | +-- Nyeinchanconcha ** --+ +-- Indopseudodontina ** | +-- Sundadontina **
Wu et al., 2021 [Publ 4215], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 2 | 20 | 15 | 3 | 685 |
+-------------------- Chamberlainiini | | +-- Indopseudodontina ** --+ +--------------+ | | +-- Pseudodontina ** | | +--+ +-- Sinosolenaia | +-----------+ | | +-- Lamprotulini | | +--+ +-- Ptychorhynchus | +--------+ | | +-- Parvasolenaia +--+ | +-------- Inversidens +--+ | +----- Gonidea +--+ | +-- Microcondylaea +--+ +-- Leguminaia
Kongim et al., 2023 [Publ 4290], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 2 | 12 | 12 | 3 | 686 |
+----------- Chamberlainiini --+ | +-------- Lamprotulini +--+ | +----- Gonideini +--+ | +-- Indopseudodontina ** +--+ +-- Pseudodontina **
Konopleva et al., 2019 [Publ 4041], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 0 | 1 | 31 | 31 | 1 | 687 |
+-- Indopseudodontina ** --+ +-- Rectidentini
Zieritz et al., 2020 [Publ 4102], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 29 | 29 | 4 | -- |
+-------------- Chamberlainiini --+ | +----------- Lamprotulini +--+ | +-------- Gonideini +--+ | +----- Rectidentini +--+ | +-- Pseudodontina ** +--+ +-- Indopseudodontina **
Hoeh et al., 2009 [Publ 2716], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 44 | 2 | 2 | -- |
1
Wu et al., 2022 [Publ 4238], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 72 | 12 | 3 | -- |
+-------- Chamberlainiini --+ | +----- Lamprotulini +--+ | +-- Pseudodontina ** +--+ +-- Gonideini
Matos et al., 2024 [Publ 4383], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 18 | 16 | 4 | -- |
+----------- Chamberlainiini --+ | +-------- Rectidentini +--+ | +----- Lamprotulini +--+ | +-- Pseudodontina ** +--+ +-- Gonideini
Dai et al., 2023 [Publ 4285], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 51 | 17 | 4 | -- |
+----------- Chamberlainiini --+ | +-------- Rectidentini +--+ | +----- Lamprotulini +--+ | +-- Pseudodontina ** +--+ +-- Gonideini
Klymus et al., 2025 [Publ 4814], fig. 3B
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 34 | 12 | 4 | -- |
+----------- Chamberlainiini --+ | +-------- Rectidentini +--+ | +----- Pseudodontina ** +--+ | +-- Lamprotulini +--+ +-- Gonideini
Hoeh et al., 2009 [Publ 2716], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 45 | 2 | 2 | -- |
+-- Gonideini --+ +-- Pseudodontina **
Hoeh et al., 2009 [Publ 2716], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 44 | 2 | 2 | -- |
1
Wu et al., 2024 [Publ 4388], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 51 | 20 | 4 | -- |
+----------- Chamberlainiini --+ | +-------- Rectidentini +--+ | +----- Lamprotulini +--+ | +-- Pseudodontina ** +--+ +-- Gonideini
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 37 | 10 | 3 | -- |
+-------- Chamberlainiini --+ | +----- Gonideini +--+ | +-- Pseudodontina ** +--+ +-- Lamprotulini
Klymus et al., 2025 [Publ 4814], fig. 3A
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 34 | 12 | 4 | -- |
+----------- Chamberlainiini --+ | +-------- Rectidentini +--+ | +----- Pseudodontina ** +--+ | +-- Lamprotulini +--+ +-- Gonideini
Wu et al., 2021 [Publ 4366], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 67 | 11 | 3 | -- |
+-------- Chamberlainiini --+ | +----- Lamprotulini +--+ | +-- Pseudodontina ** +--+ +-- Gonideini
Wu et al., 2024 [Publ 4350], fig. 8b_S2
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 90 | 19 | 4 | -- |
+----------- Chamberlainiini --+ | +-------- Rectidentini +--+ | +----- Lamprotulini +--+ | +-- Pseudodontina ** +--+ +-- Gonideini
Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 139 | 24 | 4 | -- |
+----------- Chamberlainiini --+ | +-------- Rectidentini +--+ | +----- Lamprotulini +--+ | +-- Pseudodontina ** +--+ +-- Gonideini
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 29 | 9 | 4 | -- |
+-------- Chamberlainiini | --+ +-- Gonideini | +--+ | | +-- Pseudodontina ** +--+ | +-- Rectidentini +--+ +-- Lamprotulini
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 94 | 18 | 4 | -- |
+----------- Chamberlainiini --+ | +-------- Rectidentini +--+ | +----- Lamprotulini +--+ | +-- Pseudodontina ** +--+ +-- Gonideini
Graf & Cummings, 2006 [Publ 1968], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 45 | 2 | 2 | -- |
1
Dai et al., 2024 [Publ 4358], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 51 | 9 | 3 | -- |
+-------- Chamberlainiini --+ | +----- Lamprotulini +--+ | +-- Pseudodontina ** +--+ +-- Gonideini
Fukata & Iigo, 2020 [Publ 4106], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 52 | 10 | 3 | -- |
+-------- Chamberlainiini --+ | +----- Pseudodontina ** +--+ | +-- Lamprotulini +--+ +-- Gonideini
Matos et al., 2024 [Publ 4384], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 18 | 16 | 4 | -- |
+----------- Chamberlainiini --+ | +-------- Rectidentini +--+ | +----- Lamprotulini +--+ | +-- Pseudodontina ** +--+ +-- Gonideini
Wu et al., 2026 [Publ 4811], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 94 | 26 | 4 | -- |
+----------- Chamberlainiini --+ | +-------- Rectidentini +--+ | +----- Lamprotulini +--+ | +-- Pseudodontina ** +--+ +-- Gonideini
Wu et al., 2021 [Publ 4366], fig. 3-BI
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 72 | 12 | 3 | -- |
+-------- Chamberlainiini --+ | +----- Lamprotulini +--+ | +-- Pseudodontina ** +--+ +-- Gonideini
Wu et al., 2024 [Publ 4350], fig. 8a_S1
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 90 | 19 | 4 | -- |
+----------- Chamberlainiini --+ | +-------- Rectidentini +--+ | +----- Lamprotulini +--+ | +-- Pseudodontina ** +--+ +-- Gonideini
Zhang et al., 2024 [Publ 4397], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 25 | 23 | 4 | -- |
+----------- Chamberlainiini --+ | +-------- Rectidentini +--+ | +----- Lamprotulini +--+ | +-- Pseudodontina ** +--+ +-- Gonideini
Dai et al., 2025 [Publ 4440], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 86 | 18 | 4 | -- |
+----------- Chamberlainiini --+ | +-------- Rectidentini +--+ | +----- Lamprotulini +--+ | +-- Pseudodontina ** +--+ +-- Gonideini
Froufe et al., 2019 [Publ 3936], fig. 3-M
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 17 | 6 | 3 | -- |
+-------- Chamberlainiini --+ | +----- Lamprotulini +--+ | +-- Gonideini +--+ +-- Pseudodontina **
Chen et al., 2020 [Publ 4170], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 32 | 9 | 3 | -- |
+----- Pseudodontina ** --+ | +-- Lamprotulini +--+ +-- Chamberlainiini
Chen et al., 2025 [Publ 4448], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 84 | 16 | 4 | -- |
+----------- Chamberlainiini --+ | +-------- Rectidentini +--+ | +----- Lamprotulini +--+ | +-- Pseudodontina ** +--+ +-- Gonideini
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 35 | 9 | 3 | -- |
+-------- Chamberlainiini --+ | +----- Lamprotulini +--+ | +-- Pseudodontina ** +--+ +-- Gonideini
Zieritz et al., 2021 [Publ 4231], fig. 3-M_4
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 21 | 10 | 4 | -- |
+----------- Chamberlainiini --+ | +-------- Rectidentini +--+ | +----- Pseudodontina ** +--+ | +-- Lamprotulini +--+ +-- Gonideini
Graf, 2002 [Publ 2651], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 16 | 2 | 2 | -- |
1
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 94 | 18 | 4 | -- |
+----------- Chamberlainiini --+ | +-------- Rectidentini +--+ | +----- Lamprotulini +--+ | +-- Pseudodontina ** +--+ +-- Gonideini
Froufe et al., 2019 [Publ 3936], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 42 | 11 | 3 | -- |
+-------- Chamberlainiini --+ | +----- Lamprotulini +--+ | +-- Pseudodontina ** +--+ +-- Gonideini
Campbell & Lydeard, 2026 [Publ 4810], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 45 | 13 | 4 | -- |
+----------- Chamberlainiini --+ | +-------- Rectidentini +--+ | +----- Pseudodontina ** +--+ | +-- Lamprotulini +--+ +-- Gonideini
Froufe et al., 2019 [Publ 3936], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 42 | 11 | 3 | -- |
+-------- Chamberlainiini --+ | +----- Lamprotulini +--+ | +-- Pseudodontina ** +--+ +-- Gonideini
Froufe et al., 2019 [Publ 3936], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 17 | 6 | 3 | -- |
+-------- Chamberlainiini --+ | +----- Lamprotulini +--+ | +-- Gonideini +--+ +-- Pseudodontina **
Wu et al., 2021 [Publ 4366], fig. 3-ML
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 72 | 12 | 3 | -- |
+-------- Chamberlainiini --+ | +----- Lamprotulini +--+ | +-- Pseudodontina ** +--+ +-- Gonideini
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 46 | 15 | 4 | -- |
+----------- Chamberlainiini --+ | +-------- Rectidentini +--+ | +----- Lamprotulini +--+ | +-- Pseudodontina ** +--+ +-- Gonideini
Bolotov et al., 2024 [Publ 4369], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 62 | 0 | 0 | -- |
-- Indopseudodon_kayinensis **
Dai et al., 2024 [Publ 4386], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 41 | 9 | 3 | -- |
+-------- Chamberlainiini --+ | +----- Lamprotulini +--+ | +-- Pseudodontina ** +--+ +-- Gonideini
Jeratthitikul et al., 2019 [Publ 3911], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 8 | 7 | 1 | -- |
-- Pilsbryoconcha_exilis **
Walker et al., 2006 [Publ 2666], fig. 6-M
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 13 | 1 | 1 | -- |
+-- Gonideini --+ +-- Pseudodontina **
Whelan et al., 2011 [Publ 3380], fig. 1-ML
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 35 | 5 | 3 | -- |
+-- Pseudodontina ** --+ +-- Rectidentini
Walker et al., 2006 [Publ 2666], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 35 | 1 | 1 | -- |
+-- Gonideini --+ +-- Pseudodontina **
Walker et al., 2006 [Publ 2666], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 14 | 2 | 2 | -- |
+-- Gonideini --+ +-- Pseudodontina **
Walker et al., 2006 [Publ 2666], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 14 | 2 | 1 | -- |
-- Pseudodon_vondembuschianus **
Walker et al., 2006 [Publ 2666], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 14 | 2 | 1 | -- |
-- Pseudodon_vondembuschianus **
Whelan et al., 2011 [Publ 3380], fig. 1-MP
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 35 | 5 | 3 | -- |
-- Pseudodon_mekongi **
