Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Unionini

subtribe UNIONINA Rafinesque, 1820: cladomics

1 genus

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 228 (330 total)
number of subtaxa (rank: genus) = 1
analyzed in 239 cladograms in 138 publications
maximum number of terminals in an analysis = 26 (mean = 3.45, median = 3)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 165
maximum number of included terminals = 26, (mean = 4.55 median = 3)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)
proportion of cladograms that support basic monophyly = 0.93

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Unionina cladogram distances
PCoA Ordination

NJ Tree of Unionina cladogram distances
NJ Tree

Cladograms

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
1.000 19 19 17 1 121 27 4 **
Cladomics cladogram ACULAMPROTULINA GLOBUNIONINA OXYNAIINA MIDDENDORFFINAIINA Unio

Dai et al., 2025 [Publ 4440], fig. 1

cst iT tT bi riS oT roT roS iD
0.941 2 2 0 1 42 19 4 --
Cladomics cladogram GLOBUNIONINA ACULAMPROTULINA MIDDENDORFFINAIINA Nodularia Cuneopsis Diaurora Schistodesmus Pseudocuneopsis Unio Tchangsinaia Inversiunio

Dai et al., 2025 [Publ 4440], fig. 2

cst iT tT bi riS oT roT roS iD
0.941 2 2 0 1 42 19 4 --
Cladomics cladogram GLOBUNIONINA ACULAMPROTULINA MIDDENDORFFINAIINA Nodularia Cuneopsis Diaurora Schistodesmus Pseudocuneopsis Inversiunio Tchangsinaia Unio

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.833 26 26 24 1 607 40 4 23.0
Cladomics cladogram GLOBUNIONINA ACULAMPROTULINA MIDDENDORFFINAIINA Unio OXYNAIINA

Wu et al., 2026 [Publ 4811], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 91 28 4 15.0
Cladomics cladogram GLOBUNIONINA ACULAMPROTULINA Unio MIDDENDORFFINAIINA OXYNAIINA

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
1.000 4 4 1 1 118 24 3 16.0
Cladomics cladogram ACULAMPROTULINA MIDDENDORFFINAIINA Unio OXYNAIINA

Konopleva et al., 2023 [Publ 4269], fig. S1_4

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 53 9 3 16.0
Cladomics cladogram MIDDENDORFFINAIINA ACULAMPROTULINA Unio OXYNAIINA

Chen et al., 2023 [Publ 4302], fig. 1

cst iT tT bi riS oT roT roS iD
0.933 2 2 0 1 39 16 3 --
Cladomics cladogram ACULAMPROTULINA MIDDENDORFFINAIINA Nodularia Cuneopsis Diaurora Schistodesmus Pseudocuneopsis Inversiunio Tchangsinaia Unio

Chen et al., 2023 [Publ 4302], fig. 2

cst iT tT bi riS oT roT roS iD
0.933 2 2 0 1 39 16 3 --
Cladomics cladogram ACULAMPROTULINA MIDDENDORFFINAIINA Nodularia Cuneopsis Diaurora Schistodesmus Pseudocuneopsis Inversiunio Tchangsinaia Unio

Dai et al., 2024 [Publ 4386], fig. 3

cst iT tT bi riS oT roT roS iD
0.909 1 1 0 1 13 10 3 --
Cladomics cladogram Unio MIDDENDORFFINAIINA OXYNAIINA ACULAMPROTULINA ANODONTINA Lepidodesma

Dai et al., 2024 [Publ 4386], fig. 5-BI

cst iT tT bi riS oT roT roS iD
0.917 1 1 0 1 21 11 3 --
Cladomics cladogram Unio MIDDENDORFFINAIINA OXYNAIINA

Dai et al., 2024 [Publ 4386], fig. 5-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 11 3 --
Cladomics cladogram ACULAMPROTULINA Unio MIDDENDORFFINAIINA OXYNAIINA

Hou et al., 2024 [Publ 4400], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 19 19 17 1 56 29 3 10.0
Cladomics cladogram ACULAMPROTULINA Unio MIDDENDORFFINAIINA OXYNAIINA

Dai et al., 2025 [Publ 4440], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 1 81 21 3 17.0
Cladomics cladogram GLOBUNIONINA ACULAMPROTULINA Unio OXYNAIINA

Chen et al., 2025 [Publ 4487], fig. 2

cst iT tT bi riS oT roT roS iD
0.938 2 2 0 1 39 16 3 --
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Chen et al., 2025 [Publ 4487], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 39 16 3 --
Cladomics cladogram GLOBUNIONINA ACULAMPROTULINA Unio OXYNAIINA

Wang et al., 2025 [Publ 4488], fig. 3B

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 60 28 3 15.0
Cladomics cladogram ACULAMPROTULINA Unio MIDDENDORFFINAIINA OXYNAIINA

Wu et al., 2026 [Publ 4825], fig. 6B

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 61 25 3 15.0
Cladomics cladogram ACULAMPROTULINA Unio MIDDENDORFFINAIINA OXYNAIINA

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 4 2 --
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Xue et al., 2016 [Publ 3659], fig. 1

cst iT tT bi riS oT roT roS iD
0.875 1 1 0 1 14 3 2 --
Cladomics cladogram ACULAMPROTULINA Schistodesmus Nodularia Unio

Wu et al., 2016 [Publ 3711], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 20 4 2 --
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Fonseca et al., 2016 [Publ 3775], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 29 5 2 --
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
1.000 4 4 1 1 77 7 2 16.0
Cladomics cladogram Unio ACULAMPROTULINA OXYNAIINA

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 136 7 2 16.0
Cladomics cladogram Unio ACULAMPROTULINA OXYNAIINA

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
0.800 3 3 1 1 136 7 2 16.0
Cladomics cladogram Unio ACULAMPROTULINA Schistodesmus Nodularia Cuneopsis szechenyii Tchangsinaia Cuneopsis rufescens

Bolotov et al., 2017 [Publ 3764], fig. S1

cst iT tT bi riS oT roT roS iD
0.500 3 10 0 1 122 7 2 24.0
Cladomics cladogram Unio crassus Unio pictorum Unio tumidus ACULAMPROTULINA Cuneopsis szechenyii Tchangsinaia Cuneopsis rufescens Nodularia douglasiae Nodularia nuxpersicae Schistodesmus Alasmidonta Anodonta anatina Anodonta cygnea Brachyanodon Lasmigona Pseudanodonta Pyganodon Simpsonaias Strophitus Sinanodonta lauta Cristaria plicata Cristaria radiata Pletholophus Sinanodonta lucida Sinanodonta jourdyi Lanceolaria gladiola Lanceolaria grayii Lanceolaria danzhouensis

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 30 5 2 --
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Araujo et al., 2017 [Publ 3801], fig. 2

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 13 2 2 --
Cladomics cladogram ANODONTINA Unio

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 19 6 2 15.0
Cladomics cladogram ACULAMPROTULINA OXYNAIINA Unio

Huang et al., 2018 [Publ 3828], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 38 5 2 15.0
Cladomics cladogram ACULAMPROTULINA OXYNAIINA Unio

Wu et al., 2018 [Publ 3836], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 51 12 2 --
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Wang et al., 2018 [Publ 3846], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 27 4 2 15.0
Cladomics cladogram ACULAMPROTULINA OXYNAIINA Unio

Aunins et al., 2018 [Publ 3863], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 36 7 2 15.0
Cladomics cladogram ACULAMPROTULINA OXYNAIINA Unio

Wang et al., 2018 [Publ 3865], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 27 3 2 15.0
Cladomics cladogram ACULAMPROTULINA OXYNAIINA Unio

Cha et al., 2018 [Publ 3916], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 34 6 2 15.0
Cladomics cladogram ACULAMPROTULINA OXYNAIINA Unio

Wu et al., 2018 [Publ 4143], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 5 2 --
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 197 17 2 15.0
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Huang et al., 2019 [Publ 3843], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 52 7 2 15.0
Cladomics cladogram ACULAMPROTULINA OXYNAIINA Unio

Huang et al., 2019 [Publ 3843], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 20 2 2 16.0
Cladomics cladogram OXYNAIINA ACULAMPROTULINA Unio

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 195 16 2 15.0
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.700 4 4 2 1 197 17 2 15.0
Cladomics cladogram Acuticosta Alasmidonta Anodonta anatina Anemina Anodonta cygnea Cristaria Brachyanodon Lasmigona Lepidodesma Pseudanodonta Pyganodon Simpsonaias Strophitus Utterbackia Schistodesmus Unio ACULAMPROTULINA Lanceolaria Pletholophus Sinanodonta Cuneopsis celtiformis Tchangsinaia Cuneopsis rufescens Nodularia Cuneopsis szechenyii Cuneopsis heudei

Wu et al., 2019 [Publ 3864], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 42 9 2 15.0
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Klishko et al., 2019 [Publ 3886], fig. 9

cst iT tT bi riS oT roT roS iD
1.000 4 4 1 1 7 4 2 16.0
Cladomics cladogram Unio MIDDENDORFFINAIINA OXYNAIINA

Froufe et al., 2019 [Publ 3936], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 40 6 2 15.0
Cladomics cladogram ACULAMPROTULINA OXYNAIINA Unio

Froufe et al., 2019 [Publ 3936], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 40 6 2 15.0
Cladomics cladogram ACULAMPROTULINA OXYNAIINA Unio

Fukata & Iigo, 2019 [Publ 4108], fig. 1

cst iT tT bi riS oT roT roS iD
0.909 4 4 2 1 13 9 2 15.0
Cladomics cladogram Unio OXYNAIINA

Guerra et al., 2019 [Publ 4168], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 38 2 2 16.0
Cladomics cladogram ACULAMPROTULINA OXYNAIINA Unio

Guerra et al., 2019 [Publ 4168], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 19 2 2 16.0
Cladomics cladogram OXYNAIINA ACULAMPROTULINA Unio

Guerra et al., 2019 [Publ 4168], fig. 2

cst iT tT bi riS oT roT roS iD
0.857 3 3 1 1 38 2 2 16.0
Cladomics cladogram OXYNAIINA Unio

Guerra et al., 2019 [Publ 4168], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 19 2 2 16.0
Cladomics cladogram ACULAMPROTULINA OXYNAIINA Unio

Bolotov et al., 2020 [Publ 4084], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 33 2 2 16.0
Cladomics cladogram MIDDENDORFFINAIINA OXYNAIINA Unio

Fukata & Iigo, 2020 [Publ 4106], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 50 9 2 15.0
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Wu et al., 2020 [Publ 4121], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 44 9 2 15.0
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Sano et al., 2020 [Publ 4138], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 66 10 2 16.0
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Sano et al., 2020 [Publ 4138], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 66 10 2 16.0
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Sano et al., 2020 [Publ 4138], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 66 10 2 16.0
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Wu et al., 2020 [Publ 4147], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 10 2 --
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Sayenko et al., 2021 [Publ 4189], fig. 6

cst iT tT bi riS oT roT roS iD
0.875 2 2 0 1 11 11 2 --
Cladomics cladogram Unio Inversiunio MIDDENDORFFINAIINA Nodularia

Kim et al., 2021 [Publ 4197], fig. 1

cst iT tT bi riS oT roT roS iD
0.889 4 4 2 1 14 8 2 15.0
Cladomics cladogram Unio OXYNAIINA

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 44 6 2 15.0
Cladomics cladogram ACULAMPROTULINA OXYNAIINA Unio

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 89 9 2 --
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Wu et al., 2021 [Publ 4366], fig. 3-BI

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 69 11 2 15.0
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Wu et al., 2021 [Publ 4366], fig. 3-ML

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 69 11 2 15.0
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Wu et al., 2021 [Publ 4366], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 64 11 2 15.0
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 89 9 2 --
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Wu et al., 2022 [Publ 4238], fig. 2d

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 34 19 2 15.0
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Wu et al., 2022 [Publ 4238], fig. 2e

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 15 10 2 16.0
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Wu et al., 2022 [Publ 4238], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 69 16 2 15.0
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

cst iT tT bi riS oT roT roS iD
0.900 3 3 1 1 73 7 2 16.0
Cladomics cladogram Unio ACULAMPROTULINA OXYNAIINA

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

cst iT tT bi riS oT roT roS iD
0.900 3 3 1 1 73 7 2 16.0
Cladomics cladogram Unio ACULAMPROTULINA OXYNAIINA

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 36 6 2 --
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

cst iT tT bi riS oT roT roS iD
0.889 1 1 0 1 38 6 2 --
Cladomics cladogram Unio OXYNAIINA

Dai et al., 2023 [Publ 4285], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 6 2 --
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Wu et al., 2023 [Publ 4296], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 12 2 --
Cladomics cladogram Unio ACULAMPROTULINA OXYNAIINA

Liu et al., 2023 [Publ 4310], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 23 13 2 --
Cladomics cladogram Unio ACULAMPROTULINA OXYNAIINA

Liu et al., 2023 [Publ 4310], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 23 13 2 --
Cladomics cladogram Unio ACULAMPROTULINA OXYNAIINA

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 1 89 16 2 15.0
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cst iT tT bi riS oT roT roS iD
0.909 6 6 3 1 89 16 2 14.0
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 1 85 16 2 15.0
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 1 85 16 2 15.0
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Dai et al., 2024 [Publ 4358], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 51 16 2 --
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Wu et al., 2024 [Publ 4379], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 1 47 23 2 17.0
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Dai et al., 2024 [Publ 4386], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 1 35 15 2 17.0
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Wu et al., 2024 [Publ 4388], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 8 2 --
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Hou et al., 2025 [Publ 4445], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 1 46 26 2 17.0
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Chen et al., 2025 [Publ 4448], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 81 23 2 15.0
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Klymus et al., 2025 [Publ 4814], fig. 3A

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 32 6 2 15.0
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Klymus et al., 2025 [Publ 4814], fig. 3B

cst iT tT bi riS oT roT roS iD
0.909 4 4 2 1 32 6 2 15.0
Cladomics cladogram Unio ACULAMPROTULINA Lanceolaria

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 43 10 2 15.0
Cladomics cladogram ACULAMPROTULINA Unio OXYNAIINA

Rosenberg et al., 1997 [Publ 2720], fig. 2B

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 21 1 1 --
Cladomics cladogram Gonidea Pyganodon grandis Utterbackia Pyganodon cataracta Lampsilis straminea Lampsilis teres Megalonaias Obliquaria Theliderma Quadrula Uniomerus Amblema Elliptio Fusconaia Plectomerus ACULAMPROTULINA Unio

Araujo et al., 2009 [Publ 2589], fig. 7A

cst iT tT bi riS oT roT roS iD
0.400 6 7 3 1 5 1 1 18.0
Cladomics cladogram OXYNAIINA Unio gibbus Unio tumidus ANODONTINA Unio pictorum 1 Unio pictorum 2 Unio mancus Unio tumidiformis Unio crassus

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

cst iT tT bi riS oT roT roS iD
0.400 2 7 5 1 23 1 1 25.0
Cladomics cladogram Anodonta cygnea Unio pictorum 2 Unio pictorum 1 Unio caffer Anodonta ALASMIDONTINA

Zhou et al., 2016 [Publ 3710], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 1 1 --
Cladomics cladogram OXYNAIINA Unio

Froufe et al., 2016 [Publ 3738], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 12 12 10 1 2 2 1 23.0
Cladomics cladogram OXYNAIINA Unio

Guerra et al., 2017 [Publ 3772], fig. 2a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 24 1 1 --
Cladomics cladogram ACULAMPROTULINA Unio

Guerra et al., 2017 [Publ 3772], fig. 2b

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 24 1 1 --
Cladomics cladogram Unio ACULAMPROTULINA CRISTARIINA ANODONTINA ALASMIDONTINA

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 11 1 1 --
Cladomics cladogram ACULAMPROTULINA Unio

Burzynski et al., 2017 [Publ 3774], fig. 1-top

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 11 1 1 --
Cladomics cladogram ACULAMPROTULINA Unio

Wen et al., 2017 [Publ 3795], fig. 3

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 15 2 1 --
Cladomics cladogram Lepidodesma Unio

Liu et al., 2017 [Publ 3939], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 4 4 1 1 11 7 1 16.0
Cladomics cladogram Unio OXYNAIINA

Klishko et al., 2017 [Publ 3946], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 2 2 1 15.0
Cladomics cladogram OXYNAIINA Unio

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 6 1 1 --
Cladomics cladogram Unio ACULAMPROTULINA

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 6 1 1 --
Cladomics cladogram Unio ACULAMPROTULINA

Araujo et al., 2018 [Publ 3779], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 13 13 9 1 5 1 1 19.0
Cladomics cladogram OXYNAIINA Unio

Bolotov et al., 2018 [Publ 3826], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 79 2 1 16.0
Cladomics cladogram Unio OXYNAIINA

Huang et al., 2018 [Publ 3828], fig. 3_5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 28 1 1 --
Cladomics cladogram Unio ACULAMPROTULINA

Huang et al., 2018 [Publ 3828], fig. 6-M

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 14 1 1 16.0
Cladomics cladogram ACULAMPROTULINA Unio

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 16 1 1 15.0
Cladomics cladogram ACULAMPROTULINA Unio

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 6 1 1 15.0
Cladomics cladogram ACULAMPROTULINA Unio

Klishko et al., 2018 [Publ 3945], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 4 4 1 1 4 4 1 16.0
Cladomics cladogram OXYNAIINA Unio

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 2 1 --
Cladomics cladogram Unio OXYNAIINA

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 2 1 --
Cladomics cladogram Unio OXYNAIINA

Wu et al., 2020 [Publ 4121], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 4 1 --
Cladomics cladogram Unio ACULAMPROTULINA

Wu et al., 2020 [Publ 4121], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 4 1 --
Cladomics cladogram Unio ACULAMPROTULINA

Wu et al., 2020 [Publ 4147], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 4 1 --
Cladomics cladogram Unio ACULAMPROTULINA

Choi et al., 2020 [Publ 4169], fig. 2a

cst iT tT bi riS oT roT roS iD
0.875 1 1 0 1 11 11 1 --
Cladomics cladogram Nodularia Schistodesmus Cuneopsis Unio Tchangsinaia

Choi et al., 2020 [Publ 4169], fig. 3a

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 5 5 1 --
Cladomics cladogram Nodularia Cuneopsis Unio

Sonowal et al., 2021 [Publ 4221], fig. 3a

cst iT tT bi riS oT roT roS iD
0.400 2 4 0 1 13 1 1 21.0
Cladomics cladogram Elliptio Lampsilis ANODONTINA Unio ravoisieri Unio pictorum ACULAMPROTULINA

Sonowal et al., 2021 [Publ 4221], fig. 3b

cst iT tT bi riS oT roT roS iD
0.400 2 9 0 1 13 1 1 26.0
Cladomics cladogram Triplodon Velesunio Hyridella ACULAMPROTULINA Lamprotula Lampsilis Potomida Unio pictorum ANODONTINA Unio ravoisieri Lamellidens Elliptio

Sonowal et al., 2021 [Publ 4221], fig. 4

cst iT tT bi riS oT roT roS iD
0.400 2 4 0 1 14 1 1 21.0
Cladomics cladogram ACULAMPROTULINA Unio pictorum ANODONTINA Unio ravoisieri Lamellidens

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

cst iT tT bi riS oT roT roS iD
1.000 17 17 15 1 5 2 1 22.0
Cladomics cladogram OXYNAIINA Unio

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

cst iT tT bi riS oT roT roS iD
1.000 17 17 15 1 5 2 1 12.0
Cladomics cladogram OXYNAIINA Unio

Altaba et al., 2025 [Publ 4849], fig. 10

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 9 1 1 16.0
Cladomics cladogram OXYNAIINA Unio

Altaba et al., 2025 [Publ 4849], fig. 11

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 11 1 1 16.0
Cladomics cladogram OXYNAIINA Unio

Altaba et al., 2025 [Publ 4849], fig. 12

cst iT tT bi riS oT roT roS iD
0.600 3 4 2 1 10 1 1 17.0
Cladomics cladogram Unio mancus Unio pictorum Unio tumidus OXYNAIINA Amuranodonta Cristaria Beringiana Sinanodonta woodiana 2 Sinanodonta woodiana 1 Sinanodonta lauta Sinanodonta tumens ANODONTINA

Wu et al., 2026 [Publ 4811], fig. 3

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 16 3 1 --
Cladomics cladogram Unio Lanceolaria GLOBUNIONINA Acuticosta ANODONTINA

Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5_7

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 2 0 0 18.0
Cladomics cladogram ANODONTINA Unio

Rosenberg et al., 1994 [Publ 2719], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 18 0 0 --
Cladomics cladogram Elliptio Fusconaia Plectomerus Pleurobema Unio

Rosenberg et al., 1997 [Publ 2720], fig. 2A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 0 0 --
Cladomics cladogram Gonidea Pyganodon grandis Utterbackia Pyganodon cataracta Lampsilis straminea Lampsilis teres Megalonaias Obliquaria Theliderma Quadrula Uniomerus Amblema Elliptio Fusconaia Plectomerus Pleurobema Unio

Nagel et al., 1998 [Publ 1946], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 5 0 0 17.0
Cladomics cladogram ANODONTINA Unio

Nagel et al., 1998 [Publ 1946], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 4 4 1 1 0 0 0 16.0
Cladomics cladogram Unio

Nagel et al., 1998 [Publ 1946], fig. 9

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram Unio ANODONTINA

Graf, 2000 [Publ 2647], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 16 0 0 --
Cladomics cladogram Unio Elliptio ALASMIDONTINA Lampsilis

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 28 0 0 --
Cladomics cladogram Unio ANODONTINA ALASMIDONTINA QUADRULINA Eurynia Pleurobema Fusconaia Amblema Ligumia Truncilla Ptychobranchus Sagittunio Lampsilis siliquoidea Epioblasma Ortmanniana Cambarunio Leaunio Lampsilis cardium Lampsilis fasciola

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 28 0 0 --
Cladomics cladogram Unio ANODONTINA ALASMIDONTINA QUADRULINA Eurynia Pleurobema Fusconaia Amblema Truncilla Ligumia Ptychobranchus Sagittunio Lampsilis siliquoidea Ortmanniana Lampsilis cardium Lampsilis fasciola Epioblasma Cambarunio Leaunio

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 0 0 --
Cladomics cladogram Unio ALASMIDONTINA

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 32 0 0 --
Cladomics cladogram Gonidea Unio Cumberlandia Margaritifera Toxolasma Amblema Glebula Cyrtonaias Potamilus Ortmanniana Ligumia QUADRULINA Eurynia Pleurobema Fusconaia

Hoeh et al., 2001 [Publ 2652], fig. 14f2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 29 0 0 --
Cladomics cladogram Unio Pyganodon ANODONTINA Strophitus Mutela Anodontites guanarensis Anodontites trigona Monocondylaea Diplodon Echyridella Lortiella Castalia Velesunio

Hoeh et al., 2001 [Publ 2652], fig. 14f4

cst iT tT bi riS oT roT roS iD
0.000 2 5 0 1 29 0 0 20.0
Cladomics cladogram Unio pictorum Unio tumidus ANODONTINA ALASMIDONTINA Amblema QUADRULINA Eurynia Fusconaia Pleurobema Cyrtonaias Glebula Toxolasma Potamilus Ligumia Ortmanniana

Graf, 2002 [Publ 2651], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 16 0 0 --
Cladomics cladogram Unio ALASMIDONTINA

Hoeh et al., 2002 [Publ 2653], fig. 2

cst iT tT bi riS oT roT roS iD
0.000 2 5 0 1 29 0 0 20.0
Cladomics cladogram Gonidea Unio pictorum Unio tumidus ANODONTINA ALASMIDONTINA

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 29 0 0 --
Cladomics cladogram Unio Pyganodon ANODONTINA Strophitus

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 29 0 0 --
Cladomics cladogram Unio ANODONTINA ALASMIDONTINA

Giribet & Distel, 2003 [Publ 3670], fig. 3f5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram Elliptio Lampsilis Unio

Huff et al., 2004 [Publ 2697], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 0 0 --
Cladomics cladogram Unio ANODONTINA

Huff et al., 2004 [Publ 2697], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 0 0 --
Cladomics cladogram Unio ANODONTINA

Huff et al., 2004 [Publ 2697], fig. 5-6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 0 0 --
Cladomics cladogram Unio ANODONTINA Lampsilis

Källersjö et al., 2005 [Publ 2684], fig. 1A

cst iT tT bi riS oT roT roS iD
1.000 3 3 0 1 4 0 0 17.0
Cladomics cladogram Margaritifera ANODONTINA Unio

Källersjö et al., 2005 [Publ 2684], fig. 1B-F

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 4 0 0 16.0
Cladomics cladogram Margaritifera ANODONTINA Unio

Soroka, 2005 [Publ 2726], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 1 0 0 17.0
Cladomics cladogram Pronodularia Unio

Graf & Cummings, 2006 [Publ 1968], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 45 0 0 --
Cladomics cladogram Unio ALASMIDONTINA

Taylor et al., 2007 [Publ 2696], fig. 1-3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 0 0 --
Cladomics cladogram ANODONTINA Unio

Chong et al., 2008 [Publ 2668], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 0 0 --
Cladomics cladogram Anodonta cygnea Pyganodon Alasminota Anodonta anatina Unio CRISTARIINA Brachyanodon nuttalliana Brachyanodon californiensis Pseudodontoideus Alasmidonta Pterosyna Pseudanodonta Strophitus Prolasmidonta Platynaias Brachyanodon oregonensis 4 Brachyanodon oregonensis 3 Brachyanodon oregonensis 2 Brachyanodon oregonensis 1 Brachyanodon kennerlyi

Manendo et al., 2008 [Publ 2669], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 10 0 0 18.0
Cladomics cladogram Unio Anodonta Pseudanodonta ALASMIDONTINA

Araujo et al., 2009 [Publ 2395], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 0 0 --
Cladomics cladogram Unio Cumberlandia Pseudunio Margaritifera

Araujo et al., 2009 [Publ 2589], fig. 7B

cst iT tT bi riS oT roT roS iD
1.000 7 7 6 1 4 0 0 17.0
Cladomics cladogram ANODONTINA Unio

Araujo et al., 2009 [Publ 2589], fig. 7C

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 1 4 0 0 16.0
Cladomics cladogram ANODONTINA Unio

Hoeh et al., 2009 [Publ 2716], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 45 0 0 --
Cladomics cladogram Unio ALASMIDONTINA

Hoeh et al., 2009 [Publ 2716], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 44 0 0 --
Cladomics cladogram Unio ALASMIDONTINA

Hoeh et al., 2009 [Publ 2716], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 44 0 0 --
Cladomics cladogram Unio ALASMIDONTINA

Soroka, 2010 [Publ 3404], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 0 1 4 0 0 17.0
Cladomics cladogram Unio ANODONTINA

Soroka, 2010 [Publ 3404], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 2 0 0 18.0
Cladomics cladogram ANODONTINA Unio

Soroka, 2010 [Publ 3404], fig. 3a

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 5 0 0 17.0
Cladomics cladogram ANODONTINA Unio

Soroka, 2010 [Publ 3404], fig. 3a-M

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 3 0 0 18.0
Cladomics cladogram CRISTARIINA Unio

Soroka, 2010 [Publ 3404], fig. 3b

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 5 0 0 17.0
Cladomics cladogram CRISTARIINA Unio

Soroka, 2010 [Publ 3404], fig. 3b-M

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 3 0 0 16.0
Cladomics cladogram CRISTARIINA Unio

Soroka, 2010 [Publ 3404], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 4 0 0 16.0
Cladomics cladogram ANODONTINA Unio

Soroka, 2010 [Publ 3404], fig. 5a

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 5 0 0 18.0
Cladomics cladogram Unio Anodonta anatina CRISTARIINA Anodonta cygnea Pseudanodonta

Soroka, 2010 [Publ 3404], fig. 5b

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 5 0 0 16.0
Cladomics cladogram ANODONTINA CRISTARIINA Unio

Soroka, 2010 [Publ 3404], fig. 8a

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 3 0 0 16.0
Cladomics cladogram Unio CRISTARIINA ANODONTINA

Soroka, 2010 [Publ 3404], fig. 8a-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 1 0 0 --
Cladomics cladogram CRISTARIINA Unio

Soroka, 2010 [Publ 3404], fig. 8b

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 3 0 0 16.0
Cladomics cladogram ANODONTINA Unio

Soroka, 2010 [Publ 3404], fig. 8b-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 1 0 0 --
Cladomics cladogram CRISTARIINA Unio

Khalloufi et al., 2011 [Publ 2748], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 9 9 6 1 2 0 0 18.0
Cladomics cladogram Potomida Unio

Whelan et al., 2011 [Publ 3380], fig. 1-ML

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 34 0 0 --
Cladomics cladogram ALASMIDONTINA Unio

Whelan et al., 2011 [Publ 3380], fig. 1-MP

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 34 0 0 --
Cladomics cladogram ALASMIDONTINA Unio

Plazzi et al., 2011 [Publ 3676], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 15 0 0 16.0
Cladomics cladogram Lanceolaria Anodonta Unio ALASMIDONTINA Pseudanodonta Alathyria Pronodularia Sinohyriopsis QUADRULINA Parvaspina Lampsilis Epioblasma Venustaconcha

Prie et al., 2012 [Publ 3487], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 1 1 0 0 16.0
Cladomics cladogram Pseudunio Unio

Zieritz et al., 2012 [Publ 3673], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 5 0 0 18.0
Cladomics cladogram CRISTARIINA Margaritifera Unio

Reis et al., 2013 [Publ 3580], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 1 8 0 0 15.0
Cladomics cladogram ANODONTINA Unio

Reis et al., 2013 [Publ 3580], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 7 7 6 1 8 0 0 18.0
Cladomics cladogram ANODONTINA Unio

Huang et al., 2013 [Publ 3629], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 15 0 0 --
Cladomics cladogram Unio CRISTARIINA ALASMIDONTINA

Prié & Puillandre, 2014 [Publ 3529], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 11 11 9 1 6 0 0 20.0
Cladomics cladogram ANODONTINA Unio

Graf et al., 2014 [Publ 3591], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 0 0 --
Cladomics cladogram Margaritifera Unio Leoparreysia Nitia Coelatura aegyptiaca 2 Coelatura aegyptiaca 1 Coelatura gabonensis Coelatura kunenensis Prisodontopsis Coelatura choziensis 1 Coelatura choziensis 2 Coelatura luapulaensis 1 Coelatura luapulaensis 2 Coelatura luapulaensis 3 Coelatura luapulaensis 4 Coelatura luapulaensis 5

Marshall et al., 2014 [Publ 3593], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram ALASMIDONTINA Unio

Bieler et al., 2014 [Publ 3622], fig. 26

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram Unio Aspatharia Margaritifera

Bieler et al., 2014 [Publ 3622], fig. 27

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram Unio Aspatharia Margaritifera

Bieler et al., 2014 [Publ 3622], fig. 28

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram Unio Velesunio

Bieler et al., 2014 [Publ 3622], fig. 29

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram Unio Aspatharia Margaritifera

Bieler et al., 2014 [Publ 3622], fig. 30

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram Unio Aspatharia Margaritifera

Bieler et al., 2014 [Publ 3622], fig. 31

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram Unio Margaritifera Aspatharia

Graf et al., 2015 [Publ 3619], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 0 0 --
Cladomics cladogram Unio Margaritifera

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 39 0 0 --
Cladomics cladogram Unio ALASMIDONTINA

González & Giribet, 2015 [Publ 3632], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram Unio Velesunio

González & Giribet, 2015 [Publ 3632], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram Unio Velesunio Aspatharia Margaritifera

González & Giribet, 2015 [Publ 3632], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram Unio Margaritifera

González & Giribet, 2015 [Publ 3632], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram Unio Aspatharia Margaritifera

Bolotov et al., 2015 [Publ 3646], fig. 2A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 0 0 --
Cladomics cladogram Unio Margaritifera margaritifera Margaritifera dahurica 1 Margaritifera dahurica 2 Pseudunio Cumberlandia Margaritifera falcata Margaritifera laevis Margaritifera middendorffi Margaritifera marrianae

Bolotov et al., 2015 [Publ 3646], fig. 2B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 0 0 --
Cladomics cladogram Unio Cumberlandia Pseudunio Margaritifera

Wang et al., 2016 [Publ 3714], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 2 0 0 --
Cladomics cladogram Unio Sinohyriopsis

Bolotov et al., 2016 [Publ 3743], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 28 0 0 --
Cladomics cladogram ANODONTINA Unio

Bolotov et al., 2016 [Publ 3743], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 28 0 0 --
Cladomics cladogram ANODONTINA Unio

Fonseca et al., 2016 [Publ 3775], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 0 0 --
Cladomics cladogram Unio ANODONTINA ALASMIDONTINA

Lopes-Lima et al., 2017 [Publ 3734], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 9 9 5 1 8 0 0 18.0
Cladomics cladogram ANODONTINA Unio

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 0 0 --
Cladomics cladogram Unio ANODONTINA ALASMIDONTINA

Froufe et al., 2017 [Publ 3802], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 7 0 0 18.0
Cladomics cladogram Unio CRISTARIINA ANODONTINA

Combosch et al., 2017 [Publ 3806], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 212 0 0 --
Cladomics cladogram Unio Lampsilis Scabies

Combosch et al., 2017 [Publ 3806], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 213 0 0 --
Cladomics cladogram Unio Scabies Lampsilis

Vikhrev et al., 2018 [Publ 3812], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 12 0 0 --
Cladomics cladogram Unio Lampsilis

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 27 0 0 --
Cladomics cladogram Lampsilis Unio

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 27 0 0 --
Cladomics cladogram Lampsilis Unio

Chase et al., 2018 [Publ 3847], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 37 0 0 --
Cladomics cladogram Unio CRISTARIINA ANODONTINA ALASMIDONTINA

Chase et al., 2018 [Publ 3847], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 8 0 0 --
Cladomics cladogram Unio ANODONTINA ALASMIDONTINA

Bogatov et al., 2018 [Publ 3859], fig. 1

cst iT tT bi riS oT roT roS iD
0.000 3 5 3 1 3 0 0 18.0
Cladomics cladogram Margaritifera Unio tumidus ANODONTINA Unio crassus Unio pictorum

Bogatov et al., 2018 [Publ 3859], fig. 2

cst iT tT bi riS oT roT roS iD
0.000 3 5 3 1 3 0 0 20.0
Cladomics cladogram Margaritifera Unio crassus Unio pictorum Unio tumidus ANODONTINA

Chase et al., 2018 [Publ 3861], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 9 0 0 --
Cladomics cladogram Unio ANODONTINA ALASMIDONTINA

Chase et al., 2018 [Publ 3861], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 9 0 0 --
Cladomics cladogram Unio ANODONTINA ALASMIDONTINA

Marrone et al., 2019 [Publ 3907], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 8 8 4 1 0 0 0 20.0
Cladomics cladogram Unio

Froufe et al., 2019 [Publ 3936], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 16 0 0 16.0
Cladomics cladogram Unio CRISTARIINA ANODONTINA ALASMIDONTINA

Froufe et al., 2019 [Publ 3936], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 16 0 0 16.0
Cladomics cladogram Unio CRISTARIINA ANODONTINA ALASMIDONTINA

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 0 0 --
Cladomics cladogram Unio ANODONTINA

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram Unio ANODONTINA

Riccardi et al., 2020 [Publ 4089], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 17 0 0 --
Cladomics cladogram Lanceolaria Unio

Tomilova et al., 2020 [Publ 4097], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 6 0 0 --
Cladomics cladogram Unio CRISTARIINA ANODONTINA

Kilikowska et al., 2020 [Publ 4335], fig. 4a

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 0 0 0 16.0
Cladomics cladogram Unio

Klunzinger et al., 2021 [Publ 4086], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram Margaritifera Unio

Lopes-Lima et al., 2021 [Publ 4217], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 16 16 14 1 9 0 0 21.0
Cladomics cladogram ANODONTINA Unio

Lopes-Lima et al., 2021 [Publ 4217], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 1 0 0 0 19.0
Cladomics cladogram Unio

Lopes-Lima et al., 2021 [Publ 4217], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 1 0 0 0 16.0
Cladomics cladogram Unio

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 20 0 0 16.0
Cladomics cladogram Unio CRISTARIINA ANODONTINA ALASMIDONTINA

Lyubas et al., 2022 [Publ 4255], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 23 23 20 1 8 0 0 27.0
Cladomics cladogram ANODONTINA Unio

Lyubas et al., 2022 [Publ 4255], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 23 23 14 1 8 0 0 26.0
Cladomics cladogram ANODONTINA Unio

Bashê, 2022 [Publ 4265], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 5 5 4 1 1 0 0 21.0
Cladomics cladogram Pseudunio Unio

Pfarrer et al., 2022 [Publ 4305], fig. 2-5

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 1 7 0 0 14.0
Cladomics cladogram Unio CRISTARIINA ANODONTINA

Lopes-Lima et al., 2024 [Publ 4376], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 14 14 12 1 0 0 0 16.0
Cladomics cladogram Unio

Zieritz et al., 2024 [Publ 4381], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 44 0 0 --
Cladomics cladogram ANODONTINA Unio

Wu et al., 2025 [Publ 4442], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 15 0 0 --
Cladomics cladogram Unio CRISTARIINA

Prie et al., 2026 [Publ 4818], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 11 11 9 1 0 0 0 14.0
Cladomics cladogram Unio

Lawrie et al., 2026 [Publ 4843], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 0 0 --
Cladomics cladogram Unio Castalia Triplodon Hyridella depressa Hyridella drapeta Hyridella australis Cucumerunio Westralunio Velesunio 6 Alathyria jacksoni 2 Alathyria profuga Velesunio ambiguus Velesunio 5 Alathyria pertexta 2 Velesunio 4 Alathyria jacksoni 1 Alathyria pertexta 1 Lortiella Velesunio 3 Microdontia Velesunio angasi Velesunio 1 Velesunio 2