Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Quadrulini

subtribe QUADRULINA Ihering, 1901: cladomics

4 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 20 (25 total)
number of subtaxa (rank: genus) = 4
analyzed in 193 cladograms in 116 publications
maximum number of terminals in an analysis = 17 (mean = 2.27, median = 1)
maximum number of subtaxa (genus) = 4 (mean = 1.44, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 65
maximum number of included terminals = 17, (mean = 4.77 median = 2)
maximum number of subtaxa (genus) = 4 (mean = 2.31, median = 2)
proportion of cladograms that support basic monophyly = 0.89

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 27
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 17 (mean = 7.89, median = 6)
maximum number of subtaxa (genus) = 4 (mean = 3.19, median = 3)
proportion of cladograms that support rigorous monophyly = 0.85

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Quadrulina cladogram distances
PCoA Ordination

NJ Tree of Quadrulina cladogram distances
NJ Tree

Cladograms

Neemuchwala et al., 2023 [Publ 4292], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
1.000 17 17 15 4 10 5 2 **
Cladomics cladogram Megalonaias Uniomerus Theliderma Cyclonaias Quadrula Pustulosa

Campbell et al., 2005 [Publ 2660], fig. 1

cst iT tT bi riS oT roT roS iD
0.875 9 9 6 4 93 2 2 15.0
Cladomics cladogram Megalonaias Uniomerus Theliderma Quadrula Pustulosa kieneriana Cyclonaias Pustulosa succissa Pustulosa infucata Pustulosa kleiniana

Campbell et al., 2005 [Publ 2660], fig. 2

cst iT tT bi riS oT roT roS iD
0.750 9 9 6 4 93 2 2 17.0
Cladomics cladogram Megalonaias Uniomerus Theliderma Quadrula quadrula Quadrula nobilis Quadrula verrucosa Pustulosa kieneriana Cyclonaias Pustulosa succissa Pustulosa infucata Pustulosa kleiniana

Johnson et al., 2018 [Publ 3844], fig. 2

cst iT tT bi riS oT roT roS iD
0.875 12 12 10 4 4 2 2 17.0
Cladomics cladogram Megalonaias Uniomerus Theliderma Quadrula Pustulosa kieneriana Pustulosa infucata Pustulosa kleiniana Pustulosa pustulosa Pustulosa succissa Cyclonaias Pustulosa necki Pustulosa nodulata Pustulosa petrina

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

cst iT tT bi riS oT roT roS iD
0.750 14 14 12 4 14 4 2 19.0
Cladomics cladogram Megalonaias Uniomerus Quadrula quadrula Quadrula verrucosa Quadrula nobilis Theliderma Pustulosa kieneriana Pustulosa infucata Pustulosa kleiniana Cyclonaias Pustulosa pustulosa Pustulosa succissa Pustulosa nodulata Pustulosa necki Pustulosa petrina

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

cst iT tT bi riS oT roT roS iD
0.750 17 17 15 4 12 2 2 18.0
Cladomics cladogram Uniomerus Theliderma Quadrula Pustulosa kieneriana Pustulosa infucata Pustulosa kleiniana Pustulosa pustulosa Pustulosa succissa Cyclonaias Pustulosa necki Pustulosa petrina Pustulosa nodulata

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

cst iT tT bi riS oT roT roS iD
0.750 14 14 12 4 12 2 2 19.0
Cladomics cladogram Megalonaias Uniomerus Theliderma Quadrula quadrula Quadrula verrucosa Quadrula nobilis Pustulosa kieneriana Pustulosa infucata Pustulosa kleiniana Pustulosa pustulosa Pustulosa succissa Cyclonaias Pustulosa necki Pustulosa petrina Pustulosa nodulata

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.625 16 16 14 4 617 5 2 19.0
Cladomics cladogram Megalonaias Uniomerus Theliderma johnsoni Theliderma metanevra Quadrula nobilis Quadrula verrucosa Theliderma cylindrica Quadrula fragosa Quadrula quadrula Pustulosa kieneriana Pustulosa infucata Pustulosa kleiniana Cyclonaias Pustulosa pustulosa Pustulosa succissa Pustulosa nodulata Pustulosa necki Pustulosa petrina

Boyer et al., 2011 [Publ 2718], fig. 3

cst iT tT bi riS oT roT roS iD
0.714 6 6 4 4 29 1 1 19.0
Cladomics cladogram Megalonaias Quadrula quadrula Quadrula verrucosa Theliderma Pustulosa pustulosa Cyclonaias Pustulosa nodulata

Boyer et al., 2011 [Publ 2718], fig. 4

cst iT tT bi riS oT roT roS iD
0.714 6 6 4 4 32 1 1 15.0
Cladomics cladogram Theliderma Quadrula Pustulosa pustulosa Cyclonaias Pustulosa nodulata Megalonaias Toxolasma Amblema Reginaia Obliquaria Pleurobema Elliptio Fusconaia Eurynia Potamilus Ellipsaria Truncilla Lampsilis cardium Ligumia Obovaria Epioblasma Lampsilis teres Lampsilis siliquoidea Ortmanniana Lampsilis higginsii

Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2

cst iT tT bi riS oT roT roS iD
0.714 9 9 7 4 64 2 1 18.0
Cladomics cladogram Uniomerus Theliderma Quadrula verrucosa Quadrula quadrula Quadrula nobilis Pustulosa infucata Pustulosa kleiniana Cyclonaias Pustulosa pustulosa Pustulosa succissa

Campbell & Lydeard, 2012 [Publ 3309], fig. 3

cst iT tT bi riS oT roT roS iD
0.714 9 9 6 4 64 2 1 19.0
Cladomics cladogram Uniomerus Quadrula verrucosa Quadrula nobilis Quadrula quadrula Theliderma Pustulosa infucata Pustulosa kleiniana Cyclonaias Pustulosa pustulosa Pustulosa succissa

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

cst iT tT bi riS oT roT roS iD
0.714 5 5 0 4 100 2 1 16.0
Cladomics cladogram Uniomerus Cyclonaias Pustulosa pustulosa 1 Theliderma Pustulosa pustulosa 2 Quadrula quadrula Quadrula verrucosa

Serb et al., 2003 [Publ 2659], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 13 13 10 3 16 1 1 13.0
Cladomics cladogram Megalonaias Theliderma Quadrula Pustulosa

Serb et al., 2003 [Publ 2659], fig. 3

cst iT tT bi riS oT roT roS iD
0.833 13 13 11 3 16 1 1 18.0
Cladomics cladogram Megalonaias Quadrula verrucosa Quadrula nobilis Quadrula quadrula Theliderma Pustulosa

Breton et al., 2011 [Publ 3376], fig. 3

cst iT tT bi riS oT roT roS iD
0.833 3 3 1 3 35 1 1 16.0
Cladomics cladogram Quadrula Cyclonaias Pustulosa Pronodularia Sinohyriopsis CRISTARIINA ALASMIDONTINA Megalonaias Fusconaia Reginaia Toxolasma Ellipsaria Truncilla Lemiox Potamilus Lampsilis Cambarunio Venustaconcha

Campbell & Lydeard, 2012 [Publ 3308], fig. 3

cst iT tT bi riS oT roT roS iD
0.833 6 6 4 3 37 1 1 17.0
Cladomics cladogram Uniomerus Pustulosa infucata Pustulosa kleiniana Quadrula Cyclonaias Pustulosa succissa Pustulosa pustulosa

Szumowski et al., 2012 [Publ 3569], fig. 5

cst iT tT bi riS oT roT roS iD
0.833 13 14 12 3 1 1 1 16.0
Cladomics cladogram Megalonaias Theliderma Quadrula Pustulosa

Pfeiffer et al., 2016 [Publ 3660], fig. 2

cst iT tT bi riS oT roT roS iD
0.667 5 5 3 3 33 1 1 18.0
Cladomics cladogram Uniomerus Quadrula quadrula Quadrula verrucosa Pustulosa pustulosa Pustulosa petrina Cyclonaias

Zhou et al., 2007 [Publ 2658], fig. 2

cst iT tT bi riS oT roT roS iD
0.600 7 7 5 3 11 0 0 18.0
Cladomics cladogram Lamprotula Theliderma Quadrula quadrula Pustulosa infucata Quadrula verrucosa Pustulosa kieneriana Pustulosa succissa

Campbell & Lydeard, 2012 [Publ 3308], fig. 1

cst iT tT bi riS oT roT roS iD
0.800 6 6 4 3 36 0 0 17.0
Cladomics cladogram Quadrula Pustulosa infucata Pustulosa kleiniana Cyclonaias Pustulosa succissa Pustulosa pustulosa Popenaias Amblema Cyrtonaias Reginaia Plectomerus Lemiox Lampsilis Obovaria Elliptoideus Pleurobema stabile Pleurobema pyriforme Pleurobema clava Pleurobema decisum Pleurobema cordatum Pleurobema sintoxia Pleuronaia Parvaspina Fusconaia Eurynia Hemistena Plethobasus Elliptio

Campbell & Lydeard, 2012 [Publ 3308], fig. 2

cst iT tT bi riS oT roT roS iD
0.800 6 6 4 3 36 0 0 17.0
Cladomics cladogram Quadrula Pustulosa infucata Pustulosa kleiniana Cyclonaias Pustulosa succissa Pustulosa pustulosa Reginaia Amblema Popenaias Cyrtonaias Plectomerus Lemiox Lampsilis Obovaria Elliptoideus Hemistena Plethobasus Elliptio Eurynia Parvaspina Pleuronaia Pleurobema Fusconaia

Campbell & Lydeard, 2012 [Publ 3308], fig. 4

cst iT tT bi riS oT roT roS iD
0.800 6 6 4 3 36 0 0 17.0
Cladomics cladogram Quadrula Pustulosa infucata Pustulosa kleiniana Cyclonaias Pustulosa succissa Pustulosa pustulosa Popenaias Amblema Plectomerus Cyrtonaias Reginaia Lemiox Lampsilis Obovaria Elliptoideus Pleurobema stabile Hemistena Eurynia Pleurobema cordatum Pleurobema sintoxia Pleurobema clava Pleurobema decisum Pleurobema pyriforme Parvaspina Pleuronaia Fusconaia Plethobasus Elliptio

Rosenberg et al., 1994 [Publ 2719], fig. 3

cst iT tT bi riS oT roT roS iD
0.667 2 10 0 2 17 2 2 23.0
Cladomics cladogram Elliptio Fusconaia Plectomerus Pleurobema UNIONINA Pyganodon cataracta Pyganodon grandis Utterbackia Lampsilis teres Lampsilis straminea Megalonaias Obliquaria Theliderma Quadrula Uniomerus

Lydeard et al., 1996 [Publ 2656], fig. 3

cst iT tT bi riS oT roT roS iD
0.833 2 2 0 2 25 2 2 --
Cladomics cladogram Megalonaias Quadrula Theliderma

Lydeard et al., 1996 [Publ 2656], fig. 4

cst iT tT bi riS oT roT roS iD
0.833 2 2 0 2 26 2 2 --
Cladomics cladogram Megalonaias Quadrula Theliderma

Rosenberg et al., 1997 [Publ 2720], fig. 2A

cst iT tT bi riS oT roT roS iD
0.667 2 17 0 2 20 2 2 30.0
Cladomics cladogram ANODONTINA Gonidea Pyganodon grandis Utterbackia Pyganodon cataracta Lampsilis straminea Lampsilis teres Megalonaias Obliquaria Theliderma Quadrula Uniomerus Amblema Elliptio Fusconaia Plectomerus Pleurobema UNIONINA

Rosenberg et al., 1997 [Publ 2720], fig. 2B

cst iT tT bi riS oT roT roS iD
0.667 2 17 0 2 20 2 2 30.0
Cladomics cladogram ANODONTINA Gonidea Pyganodon grandis Utterbackia Pyganodon cataracta Lampsilis straminea Lampsilis teres Megalonaias Obliquaria Theliderma Quadrula Uniomerus Amblema Elliptio Fusconaia Plectomerus ACULAMPROTULINA UNIONINA

Krebs et al., 2003 [Publ 2691], fig. 3

cst iT tT bi riS oT roT roS iD
0.667 2 2 1 2 31 2 2 --
Cladomics cladogram Megalonaias Quadrula quadrula 2 Quadrula quadrula 1 Theliderma

Pfeiffer et al., 2019 [Publ 3866], fig. 1

cst iT tT bi riS oT roT roS iD
0.833 3 3 1 2 28 2 2 16.0
Cladomics cladogram Megalonaias Uniomerus Quadrula verrucosa Cyclonaias Quadrula quadrula

Chase et al., 2018 [Publ 3847], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 37 1 1 --
Cladomics cladogram Megalonaias Cyclonaias Pustulosa

Lydeard et al., 2000 [Publ 2654], fig. 2a

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 11 0 0 14.0
Cladomics cladogram Quadrula Pustulosa Pleurobema Fusconaia

Lydeard et al., 2000 [Publ 2654], fig. 2b

cst iT tT bi riS oT roT roS iD
0.750 3 3 0 2 11 0 0 15.0
Cladomics cladogram Pleurobema Fusconaia Quadrula Pustulosa infucata Pustulosa succissa

Lydeard et al., 2000 [Publ 2654], fig. 2c

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 11 0 0 14.0
Cladomics cladogram ANODONTINA Cumberlandia Quadrula Pustulosa Obovaria unicolor Obovaria olivaria Lampsilis Reginaia Pleurobema Fusconaia

Lydeard et al., 2000 [Publ 2654], fig. 2d

cst iT tT bi riS oT roT roS iD
0.750 3 3 0 2 11 0 0 15.0
Cladomics cladogram Pleurobema Fusconaia Quadrula Pustulosa infucata Pustulosa succissa

Curole & Kocher, 2002 [Publ 2712], fig. 1-F

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 2 0 0 --
Cladomics cladogram Lampsilis Quadrula Pustulosa

Curole & Kocher, 2002 [Publ 2712], fig. 1a-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 2 0 0 --
Cladomics cladogram ALASMIDONTINA Lampsilis Quadrula Pustulosa

Curole & Kocher, 2002 [Publ 2712], fig. 1b-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 2 0 0 --
Cladomics cladogram ALASMIDONTINA Quadrula Pustulosa

Curole & Kocher, 2005 [Publ 2682], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 9 0 0 --
Cladomics cladogram Amblema Quadrula Pustulosa

Curole & Kocher, 2005 [Publ 2682], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 10 0 0 --
Cladomics cladogram Quadrula Pustulosa Plectomerus Amblema Potamilus Glebula Leaunio Lampsilis

Chapman et al., 2008 [Publ 2667], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 19 0 0 --
Cladomics cladogram Quadrula Pustulosa Pleurobema Fusconaia Amblema Plectomerus Popenaias Toxolasma Obliquaria Cyrtonaias Glebula Ptychobranchus Lemiox Venustaconcha Obovaria Hamiota Lampsilis ovata Ortmanniana Lampsilis straminea Lampsilis hydiana

Chapman et al., 2008 [Publ 2744], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 19 0 0 --
Cladomics cladogram Quadrula Pustulosa Pleurobema Fusconaia Amblema Plectomerus Popenaias Toxolasma Obliquaria Cyrtonaias Glebula Ptychobranchus Lemiox Venustaconcha Obovaria Hamiota Lampsilis ovata Ortmanniana Lampsilis straminea Lampsilis hydiana

Pieri et al., 2018 [Publ 3867], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 18 0 0 --
Cladomics cladogram Quadrula Pustulosa Eurynia Pleurobema Fusconaia

Inoue et al., 2020 [Publ 3941], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 40 0 0 --
Cladomics cladogram Pustulosa Quadrula Parvaspina Eurynia Hemistena Pleurobema Pleuronaia Fusconaia Amblema Reginaia Toxolasma Potamilus Ellipsaria Truncilla Leaunio Lampsilis bracteata Lampsilis fasciola Lampsilis ornata Lampsilis ovata Lampsilis cariosa Lampsilis cardium 3 Lampsilis cardium 2 Lampsilis satura Lampsilis cardium 1 Lampsilis higginsii Ortmanniana Lampsilis teres Lampsilis radiata Lampsilis siliquoidea Lampsilis bergmanni Lampsilis hydiana Lampsilis virescens Lampsilis straminea

Inoue et al., 2020 [Publ 3941], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 39 0 0 --
Cladomics cladogram Pustulosa Quadrula Reginaia Parvaspina Eurynia Hemistena Pleurobema Pleuronaia Fusconaia Amblema Toxolasma Ellipsaria Truncilla Potamilus Leaunio Lampsilis bracteata Lampsilis ornata Lampsilis fasciola Lampsilis ovata Lampsilis cardium 3 Lampsilis cardium 2 Lampsilis satura Lampsilis cardium 1 Lampsilis teres Lampsilis higginsii Ortmanniana Lampsilis radiata Lampsilis siliquoidea Lampsilis hydiana Lampsilis bergmanni Lampsilis virescens Lampsilis straminea

Porto-Hannes et al., 2021 [Publ 4227], fig. 4

cst iT tT bi riS oT roT roS iD
0.500 2 11 0 2 9 0 0 24.0
Cladomics cladogram Quadrula Pustulosa Pleurobema Amblema Obovaria Ortmanniana Lampsilis

Freeman et al., 2025 [Publ 4443], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 10 0 0 --
Cladomics cladogram Pleurobema Fusconaia Quadrula Pustulosa

Freeman et al., 2025 [Publ 4443], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 10 0 0 --
Cladomics cladogram Pleurobema Fusconaia Quadrula Pustulosa

Freeman et al., 2025 [Publ 4443], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 9 0 0 --
Cladomics cladogram Pleurobema Fusconaia Quadrula Pustulosa

Pfeiffer et al., 2018 [Publ 3835], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 3 2 --
Cladomics cladogram Quadrula Uniomerus Megalonaias

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 199 2 2 --
Cladomics cladogram Quadrula Megalonaias Uniomerus

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 197 2 2 --
Cladomics cladogram Quadrula Megalonaias Uniomerus

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.800 2 2 0 1 199 2 2 --
Cladomics cladogram Chamberlainia Megalonaias Uniomerus Quadrula Sinohyriopsis INDOPSEUDODONTINA PSEUDODONTINA RECTIDENTINA CONTRADENTINA Lamprotula Potomida Pronodularia Sinosolenaia carinata Gonidea Leguminaia Microcondylaea Ptychorhynchus Parvasolenaia Koreosolenaia Sinosolenaia recognita Sinosolenaia iridinea Sinosolenaia oleivora Amblema Toxolasma Obliquaria Plectomerus Elliptoideus Elliptio Eurynia Fusconaia Pleurobema Ellipsaria Potamilus leptodon Medionidus Potamilus alatus Ptychobranchus Truncilla Venustaconcha Cambarunio Leaunio Ortmanniana Lampsilis Acuticosta Alasmidonta Anodonta anatina Anemina Anodonta cygnea Cristaria Brachyanodon Lasmigona Lepidodesma Pseudanodonta Pyganodon Simpsonaias Strophitus Utterbackia Schistodesmus UNIONINA ACULAMPROTULINA Lanceolaria Pletholophus Sinanodonta Cuneopsis celtiformis Tchangsinaia Cuneopsis rufescens Nodularia Cuneopsis szechenyii Cuneopsis heudei

Huang et al., 2019 [Publ 3843], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 55 1 1 --
Cladomics cladogram Quadrula Uniomerus

Burlakova et al., 2019 [Publ 3899], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 32 1 1 --
Cladomics cladogram Quadrula Uniomerus

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 1 1 --
Cladomics cladogram Quadrula Uniomerus

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 1 1 --
Cladomics cladogram Quadrula Uniomerus

Fukata & Iigo, 2020 [Publ 4106], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 53 1 1 --
Cladomics cladogram Quadrula Uniomerus

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 18 1 1 --
Cladomics cladogram Uniomerus Quadrula

Wu et al., 2021 [Publ 4366], fig. 3-BI

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 72 1 1 --
Cladomics cladogram Uniomerus Quadrula

Wu et al., 2021 [Publ 4366], fig. 3-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 72 1 1 --
Cladomics cladogram Uniomerus Quadrula

Wu et al., 2022 [Publ 4238], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 72 1 1 --
Cladomics cladogram Uniomerus Quadrula

Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 1 1 --
Cladomics cladogram Quadrula Uniomerus

Dai et al., 2023 [Publ 4285], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 1 1 --
Cladomics cladogram Quadrula Uniomerus

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 1 1 --
Cladomics cladogram Quadrula Uniomerus

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 1 1 --
Cladomics cladogram Quadrula Uniomerus

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 1 1 --
Cladomics cladogram Quadrula Uniomerus

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 1 1 --
Cladomics cladogram Quadrula Uniomerus

Dai et al., 2024 [Publ 4358], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 1 1 --
Cladomics cladogram Quadrula Uniomerus

Wu et al., 2024 [Publ 4388], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 1 1 --
Cladomics cladogram Quadrula Uniomerus

Dai et al., 2025 [Publ 4440], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 86 1 1 --
Cladomics cladogram Quadrula Uniomerus

Chen et al., 2025 [Publ 4448], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 84 1 1 --
Cladomics cladogram Quadrula Uniomerus

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 139 1 1 --
Cladomics cladogram Quadrula Uniomerus

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 0 0 --
Cladomics cladogram Quadrula Eurynia Pleurobema Fusconaia

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 0 0 --
Cladomics cladogram Quadrula Eurynia Pleurobema Fusconaia

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 0 0 --
Cladomics cladogram Quadrula Eurynia Pleurobema Fusconaia

Turner et al., 2000 [Publ 2722], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram Quadrula ALASMIDONTINA

Hoeh et al., 2001 [Publ 2652], fig. 14f2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 --
Cladomics cladogram Amblema Coelatura Eurynia Fusconaia Gonidea Pleurobema Quadrula Ortmanniana Glebula Potamilus Ligumia Toxolasma Cyrtonaias UNIONINA Pyganodon ANODONTINA Strophitus Mutela Anodontites guanarensis Anodontites trigona Monocondylaea Diplodon Echyridella Lortiella Castalia Velesunio

Hoeh et al., 2001 [Publ 2652], fig. 14f4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 --
Cladomics cladogram Quadrula Eurynia Fusconaia Pleurobema

Kandl et al., 2001 [Publ 3384], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 0 0 --
Cladomics cladogram Amblema Quadrula Elliptio Fusconaia Pleurobema

Hoeh et al., 2002 [Publ 2653], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 --
Cladomics cladogram Quadrula Eurynia Pleurobema Fusconaia

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 --
Cladomics cladogram Quadrula Pleurobema Eurynia Ortmanniana Glebula Potamilus Ligumia Toxolasma Cyrtonaias UNIONINA Pyganodon ANODONTINA Strophitus

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 --
Cladomics cladogram Quadrula Eurynia Fusconaia Pleurobema

Krebs, 2004 [Publ 2692], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 4 0 0 --
Cladomics cladogram Quadrula ALASMIDONTINA Lampsilis Fusconaia

Graf & Cummings, 2006 [Publ 1968], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 45 0 0 --
Cladomics cladogram Quadrula Amblema Eurynia Fusconaia Pleurobema Obliquaria Truncilla Ptychobranchus Ligumia Cambarunio Epioblasma Lampsilis Ortmanniana

Hoeh et al., 2009 [Publ 2716], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 45 0 0 --
Cladomics cladogram Obliquaria Quadrula

Hoeh et al., 2009 [Publ 2716], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 44 0 0 --
Cladomics cladogram Obliquaria Quadrula

Hoeh et al., 2009 [Publ 2716], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 44 0 0 --
Cladomics cladogram Obliquaria Quadrula

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 23 0 0 --
Cladomics cladogram Quadrula Gonidea Eurynia Cambarunio Ptychobranchus Amblema Pleurobema Truncilla Lampsilis Epioblasma Ligumia Obliquaria Ortmanniana

Whelan et al., 2011 [Publ 3380], fig. 1-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 0 0 --
Cladomics cladogram Quadrula Amblema Truncilla Lampsilis

Whelan et al., 2011 [Publ 3380], fig. 1-MP

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 0 0 --
Cladomics cladogram Quadrula Pleurobema Amblema Lampsilis Truncilla

Plazzi et al., 2011 [Publ 3676], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 0 0 --
Cladomics cladogram Quadrula Parvaspina Lampsilis Epioblasma Venustaconcha

Chen et al., 2012 [Publ 3624], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 6 0 0 --
Cladomics cladogram Quadrula Lampsilis Venustaconcha

Inoue et al., 2013 [Publ 3528], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 16 0 0 --
Cladomics cladogram Eurynia Quadrula

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 --
Cladomics cladogram Quadrula Eurynia Amblema Lampsilis Truncilla

Reis et al., 2013 [Publ 3580], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram Amblema Quadrula

Reis et al., 2013 [Publ 3580], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram Amblema Theliderma

Huang et al., 2013 [Publ 3629], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 15 0 0 --
Cladomics cladogram Quadrula Toxolasma Lampsilis Venustaconcha

Huang et al., 2013 [Publ 3629], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram Quadrula Venustaconcha

Inoue et al., 2014 [Publ 3635], fig. 3A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 0 0 --
Cladomics cladogram Quadrula Gonidea ALASMIDONTINA

Sheng et al., 2014 [Publ 3704], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 0 0 --
Cladomics cladogram Venustaconcha Quadrula

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 39 0 0 --
Cladomics cladogram Quadrula Amblema Potamilus Lampsilis Truncilla

Froufe et al., 2015 [Publ 3664], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 0 0 --
Cladomics cladogram Venustaconcha Quadrula

Froufe et al., 2015 [Publ 3664], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 0 0 --
Cladomics cladogram Venustaconcha Quadrula

Xue et al., 2016 [Publ 3659], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 1 0 0 --
Cladomics cladogram Quadrula Venustaconcha

Wei et al., 2016 [Publ 3665], fig. 0

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram CRISTARIINA Lampsilis ALASMIDONTINA Quadrula

Zhou et al., 2016 [Publ 3710], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 0 0 --
Cladomics cladogram Quadrula Toxolasma Venustaconcha Lampsilis

Wu et al., 2016 [Publ 3711], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 20 0 0 --
Cladomics cladogram Quadrula Venustaconcha Lampsilis

Fonseca et al., 2016 [Publ 3775], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 --
Cladomics cladogram Quadrula Toxolasma Lampsilis Venustaconcha

Fonseca et al., 2016 [Publ 3775], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 0 0 --
Cladomics cladogram Venustaconcha Quadrula

Feng et al., 2016 [Publ 3800], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram Quadrula Potamilus Lampsilis

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 79 0 0 --
Cladomics cladogram Quadrula Elliptio Pleurobema Eurynia

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 137 0 0 --
Cladomics cladogram Quadrula Pleurobema Eurynia Elliptio

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 137 0 0 --
Cladomics cladogram Quadrula Pleurobema Elliptio Eurynia

Bolotov et al., 2017 [Publ 3764], fig. S1

cst iT tT bi riS oT roT roS iD
0.000 2 9 0 1 123 0 0 22.0
Cladomics cladogram Amblema Ortmanniana Lampsilis Cambarunio Elliptio Eurynia Pleurobema Quadrula quadrula Quadrula verrucosa Chamberlainia Sinohyriopsis Lamprotula caveata Lamprotula leaii Potomida Pronodularia Gonidea Leguminaia wheatleyi Leguminaia Microcondylaea Sinosolenaia carinata Sinosolenaia oleivora Sinosolenaia recognita

Guerra et al., 2017 [Publ 3772], fig. 2a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 24 0 0 --
Cladomics cladogram Quadrula Toxolasma Venustaconcha Lampsilis

Guerra et al., 2017 [Publ 3772], fig. 2a-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 0 0 --
Cladomics cladogram Venustaconcha Quadrula

Guerra et al., 2017 [Publ 3772], fig. 2b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 24 0 0 --
Cladomics cladogram Quadrula Toxolasma Venustaconcha Lampsilis

Guerra et al., 2017 [Publ 3772], fig. 2b-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 0 0 --
Cladomics cladogram Venustaconcha Quadrula

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 12 0 0 --
Cladomics cladogram Quadrula Venustaconcha

Burzynski et al., 2017 [Publ 3774], fig. 1-top

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 12 0 0 --
Cladomics cladogram Quadrula Venustaconcha

Zhou et al., 2017 [Publ 3788], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 15 0 0 --
Cladomics cladogram Quadrula Potamilus Lampsilis

Amaro et al., 2017 [Publ 3791], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 1 1 14 0 0 --
Cladomics cladogram Quadrula Toxolasma Venustaconcha Lampsilis

Amaro et al., 2017 [Publ 3791], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 0 0 --
Cladomics cladogram Fusconaia Quadrula

Amaro et al., 2017 [Publ 3791], fig. 3A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 0 0 --
Cladomics cladogram Quadrula Toxolasma Venustaconcha

Amaro et al., 2017 [Publ 3791], fig. 3A-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 6 0 0 --
Cladomics cladogram Quadrula Pronodularia Sinosolenaia Venustaconcha ALASMIDONTINA

Amaro et al., 2017 [Publ 3791], fig. 3B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 0 0 --
Cladomics cladogram Quadrula Pronodularia Sinosolenaia

Amaro et al., 2017 [Publ 3791], fig. 3B-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 6 0 0 --
Cladomics cladogram Quadrula Venustaconcha ALASMIDONTINA Pronodularia Sinosolenaia

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 31 0 0 --
Cladomics cladogram Quadrula Toxolasma Lampsilis Venustaconcha

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 0 0 --
Cladomics cladogram Venustaconcha Quadrula

Wen et al., 2017 [Publ 3795], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 15 0 0 --
Cladomics cladogram Quadrula Toxolasma Potamilus Lampsilis Venustaconcha

Wen et al., 2017 [Publ 3795], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram Quadrula Potamilus Venustaconcha

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 6 0 0 --
Cladomics cladogram Venustaconcha Quadrula

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 6 0 0 --
Cladomics cladogram Venustaconcha Quadrula

Bolotov et al., 2018 [Publ 3826], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 81 0 0 --
Cladomics cladogram Lamprotula Potomida Pronodularia Gonidea Leguminaia Microcondylaea Amblema Ortmanniana Lampsilis Cambarunio Elliptio Pleurobema Quadrula

Huang et al., 2018 [Publ 3828], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 41 0 0 --
Cladomics cladogram Quadrula Toxolasma Potamilus Lampsilis Venustaconcha

Huang et al., 2018 [Publ 3828], fig. 6-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 16 0 0 --
Cladomics cladogram Quadrula Venustaconcha Potamilus

Wu et al., 2018 [Publ 3836], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 0 0 --
Cladomics cladogram Quadrula Amblema Obovaria Pleurobema Fusconaia Elliptio Eurynia Potomida Lamprotula Sinosolenaia carinata Gonidea Sinosolenaia oleivora Parvasolenaia Ptychorhynchus Sinohyriopsis Lepidodesma ACULAMPROTULINA UNIONINA OXYNAIINA Acuticosta Lanceolaria CRISTARIINA Anodonta cygnea Anodonta anatina ALASMIDONTINA

Wang et al., 2018 [Publ 3846], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 --
Cladomics cladogram Quadrula Toxolasma Potamilus Venustaconcha Lampsilis

Chase et al., 2018 [Publ 3861], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 0 0 --
Cladomics cladogram Quadrula Lampsilis

Chase et al., 2018 [Publ 3861], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 0 0 --
Cladomics cladogram Quadrula Lampsilis

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 0 0 --
Cladomics cladogram Quadrula Potamilus Venustaconcha

Aunins et al., 2018 [Publ 3863], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 39 0 0 --
Cladomics cladogram Quadrula Toxolasma Lampsilis Venustaconcha

Wang et al., 2018 [Publ 3865], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 --
Cladomics cladogram Quadrula Toxolasma Potamilus Venustaconcha Lampsilis

Cha et al., 2018 [Publ 3916], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 37 0 0 --
Cladomics cladogram Quadrula Toxolasma Lampsilis Potamilus

Wu et al., 2018 [Publ 4143], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 0 0 --
Cladomics cladogram Quadrula Venustaconcha Lampsilis

Huang et al., 2019 [Publ 3843], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 0 0 --
Cladomics cladogram Quadrula Venustaconcha Potamilus

Wu et al., 2019 [Publ 3864], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 45 0 0 --
Cladomics cladogram Quadrula Toxolasma Potamilus Lampsilis Venustaconcha

Froufe et al., 2019 [Publ 3936], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 43 0 0 --
Cladomics cladogram Quadrula Toxolasma Potamilus Lampsilis Venustaconcha

Froufe et al., 2019 [Publ 3936], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 18 0 0 --
Cladomics cladogram Quadrula Potamilus Venustaconcha

Froufe et al., 2019 [Publ 3936], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 18 0 0 --
Cladomics cladogram Quadrula Potamilus Venustaconcha

Froufe et al., 2019 [Publ 3936], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 43 0 0 --
Cladomics cladogram Quadrula Toxolasma Potamilus Lampsilis Venustaconcha

Smith et al., 2019 [Publ 3951], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 0 0 --
Cladomics cladogram Quadrula Amblema Reginaia Plectomerus Toxolasma Popenaias Glebula Cyrtonaias Obliquaria Medionidus Lemiox Cyprogenia Ptychobranchus Potamilus Atlanticoncha Ellipsaria Truncilla Hamiota Villosa Ligumia Venustaconcha Obovaria Epioblasma Ortmanniana pectorosa Ortmanniana ligamentina Lampsilis

Guerra et al., 2019 [Publ 4168], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 0 0 --
Cladomics cladogram Quadrula Toxolasma Venustaconcha Potamilus

Guerra et al., 2019 [Publ 4168], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 0 0 --
Cladomics cladogram Quadrula Venustaconcha Potamilus

Guerra et al., 2019 [Publ 4168], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 0 0 --
Cladomics cladogram Quadrula Toxolasma Venustaconcha Potamilus

Guerra et al., 2019 [Publ 4168], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 0 0 --
Cladomics cladogram Quadrula Venustaconcha Potamilus

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 121 0 0 --
Cladomics cladogram Amblema Elliptio Lampsilis Quadrula

Bolotov et al., 2020 [Publ 4084], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 0 0 --
Cladomics cladogram Amblema Ortmanniana Lampsilis Cambarunio Elliptio Pleurobema Quadrula

Wu et al., 2020 [Publ 4121], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 0 0 --
Cladomics cladogram Quadrula Toxolasma Potamilus Venustaconcha Lampsilis

Inoue et al., 2020 [Publ 4128], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 0 0 --
Cladomics cladogram Quadrula Reginaia Amblema Popenaias Psoronaias Glebula Cyrtonaias Potamilus Ptychobranchus Truncilla Ortmanniana Epioblasma Lampsilis Ligumia Cambarunio Pachynaias Mesonaias

Inoue et al., 2020 [Publ 4128], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 0 0 --
Cladomics cladogram Quadrula Reginaia Amblema Popenaias Psoronaias Cyrtonaias Ptychobranchus Ligumia Potamilus Pachynaias Epioblasma Truncilla Mesonaias Cambarunio Ortmanniana Lampsilis

Smith et al., 2020 [Publ 4129], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 27 0 0 --
Cladomics cladogram Quadrula Pleurobema Elliptoideus Reginaia Amblema Plectomerus Popenaias Psoronaias Obliquaria Cyrtonaias Pachynaias Lampsilis Ellipsaria Truncilla Atlanticoncha Potamilus

Smith et al., 2020 [Publ 4129], fig. 2-3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 27 0 0 --
Cladomics cladogram Quadrula Pleurobema Elliptoideus Reginaia Amblema Plectomerus Popenaias Psoronaias Obliquaria Cyrtonaias Pachynaias Lampsilis Ellipsaria Truncilla Atlanticoncha Potamilus

Smith et al., 2020 [Publ 4136], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 0 0 --
Cladomics cladogram Quadrula Elliptoideus Elliptio Plethobasus Parvaspina Hemistena Eurynia Fusconaia Pleuronaia Pleurobema

Wu et al., 2020 [Publ 4147], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 0 0 --
Cladomics cladogram Quadrula Toxolasma Venustaconcha

Chen et al., 2020 [Publ 4170], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 0 0 --
Cladomics cladogram Quadrula Toxolasma Lampsilis ornata Potamilus Lampsilis siliquoidea Lampsilis powellii

Porto-Hannes et al., 2021 [Publ 4227], fig. 3

cst iT tT bi riS oT roT roS iD
0.000 2 15 0 1 13 0 0 28.0
Cladomics cladogram Quadrula quadrula Quadrula verrucosa Eurynia Pleurobema Amblema Obovaria Ortmanniana Obliquaria Lampsilis ornata Lampsilis radiata 2 Lampsilis siliquoidea Lampsilis radiata 1 Lampsilis virescens Lampsilis hydiana Lampsilis straminea

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 0 0 --
Cladomics cladogram Quadrula Toxolasma Potamilus Venustaconcha Lampsilis

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 0 0 --
Cladomics cladogram Quadrula Venustaconcha Potamilus

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 0 0 --
Cladomics cladogram Quadrula Elliptio Obliquaria Potamilus Lampsilis Truncilla

Wu et al., 2021 [Publ 4366], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 67 0 0 --
Cladomics cladogram Quadrula Pleurobema Amblema Popenaias Toxolasma Potamilus Lampsilis ornata Lampsilis cardium Venustaconcha Lampsilis siliquoidea Lampsilis powellii

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 0 0 --
Cladomics cladogram Quadrula Elliptio

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 74 0 0 --
Cladomics cladogram Quadrula Elliptio Pleurobema Eurynia

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 74 0 0 --
Cladomics cladogram Quadrula Pleurobema Eurynia Elliptio

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 36 0 0 --
Cladomics cladogram Quadrula Toxolasma Potamilus Lampsilis cardium Lampsilis ornata Lampsilis siliquoidea Venustaconcha

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 38 0 0 --
Cladomics cladogram Quadrula Toxolasma Potamilus Lampsilis cardium Lampsilis ornata Lampsilis siliquoidea Venustaconcha

Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram Quadrula Potamilus Venustaconcha Lampsilis

Chen et al., 2023 [Publ 4302], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 0 0 --
Cladomics cladogram Lampsilis Quadrula

Chen et al., 2023 [Publ 4302], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 0 0 --
Cladomics cladogram Lampsilis Quadrula

Hauck et al., 2023 [Publ 4333], fig. 9

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 12 0 0 --
Cladomics cladogram Quadrula Reginaia Obovaria Lampsilis Pleurobema beadleianum Pleurobema oviforme Fusconaia Pleurobema decisum Pleurobema hanleyianum 2 Pleurobema georgianum Pleurobema hanleyianum 1

Wu et al., 2023 [Publ 4339], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 18 0 0 --
Cladomics cladogram Elliptio Quadrula

Dai et al., 2024 [Publ 4386], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 0 0 --
Cladomics cladogram Quadrula Venustaconcha Lampsilis

Hou et al., 2024 [Publ 4400], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 74 0 0 --
Cladomics cladogram Lampsilis Quadrula

Xiang et al., 2025 [Publ 4433], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 12 0 0 --
Cladomics cladogram Quadrula Lanceolaria ANODONTINA CRISTARIINA

Dai et al., 2025 [Publ 4440], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 43 0 0 --
Cladomics cladogram Lampsilis Quadrula

Dai et al., 2025 [Publ 4440], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 43 0 0 --
Cladomics cladogram Lampsilis Quadrula

Hou et al., 2025 [Publ 4445], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 51 0 0 --
Cladomics cladogram Lampsilis Quadrula

Chen et al., 2025 [Publ 4487], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 0 0 --
Cladomics cladogram Lampsilis Quadrula

Chen et al., 2025 [Publ 4487], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 0 0 --
Cladomics cladogram Quadrula Lampsilis

Klymus et al., 2025 [Publ 4814], fig. 3A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 0 0 --
Cladomics cladogram Quadrula Potamilus Venustaconcha Ortmanniana Lampsilis

Klymus et al., 2025 [Publ 4814], fig. 3B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 0 0 --
Cladomics cladogram Quadrula Potamilus Venustaconcha Ortmanniana Lampsilis

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 46 0 0 --
Cladomics cladogram Quadrula Potamilus Venustaconcha Ortmanniana Lampsilis