MUSSELpdb | tribe Pseudodontini Frierson, 1927 cladomics

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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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Bivalvia | Unionoida | Unionoidea | Unionidae | Gonideinae

tribe PSEUDODONTINI Frierson, 1927: cladomics
2 subtribes

Cladomics Summary

number of species = 50 (51 total)
number of subtaxa (rank: subtribe) = 2
analyzed in 94 cladograms in 56 publications
maximum number of terminals in an analysis = 43 (mean = 8.36, median = 4)
maximum number of subtaxa (subtribe) = 2 (mean = 1.49, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 84
maximum number of included terminals = 43, (mean = 9.24 median = 5)
maximum number of subtaxa (subtribe) = 2 (mean = 1.55, median = 2)
proportion of cladograms that support basic monophyly = 0.99

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 37
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (subtribe) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 43 (mean = 13.84, median = 11)
maximum number of subtaxa (subtribe) = 2 (mean = 2, median = 2)
proportion of cladograms that support rigorous monophyly = 0.97

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ Tree

Zieritz et al., 2024 [Publ 4381], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
10	10	8	2	36	25	8	0

        +----- Ctenodesmini     
  +-----+                       
  |     |  +-- Rectidentini     
  |     +--+                    
  |        +-- Contradentini    
--+                             
  |     +----- Caudiculatus     
  |  +--+                       
  |  |  |  +-- Lamprotulini     
  |  |  +--+                    
  +--+     +-- Gonideini        
     |                          
     |  +----- Schepmaniini     
     +--+                       
        |  +-- Indopseudodontina **
        +--+                    
           +-- Pseudodontina **

Jeratthitikul et al., 2025 [Publ 4423], fig. 1

tT	bi	riS	oT	roT	roS	iD
43	43	42	2	7	7	3	40

        +-- Lamprotulini     
  +-----+                    
  |     +-- Gonideini        
--+                          
  |  +----- Schepmaniini     
  +--+                       
     |  +-- Indopseudodontina **
     +--+                    
        +-- Pseudodontina **

Bolotov et al., 2023 [Publ 4312], fig. S2

tT	bi	riS	oT	roT	roS	iD
41	41	39	2	0	0	0	36

  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

tT	bi	riS	oT	roT	roS	iD
40	40	41	2	18	18	7	37

        +-------- Chamberlainiini  
  +-----+                          
  |     |  +----- Ctenodesmini     
  |     +--+                       
  |        |  +-- Rectidentini     
--+        +--+                    
  |           +-- Contradentini    
  |                                
  |  +----------- Lamprotulini     
  +--+                             
     |  +-------- Gonideini        
     +--+                          
        |  +----- Schepmaniini     
        +--+                       
           |  +-- Indopseudodontina **
           +--+                    
              +-- Pseudodontina **

Bolotov et al., 2023 [Publ 4312], fig. S1

tT	bi	riS	oT	roT	roS	iD
39	39	37	2	0	0	0	35

  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Bolotov et al., 2023 [Publ 4312], fig. 2

tT	bi	riS	oT	roT	roS	iD
39	39	38	2	0	0	0	34

  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Konopleva et al., 2023 [Publ 4262], fig. 8

tT	bi	riS	oT	roT	roS	iD
30	30	29	2	52	38	4	25

        +-- Indopseudodontina **
  +-----+                    
  |     +-- Pseudodontina **    
  |                          
--+     +-- Gonideini        
  |  +--+                    
  |  |  +-- Lamprotulini     
  +--+                       
     |  +-- Rectidentini     
     +--+                    
        +-- Contradentini

Konopleva et al., 2021 [Publ 4213], fig. 1

tT	bi	riS	oT	roT	roS	iD
23	23	21	2	4	4	2	20

  +----- Lamprotula       
  |                       
  +----- Leguminaia       
  |                       
--+----- Potomida         
  |                       
  +----- Gonidea          
  |                       
  |  +-- Indopseudodontina **
  +--+                    
     +-- Pseudodontina **

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

tT	bi	riS	oT	roT	roS	iD
22	22	19	2	14	14	5	22

           +-- Contradentini    
  +--------+                    
  |        +-- Rectidentini     
  |                             
--+        +-- Indopseudodontina **
  |  +-----+                    
  |  |     +-- Pseudodontina **    
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Chamberlainiini  
        +--+                    
           +-- Lamprotulini

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

tT	bi	riS	oT	roT	roS	iD
22	22	20	2	14	14	5	22

           +-- Contradentini    
  +--------+                    
  |        +-- Rectidentini     
  |                             
--+        +-- Indopseudodontina **
  |  +-----+                    
  |  |     +-- Pseudodontina **    
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Chamberlainiini  
        +--+                    
           +-- Lamprotulini

Bolotov et al., 2020 [Publ 4100], fig. 1

tT	bi	riS	oT	roT	roS	iD
21	21	20	2	97	51	5	22

        +-- Rectidentini             
  +-----+                            
  |     +-- Contradentini            
  |                                  
  |     +-- Indopseudodontina **        
  |  +--+                            
  |  |  +-- Pseudodontina **            
--+  |                               
  |  |  +-- Sinosolenaia_iridinea    
  |  |  |                            
  |  |  +-- Sinosolenaia_recognita__1
  |  |  |                            
  |  |  +-- Sinosolenaia_carinata    
  |  |  |                            
  +--+  +-- Leguminaia               
     |  |                            
     |  +-- Microcondylaea           
     |  |                            
     |  +-- Gonidea                  
     |  |                            
     |  +-- Ptychorhynchus           
     |  |                            
     |  +-- Parvasolenaia            
     |  |                            
     +--+-- Lamprotula_leaii         
        |                            
        +-- Lamprotula               
        |                            
        +-- Lamprotula_caveata       
        |                            
        +-- Potomida                 
        |                            
        +-- Pronodularia             
        |                            
        +-- Sinohyriopsis            
        |                            
        +-- Chamberlainia            
        |                            
        +-- Sinosolenaia_recognita__2
        |                            
        +-- Sinosolenaia_oleivora

Bolotov et al., 2020 [Publ 4100], fig. S1

tT	bi	riS	oT	roT	roS	iD
21	21	20	2	91	51	5	22

           +-- Rectidentini     
  +--------+                    
  |        +-- Contradentini    
  |                             
--+        +-- Indopseudodontina **
  |  +-----+                    
  |  |     +-- Pseudodontina **    
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Chamberlainiini  
        +--+                    
           +-- Lamprotulini

Dai et al., 2023 [Publ 4285], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
17	17	15	2	28	26	5	19

                       +-- Indopseudodontina **
  +--------------------+                    
  |                    +-- Pseudodontina **    
  |                                         
  |                    +-- Rectidentini     
--+  +-----------------+                    
  |  |                 +-- Contradentini    
  |  |                                      
  |  |              +----- Inversidens      
  +--+  +-----------+                       
     |  |           |  +-- Chamberlainiini  
     |  |           +--+                    
     |  |              +-- Lamprotulini     
     +--+                                   
        |  +-------------- Sinosolenaia     
        |  |                                
        |  |        +----- Gonidea          
        +--+  +-----+                       
           |  |     |  +-- Leguminaia       
           |  |     +--+                    
           |  |        +-- Microcondylaea   
           +--+                             
              |        +-- Koreosolenaia    
              |  +-----+                    
              |  |     +-- Parvasolenaia    
              +--+                          
                 |  +----- Postolata        
                 +--+                       
                    |  +-- Ptychorhynchus   
                    +--+                    
                       +-- Obovalis

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

tT	bi	riS	oT	roT	roS	iD
17	17	15	2	59	14	5	17

  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Dai et al., 2023 [Publ 4285], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
17	17	14	2	28	26	5	18

                       +-- Indopseudodontina **
  +--------------------+                    
  |                    +-- Pseudodontina **    
  |                                         
--+                    +-- Rectidentini     
  |  +-----------------+                    
  |  |                 +-- Contradentini    
  |  |                                      
  +--+  +----------------- Inversidens      
     |  |                                   
     |  |              +-- Chamberlainiini  
     |  +--------------+                    
     +--+              +-- Lamprotulini     
        |                                   
        |  +-------------- Sinosolenaia     
        |  |                                
        |  |        +----- Gonidea          
        +--+  +-----+                       
           |  |     |  +-- Leguminaia       
           |  |     +--+                    
           |  |        +-- Microcondylaea   
           +--+                             
              |        +-- Koreosolenaia    
              |  +-----+                    
              |  |     +-- Parvasolenaia    
              +--+                          
                 |  +----- Postolata        
                 +--+                       
                    |  +-- Ptychorhynchus   
                    +--+                    
                       +-- Obovalis

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

tT	bi	riS	oT	roT	roS	iD
17	17	15	2	59	14	5	17

  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
15	15	9	2	184	41	5	17

  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
15	15	15	2	184	41	5	19

  +----------- Chamberlainiini  
--+                             
  |  +-------- Lamprotulini     
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Indopseudodontina **
        +--+                    
           +-- Pseudodontina **

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
15	15	16	2	182	40	5	17

  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Bolotov et al., 2017 [Publ 3764], fig. S1

tT	bi	riS	oT	roT	roS	iD
15	15	13	2	110	36	5	18

  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Bolotov et al., 2017 [Publ 3761], fig. S2

tT	bi	riS	oT	roT	roS	iD
14	14	13	2	125	35	5	19

              +----- Gonideini                 
  +-----------+                                
  |           |  +-- Chamberlainiini           
  |           +--+                             
  |              +-- Lamprotulini              
--+                                            
  |  +-------------- Pilsbryoconcha **            
  |  |                                         
  |  |  +----------- Indopseudodontina **         
  +--+  |                                      
     |  |        +-- Thaiconcha **                
     |  +--------+                             
     |  |        +-- Bineurus **                  
     +--+                                      
        |     +----- Pseudodon_vagulus **         
        |  +--+                                
        |  |  |  +-- Pseudodon_mekongi **         
        |  |  +--+                             
        |  |     +-- Pseudodon_vondembuschianus **
        +--+                                   
           |     +-- Pseudodon_inoscularis **     
           |  +--+                             
           |  |  +-- Sundadontina_tumida **       
           +--+                                
              |  +-- Nyeinchanconcha **           
              +--+                             
                 +-- Sundadontina_cumingii **

Jeratthitikul & Sutcharit, 2023 [Publ 4289], fig. 1

tT	bi	riS	oT	roT	roS	iD
13	13	11	2	0	0	0	16

  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Bolotov et al., 2017 [Publ 3761], fig. S1

tT	bi	riS	oT	roT	roS	iD
13	13	13	2	126	36	5	15

  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Konopleva et al., 2019 [Publ 3900], fig. 1

tT	bi	riS	oT	roT	roS	iD
11	11	2	2	41	27	2	13

  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Dai et al., 2024 [Publ 4389], fig. 3

tT	bi	riS	oT	roT	roS	iD
11	11	9	1	36	31	7	9

           +----- Ctenodesmini   
  +--------+                     
  |        |  +-- Rectidentini   
  |        +--+                  
--+           +-- Contradentini  
  |                              
  |  +----------- Chamberlainiini
  +--+                           
     |  +-------- Lamprotulini   
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontina **

Dai et al., 2025 [Publ 4441], fig. 1

tT	bi	riS	oT	roT	roS	iD
11	11	9	1	37	32	7	11

           +----- Ctenodesmini   
  +--------+                     
  |        |  +-- Rectidentini   
  |        +--+                  
--+           +-- Contradentini  
  |                              
  |  +----------- Chamberlainiini
  +--+                           
     |  +-------- Lamprotulini   
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontina **

Bolotov et al., 2018 [Publ 3826], fig. 2

tT	bi	riS	oT	roT	roS	iD
11	11	2	2	72	24	4	13

  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Zieritz et al., 2021 [Publ 4218], fig. 5

tT	bi	riS	oT	roT	roS	iD
9	9	7	2	23	19	7	3

  +----------------- Chamberlainiini  
  |                                   
--+  +-------------- Lamprotulini     
  |  |                                
  |  |        +----- Ctenodesmini     
  +--+  +-----+                       
     |  |     |  +-- Rectidentini     
     |  |     +--+                    
     +--+        +-- Contradentini    
        |                             
        |  +-------- Gonideini        
        +--+                          
           |  +----- Schepmaniini     
           +--+                       
              |  +-- Indopseudodontina **
              +--+                    
                 +-- Pseudodontina **

Wu et al., 2024 [Publ 4388], fig. 5

tT	bi	riS	oT	roT	roS	iD
8	8	6	2	30	28	7	4

  +----------------- Chamberlainiini  
  |                                   
  |           +----- Ctenodesmini     
--+  +--------+                       
  |  |        |  +-- Rectidentini     
  |  |        +--+                    
  +--+           +-- Contradentini    
     |                                
     |  +----------- Lamprotulini     
     +--+                             
        |  +-------- Gonideini        
        +--+                          
           |  +----- Schepmaniini     
           +--+                       
              |  +-- Indopseudodontina **
              +--+                    
                 +-- Pseudodontina **

Zieritz et al., 2024 [Publ 4381], fig. 8A

tT	bi	riS	oT	roT	roS	iD
8	8	6	1	0	0	0	12

Wu et al., 2024 [Publ 4350], fig. 5b

tT	bi	riS	oT	roT	roS	iD
8	8	6	2	29	27	7	6

  +----------------------------- Schepmaniini       
  |                                                 
  |                          +-- Indopseudodontina **  
  |  +-----------------------+                      
--+  |                       +-- Pseudodontina **      
  |  |                                              
  |  |              +----------- Hyriopsis_bialata  
  |  |              |                               
  +--+  +-----------+        +-- Rectidens          
     |  |           |  +-----+                      
     |  |           |  |     +-- Ctenodesmini       
     |  |           +--+                            
     |  |              |  +----- Ensidens           
     |  |              +--+                         
     +--+                 |  +-- Hyriopsis_desowitzi
        |                 +--+                      
        |                    +-- Contradentini      
        |                                           
        |  +-------------------- Inversidens        
        |  |                                        
        |  |              +----- Lamprotula         
        +--+  +-----------+                         
           |  |           |  +-- Chamberlainiini    
           |  |           +--+                      
           +--+              +-- Potomida           
              |                                     
              |  +-------------- Sinosolenaia       
              |  |                                  
              +--+           +-- Gonidea            
                 |  +--------+                      
                 |  |        +-- Microcondylaea     
                 +--+                               
                    |  +-------- Koreosolenaia      
                    +--+                            
                       |  +----- Parvasolenaia      
                       +--+                         
                          |  +-- Postolata          
                          +--+                      
                             +-- Ptychorhynchus

Wu et al., 2024 [Publ 4350], fig. 5c

tT	bi	riS	oT	roT	roS	iD
8	8	5	2	29	27	7	7

  +----------------------------- Schepmania_nieuwenhuisi__1
  |                                                        
  +----------------------------- Schepmania_nieuwenhuisi__2
  |                                                        
  |                          +-- Indopseudodontina **         
--+  +-----------------------+                             
  |  |                       +-- Pseudodontina **             
  |  |                                                     
  |  |                       +-- Ensidens                  
  |  |                       |                             
  |  |                       +-- Hyriopsis_desowitzi       
  +--+                       |                             
     |                       +-- Rectidens                 
     |  +--------------------+                             
     |  |                    +-- Hyriopsis_bialata         
     |  |                    |                             
     |  |                    +-- Ctenodesmini              
     |  |                    |                             
     +--+                    +-- Contradentini             
        |                                                  
        |  +-------------------- Inversidens               
        |  |                                               
        |  |              +----- Lamprotula                
        +--+  +-----------+                                
           |  |           |  +-- Chamberlainiini           
           |  |           +--+                             
           +--+              +-- Potomida                  
              |                                            
              |  +-------------- Sinosolenaia              
              |  |                                         
              +--+  +----------- Gonidea                   
                 |  |                                      
                 +--+----------- Microcondylaea            
                    |                                      
                    |  +-------- Koreosolenaia             
                    +--+                                   
                       |  +----- Parvasolenaia             
                       +--+                                
                          |  +-- Postolata                 
                          +--+                             
                             +-- Ptychorhynchus

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

tT	bi	riS	oT	roT	roS	iD
8	8	5	2	113	25	5	13

        +-- Chamberlainiini  
  +-----+                    
  |     +-- Lamprotulini     
--+                          
  |  +----- Gonideini        
  +--+                       
     |  +-- Indopseudodontina **
     +--+                    
        +-- Pseudodontina **

Liu et al., 2024 [Publ 4380], fig. 5

tT	bi	riS	oT	roT	roS	iD
8	8	6	2	29	27	7	4

           +----- Schepmaniini     
  +--------+                       
  |        |  +-- Indopseudodontina **
  |        +--+                    
--+           +-- Pseudodontina **    
  |                                
  |  +----------- Gonideini        
  |  |                             
  +--+        +-- Chamberlainiini  
     |  +-----+                    
     |  |     +-- Lamprotulini     
     +--+                          
        |  +----- Ctenodesmini     
        +--+                       
           |  +-- Rectidentini     
           +--+                    
              +-- Contradentini

Wu et al., 2024 [Publ 4350], fig. 5a

tT	bi	riS	oT	roT	roS	iD
8	8	6	2	29	27	7	4

  +-------------------------- Chamberlainiini  
  |                                            
  |                    +----- Ctenodesmini     
  |  +-----------------+                       
--+  |                 |  +-- Rectidentini     
  |  |                 +--+                    
  |  |                    +-- Contradentini    
  |  |                                         
  +--+                 +----- Schepmaniini     
     |  +--------------+                       
     |  |              |  +-- Indopseudodontina **
     |  |              +--+                    
     |  |                 +-- Pseudodontina **    
     +--+                                      
        |                 +-- Inversidens      
        |  +--------------+                    
        |  |              +-- Lamprotulini     
        +--+                                   
           |  +-------------- Sinosolenaia     
           |  |                                
           +--+           +-- Gonidea          
              |  +--------+                    
              |  |        +-- Microcondylaea   
              +--+                             
                 |  +-------- Parvasolenaia    
                 +--+                          
                    |  +----- Koreosolenaia    
                    +--+                       
                       |  +-- Postolata        
                       +--+                    
                          +-- Ptychorhynchus

Wu et al., 2024 [Publ 4350], fig. 6-7

tT	bi	riS	oT	roT	roS	iD
8	8	6	2	29	27	7	4

  +-------------- Chamberlainiini  
  |                                
  |        +----- Ctenodesmini     
--+  +-----+                       
  |  |     |  +-- Rectidentini     
  |  |     +--+                    
  |  |        +-- Contradentini    
  +--+                             
     |        +-- Lamprotulini     
     |  +-----+                    
     |  |     +-- Gonideini        
     +--+                          
        |  +----- Schepmaniini     
        +--+                       
           |  +-- Indopseudodontina **
           +--+                    
              +-- Pseudodontina **

Konopleva et al., 2017 [Publ 3807], fig. 6

tT	bi	riS	oT	roT	roS	iD
6	6	4	1	25	4	2	13

  +----- Pseudodontina **
--+                   
  |  +-- Rectidentini 
  +--+                
     +-- Contradentini

Bolotov et al., 2022 [Publ 4250], fig. 8

tT	bi	riS	oT	roT	roS	iD
6	6	4	2	43	17	3	9

  +----- Contradentini    
--+                       
  |  +-- Pseudodontina **    
  +--+                    
     +-- Indopseudodontina **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

tT	bi	riS	oT	roT	roS	iD
6	6	4	1	34	8	4	13

        +-- Rectidentini 
  +-----+                
  |     +-- Contradentini
--+                      
  |  +----- Pseudodontina **
  +--+                   
     |  +-- Gonideini    
     +--+                
        +-- Lamprotulini

Wu et al., 2021 [Publ 4366], fig. 2_I

tT	bi	riS	oT	roT	roS	iD
6	6	4	2	85	23	5	11

  +----------- Chamberlainiini  
--+                             
  |  +-------- Lamprotulini     
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Indopseudodontina **
        +--+                    
           +-- Pseudodontina **

Wu et al., 2021 [Publ 4366], fig. 6

tT	bi	riS	oT	roT	roS	iD
6	6	4	2	85	23	5	11

  +----------- Chamberlainiini  
--+                             
  |  +-------- Lamprotulini     
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Indopseudodontina **
        +--+                    
           +-- Pseudodontina **

Wu et al., 2021 [Publ 4215], fig. 3

tT	bi	riS	oT	roT	roS	iD
5	19	17	2	19	14	3	26

                 +----- Chamberlainia    
  +--------------+                       
  |              |  +-- Sundadontina **     
  |              +--+                    
  |                 +-- Sinohyriopsis    
  |                                      
--+              +----- Indopseudodontina **
  |  +-----------+                       
  |  |           |  +-- Pseudodon **        
  |  |           +--+                    
  |  |              +-- Pilsbryoconcha **   
  +--+                                   
     |              +-- Sinosolenaia     
     |  +-----------+                    
     |  |           +-- Lamprotulini     
     |  |                                
     +--+           +-- Ptychorhynchus   
        |  +--------+                    
        |  |        +-- Parvasolenaia    
        +--+                             
           |  +-------- Inversidens      
           +--+                          
              |  +----- Gonidea          
              +--+                       
                 |  +-- Microcondylaea   
                 +--+                    
                    +-- Leguminaia

Pfeiffer et al., 2019 [Publ 3937], fig. 6

tT	bi	riS	oT	roT	roS	iD
5	5	3	2	52	12	5	9

  +----------- Chamberlainiini  
  |                             
  |        +-- Rectidentini     
--+  +-----+                    
  |  |     +-- Contradentini    
  |  |                          
  +--+     +-- Gonideini        
     |  +--+                    
     |  |  +-- Lamprotulini     
     +--+                       
        |  +-- Indopseudodontina **
        +--+                    
           +-- Pseudodontina **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

tT	bi	riS	oT	roT	roS	iD
5	5	3	1	75	22	5	13

              +-- Chamberlainiini
  +-----------+                  
  |           +-- Lamprotulini   
  |                              
--+           +-- Parvasolenaia  
  |  +--------+                  
  |  |        +-- Pseudodontina **  
  +--+                           
     |  +-------- Sinosolenaia   
     +--+                        
        |  +----- Gonidea        
        +--+                     
           |  +-- Microcondylaea 
           +--+                  
              +-- Leguminaia

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

tT	bi	riS	oT	roT	roS	iD
5	5	3	2	52	12	5	9

  +-------------- Chamberlainiini  
  |                                
--+           +-- Rectidentini     
  |  +--------+                    
  |  |        +-- Contradentini    
  +--+                             
     |  +-------- Lamprotulini     
     +--+                          
        |  +----- Gonideini        
        +--+                       
           |  +-- Indopseudodontina **
           +--+                    
              +-- Pseudodontina **

Bolotov et al., 2022 [Publ 4252], fig. S1

tT	bi	riS	oT	roT	roS	iD
4	4	0	2	57	0	0	10

  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina **
  |                    
  +-- Sundadontina **

Bolotov et al., 2022 [Publ 4252], fig. 2

tT	bi	riS	oT	roT	roS	iD
4	4	0	2	57	0	0	10

  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina **
  |                    
  +-- Sundadontina **

Bolotov et al., 2022 [Publ 4239], fig. S1

tT	bi	riS	oT	roT	roS	iD
4	4	0	2	54	0	0	10

  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina **
  |                    
  +-- Sundadontina **

Bolotov et al., 2022 [Publ 4239], fig. 2

tT	bi	riS	oT	roT	roS	iD
4	4	0	2	54	0	0	10

  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina **
  |                    
  +-- Sundadontina **

Bolotov et al., 2022 [Publ 4239], fig. S2

tT	bi	riS	oT	roT	roS	iD
4	4	0	2	54	0	0	10

  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina **
  |                    
  +-- Sundadontina **

Kongim et al., 2023 [Publ 4290], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	12	12	3	10

  +----------- Chamberlainiini  
--+                             
  |  +-------- Lamprotulini     
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Indopseudodontina **
        +--+                    
           +-- Pseudodontina **

Konopleva et al., 2019 [Publ 4041], fig. 3

tT	bi	riS	oT	roT	roS	iD
3	3	0	1	31	31	2	11

  +----- Indopseudodontina **
--+                       
  |  +-- Rectidentini     
  +--+                    
     +-- Contradentini

Dai et al., 2023 [Publ 4285], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	51	17	5	--

  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Froufe et al., 2019 [Publ 3936], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	42	11	3	--

  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Chen et al., 2020 [Publ 4170], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	32	9	3	--

  +----- Pseudodontina **  
--+                     
  |  +-- Lamprotulini   
  +--+                  
     +-- Chamberlainiini

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	93	18	5	--

  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Froufe et al., 2019 [Publ 3936], fig. 3-M

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	17	6	3	--

  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Gonideini      
     +--+                  
        +-- Pseudodontina **

Wu et al., 2022 [Publ 4238], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	71	12	3	--

  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Matos et al., 2024 [Publ 4383], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	18	16	5	--

  +----------- Chamberlainiini
  |                           
--+        +-- Contradentini  
  |  +-----+                  
  |  |     +-- Rectidentini   
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Hoeh et al., 2009 [Publ 2716], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	44	2	2	--

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	29	9	4	--

  +-------- Chamberlainiini
  |                        
--+     +-- Gonideini      
  |  +--+                  
  |  |  +-- Pseudodontina **  
  +--+                     
     |  +-- Contradentini  
     +--+                  
        +-- Lamprotulini

Graf & Cummings, 2006 [Publ 1968], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	45	2	2	--

Wu et al., 2024 [Publ 4388], fig. 6

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	52	21	5	--

  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Hoeh et al., 2009 [Publ 2716], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	45	2	2	--

  +-- Gonideini    
--+                
  +-- Pseudodontina **

Wu et al., 2021 [Publ 4366], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	71	12	3	--

  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Zhang et al., 2024 [Publ 4397], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	26	24	5	--

  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	36	10	3	--

  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Lamprotulini

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	34	9	3	--

  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Zieritz et al., 2020 [Publ 4102], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	29	29	6	--

  +----------------- Chamberlainiini  
  |                                   
--+  +-------------- Lamprotulini     
  |  |                                
  +--+  +----------- Gonideini        
     |  |                             
     +--+        +-- Pseudodontina **    
        |  +-----+                    
        |  |     +-- Indopseudodontina **
        +--+                          
           |  +----- Contradentini    
           +--+                       
              |  +-- Ctenodesmini     
              +--+                    
                 +-- Rectidentini

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	46	15	5	--

  +----------- Chamberlainiini
  |                           
--+        +-- Contradentini  
  |  +-----+                  
  |  |     +-- Rectidentini   
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Froufe et al., 2019 [Publ 3936], fig. 6

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	42	11	3	--

  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Froufe et al., 2019 [Publ 3936], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	17	6	3	--

  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Gonideini      
     +--+                  
        +-- Pseudodontina **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	93	18	5	--

  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Graf, 2002 [Publ 2651], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	16	2	2	--

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	21	10	5	--

  +----------- Chamberlainiini
  |                           
--+        +-- Contradentini  
  |  +-----+                  
  |  |     +-- Rectidentini   
  +--+                        
     |  +----- Pseudodontina **  
     +--+                     
        |  +-- Lamprotulini   
        +--+                  
           +-- Gonideini

Dai et al., 2025 [Publ 4440], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	86	17	5	--

Wu et al., 2021 [Publ 4366], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	66	11	3	--

  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Fukata & Iigo, 2020 [Publ 4106], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	51	10	3	--

  +-------- Chamberlainiini
--+                        
  |  +----- Pseudodontina **  
  +--+                     
     |  +-- Lamprotulini   
     +--+                  
        +-- Gonideini

Wu et al., 2024 [Publ 4350], fig. 8b_S2

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	89	19	5	--

  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Matos et al., 2024 [Publ 4384], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	18	16	5	--

  +----------- Chamberlainiini
  |                           
--+        +-- Contradentini  
  |  +-----+                  
  |  |     +-- Rectidentini   
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Hoeh et al., 2009 [Publ 2716], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	44	2	2	--

Dai et al., 2024 [Publ 4358], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	50	9	3	--

  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Wu et al., 2021 [Publ 4366], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	71	12	3	--

  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Wu et al., 2024 [Publ 4350], fig. 8a_S1

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	89	19	5	--

  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Walker et al., 2006 [Publ 2666], fig. 6-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	13	1	1	--

  +-- Gonideini    
--+                
  +-- Pseudodontina **

Dai et al., 2024 [Publ 4386], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	39	9	3	--

  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Walker et al., 2006 [Publ 2666], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	2	1	--

-- Pseudodon_vondembuschianus **

Bolotov et al., 2024 [Publ 4369], fig. S1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	62	0	0	--

-- Indopseudodon_kayinensis **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	5	3	--

-- Pseudodon_vondembuschianus **

Walker et al., 2006 [Publ 2666], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	2	1	--

-- Pseudodon_vondembuschianus **

Walker et al., 2006 [Publ 2666], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	2	2	--

  +-- Gonideini    
--+                
  +-- Pseudodontina **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	5	3	--

  +-- Pseudodontina **
--+                
  +-- Contradentini

Walker et al., 2006 [Publ 2666], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	1	1	--

  +-- Gonideini    
--+                
  +-- Pseudodontina **

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	8	7	2	--

-- Pilsbryoconcha_exilis **

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