Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae

tribe QUADRULINI Ihering, 1901: cladomics

5 genera, 1 subtribe

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 26 (46 total)
number of subtaxa (rank: subtribe) = 6
analyzed in 202 cladograms in 122 publications
maximum number of terminals in an analysis = 22 (mean = 2.63, median = 1)
maximum number of subtaxa (subtribe) = 3 (mean = 1.36, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 89
maximum number of included terminals = 22, (mean = 4.71 median = 2)
maximum number of subtaxa (subtribe) = 3 (mean = 1.82, median = 2)
proportion of cladograms that support basic monophyly = 0.83

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 52
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (subtribe) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 22 (mean = 5.96, median = 4)
maximum number of subtaxa (subtribe) = 3 (mean = 2.35, median = 2)
proportion of cladograms that support rigorous monophyly = 0.77

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Quadrulini cladogram distances
PCoA Ordination

NJ Tree of Quadrulini cladogram distances
NJ Tree

Cladograms

Neemuchwala et al., 2023 [Publ 4292], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
1.000 22 22 20 3 5 4 4 **
Cladomics cladogram Megalonaias Uniomerus QUADRULINA PLEUROBEMINI AMBLEMINI LAMPSILINI POPENAIADINI

Campbell et al., 2005 [Publ 2660], fig. 1

cst iT tT bi riS oT roT roS iD
0.857 11 11 8 3 91 79 4 20.0
Cladomics cladogram QUADRULINA Megalonaias Uniomerus PLEUROBEMINI Plectomerus Reginaia LAMPSILINI POPENAIADINI Amblema

Campbell et al., 2005 [Publ 2660], fig. 2

cst iT tT bi riS oT roT roS iD
0.796 11 11 8 3 91 79 4 20.0
Cladomics cladogram Megalonaias Uniomerus QUADRULINA PLEUROBEMINI POPENAIADINI Amblema Reginaia Plectomerus Toxolasma Obliquaria Cyrtonaias Glebula Sagittunio Leaunio Potamilus Hamiota Ellipsaria Truncilla Lampsilis teres Ortmanniana ligamentina Lampsilis siliquoidea Villosa Ligumia Cambarunio Ortmanniana pectorosa Lampsilis ornata Lampsilis cardium Lampsilis ovata Obovaria olivaria Venustaconcha ellipsiformis 2 Epioblasma Obovaria unicolor Obovaria arkansasensis Obovaria subrotunda Ptychobranchus Venustaconcha ellipsiformis 1 Medionidus conradicus Lemiox Medionidus acutissimus Dromus Cyprogenia

Pfeiffer et al., 2019 [Publ 3866], fig. 1

cst iT tT bi riS oT roT roS iD
0.938 5 5 3 3 26 25 4 19.0
Cladomics cladogram Megalonaias Uniomerus QUADRULINA PLEUROBEMINI AMBLEMINI POPENAIADINI LAMPSILINI

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.636 21 21 21 3 612 170 4 25.0
Cladomics cladogram PLEUROBEMINI QUADRULINA Megalonaias Uniomerus Amblema Nephronaias Psoronaias Reginaia Popenaias Cyrtonaias Glebula Paetulunio Toxolasma parvum Toxolasma lividum 2 Toxolasma cylindrellus Toxolasma lividum 1 Toxolasma texasiense 2 Toxolasma glans Toxolasma paulum Toxolasma texasiense 1 Toxolasma 4 Toxolasma granulatum Toxolasma 2 Toxolasma 3 Toxolasma pullus Toxolasma germanum Toxolasma 1 Obliquaria Plectomerus Potamilus inflatus Potamilus ohiensis Potamilus amphichaenus Potamilus streckersoni Potamilus fragilis 2 Potamilus leptodon Potamilus fragilis 1 Potamilus metnecktayi Potamilus alatus 3 Potamilus alatus 2 Potamilus purpuratus 2 Potamilus alatus 1 Potamilus purpuratus 1 Villosa villosa Atlanticoncha Cambarunio nebulosus Ligumia Leaunio umbrans Cambarunio taeniatus Cambarunio iris Cambarunio simus Hamiota Pachynaias Sagittunio Ellipsaria Potamilus capax Truncilla Medionidus acutissimus Lemiox Medionidus penicillatus Ptychobranchus Cyprogenia aberti 2 Dromus Cyprogenia stegaria 2 Cyprogenia aberti 1 Cyprogenia stegaria 1 Villosa delumbis Villosa vibex Obovaria olivaria Venustaconcha Epioblasma Obovaria retusa Obovaria choctawensis Obovaria unicolor Obovaria subrotunda Mesonaias Obovaria arkansasensis Lampsilis splendida Lampsilis bracteata Lampsilis fasciola Ortmanniana pectorosa Lampsilis ornata Lampsilis ovata 2 Lampsilis dolabraeformis Lampsilis cardium 2 Lampsilis cardium 1 Lampsilis satura Lampsilis cariosa 2 Lampsilis ovata 1 Lampsilis cariosa 1 Ortmanniana ligamentina 2 Lampsilis abrupta Lampsilis higginsii 2 Ortmanniana ligamentina 1 Lampsilis higginsii 1 Medionidus conradicus Leaunio lienosus Leaunio vanuxemensis Lampsilis floridensis Lampsilis teres Lampsilis siliquoidea 1 Lampsilis siliquoidea 2 Lampsilis powellii 1 Lampsilis powellii 2 Lampsilis sietmani Lampsilis straminea 2 Lampsilis virescens Lampsilis hydiana 2 Lampsilis hydiana 1 Lampsilis bergmanni Lampsilis straminea 1

Rosenberg et al., 1994 [Publ 2719], fig. 3

cst iT tT bi riS oT roT roS iD
0.625 4 10 0 3 15 7 3 26.0
Cladomics cladogram Elliptio Fusconaia AMBLEMINI Pleurobema UNIONINI Pyganodon cataracta Pyganodon grandis Utterbackia Lampsilis teres Lampsilis straminea Megalonaias Obliquaria Theliderma Quadrula Uniomerus

Lydeard et al., 1996 [Publ 2656], fig. 3

cst iT tT bi riS oT roT roS iD
0.773 4 18 14 3 23 14 3 34.0
Cladomics cladogram ANODONTINI Megalonaias QUADRULINA Elliptoideus Eurynia Pleurobema Uniomerus Amblema Plectomerus Obliquaria Potamilus fragilis Ptychobranchus Medionidus Potamilus purpuratus Lampsilis Ellipsaria Villosa

Lydeard et al., 1996 [Publ 2656], fig. 4

cst iT tT bi riS oT roT roS iD
0.818 4 18 16 3 24 14 3 34.0
Cladomics cladogram ANODONTINI Megalonaias QUADRULINA Amblema Plectomerus LAMPSILINI Elliptoideus Uniomerus Eurynia Pleurobema

Rosenberg et al., 1997 [Publ 2720], fig. 2A

cst iT tT bi riS oT roT roS iD
0.588 4 17 0 3 18 8 3 33.0
Cladomics cladogram ANODONTINA GONIDEINI Pyganodon grandis Utterbackia Pyganodon cataracta Lampsilis straminea Lampsilis teres Megalonaias Obliquaria Theliderma Quadrula Uniomerus Amblema Elliptio Fusconaia Plectomerus Pleurobema UNIONINI

Rosenberg et al., 1997 [Publ 2720], fig. 2B

cst iT tT bi riS oT roT roS iD
0.562 4 17 0 3 18 7 3 33.0
Cladomics cladogram ANODONTINA GONIDEINI Pyganodon grandis Utterbackia Pyganodon cataracta Lampsilis straminea Lampsilis teres Megalonaias Obliquaria Theliderma Quadrula Uniomerus Amblema Elliptio Fusconaia Plectomerus ACULAMPROTULINA UNIONINA

Krebs et al., 2003 [Publ 2691], fig. 3

cst iT tT bi riS oT roT roS iD
0.708 4 21 20 3 29 17 3 38.0
Cladomics cladogram ANODONTINI Megalonaias QUADRULINA Elliptoideus Pleurobema Uniomerus Fusconaia Eurynia Amblema Toxolasma Plectomerus Obliquaria Ptychobranchus Potamilus fragilis Medionidus Villosa Ellipsaria Lampsilis teres Lampsilis siliquoidea Potamilus purpuratus Potamilus alatus

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
0.893 4 4 2 3 197 21 3 20.0
Cladomics cladogram QUADRULINA Megalonaias Uniomerus PLEUROBEMINI Amblema Toxolasma Obliquaria Plectomerus Potamilus leptodon Potamilus alatus Venustaconcha Leaunio Ellipsaria Truncilla Medionidus Ptychobranchus Cambarunio Ortmanniana Lampsilis

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
0.893 4 4 2 3 195 21 3 20.0
Cladomics cladogram QUADRULINA Megalonaias Uniomerus PLEUROBEMINI Amblema Toxolasma Obliquaria Plectomerus Potamilus leptodon Potamilus alatus Medionidus Ptychobranchus Ellipsaria Truncilla Leaunio Venustaconcha Cambarunio Ortmanniana Lampsilis

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.821 4 128 73 3 197 21 3 145.0
Cladomics cladogram PARREYSIINI LAMELLIDENTINI LEOPARREYSIINI COELATURINI INDOCHINELLINI Chamberlainia Megalonaias Uniomerus QUADRULINA Sinohyriopsis LAMPROTULINI GONIDEINI PSEUDODONTINI RECTIDENTINI Amblema Toxolasma PLEUROBEMINI Obliquaria Plectomerus Ellipsaria Potamilus leptodon Medionidus Potamilus alatus Ptychobranchus Truncilla Venustaconcha Cambarunio Leaunio Ortmanniana Lampsilis ACUTICOSTINI Alasmidonta Anodonta anatina Anemina Anodonta cygnea Cristaria Brachyanodon Lasmigona LEPIDODESMINI Pseudanodonta Pyganodon Simpsonaias Strophitus Utterbackia Schistodesmus UNIONINA ACULAMPROTULINA LANCEOLARIINI Pletholophus Sinanodonta Cuneopsis celtiformis Tchangsinaia Cuneopsis rufescens Nodularia Cuneopsis szechenyii Cuneopsis heudei

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

cst iT tT bi riS oT roT roS iD
0.882 18 18 15 3 10 6 3 23.0
Cladomics cladogram AMBLEMINI LAMPSILINI PLEUROBEMINI QUADRULINA Megalonaias Uniomerus

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

cst iT tT bi riS oT roT roS iD
0.875 19 25 23 3 10 6 3 27.0
Cladomics cladogram ANODONTINI Megalonaias Uniomerus QUADRULINA PLEUROBEMINI LAMPSILINI AMBLEMINI

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

cst iT tT bi riS oT roT roS iD
0.875 16 16 14 3 10 6 3 22.0
Cladomics cladogram AMBLEMINI PLEUROBEMINI LAMPSILINI Megalonaias Uniomerus QUADRULINA

Johnson et al., 2018 [Publ 3844], fig. 2

cst iT tT bi riS oT roT roS iD
0.923 14 14 12 3 2 2 2 20.0
Cladomics cladogram AMBLEMINI PLEUROBEMINI Megalonaias Uniomerus QUADRULINA

Pfeiffer et al., 2018 [Publ 3835], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 3 0 0 0 22.0
Cladomics cladogram QUADRULINA Uniomerus Megalonaias

Campbell & Lydeard, 2012 [Publ 3308], fig. 3

cst iT tT bi riS oT roT roS iD
0.786 7 7 5 2 36 36 4 21.0
Cladomics cladogram Uniomerus QUADRULINA Elliptoideus Plectomerus Obovaria unicolor Obovaria olivaria Lampsilis Lemiox Cyrtonaias Reginaia POPENAIADINI Amblema Elliptio Eurynia Parvaspina Hemistena Plethobasus Pleuronaia Pleurobema cordatum Fusconaia Pleurobema stabile Pleurobema sintoxia Pleurobema decisum Pleurobema clava Pleurobema pyriforme

Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2

cst iT tT bi riS oT roT roS iD
0.806 11 11 9 2 62 62 4 21.0
Cladomics cladogram Uniomerus QUADRULINA PLEUROBEMINI Plectomerus Reginaia LAMPSILINI POPENAIADINI Amblema

Campbell & Lydeard, 2012 [Publ 3309], fig. 3

cst iT tT bi riS oT roT roS iD
0.833 11 11 8 2 62 62 4 22.0
Cladomics cladogram Uniomerus QUADRULINA PLEUROBEMINI Amblema Reginaia POPENAIADINI Glebula Paetulunio Toxolasma Plectomerus Obliquaria Potamilus Dromus Lemiox Lampsilis ovata Lampsilis virescens Epioblasma Obovaria arkansasensis Obovaria olivaria Venustaconcha

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

cst iT tT bi riS oT roT roS iD
0.758 7 7 2 2 98 76 4 19.0
Cladomics cladogram Reginaia Amblema PLEUROBEMINI Uniomerus QUADRULINA POPENAIADINI Dromus Epioblasma Glebula Lampsilis ovata Lampsilis virescens Lemiox Obliquaria Obovaria olivaria Obovaria unicolor Plectomerus Potamilus alatus Potamilus capax

Burlakova et al., 2019 [Publ 3899], fig. 1

cst iT tT bi riS oT roT roS iD
0.906 2 2 0 2 31 31 4 --
Cladomics cladogram QUADRULINA Uniomerus Plectomerus POPENAIADINI PLEUROBEMINI Cyrtonaias Glebula Reginaia Obliquaria Toxolasma Potamilus Ellipsaria Truncilla Medionidus Lemiox Dromus Cambarunio Leaunio Villosa Sagittunio Lampsilis ovata Venustaconcha Epioblasma Obovaria Lampsilis teres Lampsilis siliquoidea Lampsilis hydiana Lampsilis straminea

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 55 13 4 --
Cladomics cladogram QUADRULINA Uniomerus PLEUROBEMINI AMBLEMINI POPENAIADINI LAMPSILINI

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 55 13 4 --
Cladomics cladogram QUADRULINA Uniomerus PLEUROBEMINI AMBLEMINI POPENAIADINI LAMPSILINI

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

cst iT tT bi riS oT roT roS iD
0.941 2 2 0 2 17 12 4 --
Cladomics cladogram Uniomerus QUADRULINA PLEUROBEMINI AMBLEMINI POPENAIADINI LAMPSILINI

Wu et al., 2021 [Publ 4366], fig. 3-BI

cst iT tT bi riS oT roT roS iD
0.944 2 2 0 2 71 13 4 --
Cladomics cladogram Uniomerus QUADRULINA PLEUROBEMINI AMBLEMINI POPENAIADINI LAMPSILINI

Wu et al., 2021 [Publ 4366], fig. 3-ML

cst iT tT bi riS oT roT roS iD
0.944 2 2 0 2 71 13 4 --
Cladomics cladogram Uniomerus QUADRULINA PLEUROBEMINI AMBLEMINI POPENAIADINI LAMPSILINI

Wu et al., 2022 [Publ 4238], fig. 3

cst iT tT bi riS oT roT roS iD
0.944 2 2 0 2 71 13 4 --
Cladomics cladogram Uniomerus QUADRULINA PLEUROBEMINI AMBLEMINI POPENAIADINI LAMPSILINI

Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F

cst iT tT bi riS oT roT roS iD
0.882 2 2 0 2 13 13 4 --
Cladomics cladogram Toxolasma POPENAIADINI AMBLEMINI PLEUROBEMINI QUADRULINA Uniomerus Potamilus Lampsilis cardium Lampsilis ornata Venustaconcha Lampsilis siliquoidea Lampsilis powellii

Dai et al., 2023 [Publ 4285], fig. 5

cst iT tT bi riS oT roT roS iD
0.944 2 2 0 2 51 13 4 --
Cladomics cladogram QUADRULINA Uniomerus PLEUROBEMINI AMBLEMINI POPENAIADINI LAMPSILINI

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cst iT tT bi riS oT roT roS iD
0.944 2 2 0 2 93 15 4 --
Cladomics cladogram QUADRULINA Uniomerus PLEUROBEMINI AMBLEMINI POPENAIADINI LAMPSILINI

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cst iT tT bi riS oT roT roS iD
0.944 2 2 0 2 93 15 4 --
Cladomics cladogram QUADRULINA Uniomerus PLEUROBEMINI AMBLEMINI POPENAIADINI LAMPSILINI

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cst iT tT bi riS oT roT roS iD
0.944 2 2 0 2 89 15 4 --
Cladomics cladogram QUADRULINA Uniomerus PLEUROBEMINI AMBLEMINI POPENAIADINI LAMPSILINI

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cst iT tT bi riS oT roT roS iD
0.944 2 2 0 2 89 15 4 --
Cladomics cladogram QUADRULINA Uniomerus PLEUROBEMINI AMBLEMINI POPENAIADINI LAMPSILINI

Dai et al., 2024 [Publ 4358], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 51 10 4 --
Cladomics cladogram QUADRULINA Uniomerus PLEUROBEMINI AMBLEMINI POPENAIADINI LAMPSILINI

Dai et al., 2025 [Publ 4440], fig. 3

cst iT tT bi riS oT roT roS iD
0.944 2 2 0 2 85 14 4 --
Cladomics cladogram QUADRULINA Uniomerus PLEUROBEMINI AMBLEMINI POPENAIADINI LAMPSILINI

Chen et al., 2025 [Publ 4448], fig. 3

cst iT tT bi riS oT roT roS iD
0.944 2 2 0 2 83 13 4 --
Cladomics cladogram QUADRULINA Uniomerus PLEUROBEMINI AMBLEMINI POPENAIADINI LAMPSILINI

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cst iT tT bi riS oT roT roS iD
0.947 2 2 0 2 138 16 4 --
Cladomics cladogram QUADRULINA Uniomerus PLEUROBEMINI LAMPSILINI AMBLEMINI POPENAIADINI

Huang et al., 2002 [Publ 2657], fig. 1

cst iT tT bi riS oT roT roS iD
0.722 2 14 0 2 32 12 3 32.0
Cladomics cladogram GONIDEINI Megalonaias Elliptoideus Uniomerus Pleurobema Amblema Obliquaria Plectomerus Medionidus Potamilus fragilis Ptychobranchus Potamilus purpuratus Lampsilis Villosa Ellipsaria

Huang et al., 2002 [Publ 2657], fig. 2

cst iT tT bi riS oT roT roS iD
0.722 2 14 0 2 32 12 3 32.0
Cladomics cladogram Ptychorhynchus CHAMBERLAINIINI LAMPROTULINI Sinosolenaia Megalonaias Elliptoideus Uniomerus Pleurobema Potamilus fragilis Medionidus Amblema Obliquaria Plectomerus Villosa Ellipsaria Lampsilis Potamilus purpuratus Ptychobranchus

Serb et al., 2003 [Publ 2659], fig. 2

cst iT tT bi riS oT roT roS iD
0.870 14 14 11 2 15 15 3 17.0
Cladomics cladogram Lampsilis Obliquaria Amblema Reginaia Potamilus Elliptoideus Plectomerus Megalonaias QUADRULINA Medionidus Dromus Cyprogenia Hemistena Elliptio Fusconaia

Serb et al., 2003 [Publ 2659], fig. 3

cst iT tT bi riS oT roT roS iD
0.826 14 14 12 2 15 15 3 22.0
Cladomics cladogram Lampsilis Potamilus Medionidus Dromus Cyprogenia Obliquaria Amblema Reginaia Elliptoideus Plectomerus Megalonaias QUADRULINA Hemistena Elliptio Fusconaia

Boyer et al., 2011 [Publ 2718], fig. 3

cst iT tT bi riS oT roT roS iD
0.852 7 7 5 2 28 22 3 23.0
Cladomics cladogram Reginaia Amblema LAMPSILINI PLEUROBEMINI Megalonaias QUADRULINA

Boyer et al., 2011 [Publ 2718], fig. 4

cst iT tT bi riS oT roT roS iD
0.815 7 29 25 2 31 22 3 41.0
Cladomics cladogram ANODONTINI QUADRULINA Megalonaias Toxolasma Amblema PLEUROBEMINI Reginaia Obliquaria Potamilus Ellipsaria Truncilla Lampsilis cardium Ligumia Obovaria Epioblasma Lampsilis teres Lampsilis siliquoidea Ortmanniana Lampsilis higginsii

Breton et al., 2011 [Publ 3376], fig. 3

cst iT tT bi riS oT roT roS iD
0.900 4 33 32 2 34 15 3 50.0
Cladomics cladogram Cumberlandia Margaritifera QUADRULINA LAMPROTULINI CHAMBERLAINIINI ANODONTINI Megalonaias PLEUROBEMINI AMBLEMINI LAMPSILINI

Pfeiffer et al., 2016 [Publ 3660], fig. 2

cst iT tT bi riS oT roT roS iD
0.852 6 6 4 2 32 31 3 22.0
Cladomics cladogram AMBLEMINI LAMPSILINI Elliptoideus Uniomerus QUADRULINA Elliptio Parvaspina Plethobasus Hemistena Eurynia Fusconaia Pleurobema riddellii Pleurobema sintoxia Pleurobema cordatum Pleuronaia Pleurobema pyriforme Pleurobema strodeanum Pleurobema clava

Chase et al., 2018 [Publ 3847], fig. 4

cst iT tT bi riS oT roT roS iD
0.947 3 3 1 2 36 15 3 19.0
Cladomics cladogram PLEUROBEMINI Megalonaias QUADRULINA

Smith, 2021 [Publ 4331], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 18 7 3 --
Cladomics cladogram Uniomerus Megalonaias PLEUROBEMINI AMBLEMINI LAMPSILINI

Wu et al., 2024 [Publ 4388], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 51 8 3 --
Cladomics cladogram QUADRULINA Uniomerus PLEUROBEMINI POPENAIADINI LAMPSILINI

Huang et al., 2019 [Publ 3843], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 54 8 2 --
Cladomics cladogram QUADRULINA Uniomerus PLEUROBEMINI LAMPSILINI

Fukata & Iigo, 2020 [Publ 4106], fig. 1

cst iT tT bi riS oT roT roS iD
0.917 2 2 0 2 52 9 2 --
Cladomics cladogram QUADRULINA Uniomerus PLEUROBEMINI LAMPSILINI

Szumowski et al., 2012 [Publ 3569], fig. 5

cst iT tT bi riS oT roT roS iD
0.857 14 14 12 2 0 0 0 20.0
Cladomics cladogram Megalonaias Theliderma Quadrula Pustulosa

Chapman et al., 2008 [Publ 2667], fig. 1

cst iT tT bi riS oT roT roS iD
0.920 2 2 0 1 19 18 4 --
Cladomics cladogram QUADRULINA PLEUROBEMINI Amblema Plectomerus POPENAIADINI LAMPSILINI

Chapman et al., 2008 [Publ 2744], fig. 2

cst iT tT bi riS oT roT roS iD
0.920 2 2 0 1 19 18 4 --
Cladomics cladogram QUADRULINA PLEUROBEMINI Amblema Plectomerus POPENAIADINI LAMPSILINI

Campbell & Lydeard, 2012 [Publ 3308], fig. 1

cst iT tT bi riS oT roT roS iD
0.852 6 6 4 1 36 36 4 22.0
Cladomics cladogram QUADRULINA PLEUROBEMINI POPENAIADINI Amblema Cyrtonaias Reginaia Plectomerus Lemiox Lampsilis Obovaria

Campbell & Lydeard, 2012 [Publ 3308], fig. 2

cst iT tT bi riS oT roT roS iD
0.889 6 6 4 1 36 36 4 22.0
Cladomics cladogram QUADRULINA PLEUROBEMINI Reginaia Amblema POPENAIADINI Cyrtonaias Plectomerus Lemiox Lampsilis Obovaria

Campbell & Lydeard, 2012 [Publ 3308], fig. 4

cst iT tT bi riS oT roT roS iD
0.852 6 6 4 1 36 36 4 22.0
Cladomics cladogram QUADRULINA PLEUROBEMINI POPENAIADINI Amblema Plectomerus Cyrtonaias Reginaia Lemiox Lampsilis Obovaria

Smith et al., 2020 [Publ 4129], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 27 27 4 --
Cladomics cladogram QUADRULINA PLEUROBEMINI AMBLEMINI POPENAIADINI LAMPSILINI

Smith et al., 2020 [Publ 4129], fig. 2-3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 27 27 4 --
Cladomics cladogram QUADRULINA PLEUROBEMINI AMBLEMINI POPENAIADINI LAMPSILINI

Wu et al., 2021 [Publ 4366], fig. 4

cst iT tT bi riS oT roT roS iD
0.933 1 1 0 1 67 11 4 --
Cladomics cladogram QUADRULINA PLEUROBEMINI AMBLEMINI POPENAIADINI LAMPSILINI

Freeman et al., 2025 [Publ 4443], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 10 10 4 --
Cladomics cladogram POPENAIADINI LAMPSILINI AMBLEMINI PLEUROBEMINI QUADRULINA

Freeman et al., 2025 [Publ 4443], fig. 4

cst iT tT bi riS oT roT roS iD
0.947 2 2 0 1 10 10 4 --
Cladomics cladogram AMBLEMINI PLEUROBEMINI QUADRULINA Obliquaria Toxolasma POPENAIADINI Glebula Cyrtonaias

Freeman et al., 2025 [Publ 4443], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 9 9 4 --
Cladomics cladogram POPENAIADINI LAMPSILINI AMBLEMINI PLEUROBEMINI QUADRULINA

Barraza et al., 2026 [Publ 4851], fig. 3

cst iT tT bi riS oT roT roS iD
0.824 1 1 0 1 18 11 4 --
Cladomics cladogram Amblema Psoronaias Nephronaias Reginaia QUADRULINA PLEUROBEMINI Mesonaias Pachynaias Cyrtonaias Popenaias

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

cst iT tT bi riS oT roT roS iD
0.950 1 1 0 1 29 16 3 --
Cladomics cladogram QUADRULINA PLEUROBEMINI AMBLEMINI LAMPSILINI

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

cst iT tT bi riS oT roT roS iD
0.950 1 1 0 1 29 16 3 --
Cladomics cladogram QUADRULINA PLEUROBEMINI AMBLEMINI LAMPSILINI

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

cst iT tT bi riS oT roT roS iD
0.941 1 1 0 1 33 10 3 --
Cladomics cladogram Toxolasma AMBLEMINI Glebula Cyrtonaias QUADRULINA PLEUROBEMINI Potamilus Ortmanniana Ligumia

Lydeard et al., 2000 [Publ 2654], fig. 2a

cst iT tT bi riS oT roT roS iD
0.923 3 3 1 1 11 9 3 19.0
Cladomics cladogram QUADRULINA PLEUROBEMINI

Lydeard et al., 2000 [Publ 2654], fig. 2b

cst iT tT bi riS oT roT roS iD
0.923 3 3 0 1 11 9 3 20.0
Cladomics cladogram LAMPSILINI AMBLEMINI QUADRULINA PLEUROBEMINI

Lydeard et al., 2000 [Publ 2654], fig. 2c

cst iT tT bi riS oT roT roS iD
0.846 3 3 1 1 11 9 3 19.0
Cladomics cladogram ANODONTINI Cumberlandia QUADRULINA LAMPSILINI AMBLEMINI PLEUROBEMINI

Lydeard et al., 2000 [Publ 2654], fig. 2d

cst iT tT bi riS oT roT roS iD
0.923 3 3 0 1 11 9 3 20.0
Cladomics cladogram AMBLEMINI LAMPSILINI QUADRULINA PLEUROBEMINI

Turner et al., 2000 [Publ 2722], fig. 2

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 13 11 3 --
Cladomics cladogram QUADRULINA ANODONTINI

Hoeh et al., 2001 [Publ 2652], fig. 14f2

cst iT tT bi riS oT roT roS iD
0.824 1 1 0 1 30 10 3 --
Cladomics cladogram AMBLEMINI COELATURINI Eurynia Fusconaia GONIDEINI Pleurobema QUADRULINA Ortmanniana Glebula Potamilus Ligumia Toxolasma Cyrtonaias UNIONINI ANODONTINI Mutela Anodontites guanarensis Anodontites trigona Monocondylaea RHIPIDODONTINI Echyridella Lortiella CASTALIINI Velesunio

Hoeh et al., 2001 [Publ 2652], fig. 14f4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 10 3 --
Cladomics cladogram AMBLEMINI LAMPSILINI QUADRULINA PLEUROBEMINI

Hoeh et al., 2002 [Publ 2653], fig. 2

cst iT tT bi riS oT roT roS iD
0.941 1 1 0 1 30 10 3 --
Cladomics cladogram Glebula Cyrtonaias Toxolasma Potamilus Ligumia Ortmanniana AMBLEMINI QUADRULINA PLEUROBEMINI

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

cst iT tT bi riS oT roT roS iD
0.824 1 1 0 1 30 10 3 --
Cladomics cladogram QUADRULINA Pleurobema Eurynia Cyrtonaias UNIONINI ANODONTINI Ortmanniana Glebula Potamilus Ligumia Toxolasma

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 10 3 --
Cladomics cladogram AMBLEMINI LAMPSILINI QUADRULINA PLEUROBEMINI

Graf & Cummings, 2006 [Publ 1968], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 45 12 3 --
Cladomics cladogram QUADRULINA AMBLEMINI PLEUROBEMINI LAMPSILINI

Campbell et al., 2008 [Publ 2675], fig. 4

cst iT tT bi riS oT roT roS iD
0.812 1 1 0 1 24 24 3 --
Cladomics cladogram Uniomerus AMBLEMINI LAMPSILINI Eurynia Fusconaia cerina Fusconaia flava Hemistena Plethobasus Parvaspina Pleurobema cordatum Pleuronaia Pleurobema oviforme 2 Pleurobema stabile Elliptio Pleurobema decisum Pleurobema sintoxia Pleurobema clava Pleurobema oviforme 1 Pleurobema beadleianum Pleurobema pyriforme Pleurobema rubellum Pleurobema hanleyianum 2 Pleurobema georgianum 2 Pleurobema hanleyianum 1 Pleurobema georgianum 1 Pleurobema athearni

Campbell et al., 2008 [Publ 2675], fig. 5

cst iT tT bi riS oT roT roS iD
0.750 1 1 0 1 24 24 3 --
Cladomics cladogram Uniomerus AMBLEMINI Elliptio arctata Elliptio crassidens Pleurobema decisum Pleurobema rubellum Pleurobema georgianum 2 Pleurobema stabile Pleurobema pyriforme Pleurobema oviforme 2 Pleurobema hanleyianum 2 Pleuronaia Pleurobema georgianum 3 Pleurobema cordatum Pleurobema beadleianum Hemistena Fusconaia flava Fusconaia cerina Eurynia Elliptio arca LAMPSILINI Plethobasus Parvaspina Pleurobema sintoxia Pleurobema oviforme 1 Pleurobema clava Pleurobema athearni Pleurobema georgianum 1 Pleurobema hanleyianum 1

Hoeh et al., 2009 [Publ 2716], fig. 3

cst iT tT bi riS oT roT roS iD
0.947 2 2 0 1 45 12 3 --
Cladomics cladogram AMBLEMINI PLEUROBEMINI Obliquaria QUADRULINA Ptychobranchus Truncilla Ortmanniana Lampsilis Cambarunio Ligumia Epioblasma

Hoeh et al., 2009 [Publ 2716], fig. 5

cst iT tT bi riS oT roT roS iD
0.944 2 2 0 1 44 11 3 --
Cladomics cladogram Obliquaria QUADRULINA AMBLEMINI PLEUROBEMINI Truncilla Ptychobranchus Ortmanniana Cambarunio Ligumia Epioblasma

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

cst iT tT bi riS oT roT roS iD
0.833 2 2 0 1 23 11 3 --
Cladomics cladogram QUADRULINA GONIDEINI Eurynia Cambarunio Ptychobranchus AMBLEMINI Pleurobema Truncilla Lampsilis Epioblasma Ligumia Obliquaria Ortmanniana

Whelan et al., 2011 [Publ 3380], fig. 1-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 4 3 --
Cladomics cladogram PLEUROBEMINI QUADRULINA AMBLEMINI LAMPSILINI

Whelan et al., 2011 [Publ 3380], fig. 1-MP

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 4 3 --
Cladomics cladogram QUADRULINA PLEUROBEMINI AMBLEMINI LAMPSILINI

Inoue et al., 2013 [Publ 3528], fig. 3

cst iT tT bi riS oT roT roS iD
0.938 1 1 0 1 16 14 3 --
Cladomics cladogram PLEUROBEMINI QUADRULINA Paetulunio AMBLEMINI Lampsilis Leaunio Obovaria Villosa Cambarunio

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 4 3 --
Cladomics cladogram QUADRULINA PLEUROBEMINI AMBLEMINI LAMPSILINI

Inoue et al., 2014 [Publ 3635], fig. 3A

cst iT tT bi riS oT roT roS iD
0.900 1 1 0 1 19 3 3 --
Cladomics cladogram QUADRULINA GONIDEINI ANODONTINI

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 79 7 3 --
Cladomics cladogram AMBLEMINI LAMPSILINI QUADRULINA PLEUROBEMINI

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 137 7 3 --
Cladomics cladogram AMBLEMINI LAMPSILINI QUADRULINA PLEUROBEMINI

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 137 7 3 --
Cladomics cladogram AMBLEMINI LAMPSILINI QUADRULINA PLEUROBEMINI

Bolotov et al., 2017 [Publ 3764], fig. S1

cst iT tT bi riS oT roT roS iD
0.643 2 9 0 1 123 7 3 27.0
Cladomics cladogram AMBLEMINI Ortmanniana Lampsilis Cambarunio Elliptio Eurynia Pleurobema Quadrula quadrula Quadrula verrucosa Chamberlainia Sinohyriopsis Lamprotula caveata Lamprotula leaii Potomida Pronodularia Gonidea Leguminaia wheatleyi Leguminaia Microcondylaea Sinosolenaia carinata Sinosolenaia oleivora Sinosolenaia recognita

Bolotov et al., 2018 [Publ 3826], fig. 2

cst iT tT bi riS oT roT roS iD
0.769 1 1 0 1 81 6 3 --
Cladomics cladogram Lamprotula Potomida Pronodularia Gonidea Leguminaia Microcondylaea AMBLEMINI Ortmanniana Lampsilis Cambarunio Elliptio Pleurobema QUADRULINA

Wu et al., 2018 [Publ 3836], fig. 2

cst iT tT bi riS oT roT roS iD
0.923 1 1 0 1 52 8 3 --
Cladomics cladogram QUADRULINA PLEUROBEMINI AMBLEMINI LAMPSILINI LAMPROTULINI GONIDEINI CHAMBERLAINIINI LEPIDODESMINI UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Smith et al., 2019 [Publ 3951], fig. 2

cst iT tT bi riS oT roT roS iD
0.903 1 1 0 1 40 40 3 --
Cladomics cladogram QUADRULINA Amblema Reginaia Plectomerus Toxolasma POPENAIADINI Glebula Cyrtonaias Obliquaria Medionidus Lemiox Cyprogenia Ptychobranchus Potamilus Atlanticoncha Ellipsaria Truncilla Hamiota Villosa Ligumia Venustaconcha Obovaria Epioblasma Ortmanniana pectorosa Ortmanniana ligamentina Lampsilis

Inoue et al., 2020 [Publ 3941], fig. 3

cst iT tT bi riS oT roT roS iD
0.957 2 2 0 1 40 39 3 --
Cladomics cladogram QUADRULINA PLEUROBEMINI AMBLEMINI LAMPSILINI

Inoue et al., 2020 [Publ 3941], fig. 4

cst iT tT bi riS oT roT roS iD
0.913 2 2 0 1 39 38 3 --
Cladomics cladogram QUADRULINA Reginaia PLEUROBEMINI Amblema LAMPSILINI

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 121 3 3 --
Cladomics cladogram AMBLEMINI PLEUROBEMINI LAMPSILINI QUADRULINA

Bolotov et al., 2020 [Publ 4084], fig. 1

cst iT tT bi riS oT roT roS iD
0.846 1 1 0 1 35 6 3 --
Cladomics cladogram AMBLEMINI Ortmanniana Lampsilis Cambarunio Elliptio Pleurobema QUADRULINA

Inoue et al., 2020 [Publ 4128], fig. 2

cst iT tT bi riS oT roT roS iD
0.957 1 1 0 1 22 22 3 --
Cladomics cladogram QUADRULINA Reginaia Amblema POPENAIADINI LAMPSILINI

Inoue et al., 2020 [Publ 4128], fig. 3

cst iT tT bi riS oT roT roS iD
0.955 1 1 0 1 21 21 3 --
Cladomics cladogram QUADRULINA Reginaia Amblema POPENAIADINI LAMPSILINI

Porto-Hannes et al., 2021 [Publ 4227], fig. 3

cst iT tT bi riS oT roT roS iD
0.714 2 15 0 1 13 13 3 33.0
Cladomics cladogram Quadrula quadrula Quadrula verrucosa PLEUROBEMINI AMBLEMINI LAMPSILINI

Porto-Hannes et al., 2021 [Publ 4227], fig. 4

cst iT tT bi riS oT roT roS iD
0.846 2 11 0 1 9 9 3 29.0
Cladomics cladogram Quadrula Pustulosa PLEUROBEMINI AMBLEMINI LAMPSILINI

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 74 7 3 --
Cladomics cladogram AMBLEMINI LAMPSILINI QUADRULINA PLEUROBEMINI

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 74 7 3 --
Cladomics cladogram AMBLEMINI LAMPSILINI QUADRULINA PLEUROBEMINI

Hauck et al., 2023 [Publ 4333], fig. 9

cst iT tT bi riS oT roT roS iD
0.917 1 1 0 1 12 12 3 --
Cladomics cladogram QUADRULINA PLEUROBEMINI AMBLEMINI LAMPSILINI

Kandl et al., 2001 [Publ 3384], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 8 2 --
Cladomics cladogram AMBLEMINI QUADRULINA PLEUROBEMINI

Krebs, 2004 [Publ 2692], fig. 2

cst iT tT bi riS oT roT roS iD
0.875 1 1 0 1 4 2 2 --
Cladomics cladogram QUADRULINA ANODONTINI LAMPSILINI PLEUROBEMINI

Curole & Kocher, 2005 [Publ 2682], fig. 1

cst iT tT bi riS oT roT roS iD
0.769 2 2 0 1 9 7 2 --
Cladomics cladogram Glebula Amblema QUADRULINA Plectomerus Leaunio Lampsilis straminea Lampsilis teres Potamilus

Curole & Kocher, 2005 [Publ 2682], fig. 1-M

cst iT tT bi riS oT roT roS iD
0.923 2 2 0 1 10 7 2 --
Cladomics cladogram QUADRULINA Plectomerus Amblema LAMPSILINI

Hoeh et al., 2009 [Publ 2716], fig. 4

cst iT tT bi riS oT roT roS iD
0.941 2 2 0 1 44 11 2 --
Cladomics cladogram PLEUROBEMINI Obliquaria QUADRULINA Ptychobranchus Truncilla Ortmanniana Lampsilis Cambarunio Ligumia Epioblasma

Plazzi et al., 2011 [Publ 3676], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 4 2 --
Cladomics cladogram QUADRULINA PLEUROBEMINI LAMPSILINI

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 39 4 2 --
Cladomics cladogram QUADRULINA AMBLEMINI LAMPSILINI

Perkins et al., 2017 [Publ 3777], fig. 3

cst iT tT bi riS oT roT roS iD
0.769 1 1 0 1 23 23 2 --
Cladomics cladogram AMBLEMINI Uniomerus PLEUROBEMINI

Amaro et al., 2017 [Publ 3791], fig. 2

cst iT tT bi riS oT roT roS iD
0.900 2 2 1 1 14 4 2 --
Cladomics cladogram LAMPSILINI QUADRULINA

Amaro et al., 2017 [Publ 3791], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 3 2 --
Cladomics cladogram LAMPSILINI PLEUROBEMINI QUADRULINA

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 5 2 --
Cladomics cladogram QUADRULINA PLEUROBEMINI LAMPSILINI

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 5 2 --
Cladomics cladogram LAMPSILINI QUADRULINA PLEUROBEMINI

Wu et al., 2023 [Publ 4339], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 18 5 2 --
Cladomics cladogram LAMPSILINI PLEUROBEMINI QUADRULINA

Mulvey et al., 1997 [Publ 3390], fig. 1_3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 4 3 1 --
Cladomics cladogram Megalonaias AMBLEMINI

Kandl et al., 2001 [Publ 3384], fig. 4

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 5 5 1 --
Cladomics cladogram Elliptio Uniomerus Pleurobema

Curole & Kocher, 2002 [Publ 2712], fig. 1-F

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 2 1 1 --
Cladomics cladogram LAMPSILINI QUADRULINA

Curole & Kocher, 2002 [Publ 2712], fig. 1a-M

cst iT tT bi riS oT roT roS iD
0.857 2 2 0 1 2 1 1 --
Cladomics cladogram QUADRULINA ANODONTINI LAMPSILINI

Curole & Kocher, 2002 [Publ 2712], fig. 1b-M

cst iT tT bi riS oT roT roS iD
0.857 2 2 0 1 2 1 1 --
Cladomics cladogram ANODONTINI QUADRULINA

Chen et al., 2012 [Publ 3624], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 6 2 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Reis et al., 2013 [Publ 3580], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 1 1 --
Cladomics cladogram AMBLEMINI QUADRULINA

Reis et al., 2013 [Publ 3580], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 1 1 --
Cladomics cladogram AMBLEMINI QUADRULINA

Huang et al., 2013 [Publ 3629], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 15 3 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Huang et al., 2013 [Publ 3629], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 1 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Sheng et al., 2014 [Publ 3704], fig. 2

cst iT tT bi riS oT roT roS iD
0.714 1 1 0 1 7 2 1 --
Cladomics cladogram LAMPSILINI QUADRULINA

Froufe et al., 2015 [Publ 3664], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 1 1 --
Cladomics cladogram LAMPSILINI QUADRULINA

Froufe et al., 2015 [Publ 3664], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 1 1 --
Cladomics cladogram LAMPSILINI QUADRULINA

Xue et al., 2016 [Publ 3659], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 1 1 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Wei et al., 2016 [Publ 3665], fig. 0

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 5 1 1 --
Cladomics cladogram CRISTARIINA LAMPSILINI ALASMIDONTINA QUADRULINA

Zhou et al., 2016 [Publ 3710], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 3 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Wu et al., 2016 [Publ 3711], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 20 2 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Fonseca et al., 2016 [Publ 3775], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 3 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Fonseca et al., 2016 [Publ 3775], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 1 1 --
Cladomics cladogram LAMPSILINI QUADRULINA

Feng et al., 2016 [Publ 3800], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 2 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Guerra et al., 2017 [Publ 3772], fig. 2a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 24 3 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Guerra et al., 2017 [Publ 3772], fig. 2a-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 1 1 --
Cladomics cladogram LAMPSILINI QUADRULINA

Guerra et al., 2017 [Publ 3772], fig. 2b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 24 3 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Guerra et al., 2017 [Publ 3772], fig. 2b-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 1 1 --
Cladomics cladogram LAMPSILINI QUADRULINA

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 12 1 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Burzynski et al., 2017 [Publ 3774], fig. 1-top

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 12 1 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Zhou et al., 2017 [Publ 3788], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 15 2 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Amaro et al., 2017 [Publ 3791], fig. 3A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 2 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Amaro et al., 2017 [Publ 3791], fig. 3A-M

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 6 1 1 --
Cladomics cladogram QUADRULINA LAMPROTULINI GONIDEINI LAMPSILINI ANODONTINI

Amaro et al., 2017 [Publ 3791], fig. 3B

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 11 2 1 --
Cladomics cladogram QUADRULINA LAMPROTULINI GONIDEINI

Amaro et al., 2017 [Publ 3791], fig. 3B-M

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 6 1 1 --
Cladomics cladogram QUADRULINA LAMPSILINI ANODONTINI LAMPROTULINI GONIDEINI

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 31 3 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 1 1 --
Cladomics cladogram LAMPSILINI QUADRULINA

Wen et al., 2017 [Publ 3795], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 15 5 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Wen et al., 2017 [Publ 3795], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 2 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 6 1 1 --
Cladomics cladogram LAMPSILINI QUADRULINA

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 6 1 1 --
Cladomics cladogram LAMPSILINI QUADRULINA

Huang et al., 2018 [Publ 3828], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 41 5 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Huang et al., 2018 [Publ 3828], fig. 6-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 16 2 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Wang et al., 2018 [Publ 3846], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 5 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Chase et al., 2018 [Publ 3861], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 2 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Chase et al., 2018 [Publ 3861], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 2 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 2 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Aunins et al., 2018 [Publ 3863], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 39 3 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Wang et al., 2018 [Publ 3865], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 5 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Pieri et al., 2018 [Publ 3867], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 18 17 1 --
Cladomics cladogram QUADRULINA PLEUROBEMINI

Cha et al., 2018 [Publ 3916], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 37 4 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Wu et al., 2018 [Publ 4143], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 2 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Huang et al., 2019 [Publ 3843], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 2 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Wu et al., 2019 [Publ 3864], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 45 5 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Froufe et al., 2019 [Publ 3936], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 43 5 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Froufe et al., 2019 [Publ 3936], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 18 2 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Froufe et al., 2019 [Publ 3936], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 18 2 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Froufe et al., 2019 [Publ 3936], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 43 5 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Guerra et al., 2019 [Publ 4168], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 3 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Guerra et al., 2019 [Publ 4168], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 2 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Guerra et al., 2019 [Publ 4168], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 3 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Guerra et al., 2019 [Publ 4168], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 2 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Wu et al., 2020 [Publ 4121], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 5 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Smith et al., 2020 [Publ 4136], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 19 1 --
Cladomics cladogram QUADRULINA PLEUROBEMINI

Wu et al., 2020 [Publ 4147], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 2 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Chen et al., 2020 [Publ 4170], fig. 1

cst iT tT bi riS oT roT roS iD
0.500 1 1 0 1 33 6 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 5 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 2 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

cst iT tT bi riS oT roT roS iD
0.889 1 1 0 1 36 6 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

cst iT tT bi riS oT roT roS iD
0.889 1 1 0 1 38 6 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 5 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Chen et al., 2023 [Publ 4302], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 1 1 --
Cladomics cladogram LAMPSILINI QUADRULINA

Chen et al., 2023 [Publ 4302], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 1 1 --
Cladomics cladogram LAMPSILINI QUADRULINA

Dai et al., 2024 [Publ 4386], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 3 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Hou et al., 2024 [Publ 4400], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 74 1 1 --
Cladomics cladogram LAMPSILINI QUADRULINA

Dai et al., 2025 [Publ 4440], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 43 1 1 --
Cladomics cladogram LAMPSILINI QUADRULINA

Dai et al., 2025 [Publ 4440], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 43 1 1 --
Cladomics cladogram LAMPSILINI QUADRULINA

Hou et al., 2025 [Publ 4445], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 51 1 1 --
Cladomics cladogram LAMPSILINI QUADRULINA

Chen et al., 2025 [Publ 4487], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 1 1 --
Cladomics cladogram LAMPSILINI QUADRULINA

Chen et al., 2025 [Publ 4487], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 1 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Klymus et al., 2025 [Publ 4814], fig. 3A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 5 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Klymus et al., 2025 [Publ 4814], fig. 3B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 5 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 46 6 1 --
Cladomics cladogram QUADRULINA LAMPSILINI

Zhou et al., 2007 [Publ 2658], fig. 2

cst iT tT bi riS oT roT roS iD
0.667 7 7 5 1 11 0 0 23.0
Cladomics cladogram LAMPROTULINI QUADRULINA

Xiang et al., 2025 [Publ 4433], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 12 0 0 --
Cladomics cladogram QUADRULINA LANCEOLARIINI ANODONTINI