MUSSELp

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Unionidae

Margaritiferidae

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Data last updated
Mon 1 Dec 2025

Bivalvia | Unionoida | Unionoidea | Unionidae | Gonideinae

tribe PSEUDODONTINI Frierson, 1927: cladomics

2 subtribes

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 51 (52 total)
number of subtaxa (rank: subtribe) = 2
analyzed in 97 cladograms in 59 publications
maximum number of terminals in an analysis = 44 (mean = 8.74, median = 4)
maximum number of subtaxa (subtribe) = 2 (mean = 1.48, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 87
maximum number of included terminals = 44, (mean = 9.63 median = 5)
maximum number of subtaxa (subtribe) = 2 (mean = 1.54, median = 2)
proportion of cladograms that support basic monophyly = 0.99

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 37
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (subtribe) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 43 (mean = 13.84, median = 11)
maximum number of subtaxa (subtribe) = 2 (mean = 2, median = 2)
proportion of cladograms that support rigorous monophyly = 0.97

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Pseudodontini cladogram distances
PCoA Ordination

NJ Tree of Pseudodontini cladogram distances
NJ Tree

Cladograms

Zieritz et al., 2024 [Publ 4381], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
10 10 8 2 36 25 8 0 
        +----- Ctenodesmini     
  +-----+                       
  |     |  +-- Rectidentini     
  |     +--+                    
  |        +-- Contradentini    
--+                             
  |     +----- Caudiculatus     
  |  +--+                       
  |  |  |  +-- Lamprotulini     
  |  |  +--+                    
  +--+     +-- Gonideini        
     |                          
     |  +----- Schepmaniini     
     +--+                       
        |  +-- Indopseudodontina **
        +--+                    
           +-- Pseudodontina **

Konopleva et al., 2025 [Publ 4446], fig. 4

iTtTbiriSoTroTroSiD
44 44 43 2 0 0 0 41 
  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Jeratthitikul et al., 2025 [Publ 4423], fig. 1

iTtTbiriSoTroTroSiD
43 43 42 2 7 7 3 40 
        +-- Lamprotulini     
  +-----+                    
  |     +-- Gonideini        
--+                          
  |  +----- Schepmaniini     
  +--+                       
     |  +-- Indopseudodontina **
     +--+                    
        +-- Pseudodontina **

Bolotov et al., 2023 [Publ 4312], fig. S2

iTtTbiriSoTroTroSiD
41 41 39 2 0 0 0 36 
  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iTtTbiriSoTroTroSiD
40 40 41 2 18 18 7 37 
        +-------- Chamberlainiini  
  +-----+                          
  |     |  +----- Ctenodesmini     
  |     +--+                       
  |        |  +-- Rectidentini     
--+        +--+                    
  |           +-- Contradentini    
  |                                
  |  +----------- Lamprotulini     
  +--+                             
     |  +-------- Gonideini        
     +--+                          
        |  +----- Schepmaniini     
        +--+                       
           |  +-- Indopseudodontina **
           +--+                    
              +-- Pseudodontina **

Bolotov et al., 2023 [Publ 4312], fig. 2

iTtTbiriSoTroTroSiD
39 39 38 2 0 0 0 34 
  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Bolotov et al., 2023 [Publ 4312], fig. S1

iTtTbiriSoTroTroSiD
39 39 37 2 0 0 0 35 
  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
30 30 29 2 52 38 4 25 
        +-- Indopseudodontina **
  +-----+                    
  |     +-- Pseudodontina **    
  |                          
--+     +-- Gonideini        
  |  +--+                    
  |  |  +-- Lamprotulini     
  +--+                       
     |  +-- Rectidentini     
     +--+                    
        +-- Contradentini

Konopleva et al., 2021 [Publ 4213], fig. 1

iTtTbiriSoTroTroSiD
23 23 21 2 4 4 2 20 
  +----- Lamprotula       
  |                       
  +----- Leguminaia       
  |                       
--+----- Potomida         
  |                       
  +----- Gonidea          
  |                       
  |  +-- Indopseudodontina **
  +--+                    
     +-- Pseudodontina **

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iTtTbiriSoTroTroSiD
22 22 20 2 14 14 5 22 
           +-- Contradentini    
  +--------+                    
  |        +-- Rectidentini     
  |                             
--+        +-- Indopseudodontina **
  |  +-----+                    
  |  |     +-- Pseudodontina **    
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Chamberlainiini  
        +--+                    
           +-- Lamprotulini

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iTtTbiriSoTroTroSiD
22 22 19 2 14 14 5 22 
           +-- Contradentini    
  +--------+                    
  |        +-- Rectidentini     
  |                             
--+        +-- Indopseudodontina **
  |  +-----+                    
  |  |     +-- Pseudodontina **    
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Chamberlainiini  
        +--+                    
           +-- Lamprotulini

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
21 21 20 2 91 51 5 22 
           +-- Rectidentini     
  +--------+                    
  |        +-- Contradentini    
  |                             
--+        +-- Indopseudodontina **
  |  +-----+                    
  |  |     +-- Pseudodontina **    
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Chamberlainiini  
        +--+                    
           +-- Lamprotulini

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
21 21 20 2 97 51 5 22 
        +-- Rectidentini             
  +-----+                            
  |     +-- Contradentini            
  |                                  
  |     +-- Indopseudodontina **        
  |  +--+                            
  |  |  +-- Pseudodontina **            
--+  |                               
  |  |  +-- Sinosolenaia_iridinea    
  |  |  |                            
  |  |  +-- Sinosolenaia_recognita__1
  |  |  |                            
  |  |  +-- Sinosolenaia_carinata    
  |  |  |                            
  +--+  +-- Leguminaia               
     |  |                            
     |  +-- Microcondylaea           
     |  |                            
     |  +-- Gonidea                  
     |  |                            
     |  +-- Ptychorhynchus           
     |  |                            
     |  +-- Parvasolenaia            
     |  |                            
     +--+-- Lamprotula_leaii         
        |                            
        +-- Lamprotula               
        |                            
        +-- Lamprotula_caveata       
        |                            
        +-- Potomida                 
        |                            
        +-- Pronodularia             
        |                            
        +-- Sinohyriopsis            
        |                            
        +-- Chamberlainia            
        |                            
        +-- Sinosolenaia_recognita__2
        |                            
        +-- Sinosolenaia_oleivora

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
17 17 15 2 59 14 5 17 
  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
17 17 14 2 28 26 5 18 
                       +-- Indopseudodontina **
  +--------------------+                    
  |                    +-- Pseudodontina **    
  |                                         
--+                    +-- Rectidentini     
  |  +-----------------+                    
  |  |                 +-- Contradentini    
  |  |                                      
  +--+  +----------------- Inversidens      
     |  |                                   
     |  |              +-- Chamberlainiini  
     |  +--------------+                    
     +--+              +-- Lamprotulini     
        |                                   
        |  +-------------- Sinosolenaia     
        |  |                                
        |  |        +----- Gonidea          
        +--+  +-----+                       
           |  |     |  +-- Leguminaia       
           |  |     +--+                    
           |  |        +-- Microcondylaea   
           +--+                             
              |        +-- Koreosolenaia    
              |  +-----+                    
              |  |     +-- Parvasolenaia    
              +--+                          
                 |  +----- Postolata        
                 +--+                       
                    |  +-- Ptychorhynchus   
                    +--+                    
                       +-- Obovalis

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
17 17 15 2 28 26 5 19 
                       +-- Indopseudodontina **
  +--------------------+                    
  |                    +-- Pseudodontina **    
  |                                         
  |                    +-- Rectidentini     
--+  +-----------------+                    
  |  |                 +-- Contradentini    
  |  |                                      
  |  |              +----- Inversidens      
  +--+  +-----------+                       
     |  |           |  +-- Chamberlainiini  
     |  |           +--+                    
     |  |              +-- Lamprotulini     
     +--+                                   
        |  +-------------- Sinosolenaia     
        |  |                                
        |  |        +----- Gonidea          
        +--+  +-----+                       
           |  |     |  +-- Leguminaia       
           |  |     +--+                    
           |  |        +-- Microcondylaea   
           +--+                             
              |        +-- Koreosolenaia    
              |  +-----+                    
              |  |     +-- Parvasolenaia    
              +--+                          
                 |  +----- Postolata        
                 +--+                       
                    |  +-- Ptychorhynchus   
                    +--+                    
                       +-- Obovalis

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
17 17 15 2 59 14 5 17 
  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Jeratthitikul et al., 2025 [Publ 4476], fig. 2

iTtTbiriSoTroTroSiD
16 16 14 1 0 0 0 19 
1

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
15 15 15 2 186 41 5 19 
  +----------- Chamberlainiini  
--+                             
  |  +-------- Lamprotulini     
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Indopseudodontina **
        +--+                    
           +-- Pseudodontina **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
15 15 9 2 186 41 5 17 
  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
15 15 13 2 110 36 5 18 
  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
15 15 16 2 184 40 5 17 
  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
14 14 13 2 125 35 5 19 
              +----- Gonideini                 
  +-----------+                                
  |           |  +-- Chamberlainiini           
  |           +--+                             
  |              +-- Lamprotulini              
--+                                            
  |  +-------------- Pilsbryoconcha **            
  |  |                                         
  |  |  +----------- Indopseudodontina **         
  +--+  |                                      
     |  |        +-- Thaiconcha **                
     |  +--------+                             
     |  |        +-- Bineurus **                  
     +--+                                      
        |     +----- Pseudodon_cambodjensis **    
        |  +--+                                
        |  |  |  +-- Pseudodon_mekongi **         
        |  |  +--+                             
        |  |     +-- Pseudodon_vondembuschianus **
        +--+                                   
           |     +-- Pseudodon_inoscularis **     
           |  +--+                             
           |  |  +-- Sundadontina_tumida **       
           +--+                                
              |  +-- Nyeinchanconcha **           
              +--+                             
                 +-- Sundadontina_cumingii **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
13 13 13 2 126 36 5 15 
  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Jeratthitikul & Sutcharit, 2023 [Publ 4289], fig. 1

iTtTbiriSoTroTroSiD
13 13 11 2 0 0 0 16 
  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
11 11 9 1 37 32 7 11 
           +----- Ctenodesmini   
  +--------+                     
  |        |  +-- Rectidentini   
  |        +--+                  
--+           +-- Contradentini  
  |                              
  |  +----------- Chamberlainiini
  +--+                           
     |  +-------- Lamprotulini   
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontina **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
11 11 2 2 72 24 4 13 
  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Konopleva et al., 2019 [Publ 3900], fig. 1

iTtTbiriSoTroTroSiD
11 11 2 2 41 27 2 13 
  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
11 11 9 1 36 31 7 9 
           +----- Ctenodesmini   
  +--------+                     
  |        |  +-- Rectidentini   
  |        +--+                  
--+           +-- Contradentini  
  |                              
  |  +----------- Chamberlainiini
  +--+                           
     |  +-------- Lamprotulini   
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontina **

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
9 9 7 2 23 19 7 3 
  +----------------- Chamberlainiini  
  |                                   
--+  +-------------- Lamprotulini     
  |  |                                
  |  |        +----- Ctenodesmini     
  +--+  +-----+                       
     |  |     |  +-- Rectidentini     
     |  |     +--+                    
     +--+        +-- Contradentini    
        |                             
        |  +-------- Gonideini        
        +--+                          
           |  +----- Schepmaniini     
           +--+                       
              |  +-- Indopseudodontina **
              +--+                    
                 +-- Pseudodontina **

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
8 8 6 2 29 27 7 4 
  +-------------- Chamberlainiini  
  |                                
  |        +----- Ctenodesmini     
--+  +-----+                       
  |  |     |  +-- Rectidentini     
  |  |     +--+                    
  |  |        +-- Contradentini    
  +--+                             
     |        +-- Lamprotulini     
     |  +-----+                    
     |  |     +-- Gonideini        
     +--+                          
        |  +----- Schepmaniini     
        +--+                       
           |  +-- Indopseudodontina **
           +--+                    
              +-- Pseudodontina **

Zieritz et al., 2024 [Publ 4381], fig. 8A

iTtTbiriSoTroTroSiD
8 8 6 1 0 0 0 12 
1

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
8 8 6 2 29 27 7 4 
  +-------------------------- Chamberlainiini  
  |                                            
  |                    +----- Ctenodesmini     
  |  +-----------------+                       
--+  |                 |  +-- Rectidentini     
  |  |                 +--+                    
  |  |                    +-- Contradentini    
  |  |                                         
  +--+                 +----- Schepmaniini     
     |  +--------------+                       
     |  |              |  +-- Indopseudodontina **
     |  |              +--+                    
     |  |                 +-- Pseudodontina **    
     +--+                                      
        |                 +-- Inversidens      
        |  +--------------+                    
        |  |              +-- Lamprotulini     
        +--+                                   
           |  +-------------- Sinosolenaia     
           |  |                                
           +--+           +-- Gonidea          
              |  +--------+                    
              |  |        +-- Microcondylaea   
              +--+                             
                 |  +-------- Parvasolenaia    
                 +--+                          
                    |  +----- Koreosolenaia    
                    +--+                       
                       |  +-- Postolata        
                       +--+                    
                          +-- Ptychorhynchus

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
8 8 6 2 29 27 7 6 
  +----------------------------- Schepmaniini       
  |                                                 
  |                          +-- Indopseudodontina **  
  |  +-----------------------+                      
--+  |                       +-- Pseudodontina **      
  |  |                                              
  |  |              +----------- Hyriopsis_bialata  
  |  |              |                               
  +--+  +-----------+        +-- Rectidens          
     |  |           |  +-----+                      
     |  |           |  |     +-- Ctenodesmini       
     |  |           +--+                            
     |  |              |  +----- Ensidens           
     |  |              +--+                         
     +--+                 |  +-- Hyriopsis_desowitzi
        |                 +--+                      
        |                    +-- Contradentini      
        |                                           
        |  +-------------------- Inversidens        
        |  |                                        
        |  |              +----- Lamprotula         
        +--+  +-----------+                         
           |  |           |  +-- Chamberlainiini    
           |  |           +--+                      
           +--+              +-- Potomida           
              |                                     
              |  +-------------- Sinosolenaia       
              |  |                                  
              +--+           +-- Gonidea            
                 |  +--------+                      
                 |  |        +-- Microcondylaea     
                 +--+                               
                    |  +-------- Koreosolenaia      
                    +--+                            
                       |  +----- Parvasolenaia      
                       +--+                         
                          |  +-- Postolata          
                          +--+                      
                             +-- Ptychorhynchus

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
8 8 6 2 30 28 7 4 
  +----------------- Chamberlainiini  
  |                                   
  |           +----- Ctenodesmini     
--+  +--------+                       
  |  |        |  +-- Rectidentini     
  |  |        +--+                    
  +--+           +-- Contradentini    
     |                                
     |  +----------- Lamprotulini     
     +--+                             
        |  +-------- Gonideini        
        +--+                          
           |  +----- Schepmaniini     
           +--+                       
              |  +-- Indopseudodontina **
              +--+                    
                 +-- Pseudodontina **

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
8 8 6 2 29 27 7 4 
           +----- Schepmaniini     
  +--------+                       
  |        |  +-- Indopseudodontina **
  |        +--+                    
--+           +-- Pseudodontina **    
  |                                
  |  +----------- Gonideini        
  |  |                             
  +--+        +-- Chamberlainiini  
     |  +-----+                    
     |  |     +-- Lamprotulini     
     +--+                          
        |  +----- Ctenodesmini     
        +--+                       
           |  +-- Rectidentini     
           +--+                    
              +-- Contradentini

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
8 8 5 2 29 27 7 7 
  +----------------------------- Schepmania_nieuwenhuisi__1
  |                                                        
  +----------------------------- Schepmania_nieuwenhuisi__2
  |                                                        
  |                          +-- Indopseudodontina **         
--+  +-----------------------+                             
  |  |                       +-- Pseudodontina **             
  |  |                                                     
  |  |                       +-- Ensidens                  
  |  |                       |                             
  |  |                       +-- Hyriopsis_desowitzi       
  +--+                       |                             
     |                       +-- Rectidens                 
     |  +--------------------+                             
     |  |                    +-- Hyriopsis_bialata         
     |  |                    |                             
     |  |                    +-- Ctenodesmini              
     |  |                    |                             
     +--+                    +-- Contradentini             
        |                                                  
        |  +-------------------- Inversidens               
        |  |                                               
        |  |              +----- Lamprotula                
        +--+  +-----------+                                
           |  |           |  +-- Chamberlainiini           
           |  |           +--+                             
           +--+              +-- Potomida                  
              |                                            
              |  +-------------- Sinosolenaia              
              |  |                                         
              +--+  +----------- Gonidea                   
                 |  |                                      
                 +--+----------- Microcondylaea            
                    |                                      
                    |  +-------- Koreosolenaia             
                    +--+                                   
                       |  +----- Parvasolenaia             
                       +--+                                
                          |  +-- Postolata                 
                          +--+                             
                             +-- Ptychorhynchus

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
8 8 5 2 114 25 5 13 
        +-- Chamberlainiini  
  +-----+                    
  |     +-- Lamprotulini     
--+                          
  |  +----- Gonideini        
  +--+                       
     |  +-- Indopseudodontina **
     +--+                    
        +-- Pseudodontina **

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
6 6 4 1 25 4 2 13 
  +----- Pseudodontina **
--+                   
  |  +-- Rectidentini 
  +--+                
     +-- Contradentini

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
6 6 4 2 85 23 5 11 
  +----------- Chamberlainiini  
--+                             
  |  +-------- Lamprotulini     
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Indopseudodontina **
        +--+                    
           +-- Pseudodontina **

Bolotov et al., 2022 [Publ 4250], fig. 8

iTtTbiriSoTroTroSiD
6 6 4 2 43 17 3 9 
  +----- Contradentini    
--+                       
  |  +-- Pseudodontina **    
  +--+                    
     +-- Indopseudodontina **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
6 6 4 2 85 23 5 11 
  +----------- Chamberlainiini  
--+                             
  |  +-------- Lamprotulini     
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Indopseudodontina **
        +--+                    
           +-- Pseudodontina **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
6 6 4 1 34 8 4 13 
        +-- Rectidentini 
  +-----+                
  |     +-- Contradentini
--+                      
  |  +----- Pseudodontina **
  +--+                   
     |  +-- Gonideini    
     +--+                
        +-- Lamprotulini

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
5 5 3 2 52 12 5 9 
  +----------- Chamberlainiini  
  |                             
  |        +-- Rectidentini     
--+  +-----+                    
  |  |     +-- Contradentini    
  |  |                          
  +--+     +-- Gonideini        
     |  +--+                    
     |  |  +-- Lamprotulini     
     +--+                       
        |  +-- Indopseudodontina **
        +--+                    
           +-- Pseudodontina **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
5 5 3 2 52 12 5 9 
  +-------------- Chamberlainiini  
  |                                
--+           +-- Rectidentini     
  |  +--------+                    
  |  |        +-- Contradentini    
  +--+                             
     |  +-------- Lamprotulini     
     +--+                          
        |  +----- Gonideini        
        +--+                       
           |  +-- Indopseudodontina **
           +--+                    
              +-- Pseudodontina **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
5 5 3 1 75 22 5 13 
              +-- Chamberlainiini
  +-----------+                  
  |           +-- Lamprotulini   
  |                              
--+           +-- Parvasolenaia  
  |  +--------+                  
  |  |        +-- Pseudodontina **  
  +--+                           
     |  +-------- Sinosolenaia   
     +--+                        
        |  +----- Gonidea        
        +--+                     
           |  +-- Microcondylaea 
           +--+                  
              +-- Leguminaia

Wu et al., 2021 [Publ 4215], fig. 3

iTtTbiriSoTroTroSiD
5 19 17 2 19 14 3 26 
                 +----- Chamberlainia    
  +--------------+                       
  |              |  +-- Sundadontina **     
  |              +--+                    
  |                 +-- Sinohyriopsis    
  |                                      
--+              +----- Indopseudodontina **
  |  +-----------+                       
  |  |           |  +-- Pseudodon **        
  |  |           +--+                    
  |  |              +-- Pilsbryoconcha **   
  +--+                                   
     |              +-- Sinosolenaia     
     |  +-----------+                    
     |  |           +-- Lamprotulini     
     |  |                                
     +--+           +-- Ptychorhynchus   
        |  +--------+                    
        |  |        +-- Parvasolenaia    
        +--+                             
           |  +-------- Inversidens      
           +--+                          
              |  +----- Gonidea          
              +--+                       
                 |  +-- Microcondylaea   
                 +--+                    
                    +-- Leguminaia

Bolotov et al., 2022 [Publ 4252], fig. S1

iTtTbiriSoTroTroSiD
4 4 0 2 57 0 0 10 
  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina **
  |                    
  +-- Sundadontina **

Bolotov et al., 2022 [Publ 4239], fig. S1

iTtTbiriSoTroTroSiD
4 4 0 2 54 0 0 10 
  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina **
  |                    
  +-- Sundadontina **

Bolotov et al., 2022 [Publ 4239], fig. S2

iTtTbiriSoTroTroSiD
4 4 0 2 54 0 0 10 
  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina **
  |                    
  +-- Sundadontina **

Bolotov et al., 2022 [Publ 4252], fig. 2

iTtTbiriSoTroTroSiD
4 4 0 2 57 0 0 10 
  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina **
  |                    
  +-- Sundadontina **

Bolotov et al., 2022 [Publ 4239], fig. 2

iTtTbiriSoTroTroSiD
4 4 0 2 54 0 0 10 
  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina **
  |                    
  +-- Sundadontina **

Kongim et al., 2023 [Publ 4290], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 2 12 12 3 10 
  +----------- Chamberlainiini  
--+                             
  |  +-------- Lamprotulini     
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Indopseudodontina **
        +--+                    
           +-- Pseudodontina **

Konopleva et al., 2019 [Publ 4041], fig. 3

iTtTbiriSoTroTroSiD
3 3 0 1 31 31 2 11 
  +----- Indopseudodontina **
--+                       
  |  +-- Rectidentini     
  +--+                    
     +-- Contradentini

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
2 2 0 1 90 19 5 -- 
  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 1 52 21 5 -- 
  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
2 2 0 1 37 10 3 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Lamprotulini

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 72 12 3 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 1 51 9 3 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Matos et al., 2024 [Publ 4383], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 18 16 5 -- 
  +----------- Chamberlainiini
  |                           
--+        +-- Contradentini  
  |  +-----+                  
  |  |     +-- Rectidentini   
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 1 44 2 2 -- 
1

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 1 44 2 2 -- 
1

Froufe et al., 2019 [Publ 3936], fig. 3-M

iTtTbiriSoTroTroSiD
2 2 0 1 17 6 3 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Gonideini      
     +--+                  
        +-- Pseudodontina **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
2 2 0 1 94 18 5 -- 
  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 45 2 2 -- 
  +-- Gonideini    
--+                
  +-- Pseudodontina **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
2 2 0 1 94 18 5 -- 
  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Graf, 2002 [Publ 2651], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 16 2 2 -- 
1

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 1 35 9 3 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
2 2 0 1 72 12 3 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 52 10 3 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Pseudodontina **  
  +--+                     
     |  +-- Lamprotulini   
     +--+                  
        +-- Gonideini

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
2 2 0 1 46 15 5 -- 
  +----------- Chamberlainiini
  |                           
--+        +-- Contradentini  
  |  +-----+                  
  |  |     +-- Rectidentini   
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Matos et al., 2024 [Publ 4384], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 18 16 5 -- 
  +----------- Chamberlainiini
  |                           
--+        +-- Contradentini  
  |  +-----+                  
  |  |     +-- Rectidentini   
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Froufe et al., 2019 [Publ 3936], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 1 17 6 3 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Gonideini      
     +--+                  
        +-- Pseudodontina **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
2 2 0 1 72 12 3 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 1 42 11 3 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 86 17 5 -- 
1

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 32 9 3 -- 
  +----- Pseudodontina **  
--+                     
  |  +-- Lamprotulini   
  +--+                  
     +-- Chamberlainiini

Zieritz et al., 2020 [Publ 4102], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 29 29 6 -- 
  +----------------- Chamberlainiini  
  |                                   
--+  +-------------- Lamprotulini     
  |  |                                
  +--+  +----------- Gonideini        
     |  |                             
     +--+        +-- Pseudodontina **    
        |  +-----+                    
        |  |     +-- Indopseudodontina **
        +--+                          
           |  +----- Contradentini    
           +--+                       
              |  +-- Ctenodesmini     
              +--+                    
                 +-- Rectidentini

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
2 2 0 1 21 10 5 -- 
  +----------- Chamberlainiini
  |                           
--+        +-- Contradentini  
  |  +-----+                  
  |  |     +-- Rectidentini   
  +--+                        
     |  +----- Pseudodontina **  
     +--+                     
        |  +-- Lamprotulini   
        +--+                  
           +-- Gonideini

Zhang et al., 2024 [Publ 4397], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 1 26 24 5 -- 
  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 84 16 5 -- 
  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 29 9 4 -- 
  +-------- Chamberlainiini
  |                        
--+     +-- Gonideini      
  |  +--+                  
  |  |  +-- Pseudodontina **  
  +--+                     
     |  +-- Contradentini  
     +--+                  
        +-- Lamprotulini

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 42 11 3 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
2 2 0 1 90 19 5 -- 
  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 1 67 11 3 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 45 2 2 -- 
1

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 1 51 17 5 -- 
  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Bolotov et al., 2024 [Publ 4369], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 62 0 0 -- 
-- Indopseudodon_kayinensis **

Walker et al., 2006 [Publ 2666], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 2 2 -- 
  +-- Gonideini    
--+                
  +-- Pseudodontina **

Walker et al., 2006 [Publ 2666], fig. 6-M

iTtTbiriSoTroTroSiD
1 1 0 1 13 1 1 -- 
  +-- Gonideini    
--+                
  +-- Pseudodontina **

Walker et al., 2006 [Publ 2666], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 35 1 1 -- 
  +-- Gonideini    
--+                
  +-- Pseudodontina **

Walker et al., 2006 [Publ 2666], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 14 2 1 -- 
-- Pseudodon_vondembuschianus **

Walker et al., 2006 [Publ 2666], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 14 2 1 -- 
-- Pseudodon_vondembuschianus **

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 8 7 2 -- 
-- Pilsbryoconcha_exilis **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
1 1 0 1 35 5 3 -- 
  +-- Pseudodontina **
--+                
  +-- Contradentini

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
1 1 0 1 35 5 3 -- 
-- Pseudodon_vondembuschianus **

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 40 9 3 -- 
  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini
 
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