Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae

tribe UNIONINI Rafinesque, 1820: cladomics

5 subtribes, 9 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 94 (159 total)
number of subtaxa (rank: subtribe) = 14
analyzed in 285 cladograms in 158 publications
maximum number of terminals in an analysis = 66 (mean = 7.89, median = 5)
maximum number of subtaxa (subtribe) = 5 (mean = 1.91, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 238
maximum number of included terminals = 66, (mean = 9.25 median = 7)
maximum number of subtaxa (subtribe) = 5 (mean = 2.08, median = 2)
proportion of cladograms that support basic monophyly = 0.84

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 131
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (subtribe) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 66 (mean = 12.53, median = 10)
maximum number of subtaxa (subtribe) = 5 (mean = 2.85, median = 3)
proportion of cladograms that support rigorous monophyly = 0.78

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Unionini cladogram distances
PCoA Ordination

NJ Tree of Unionini cladogram distances
NJ Tree

Cladograms

Wu et al., 2026 [Publ 4811], fig. 4

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
0.977 32 32 30 5 63 33 4 **
Cladomics cladogram LEPIDODESMINI ANODONTINI ACUTICOSTINI LANCEOLARIINI GLOBUNIONINA ACULAMPROTULINA UNIONINA MIDDENDORFFINAIINA OXYNAIINA

Dai et al., 2025 [Publ 4440], fig. 1

cst iT tT bi riS oT roT roS iD
0.895 21 21 16 5 23 14 4 33.0
Cladomics cladogram Pletholophus Sinanodonta Beringiana Cristaria ALASMIDONTINA ANODONTINA LANCEOLARIINI LEPIDODESMINI ACUTICOSTINI GLOBUNIONINA ACULAMPROTULINA MIDDENDORFFINAIINA Nodularia Cuneopsis Diaurora Schistodesmus Pseudocuneopsis UNIONINA Tchangsinaia Inversiunio

Dai et al., 2025 [Publ 4440], fig. 2

cst iT tT bi riS oT roT roS iD
0.868 21 21 18 5 23 14 4 32.0
Cladomics cladogram Anemina Amuranodonta Pletholophus Sinanodonta Beringiana ALASMIDONTINA ANODONTINA Lanceolaria gladiola Lanceolaria oxyrhyncha Cristaria ACUTICOSTINI LEPIDODESMINI GLOBUNIONINA ACULAMPROTULINA MIDDENDORFFINAIINA Nodularia Cuneopsis Diaurora Schistodesmus Pseudocuneopsis Inversiunio Tchangsinaia UNIONINA

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cst iT tT bi riS oT roT roS iD
0.947 46 46 44 5 94 22 4 34.0
Cladomics cladogram LEPIDODESMINI LANCEOLARIINI ACUTICOSTINI ANODONTINI ACULAMPROTULINA GLOBUNIONINA OXYNAIINA MIDDENDORFFINAIINA UNIONINA

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.818 66 74 77 5 567 90 4 70.0
Cladomics cladogram Amuranodonta Anemina Buldowskia Beringiana Pletholophus Mimo Acudonta Sinanodonta Cristaria LEPIDODESMINI LANCEOLARIINI Pseudoacuticosta retiaria 2 Acuticosta chinensis 2 Acuticosta chinensis 1 Pseudoacuticosta retiaria 1 Globunio ACULAMPROTULINA MIDDENDORFFINAIINA UNIONINA OXYNAIINA Pseudodontoideus Lasmigona Pterosyna Arcidens ANODONTINA Utterbackia Pyganodon Utterbackiana Simpsonaias Alasminota Alasmidonta Prolasmidonta Platynaias Anodontoides Strophitus

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
0.949 28 28 24 4 94 43 4 33.0
Cladomics cladogram ACUTICOSTINI LEPIDODESMINI LANCEOLARIINI ANODONTINI ACULAMPROTULINA MIDDENDORFFINAIINA UNIONINA OXYNAIINA

Konopleva et al., 2023 [Publ 4269], fig. S1_4

cst iT tT bi riS oT roT roS iD
0.966 12 12 10 4 44 21 4 28.0
Cladomics cladogram MIDDENDORFFINAIINA ACULAMPROTULINA UNIONINA OXYNAIINA LEPIDODESMINI LANCEOLARIINI ACUTICOSTINI ANODONTINI

Chen et al., 2023 [Publ 4302], fig. 1

cst iT tT bi riS oT roT roS iD
0.861 18 18 15 4 23 14 4 36.0
Cladomics cladogram Anemina Amuranodonta Pletholophus Sinanodonta Beringiana Lanceolaria gladiola ACUTICOSTINI LEPIDODESMINI Lanceolaria oxyrhyncha Cristaria ALASMIDONTINA ANODONTINA ACULAMPROTULINA MIDDENDORFFINAIINA Nodularia Cuneopsis Diaurora Schistodesmus Pseudocuneopsis Inversiunio Tchangsinaia UNIONINA

Chen et al., 2023 [Publ 4302], fig. 2

cst iT tT bi riS oT roT roS iD
0.972 18 18 15 4 23 14 4 36.0
Cladomics cladogram ACUTICOSTINI LEPIDODESMINI LANCEOLARIINI ANODONTINI ACULAMPROTULINA MIDDENDORFFINAIINA Nodularia Cuneopsis Diaurora Schistodesmus Pseudocuneopsis Inversiunio Tchangsinaia UNIONINA

Dai et al., 2024 [Publ 4386], fig. 5-BI

cst iT tT bi riS oT roT roS iD
0.917 12 14 10 4 10 5 4 36.0
Cladomics cladogram ACUTICOSTINI CRISTARIINA ANODONTINA LANCEOLARIINI ACULAMPROTULINA LEPIDODESMINI UNIONINA MIDDENDORFFINAIINA OXYNAIINA

Dai et al., 2024 [Publ 4386], fig. 5-ML

cst iT tT bi riS oT roT roS iD
0.958 12 12 9 4 10 5 4 33.0
Cladomics cladogram ANODONTINA ACUTICOSTINI CRISTARIINA LEPIDODESMINI LANCEOLARIINI ACULAMPROTULINA UNIONINA MIDDENDORFFINAIINA OXYNAIINA

Dai et al., 2025 [Publ 4440], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 27 27 25 4 60 15 4 21.0
Cladomics cladogram LEPIDODESMINI ANODONTINI ACUTICOSTINI LANCEOLARIINI GLOBUNIONINA ACULAMPROTULINA UNIONINA OXYNAIINA

Chen et al., 2025 [Publ 4487], fig. 2

cst iT tT bi riS oT roT roS iD
0.973 18 32 30 4 23 14 4 45.0
Cladomics cladogram PARREYSIINI COELATURINI INDOCHINELLINI GLOBUNIONINA ACULAMPROTULINA UNIONINA OXYNAIINA ACUTICOSTINI LEPIDODESMINI LANCEOLARIINI ANODONTINI

Chen et al., 2025 [Publ 4487], fig. 3

cst iT tT bi riS oT roT roS iD
0.946 18 18 16 4 23 14 4 31.0
Cladomics cladogram GLOBUNIONINA ACULAMPROTULINA UNIONINA OXYNAIINA ACUTICOSTINI ALASMIDONTINA ANODONTINA LEPIDODESMINI Cristaria LANCEOLARIINI Anemina Amuranodonta Pletholophus Sinanodonta Beringiana

Hou et al., 2024 [Publ 4400], fig. 4

cst iT tT bi riS oT roT roS iD
0.971 48 48 46 4 27 23 3 32.0
Cladomics cladogram LEPIDODESMINI LANCEOLARIINI ANODONTINI ACULAMPROTULINA UNIONINA MIDDENDORFFINAIINA OXYNAIINA

Wang et al., 2025 [Publ 4488], fig. 3B

cst iT tT bi riS oT roT roS iD
0.974 32 32 30 4 32 30 3 6.0
Cladomics cladogram LEPIDODESMINI ACUTICOSTINI ANODONTINI ACULAMPROTULINA UNIONINA MIDDENDORFFINAIINA OXYNAIINA

Wu et al., 2026 [Publ 4825], fig. 6B

cst iT tT bi riS oT roT roS iD
0.974 29 29 27 4 36 34 3 3.0
Cladomics cladogram LEPIDODESMINI LANCEOLARIINI ANODONTINI ACULAMPROTULINA UNIONINA MIDDENDORFFINAIINA OXYNAIINA

Dai et al., 2024 [Publ 4386], fig. 3

cst iT tT bi riS oT roT roS iD
0.941 11 13 11 4 3 2 2 36.0
Cladomics cladogram INDOCHINELLINI UNIONINA MIDDENDORFFINAIINA OXYNAIINA ACULAMPROTULINA ANODONTINI LEPIDODESMINI

Wu et al., 2018 [Publ 3836], fig. 2

cst iT tT bi riS oT roT roS iD
0.931 14 14 12 3 39 19 4 25.0
Cladomics cladogram LEPIDODESMINI ACULAMPROTULINA UNIONINA OXYNAIINA ANODONTINI ACUTICOSTINI LANCEOLARIINI

Aunins et al., 2018 [Publ 3863], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 11 11 9 3 29 15 4 27.0
Cladomics cladogram LEPIDODESMINI ACULAMPROTULINA OXYNAIINA UNIONINA ANODONTINI ACUTICOSTINI LANCEOLARIINI

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
0.912 21 21 17 3 180 27 4 27.0
Cladomics cladogram ACUTICOSTINI LEPIDODESMINI LANCEOLARIINI ANODONTINI ACULAMPROTULINA UNIONINA OXYNAIINA

Huang et al., 2019 [Publ 3843], fig. 3

cst iT tT bi riS oT roT roS iD
0.926 11 11 9 3 45 15 4 25.0
Cladomics cladogram LEPIDODESMINI ACULAMPROTULINA OXYNAIINA UNIONINA LANCEOLARIINI ACUTICOSTINI ANODONTINI

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
0.912 20 20 18 3 179 27 4 26.0
Cladomics cladogram ACUTICOSTINI ACULAMPROTULINA UNIONINA OXYNAIINA LEPIDODESMINI LANCEOLARIINI ANODONTINI

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.765 21 48 25 3 180 27 4 59.0
Cladomics cladogram Chamberlainia Megalonaias Uniomerus QUADRULINA Sinohyriopsis LAMPROTULINI GONIDEINI PSEUDODONTINI LIMNOSCAPHINI Amblema Toxolasma PLEUROBEMINI Obliquaria Plectomerus Ellipsaria Potamilus leptodon Medionidus Potamilus alatus Ptychobranchus Truncilla Venustaconcha Cambarunio Leaunio Ortmanniana Lampsilis ACUTICOSTINI Alasmidonta Anodonta anatina Anemina Anodonta cygnea Cristaria Brachyanodon Lasmigona LEPIDODESMINI Pseudanodonta Pyganodon Simpsonaias Strophitus Utterbackia Schistodesmus UNIONINA ACULAMPROTULINA LANCEOLARIINI Pletholophus Sinanodonta Cuneopsis celtiformis Tchangsinaia Cuneopsis rufescens Nodularia Cuneopsis szechenyii Cuneopsis heudei

Wu et al., 2019 [Publ 3864], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 13 13 11 3 33 13 4 25.0
Cladomics cladogram LEPIDODESMINI ANODONTINI ACUTICOSTINI LANCEOLARIINI ACULAMPROTULINA UNIONINA OXYNAIINA

Fukata & Iigo, 2020 [Publ 4106], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 13 13 11 3 41 15 4 29.0
Cladomics cladogram LEPIDODESMINI LANCEOLARIINI ACUTICOSTINI ANODONTINI ACULAMPROTULINA UNIONINA OXYNAIINA

Wu et al., 2020 [Publ 4121], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 13 13 11 3 35 13 4 25.0
Cladomics cladogram LEPIDODESMINI ANODONTINI ACUTICOSTINI LANCEOLARIINI ACULAMPROTULINA UNIONINA OXYNAIINA

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
0.931 11 11 9 3 80 15 4 25.0
Cladomics cladogram ACUTICOSTINI LANCEOLARIINI ANODONTINI LEPIDODESMINI ACULAMPROTULINA UNIONINA OXYNAIINA

Wu et al., 2021 [Publ 4366], fig. 3-BI

cst iT tT bi riS oT roT roS iD
0.935 15 15 13 3 58 18 4 23.0
Cladomics cladogram LEPIDODESMINI ACULAMPROTULINA UNIONINA OXYNAIINA ANODONTINI ACUTICOSTINI LANCEOLARIINI

Wu et al., 2021 [Publ 4366], fig. 3-ML

cst iT tT bi riS oT roT roS iD
0.935 15 15 13 3 58 18 4 25.0
Cladomics cladogram LEPIDODESMINI ACULAMPROTULINA UNIONINA OXYNAIINA ANODONTINI ACUTICOSTINI LANCEOLARIINI

Wu et al., 2021 [Publ 4366], fig. 4

cst iT tT bi riS oT roT roS iD
0.935 15 15 13 3 53 17 4 23.0
Cladomics cladogram LEPIDODESMINI ACULAMPROTULINA UNIONINA OXYNAIINA ANODONTINI ACUTICOSTINI LANCEOLARIINI

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
0.897 11 11 9 3 80 15 4 25.0
Cladomics cladogram ACUTICOSTINI ACULAMPROTULINA UNIONINA OXYNAIINA LANCEOLARIINI LEPIDODESMINI ANODONTINI

Wu et al., 2022 [Publ 4238], fig. 2d

cst iT tT bi riS oT roT roS iD
1.000 23 23 21 3 15 15 4 23.0
Cladomics cladogram LEPIDODESMINI LANCEOLARIINI ACUTICOSTINI ANODONTINI ACULAMPROTULINA UNIONINA OXYNAIINA

Wu et al., 2022 [Publ 4238], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 20 20 18 3 53 15 4 20.0
Cladomics cladogram LEPIDODESMINI ANODONTINI ACUTICOSTINI LANCEOLARIINI ACULAMPROTULINA UNIONINA OXYNAIINA

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 3 30 9 4 29.0
Cladomics cladogram LEPIDODESMINI ACULAMPROTULINA UNIONINA OXYNAIINA LANCEOLARIINI ACUTICOSTINI ANODONTINI

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

cst iT tT bi riS oT roT roS iD
0.960 7 8 6 3 32 10 4 30.0
Cladomics cladogram ANODONTINI ACUTICOSTINI LANCEOLARIINI LEPIDODESMINI ACULAMPROTULINA UNIONINA OXYNAIINA

Dai et al., 2023 [Publ 4285], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 3 46 10 4 27.0
Cladomics cladogram LEPIDODESMINI ACULAMPROTULINA UNIONINA OXYNAIINA ANODONTINI ACUTICOSTINI LANCEOLARIINI

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cst iT tT bi riS oT roT roS iD
1.000 22 22 20 3 73 24 4 24.0
Cladomics cladogram LEPIDODESMINI ANODONTINI ACUTICOSTINI LANCEOLARIINI ACULAMPROTULINA UNIONINA OXYNAIINA

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cst iT tT bi riS oT roT roS iD
0.914 22 22 19 3 73 24 4 26.0
Cladomics cladogram LEPIDODESMINI ANODONTINI ACUTICOSTINI LANCEOLARIINI ACULAMPROTULINA UNIONINA OXYNAIINA

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cst iT tT bi riS oT roT roS iD
0.971 22 22 19 3 69 24 4 21.0
Cladomics cladogram LEPIDODESMINI ANODONTINI ACUTICOSTINI LANCEOLARIINI ACULAMPROTULINA UNIONINA OXYNAIINA

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cst iT tT bi riS oT roT roS iD
0.971 22 22 19 3 69 24 4 21.0
Cladomics cladogram LEPIDODESMINI ANODONTINI ACUTICOSTINI LANCEOLARIINI ACULAMPROTULINA UNIONINA OXYNAIINA

Dai et al., 2024 [Publ 4358], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 18 18 16 3 35 10 4 22.0
Cladomics cladogram LEPIDODESMINI LANCEOLARIINI ACUTICOSTINI ANODONTINI ACULAMPROTULINA UNIONINA OXYNAIINA

Dai et al., 2024 [Publ 4386], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 21 21 19 3 20 6 4 23.0
Cladomics cladogram LEPIDODESMINI LANCEOLARIINI ACUTICOSTINI ANODONTINI ACULAMPROTULINA UNIONINA OXYNAIINA

Chen et al., 2025 [Publ 4448], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 27 27 25 3 58 16 4 21.0
Cladomics cladogram LEPIDODESMINI LANCEOLARIINI ACUTICOSTINI ANODONTINI ACULAMPROTULINA UNIONINA OXYNAIINA

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 3 13 13 3 28.0
Cladomics cladogram ANODONTINI LANCEOLARIINI LEPIDODESMINI ACULAMPROTULINA OXYNAIINA UNIONINA

Wang et al., 2018 [Publ 3865], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 3 24 10 3 29.0
Cladomics cladogram ACULAMPROTULINA OXYNAIINA UNIONINA ANODONTINI ACUTICOSTINI LANCEOLARIINI

Cha et al., 2018 [Publ 3916], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 3 28 11 3 28.5
Cladomics cladogram LEPIDODESMINI LANCEOLARIINI ANODONTINI ACULAMPROTULINA OXYNAIINA UNIONINA

Froufe et al., 2019 [Publ 3936], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 3 34 12 3 26.0
Cladomics cladogram LEPIDODESMINI LANCEOLARIINI ANODONTINI ACULAMPROTULINA OXYNAIINA UNIONINA

Froufe et al., 2019 [Publ 3936], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 3 34 12 3 26.0
Cladomics cladogram LEPIDODESMINI LANCEOLARIINI ANODONTINI ACULAMPROTULINA OXYNAIINA UNIONINA

Sano et al., 2020 [Publ 4138], fig. 2

cst iT tT bi riS oT roT roS iD
0.935 13 13 13 3 56 23 3 31.0
Cladomics cladogram Anemina Buldowskia Pletholophus Beringiana Sinanodonta Cristaria ANODONTINA ALASMIDONTINA LANCEOLARIINI LEPIDODESMINI ACULAMPROTULINA UNIONINA OXYNAIINA

Sano et al., 2020 [Publ 4138], fig. 3

cst iT tT bi riS oT roT roS iD
0.935 13 13 14 3 56 23 3 31.0
Cladomics cladogram Anemina Buldowskia Sinanodonta Pletholophus Beringiana Cristaria ANODONTINA ALASMIDONTINA LANCEOLARIINI LEPIDODESMINI ACULAMPROTULINA UNIONINA OXYNAIINA

Sano et al., 2020 [Publ 4138], fig. 4

cst iT tT bi riS oT roT roS iD
0.935 13 13 14 3 56 23 3 31.0
Cladomics cladogram Anemina Buldowskia Sinanodonta Pletholophus Beringiana Cristaria ANODONTINA ALASMIDONTINA LEPIDODESMINI LANCEOLARIINI ACULAMPROTULINA UNIONINA OXYNAIINA

Kim et al., 2021 [Publ 4197], fig. 1

cst iT tT bi riS oT roT roS iD
0.875 12 16 14 3 6 5 3 34.0
Cladomics cladogram CHAMBERLAINIINI LEPIDODESMINI ACULAMPROTULINA ACUTICOSTINI LANCEOLARIINI UNIONINA OXYNAIINA

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 3 38 12 3 26.0
Cladomics cladogram LEPIDODESMINI LANCEOLARIINI ANODONTINI ACULAMPROTULINA OXYNAIINA UNIONINA

Wu et al., 2023 [Publ 4296], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 13 13 11 3 10 8 3 29.0
Cladomics cladogram ACUTICOSTINI LANCEOLARIINI ANODONTINI UNIONINA ACULAMPROTULINA OXYNAIINA

Liu et al., 2023 [Publ 4310], fig. 4

cst iT tT bi riS oT roT roS iD
0.917 14 14 12 3 10 8 3 28.0
Cladomics cladogram ACUTICOSTINI Lanceolaria lanceolata Lanceolaria grayii Lanceolaria gladiola ANODONTINI UNIONINA ACULAMPROTULINA OXYNAIINA

Liu et al., 2023 [Publ 4310], fig. 5

cst iT tT bi riS oT roT roS iD
0.875 14 14 9 3 10 8 3 26.0
Cladomics cladogram ACUTICOSTINI Lanceolaria grayii Lanceolaria gladiola Lanceolaria lanceolata ANODONTINI UNIONINA ACULAMPROTULINA OXYNAIINA

Wu et al., 2024 [Publ 4379], fig. 5

cst iT tT bi riS oT roT roS iD
0.970 29 29 27 3 24 22 3 25.0
Cladomics cladogram LEPIDODESMINI LANCEOLARIINI ANODONTINI ACULAMPROTULINA UNIONINA OXYNAIINA

Wu et al., 2024 [Publ 4388], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 9 9 7 3 44 9 3 29.0
Cladomics cladogram LEPIDODESMINI LANCEOLARIINI ANODONTINI ACULAMPROTULINA UNIONINA OXYNAIINA

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 3 26 6 2 28.0
Cladomics cladogram ANODONTINI ACUTICOSTINI ACULAMPROTULINA UNIONINA OXYNAIINA

Xue et al., 2016 [Publ 3659], fig. 1

cst iT tT bi riS oT roT roS iD
0.889 4 4 2 3 11 7 2 31.0
Cladomics cladogram ANODONTINI LANCEOLARIINI ACULAMPROTULINA Schistodesmus Nodularia UNIONINA

Fonseca et al., 2016 [Publ 3775], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 3 24 11 2 30.0
Cladomics cladogram ANODONTINI LANCEOLARIINI ACULAMPROTULINA UNIONINA OXYNAIINA

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
0.966 11 11 8 3 70 18 2 31.0
Cladomics cladogram LANCEOLARIINI ANODONTINI UNIONINA ACULAMPROTULINA OXYNAIINA

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
0.929 10 10 8 3 129 18 2 32.0
Cladomics cladogram LANCEOLARIINI ANODONTINI UNIONINA ACULAMPROTULINA OXYNAIINA

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
0.821 10 10 6 3 129 18 2 34.0
Cladomics cladogram UNIONINA ACULAMPROTULINA Schistodesmus Nodularia Cuneopsis szechenyii Tchangsinaia Cuneopsis rufescens LANCEOLARIINI CRISTARIINA Anodonta anatina Pseudanodonta ALASMIDONTINA Anodonta cygnea Brachyanodon

Bolotov et al., 2017 [Publ 3764], fig. S1

cst iT tT bi riS oT roT roS iD
0.500 10 10 0 3 115 18 2 33.0
Cladomics cladogram Unio crassus Unio pictorum Unio tumidus ACULAMPROTULINA Cuneopsis szechenyii Tchangsinaia Cuneopsis rufescens Nodularia douglasiae Nodularia nuxpersicae Schistodesmus Alasmidonta Anodonta anatina Anodonta cygnea Brachyanodon Lasmigona Pseudanodonta Pyganodon Simpsonaias Strophitus Sinanodonta lauta Cristaria plicata Cristaria radiata Pletholophus Sinanodonta lucida Sinanodonta jourdyi Lanceolaria gladiola Lanceolaria grayii Lanceolaria danzhouensis

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 3 25 11 2 30.0
Cladomics cladogram ANODONTINI LANCEOLARIINI ACULAMPROTULINA UNIONINA OXYNAIINA

Wang et al., 2018 [Publ 3846], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 3 23 9 2 28.0
Cladomics cladogram ANODONTINI LANCEOLARIINI ACULAMPROTULINA OXYNAIINA UNIONINA

Huang et al., 2019 [Publ 3843], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 3 18 5 2 31.0
Cladomics cladogram ANODONTINI LANCEOLARIINI OXYNAIINA ACULAMPROTULINA UNIONINA

Fukata & Iigo, 2019 [Publ 4108], fig. 1

cst iT tT bi riS oT roT roS iD
0.938 13 17 15 3 4 4 2 35.0
Cladomics cladogram UNIONINA OXYNAIINA ACULAMPROTULINA ACUTICOSTINI LANCEOLARIINI

Guerra et al., 2019 [Publ 4168], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 3 36 8 2 29.0
Cladomics cladogram ANODONTINI LANCEOLARIINI ACULAMPROTULINA OXYNAIINA UNIONINA

Guerra et al., 2019 [Publ 4168], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 3 17 5 2 31.0
Cladomics cladogram ANODONTINI LANCEOLARIINI OXYNAIINA ACULAMPROTULINA UNIONINA

Guerra et al., 2019 [Publ 4168], fig. 2

cst iT tT bi riS oT roT roS iD
0.947 5 13 11 3 36 8 2 37.0
Cladomics cladogram GONIDEINI LAMPROTULINI ACULAMPROTULINA OXYNAIINA UNIONINA LANCEOLARIINI ANODONTINI

Guerra et al., 2019 [Publ 4168], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 3 17 5 2 29.0
Cladomics cladogram ANODONTINI LANCEOLARIINI ACULAMPROTULINA OXYNAIINA UNIONINA

Bolotov et al., 2020 [Publ 4084], fig. 1

cst iT tT bi riS oT roT roS iD
0.952 5 5 3 3 31 14 2 33.0
Cladomics cladogram LANCEOLARIINI ANODONTINI MIDDENDORFFINAIINA OXYNAIINA UNIONINA

Wu et al., 2022 [Publ 4238], fig. 2e

cst iT tT bi riS oT roT roS iD
1.000 13 13 11 3 5 5 2 23.0
Cladomics cladogram ANODONTINI LANCEOLARIINI ACULAMPROTULINA UNIONINA OXYNAIINA

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

cst iT tT bi riS oT roT roS iD
0.964 10 10 8 3 66 16 2 32.0
Cladomics cladogram LANCEOLARIINI ANODONTINI UNIONINA ACULAMPROTULINA OXYNAIINA

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

cst iT tT bi riS oT roT roS iD
0.929 10 10 8 3 66 16 2 32.0
Cladomics cladogram LANCEOLARIINI ANODONTINI UNIONINA ACULAMPROTULINA OXYNAIINA

Klymus et al., 2025 [Publ 4814], fig. 3A

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 3 26 5 2 31.0
Cladomics cladogram LANCEOLARIINI ANODONTINI ACULAMPROTULINA UNIONINA OXYNAIINA

Klymus et al., 2025 [Publ 4814], fig. 3B

cst iT tT bi riS oT roT roS iD
0.955 10 11 9 3 26 5 2 31.0
Cladomics cladogram ANODONTINI OXYNAIINA UNIONINA ACULAMPROTULINA LANCEOLARIINI

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 14 14 12 3 33 5 2 24.0
Cladomics cladogram ANODONTINI LANCEOLARIINI ACULAMPROTULINA UNIONINA OXYNAIINA

Wu et al., 2016 [Publ 3711], fig. 1

cst iT tT bi riS oT roT roS iD
0.944 5 5 3 3 16 7 1 28.0
Cladomics cladogram ANODONTINI ACULAMPROTULINA UNIONINA OXYNAIINA

Araujo et al., 2017 [Publ 3801], fig. 2

cst iT tT bi riS oT roT roS iD
0.909 3 4 2 3 11 1 1 33.0
Cladomics cladogram GONIDEINI LAMPROTULINI OXYNAIINA ACULAMPROTULINA ANODONTINI UNIONINA

Huang et al., 2018 [Publ 3828], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 9 9 7 3 33 9 1 29.0
Cladomics cladogram ANODONTINI ACULAMPROTULINA OXYNAIINA UNIONINA

Wu et al., 2018 [Publ 4143], fig. 2

cst iT tT bi riS oT roT roS iD
0.944 6 6 4 3 16 7 1 27.0
Cladomics cladogram ANODONTINI ACULAMPROTULINA UNIONINA OXYNAIINA

Klishko et al., 2019 [Publ 3886], fig. 9

cst iT tT bi riS oT roT roS iD
1.000 8 8 5 3 3 2 1 32.0
Cladomics cladogram ANODONTINI UNIONINA MIDDENDORFFINAIINA OXYNAIINA

Wu et al., 2020 [Publ 4147], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 11 11 9 3 19 7 1 22.0
Cladomics cladogram ANODONTINI ACULAMPROTULINA UNIONINA OXYNAIINA

Hou et al., 2025 [Publ 4445], fig. 5

cst iT tT bi riS oT roT roS iD
0.966 32 32 30 3 20 18 1 22.0
Cladomics cladogram ANODONTINI ACULAMPROTULINA UNIONINA OXYNAIINA

Sayenko et al., 2021 [Publ 4189], fig. 6

cst iT tT bi riS oT roT roS iD
0.889 13 13 11 3 0 0 0 39.0
Cladomics cladogram UNIONINA Inversiunio MIDDENDORFFINAIINA Nodularia

Huang et al., 2019 [Publ 3843], fig. 1

cst iT tT bi riS oT roT roS iD
0.800 17 24 22 2 29 15 4 32.0
Cladomics cladogram Anemina ACUTICOSTINI Sinanodonta ACULAMPROTULINA Cristaria LANCEOLARIINI LEPIDODESMINI OXYNAIINA

Huang et al., 2019 [Publ 3843], fig. S1

cst iT tT bi riS oT roT roS iD
0.900 17 17 15 2 30 16 4 25.0
Cladomics cladogram Anemina Sinanodonta ACUTICOSTINI LEPIDODESMINI Cristaria LANCEOLARIINI ACULAMPROTULINA OXYNAIINA

Sano et al., 2020 [Publ 4138], fig. 1

cst iT tT bi riS oT roT roS iD
0.565 7 22 5 2 27 15 4 49.0
Cladomics cladogram GONIDEINI LAMPROTULINI CHAMBERLAINIINI Lanceolaria oxyrhyncha 1 Lanceolaria oxyrhyncha 2 Cristaria plicata 1 Cristaria plicata 2 Sinanodonta lauta Beringiana japonica Beringiana fukuharai 1 Beringiana beringiana Beringiana fukuharai 2 Beringiana fukuharai 3 Buldowskia iwakawai 1 Buldowskia iwakawai 2 Sinanodonta tumens 1 Sinanodonta tumens 2 Pletholophus Anemina Lanceolaria lanceolata Cristaria plicata 3 Lanceolaria gladiola Sinanodonta angula LEPIDODESMINI ACULAMPROTULINA ACUTICOSTINI Cuneopsis Nodularia Schistodesmus Inversiunio

Zhou et al., 2007 [Publ 2658], fig. 1

cst iT tT bi riS oT roT roS iD
0.812 7 11 9 2 14 6 3 33.0
Cladomics cladogram CHAMBERLAINIINI Lamprotula leaii Lamprotula caveata Gibbosula Lamprotula cornuumlunae ACULAMPROTULINA Tchangsinaia Nodularia Schistodesmus LANCEOLARIINI ACUTICOSTINI

Ouyang et al., 2011 [Publ 3408], fig. 1

cst iT tT bi riS oT roT roS iD
0.800 3 6 3 2 10 6 3 32.0
Cladomics cladogram ANODONTINI OXYNAIINA Lanceolaria grayii ACUTICOSTINI Lanceolaria gladiola ACULAMPROTULINA

Ouyang et al., 2015 [Publ 4493], fig. 1

cst iT tT bi riS oT roT roS iD
0.882 8 8 6 2 23 8 3 34.0
Cladomics cladogram LANCEOLARIINI ACUTICOSTINI ANODONTINI Schistodesmus ACULAMPROTULINA Tchangsinaia Cuneopsis rufescens Nodularia Cuneopsis szechenyii Cuneopsis celtiformis

Ouyang et al., 2015 [Publ 4493], fig. 2

cst iT tT bi riS oT roT roS iD
0.882 8 14 19 2 23 8 3 41.0
Cladomics cladogram LANCEOLARIINI ACULAMPROTULINA ACUTICOSTINI ANODONTINI OXYNAIINA

Liu et al., 2017 [Publ 3939], fig. 2

cst iT tT bi riS oT roT roS iD
0.933 11 11 8 2 4 3 3 35.0
Cladomics cladogram LANCEOLARIINI UNIONINA OXYNAIINA

Wu et al., 2026 [Publ 4811], fig. 3

cst iT tT bi riS oT roT roS iD
0.933 4 10 8 2 13 6 3 37.0
Cladomics cladogram GONIDEINI UNIONINA LANCEOLARIINI GLOBUNIONINA ACUTICOSTINI ANODONTINI

Zhou et al., 2007 [Publ 2658], fig. 2

cst iT tT bi riS oT roT roS iD
0.692 5 6 4 2 13 3 2 28.0
Cladomics cladogram LAMPROTULINI QUADRULINI ACULAMPROTULINA ACUTICOSTINI OXYNAIINA

Zhou et al., 2016 [Publ 3710], fig. 1

cst iT tT bi riS oT roT roS iD
0.938 2 2 0 2 18 7 2 --
Cladomics cladogram ANODONTINI LEPIDODESMINI OXYNAIINA UNIONINA

Zhou et al., 2017 [Publ 3788], fig. 2

cst iT tT bi riS oT roT roS iD
0.917 4 4 2 2 12 2 2 28.0
Cladomics cladogram LANCEOLARIINI OXYNAIINA ACULAMPROTULINA

Wen et al., 2017 [Publ 3795], fig. 3

cst iT tT bi riS oT roT roS iD
0.929 3 4 2 2 13 4 2 31.0
Cladomics cladogram ANODONTINI ACULAMPROTULINA LEPIDODESMINI UNIONINA

Bolotov et al., 2018 [Publ 3826], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 5 5 4 2 77 12 2 34.0
Cladomics cladogram UNIONINA OXYNAIINA LANCEOLARIINI ANODONTINI

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 2 15 5 2 31.0
Cladomics cladogram ANODONTINI LANCEOLARIINI ACULAMPROTULINA UNIONINA

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 2 5 5 2 31.0
Cladomics cladogram LANCEOLARIINI ANODONTINI ACULAMPROTULINA UNIONINA

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 54 5 2 30.0
Cladomics cladogram LANCEOLARIINI ANODONTINI UNIONINA OXYNAIINA

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 54 5 2 30.0
Cladomics cladogram ANODONTINI LANCEOLARIINI UNIONINA OXYNAIINA

Chen et al., 2020 [Publ 4170], fig. 1

cst iT tT bi riS oT roT roS iD
0.714 3 4 2 2 31 12 2 30.0
Cladomics cladogram ANODONTINI OXYNAIINA LEPIDODESMINI ACULAMPROTULINA

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

cst iT tT bi riS oT roT roS iD
1.000 19 19 17 2 3 3 2 41.0
Cladomics cladogram LANCEOLARIINI ANODONTINI OXYNAIINA UNIONINA

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

cst iT tT bi riS oT roT roS iD
1.000 19 19 17 2 3 3 2 42.0
Cladomics cladogram LANCEOLARIINI ANODONTINI OXYNAIINA UNIONINA

Rosenberg et al., 1997 [Publ 2720], fig. 2B

cst iT tT bi riS oT roT roS iD
0.583 2 17 0 2 20 4 1 46.0
Cladomics cladogram ANODONTINA GONIDEINI Pyganodon grandis Utterbackia Pyganodon cataracta Lampsilis straminea Lampsilis teres Megalonaias Obliquaria Theliderma Quadrula Uniomerus Amblema Elliptio Fusconaia Plectomerus ACULAMPROTULINA UNIONINA

Araujo et al., 2009 [Publ 2589], fig. 7A

cst iT tT bi riS oT roT roS iD
0.667 7 8 4 2 4 1 1 36.0
Cladomics cladogram LAMPROTULINI OXYNAIINA Unio gibbus Unio tumidus ANODONTINI Unio pictorum 1 Unio pictorum 2 Unio mancus Unio tumidiformis Unio crassus

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

cst iT tT bi riS oT roT roS iD
0.571 3 9 7 2 22 6 1 39.0
Cladomics cladogram OXYNAIINA Anodonta cygnea Unio pictorum 2 Unio pictorum 1 Unio caffer Anodonta ALASMIDONTINA LAMPROTULINI QUADRULINI GONIDEINI Eurynia Cambarunio Ptychobranchus AMBLEMINI Pleurobema Truncilla Lampsilis Epioblasma Ligumia Obliquaria Ortmanniana

Guerra et al., 2017 [Publ 3772], fig. 2a

cst iT tT bi riS oT roT roS iD
0.938 2 2 0 2 23 8 1 --
Cladomics cladogram ANODONTINI ACULAMPROTULINA UNIONINA

Guerra et al., 2017 [Publ 3772], fig. 2b

cst iT tT bi riS oT roT roS iD
0.938 2 10 0 2 23 8 1 39.0
Cladomics cladogram CHAMBERLAINIINI LAMPROTULINI GONIDEINI UNIONINA ACULAMPROTULINA ANODONTINI

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 10 3 1 29.0
Cladomics cladogram ANODONTINI ACULAMPROTULINA UNIONINA

Burzynski et al., 2017 [Publ 3774], fig. 1-top

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 10 3 1 29.0
Cladomics cladogram ANODONTINI ACULAMPROTULINA UNIONINA

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 5 3 1 --
Cladomics cladogram ANODONTINI UNIONINA ACULAMPROTULINA

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 5 3 1 --
Cladomics cladogram ANODONTINI UNIONINA ACULAMPROTULINA

Araujo et al., 2018 [Publ 3779], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 14 14 10 2 4 3 1 40.0
Cladomics cladogram ANODONTINI OXYNAIINA UNIONINA

Huang et al., 2018 [Publ 3828], fig. 6-M

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 2 13 4 1 32.0
Cladomics cladogram ANODONTINI ACULAMPROTULINA UNIONINA

Sonowal et al., 2021 [Publ 4221], fig. 3a

cst iT tT bi riS oT roT roS iD
0.667 3 4 0 2 12 1 1 33.0
Cladomics cladogram PLEUROBEMINI LAMPSILINI ANODONTINI Unio ravoisieri Unio pictorum ACULAMPROTULINA

Sonowal et al., 2021 [Publ 4221], fig. 3b

cst iT tT bi riS oT roT roS iD
0.556 3 9 0 2 12 1 1 38.0
Cladomics cladogram HYRIINI Velesunio HYRIDELLINI ACULAMPROTULINA Lamprotula LAMPSILINI Potomida Unio pictorum ANODONTINI Unio ravoisieri LAMELLIDENTINI PLEUROBEMINI

Sonowal et al., 2021 [Publ 4221], fig. 4

cst iT tT bi riS oT roT roS iD
0.556 3 5 3 2 13 1 1 34.0
Cladomics cladogram PLEUROBEMINI ACULAMPROTULINA Unio pictorum ANODONTINI Unio ravoisieri LAMELLIDENTINI

Wu et al., 2024 [Publ 4379], fig. 4_6

cst iT tT bi riS oT roT roS iD
0.909 15 18 16 2 8 3 1 34.0
Cladomics cladogram LAMPROTULINI ACULAMPROTULINA ANODONTINI OXYNAIINA

Hou et al., 2025 [Publ 4401], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 26 26 24 2 2 2 1 24.0
Cladomics cladogram LEPIDODESMINI MIDDENDORFFINAIINA OXYNAIINA

Hou et al., 2025 [Publ 4432], fig. 3

cst iT tT bi riS oT roT roS iD
0.900 10 10 8 2 1 1 1 28.0
Cladomics cladogram LEPIDODESMINI Tchangsinaia Pseudocuneopsis MIDDENDORFFINAIINA Cuneopsis

Hou et al., 2025 [Publ 4432], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 26 26 24 2 2 2 1 24.0
Cladomics cladogram LEPIDODESMINI MIDDENDORFFINAIINA OXYNAIINA

Altaba et al., 2025 [Publ 4849], fig. 10

cst iT tT bi riS oT roT roS iD
0.867 4 4 2 2 8 8 1 32.0
Cladomics cladogram ANODONTINI OXYNAIINA UNIONINA

Altaba et al., 2025 [Publ 4849], fig. 11

cst iT tT bi riS oT roT roS iD
0.867 4 4 2 2 10 10 1 32.0
Cladomics cladogram ANODONTINI OXYNAIINA UNIONINA

Altaba et al., 2025 [Publ 4849], fig. 12

cst iT tT bi riS oT roT roS iD
0.714 4 4 2 2 9 9 1 32.0
Cladomics cladogram ANODONTINI Unio mancus Unio pictorum Unio tumidus OXYNAIINA

Wu et al., 2026 [Publ 4825], fig. 5

cst iT tT bi riS oT roT roS iD
0.889 7 17 15 2 12 10 1 37.0
Cladomics cladogram LAMPROTULINI OXYNAIINA ACULAMPROTULINA LANCEOLARIINI

Froufe et al., 2016 [Publ 3738], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 14 14 12 2 0 0 0 40.0
Cladomics cladogram OXYNAIINA UNIONINA

Klishko et al., 2017 [Publ 3946], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 2 0 0 0 32.0
Cladomics cladogram OXYNAIINA UNIONINA

Huang et al., 2018 [Publ 3828], fig. 3_5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 27 0 0 --
Cladomics cladogram LAMPSILINI INDOCHINELLINI UNIONINA ACULAMPROTULINA

Klishko et al., 2018 [Publ 3945], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 8 8 5 2 0 0 0 34.0
Cladomics cladogram OXYNAIINA UNIONINA

Lopes-Lima et al., 2020 [Publ 4080], fig. 13

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 2 2 0 0 34.0
Cladomics cladogram Margaritifera MIDDENDORFFINAIINA OXYNAIINA

Lopes-Lima et al., 2020 [Publ 4080], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 2 2 0 0 31.0
Cladomics cladogram Margaritifera OXYNAIINA ACULAMPROTULINA

Wu et al., 2020 [Publ 4121], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 2 29 0 0 30.0
Cladomics cladogram GONIDEINI LAMPROTULINI UNIONINA ACULAMPROTULINA

Wu et al., 2020 [Publ 4121], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 2 29 0 0 30.0
Cladomics cladogram GONIDEINI LAMPROTULINI UNIONINA ACULAMPROTULINA

Wu et al., 2020 [Publ 4147], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 2 9 0 0 30.0
Cladomics cladogram GONIDEINI LAMPROTULINI UNIONINA ACULAMPROTULINA

Choi et al., 2020 [Publ 4169], fig. 2a

cst iT tT bi riS oT roT roS iD
0.889 12 12 11 2 0 0 0 37.0
Cladomics cladogram Nodularia Schistodesmus Cuneopsis UNIONINA Tchangsinaia

Choi et al., 2020 [Publ 4169], fig. 3a

cst iT tT bi riS oT roT roS iD
0.857 6 6 5 2 0 0 0 36.0
Cladomics cladogram Nodularia Cuneopsis UNIONINA

Wu et al., 2022 [Publ 4238], fig. 2a_S1

cst iT tT bi riS oT roT roS iD
1.000 12 12 10 2 0 0 0 27.0
Cladomics cladogram ACULAMPROTULINA OXYNAIINA

Hou et al., 2024 [Publ 4400], fig. 1

cst iT tT bi riS oT roT roS iD
0.750 13 13 11 2 2 0 0 29.0
Cladomics cladogram LAMPROTULINI Cuneopsis rufescens MIDDENDORFFINAIINA Pseudocuneopsis Tchangsinaia Cuneopsis kiangsiensis Cuneopsis celtiformis Cuneopsis szechenyii Cuneopsis heudei Cuneopsis demangei

Chen et al., 2026 [Publ 4827], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 2 4 0 0 30.0
Cladomics cladogram Gibbosula GLOBUNIONINA ACULAMPROTULINA

Huang et al., 2002 [Publ 2657], fig. 1

cst iT tT bi riS oT roT roS iD
0.909 3 7 5 1 31 11 4 35.0
Cladomics cladogram ANODONTINI ACUTICOSTINI Nodularia Tchangsinaia LANCEOLARIINI Schistodesmus LEPIDODESMINI

Huang et al., 2002 [Publ 2657], fig. 2

cst iT tT bi riS oT roT roS iD
0.818 3 14 5 1 31 11 4 42.0
Cladomics cladogram Strophitus Schistodesmus LEPIDODESMINI ACUTICOSTINI CRISTARIINA LANCEOLARIINI Nodularia Tchangsinaia Utterbackia Utterbackiana Alasmidonta Lasmigona Ptychorhynchus CHAMBERLAINIINI LAMPROTULINI Sinosolenaia Megalonaias Elliptoideus Uniomerus Pleurobema Potamilus fragilis Medionidus Amblema Obliquaria Plectomerus Villosa Ellipsaria Lampsilis Potamilus purpuratus Ptychobranchus

Sano et al., 2017 [Publ 3799], fig. 2

cst iT tT bi riS oT roT roS iD
0.900 8 12 13 1 25 13 4 40.0
Cladomics cladogram ANODONTINI Tchangsinaia Nodularia Schistodesmus ACUTICOSTINI Inversiunio LEPIDODESMINI LANCEOLARIINI

Zhang et al., 2004 [Publ 2735], fig. 1

cst iT tT bi riS oT roT roS iD
0.938 7 7 5 1 7 5 3 33.0
Cladomics cladogram ANODONTINI OXYNAIINA ACUTICOSTINI LANCEOLARIINI

Plazzi et al., 2011 [Publ 3676], fig. 5

cst iT tT bi riS oT roT roS iD
0.786 3 3 1 1 15 6 2 33.0
Cladomics cladogram LANCEOLARIINI Anodonta UNIONINA ALASMIDONTINA Pseudanodonta Alathyria LAMPROTULINI CHAMBERLAINIINI QUADRULINI PLEUROBEMINI LAMPSILINI

Sano et al., 2017 [Publ 3799], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 4 4 5 1 17 9 2 35.0
Cladomics cladogram ANODONTINI LANCEOLARIINI OXYNAIINA

Kano et al., 2019 [Publ 4194], fig. 6

cst iT tT bi riS oT roT roS iD
0.917 2 2 0 1 8 4 2 --
Cladomics cladogram LANCEOLARIINI OXYNAIINA

Riccardi et al., 2020 [Publ 4089], fig. 2

cst iT tT bi riS oT roT roS iD
0.895 2 2 0 1 17 15 2 --
Cladomics cladogram ANODONTINI LANCEOLARIINI UNIONINA

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

cst iT tT bi riS oT roT roS iD
0.667 2 3 0 1 10 6 2 33.0
Cladomics cladogram ANODONTINI Nodularia LANCEOLARIINI Inversiunio

Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5_7

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 2 2 1 33.0
Cladomics cladogram ANODONTINI UNIONINA

Rosenberg et al., 1994 [Publ 2719], fig. 3

cst iT tT bi riS oT roT roS iD
0.500 1 1 0 1 18 3 1 --
Cladomics cladogram Elliptio Fusconaia AMBLEMINI Pleurobema UNIONINA

Rosenberg et al., 1997 [Publ 2720], fig. 2A

cst iT tT bi riS oT roT roS iD
0.600 1 1 0 1 21 4 1 --
Cladomics cladogram GONIDEINI Pyganodon grandis Utterbackia Pyganodon cataracta Lampsilis straminea Lampsilis teres Megalonaias Obliquaria Theliderma Quadrula Uniomerus Amblema Elliptio Fusconaia Plectomerus Pleurobema UNIONINA

Nagel et al., 1998 [Publ 1946], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 5 3 1 34.0
Cladomics cladogram ANODONTINI UNIONINA

Nagel et al., 1998 [Publ 1946], fig. 9

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 2 1 --
Cladomics cladogram UNIONINA ANODONTINI

Graf, 2000 [Publ 2647], fig. 1

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 16 1 1 --
Cladomics cladogram UNIONINA PLEUROBEMINI ANODONTINI LAMPSILINI

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

cst iT tT bi riS oT roT roS iD
0.917 2 2 0 1 28 6 1 --
Cladomics cladogram UNIONINA ANODONTINI QUADRULINI PLEUROBEMINI AMBLEMINI LAMPSILINI

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

cst iT tT bi riS oT roT roS iD
0.917 2 2 0 1 28 6 1 --
Cladomics cladogram UNIONINA ANODONTINI QUADRULINI PLEUROBEMINI AMBLEMINI LAMPSILINI

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 1 1 --
Cladomics cladogram UNIONINA ANODONTINI

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

cst iT tT bi riS oT roT roS iD
0.900 2 2 0 1 32 3 1 --
Cladomics cladogram GONIDEINI UNIONINA Cumberlandia Margaritifera Toxolasma AMBLEMINI Glebula Cyrtonaias QUADRULINI PLEUROBEMINI Potamilus Ortmanniana Ligumia

Hoeh et al., 2001 [Publ 2652], fig. 14f2

cst iT tT bi riS oT roT roS iD
0.800 2 2 0 1 29 3 1 --
Cladomics cladogram UNIONINA ANODONTINI Mutela Anodontites guanarensis Anodontites trigona Monocondylaea RHIPIDODONTINI Echyridella Lortiella CASTALIINI Velesunio

Hoeh et al., 2001 [Publ 2652], fig. 14f4

cst iT tT bi riS oT roT roS iD
0.700 2 5 0 1 29 3 1 35.0
Cladomics cladogram Unio pictorum Unio tumidus ANODONTINI AMBLEMINI LAMPSILINI QUADRULINI PLEUROBEMINI

Graf, 2002 [Publ 2651], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 16 2 1 --
Cladomics cladogram UNIONINA ANODONTINI

Hoeh et al., 2002 [Publ 2653], fig. 2

cst iT tT bi riS oT roT roS iD
0.700 2 5 0 1 29 3 1 35.0
Cladomics cladogram GONIDEINI Unio pictorum Unio tumidus ANODONTINI

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

cst iT tT bi riS oT roT roS iD
0.900 2 2 0 1 29 3 1 --
Cladomics cladogram UNIONINA ANODONTINI

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 29 3 1 --
Cladomics cladogram UNIONINA ANODONTINI

Huff et al., 2004 [Publ 2697], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 1 1 --
Cladomics cladogram UNIONINA ANODONTINI

Huff et al., 2004 [Publ 2697], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 1 1 --
Cladomics cladogram UNIONINA ANODONTINI

Huff et al., 2004 [Publ 2697], fig. 5-6

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 11 1 1 --
Cladomics cladogram UNIONINA ANODONTINI LAMPSILINI

Källersjö et al., 2005 [Publ 2684], fig. 1A

cst iT tT bi riS oT roT roS iD
0.750 3 3 0 1 4 3 1 33.0
Cladomics cladogram Margaritifera ANODONTINI UNIONINA

Källersjö et al., 2005 [Publ 2684], fig. 1B-F

cst iT tT bi riS oT roT roS iD
0.750 3 3 1 1 4 3 1 33.0
Cladomics cladogram Margaritifera ANODONTINI UNIONINA

Graf & Cummings, 2006 [Publ 1968], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 45 4 1 --
Cladomics cladogram UNIONINA ANODONTINI

Taylor et al., 2007 [Publ 2696], fig. 1-3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 1 1 --
Cladomics cladogram ANODONTINI UNIONINA

Chong et al., 2008 [Publ 2668], fig. 3

cst iT tT bi riS oT roT roS iD
0.762 1 1 0 1 17 17 1 --
Cladomics cladogram Anodonta cygnea Pyganodon Alasminota Anodonta anatina UNIONINA CRISTARIINA Brachyanodon nuttalliana Brachyanodon californiensis Pseudodontoideus Alasmidonta Pterosyna Pseudanodonta Strophitus Prolasmidonta Platynaias Brachyanodon oregonensis 4 Brachyanodon oregonensis 3 Brachyanodon oregonensis 2 Brachyanodon oregonensis 1 Brachyanodon kennerlyi

Manendo et al., 2008 [Publ 2669], fig. 1

cst iT tT bi riS oT roT roS iD
0.909 3 3 1 1 10 5 1 33.0
Cladomics cladogram UNIONINA ANODONTINI

Araujo et al., 2009 [Publ 2589], fig. 7B

cst iT tT bi riS oT roT roS iD
1.000 7 7 6 1 4 1 1 37.0
Cladomics cladogram ANODONTINI UNIONINA

Araujo et al., 2009 [Publ 2589], fig. 7C

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 1 4 1 1 36.0
Cladomics cladogram ANODONTINI UNIONINA

Hoeh et al., 2009 [Publ 2716], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 45 4 1 --
Cladomics cladogram UNIONINA ANODONTINI

Hoeh et al., 2009 [Publ 2716], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 44 4 1 --
Cladomics cladogram UNIONINA ANODONTINI

Hoeh et al., 2009 [Publ 2716], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 44 4 1 --
Cladomics cladogram UNIONINA ANODONTINI

Soroka, 2010 [Publ 3404], fig. 2

cst iT tT bi riS oT roT roS iD
0.800 3 3 0 1 4 4 1 31.0
Cladomics cladogram CRISTARIINA UNIONINA ANODONTINA

Soroka, 2010 [Publ 3404], fig. 2-M

cst iT tT bi riS oT roT roS iD
0.889 3 3 1 1 2 2 1 31.0
Cladomics cladogram CRISTARIINA ANODONTINA UNIONINA

Soroka, 2010 [Publ 3404], fig. 3a

cst iT tT bi riS oT roT roS iD
0.800 4 4 2 1 5 5 1 34.0
Cladomics cladogram CRISTARIINA ANODONTINA UNIONINA

Soroka, 2010 [Publ 3404], fig. 3a-M

cst iT tT bi riS oT roT roS iD
0.889 3 3 1 1 3 3 1 31.0
Cladomics cladogram ANODONTINA CRISTARIINA UNIONINA

Soroka, 2010 [Publ 3404], fig. 3b

cst iT tT bi riS oT roT roS iD
0.700 4 4 2 1 5 5 1 34.0
Cladomics cladogram Anodonta anatina Anodonta cygnea Pseudanodonta CRISTARIINA UNIONINA

Soroka, 2010 [Publ 3404], fig. 3b-M

cst iT tT bi riS oT roT roS iD
0.889 3 3 1 1 3 3 1 31.0
Cladomics cladogram ANODONTINA CRISTARIINA UNIONINA

Soroka, 2010 [Publ 3404], fig. 4

cst iT tT bi riS oT roT roS iD
0.700 3 3 1 1 4 4 1 31.0
Cladomics cladogram Anodonta anatina UNIONINA CRISTARIINA Anodonta cygnea Pseudanodonta

Soroka, 2010 [Publ 3404], fig. 5a

cst iT tT bi riS oT roT roS iD
0.800 3 3 1 1 5 5 1 31.0
Cladomics cladogram UNIONINA ANODONTINI

Soroka, 2010 [Publ 3404], fig. 5b

cst iT tT bi riS oT roT roS iD
0.700 3 3 1 1 5 5 1 31.0
Cladomics cladogram Anodonta anatina CRISTARIINA UNIONINA Anodonta cygnea Pseudanodonta

Soroka, 2010 [Publ 3404], fig. 8a

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 3 3 1 31.0
Cladomics cladogram UNIONINA ANODONTINI

Soroka, 2010 [Publ 3404], fig. 8a-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 1 1 1 --
Cladomics cladogram ANODONTINI UNIONINA

Soroka, 2010 [Publ 3404], fig. 8b

cst iT tT bi riS oT roT roS iD
0.889 3 3 1 1 3 3 1 31.0
Cladomics cladogram CRISTARIINA ANODONTINA UNIONINA

Soroka, 2010 [Publ 3404], fig. 8b-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 1 1 1 --
Cladomics cladogram ANODONTINI UNIONINA

Whelan et al., 2011 [Publ 3380], fig. 1-ML

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 34 2 1 --
Cladomics cladogram ANODONTINI UNIONINA

Whelan et al., 2011 [Publ 3380], fig. 1-MP

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 34 2 1 --
Cladomics cladogram ANODONTINI UNIONINA

Zieritz et al., 2012 [Publ 3673], fig. 4

cst iT tT bi riS oT roT roS iD
0.700 3 3 1 1 5 4 1 33.0
Cladomics cladogram ANODONTINI Margaritifera UNIONINA

Reis et al., 2013 [Publ 3580], fig. 4

cst iT tT bi riS oT roT roS iD
0.875 6 6 4 1 8 3 1 36.0
Cladomics cladogram ANODONTINI UNIONINA

Reis et al., 2013 [Publ 3580], fig. 5

cst iT tT bi riS oT roT roS iD
0.875 7 7 6 1 8 3 1 37.0
Cladomics cladogram ANODONTINI UNIONINA

Huang et al., 2013 [Publ 3629], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 15 6 1 --
Cladomics cladogram UNIONINA ANODONTINI

Prié & Puillandre, 2014 [Publ 3529], fig. 2

cst iT tT bi riS oT roT roS iD
0.875 11 11 9 1 6 3 1 39.0
Cladomics cladogram ANODONTINI UNIONINA

Marshall et al., 2014 [Publ 3593], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 1 1 --
Cladomics cladogram ANODONTINI UNIONINA

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 39 2 1 --
Cladomics cladogram UNIONINA ANODONTINI

Bolotov et al., 2016 [Publ 3743], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 28 1 1 --
Cladomics cladogram ANODONTINI UNIONINA

Bolotov et al., 2016 [Publ 3743], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 28 1 1 --
Cladomics cladogram ANODONTINI UNIONINA

Fonseca et al., 2016 [Publ 3775], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 3 1 --
Cladomics cladogram UNIONINA ANODONTINI

Feng et al., 2016 [Publ 3800], fig. 2

cst iT tT bi riS oT roT roS iD
0.769 1 1 0 1 13 6 1 --
Cladomics cladogram OXYNAIINA ANODONTINI

Lopes-Lima et al., 2017 [Publ 3734], fig. 1

cst iT tT bi riS oT roT roS iD
0.875 9 9 5 1 8 3 1 37.0
Cladomics cladogram ANODONTINI UNIONINA

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 3 1 --
Cladomics cladogram UNIONINA ANODONTINI

Froufe et al., 2017 [Publ 3802], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 7 4 1 33.0
Cladomics cladogram UNIONINA ANODONTINI

Chase et al., 2018 [Publ 3847], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 37 14 1 --
Cladomics cladogram UNIONINA ANODONTINI

Chase et al., 2018 [Publ 3847], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 8 2 1 --
Cladomics cladogram ANODONTINI UNIONINA

Bogatov et al., 2018 [Publ 3859], fig. 1

cst iT tT bi riS oT roT roS iD
0.625 3 5 3 1 3 2 1 35.0
Cladomics cladogram Margaritifera Unio tumidus ANODONTINI Unio crassus Unio pictorum

Bogatov et al., 2018 [Publ 3859], fig. 2

cst iT tT bi riS oT roT roS iD
0.625 3 5 3 1 3 2 1 35.0
Cladomics cladogram Margaritifera Unio crassus Unio pictorum Unio tumidus ANODONTINI

Chase et al., 2018 [Publ 3861], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 9 3 1 --
Cladomics cladogram UNIONINA ANODONTINI

Chase et al., 2018 [Publ 3861], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 9 3 1 --
Cladomics cladogram UNIONINA ANODONTINI

Froufe et al., 2019 [Publ 3936], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 16 4 1 31.0
Cladomics cladogram UNIONINA ANODONTINI

Froufe et al., 2019 [Publ 3936], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 16 4 1 31.0
Cladomics cladogram UNIONINA ANODONTINI

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 1 1 --
Cladomics cladogram UNIONINA ANODONTINI

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 1 1 --
Cladomics cladogram UNIONINA ANODONTINI

Lopes-Lima et al., 2020 [Publ 4080], fig. 11

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 10 8 1 --
Cladomics cladogram OXYNAIINA LANCEOLARIINI

Tomilova et al., 2020 [Publ 4097], fig. 3

cst iT tT bi riS oT roT roS iD
0.800 2 2 0 1 6 6 1 --
Cladomics cladogram UNIONINA CRISTARIINA ANODONTINA

Wu et al., 2021 [Publ 4215], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 21 1 1 33.0
Cladomics cladogram ACUTICOSTINI OXYNAIINA

Lopes-Lima et al., 2021 [Publ 4217], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 16 16 14 1 9 3 1 42.0
Cladomics cladogram ANODONTINI UNIONINA

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 20 4 1 31.0
Cladomics cladogram UNIONINA ANODONTINI

Lyubas et al., 2022 [Publ 4255], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 23 23 20 1 8 5 1 49.0
Cladomics cladogram ANODONTINI UNIONINA

Lyubas et al., 2022 [Publ 4255], fig. S1

cst iT tT bi riS oT roT roS iD
0.875 23 23 14 1 8 5 1 49.0
Cladomics cladogram ANODONTINI UNIONINA

Pfarrer et al., 2022 [Publ 4305], fig. 2-5

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 1 7 5 1 31.0
Cladomics cladogram UNIONINA ANODONTINI

Bogan et al., 2023 [Publ 4295], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 28 1 --
Cladomics cladogram OXYNAIINA ANODONTINI

Gomes-dos-Santos et al., 2023 [Publ 4463], fig. 2a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 20 1 1 --
Cladomics cladogram ANODONTINI OXYNAIINA

Gomes-dos-Santos et al., 2023 [Publ 4463], fig. 2b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 1 1 --
Cladomics cladogram ANODONTINI OXYNAIINA

Zieritz et al., 2024 [Publ 4381], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 44 1 1 --
Cladomics cladogram ANODONTINI UNIONINA

Wu et al., 2025 [Publ 4442], fig. 2

cst iT tT bi riS oT roT roS iD
0.929 2 2 0 1 33 33 1 --
Cladomics cladogram OXYNAIINA ANODONTINI

Wu et al., 2025 [Publ 4442], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 15 15 1 --
Cladomics cladogram UNIONINA ANODONTINI

Chen et al., 2026 [Publ 4805], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 33 1 --
Cladomics cladogram OXYNAIINA ANODONTINI

Wu et al., 2026 [Publ 4825], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 10 10 1 --
Cladomics cladogram OXYNAIINA LANCEOLARIINI

Nagel et al., 1998 [Publ 1946], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 4 4 1 1 0 0 0 34.0
Cladomics cladogram Unio

Giribet & Distel, 2003 [Publ 3670], fig. 3f5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram PLEUROBEMINI LAMPSILINI UNIONINA

Soroka, 2005 [Publ 2726], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 1 0 0 32.0
Cladomics cladogram LAMPROTULINI UNIONINA

Araujo et al., 2009 [Publ 2395], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 0 0 --
Cladomics cladogram UNIONINA Cumberlandia Pseudunio Margaritifera

Khalloufi et al., 2011 [Publ 2748], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 9 9 6 1 2 0 0 39.0
Cladomics cladogram LAMPROTULINI UNIONINA

Prie et al., 2012 [Publ 3487], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 1 1 0 0 33.0
Cladomics cladogram Pseudunio UNIONINA

Graf et al., 2014 [Publ 3591], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 0 0 --
Cladomics cladogram Margaritifera UNIONINA LEOPARREYSIINI COELATURINI

Bieler et al., 2014 [Publ 3622], fig. 26

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram UNIONINA Aspatharia Margaritifera

Bieler et al., 2014 [Publ 3622], fig. 27

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram UNIONINA Aspatharia Margaritifera

Bieler et al., 2014 [Publ 3622], fig. 28

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram UNIONINA Velesunio

Bieler et al., 2014 [Publ 3622], fig. 29

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram UNIONINA Aspatharia Margaritifera

Bieler et al., 2014 [Publ 3622], fig. 30

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram UNIONINA Aspatharia Margaritifera

Bieler et al., 2014 [Publ 3622], fig. 31

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram UNIONINA Margaritifera Aspatharia

Graf et al., 2015 [Publ 3619], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 0 0 --
Cladomics cladogram UNIONINA Margaritifera

González & Giribet, 2015 [Publ 3632], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram UNIONINA Velesunio

González & Giribet, 2015 [Publ 3632], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram UNIONINA Velesunio Aspatharia Margaritifera

González & Giribet, 2015 [Publ 3632], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram UNIONINA Margaritifera

González & Giribet, 2015 [Publ 3632], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram UNIONINA Aspatharia Margaritifera

Bolotov et al., 2015 [Publ 3646], fig. 2A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 0 0 --
Cladomics cladogram UNIONINA Margaritifera margaritifera Margaritifera dahurica 1 Margaritifera dahurica 2 Pseudunio Cumberlandia Margaritifera falcata Margaritifera laevis Margaritifera middendorffi Margaritifera marrianae

Bolotov et al., 2015 [Publ 3646], fig. 2B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 0 0 --
Cladomics cladogram UNIONINA Cumberlandia Pseudunio Margaritifera

Wang et al., 2016 [Publ 3714], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 2 0 0 --
Cladomics cladogram UNIONINA CHAMBERLAINIINI

Combosch et al., 2017 [Publ 3806], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 212 0 0 --
Cladomics cladogram UNIONINA LAMPSILINI INDOCHINELLINI

Combosch et al., 2017 [Publ 3806], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 213 0 0 --
Cladomics cladogram UNIONINA INDOCHINELLINI LAMPSILINI

Vikhrev et al., 2018 [Publ 3812], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 12 0 0 --
Cladomics cladogram UNIONINA LAMPSILINI

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 27 0 0 --
Cladomics cladogram LAMPSILINI UNIONINA

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 27 0 0 --
Cladomics cladogram LAMPSILINI UNIONINA

Marrone et al., 2019 [Publ 3907], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 8 8 4 1 0 0 0 36.0
Cladomics cladogram Unio

Lopes-Lima et al., 2020 [Publ 4080], fig. 12

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 1 2 0 0 36.0
Cladomics cladogram Margaritifera OXYNAIINA

Choi et al., 2020 [Publ 4169], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 1 3 0 0 35.0
Cladomics cladogram OXYNAIINA INDOCHINELLINI LAMELLIDENTINI LEOPARREYSIINI

Kilikowska et al., 2020 [Publ 4335], fig. 4a

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 0 0 0 31.0
Cladomics cladogram Unio

Klunzinger et al., 2021 [Publ 4086], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram Margaritifera UNIONINA

Lopes-Lima et al., 2021 [Publ 4217], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 1 0 0 0 38.0
Cladomics cladogram Unio

Lopes-Lima et al., 2021 [Publ 4217], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 1 0 0 0 35.0
Cladomics cladogram Unio

Mabuchi et al., 2021 [Publ 4233], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 1 0 0 0 33.0
Cladomics cladogram OXYNAIINA

Bashê, 2022 [Publ 4265], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 5 5 4 1 1 0 0 35.0
Cladomics cladogram Pseudunio UNIONINA

Dai et al., 2024 [Publ 4358], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 1 0 0 0 28.0
Cladomics cladogram OXYNAIINA

Lopes-Lima et al., 2024 [Publ 4376], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 14 14 12 1 0 0 0 42.0
Cladomics cladogram Unio

Wu et al., 2024 [Publ 4379], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 13 13 10 1 0 0 0 36.0
Cladomics cladogram OXYNAIINA

Wu et al., 2024 [Publ 4388], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 1 4 0 0 35.0
Cladomics cladogram GONIDEINI OXYNAIINA

Hou et al., 2025 [Publ 4401], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 11 11 9 1 0 0 0 37.0
Cladomics cladogram Nodularia

Hou et al., 2025 [Publ 4445], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 1 0 0 0 27.0
Cladomics cladogram OXYNAIINA

Chen et al., 2025 [Publ 4448], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 1 1 0 0 30.0
Cladomics cladogram GONIDEINI OXYNAIINA

Prie et al., 2026 [Publ 4818], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 11 11 9 1 0 0 0 39.0
Cladomics cladogram Unio

Lawrie et al., 2026 [Publ 4843], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 0 0 --
Cladomics cladogram UNIONINA HYRIDELLINI CASTALIINI HYRIINI Westralunio Velesunio 6 Alathyria jacksoni 2 Alathyria profuga Velesunio ambiguus Velesunio 5 Alathyria pertexta 2 Velesunio 4 Alathyria jacksoni 1 Alathyria pertexta 1 Lortiella Velesunio 3 Microdontia Velesunio angasi Velesunio 1 Velesunio 2

Wu et al., 2026 [Publ 4852], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 11 11 9 1 9 0 0 35.0
Cladomics cladogram OXYNAIINA GONIDEINI LAMPROTULINI