Bivalvia | Unionida | Unionoidea | Unionidae

subfamily UNIONINAE Rafinesque, 1820: cladomics

5 tribes

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 228 (289 total)
number of subtaxa (rank: tribe) = 5
analyzed in 436 cladograms in 242 publications
maximum number of terminals in an analysis = 156 (mean = 11.94, median = 7)
maximum number of subtaxa (tribe) = 5 (mean = 2.04, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 383
maximum number of included terminals = 156, (mean = 13.46 median = 8)
maximum number of subtaxa (tribe) = 5 (mean = 2.19, median = 2)
proportion of cladograms that support basic monophyly = 0.93

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 180
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (tribe) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 156 (mean = 17.94, median = 13)
maximum number of subtaxa (tribe) = 5 (mean = 3.17, median = 3)
proportion of cladograms that support rigorous monophyly = 0.88

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Unioninae cladogram distances
PCoA Ordination

NJ Tree of Unioninae cladogram distances
NJ Tree

Cladograms

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
0.988 46 46 44 5 49 38 3 **
Cladomics cladogram AMBLEMINAE GONIDEINAE PARREYSIINAE LEPIDODESMINI UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
0.922 48 48 37 5 153 114 3 49.0
Cladomics cladogram PARREYSIINAE ACUTICOSTINI LEPIDODESMINI UNIONINI LANCEOLARIINI ANODONTINI Rectidentini AMBLEMINAE CHAMBERLAINIINI LAMPROTULINI GONIDEINI PSEUDODONTINI

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
0.913 47 47 44 5 152 113 3 51.0
Cladomics cladogram PARREYSIINAE PSEUDODONTINI Rectidentini CHAMBERLAINIINI AMBLEMINAE LAMPROTULINI GONIDEINI ACUTICOSTINI UNIONINI LEPIDODESMINI LANCEOLARIINI ANODONTINI

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.826 48 48 25 5 153 114 3 53.0
Cladomics cladogram PARREYSIINAE Chamberlainia Megalonaias Uniomerus QUADRULINA Sinohyriopsis LAMPROTULINI GONIDEINI PSEUDODONTINI Rectidentini Amblema Toxolasma PLEUROBEMINI Obliquaria Plectomerus Ellipsaria Potamilus leptodon Medionidus Potamilus alatus Ptychobranchus Truncilla Venustaconcha Cambarunio Leaunio Ortmanniana Lampsilis ACUTICOSTINI Alasmidonta Anodonta anatina Anemina Anodonta cygnea Cristaria Brachyanodon Lasmigona LEPIDODESMINI Pseudanodonta Pyganodon Simpsonaias Strophitus Utterbackia Schistodesmus UNIONINA ACULAMPROTULINA LANCEOLARIINI Pletholophus Sinanodonta Cuneopsis celtiformis Tchangsinaia Cuneopsis rufescens Nodularia Cuneopsis szechenyii Cuneopsis heudei

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
0.946 71 71 59 5 51 40 3 67.0
Cladomics cladogram PARREYSIINAE ACUTICOSTINI LEPIDODESMINI UNIONINI LANCEOLARIINI ANODONTINI Rectidentini AMBLEMINAE CHAMBERLAINIINI LAMPROTULINI PSEUDODONTINI GONIDEINI

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
0.952 26 26 24 5 65 48 3 38.0
Cladomics cladogram PARREYSIINAE ACUTICOSTINI LEPIDODESMINI UNIONINI LANCEOLARIINI ANODONTINI Rectidentini AMBLEMINAE CHAMBERLAINIINI LAMPROTULINI PSEUDODONTINI GONIDEINI

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
0.928 26 26 24 5 65 48 3 40.0
Cladomics cladogram PARREYSIINAE ACUTICOSTINI UNIONINI LANCEOLARIINI LEPIDODESMINI ANODONTINI Rectidentini AMBLEMINAE CHAMBERLAINIINI LAMPROTULINI PSEUDODONTINI GONIDEINI

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

cst iT tT bi riS oT roT roS iD
0.981 16 16 14 5 21 19 3 37.0
Cladomics cladogram AMBLEMINAE GONIDEINAE PARREYSIINAE LEPIDODESMINI UNIONINI LANCEOLARIINI ACUTICOSTINI ANODONTINI

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

cst iT tT bi riS oT roT roS iD
0.963 17 17 15 5 22 20 3 40.0
Cladomics cladogram AMBLEMINAE GONIDEINAE PARREYSIINAE ANODONTINI ACUTICOSTINI LANCEOLARIINI LEPIDODESMINI ACULAMPROTULINA UNIONINA OXYNAIINA

Konopleva et al., 2023 [Publ 4269], fig. S1_4

cst iT tT bi riS oT roT roS iD
0.984 33 33 31 5 23 22 3 50.0
Cladomics cladogram PARREYSIINAE AMBLEMINAE GONIDEINAE UNIONINI LEPIDODESMINI LANCEOLARIINI ACUTICOSTINI ANODONTINI

Chen et al., 2023 [Publ 4302], fig. 1

cst iT tT bi riS oT roT roS iD
0.907 32 32 24 5 9 7 3 55.0
Cladomics cladogram GONIDEINAE AMBLEMINAE PARREYSIINAE Anemina Amuranodonta Pletholophus Sinanodonta Beringiana Lanceolaria gladiola ACUTICOSTINI LEPIDODESMINI Lanceolaria oxyrhyncha Cristaria UNIONINI ALASMIDONTINA ANODONTINA

Chen et al., 2023 [Publ 4302], fig. 2

cst iT tT bi riS oT roT roS iD
0.981 32 32 28 5 9 7 3 54.0
Cladomics cladogram GONIDEINAE AMBLEMINAE PARREYSIINAE ACUTICOSTINI LEPIDODESMINI UNIONINI LANCEOLARIINI ANODONTINI

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cst iT tT bi riS oT roT roS iD
0.951 46 46 41 5 49 38 3 19.0
Cladomics cladogram AMBLEMINAE GONIDEINAE PARREYSIINAE LEPIDODESMINI UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cst iT tT bi riS oT roT roS iD
0.976 46 46 43 5 45 39 3 11.0
Cladomics cladogram AMBLEMINAE GONIDEINAE PARREYSIINAE LEPIDODESMINI UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cst iT tT bi riS oT roT roS iD
0.976 46 46 43 5 45 39 3 11.0
Cladomics cladogram AMBLEMINAE GONIDEINAE PARREYSIINAE LEPIDODESMINI UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Dai et al., 2025 [Publ 4440], fig. 1

cst iT tT bi riS oT roT roS iD
0.929 35 38 31 5 9 7 3 58.0
Cladomics cladogram GONIDEINAE AMBLEMINAE PARREYSIINAE Anemina Amuranodonta Pletholophus Sinanodonta Beringiana Cristaria ALASMIDONTINA ANODONTINA LANCEOLARIINI LEPIDODESMINI ACUTICOSTINI UNIONINI

Dai et al., 2025 [Publ 4440], fig. 2

cst iT tT bi riS oT roT roS iD
0.911 35 35 29 5 9 7 3 54.0
Cladomics cladogram GONIDEINAE AMBLEMINAE PARREYSIINAE Anemina Amuranodonta Pletholophus Sinanodonta Beringiana ALASMIDONTINA ANODONTINA Lanceolaria gladiola Lanceolaria oxyrhyncha Cristaria ACUTICOSTINI LEPIDODESMINI UNIONINI

Chen et al., 2025 [Publ 4487], fig. 2

cst iT tT bi riS oT roT roS iD
0.982 32 32 30 5 9 7 3 54.0
Cladomics cladogram GONIDEINAE AMBLEMINAE PARREYSIINAE GLOBUNIONINA ACULAMPROTULINA UNIONINA OXYNAIINA ACUTICOSTINI LEPIDODESMINI LANCEOLARIINI ANODONTINI

Chen et al., 2025 [Publ 4487], fig. 3

cst iT tT bi riS oT roT roS iD
0.964 32 32 30 5 9 7 3 58.0
Cladomics cladogram AMBLEMINAE GONIDEINAE PARREYSIINAE UNIONINI ACUTICOSTINI ALASMIDONTINA ANODONTINA LEPIDODESMINI Cristaria LANCEOLARIINI Anemina Amuranodonta Pletholophus Sinanodonta Beringiana

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cst iT tT bi riS oT roT roS iD
0.966 68 68 66 5 72 45 3 46.0
Cladomics cladogram AMBLEMINAE GONIDEINAE PARREYSIINAE LEPIDODESMINI UNIONINI LANCEOLARIINI ACUTICOSTINI ANODONTINI

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.753 156 156 164 5 477 411 3 158.0
Cladomics cladogram Lamellidens Balwantia Trapezidens LEOPARREYSIINI Pseudomulleria INDOCHINELLINI PARREYSIINI COELATURINI Amuranodonta Anemina Buldowskia Beringiana Pletholophus Mimo Acudonta Sinanodonta Cristaria LEPIDODESMINI LANCEOLARIINI Pseudoacuticosta retiaria 2 Acuticosta chinensis 2 Acuticosta chinensis 1 Pseudoacuticosta retiaria 1 Globunio ACULAMPROTULINA MIDDENDORFFINAIINA UNIONINA OXYNAIINA Pseudodontoideus Lasmigona Pterosyna Arcidens ANODONTINA Utterbackia Pyganodon Utterbackiana Simpsonaias Alasminota Alasmidonta Prolasmidonta Platynaias Anodontoides Strophitus

Huang et al., 2002 [Publ 2657], fig. 1

cst iT tT bi riS oT roT roS iD
0.816 14 14 12 5 20 18 2 50.0
Cladomics cladogram Sinosolenaia LAMPROTULINI CHAMBERLAINIINI Ptychorhynchus AMBLEMINAE ANODONTINI ACUTICOSTINI Nodularia Tchangsinaia LANCEOLARIINI Schistodesmus LEPIDODESMINI

Huang et al., 2002 [Publ 2657], fig. 2

cst iT tT bi riS oT roT roS iD
0.776 14 14 5 5 20 18 2 47.0
Cladomics cladogram Ptychorhynchus CHAMBERLAINIINI LAMPROTULINI Sinosolenaia AMBLEMINAE Strophitus Schistodesmus LEPIDODESMINI ACUTICOSTINI CRISTARIINA LANCEOLARIINI Nodularia Tchangsinaia Utterbackia Utterbackiana Alasmidonta Lasmigona

Wu et al., 2018 [Publ 3836], fig. 2

cst iT tT bi riS oT roT roS iD
0.907 33 33 28 5 20 18 2 39.0
Cladomics cladogram QUADRULINI PLEUROBEMINI AMBLEMINI LAMPSILINI LAMPROTULINI GONIDEINI CHAMBERLAINIINI LEPIDODESMINI UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Aunins et al., 2018 [Publ 3863], fig. 1

cst iT tT bi riS oT roT roS iD
0.979 26 26 24 5 14 11 2 22.0
Cladomics cladogram AMBLEMINAE GONIDEINAE LEPIDODESMINI UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Huang et al., 2019 [Publ 3843], fig. 3

cst iT tT bi riS oT roT roS iD
0.962 26 26 24 5 30 20 2 28.0
Cladomics cladogram GONIDEINAE AMBLEMINAE LEPIDODESMINI UNIONINI LANCEOLARIINI ACUTICOSTINI ANODONTINI

Wu et al., 2019 [Publ 3864], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 26 26 24 5 20 14 2 24.0
Cladomics cladogram AMBLEMINAE GONIDEINAE LEPIDODESMINI UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Fukata & Iigo, 2020 [Publ 4106], fig. 1

cst iT tT bi riS oT roT roS iD
0.983 28 28 26 5 26 23 2 28.0
Cladomics cladogram AMBLEMINAE GONIDEINAE LEPIDODESMINI LANCEOLARIINI UNIONINI ACUTICOSTINI ANODONTINI

Wu et al., 2020 [Publ 4121], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 26 26 24 5 22 14 2 24.0
Cladomics cladogram AMBLEMINAE GONIDEINAE LEPIDODESMINI UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Wu et al., 2021 [Publ 4366], fig. 3-BI

cst iT tT bi riS oT roT roS iD
0.955 33 33 31 5 40 29 2 23.0
Cladomics cladogram AMBLEMINAE GONIDEINAE LEPIDODESMINI UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Wu et al., 2021 [Publ 4366], fig. 3-ML

cst iT tT bi riS oT roT roS iD
0.955 33 33 31 5 40 29 2 21.0
Cladomics cladogram AMBLEMINAE GONIDEINAE LEPIDODESMINI UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Wu et al., 2021 [Publ 4366], fig. 4

cst iT tT bi riS oT roT roS iD
0.952 32 32 30 5 36 25 2 22.0
Cladomics cladogram AMBLEMINAE GONIDEINAE LEPIDODESMINI UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Wu et al., 2022 [Publ 4238], fig. 3

cst iT tT bi riS oT roT roS iD
0.985 35 35 33 5 38 29 2 17.0
Cladomics cladogram AMBLEMINAE GONIDEINAE LEPIDODESMINI UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Dai et al., 2023 [Publ 4285], fig. 5

cst iT tT bi riS oT roT roS iD
0.986 17 17 15 5 36 34 2 33.0
Cladomics cladogram AMBLEMINAE GONIDEINAE LEPIDODESMINI UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Dai et al., 2024 [Publ 4358], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 28 28 26 5 25 23 2 30.0
Cladomics cladogram AMBLEMINAE GONIDEINAE LEPIDODESMINI UNIONINI LANCEOLARIINI ACUTICOSTINI ANODONTINI

Dai et al., 2024 [Publ 4386], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 27 27 25 5 14 14 2 35.0
Cladomics cladogram AMBLEMINAE GONIDEINAE LEPIDODESMINI UNIONINI LANCEOLARIINI ACUTICOSTINI ANODONTINI

Dai et al., 2025 [Publ 4440], fig. 3

cst iT tT bi riS oT roT roS iD
0.987 42 42 40 5 45 36 2 24.0
Cladomics cladogram AMBLEMINAE GONIDEINAE LEPIDODESMINI UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Chen et al., 2025 [Publ 4448], fig. 3

cst iT tT bi riS oT roT roS iD
0.986 43 43 41 5 42 33 2 27.0
Cladomics cladogram AMBLEMINAE GONIDEINAE LEPIDODESMINI UNIONINI LANCEOLARIINI ACUTICOSTINI ANODONTINI

Sano et al., 2017 [Publ 3799], fig. 2

cst iT tT bi riS oT roT roS iD
0.906 21 21 22 5 12 8 1 55.0
Cladomics cladogram GONIDEINAE ANODONTINI Tchangsinaia Nodularia Schistodesmus ACUTICOSTINI Inversiunio LEPIDODESMINI LANCEOLARIINI

Huang et al., 2019 [Publ 3843], fig. 1

cst iT tT bi riS oT roT roS iD
0.833 32 32 30 5 14 12 1 54.0
Cladomics cladogram LAMPROTULINI GONIDEINI CHAMBERLAINIINI Anemina ACUTICOSTINI Sinanodonta ACULAMPROTULINA Cristaria LANCEOLARIINI LEPIDODESMINI OXYNAIINA

Huang et al., 2019 [Publ 3843], fig. S1

cst iT tT bi riS oT roT roS iD
0.833 33 33 36 5 14 12 1 51.0
Cladomics cladogram GONIDEINAE Anemina Sinanodonta ACUTICOSTINI LEPIDODESMINI Cristaria LANCEOLARIINI UNIONINI

Sano et al., 2020 [Publ 4138], fig. 1

cst iT tT bi riS oT roT roS iD
0.686 22 22 5 5 12 8 1 59.0
Cladomics cladogram GONIDEINAE Lanceolaria oxyrhyncha 1 Lanceolaria oxyrhyncha 2 Cristaria plicata 1 Cristaria plicata 2 Sinanodonta lauta Beringiana japonica Beringiana fukuharai 1 Beringiana beringiana Beringiana fukuharai 2 Beringiana fukuharai 3 Buldowskia iwakawai 1 Buldowskia iwakawai 2 Sinanodonta tumens 1 Sinanodonta tumens 2 Pletholophus Anemina Lanceolaria lanceolata Cristaria plicata 3 Lanceolaria gladiola Sinanodonta angula LEPIDODESMINI ACULAMPROTULINA ACUTICOSTINI Cuneopsis Nodularia Schistodesmus Inversiunio

Dai et al., 2024 [Publ 4386], fig. 5-BI

cst iT tT bi riS oT roT roS iD
0.933 17 17 11 5 5 3 1 50.0
Cladomics cladogram PARREYSIINAE ACUTICOSTINI CRISTARIINA ANODONTINA LANCEOLARIINI ACULAMPROTULINA LEPIDODESMINI UNIONINA MIDDENDORFFINAIINA OXYNAIINA

Dai et al., 2024 [Publ 4386], fig. 5-ML

cst iT tT bi riS oT roT roS iD
0.967 17 17 14 5 5 3 1 50.0
Cladomics cladogram PARREYSIINAE ANODONTINA ACUTICOSTINI CRISTARIINA LEPIDODESMINI LANCEOLARIINI UNIONINI

Wu et al., 2026 [Publ 4811], fig. 4

cst iT tT bi riS oT roT roS iD
0.986 65 65 63 5 30 28 1 41.0
Cladomics cladogram GONIDEINAE LEPIDODESMINI UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Wu et al., 2022 [Publ 4238], fig. 2d

cst iT tT bi riS oT roT roS iD
1.000 38 38 36 5 0 0 0 22.0
Cladomics cladogram LEPIDODESMINI UNIONINI LANCEOLARIINI ACUTICOSTINI ANODONTINI

Sano et al., 2020 [Publ 4138], fig. 2

cst iT tT bi riS oT roT roS iD
0.939 36 36 36 4 33 10 3 42.0
Cladomics cladogram PARREYSIINAE AMBLEMINAE GONIDEINAE Anemina Buldowskia Pletholophus Beringiana Sinanodonta Cristaria ANODONTINA ALASMIDONTINA LANCEOLARIINI LEPIDODESMINI UNIONINI

Sano et al., 2020 [Publ 4138], fig. 3

cst iT tT bi riS oT roT roS iD
0.939 36 36 37 4 33 10 3 42.0
Cladomics cladogram PARREYSIINAE CHAMBERLAINIINI AMBLEMINAE LAMPROTULINI GONIDEINI Anemina Buldowskia Sinanodonta Pletholophus Beringiana Cristaria ANODONTINA ALASMIDONTINA LANCEOLARIINI LEPIDODESMINI UNIONINI

Sano et al., 2020 [Publ 4138], fig. 4

cst iT tT bi riS oT roT roS iD
0.939 36 36 37 4 33 10 3 42.0
Cladomics cladogram PARREYSIINAE AMBLEMINAE GONIDEINAE Anemina Buldowskia Sinanodonta Pletholophus Beringiana Cristaria ANODONTINA ALASMIDONTINA LEPIDODESMINI LANCEOLARIINI UNIONINI

Wu et al., 2024 [Publ 4388], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 18 18 16 4 35 33 3 34.0
Cladomics cladogram AMBLEMINAE GONIDEINAE PARREYSIINAE LEPIDODESMINI UNIONINI LANCEOLARIINI ANODONTINI

Ouyang et al., 2015 [Publ 4493], fig. 1

cst iT tT bi riS oT roT roS iD
0.917 16 16 14 4 15 15 2 52.0
Cladomics cladogram GONIDEINAE AMBLEMINAE UNIONINI LANCEOLARIINI ACUTICOSTINI ANODONTINI

Ouyang et al., 2015 [Publ 4493], fig. 2

cst iT tT bi riS oT roT roS iD
0.917 16 16 21 4 15 15 2 55.0
Cladomics cladogram CHAMBERLAINIINI LAMPROTULINI GONIDEINI AMBLEMINAE LANCEOLARIINI ACULAMPROTULINA ACUTICOSTINI ANODONTINI OXYNAIINA

Wang et al., 2018 [Publ 3865], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 17 17 15 4 14 14 2 31.0
Cladomics cladogram AMBLEMINAE GONIDEINAE UNIONINI ANODONTINI ACUTICOSTINI LANCEOLARIINI

Cha et al., 2018 [Publ 3916], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 21 21 19 4 17 14 2 29.5
Cladomics cladogram AMBLEMINAE GONIDEINAE LEPIDODESMINI UNIONINI LANCEOLARIINI ANODONTINI

Froufe et al., 2019 [Publ 3936], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 22 22 20 4 22 19 2 26.0
Cladomics cladogram AMBLEMINAE GONIDEINAE LEPIDODESMINI UNIONINI LANCEOLARIINI ANODONTINI

Froufe et al., 2019 [Publ 3936], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 22 22 20 4 22 19 2 26.0
Cladomics cladogram AMBLEMINAE GONIDEINAE LEPIDODESMINI UNIONINI LANCEOLARIINI ANODONTINI

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

cst iT tT bi riS oT roT roS iD
1.000 22 22 20 4 26 23 2 26.0
Cladomics cladogram AMBLEMINAE GONIDEINAE LEPIDODESMINI UNIONINI LANCEOLARIINI ANODONTINI

Wu et al., 2026 [Publ 4811], fig. 3

cst iT tT bi riS oT roT roS iD
0.957 10 10 8 4 7 5 2 48.0
Cladomics cladogram AMBLEMINAE GONIDEINAE UNIONINA LANCEOLARIINI GLOBUNIONINA ACUTICOSTINI ANODONTINI

Zhang et al., 2004 [Publ 2735], fig. 1

cst iT tT bi riS oT roT roS iD
0.955 12 12 10 4 2 2 1 44.0
Cladomics cladogram GONIDEINAE ANODONTINI UNIONINI ACUTICOSTINI LANCEOLARIINI

Zhou et al., 2007 [Publ 2658], fig. 1

cst iT tT bi riS oT roT roS iD
0.682 13 19 17 4 8 5 1 51.0
Cladomics cladogram MARGARITIFERINAE ANODONTINI CHAMBERLAINIINI Lamprotula leaii Lamprotula caveata GIBBOSULINAE Lamprotula cornuumlunae ACULAMPROTULINA Tchangsinaia Nodularia Schistodesmus LANCEOLARIINI ACUTICOSTINI

Ouyang et al., 2011 [Publ 3408], fig. 1

cst iT tT bi riS oT roT roS iD
0.818 9 9 6 4 4 4 1 47.0
Cladomics cladogram GONIDEINAE ANODONTINI OXYNAIINA Lanceolaria grayii ACUTICOSTINI Lanceolaria gladiola ACULAMPROTULINA

Liu et al., 2017 [Publ 3939], fig. 2

cst iT tT bi riS oT roT roS iD
0.947 14 15 12 4 1 1 1 49.0
Cladomics cladogram LANCEOLARIINI UNIONINI ACUTICOSTINI ANODONTINI GONIDEINAE

Kim et al., 2021 [Publ 4197], fig. 1

cst iT tT bi riS oT roT roS iD
0.900 17 18 16 4 1 1 1 36.0
Cladomics cladogram GONIDEINAE LEPIDODESMINI ACULAMPROTULINA ACUTICOSTINI LANCEOLARIINI UNIONINA OXYNAIINA

Wu et al., 2023 [Publ 4296], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 21 21 19 4 2 2 1 47.0
Cladomics cladogram GONIDEINAE ACUTICOSTINI LANCEOLARIINI ANODONTINI UNIONINI

Liu et al., 2023 [Publ 4310], fig. 4

cst iT tT bi riS oT roT roS iD
0.933 22 22 20 4 2 2 1 50.0
Cladomics cladogram GONIDEINAE ACUTICOSTINI Lanceolaria lanceolata Lanceolaria grayii Lanceolaria gladiola ANODONTINI UNIONINI

Liu et al., 2023 [Publ 4310], fig. 5

cst iT tT bi riS oT roT roS iD
0.900 22 22 15 4 2 2 1 48.0
Cladomics cladogram GONIDEINAE ACUTICOSTINI Lanceolaria grayii Lanceolaria gladiola Lanceolaria lanceolata ANODONTINI UNIONINI

Hou et al., 2024 [Publ 4400], fig. 4

cst iT tT bi riS oT roT roS iD
0.976 71 71 69 4 4 2 1 41.0
Cladomics cladogram AMBLEMINAE LEPIDODESMINI UNIONINI LANCEOLARIINI ANODONTINI

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 23 23 21 4 0 0 0 31.0
Cladomics cladogram ANODONTINI LANCEOLARIINI LEPIDODESMINI UNIONINI

Wu et al., 2024 [Publ 4379], fig. 5

cst iT tT bi riS oT roT roS iD
0.971 51 51 49 4 2 0 0 27.0
Cladomics cladogram GIBBOSULINAE MARGARITIFERINAE LEPIDODESMINI UNIONINI LANCEOLARIINI ANODONTINI

Wang et al., 2025 [Publ 4488], fig. 3B

cst iT tT bi riS oT roT roS iD
0.974 62 62 60 4 2 0 0 44.0
Cladomics cladogram GIBBOSULINAE MARGARITIFERINAE LEPIDODESMINI UNIONINI ACUTICOSTINI ANODONTINI

Wu et al., 2026 [Publ 4825], fig. 6B

cst iT tT bi riS oT roT roS iD
0.974 63 63 62 4 2 0 0 44.0
Cladomics cladogram GIBBOSULINAE MARGARITIFERINAE LEPIDODESMINI UNIONINI LANCEOLARIINI ANODONTINI

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 11 11 9 3 20 18 3 39.0
Cladomics cladogram PARREYSIINAE AMBLEMINAE GONIDEINAE ANODONTINI ACUTICOSTINI UNIONINI

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
0.951 29 29 25 3 52 41 3 47.0
Cladomics cladogram PARREYSIINAE UNIONINI LANCEOLARIINI ANODONTINI Rectidentini AMBLEMINAE CHAMBERLAINIINI LAMPROTULINI Parvasolenaia PSEUDODONTINI Sinosolenaia Gonidea Microcondylaea Leguminaia

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
0.924 28 28 26 3 111 86 3 51.0
Cladomics cladogram PARREYSIINAE Rectidentini PSEUDODONTINI UNIONINI LANCEOLARIINI ANODONTINI AMBLEMINAE GONIDEINI CHAMBERLAINIINI LAMPROTULINI

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
0.837 28 28 19 3 111 86 3 52.0
Cladomics cladogram PARREYSIINAE Rectidentini AMBLEMINAE PSEUDODONTINI GONIDEINI CHAMBERLAINIINI LAMPROTULINI UNIONINI LANCEOLARIINI CRISTARIINA Anodonta anatina Pseudanodonta ALASMIDONTINA Anodonta cygnea Brachyanodon

Bolotov et al., 2017 [Publ 3764], fig. S1

cst iT tT bi riS oT roT roS iD
0.630 28 28 2 3 97 97 3 54.0
Cladomics cladogram PARREYSIINAE PSEUDODONTINI Rectidentini AMBLEMINAE Chamberlainia Sinohyriopsis Lamprotula caveata Lamprotula leaii Potomida Pronodularia Gonidea Leguminaia wheatleyi Leguminaia Microcondylaea Sinosolenaia carinata Sinosolenaia oleivora Sinosolenaia recognita UNIONINI Alasmidonta Anodonta anatina Anodonta cygnea Brachyanodon Lasmigona Pseudanodonta Pyganodon Simpsonaias Strophitus Sinanodonta lauta Cristaria plicata Cristaria radiata Pletholophus Sinanodonta lucida Sinanodonta jourdyi Lanceolaria gladiola Lanceolaria grayii Lanceolaria danzhouensis

Bolotov et al., 2018 [Publ 3826], fig. 2

cst iT tT bi riS oT roT roS iD
0.813 17 17 16 3 65 65 3 46.0
Cladomics cladogram PARREYSIINAE PSEUDODONTINI Rectidentini UNIONINI LANCEOLARIINI ANODONTINI Lamprotula Potomida Pronodularia Gonidea Leguminaia Microcondylaea AMBLEMINI Ortmanniana Lampsilis Cambarunio Elliptio Pleurobema QUADRULINI

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 3 49 42 3 41.0
Cladomics cladogram AMBLEMINAE GONIDEINAE PARREYSIINAE LANCEOLARIINI UNIONINI ANODONTINI

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 3 49 42 3 41.0
Cladomics cladogram AMBLEMINAE GONIDEINAE PARREYSIINAE ANODONTINI LANCEOLARIINI UNIONINI

Zhou et al., 2007 [Publ 2658], fig. 2

cst iT tT bi riS oT roT roS iD
0.739 8 18 16 3 10 10 2 54.0
Cladomics cladogram Sinanodonta Cristaria GONIDEINAE AMBLEMINAE ACULAMPROTULINA ACUTICOSTINI OXYNAIINA

Plazzi et al., 2011 [Publ 3676], fig. 5

cst iT tT bi riS oT roT roS iD
0.833 9 18 11 3 9 8 2 54.0
Cladomics cladogram CRISTARIINA LANCEOLARIINI Anodonta UNIONINI ALASMIDONTINA Pseudanodonta AMBLEMINAE VELESUNIONINAE LAMPROTULINI CHAMBERLAINIINI

Xue et al., 2016 [Publ 3659], fig. 1

cst iT tT bi riS oT roT roS iD
0.929 11 11 9 3 4 4 2 43.0
Cladomics cladogram GONIDEINAE AMBLEMINAE ANODONTINI LANCEOLARIINI UNIONINI

Zhou et al., 2016 [Publ 3710], fig. 1

cst iT tT bi riS oT roT roS iD
0.939 9 9 7 3 11 10 2 41.0
Cladomics cladogram ANODONTINI LEPIDODESMINI UNIONINI MARGARITIFERINAE AMBLEMINAE GONIDEINAE

Fonseca et al., 2016 [Publ 3775], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 18 18 16 3 13 11 2 34.0
Cladomics cladogram AMBLEMINAE GONIDEINAE ANODONTINI LANCEOLARIINI UNIONINI

Zhou et al., 2017 [Publ 3788], fig. 2

cst iT tT bi riS oT roT roS iD
0.964 6 16 14 3 10 10 2 52.0
Cladomics cladogram ANODONTINI LANCEOLARIINI UNIONINI AMBLEMINAE GONIDEINAE

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 18 18 16 3 14 11 2 34.0
Cladomics cladogram AMBLEMINAE GONIDEINAE ANODONTINI LANCEOLARIINI UNIONINI

Wen et al., 2017 [Publ 3795], fig. 3

cst iT tT bi riS oT roT roS iD
0.963 7 7 5 3 9 7 2 43.0
Cladomics cladogram AMBLEMINAE GONIDEINAE ANODONTINI ACULAMPROTULINA LEPIDODESMINI UNIONINA

Wang et al., 2018 [Publ 3846], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 17 17 15 3 14 14 2 33.0
Cladomics cladogram AMBLEMINAE GONIDEINAE ANODONTINI LANCEOLARIINI UNIONINI

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

cst iT tT bi riS oT roT roS iD
1.000 10 10 7 3 10 8 2 39.0
Cladomics cladogram AMBLEMINAE GONIDEINAE ANODONTINI LANCEOLARIINI UNIONINI

Huang et al., 2019 [Publ 3843], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 3 13 10 2 40.0
Cladomics cladogram AMBLEMINAE GONIDEINAE ANODONTINI LANCEOLARIINI UNIONINI

Guerra et al., 2019 [Publ 4168], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 13 13 11 3 28 8 2 37.0
Cladomics cladogram AMBLEMINAE GONIDEINAE ANODONTINI LANCEOLARIINI UNIONINI

Guerra et al., 2019 [Publ 4168], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 3 12 7 2 40.0
Cladomics cladogram AMBLEMINAE GONIDEINAE ANODONTINI LANCEOLARIINI UNIONINI

Guerra et al., 2019 [Publ 4168], fig. 2

cst iT tT bi riS oT roT roS iD
0.971 13 13 11 3 28 8 2 37.0
Cladomics cladogram AMBLEMINAE GONIDEINAE ACULAMPROTULINA OXYNAIINA UNIONINA LANCEOLARIINI ANODONTINI

Guerra et al., 2019 [Publ 4168], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 3 12 7 2 40.0
Cladomics cladogram AMBLEMINAE GONIDEINAE ANODONTINI LANCEOLARIINI UNIONINI

Bolotov et al., 2020 [Publ 4084], fig. 1

cst iT tT bi riS oT roT roS iD
0.864 19 19 17 3 17 13 2 48.0
Cladomics cladogram GONIDEINAE UNIONINI LANCEOLARIINI ANODONTINI

Chen et al., 2020 [Publ 4170], fig. 1

cst iT tT bi riS oT roT roS iD
0.651 15 34 32 3 19 18 2 72.0
Cladomics cladogram Prolasmidonta Alasmidonta Anemina PSEUDODONTINI Lamprotula caveata Chamberlainia OXYNAIINA LEPIDODESMINI ACULAMPROTULINA Platynaias Utterbackia imbecillis Pyganodon Sinanodonta Cristaria ANODONTINA Utterbackia peninsularis Potamilus alatus Lamprotula leaii Sinohyriopsis GONIDEINI GIBBOSULINAE QUADRULINI Toxolasma Lampsilis ornata Potamilus leptodon Lampsilis siliquoidea Lampsilis powellii

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

cst iT tT bi riS oT roT roS iD
0.973 26 26 24 3 50 40 2 47.0
Cladomics cladogram PSEUDODONTINI Rectidentini GONIDEINI CHAMBERLAINIINI LAMPROTULINI AMBLEMINAE UNIONINI LANCEOLARIINI ANODONTINI

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

cst iT tT bi riS oT roT roS iD
0.960 26 26 24 3 50 40 2 49.0
Cladomics cladogram PSEUDODONTINI Rectidentini GONIDEINI CHAMBERLAINIINI LAMPROTULINI AMBLEMINAE UNIONINI LANCEOLARIINI ANODONTINI

Klymus et al., 2025 [Publ 4814], fig. 3A

cst iT tT bi riS oT roT roS iD
1.000 15 15 13 3 21 20 2 39.0
Cladomics cladogram AMBLEMINAE GONIDEINAE LANCEOLARIINI ANODONTINI UNIONINI

Klymus et al., 2025 [Publ 4814], fig. 3B

cst iT tT bi riS oT roT roS iD
0.981 15 15 13 3 21 20 2 39.0
Cladomics cladogram AMBLEMINAE GONIDEINAE ANODONTINI OXYNAIINA UNIONINA ACULAMPROTULINA LANCEOLARIINI

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 19 19 17 3 28 22 2 31.0
Cladomics cladogram AMBLEMINAE GONIDEINAE ANODONTINI LANCEOLARIINI UNIONINI

Sano et al., 2017 [Publ 3799], fig. 1

cst iT tT bi riS oT roT roS iD
0.957 13 13 14 3 8 4 1 52.0
Cladomics cladogram GONIDEINAE ANODONTINI LANCEOLARIINI UNIONINI

Kano et al., 2019 [Publ 4194], fig. 6

cst iT tT bi riS oT roT roS iD
0.895 6 9 7 3 4 3 1 52.0
Cladomics cladogram MARGARITIFERINAE ANODONTINI GONIDEINAE LANCEOLARIINI UNIONINI

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

cst iT tT bi riS oT roT roS iD
0.762 8 8 8 3 4 4 1 48.0
Cladomics cladogram GONIDEINAE ANODONTINI Nodularia LANCEOLARIINI Inversiunio

Dai et al., 2024 [Publ 4386], fig. 3

cst iT tT bi riS oT roT roS iD
0.952 13 13 11 3 1 1 1 52.0
Cladomics cladogram PARREYSIINAE UNIONINA MIDDENDORFFINAIINA OXYNAIINA ACULAMPROTULINA ANODONTINI LEPIDODESMINI

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 3 0 0 0 38.0
Cladomics cladogram LANCEOLARIINI ANODONTINI UNIONINI

Fukata & Iigo, 2019 [Publ 4108], fig. 1

cst iT tT bi riS oT roT roS iD
0.941 17 17 15 3 0 0 0 39.0
Cladomics cladogram UNIONINA OXYNAIINA ACULAMPROTULINA ACUTICOSTINI LANCEOLARIINI

Riccardi et al., 2020 [Publ 4089], fig. 2

cst iT tT bi riS oT roT roS iD
0.900 17 17 15 3 2 0 0 39.0
Cladomics cladogram MARGARITIFERINAE ANODONTINI LANCEOLARIINI UNIONINI

Wu et al., 2022 [Publ 4238], fig. 2e

cst iT tT bi riS oT roT roS iD
1.000 18 18 16 3 0 0 0 32.0
Cladomics cladogram ANODONTINI LANCEOLARIINI UNIONINI

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

cst iT tT bi riS oT roT roS iD
1.000 22 22 20 3 0 0 0 50.0
Cladomics cladogram UNIONINI LANCEOLARIINI ANODONTINI

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

cst iT tT bi riS oT roT roS iD
1.000 22 22 20 3 0 0 0 51.0
Cladomics cladogram UNIONINI LANCEOLARIINI ANODONTINI

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

cst iT tT bi riS oT roT roS iD
0.912 5 19 12 2 29 13 3 57.0
Cladomics cladogram Pseudomulleria ANODONTINI GONIDEINAE UNIONINI MARGARITIFERINAE AMBLEMINAE

Hoeh et al., 2001 [Publ 2652], fig. 14f2

cst iT tT bi riS oT roT roS iD
0.824 5 15 10 2 26 13 3 53.0
Cladomics cladogram AMBLEMINI PARREYSIINAE Eurynia Fusconaia GONIDEINAE Pleurobema QUADRULINI Ortmanniana Glebula Potamilus Ligumia Toxolasma Cyrtonaias UNIONINI ANODONTINI IRIDININAE Anodontites guanarensis Anodontites trigona MONOCONDYLAEINAE RHIPIDODONTINI Echyridella Lortiella CASTALIINI Velesunio

Hoeh et al., 2001 [Publ 2652], fig. 14f4

cst iT tT bi riS oT roT roS iD
0.882 5 5 2 2 26 13 3 44.0
Cladomics cladogram GONIDEINAE AMBLEMINAE Unio pictorum Unio tumidus ANODONTINI

Hoeh et al., 2002 [Publ 2653], fig. 2

cst iT tT bi riS oT roT roS iD
0.853 5 5 3 2 26 13 3 45.0
Cladomics cladogram AMBLEMINAE GONIDEINAE Unio pictorum Unio tumidus ANODONTINI

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

cst iT tT bi riS oT roT roS iD
0.882 5 5 3 2 26 13 3 43.0
Cladomics cladogram GONIDEINAE PARREYSIINAE Fusconaia AMBLEMINI QUADRULINI Pleurobema Eurynia Cyrtonaias UNIONINI ANODONTINI Ortmanniana Glebula Potamilus Ligumia Toxolasma

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 2 26 13 3 43.0
Cladomics cladogram PARREYSIINAE UNIONINI ANODONTINI GONIDEINAE AMBLEMINAE

Graf & Cummings, 2006 [Publ 1968], fig. 2

cst iT tT bi riS oT roT roS iD
0.977 6 6 4 2 41 19 3 47.0
Cladomics cladogram PSEUDODONTINI PARREYSIINAE Rectidentini UNIONINI ANODONTINI GONIDEINI AMBLEMINAE

Hoeh et al., 2009 [Publ 2716], fig. 3

cst iT tT bi riS oT roT roS iD
0.930 6 6 4 2 41 19 3 47.0
Cladomics cladogram UNIONINI ANODONTINI Rectidentini AMBLEMINAE GONIDEINI PSEUDODONTINI

Hoeh et al., 2009 [Publ 2716], fig. 4

cst iT tT bi riS oT roT roS iD
0.927 6 6 4 2 40 18 3 47.0
Cladomics cladogram PSEUDODONTINI Rectidentini UNIONINI ANODONTINI GONIDEINI AMBLEMINAE

Hoeh et al., 2009 [Publ 2716], fig. 5

cst iT tT bi riS oT roT roS iD
0.929 6 6 4 2 40 18 3 47.0
Cladomics cladogram PSEUDODONTINI Rectidentini UNIONINI ANODONTINI GONIDEINI AMBLEMINAE

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

cst iT tT bi riS oT roT roS iD
0.756 9 9 7 2 16 16 3 51.0
Cladomics cladogram PARREYSIINAE OXYNAIINA Anodonta cygnea Unio pictorum 2 Unio pictorum 1 Unio caffer Anodonta ALASMIDONTINA LAMPROTULINI QUADRULINI GONIDEINI Eurynia Cambarunio Ptychobranchus AMBLEMINI Pleurobema Truncilla Lampsilis Epioblasma Ligumia Obliquaria Ortmanniana

Whelan et al., 2011 [Publ 3380], fig. 1-ML

cst iT tT bi riS oT roT roS iD
0.956 4 4 2 2 32 20 3 46.0
Cladomics cladogram PARREYSIINAE PSEUDODONTINI Rectidentini ANODONTINI UNIONINI GONIDEINI LAMPROTULINI AMBLEMINAE

Whelan et al., 2011 [Publ 3380], fig. 1-MP

cst iT tT bi riS oT roT roS iD
0.956 4 4 2 2 32 20 3 46.0
Cladomics cladogram PARREYSIINAE Rectidentini PSEUDODONTINI ANODONTINI UNIONINI GONIDEINI LAMPROTULINI AMBLEMINAE

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 37 27 3 45.0
Cladomics cladogram PARREYSIINAE UNIONINI ANODONTINI AMBLEMINAE GONIDEINAE

Araujo et al., 2017 [Publ 3801], fig. 2

cst iT tT bi riS oT roT roS iD
0.967 4 4 2 2 10 9 3 46.0
Cladomics cladogram PARREYSIINAE AMBLEMINAE GONIDEINAE OXYNAIINA ACULAMPROTULINA ANODONTINI UNIONINA

Sonowal et al., 2021 [Publ 4221], fig. 3a

cst iT tT bi riS oT roT roS iD
0.773 4 4 0 2 11 5 3 45.0
Cladomics cladogram PLEUROBEMINI LAMPSILINI ANODONTINI Unio ravoisieri Unio pictorum ACULAMPROTULINA

Sonowal et al., 2021 [Publ 4221], fig. 3b

cst iT tT bi riS oT roT roS iD
0.682 4 9 0 2 11 5 3 50.0
Cladomics cladogram HYRIINI VELESUNIONINAE HYRIDELLINI ACULAMPROTULINA Lamprotula LAMPSILINI Potomida Unio pictorum ANODONTINI Unio ravoisieri PARREYSIINAE PLEUROBEMINI

Sonowal et al., 2021 [Publ 4221], fig. 4

cst iT tT bi riS oT roT roS iD
0.545 4 5 3 2 12 6 3 47.0
Cladomics cladogram Lamellidens phenchooganjensis Lamprotula Potomida LAMPSILINI PLEUROBEMINI ACULAMPROTULINA Unio pictorum ANODONTINI Unio ravoisieri Lamellidens corrianus

Zieritz et al., 2024 [Publ 4381], fig. 2

cst iT tT bi riS oT roT roS iD
0.965 2 2 0 2 43 41 3 --
Cladomics cladogram AMBLEMINAE PARREYSIINAE ANODONTINI UNIONINI CHAMBERLAINIINI Discomya Rectidentini SCHEPMANIINI PSEUDODONTINI Caudiculatus Potomida GONIDEINI

Rosenberg et al., 1994 [Publ 2719], fig. 3

cst iT tT bi riS oT roT roS iD
0.643 4 15 2 2 15 12 2 55.0
Cladomics cladogram GONIDEINAE Elliptio Fusconaia AMBLEMINI Pleurobema UNIONINI Pyganodon cataracta Pyganodon grandis Utterbackia Lampsilis teres Lampsilis straminea Megalonaias Obliquaria Theliderma Quadrula Uniomerus

Rosenberg et al., 1997 [Publ 2720], fig. 2A

cst iT tT bi riS oT roT roS iD
0.645 5 18 1 2 17 13 2 59.0
Cladomics cladogram None 1 ANODONTINA GONIDEINAE Pyganodon grandis Utterbackia Pyganodon cataracta Lampsilis straminea Lampsilis teres Megalonaias Obliquaria Theliderma Quadrula Uniomerus Amblema Elliptio Fusconaia Plectomerus Pleurobema UNIONINI

Rosenberg et al., 1997 [Publ 2720], fig. 2B

cst iT tT bi riS oT roT roS iD
0.625 6 18 1 2 16 12 2 58.5
Cladomics cladogram Cumberlandia None 1 None 2 Margaritifera ANODONTINA GONIDEINAE Pyganodon grandis Utterbackia Pyganodon cataracta Lampsilis straminea Lampsilis teres Megalonaias Obliquaria Theliderma Quadrula Uniomerus Amblema Elliptio Fusconaia Plectomerus ACULAMPROTULINA UNIONINA

Graf, 2000 [Publ 2647], fig. 1

cst iT tT bi riS oT roT roS iD
0.778 2 4 0 2 15 4 2 47.0
Cladomics cladogram UNIONINI PLEUROBEMINI ANODONTINI LAMPSILINI PARREYSIINI COELATURINI CASTALIINI RHIPIDODONTINI HYRIDELLINI VELESUNIONINAE IRIDININAE ANODONTITINAE MYCETOPODINAE Etheria Mulleria

Graf, 2002 [Publ 2651], fig. 2

cst iT tT bi riS oT roT roS iD
0.963 4 4 2 2 14 8 2 45.0
Cladomics cladogram PSEUDODONTINI Rectidentini UNIONINI ANODONTINI GONIDEINI AMBLEMINAE

Araujo et al., 2009 [Publ 2589], fig. 7A

cst iT tT bi riS oT roT roS iD
0.812 8 8 4 2 3 2 2 49.0
Cladomics cladogram AMBLEMINAE GONIDEINAE OXYNAIINA Unio gibbus Unio tumidus ANODONTINI Unio pictorum 1 Unio pictorum 2 Unio mancus Unio tumidiformis Unio crassus

Araujo et al., 2009 [Publ 2589], fig. 7B

cst iT tT bi riS oT roT roS iD
1.000 8 8 7 2 3 2 2 47.0
Cladomics cladogram GONIDEINAE AMBLEMINAE ANODONTINI UNIONINI

Araujo et al., 2009 [Publ 2589], fig. 7C

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 2 3 2 2 46.0
Cladomics cladogram GONIDEINAE AMBLEMINAE ANODONTINI UNIONINI

Reis et al., 2013 [Publ 3580], fig. 4

cst iT tT bi riS oT roT roS iD
0.944 9 9 7 2 5 3 2 45.0
Cladomics cladogram AMBLEMINAE GONIDEINAE ANODONTINI UNIONINI

Reis et al., 2013 [Publ 3580], fig. 5

cst iT tT bi riS oT roT roS iD
0.944 10 10 8 2 5 3 2 43.0
Cladomics cladogram AMBLEMINAE GONIDEINAE ANODONTINI UNIONINI

Huang et al., 2013 [Publ 3629], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 2 9 8 2 39.0
Cladomics cladogram UNIONINI ANODONTINI AMBLEMINAE GONIDEINAE

Prié & Puillandre, 2014 [Publ 3529], fig. 2

cst iT tT bi riS oT roT roS iD
0.933 14 14 12 2 3 2 2 50.0
Cladomics cladogram AMBLEMINAE GONIDEINAE ANODONTINI UNIONINI

Wu et al., 2016 [Publ 3711], fig. 1

cst iT tT bi riS oT roT roS iD
0.971 12 12 10 2 9 9 2 38.0
Cladomics cladogram AMBLEMINAE GONIDEINAE UNIONINI ANODONTINI

Fonseca et al., 2016 [Publ 3775], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 2 5 5 2 42.0
Cladomics cladogram AMBLEMINAE GONIDEINAE UNIONINI ANODONTINI

Feng et al., 2016 [Publ 3800], fig. 2

cst iT tT bi riS oT roT roS iD
0.875 7 12 10 2 7 5 2 50.0
Cladomics cladogram MARGARITIFERINAE ANODONTINA Utterbackia GONIDEINAE AMBLEMINAE UNIONINI Pyganodon CRISTARIINA

Guerra et al., 2017 [Publ 3772], fig. 2a

cst iT tT bi riS oT roT roS iD
0.969 10 10 8 2 15 9 2 40.0
Cladomics cladogram AMBLEMINAE GONIDEINAE UNIONINI ANODONTINI

Guerra et al., 2017 [Publ 3772], fig. 2b

cst iT tT bi riS oT roT roS iD
0.969 10 10 8 2 15 9 2 40.0
Cladomics cladogram AMBLEMINAE GONIDEINAE UNIONINA ACULAMPROTULINA ANODONTINI

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 2 7 7 2 40.0
Cladomics cladogram UNIONINI ANODONTINI AMBLEMINAE GONIDEINAE

Burzynski et al., 2017 [Publ 3774], fig. 1-top

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 2 7 7 2 40.0
Cladomics cladogram UNIONINI ANODONTINI AMBLEMINAE GONIDEINAE

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 2 6 5 2 42.0
Cladomics cladogram AMBLEMINAE GONIDEINAE UNIONINI ANODONTINI

Huang et al., 2018 [Publ 3828], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 18 18 16 2 24 14 2 32.0
Cladomics cladogram AMBLEMINAE GONIDEINAE ANODONTINI UNIONINI

Huang et al., 2018 [Publ 3828], fig. 6-M

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 2 9 7 2 40.0
Cladomics cladogram AMBLEMINAE GONIDEINAE ANODONTINI UNIONINI

Chase et al., 2018 [Publ 3847], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 2 6 4 2 42.0
Cladomics cladogram GONIDEINAE AMBLEMINAE ANODONTINI UNIONINI

Chase et al., 2018 [Publ 3861], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 2 6 4 2 41.0
Cladomics cladogram AMBLEMINAE GONIDEINAE UNIONINI ANODONTINI

Chase et al., 2018 [Publ 3861], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 2 6 4 2 41.0
Cladomics cladogram GONIDEINAE AMBLEMINAE UNIONINI ANODONTINI

Wu et al., 2018 [Publ 4143], fig. 2

cst iT tT bi riS oT roT roS iD
0.971 13 13 11 2 9 9 2 37.0
Cladomics cladogram AMBLEMINAE GONIDEINAE UNIONINI ANODONTINI

Froufe et al., 2019 [Publ 3936], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 2 12 11 2 39.0
Cladomics cladogram AMBLEMINAE GONIDEINAE UNIONINI ANODONTINI

Froufe et al., 2019 [Publ 3936], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 2 12 11 2 39.0
Cladomics cladogram AMBLEMINAE GONIDEINAE UNIONINI ANODONTINI

Wu et al., 2020 [Publ 4147], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 18 18 16 2 12 11 2 34.0
Cladomics cladogram AMBLEMINAE GONIDEINAE ANODONTINI UNIONINI

Wu et al., 2021 [Publ 4215], fig. 3

cst iT tT bi riS oT roT roS iD
0.970 4 4 2 2 20 20 2 46.0
Cladomics cladogram PARREYSIINAE GONIDEINAE ACUTICOSTINI UNIONINI

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 2 16 15 2 39.0
Cladomics cladogram AMBLEMINAE GONIDEINAE UNIONINI ANODONTINI

Lyubas et al., 2022 [Publ 4255], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 28 28 25 2 3 3 2 60.0
Cladomics cladogram PARREYSIINAE GONIDEINAE ANODONTINI UNIONINI

Lyubas et al., 2022 [Publ 4255], fig. S1

cst iT tT bi riS oT roT roS iD
0.933 28 28 19 2 3 3 2 60.0
Cladomics cladogram PARREYSIINAE GONIDEINAE ANODONTINI UNIONINI

Gomes-dos-Santos et al., 2023 [Publ 4463], fig. 2a

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 19 2 2 --
Cladomics cladogram AMBLEMINAE GONIDEINAE ANODONTINI UNIONINI

Gomes-dos-Santos et al., 2023 [Publ 4463], fig. 2b

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 20 2 2 --
Cladomics cladogram AMBLEMINAE GONIDEINAE ANODONTINI UNIONINI

Nagel et al., 1998 [Publ 1946], fig. 1

cst iT tT bi riS oT roT roS iD
0.929 7 7 5 2 2 2 1 41.0
Cladomics cladogram GONIDEINI LAMPROTULINI ANODONTINI UNIONINI

Nagel et al., 1998 [Publ 1946], fig. 9

cst iT tT bi riS oT roT roS iD
0.929 3 3 1 2 3 2 1 44.5
Cladomics cladogram LAMPROTULINI GONIDEINI UNIONINI ANODONTINI

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

cst iT tT bi riS oT roT roS iD
0.939 8 25 19 2 22 17 1 64.0
Cladomics cladogram MARGARITIFERINAE UNIONINI ANODONTINI AMBLEMINAE

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

cst iT tT bi riS oT roT roS iD
0.939 8 25 23 2 22 17 1 64.0
Cladomics cladogram MARGARITIFERINAE UNIONINI ANODONTINI AMBLEMINAE

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 9 2 1 --
Cladomics cladogram AMBLEMINAE UNIONINI ANODONTINI

Huff et al., 2004 [Publ 2697], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 10 1 1 --
Cladomics cladogram AMBLEMINAE UNIONINI ANODONTINI

Huff et al., 2004 [Publ 2697], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 9 1 1 --
Cladomics cladogram AMBLEMINAE UNIONINI ANODONTINI

Huff et al., 2004 [Publ 2697], fig. 5-6

cst iT tT bi riS oT roT roS iD
0.909 2 3 0 2 10 1 1 46.5
Cladomics cladogram MARGARITIFERINAE UNIONINI ANODONTINI AMBLEMINAE

Manendo et al., 2008 [Publ 2669], fig. 1

cst iT tT bi riS oT roT roS iD
0.952 8 8 6 2 5 5 1 43.0
Cladomics cladogram AMBLEMINAE UNIONINI ANODONTINI

Marshall et al., 2014 [Publ 3593], fig. 1

cst iT tT bi riS oT roT roS iD
0.909 2 2 0 2 12 1 1 --
Cladomics cladogram MARGARITIFERINAE ANODONTINI UNIONINI

Bolotov et al., 2016 [Publ 3743], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 27 1 1 45.0
Cladomics cladogram AMBLEMINAE ANODONTINI UNIONINI

Bolotov et al., 2016 [Publ 3743], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 27 1 1 45.0
Cladomics cladogram AMBLEMINAE ANODONTINI UNIONINI

Lopes-Lima et al., 2017 [Publ 3734], fig. 1

cst iT tT bi riS oT roT roS iD
0.929 12 12 8 2 5 2 1 46.0
Cladomics cladogram GONIDEINAE ANODONTINI UNIONINI

Froufe et al., 2017 [Publ 3802], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 2 3 2 1 41.0
Cladomics cladogram GONIDEINAE UNIONINI ANODONTINI

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 2 2 2 1 41.0
Cladomics cladogram AMBLEMINAE UNIONINI ANODONTINI

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 2 2 2 1 41.0
Cladomics cladogram AMBLEMINAE UNIONINI ANODONTINI

Chase et al., 2018 [Publ 3847], fig. 4

cst iT tT bi riS oT roT roS iD
0.946 16 16 15 2 23 18 1 42.0
Cladomics cladogram UNIONINI ANODONTINI LAMPSILINI HYRIINAE AMBLEMINI PLEUROBEMINI QUADRULINI

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 8 1 1 --
Cladomics cladogram GONIDEINAE UNIONINI ANODONTINI

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 4 1 1 --
Cladomics cladogram GONIDEINAE UNIONINI ANODONTINI

Lopes-Lima et al., 2021 [Publ 4217], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 19 19 17 2 6 6 1 53.0
Cladomics cladogram GONIDEINAE ANODONTINI UNIONINI

Pfarrer et al., 2022 [Publ 4305], fig. 2-5

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 2 2 2 1 36.0
Cladomics cladogram GONIDEINAE UNIONINI ANODONTINI

Wu et al., 2023 [Publ 4339], fig. 5

cst iT tT bi riS oT roT roS iD
0.960 11 11 9 2 8 6 1 41.0
Cladomics cladogram AMBLEMINAE LEPIDODESMINI ANODONTINI

Wu et al., 2024 [Publ 4379], fig. 4_6

cst iT tT bi riS oT roT roS iD
0.938 18 18 16 2 5 3 1 46.0
Cladomics cladogram GONIDEINAE ACULAMPROTULINA ANODONTINI OXYNAIINA

Xiang et al., 2025 [Publ 4433], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 12 12 10 2 1 1 1 38.0
Cladomics cladogram AMBLEMINAE LANCEOLARIINI ANODONTINI

Hou et al., 2025 [Publ 4445], fig. 5

cst iT tT bi riS oT roT roS iD
0.972 50 50 48 2 2 2 1 32.0
Cladomics cladogram AMBLEMINAE ANODONTINI UNIONINI

Wu et al., 2026 [Publ 4825], fig. 5

cst iT tT bi riS oT roT roS iD
0.929 17 17 15 2 2 2 1 49.0
Cladomics cladogram GONIDEINAE OXYNAIINA ACULAMPROTULINA LANCEOLARIINI

Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5_7

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 2 0 0 0 43.0
Cladomics cladogram ANODONTINI UNIONINI

Källersjö et al., 2005 [Publ 2684], fig. 1A

cst iT tT bi riS oT roT roS iD
0.667 6 7 2 2 1 0 0 44.0
Cladomics cladogram MARGARITIFERINAE ANODONTINI UNIONINI

Källersjö et al., 2005 [Publ 2684], fig. 1B-F

cst iT tT bi riS oT roT roS iD
0.667 6 7 4 2 1 0 0 45.0
Cladomics cladogram MARGARITIFERINAE ANODONTINI UNIONINI

Taylor et al., 2007 [Publ 2696], fig. 1-3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 2 0 0 --
Cladomics cladogram MARGARITIFERINAE ANODONTINI UNIONINI

Chong et al., 2008 [Publ 2668], fig. 3

cst iT tT bi riS oT roT roS iD
0.773 18 18 10 2 0 0 0 55.0
Cladomics cladogram Anodonta cygnea Pyganodon Alasminota Anodonta anatina UNIONINI CRISTARIINA Brachyanodon nuttalliana Brachyanodon californiensis Pseudodontoideus Alasmidonta Pterosyna Pseudanodonta Strophitus Prolasmidonta Platynaias Brachyanodon oregonensis 4 Brachyanodon oregonensis 3 Brachyanodon oregonensis 2 Brachyanodon oregonensis 1 Brachyanodon kennerlyi

Soroka, 2010 [Publ 3404], fig. 2

cst iT tT bi riS oT roT roS iD
0.818 7 7 4 2 0 0 0 44.0
Cladomics cladogram CRISTARIINA UNIONINI ANODONTINA

Soroka, 2010 [Publ 3404], fig. 2-M

cst iT tT bi riS oT roT roS iD
0.900 5 5 3 2 0 0 0 45.0
Cladomics cladogram CRISTARIINA ANODONTINA UNIONINI

Soroka, 2010 [Publ 3404], fig. 3a

cst iT tT bi riS oT roT roS iD
0.818 9 9 7 2 0 0 0 45.0
Cladomics cladogram CRISTARIINA ANODONTINA UNIONINI

Soroka, 2010 [Publ 3404], fig. 3a-M

cst iT tT bi riS oT roT roS iD
0.900 6 6 4 2 0 0 0 46.0
Cladomics cladogram ANODONTINA CRISTARIINA UNIONINI

Soroka, 2010 [Publ 3404], fig. 3b

cst iT tT bi riS oT roT roS iD
0.727 9 9 5 2 0 0 0 45.0
Cladomics cladogram Anodonta anatina Anodonta cygnea Pseudanodonta CRISTARIINA UNIONINI

Soroka, 2010 [Publ 3404], fig. 3b-M

cst iT tT bi riS oT roT roS iD
0.900 6 6 4 2 0 0 0 44.0
Cladomics cladogram ANODONTINA CRISTARIINA UNIONINI

Soroka, 2010 [Publ 3404], fig. 4

cst iT tT bi riS oT roT roS iD
0.727 7 7 5 2 0 0 0 45.0
Cladomics cladogram Anodonta anatina UNIONINI CRISTARIINA Anodonta cygnea Pseudanodonta

Soroka, 2010 [Publ 3404], fig. 5a

cst iT tT bi riS oT roT roS iD
0.818 8 8 6 2 0 0 0 46.0
Cladomics cladogram UNIONINI ANODONTINI

Soroka, 2010 [Publ 3404], fig. 5b

cst iT tT bi riS oT roT roS iD
0.727 8 8 5 2 0 0 0 45.0
Cladomics cladogram Anodonta anatina CRISTARIINA UNIONINI Anodonta cygnea Pseudanodonta

Soroka, 2010 [Publ 3404], fig. 8a

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 2 0 0 0 40.0
Cladomics cladogram UNIONINI ANODONTINI

Soroka, 2010 [Publ 3404], fig. 8a-M

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 0 0 0 43.0
Cladomics cladogram ANODONTINI UNIONINI

Soroka, 2010 [Publ 3404], fig. 8b

cst iT tT bi riS oT roT roS iD
0.900 6 6 4 2 0 0 0 42.0
Cladomics cladogram CRISTARIINA ANODONTINA UNIONINI

Soroka, 2010 [Publ 3404], fig. 8b-M

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 0 0 0 43.0
Cladomics cladogram ANODONTINI UNIONINI

Zieritz et al., 2012 [Publ 3673], fig. 4

cst iT tT bi riS oT roT roS iD
0.636 7 8 4 2 1 0 0 45.0
Cladomics cladogram ANODONTINA CRISTARIINA MARGARITIFERINAE UNIONINI

Araujo et al., 2018 [Publ 3779], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 17 17 13 2 1 0 0 49.0
Cladomics cladogram MARGARITIFERINAE ANODONTINI UNIONINI

Bogatov et al., 2018 [Publ 3859], fig. 1

cst iT tT bi riS oT roT roS iD
0.667 5 5 3 2 1 0 0 45.0
Cladomics cladogram MARGARITIFERINAE Unio tumidus ANODONTINI Unio crassus Unio pictorum

Bogatov et al., 2018 [Publ 3859], fig. 2

cst iT tT bi riS oT roT roS iD
0.667 5 5 3 2 1 0 0 45.0
Cladomics cladogram MARGARITIFERINAE Unio crassus Unio pictorum Unio tumidus ANODONTINI

Klishko et al., 2019 [Publ 3886], fig. 9

cst iT tT bi riS oT roT roS iD
1.000 10 10 7 2 1 0 0 42.0
Cladomics cladogram MARGARITIFERINAE ANODONTINI UNIONINI

Lopes-Lima et al., 2020 [Publ 4080], fig. 11

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 2 2 0 0 46.0
Cladomics cladogram MARGARITIFERINAE UNIONINI LANCEOLARIINI

Tomilova et al., 2020 [Publ 4097], fig. 3

cst iT tT bi riS oT roT roS iD
0.818 8 8 5 2 0 0 0 43.0
Cladomics cladogram UNIONINI CRISTARIINA ANODONTINA

Bogan et al., 2023 [Publ 4295], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 29 29 26 2 2 0 0 61.0
Cladomics cladogram GIBBOSULINAE MARGARITIFERINAE UNIONINI ANODONTINI

Hou et al., 2025 [Publ 4401], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 28 28 26 2 0 0 0 52.0
Cladomics cladogram LEPIDODESMINI UNIONINI

Hou et al., 2025 [Publ 4432], fig. 3

cst iT tT bi riS oT roT roS iD
0.909 11 11 9 2 0 0 0 46.0
Cladomics cladogram LEPIDODESMINI UNIONINI

Hou et al., 2025 [Publ 4432], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 28 28 26 2 0 0 0 52.0
Cladomics cladogram LEPIDODESMINI UNIONINI

Wu et al., 2025 [Publ 4442], fig. 2

cst iT tT bi riS oT roT roS iD
0.933 35 35 33 2 0 0 0 67.0
Cladomics cladogram UNIONINI ANODONTINI

Wu et al., 2025 [Publ 4442], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 17 17 15 2 0 0 0 51.0
Cladomics cladogram UNIONINI ANODONTINI

Altaba et al., 2025 [Publ 4849], fig. 10

cst iT tT bi riS oT roT roS iD
0.875 12 12 10 2 0 0 0 41.0
Cladomics cladogram UNIONINI ANODONTINI

Altaba et al., 2025 [Publ 4849], fig. 11

cst iT tT bi riS oT roT roS iD
0.875 14 14 12 2 0 0 0 42.0
Cladomics cladogram UNIONINI ANODONTINI

Altaba et al., 2025 [Publ 4849], fig. 12

cst iT tT bi riS oT roT roS iD
0.733 13 13 12 2 0 0 0 49.0
Cladomics cladogram UNIONINI ANODONTINI

Chen et al., 2026 [Publ 4805], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 34 34 32 2 0 0 0 70.0
Cladomics cladogram UNIONINI ANODONTINI

Wu et al., 2026 [Publ 4825], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 12 12 10 2 0 0 0 48.0
Cladomics cladogram UNIONINI LANCEOLARIINI

Huang et al., 2018 [Publ 3828], fig. 3_5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 27 4 3 --
Cladomics cladogram GONIDEINAE UNIONINI AMBLEMINAE PARREYSIINAE

Wu et al., 2020 [Publ 4121], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 1 29 6 3 43.0
Cladomics cladogram AMBLEMINAE PARREYSIINAE GONIDEINAE UNIONINI

Wu et al., 2020 [Publ 4121], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 1 29 6 3 43.0
Cladomics cladogram AMBLEMINAE PARREYSIINAE GONIDEINAE UNIONINI

Smith, 2021 [Publ 4331], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 17 13 3 43.0
Cladomics cladogram AMBLEMINAE GONIDEINAE PARREYSIINAE ANODONTINI

Campbell et al., 2005 [Publ 2660], fig. 1

cst iT tT bi riS oT roT roS iD
0.853 6 6 4 1 96 94 2 49.0
Cladomics cladogram CHAMBERLAINIINI ANODONTINI AMBLEMINAE GONIDEINI LAMPROTULINI

Campbell et al., 2005 [Publ 2660], fig. 2

cst iT tT bi riS oT roT roS iD
0.824 6 6 3 1 96 94 2 47.0
Cladomics cladogram CHAMBERLAINIINI ANODONTINI AMBLEMINAE GONIDEINI LAMPROTULINI

Walker et al., 2006 [Publ 2666], fig. 2

cst iT tT bi riS oT roT roS iD
0.875 3 3 1 1 12 9 2 45.5
Cladomics cladogram ANODONTINI LAMPROTULINI GONIDEINI PSEUDODONTINI LAMPSILINI PLEUROBEMINI MARGARITIFERINAE

Walker et al., 2006 [Publ 2666], fig. 3

cst iT tT bi riS oT roT roS iD
0.913 3 3 1 1 12 9 2 45.5
Cladomics cladogram PSEUDODONTINI Gonidea Inversidens ANODONTINI AMBLEMINAE

Walker et al., 2006 [Publ 2666], fig. 4

cst iT tT bi riS oT roT roS iD
0.913 3 3 1 1 12 9 2 45.5
Cladomics cladogram PSEUDODONTINI Gonidea Inversidens ANODONTINI AMBLEMINAE

Walker et al., 2006 [Publ 2666], fig. 6

cst iT tT bi riS oT roT roS iD
0.900 2 2 0 1 34 8 2 --
Cladomics cladogram ANODONTINI MARGARITIFERINAE GONIDEINAE AMBLEMINAE

Walker et al., 2006 [Publ 2666], fig. 6-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 12 8 2 --
Cladomics cladogram GONIDEINAE ANODONTINI AMBLEMINAE

Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 3 2 --
Cladomics cladogram ANODONTINI AMBLEMINAE GONIDEINAE

Jiang et al., 2010 [Publ 3438], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 3 2 --
Cladomics cladogram AMBLEMINAE ANODONTINI GONIDEINAE

Khalloufi et al., 2011 [Publ 2748], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 9 9 6 1 2 2 2 47.0
Cladomics cladogram AMBLEMINAE GONIDEINAE UNIONINI

Breton et al., 2011 [Publ 3376], fig. 3

cst iT tT bi riS oT roT roS iD
0.921 12 12 11 1 26 21 2 48.0
Cladomics cladogram QUADRULINA LAMPROTULINI CHAMBERLAINIINI ANODONTINI Megalonaias PLEUROBEMINI AMBLEMINI LAMPSILINI

Chen et al., 2012 [Publ 3624], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 5 5 2 --
Cladomics cladogram AMBLEMINAE ANODONTINI GONIDEINAE

Inoue et al., 2013 [Publ 3528], fig. 3

cst iT tT bi riS oT roT roS iD
0.958 1 1 0 1 16 16 2 --
Cladomics cladogram GONIDEINAE ANODONTINI AMBLEMINAE

Huang et al., 2013 [Publ 3629], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 4 4 2 --
Cladomics cladogram ANODONTINI GONIDEINAE AMBLEMINAE

Inoue et al., 2014 [Publ 3635], fig. 3A

cst iT tT bi riS oT roT roS iD
0.962 14 14 13 1 6 5 2 51.0
Cladomics cladogram AMBLEMINI PLEUROBEMINI LAMPSILINI QUADRULINI GONIDEINAE ANODONTINI

Sheng et al., 2014 [Publ 3704], fig. 2

cst iT tT bi riS oT roT roS iD
0.789 2 8 0 1 6 6 2 50.0
Cladomics cladogram CRISTARIINA ALASMIDONTINA Lampsilis GONIDEINAE Venustaconcha QUADRULINI

Froufe et al., 2015 [Publ 3664], fig. 1

cst iT tT bi riS oT roT roS iD
0.957 3 3 1 1 9 9 2 43.0
Cladomics cladogram GONIDEINI LAMPROTULINI CHAMBERLAINIINI AMBLEMINAE ANODONTINI

Froufe et al., 2015 [Publ 3664], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 5 5 2 43.0
Cladomics cladogram ANODONTINI AMBLEMINAE GONIDEINAE

Wei et al., 2016 [Publ 3665], fig. 0

cst iT tT bi riS oT roT roS iD
0.812 2 4 0 1 4 4 2 46.0
Cladomics cladogram GONIDEINAE CRISTARIINA LAMPSILINI ALASMIDONTINA QUADRULINI

Guerra et al., 2017 [Publ 3772], fig. 2a-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 6 4 2 --
Cladomics cladogram GONIDEINAE ANODONTINI AMBLEMINAE

Guerra et al., 2017 [Publ 3772], fig. 2b-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 6 4 2 --
Cladomics cladogram GONIDEINAE ANODONTINI AMBLEMINAE

Amaro et al., 2017 [Publ 3791], fig. 2

cst iT tT bi riS oT roT roS iD
0.957 5 5 3 1 11 8 2 43.0
Cladomics cladogram ANODONTINI PLEUROBEMINI GONIDEINAE LAMPSILINI QUADRULINI

Amaro et al., 2017 [Publ 3791], fig. 2-M

cst iT tT bi riS oT roT roS iD
0.957 4 4 1 1 6 6 2 46.0
Cladomics cladogram AMBLEMINAE GONIDEINAE ANODONTINI

Amaro et al., 2017 [Publ 3791], fig. 3A

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 8 5 2 42.0
Cladomics cladogram ANODONTINI GONIDEINAE AMBLEMINAE

Amaro et al., 2017 [Publ 3791], fig. 3A-M

cst iT tT bi riS oT roT roS iD
0.941 2 2 0 1 5 4 2 --
Cladomics cladogram QUADRULINI GONIDEINAE LAMPSILINI ANODONTINI

Amaro et al., 2017 [Publ 3791], fig. 3B

cst iT tT bi riS oT roT roS iD
0.947 4 4 2 1 8 5 2 42.0
Cladomics cladogram ANODONTINI LAMPSILINI QUADRULINI GONIDEINAE

Amaro et al., 2017 [Publ 3791], fig. 3B-M

cst iT tT bi riS oT roT roS iD
0.941 2 2 0 1 5 4 2 --
Cladomics cladogram QUADRULINI LAMPSILINI ANODONTINI GONIDEINAE

Combosch et al., 2017 [Publ 3806], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 212 2 2 --
Cladomics cladogram UNIONINI AMBLEMINAE PARREYSIINAE

Combosch et al., 2017 [Publ 3806], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 213 2 2 --
Cladomics cladogram UNIONINI PARREYSIINAE AMBLEMINAE

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 27 2 2 --
Cladomics cladogram GONIDEINAE AMBLEMINAE UNIONINI

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 27 2 2 --
Cladomics cladogram GONIDEINAE AMBLEMINAE UNIONINI

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

cst iT tT bi riS oT roT roS iD
0.704 20 20 0 1 85 84 2 58.0
Cladomics cladogram ANODONTINI GONIDEINAE AMBLEMINAE

Wu et al., 2020 [Publ 4147], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 1 9 6 2 43.0
Cladomics cladogram AMBLEMINAE GONIDEINAE UNIONINI

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

cst iT tT bi riS oT roT roS iD
0.962 1 1 0 1 18 16 2 --
Cladomics cladogram AMBLEMINAE LANCEOLARIINI GONIDEINAE

Wu et al., 2026 [Publ 4852], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 11 11 9 1 9 7 2 49.0
Cladomics cladogram PARREYSIINAE GONIDEINAE UNIONINI

Lydeard et al., 1996 [Publ 2656], fig. 3

cst iT tT bi riS oT roT roS iD
0.839 6 6 4 1 21 18 1 49.0
Cladomics cladogram ANODONTINI AMBLEMINAE

Lydeard et al., 1996 [Publ 2656], fig. 4

cst iT tT bi riS oT roT roS iD
0.848 6 6 4 1 22 18 1 49.5
Cladomics cladogram ANODONTINI AMBLEMINAE

Hoeh et al., 1996 [Publ 2694], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 2 1 1 --
Cladomics cladogram None 1 None 2 AMBLEMINAE ANODONTINI

Mulvey et al., 1997 [Publ 3390], fig. 1_3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 4 4 1 --
Cladomics cladogram ANODONTINI AMBLEMINAE

Hoeh et al., 1998 [Publ 2695], fig. 3

cst iT tT bi riS oT roT roS iD
0.875 1 1 0 1 4 1 1 --
Cladomics cladogram ANODONTINI MARGARITIFERINAE AMBLEMINAE

Hoeh et al., 1998 [Publ 2695], fig. 4

cst iT tT bi riS oT roT roS iD
0.875 1 1 0 1 4 1 1 --
Cladomics cladogram MARGARITIFERINAE ANODONTINI

Lydeard et al., 2000 [Publ 2654], fig. 2a

cst iT tT bi riS oT roT roS iD
0.889 1 1 0 1 13 12 1 --
Cladomics cladogram ANODONTINI MARGARITIFERINAE LAMPSILINI AMBLEMINI QUADRULINI PLEUROBEMINI

Lydeard et al., 2000 [Publ 2654], fig. 2b

cst iT tT bi riS oT roT roS iD
0.889 1 1 0 1 13 12 1 --
Cladomics cladogram ANODONTINI MARGARITIFERINAE AMBLEMINAE

Lydeard et al., 2000 [Publ 2654], fig. 2c

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 13 12 1 --
Cladomics cladogram ANODONTINI MARGARITIFERINAE QUADRULINI LAMPSILINI AMBLEMINI PLEUROBEMINI

Lydeard et al., 2000 [Publ 2654], fig. 2d

cst iT tT bi riS oT roT roS iD
0.889 1 1 0 1 13 12 1 --
Cladomics cladogram ANODONTINI MARGARITIFERINAE AMBLEMINAE

Turner et al., 2000 [Publ 2722], fig. 2

cst iT tT bi riS oT roT roS iD
0.882 1 1 0 1 13 12 1 --
Cladomics cladogram QUADRULINI ANODONTINI LAMPSILINI AMBLEMINI PLEUROBEMINI

Campbell, 2000 [Publ 2724], fig. 1-5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 2 1 1 --
Cladomics cladogram ANODONTINI AMBLEMINAE

Hoeh et al., 2002 [Publ 2673], fig. 1-F

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 7 6 1 --
Cladomics cladogram ANODONTINI AMBLEMINAE

Hoeh et al., 2002 [Publ 2673], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 7 6 1 --
Cladomics cladogram ANODONTINI AMBLEMINAE

Curole & Kocher, 2002 [Publ 2712], fig. 1a-M

cst iT tT bi riS oT roT roS iD
0.917 1 1 0 1 3 3 1 --
Cladomics cladogram QUADRULINI ANODONTINI LAMPSILINI

Curole & Kocher, 2002 [Publ 2712], fig. 1b-M

cst iT tT bi riS oT roT roS iD
0.917 1 1 0 1 3 3 1 --
Cladomics cladogram LAMPSILINI ANODONTINI QUADRULINI

Krebs et al., 2003 [Publ 2691], fig. 3

cst iT tT bi riS oT roT roS iD
0.789 10 10 5 1 23 21 1 51.5
Cladomics cladogram ANODONTINI AMBLEMINAE

Giribet & Distel, 2003 [Publ 3670], fig. 3f5

cst iT tT bi riS oT roT roS iD
0.900 1 1 0 1 5 2 1 --
Cladomics cladogram PLEUROBEMINI LAMPSILINI UNIONINI

Krebs, 2004 [Publ 2692], fig. 2

cst iT tT bi riS oT roT roS iD
0.929 2 2 0 1 3 3 1 --
Cladomics cladogram QUADRULINI ANODONTINI LAMPSILINI PLEUROBEMINI

Krebs, 2004 [Publ 2692], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 2 2 1 --
Cladomics cladogram ANODONTINI AMBLEMINAE

Curole & Kocher, 2005 [Publ 2682], fig. 1

cst iT tT bi riS oT roT roS iD
0.778 1 1 0 1 10 9 1 --
Cladomics cladogram ANODONTINI MARGARITIFERINAE

Curole & Kocher, 2005 [Publ 2682], fig. 1-M

cst iT tT bi riS oT roT roS iD
0.947 2 2 0 1 10 9 1 --
Cladomics cladogram ANODONTINI AMBLEMINAE

Soroka, 2005 [Publ 2726], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 1 1 1 44.0
Cladomics cladogram GONIDEINAE UNIONINI

Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 2 1 1 46.0
Cladomics cladogram GONIDEINAE ANODONTINI

Zanatta & Murphy, 2006 [Publ 2672], fig. 1

cst iT tT bi riS oT roT roS iD
0.923 1 1 0 1 55 54 1 --
Cladomics cladogram ANODONTINI AMBLEMINAE

Zanatta & Murphy, 2006 [Publ 2672], fig. 2

cst iT tT bi riS oT roT roS iD
0.875 1 1 0 1 56 55 1 --
Cladomics cladogram ANODONTINI MARGARITIFERINAE AMBLEMINAE

Cyr et al., 2007 [Publ 2745], fig. 2A

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 1 1 1 48.0
Cladomics cladogram AMBLEMINAE ANODONTINI

Soroka, 2008 [Publ 3695], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 1 1 1 43.0
Cladomics cladogram AMBLEMINAE ANODONTINI

Soroka, 2008 [Publ 3695], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 1 1 1 43.0
Cladomics cladogram AMBLEMINAE ANODONTINI

Boyer et al., 2011 [Publ 2718], fig. 3

cst iT tT bi riS oT roT roS iD
0.892 6 6 2 1 29 29 1 47.0
Cladomics cladogram ANODONTINI AMBLEMINAE

Boyer et al., 2011 [Publ 2718], fig. 4

cst iT tT bi riS oT roT roS iD
0.875 9 9 6 1 29 29 1 51.0
Cladomics cladogram ANODONTINI AMBLEMINAE

Moyer & Díaz-Ferguson, 2012 [Publ 3375], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 3 3 1 47.0
Cladomics cladogram ANODONTINI AMBLEMINAE

Peng et al., 2012 [Publ 3689], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 2 2 1 --
Cladomics cladogram ANODONTINI GONIDEINAE

Zanatta & Harris, 2013 [Publ 3530], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 4 4 1 --
Cladomics cladogram ANODONTINI AMBLEMINAE

Zhang et al., 2013 [Publ 4214], fig. 5

cst iT tT bi riS oT roT roS iD
0.556 2 4 0 1 2 1 1 47.0
Cladomics cladogram Cristaria Sinanodonta MARGARITIFERINAE GONIDEINAE

Graf et al., 2014 [Publ 3591], fig. 3

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 10 9 1 --
Cladomics cladogram MARGARITIFERINAE UNIONINI PARREYSIINAE

Inoue et al., 2014 [Publ 3635], fig. 3B

cst iT tT bi riS oT roT roS iD
0.900 7 7 2 1 1 1 1 50.0
Cladomics cladogram AMBLEMINAE ANODONTINI

Pfeiffer et al., 2016 [Publ 3660], fig. 2

cst iT tT bi riS oT roT roS iD
0.875 1 1 0 1 37 37 1 --
Cladomics cladogram ANODONTINI AMBLEMINAE

Xia et al., 2016 [Publ 3712], fig. 7B

cst iT tT bi riS oT roT roS iD
0.667 2 3 0 1 1 1 1 45.0
Cladomics cladogram Sinanodonta Cristaria GONIDEINAE

Wang et al., 2016 [Publ 3714], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 2 2 1 --
Cladomics cladogram UNIONINI GONIDEINAE

Wen et al., 2017 [Publ 3795], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 3 3 1 43.0
Cladomics cladogram ANODONTINI AMBLEMINAE

Hewitt et al., 2018 [Publ 3758], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 6 6 3 1 3 2 1 48.0
Cladomics cladogram AMBLEMINAE ANODONTINI

Vikhrev et al., 2018 [Publ 3812], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 12 1 1 --
Cladomics cladogram UNIONINI AMBLEMINAE

Pieri et al., 2018 [Publ 3867], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 19 1 --
Cladomics cladogram ANODONTINI AMBLEMINAE

Metzger et al., 2018 [Publ 3952], fig. 1C

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 2 2 1 --
Cladomics cladogram ANODONTINI AMBLEMINAE

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

cst iT tT bi riS oT roT roS iD
0.917 2 2 0 1 26 24 1 --
Cladomics cladogram ANODONTINI AMBLEMINAE

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

cst iT tT bi riS oT roT roS iD
0.913 2 2 0 1 27 25 1 --
Cladomics cladogram ANODONTINI AMBLEMINAE

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

cst iT tT bi riS oT roT roS iD
0.913 2 2 0 1 24 22 1 --
Cladomics cladogram ANODONTINI AMBLEMINAE

Hewitt et al., 2019 [Publ 3962], fig. 2

cst iT tT bi riS oT roT roS iD
0.909 1 1 0 1 9 8 1 --
Cladomics cladogram ANODONTINI AMBLEMINAE

Inoue et al., 2020 [Publ 3941], fig. 3

cst iT tT bi riS oT roT roS iD
0.964 1 1 0 1 41 41 1 --
Cladomics cladogram ANODONTINI AMBLEMINAE

Inoue et al., 2020 [Publ 3941], fig. 4

cst iT tT bi riS oT roT roS iD
0.929 1 1 0 1 40 40 1 --
Cladomics cladogram ANODONTINI AMBLEMINAE

Choi et al., 2020 [Publ 4169], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 1 3 3 1 49.0
Cladomics cladogram PARREYSIINAE UNIONINI

Stewart et al., 2021 [Publ 4451], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 10 10 9 1 6 3 1 50.0
Cladomics cladogram AMBLEMINAE ANODONTINI

Dai et al., 2024 [Publ 4385], fig. 3-BI

cst iT tT bi riS oT roT roS iD
1.000 27 27 24 1 4 2 1 63.0
Cladomics cladogram PARREYSIINAE ANODONTINI

Dai et al., 2024 [Publ 4385], fig. 3-ML

cst iT tT bi riS oT roT roS iD
1.000 27 27 25 1 4 2 1 63.0
Cladomics cladogram PARREYSIINAE ANODONTINI

Wu et al., 2024 [Publ 4388], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 1 4 4 1 47.0
Cladomics cladogram GONIDEINAE UNIONINI

Dai et al., 2024 [Publ 4389], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 43 41 1 43.0
Cladomics cladogram ANODONTINI GONIDEINAE

Hou et al., 2024 [Publ 4400], fig. 1

cst iT tT bi riS oT roT roS iD
0.833 13 13 11 1 2 2 1 49.0
Cladomics cladogram GONIDEINAE UNIONINI

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

cst iT tT bi riS oT roT roS iD
0.667 13 13 11 1 15 11 1 51.0
Cladomics cladogram ANODONTINI IRIDININAE ASPATHARIINAE

Xiang et al., 2025 [Publ 4433], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 27 27 25 1 1 1 1 61.0
Cladomics cladogram PARREYSIINAE ANODONTINI

Dai et al., 2025 [Publ 4441], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 44 42 1 43.0
Cladomics cladogram ANODONTINI GONIDEINAE

Chen et al., 2025 [Publ 4448], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 1 1 1 1 48.0
Cladomics cladogram GONIDEINAE UNIONINI

Chen et al., 2025 [Publ 4663], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 45 43 1 43.0
Cladomics cladogram ANODONTINI GONIDEINAE

Dai et al., 2026 [Publ 4757], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 30 30 29 1 4 2 1 66.0
Cladomics cladogram PARREYSIINAE ANODONTINI

Barraza et al., 2026 [Publ 4851], fig. 3

cst iT tT bi riS oT roT roS iD
0.864 1 1 0 1 18 12 1 --
Cladomics cladogram ANODONTINI AMBLEMINAE

Hoeh, 1990 [Publ 2717], fig. 2

cst iT tT bi riS oT roT roS iD
0.846 15 15 13 1 0 0 0 52.0
Cladomics cladogram ANODONTINI

Hoeh, 1990 [Publ 2717], fig. 3

cst iT tT bi riS oT roT roS iD
0.615 15 15 2 1 0 0 0 55.0
Cladomics cladogram ANODONTINI

Hoeh, 1990 [Publ 2717], fig. 4

cst iT tT bi riS oT roT roS iD
0.846 15 15 13 1 0 0 0 52.0
Cladomics cladogram ANODONTINI

Hoeh et al., 1995 [Publ 3391], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 7 7 4 1 0 0 0 47.0
Cladomics cladogram ALASMIDONTINA

Nagel et al., 1996 [Publ 3400], fig. 351

cst iT tT bi riS oT roT roS iD
0.750 5 5 4 1 0 0 0 47.0
Cladomics cladogram ANODONTINI

Nagel et al., 1998 [Publ 1946], fig. 7

cst iT tT bi riS oT roT roS iD
0.750 5 5 4 1 0 0 0 47.0
Cladomics cladogram ANODONTINI

Nagel et al., 1998 [Publ 1946], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 4 4 1 1 0 0 0 44.0
Cladomics cladogram Unio

King et al., 1999 [Publ 2699], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 9 9 7 1 0 0 0 49.0
Cladomics cladogram ALASMIDONTINA

King et al., 1999 [Publ 2699], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 6 6 3 1 0 0 0 48.0
Cladomics cladogram ALASMIDONTINA

Mock et al., 2004 [Publ 2686], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 0 0 0 --
Cladomics cladogram ANODONTINI

Mock et al., 2004 [Publ 2686], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 0 0 0 --
Cladomics cladogram ANODONTINI

Huff et al., 2004 [Publ 2697], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 0 0 --
Cladomics cladogram ANODONTINI MARGARITIFERINAE

Zanatta et al., 2007 [Publ 2698], fig. 1A

cst iT tT bi riS oT roT roS iD
0.846 16 16 10 1 0 0 0 55.0
Cladomics cladogram ANODONTINI

Zanatta et al., 2007 [Publ 2698], fig. 1B

cst iT tT bi riS oT roT roS iD
0.846 16 16 13 1 0 0 0 53.0
Cladomics cladogram ANODONTINI

Cyr et al., 2007 [Publ 2745], fig. 2B

cst iT tT bi riS oT roT roS iD
1.000 5 5 4 1 0 0 0 47.0
Cladomics cladogram ANODONTINI

Cyr et al., 2007 [Publ 2745], fig. 2C

cst iT tT bi riS oT roT roS iD
1.000 5 5 4 1 0 0 0 47.0
Cladomics cladogram ANODONTINI

Cyr et al., 2007 [Publ 2745], fig. 2D

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 0 0 0 48.0
Cladomics cladogram Pyganodon

Araujo et al., 2009 [Publ 2395], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 0 0 --
Cladomics cladogram UNIONINI MARGARITIFERINAE

Soroka, 2010 [Publ 3404], fig. 4-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 0 0 0 --
Cladomics cladogram ANODONTINI

Soroka, 2010 [Publ 3404], fig. 5a-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 0 0 0 --
Cladomics cladogram ANODONTINI

Soroka, 2010 [Publ 3404], fig. 5b-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 0 0 0 --
Cladomics cladogram ANODONTINI

Soroka, 2010 [Publ 3404], fig. 9a

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 1 0 0 0 48.0
Cladomics cladogram CRISTARIINA

Soroka, 2010 [Publ 3404], fig. 9b

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 0 0 0 48.0
Cladomics cladogram CRISTARIINA

Soroka, 2010 [Publ 3404], fig. 9c

cst iT tT bi riS oT roT roS iD
1.000 4 4 3 1 0 0 0 49.0
Cladomics cladogram CRISTARIINA

Prie et al., 2012 [Publ 3487], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 1 1 0 0 45.0
Cladomics cladogram MARGARITIFERINAE UNIONINI

Hinzmann et al., 2013 [Publ 3541], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 0 0 0 43.0
Cladomics cladogram ANODONTINA

Inoue et al., 2013 [Publ 3579], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 0 0 --
Cladomics cladogram ANODONTINI MARGARITIFERINAE

Froufe et al., 2014 [Publ 3592], fig. 2

cst iT tT bi riS oT roT roS iD
0.833 3 3 1 1 0 0 0 45.0
Cladomics cladogram ANODONTINA

Bieler et al., 2014 [Publ 3622], fig. 26

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram UNIONINI ASPATHARIINAE MARGARITIFERINAE

Bieler et al., 2014 [Publ 3622], fig. 27

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram UNIONINI ASPATHARIINAE MARGARITIFERINAE

Bieler et al., 2014 [Publ 3622], fig. 28

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram UNIONINI VELESUNIONINAE

Bieler et al., 2014 [Publ 3622], fig. 29

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram UNIONINI ASPATHARIINAE MARGARITIFERINAE

Bieler et al., 2014 [Publ 3622], fig. 30

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram UNIONINI ASPATHARIINAE MARGARITIFERINAE

Bieler et al., 2014 [Publ 3622], fig. 31

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram UNIONINI MARGARITIFERINAE ASPATHARIINAE

Klishko et al., 2014 [Publ 3666], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 0 0 0 45.0
Cladomics cladogram CRISTARIINA

Mezhzherin et al., 2014 [Publ 3705], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 0 0 0 42.0
Cladomics cladogram ANODONTINI

Mezhzherin et al., 2014 [Publ 3705], fig. 4

cst iT tT bi riS oT roT roS iD
0.875 4 4 2 1 0 0 0 44.0
Cladomics cladogram ANODONTINI

Graf et al., 2015 [Publ 3619], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 0 0 --
Cladomics cladogram UNIONINI MARGARITIFERINAE

González & Giribet, 2015 [Publ 3632], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram UNIONINI VELESUNIONINAE

González & Giribet, 2015 [Publ 3632], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram UNIONINI VELESUNIONINAE ASPATHARIINAE MARGARITIFERINAE

González & Giribet, 2015 [Publ 3632], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram UNIONINI MARGARITIFERINAE

González & Giribet, 2015 [Publ 3632], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram UNIONINI ASPATHARIINAE MARGARITIFERINAE

Bolotov et al., 2015 [Publ 3646], fig. 2A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 0 0 --
Cladomics cladogram UNIONINI MARGARITIFERINAE

Bolotov et al., 2015 [Publ 3646], fig. 2B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 0 0 --
Cladomics cladogram UNIONINI MARGARITIFERINAE

Bolotov et al., 2016 [Publ 3732], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 6 6 5 1 2 0 0 49.0
Cladomics cladogram ANODONTINI GIBBOSULINAE MARGARITIFERINAE

Klishko et al., 2016 [Publ 3733], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 0 0 0 45.0
Cladomics cladogram CRISTARIINA

Froufe et al., 2016 [Publ 3738], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 14 14 12 1 0 0 0 50.0
Cladomics cladogram UNIONINI

Lopes-Lima et al., 2016 [Publ 3757], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 0 0 0 43.0
Cladomics cladogram ANODONTINA

Sayenko et al., 2017 [Publ 3760], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 0 0 0 47.0
Cladomics cladogram CRISTARIINA

Froufe et al., 2017 [Publ 3802], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 0 0 0 42.0
Cladomics cladogram ANODONTINI

Froufe et al., 2017 [Publ 3802], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 12 12 9 1 0 0 0 47.0
Cladomics cladogram ANODONTINI

Vikhrev et al., 2017 [Publ 3810], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 1 2 0 0 48.0
Cladomics cladogram ANODONTINI GIBBOSULINAE MARGARITIFERINAE

Klishko et al., 2017 [Publ 3946], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 1 0 0 0 42.0
Cladomics cladogram UNIONINI

Smith et al., 2018 [Publ 3790], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 18 18 17 1 0 0 0 58.0
Cladomics cladogram ANODONTINI

Bespalaya et al., 2018 [Publ 3811], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 8 8 7 1 2 0 0 49.0
Cladomics cladogram ANODONTINI GIBBOSULINAE MARGARITIFERINAE

Burzynski & Soroka, 2018 [Publ 3862], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 0 0 0 42.0
Cladomics cladogram ANODONTINI

Kondakov et al., 2018 [Publ 3914], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 1 0 0 0 48.0
Cladomics cladogram CRISTARIINA

Klishko et al., 2018 [Publ 3945], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 8 8 5 1 0 0 0 44.0
Cladomics cladogram UNIONINI

Marrone et al., 2019 [Publ 3907], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 8 8 4 1 0 0 0 47.0
Cladomics cladogram Unio

Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 1 0 0 0 43.0
Cladomics cladogram ANODONTINI

Krebs et al., 2019 [Publ 4092], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 0 0 0 47.0
Cladomics cladogram Pyganodon

Lopes-Lima et al., 2020 [Publ 4080], fig. 10

cst iT tT bi riS oT roT roS iD
1.000 12 12 8 1 2 0 0 51.0
Cladomics cladogram MARGARITIFERINAE ANODONTINI

Lopes-Lima et al., 2020 [Publ 4080], fig. 12

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 1 2 0 0 48.0
Cladomics cladogram MARGARITIFERINAE UNIONINI

Lopes-Lima et al., 2020 [Publ 4080], fig. 13

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 1 2 0 0 50.0
Cladomics cladogram MARGARITIFERINAE UNIONINI

Lopes-Lima et al., 2020 [Publ 4080], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 1 2 0 0 43.0
Cladomics cladogram MARGARITIFERINAE UNIONINI

Lopes-Lima et al., 2020 [Publ 4080], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 9 9 7 1 2 0 0 51.0
Cladomics cladogram MARGARITIFERINAE ANODONTINI

Lopes-Lima et al., 2020 [Publ 4080], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 1 2 0 0 46.0
Cladomics cladogram MARGARITIFERINAE ANODONTINI

Lopes-Lima et al., 2020 [Publ 4080], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 1 2 0 0 45.0
Cladomics cladogram MARGARITIFERINAE ANODONTINI

Lopes-Lima et al., 2020 [Publ 4080], fig. 9

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 2 0 0 47.0
Cladomics cladogram MARGARITIFERINAE ANODONTINI

Riccardi et al., 2020 [Publ 4089], fig. 1

cst iT tT bi riS oT roT roS iD
0.889 6 6 4 1 0 0 0 42.0
Cladomics cladogram ANODONTINI

Choi et al., 2020 [Publ 4169], fig. 2a

cst iT tT bi riS oT roT roS iD
0.900 12 12 11 1 0 0 0 50.0
Cladomics cladogram UNIONINI

Choi et al., 2020 [Publ 4169], fig. 3a

cst iT tT bi riS oT roT roS iD
0.875 6 6 5 1 0 0 0 47.0
Cladomics cladogram UNIONINI

Kilikowska et al., 2020 [Publ 4335], fig. 4a

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 0 0 0 45.0
Cladomics cladogram Unio

Klunzinger et al., 2021 [Publ 4086], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram MARGARITIFERINAE UNIONINI

Sayenko et al., 2021 [Publ 4189], fig. 6

cst iT tT bi riS oT roT roS iD
0.900 13 13 11 1 0 0 0 51.0
Cladomics cladogram UNIONINI

Lopes-Lima et al., 2021 [Publ 4217], fig. 11

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 0 0 0 44.0
Cladomics cladogram Anodonta

Lopes-Lima et al., 2021 [Publ 4217], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 1 0 0 0 50.0
Cladomics cladogram Unio

Lopes-Lima et al., 2021 [Publ 4217], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 1 0 0 0 47.0
Cladomics cladogram Unio

Mabuchi et al., 2021 [Publ 4233], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 1 0 0 0 47.0
Cladomics cladogram OXYNAIINA

Bauer et al., 2021 [Publ 4365], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 8 8 5 1 0 0 0 48.0
Cladomics cladogram CRISTARIINA

Wu et al., 2022 [Publ 4238], fig. 2a_S1

cst iT tT bi riS oT roT roS iD
1.000 12 12 10 1 0 0 0 43.0
Cladomics cladogram UNIONINI

Bashê, 2022 [Publ 4265], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 5 5 4 1 1 0 0 48.0
Cladomics cladogram MARGARITIFERINAE UNIONINI

Sano et al., 2022 [Publ 4311], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 12 12 12 1 0 0 0 52.0
Cladomics cladogram CRISTARIINA

Sano et al., 2022 [Publ 4311], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 18 18 16 1 0 0 0 54.0
Cladomics cladogram CRISTARIINA

Whelan et al., 2023 [Publ 4287], fig. 2

cst iT tT bi riS oT roT roS iD
0.920 41 41 42 1 0 0 0 67.0
Cladomics cladogram ANODONTINI

Whelan et al., 2023 [Publ 4287], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 28 28 27 1 0 0 0 65.0
Cladomics cladogram ANODONTINI

Whelan et al., 2023 [Publ 4287], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 27 27 25 1 0 0 0 64.0
Cladomics cladogram ANODONTINI

Smith et al., 2023 [Publ 4291], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 1 0 0 0 52.0
Cladomics cladogram ALASMIDONTINA

Bogan et al., 2023 [Publ 4295], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 1 1 0 0 49.0
Cladomics cladogram GIBBOSULINAE ANODONTINI

Bogan et al., 2023 [Publ 4295], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 1 1 0 0 49.0
Cladomics cladogram GIBBOSULINAE ANODONTINI

Bogan et al., 2023 [Publ 4295], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 12 12 9 1 1 0 0 51.0
Cladomics cladogram GIBBOSULINAE ANODONTINI

Bulakhova et al., 2023 [Publ 4325], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 0 0 0 48.0
Cladomics cladogram Beringiana

Dai et al., 2024 [Publ 4358], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 1 0 0 0 46.0
Cladomics cladogram OXYNAIINA

Douda et al., 2024 [Publ 4364], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 13 13 12 1 1 0 0 52.0
Cladomics cladogram MARGARITIFERINAE ANODONTINI

Lopes-Lima et al., 2024 [Publ 4376], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 14 14 12 1 0 0 0 54.0
Cladomics cladogram Unio

Wu et al., 2024 [Publ 4379], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 13 13 10 1 0 0 0 52.0
Cladomics cladogram OXYNAIINA

Kashiri et al., 2024 [Publ 4822], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 1 2 0 0 48.0
Cladomics cladogram ANODONTINI GIBBOSULINAE MARGARITIFERINAE

Hou et al., 2025 [Publ 4401], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 11 11 9 1 0 0 0 53.0
Cladomics cladogram Nodularia

Hou et al., 2025 [Publ 4445], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 1 0 0 0 43.0
Cladomics cladogram OXYNAIINA

Perkins et al., 2025 [Publ 4447], fig. 2_mtDNA-ITS

cst iT tT bi riS oT roT roS iD
0.955 36 36 37 1 0 0 0 72.0
Cladomics cladogram ANODONTINI

Prie et al., 2026 [Publ 4818], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 11 11 9 1 0 0 0 55.0
Cladomics cladogram Unio

Chen et al., 2026 [Publ 4827], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 1 4 0 0 48.0
Cladomics cladogram GIBBOSULINAE UNIONINI

Itoh et al., 2026 [Publ 4842], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 1 0 0 0 48.0
Cladomics cladogram CRISTARIINA

Lawrie et al., 2026 [Publ 4843], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 0 0 --
Cladomics cladogram UNIONINI HYRIDELLINI VELESUNIONINAE CASTALIINI HYRIINI

Tumpeesuwan et al., 2026 [Publ 4845], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 1 0 0 0 52.0
Cladomics cladogram CRISTARIINA