MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Sun 12 April 2026
superfamily UNIONOIDEA Rafinesque, 1820: cladomics
Cladomics pages are experimental — even more so than the rest of this site!
The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.
The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.
Following the summary, the available published clades representing the taxon are listed.
For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!
Cladomics Summary
number of species = 822 (1123 total)
number of subtaxa (rank: family) = 2
analyzed in 588 cladograms in 326 publications
maximum number of terminals in an analysis = 581 (mean = 23.83, median = 14)
maximum number of subtaxa (family) = 2 (mean = 1.46, median = 1)
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 586
maximum number of included terminals = 581, (mean = 23.91 median = 14)
maximum number of subtaxa (family) = 2 (mean = 1.46, median = 1)
proportion of cladograms that support basic monophyly = 0.94
Rigorous Monophyly
number of cladograms that test rigorous monophyly = 94
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (family) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 581 (mean = 43.86, median = 23)
maximum number of subtaxa (family) = 2 (mean = 2, median = 2)
proportion of cladograms that support rigorous monophyly = 0.69
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Cladograms
Graf, 2000 [Publ 2647], fig. 1
** Top Monophyly Cladogram **
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 16 | 14 | 2 | 10 | 9 | 2 | 0 |
+-------------- Margaritiferidae ** | | +-------- Unionini ** --+ +--+ | | | +----- Pleurobemini ** | | +--+ | | | +-- Anodontini ** +--+ +--+ | +-- Lampsilini ** | | +-------- Parreysiini ** +--+ | +----- Coelaturini ** +--+ | +-- Hyrioidea +--+ +-- Etherioidea
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14
** Top Sister-Group Cladogram **
| tT | bi | riS | oT | roT | roS | iD | |
| 581 | 582 | 651 | 2 | 51 | 49 | 2 | 701.5 |
+----------------------------- Iridininae
|
+-----------------------------------------------------------------+ +-------------------------- Aspathariinae
| | |
| +--+ +----------------------- Etheria
| | |
| +--+ +-------------------- Anodontites_trapesialis
| | |
| +--+ +----------------- Mulleria
| | |
| +--+ +-------------- Lamproscapha
| | |
| +--+ +-- Monocondylaea
| | +--------+
| | | +-- Tamsiella
| +--+
| | +-------- Mycetopodinae
| | |
| +--+ +-- Anodontites_guanarensis
| | +--+
| | | +-- Fossula
| +--+
| | +-- Anodontites_trigona
| +--+
| +-- Anodontites_elongata
|
| +-------------------------------------------------------------------------------------------- Hyrioidea
| |
| | +-------------------------------- Lamellidens **
| | |
| | +-----+ +-- Balwantia **
| | | | +--------------------------+
--+ | | | | +-- Trapezidens **
| | | +--+
| | | | +-------------------------- Leoparreysiini **
| | | | |
| | | +--+ +----------------------- Pseudomulleria
| | | | |
| | | | | +-- Parreysiini **
| | | +--+ +-----------------+
| | | | | +-- Coelaturini **
| | | | |
| | | +--+ +-- Pseudoscabies **
| | | | +--------------+
| | | | | +-- Indonaia **
| | | +--+
| | | | +-------------- Indochinella **
| | | | |
| | | +--+ +----------- Radiatula **
| | +--------------------------------------------------+ | |
| | | | +--+ +-- Songkhlanaia_songkhramensis__2 **
| | | | | +-----+
| | | | | | +-- Unionetta_fabagina__2 **
| | | | +--+
| | | | | +----- Harmandia **
| | | | +--+
| | | | | +-- Unionetta_fabagina__1 **
| | | | +--+
| | | | +-- Scabies **
| | | |
| | | | +----- Amuranodonta **
| | | | +-----------------------------+
| | | | | | +-- Anemina **
+--+ | | | +--+
| | | | +-- Buldowskia **
| | | |
| | | | +-- Beringiana **
| | | | +-----+
| | | | | +-- Pletholophus **
| | +--+ +-----------------------+
| | | | | +----- Mimo **
| | | | +--+
| | | | | +-- Acudonta **
| | | | +--+
| | | | +-- Sinanodonta **
| | | |
| | | | +----------------------------- Cristaria **
| | +--+ |
| | | | +----------------- Lepidodesmini **
| | | | |
| | | | | +----------- Lanceolariini **
| | | | | +--+
| | | | +--------+ | | +-------- Pseudoacuticosta_retiaria__2 **
| | | | | | | +--+
| | | | | | | | +----- Acuticosta_chinensis__2 **
| | +--+ | | | +--+
| | | | +--+ | +-- Acuticosta_chinensis__1 **
| | | | | +--+
| | | | | +-- Pseudoacuticosta_retiaria__1 **
| | | | |
| | | | | +----------- Globunio **
| | | | +--+
| | | | | +-------- Aculamprotulina **
| | | | +--+
| | +--+ | +----- Middendorffinaiina **
| | | +--+
| | | | +-- Unionina **
| | | +--+
| | | +-- Oxynaiina **
| | |
| | | +-------- Pseudodontoideus **
+--+ | +--------------+
| | | | +----- Lasmigona **
| | | +--+
| | | | +-- Pterosyna **
| | | +--+
| +--+ +-- Arcidens **
| |
| | +-------------------- Anodontina **
| | |
| | | +-------------- Utterbackia **
| | | +--+
| +--+ | | +----------- Pyganodon_grandis **
| | | +--+
| | | | +-------- Pyganodon_lacustris **
| | | +--+
| | | | +----- Pyganodon_cataracta__2 **
| +--+ +--+
| | | +-- Pyganodon_cataracta__1 **
| | +--+
| | +-- Potamilus_fragilis__3 **
| |
| | +-------------- Utterbackiana **
| | |
| +--+ +----------- Simpsonaias **
| | |
| | | +-- Alasminota **
| +--+ +-----+
| | | +-- Alasmidonta **
| | |
| +--+ +-- Prolasmidonta **
| | +--+
| | | +-- Platynaias **
| +--+
| | +-- Anodontoides **
| +--+
| +-- Strophitus **
|
| +-------------------------------------------------------------------------------------- Rectidentini **
| |
| | +----------------------------------------------------------------------------------- Margaritiferidae **
| | |
| | | +-------------------- Chamberlainiini **
| | | |
| | | +--+ +----------------- Parvasolenaia **
| | | | | |
| | | | +--+ +-------------- Pseudopostolata **
| | | | | |
| | | | +--+ +----------- Koreosolenaia **
| | | | | |
| | | | +--+ +-- Cosmopseudodon **
| | | | | +-----+
| | | | | | +-- Guiunio **
+--+ | | +--+
| | +--------+ | +----- Ptychorhynchus **
| | | | +--+
| | | | | +-- Obovalis **
| | | | +--+
| | | | +-- Postolata **
| | | |
| | | | +-- Caudiculatus **
| | | | +--------------+
| | | | | +-- Lamprotulini **
| | | | |
| | | +-----+ +-------------- Sinosolenaia_carinata **
| | | | |
| | | | | +----- Gonidea **
| | | +--+ +-----+
+--+ +-----------------------------------------------+ | | | +-- Microcondylaea **
| | | | | +--+
| | | +--+ +-- Leguminaia **
| | | |
| | | | +-------- Inversidens **
| | | +--+
| | | | +----- Sinosolenaia_recognita **
| | | +--+
| | | | +-- Sinosolenaia_iridinea **
| | | +--+
| | | +-- Sinosolenaia_oleivora **
| | |
| | | +----------------------------- Indopseudodontina **
| | | |
| | | | +-- Lannanaia **
| | +--+ +-----------------------+
| | | | +-- Pilsbryoconcha **
| | | |
| | | | +----- Songkhlanaia_tamodienica **
| | +--+ +-----------------+
| | | | | +-- Songkhlanaia_moreleti **
| | | | +--+
| | | | +-- Songkhlanaia_songkhramensis__1 **
| | | |
| | +--+ +-- Nyeinchanconcha **
| | | +-----------------+
| | | | +-- Sundadontina **
| | | |
+--+ | | +-- Isannaia **
| +--+ +--+
| | | +-- Namkongnaia **
| | +-----------+
| | | | +-- Thaiconcha **
| | | +--+
| +--+ +-- Bineurus **
| |
| | +-------------- Pseudodon_walpolei **
| | |
| +--+ +----------- Pseudodon_vagulus **
| | |
| | | +----- Pseudodon_vondembuschianus **
| +--+ +--+
| | | | +-- Pseudodon_mekongi **
| | | +--+
| +--+ +-- Schepmaniini **
| |
| | +----- Pseudodon_lenyanensis__2 **
| +--+
| | +-- Pseudodon_inoscularis **
| +--+
| +-- Pseudodon_lenyanensis__1 **
|
| +-- Quadrulini **
| +--------------------------------------------------------------------------+
| | +-- Pleurobemini **
| |
| | +----- Amblema **
+--+ +--------------------------------------------------------------------+
| | | +-- Nephronaias **
| | +--+
| | +-- Psoronaias **
| |
+--+ +----------------------------------------------------------------------- Reginaia **
| |
| | +----- Popenaias **
| | +--------------------------------------------------------------+
| | | | +-- Cyrtonaias **
+--+ | +--+
| | +-- Glebula **
| |
| | +----------------------------------------------------------------- Paetulunio **
| | |
+--+ | +----------------- Toxolasma_parvum **
| | |
| | | +----- Toxolasma_lividum__2 **
| | | +--------+
| | +--------------------------------------------+ | | +-- Toxolasma_cylindrellus **
| | | | | +--+
| | | | | +-- Toxolasma_lividum__1 **
| | | | |
+--+ | +--+ +-------- Toxolasma_texasiense__2 **
| | | +--+
| | | | | +----- Toxolasma_glans **
| | | | +--+
| | | | | +-- Toxolasma_paulum **
| | +--+ +--+
| | | +-- Toxolasma_texasiense__1 **
| | |
| | | +-------- Toxolasma__4 **
| | +--+
+--+ | +----- Toxolasma_granulatum **
| +--+
| | +-- Toxolasma__2 **
| +--+
| +-- Toxolasma__3 **
|
| +----- Toxolasma_pullus **
| +-----------------------------------------------------+
| | | +-- Toxolasma_germanum **
| | +--+
| | +-- Toxolasma__1 **
| |
| | +-- Obliquaria **
| | +-----------------------------------------------------+
| | | +-- Plectomerus **
+--+ |
| | +-------- Potamilus_inflatus **
| | +--------+
| | | | +----- Potamilus_ohiensis **
| | | +--+
| | | | +-- Potamilus_amphichaenus **
| | | +--+
| | +-----------------------------------+ +-- Potamilus_streckersoni **
| | | |
+--+ | | +----- Potamilus_fragilis__2 **
| | | +--------+
| | | | | +-- Potamilus_leptodon **
| | | | +--+
| | +--+ +-- Potamilus_fragilis__1 **
| | |
| | | +----------- Potamilus_metnecktayi **
| | | |
| | +--+ +-------- Potamilus_alatus__3 **
| | | |
| | +--+ +----- Potamilus_alatus__2 **
+--+ | |
| +--+----- Potamilus_purpuratus__2 **
| |
| | +-- Potamilus_alatus__1 **
| +--+
| +-- Potamilus_purpuratus__1 **
|
| +-- Villosa_villosa **
| +-----------------------------------------------+
| | +-- Atlanticoncha **
| |
| | +-- Cambarunio_nebulosus **
| | +--------------------------------------------+
+--+ | +-- Ligumia **
| |
| | +-------- Leaunio_umbrans **
| | +-----------------------------------+
| | | | +----- Cambarunio_taeniatus **
+--+ | +--+
| | | +-- Cambarunio_iris **
| | +--+
| | +-- Cambarunio_simus **
| |
| | +----------------------------------------- Hamiota **
| | |
+--+ | +----------------------- Pachynaias **
| | |
| | +--------------+ +-------------------- Sagittunio **
| | | | |
| | | | | +----- Ellipsaria **
| | | +--+ +-----------+
| | | | | | +-- Potamilus_capax **
| | | | | +--+
| | | | | +-- Truncilla **
+--+ | +--+
| | | +----- Medionidus_acutissimus **
| | | +--------+
| | | | | +-- Lemiox **
| | | | +--+
| | +--+ +-- Medionidus_penicillatus **
| | |
| | | +----------- Ptychobranchus **
| | | |
| | +--+ +-- Cyprogenia_aberti__2 **
| | | +-----+
+--+ | | +-- Dromus **
| +--+
| | +----- Cyprogenia_stegaria__2 **
| +--+
| | +-- Cyprogenia_aberti__1 **
| +--+
| +-- Cyprogenia_stegaria__1 **
|
| +-- Villosa_delumbis **
| +--------------------------------+
| | +-- Villosa_vibex **
| |
| | +-------------- Obovaria_olivaria **
| | |
| | +-----------------+ +-- Venustaconcha **
| | | | +--------+
| | | | | +-- Epioblasma **
| | | +--+
| | | | +-------- Obovaria_retusa **
+--+ | +--+
| | | +----- Obovaria_choctawensis **
| | +--+
| | | +-- Obovaria_unicolor **
| | +--+
| | +-- Obovaria_subrotunda **
| |
| | +-- Mesonaias **
| | +-----------------------+
| | | +-- Obovaria_arkansasensis **
| | |
+--+ +--+ +-- Lampsilis_splendida **
| | | +--------------------+
| | | | +-- Lampsilis_bracteata **
| | | |
| | +--+ +-- Lampsilis_fasciola **
| | | +-----------------+
| | | | +-- Ortmanniana_pectorosa **
| | +--+
| | | +----------------- Lampsilis_ornata **
| | | |
| | +--+ +-------------- Lampsilis_ovata__2 **
| | | |
| | +--+ +----------- Lampsilis_dolabraeformis **
| | | |
| | | | +----- Lampsilis_cardium__2 **
| | +--+ +--+
| | | | | +-- Lampsilis_cardium__1 **
+--+ | | +--+
| +--+ +-- Lampsilis_satura **
| |
| | +----- Lampsilis_cariosa__2 **
| +--+
| | +-- Lampsilis_ovata__1 **
| +--+
| +-- Lampsilis_cariosa__1 **
|
| +----------- Ortmanniana_ligamentina__2 **
| +--------------+
| | | +-------- Lampsilis_abrupta **
| | +--+
| | | +----- Lampsilis_higginsii__2 **
| | +--+
| | | +-- Ortmanniana_ligamentina__1 **
| | +--+
+--+ +-- Lampsilis_higginsii__1 **
|
| +----- Medionidus_conradicus **
| +-----------------+
| | | +-- Leaunio_lienosus **
| | +--+
| | +-- Leaunio_vanuxemensis **
| |
+--+ +-- Lampsilis_floridensis **
| +-----------------+
| | +-- Lampsilis_teres **
| |
| | +-- Lampsilis_siliquoidea__1 **
+--+ |
| +-- Lampsilis_siliquoidea__2 **
| +--------------+
| | +-- Lampsilis_powellii__1 **
| | |
+--+ +-- Lampsilis_powellii__2 **
|
| +-------------- Lampsilis_sietmani **
| |
+--+ +-- Lampsilis_straminea__2 **
| +--------+
| | +-- Lampsilis_virescens **
+--+
| +-------- Lampsilis_hydiana__2 **
+--+
| +----- Lampsilis_hydiana__1 **
+--+
| +-- Lampsilis_bergmanni **
+--+
+-- Lampsilis_straminea__1 **
Huang et al., 2019 [Publ 3843], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 174 | 174 | 154 | 2 | 26 | 24 | 2 | 190 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Huang et al., 2019 [Publ 3843], fig. S4
| tT | bi | riS | oT | roT | roS | iD | |
| 174 | 174 | 110 | 2 | 26 | 24 | 2 | 188 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Huang et al., 2019 [Publ 3843], fig. S3
| tT | bi | riS | oT | roT | roS | iD | |
| 172 | 172 | 169 | 2 | 26 | 24 | 2 | 186 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5
| tT | bi | riS | oT | roT | roS | iD | |
| 128 | 128 | 128 | 2 | 13 | 12 | 2 | 143.5 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Bolotov et al., 2017 [Publ 3764], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 124 | 124 | 71 | 1 | 0 | 0 | 0 | 141.5 |
1
Bolotov et al., 2017 [Publ 3761], fig. S2
| tT | bi | riS | oT | roT | roS | iD | |
| 123 | 136 | 122 | 2 | 15 | 13 | 2 | 149 |
+-- Chamberlainia **
+--------------------+
| +-- Margaritiferidae **
|
| +-- Etherioidea
--+--------------------+
| +-- Hyrioidea
|
| +-------------------- Parreysiinae **
+--+
| +----------------- Unioninae **
+--+
| +-------------- Rectidentini **
+--+
| +----------- Ambleminae **
+--+
| +-------- Pseudodontini **
+--+
| +----- Gonideini **
+--+
| +-- Sinohyriopsis **
+--+
+-- Lamprotulini **
Bolotov et al., 2017 [Publ 3761], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 123 | 136 | 136 | 2 | 15 | 13 | 2 | 150 |
+-------- Margaritiferidae ** --+ | +----- Unionidae ** +--+ | +-- Etherioidea +--+ +-- Hyrioidea
Bolotov et al., 2020 [Publ 4100], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 117 | 117 | 99 | 2 | 0 | 0 | 0 | 131 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
| tT | bi | riS | oT | roT | roS | iD | |
| 113 | 118 | 100 | 2 | 9 | 7 | 2 | 132.5 |
+-------- Hyrioidea --+ | +----- Margaritiferidae ** +--+ | +-- Etherioidea +--+ +-- Unionidae **
Bolotov et al., 2020 [Publ 4100], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 111 | 111 | 99 | 1 | 0 | 0 | 0 | 126.5 |
1
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c
| tT | bi | riS | oT | roT | roS | iD | |
| 105 | 105 | 63 | 2 | 0 | 0 | 0 | 202.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Campbell et al., 2005 [Publ 2660], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 102 | 102 | 103 | 2 | 0 | 0 | 0 | 125 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Campbell et al., 2005 [Publ 2660], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 102 | 102 | 92 | 2 | 0 | 0 | 0 | 124 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wu et al., 2026 [Publ 4811], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 96 | 96 | 95 | 2 | 0 | 0 | 0 | 108.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa
| tT | bi | riS | oT | roT | roS | iD | |
| 92 | 92 | 90 | 2 | 4 | 3 | 2 | 105.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt
| tT | bi | riS | oT | roT | roS | iD | |
| 92 | 92 | 87 | 2 | 4 | 3 | 2 | 105.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Wu et al., 2024 [Publ 4350], fig. 8b_S2
| tT | bi | riS | oT | roT | roS | iD | |
| 88 | 88 | 84 | 2 | 4 | 3 | 2 | 101.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Wu et al., 2024 [Publ 4350], fig. 8a_S1
| tT | bi | riS | oT | roT | roS | iD | |
| 88 | 88 | 85 | 2 | 4 | 3 | 2 | 101.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Dai et al., 2025 [Publ 4440], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 85 | 85 | 83 | 2 | 3 | 3 | 2 | 97.5 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Chen et al., 2025 [Publ 4448], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 83 | 84 | 82 | 2 | 3 | 3 | 2 | 97.5 |
+-------- Etherioidea --+ | +----- Unionidae ** +--+ | +-- Hyrioidea +--+ +-- Margaritiferidae **
Bolotov et al., 2018 [Publ 3826], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 82 | 82 | 50 | 1 | 0 | 0 | 0 | 96.5 |
1
Konopleva et al., 2023 [Publ 4262], fig. 8
| tT | bi | riS | oT | roT | roS | iD | |
| 82 | 82 | 79 | 1 | 0 | 0 | 0 | 98 |
1
Wu et al., 2021 [Publ 4366], fig. 2_I
| tT | bi | riS | oT | roT | roS | iD | |
| 81 | 81 | 79 | 2 | 10 | 8 | 2 | 92.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Wu et al., 2021 [Publ 4366], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 81 | 81 | 79 | 2 | 10 | 8 | 2 | 92.5 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2
| tT | bi | riS | oT | roT | roS | iD | |
| 76 | 76 | 74 | 2 | 0 | 0 | 0 | 88.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Hou et al., 2024 [Publ 4400], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 76 | 76 | 74 | 2 | 0 | 0 | 0 | 88 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1
| tT | bi | riS | oT | roT | roS | iD | |
| 76 | 76 | 73 | 2 | 0 | 0 | 0 | 88.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2
| tT | bi | riS | oT | roT | roS | iD | |
| 73 | 73 | 63 | 1 | 0 | 0 | 0 | 86 |
1
Campbell & Lydeard, 2012 [Publ 3309], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 73 | 73 | 67 | 1 | 0 | 0 | 0 | 86 |
1
Wu et al., 2022 [Publ 4238], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 71 | 71 | 69 | 2 | 3 | 3 | 2 | 83.5 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
| tT | bi | riS | oT | roT | roS | iD | |
| 71 | 77 | 71 | 2 | 9 | 8 | 2 | 88 |
+-------- Hyrioidea --+ | +----- Margaritiferidae ** +--+ | +-- Etherioidea +--+ +-- Unionidae **
Wu et al., 2021 [Publ 4366], fig. 3-ML
| tT | bi | riS | oT | roT | roS | iD | |
| 70 | 70 | 68 | 2 | 4 | 3 | 2 | 82.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Wu et al., 2021 [Publ 4366], fig. 3-BI
| tT | bi | riS | oT | roT | roS | iD | |
| 70 | 70 | 68 | 2 | 4 | 3 | 2 | 82.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Wang et al., 2025 [Publ 4488], fig. 3B
| tT | bi | riS | oT | roT | roS | iD | |
| 65 | 65 | 63 | 2 | 0 | 0 | 0 | 76.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wu et al., 2021 [Publ 4366], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 65 | 65 | 63 | 2 | 4 | 3 | 2 | 78 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Bolotov et al., 2024 [Publ 4369], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 63 | 63 | 61 | 1 | 0 | 0 | 0 | 76.5 |
1
Bolotov et al., 2024 [Publ 4369], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 62 | 62 | 60 | 1 | 0 | 0 | 0 | 75.5 |
1
Bolotov et al., 2022 [Publ 4252], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 61 | 61 | 55 | 1 | 0 | 0 | 0 | 74.5 |
1
Bolotov et al., 2022 [Publ 4252], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 61 | 61 | 49 | 1 | 0 | 0 | 0 | 74.5 |
1
Jeratthitikul et al., 2024 [Publ 4387], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 59 | 59 | 57 | 1 | 0 | 0 | 0 | 74 |
1
Bolotov et al., 2022 [Publ 4239], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 58 | 58 | 52 | 1 | 0 | 0 | 0 | 71.5 |
1
Bolotov et al., 2022 [Publ 4239], fig. S2
| tT | bi | riS | oT | roT | roS | iD | |
| 58 | 58 | 54 | 1 | 0 | 0 | 0 | 71.5 |
1
Bolotov et al., 2022 [Publ 4239], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 58 | 58 | 54 | 1 | 0 | 0 | 0 | 71.5 |
1
Sano et al., 2020 [Publ 4138], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 57 | 57 | 60 | 2 | 11 | 9 | 2 | 70 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Sano et al., 2020 [Publ 4138], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 57 | 57 | 59 | 2 | 11 | 9 | 2 | 70 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Sano et al., 2020 [Publ 4138], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 57 | 57 | 60 | 2 | 11 | 9 | 2 | 70 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Zanatta & Murphy, 2006 [Publ 2672], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 57 | 57 | 48 | 2 | 0 | 0 | 0 | 71 |
+-- Unioninae ** | --+-- Margaritiferidae ** | +-- Ambleminae **
Konopleva et al., 2023 [Publ 4269], fig. S1_4
| tT | bi | riS | oT | roT | roS | iD | |
| 56 | 56 | 54 | 2 | 0 | 0 | 0 | 67.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Zanatta & Murphy, 2006 [Publ 2672], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 56 | 56 | 51 | 2 | 0 | 0 | 0 | 69 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wu et al., 2024 [Publ 4379], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 54 | 54 | 52 | 2 | 0 | 0 | 0 | 65.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Fukata & Iigo, 2020 [Publ 4106], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 54 | 54 | 52 | 2 | 0 | 0 | 0 | 67 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Dai et al., 2023 [Publ 4285], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 53 | 53 | 51 | 2 | 0 | 0 | 0 | 65.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wu et al., 2024 [Publ 4388], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 53 | 53 | 51 | 2 | 0 | 0 | 0 | 67 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Dai et al., 2024 [Publ 4358], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 53 | 53 | 51 | 2 | 0 | 0 | 0 | 65.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Hou et al., 2025 [Publ 4445], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 53 | 53 | 51 | 1 | 0 | 0 | 0 | 66.5 |
1
Wu et al., 2018 [Publ 3836], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 53 | 53 | 45 | 2 | 0 | 0 | 0 | 65 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Huang et al., 2019 [Publ 3843], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 52 | 52 | 50 | 2 | 4 | 3 | 2 | 64.5 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
| tT | bi | riS | oT | roT | roS | iD | |
| 52 | 52 | 50 | 2 | 5 | 4 | 2 | 65.5 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Pfeiffer et al., 2019 [Publ 3937], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 52 | 53 | 50 | 2 | 5 | 4 | 2 | 69 |
+-------- Etherioidea --+ | +----- Unionidae ** +--+ | +-- Hyrioidea +--+ +-- Margaritiferidae **
Konopleva et al., 2019 [Publ 3900], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 51 | 51 | 41 | 2 | 0 | 0 | 0 | 65.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Jeratthitikul et al., 2025 [Publ 4423], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 50 | 50 | 48 | 1 | 0 | 0 | 0 | 65 |
1
Bolotov et al., 2022 [Publ 4250], fig. 8
| tT | bi | riS | oT | roT | roS | iD | |
| 49 | 49 | 42 | 2 | 0 | 0 | 0 | 62 |
+----- Parreysiinae **
+--+
| | +-- Pseudodontini **
| +--+
| +-- Rectidentini **
|
--+ +-- Gibbosulinae **
| |
| +-- Margaritiferinae **
| |
| +-- Sinosolenaia **
| |
+-----+-- Leguminaia **
|
+-- Gonidea **
|
+-- Lamprotula **
|
+-- Potomida **
Chen et al., 2025 [Publ 4663], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 48 | 48 | 47 | 2 | 0 | 0 | 0 | 62.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5
| tT | bi | riS | oT | roT | roS | iD | |
| 48 | 48 | 46 | 2 | 0 | 0 | 0 | 60 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Dai et al., 2025 [Publ 4441], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 47 | 47 | 46 | 2 | 0 | 0 | 0 | 61.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Huang et al., 2019 [Publ 3843], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 47 | 47 | 55 | 2 | 0 | 0 | 0 | 78.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Dai et al., 2024 [Publ 4389], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 46 | 46 | 44 | 2 | 0 | 0 | 0 | 59.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Huang et al., 2019 [Publ 3843], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 45 | 45 | 44 | 2 | 0 | 0 | 0 | 59.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Dai et al., 2025 [Publ 4440], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 45 | 45 | 38 | 2 | 0 | 0 | 0 | 57 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Dai et al., 2023 [Publ 4285], fig. 3-BI
| tT | bi | riS | oT | roT | roS | iD | |
| 45 | 45 | 41 | 2 | 0 | 0 | 0 | 58.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Zieritz et al., 2024 [Publ 4381], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 45 | 45 | 43 | 2 | 0 | 0 | 0 | 58.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Dai et al., 2023 [Publ 4285], fig. 3-ML
| tT | bi | riS | oT | roT | roS | iD | |
| 45 | 45 | 43 | 2 | 0 | 0 | 0 | 58.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Dai et al., 2025 [Publ 4440], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 45 | 45 | 38 | 2 | 0 | 0 | 0 | 57 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wu et al., 2020 [Publ 4121], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 45 | 45 | 43 | 2 | 4 | 3 | 2 | 57 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Froufe et al., 2019 [Publ 3936], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 44 | 44 | 42 | 2 | 0 | 0 | 0 | 56 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Konopleva et al., 2025 [Publ 4446], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 44 | 44 | 42 | 1 | 0 | 0 | 0 | 59 |
1
Froufe et al., 2019 [Publ 3936], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 44 | 44 | 42 | 2 | 0 | 0 | 0 | 56 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wu et al., 2019 [Publ 3864], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 44 | 46 | 44 | 2 | 2 | 2 | 2 | 58.5 |
+-------- Unionidae ** --+ | +----- Margaritiferidae ** +--+ | +-- Hyrioidea +--+ +-- Etherioidea
Campbell & Lydeard, 2026 [Publ 4810], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 44 | 44 | 42 | 2 | 3 | 2 | 1 | 56 |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Campbell & Lydeard, 2012 [Publ 3308], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 43 | 43 | 42 | 1 | 0 | 0 | 0 | 57 |
1
Inoue et al., 2020 [Publ 3941], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 42 | 42 | 42 | 1 | 0 | 0 | 0 | 56.5 |
1
Campbell & Lydeard, 2012 [Publ 3308], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 42 | 42 | 36 | 1 | 0 | 0 | 0 | 55 |
1
Dai et al., 2024 [Publ 4386], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 42 | 42 | 40 | 1 | 0 | 0 | 0 | 55.5 |
1
Campbell & Lydeard, 2012 [Publ 3308], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 42 | 42 | 40 | 1 | 0 | 0 | 0 | 55 |
1
Campbell & Lydeard, 2012 [Publ 3308], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 42 | 42 | 41 | 1 | 0 | 0 | 0 | 55 |
1
Smith et al., 2019 [Publ 3951], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 41 | 41 | 40 | 1 | 0 | 0 | 0 | 55.5 |
1
Chen et al., 2023 [Publ 4302], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 41 | 41 | 32 | 2 | 0 | 0 | 0 | 53 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Chen et al., 2025 [Publ 4487], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 41 | 41 | 39 | 2 | 0 | 0 | 0 | 53 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Chen et al., 2025 [Publ 4487], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 41 | 41 | 39 | 2 | 0 | 0 | 0 | 53 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Inoue et al., 2020 [Publ 3941], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 41 | 41 | 41 | 1 | 0 | 0 | 0 | 55.5 |
1
Burlakova et al., 2012 [Publ 3486], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 41 | 41 | 42 | 1 | 0 | 0 | 0 | 56 |
1
Chen et al., 2023 [Publ 4302], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 41 | 41 | 37 | 2 | 0 | 0 | 0 | 53 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Whelan et al., 2023 [Publ 4287], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 41 | 41 | 42 | 1 | 0 | 0 | 0 | 57.5 |
1
Bolotov et al., 2023 [Publ 4312], fig. S2
| tT | bi | riS | oT | roT | roS | iD | |
| 40 | 40 | 38 | 1 | 0 | 0 | 0 | 56 |
1
Aunins et al., 2018 [Publ 3863], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 40 | 40 | 38 | 2 | 0 | 0 | 0 | 52 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5
| tT | bi | riS | oT | roT | roS | iD | |
| 39 | 39 | 37 | 2 | 0 | 0 | 0 | 52 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wu et al., 2022 [Publ 4238], fig. 2d
| tT | bi | riS | oT | roT | roS | iD | |
| 39 | 39 | 37 | 1 | 0 | 0 | 0 | 52 |
1
Boyer et al., 2011 [Publ 2718], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 38 | 38 | 33 | 1 | 0 | 0 | 0 | 51.5 |
1
Bolotov et al., 2023 [Publ 4312], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 38 | 38 | 36 | 1 | 0 | 0 | 0 | 53 |
1
Bolotov et al., 2023 [Publ 4312], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 38 | 38 | 36 | 1 | 0 | 0 | 0 | 53 |
1
Chase et al., 2018 [Publ 3847], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 38 | 39 | 38 | 2 | 1 | 1 | 1 | 53 |
+-------------- Margaritiferidae ** | --+ +----------- Unioninae ** | | +--+ +-------- Lampsilini ** | | +--+ +-- Hyrioidea | +--+ | | +-- Amblemini ** +--+ | +-- Pleurobemini ** +--+ +-- Quadrulini **
Huang et al., 2018 [Publ 3828], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 38 | 38 | 36 | 2 | 4 | 3 | 2 | 50 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Cha et al., 2018 [Publ 3916], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 38 | 38 | 36 | 2 | 0 | 0 | 0 | 50 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Pfeiffer et al., 2016 [Publ 3660], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 38 | 38 | 34 | 1 | 0 | 0 | 0 | 52 |
1
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 37 | 37 | 35 | 2 | 0 | 0 | 0 | 50 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Breton et al., 2011 [Publ 3376], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 37 | 37 | 36 | 2 | 1 | 1 | 1 | 50.5 |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Perkins et al., 2017 [Publ 3777], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 37 | 37 | 34 | 1 | 0 | 0 | 0 | 51.5 |
1
Wu et al., 2024 [Publ 4350], fig. 5c
| tT | bi | riS | oT | roT | roS | iD | |
| 36 | 36 | 24 | 2 | 0 | 0 | 0 | 50.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Perkins et al., 2025 [Publ 4447], fig. 2_mtDNA-ITS
| tT | bi | riS | oT | roT | roS | iD | |
| 36 | 36 | 37 | 1 | 0 | 0 | 0 | 53.5 |
1
Wu et al., 2024 [Publ 4350], fig. 6-7
| tT | bi | riS | oT | roT | roS | iD | |
| 36 | 36 | 34 | 2 | 0 | 0 | 0 | 49.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Bolotov et al., 2020 [Publ 4084], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 36 | 36 | 22 | 2 | 0 | 0 | 0 | 50 |
+----- Margaritiferidae ** | --+----- Ambleminae ** | | +-- Gonideinae ** +--+ +-- Unioninae **
Burlakova et al., 2012 [Publ 3486], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 36 | 36 | 26 | 1 | 0 | 0 | 0 | 52.5 |
1
Wu et al., 2024 [Publ 4350], fig. 5b
| tT | bi | riS | oT | roT | roS | iD | |
| 36 | 36 | 34 | 2 | 0 | 0 | 0 | 49.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wu et al., 2024 [Publ 4388], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 36 | 36 | 34 | 2 | 0 | 0 | 0 | 49.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 36 | 36 | 22 | 1 | 0 | 0 | 0 | 50.5 |
1
Liu et al., 2024 [Publ 4380], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 36 | 36 | 34 | 2 | 0 | 0 | 0 | 49.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Klymus et al., 2025 [Publ 4814], fig. 3B
| tT | bi | riS | oT | roT | roS | iD | |
| 36 | 36 | 34 | 2 | 0 | 0 | 0 | 48.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Klymus et al., 2025 [Publ 4814], fig. 3A
| tT | bi | riS | oT | roT | roS | iD | |
| 36 | 36 | 34 | 2 | 0 | 0 | 0 | 48.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 36 | 36 | 20 | 1 | 0 | 0 | 0 | 50.5 |
1
Wu et al., 2024 [Publ 4350], fig. 5a
| tT | bi | riS | oT | roT | roS | iD | |
| 36 | 36 | 34 | 2 | 0 | 0 | 0 | 49.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Burlakova et al., 2012 [Publ 3486], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 35 | 35 | 30 | 1 | 0 | 0 | 0 | 53.5 |
1
Boyer et al., 2011 [Publ 2718], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 35 | 35 | 26 | 1 | 0 | 0 | 0 | 48.5 |
1
Jeratthitikul et al., 2021 [Publ 4219], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 35 | 35 | 33 | 1 | 0 | 0 | 0 | 50 |
1
Wu et al., 2025 [Publ 4442], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 35 | 35 | 33 | 1 | 0 | 0 | 0 | 49 |
1
Huang et al., 2002 [Publ 2657], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 34 | 34 | 31 | 2 | 0 | 0 | 0 | 48 |
+-- Cumberlandia ** | --+-- Margaritifera ** | +-- Unionidae **
Chen et al., 2026 [Publ 4805], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 34 | 34 | 32 | 1 | 0 | 0 | 0 | 48 |
1
Dai et al., 2026 [Publ 4757], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 34 | 34 | 33 | 2 | 0 | 0 | 0 | 46.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Huang et al., 2002 [Publ 2657], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 34 | 34 | 15 | 2 | 0 | 0 | 0 | 48 |
+-- Cumberlandia ** | --+-- Margaritifera ** | +-- Unionidae **
Konopleva et al., 2019 [Publ 4041], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 34 | 34 | 21 | 1 | 0 | 0 | 0 | 50 |
1
Chen et al., 2020 [Publ 4170], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 34 | 34 | 32 | 2 | 0 | 0 | 0 | 49 |
+----- Prolasmidonta **
+--+
| | +-- Alasmidonta **
| +--+
+-----------------+ +-- Anemina **
| |
| | +----- Pseudodontini **
| +--+
| | +-- Lamprotula_caveata **
| +--+
| +-- Chamberlainia **
|
| +----- Oxynaiina **
| +-----+
| | | +-- Lepidodesmini **
--+ | +--+
| | +-- Aculamprotulina **
| +-----------+
| | | +----- Platynaias **
| | | +--+
| | | | | +-- Utterbackia_imbecillis **
| | | | +--+
| | +--+ +-- Pyganodon **
| | |
| | | +----- Sinanodonta **
| | +--+
| | | +-- Cristaria **
+--+ +--+
| +-- Anodontina **
|
| +-- Utterbackia_peninsularis **
| +-----+
| | +-- Potamilus_alatus **
| +-----------+
| | | +----- Lamprotula_leaii **
| | +--+
| | | +-- Sinohyriopsis **
+--+ +--+
| +-- Gonideini **
|
| +----------------- Margaritiferidae **
+--+
| +-------------- Quadrulini **
+--+
| +----------- Toxolasma **
+--+
| +-------- Lampsilis_ornata **
+--+
| +----- Potamilus_leptodon **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Lampsilis_powellii **
Jeratthitikul et al., 2021 [Publ 4219], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 34 | 34 | 32 | 1 | 0 | 0 | 0 | 50 |
1
Sano et al., 2020 [Publ 4138], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 33 | 33 | 16 | 2 | 1 | 0 | 0 | 56.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Burlakova et al., 2019 [Publ 3899], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 33 | 33 | 21 | 1 | 0 | 0 | 0 | 46 |
1
Krebs et al., 2003 [Publ 2691], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 33 | 33 | 29 | 2 | 0 | 0 | 0 | 46.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Lopes-Lima et al., 2017 [Publ 3792], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 32 | 32 | 30 | 2 | 0 | 0 | 0 | 44 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 32 | 32 | 30 | 2 | 8 | 7 | 2 | 43 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Sano et al., 2017 [Publ 3799], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 32 | 2 | 2 | 0 | 0 | 45.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Fonseca et al., 2016 [Publ 3775], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 29 | 2 | 0 | 0 | 0 | 43 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wang et al., 2018 [Publ 3865], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 29 | 1 | 0 | 0 | 0 | 44.5 |
1
Zieritz et al., 2021 [Publ 4218], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 29 | 2 | 0 | 0 | 0 | 44 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wang et al., 2018 [Publ 3846], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 29 | 1 | 0 | 0 | 0 | 44.5 |
1
Ouyang et al., 2015 [Publ 4493], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 29 | 1 | 0 | 0 | 0 | 46 |
1
Pfeiffer et al., 2019 [Publ 3866], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 30 | 1 | 0 | 0 | 0 | 44 |
1
Zieritz et al., 2020 [Publ 4102], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 27 | 1 | 0 | 0 | 0 | 46 |
1
Dai et al., 2024 [Publ 4385], fig. 3-ML
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 29 | 2 | 0 | 0 | 0 | 43.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Lyubas et al., 2022 [Publ 4255], fig. 7
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 28 | 1 | 0 | 0 | 0 | 43 |
1
Dai et al., 2024 [Publ 4385], fig. 3-BI
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 28 | 2 | 0 | 0 | 0 | 43.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Ouyang et al., 2015 [Publ 4493], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 38 | 1 | 0 | 0 | 0 | 54 |
1
Bogan et al., 2023 [Publ 4295], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 28 | 2 | 0 | 0 | 0 | 43.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Lyubas et al., 2022 [Publ 4255], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 22 | 1 | 0 | 0 | 0 | 43 |
1
Burdick & White, 2007 [Publ 2679], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 30 | 30 | 31 | 1 | 0 | 0 | 0 | 46 |
1
Wu et al., 2020 [Publ 4147], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 30 | 30 | 29 | 2 | 0 | 0 | 0 | 42.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 30 | 31 | 29 | 2 | 1 | 1 | 1 | 44.5 |
+-- Margaritiferidae ** | --+-- Hyrioidea | +-- Unionidae **
Serb et al., 2003 [Publ 2659], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 29 | 29 | 20 | 1 | 0 | 0 | 0 | 42 |
1
Serb et al., 2003 [Publ 2659], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 29 | 29 | 27 | 1 | 0 | 0 | 0 | 42 |
1
Lopes-Lima et al., 2019 [Publ 3901], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 29 | 29 | 27 | 2 | 0 | 0 | 0 | 41.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Pfeiffer et al., 2021 [Publ 4126], fig. 9
| tT | bi | riS | oT | roT | roS | iD | |
| 29 | 29 | 27 | 1 | 0 | 0 | 0 | 44 |
1
Whelan et al., 2023 [Publ 4287], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 28 | 28 | 27 | 1 | 0 | 0 | 0 | 42.5 |
1
Hou et al., 2025 [Publ 4401], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 28 | 28 | 26 | 1 | 0 | 0 | 0 | 43 |
1
Guerra et al., 2019 [Publ 4168], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 28 | 28 | 26 | 2 | 13 | 12 | 2 | 40.5 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Hou et al., 2025 [Publ 4432], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 28 | 28 | 26 | 1 | 0 | 0 | 0 | 43 |
1
Pfeiffer et al., 2018 [Publ 3827], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 28 | 28 | 26 | 2 | 0 | 0 | 0 | 40 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Guerra et al., 2019 [Publ 4168], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 28 | 28 | 26 | 2 | 13 | 12 | 2 | 40.5 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Xiang et al., 2025 [Publ 4433], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 28 | 28 | 26 | 1 | 0 | 0 | 0 | 42 |
1
Lopes-Lima et al., 2019 [Publ 3901], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 28 | 28 | 25 | 2 | 0 | 0 | 0 | 40 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Smith et al., 2020 [Publ 4129], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 28 | 28 | 26 | 1 | 0 | 0 | 0 | 41.5 |
1
Smith et al., 2020 [Publ 4129], fig. 2-3
| tT | bi | riS | oT | roT | roS | iD | |
| 28 | 28 | 26 | 1 | 0 | 0 | 0 | 41.5 |
1
Whelan et al., 2023 [Publ 4287], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 27 | 27 | 25 | 1 | 0 | 0 | 0 | 41.5 |
1
Lydeard et al., 1996 [Publ 2656], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 27 | 27 | 25 | 2 | 1 | 0 | 0 | 40 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A
| tT | bi | riS | oT | roT | roS | iD | |
| 27 | 27 | 21 | 2 | 3 | 2 | 2 | 38 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Graf & Cummings, 2006 [Publ 1968], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 27 | 27 | 21 | 2 | 20 | 19 | 2 | 40 |
+-- Hyrioidea
+--+
| +-- Etherioidea
--+
| +-- Margaritiferidae **
+--+
+-- Unionidae **
Neemuchwala et al., 2023 [Publ 4292], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 27 | 27 | 25 | 1 | 0 | 0 | 0 | 40.5 |
1
Hoeh et al., 2009 [Publ 2716], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 27 | 28 | 26 | 2 | 20 | 19 | 2 | 41 |
+-------------------------- Hyrioidea | | +-------- Iridinidae --+ +--------------+ | | | +----- Etheria | | +--+ | | | +-- Mulleria +--+ +--+ | +-- Mycetopodidae | | +-------------------- Margaritiferidae ** +--+ | +----------------- Parreysiinae ** +--+ | +-------------- Pseudomulleria +--+ | +----------- Unioninae ** +--+ | +-------- Rectidentini ** +--+ | +----- Ambleminae ** +--+ | +-- Gonideini ** +--+ +-- Pseudodontini **
Lydeard et al., 1996 [Publ 2656], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 27 | 27 | 23 | 2 | 0 | 0 | 0 | 40 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B
| tT | bi | riS | oT | roT | roS | iD | |
| 27 | 27 | 25 | 2 | 3 | 2 | 2 | 38 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Zhang et al., 2024 [Publ 4397], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 27 | 27 | 25 | 2 | 0 | 0 | 0 | 40.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Campbell et al., 2008 [Publ 2675], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 26 | 26 | 23 | 1 | 0 | 0 | 0 | 40.5 |
1
Campbell et al., 2008 [Publ 2675], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 26 | 26 | 18 | 1 | 0 | 0 | 0 | 41.5 |
1
Hoeh et al., 2009 [Publ 2716], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 26 | 27 | 24 | 2 | 20 | 19 | 2 | 42 |
+-------------------------- Hyrioidea | | +-- Etheria | +--+ --+ | +-- Mulleria | +-----------------+ | | | +-- Mycetopodidae | | +--+ +--+ +-- Iridinidae | | +-------------------- Parreysiinae ** +--+ | +----------------- Pseudomulleria +--+ | +-------------- Margaritiferidae ** +--+ | +----------- Pseudodontini ** +--+ | +-------- Rectidentini ** +--+ | +----- Unioninae ** +--+ | +-- Gonideini ** +--+ +-- Ambleminae **
Whelan et al., 2011 [Publ 3380], fig. 1-ML
| tT | bi | riS | oT | roT | roS | iD | |
| 26 | 31 | 30 | 2 | 10 | 9 | 2 | 44 |
+-------- Hyrioidea --+ | +----- Margaritiferidae ** +--+ | +-- Etherioidea +--+ +-- Unionidae **
Whelan et al., 2011 [Publ 3380], fig. 1-MP
| tT | bi | riS | oT | roT | roS | iD | |
| 26 | 30 | 27 | 2 | 10 | 9 | 2 | 44.5 |
+-------- Etherioidea --+ | +----- Unionidae ** +--+ | +-- Margaritiferidae ** +--+ +-- Hyrioidea
Hoeh et al., 2009 [Publ 2716], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 26 | 45 | 39 | 2 | 20 | 19 | 2 | 58 |
+----------------- Parreysiinae ** | | +----- Margaritiferidae ** --+ +--------+ | | | +-- Hyrioidea | | +--+ +--+ +-- Etherioidea | | +----------- Pseudodontini ** +--+ | +-------- Rectidentini ** +--+ | +----- Unioninae ** +--+ | +-- Gonideini ** +--+ +-- Ambleminae **
Lopes-Lima et al., 2019 [Publ 3901], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 26 | 26 | 24 | 2 | 0 | 0 | 0 | 38.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Konopleva et al., 2021 [Publ 4213], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 26 | 26 | 21 | 1 | 0 | 0 | 0 | 42 |
1
Campbell et al., 2008 [Publ 2675], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 25 | 25 | 9 | 1 | 0 | 0 | 0 | 39.5 |
1
Lopes-Lima et al., 2021 [Publ 4217], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 25 | 25 | 23 | 1 | 0 | 0 | 0 | 38.5 |
1
Campbell et al., 2008 [Publ 2675], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 25 | 25 | 6 | 1 | 0 | 0 | 0 | 40.5 |
1
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11
| tT | bi | riS | oT | roT | roS | iD | |
| 25 | 25 | 23 | 1 | 0 | 0 | 0 | 39.5 |
1
Perkins et al., 2017 [Publ 3777], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 24 | 24 | 10 | 1 | 0 | 0 | 0 | 41.5 |
1
Galal-Khallaf et al., 2024 [Publ 4459], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 24 | 28 | 29 | 1 | 4 | 4 | 1 | 53.5 |
+----- Parreysiinae ** --+ | +-- Etherioidea +--+ +-- Unioninae **
Liu et al., 2023 [Publ 4310], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 24 | 24 | 17 | 1 | 0 | 0 | 0 | 37 |
1
Wu et al., 2021 [Publ 4215], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 24 | 24 | 22 | 1 | 0 | 0 | 0 | 38 |
1
Liu et al., 2023 [Publ 4310], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 24 | 24 | 22 | 1 | 0 | 0 | 0 | 37 |
1
Jeratthitikul et al., 2019 [Publ 3922], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 23 | 23 | 21 | 1 | 0 | 0 | 0 | 36.5 |
1
Konopleva et al., 2017 [Publ 3807], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 23 | 23 | 21 | 2 | 8 | 6 | 2 | 36.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Wu et al., 2020 [Publ 4121], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 23 | 23 | 21 | 2 | 11 | 9 | 2 | 35.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Unionidae ** +--+ +-- Margaritiferidae **
Zieritz et al., 2021 [Publ 4231], fig. 3-M_4
| tT | bi | riS | oT | roT | roS | iD | |
| 23 | 23 | 21 | 2 | 0 | 0 | 0 | 36 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wu et al., 2023 [Publ 4296], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 23 | 23 | 21 | 1 | 0 | 0 | 0 | 36 |
1
Wu et al., 2024 [Publ 4379], fig. 4_6
| tT | bi | riS | oT | roT | roS | iD | |
| 23 | 23 | 21 | 2 | 0 | 0 | 0 | 35.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wu et al., 2020 [Publ 4121], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 23 | 23 | 21 | 2 | 11 | 9 | 2 | 35.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Unionidae ** +--+ +-- Margaritiferidae **
Inoue et al., 2020 [Publ 4128], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 23 | 23 | 16 | 1 | 0 | 0 | 0 | 39.5 |
1
Soroka & Burzynski, 2017 [Publ 3837], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 23 | 23 | 21 | 1 | 0 | 0 | 0 | 36 |
1
Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE
| tT | bi | riS | oT | roT | roS | iD | |
| 22 | 22 | 20 | 1 | 0 | 0 | 0 | 35.5 |
1
Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG
| tT | bi | riS | oT | roT | roS | iD | |
| 22 | 22 | 20 | 1 | 0 | 0 | 0 | 35.5 |
1
Huang et al., 2019 [Publ 3843], fig. 3-M
| tT | bi | riS | oT | roT | roS | iD | |
| 22 | 22 | 20 | 2 | 1 | 1 | 1 | 34.5 |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Dai et al., 2024 [Publ 4386], fig. 5-ML
| tT | bi | riS | oT | roT | roS | iD | |
| 22 | 22 | 19 | 2 | 0 | 0 | 0 | 33 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wu et al., 2018 [Publ 4143], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 22 | 22 | 20 | 1 | 0 | 0 | 0 | 35.5 |
1
Inoue et al., 2020 [Publ 4128], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 22 | 22 | 24 | 1 | 0 | 0 | 0 | 37.5 |
1
Dai et al., 2024 [Publ 4386], fig. 5-BI
| tT | bi | riS | oT | roT | roS | iD | |
| 22 | 22 | 16 | 2 | 0 | 0 | 0 | 33 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Rosenberg et al., 1997 [Publ 2720], fig. 2B
| tT | bi | riS | oT | roT | roS | iD | |
| 21 | 22 | 3 | 2 | 1 | 0 | 0 | 31.5 |
+-- Cumberlandia ** | +-- None__1 | --+-- None__2 | +-- Margaritifera ** | +-- Unionidae **
Chapman et al., 2008 [Publ 2744], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 21 | 21 | 19 | 1 | 0 | 0 | 0 | 34.5 |
1
Zhou et al., 2007 [Publ 2658], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 21 | 21 | 19 | 2 | 0 | 0 | 0 | 33.5 |
+-------------------- Margaritiferinae ** | | +----------------- Anodontini ** --+ | | | +----------- Chamberlainiini ** | | +--+ +--+ | | +-------- Lamprotula_leaii ** | | +--+ | | | +----- Lamprotula_caveata ** | | +--+ | | | +-- Gibbosulinae ** +--+ +--+ | +-- Lamprotula_cornuumlunae ** | | +----- Aculamprotulina ** | +--+ | | | +-- Tchangsinaia ** | | +--+ +-----+ +-- Nodularia ** | | +----- Schistodesmus ** +--+ | +-- Lanceolariini ** +--+ +-- Acuticostini **
Jeratthitikul et al., 2021 [Publ 4212], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 21 | 21 | 19 | 1 | 0 | 0 | 0 | 36 |
1
Wu et al., 2016 [Publ 3711], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 21 | 21 | 19 | 1 | 0 | 0 | 0 | 34.5 |
1
Chapman et al., 2008 [Publ 2667], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 21 | 21 | 19 | 1 | 0 | 0 | 0 | 34.5 |
1
Rosenberg et al., 1997 [Publ 2720], fig. 2A
| tT | bi | riS | oT | roT | roS | iD | |
| 21 | 22 | 4 | 2 | 1 | 0 | 0 | 31.5 |
+----- Cumberlandia ** | +----- None__2 --+ +----- Margaritifera ** | | +-- None__1 +--+ +-- Unionidae **
Guerra et al., 2017 [Publ 3772], fig. 2a
| tT | bi | riS | oT | roT | roS | iD | |
| 21 | 21 | 19 | 2 | 4 | 3 | 2 | 33 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Guerra et al., 2017 [Publ 3772], fig. 2b
| tT | bi | riS | oT | roT | roS | iD | |
| 21 | 21 | 19 | 2 | 4 | 3 | 2 | 33 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Matos et al., 2024 [Publ 4383], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 18 | 2 | 0 | 0 | 0 | 33.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Roe & Hoeh, 2003 [Publ 2655], fig. 4f2
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 16 | 2 | 11 | 10 | 2 | 31.5 |
+-- Etherioidea
+--+
| +-- Hyrioidea
--+
| +-- Margaritiferidae **
+--+
+-- Unionidae **
Hoeh et al., 2001 [Publ 2652], fig. 14f4
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 17 | 2 | 11 | 10 | 2 | 34.5 |
+----------------- Hyrioidea --+ | +-------------- Etherioidea +--+ | +----------- Parreysiinae ** +--+ | +-------- Margaritiferidae ** +--+ | +----- Gonideinae ** +--+ | +-- Unioninae ** +--+ +-- Ambleminae **
Guerra et al., 2019 [Publ 4168], fig. 2-M
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 18 | 2 | 2 | 2 | 1 | 32.5 |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Hoeh et al., 2001 [Publ 2652], fig. 14f2
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 30 | 18 | 2 | 11 | 10 | 2 | 40 |
+----------------- Margaritiferidae ** | | +-------------- Amblemini ** | | | +-------------- Parreysiinae ** | | | +-------------- Eurynia ** --+ | | +-------------- Fusconaia ** | | | +-------------- Gonideinae ** | | | +-------------- Pleurobema ** | | +--+-------------- Quadrulini ** | | +----------- Ortmanniana ** +--+ | | +-------- Glebula ** | +--+ | | +----- Potamilus ** | +--+ | | +-- Ligumia ** | +--+ | +-- Toxolasma ** | | +----------- Cyrtonaias ** +--+ | +-------- Unionini ** +--+ | +----- Anodontini ** +--+ | +-- Hyrioidea +--+ +-- Etherioidea
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 17 | 2 | 0 | 0 | 0 | 33 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Matos et al., 2024 [Publ 4384], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 18 | 2 | 0 | 0 | 0 | 33.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Zhou et al., 2016 [Publ 3710], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 18 | 2 | 0 | 0 | 0 | 34 |
+-------- Unioninae ** --+ | +----- Margaritiferidae ** +--+ | +-- Ambleminae ** +--+ +-- Gonideinae **
Smith et al., 2020 [Publ 4136], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 13 | 1 | 0 | 0 | 0 | 34.5 |
1
Bogan & Hoeh, 2000 [Publ 2650], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 21 | 14 | 2 | 14 | 13 | 2 | 38.5 |
+-------------------- Hyrioidea | | +-------- Iridinidae --+ +--------+ | | | +----- Etheria | | +--+ | | | +-- Mulleria +--+ +--+ | +-- Mycetopodidae | | +-------------- Parreysiinae ** | | +--+ +----------- Pseudomulleria | | +--+ +-------- Anodontini ** | | +--+ +----- Gonideinae ** | | +--+----- Unionini ** | | +-- Margaritiferidae ** +--+ +-- Ambleminae **
Inoue et al., 2014 [Publ 3635], fig. 3A
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 17 | 2 | 0 | 0 | 0 | 32 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Roe & Hoeh, 2003 [Publ 2655], fig. 4f3
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 30 | 28 | 2 | 11 | 10 | 2 | 40 |
+-------- Margaritiferidae ** --+ | +----- Unionidae ** +--+ | +-- Etherioidea +--+ +-- Hyrioidea
Pieri et al., 2018 [Publ 3867], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 23 | 1 | 0 | 0 | 0 | 39 |
1
Hoeh et al., 2002 [Publ 2653], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 17 | 2 | 11 | 10 | 2 | 34.5 |
+----------------- Hyrioidea --+ | +-------------- Etherioidea +--+ | +----------- Parreysiinae ** +--+ | +-------- Margaritiferidae ** +--+ | +----- Ambleminae ** +--+ | +-- Gonideinae ** +--+ +-- Unioninae **
Guerra et al., 2019 [Publ 4168], fig. 1-M
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 18 | 2 | 2 | 2 | 1 | 32.5 |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 19 | 19 | 17 | 2 | 0 | 0 | 0 | 31.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Froufe et al., 2019 [Publ 3936], fig. 3-M
| tT | bi | riS | oT | roT | roS | iD | |
| 19 | 19 | 17 | 2 | 0 | 0 | 0 | 32 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Riccardi et al., 2020 [Publ 4089], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 19 | 19 | 17 | 2 | 0 | 0 | 0 | 31 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Froufe et al., 2019 [Publ 3936], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 19 | 19 | 17 | 2 | 0 | 0 | 0 | 32 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wu et al., 2023 [Publ 4339], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 19 | 19 | 17 | 2 | 0 | 0 | 0 | 31.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Sano et al., 2017 [Publ 3799], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 19 | 19 | 22 | 2 | 2 | 0 | 0 | 34.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Rosenberg et al., 1994 [Publ 2719], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 19 | 19 | 5 | 2 | 0 | 0 | 0 | 29.5 |
+-- Cumberlandia ** | --+-- Margaritifera ** | +-- Unionidae **
Bolotov et al., 2019 [Publ 3974], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 18 | 18 | 16 | 1 | 0 | 0 | 0 | 33 |
1
Chong et al., 2008 [Publ 2668], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 18 | 18 | 10 | 1 | 0 | 0 | 0 | 31 |
1
Porto-Hannes et al., 2019 [Publ 3969], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 18 | 18 | 16 | 1 | 0 | 0 | 0 | 32.5 |
1
Porto-Hannes et al., 2019 [Publ 3969], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 18 | 18 | 13 | 1 | 0 | 0 | 0 | 32.5 |
1
Huang et al., 2018 [Publ 3828], fig. 3_5
| tT | bi | riS | oT | roT | roS | iD | |
| 18 | 18 | 16 | 2 | 11 | 9 | 2 | 30.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Unionidae ** +--+ +-- Margaritiferidae **
Araujo et al., 2018 [Publ 3779], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 18 | 18 | 14 | 2 | 0 | 0 | 0 | 30.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Smith et al., 2018 [Publ 3790], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 18 | 18 | 17 | 1 | 0 | 0 | 0 | 32.5 |
1
Wu et al., 2022 [Publ 4238], fig. 2e
| tT | bi | riS | oT | roT | roS | iD | |
| 18 | 18 | 16 | 1 | 0 | 0 | 0 | 31 |
1
Zhou et al., 2007 [Publ 2658], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 18 | 18 | 16 | 1 | 0 | 0 | 0 | 33 |
1
Sano et al., 2022 [Publ 4311], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 18 | 18 | 16 | 1 | 0 | 0 | 0 | 33 |
1
Kim et al., 2021 [Publ 4197], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 18 | 18 | 16 | 1 | 0 | 0 | 0 | 32.5 |
1
Wu et al., 2026 [Publ 4811], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 17 | 17 | 15 | 2 | 0 | 0 | 0 | 30 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Prié & Puillandre, 2014 [Publ 3529], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 17 | 17 | 15 | 2 | 0 | 0 | 0 | 27.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Smith, 2021 [Publ 4331], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 17 | 17 | 15 | 2 | 3 | 2 | 1 | 30 |
+----- Etherioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Fukata & Iigo, 2019 [Publ 4108], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 17 | 17 | 15 | 1 | 0 | 0 | 0 | 31.5 |
1
Chen et al., 2025 [Publ 4455], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 17 | 17 | 15 | 2 | 9 | 7 | 2 | 30 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Unionidae ** +--+ +-- Margaritiferidae **
Inoue et al., 2013 [Publ 3528], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 17 | 17 | 13 | 1 | 0 | 0 | 0 | 31 |
1
Grobler et al., 2011 [Publ 3359], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 17 | 17 | 13 | 1 | 0 | 0 | 0 | 31.5 |
1
Plazzi et al., 2011 [Publ 3676], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 17 | 18 | 11 | 1 | 1 | 1 | 1 | 32.5 |
+----------- Cristariina ** | | +-------- Lanceolariini ** | | --+ +-------- Anodonta ** | | | +-------- Unionini ** | | +--+ +-- Alasmidontina ** +-----+ | +-- Pseudanodonta ** | | +----- Ambleminae ** | | +--+ +-- Hyrioidea | | +--+-- Lamprotulini ** | +-- Chamberlainiini **
Lopes-Lima et al., 2018 [Publ 3829], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 17 | 17 | 15 | 2 | 11 | 9 | 2 | 28.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Unionidae ** +--+ +-- Margaritiferidae **
Lopes-Lima et al., 2018 [Publ 3829], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 17 | 17 | 15 | 2 | 11 | 9 | 2 | 28.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Unionidae ** +--+ +-- Margaritiferidae **
Wu et al., 2025 [Publ 4442], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 17 | 17 | 15 | 1 | 0 | 0 | 0 | 30.5 |
1
Stewart et al., 2021 [Publ 4451], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 16 | 16 | 15 | 2 | 0 | 0 | 0 | 28 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Zhou et al., 2017 [Publ 3788], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 16 | 16 | 14 | 1 | 0 | 0 | 0 | 30.5 |
1
Szumowski et al., 2012 [Publ 3569], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 16 | 16 | 12 | 1 | 0 | 0 | 0 | 30.5 |
1
Zanatta et al., 2007 [Publ 2698], fig. 1B
| tT | bi | riS | oT | roT | roS | iD | |
| 16 | 16 | 13 | 1 | 0 | 0 | 0 | 30.5 |
1
Zanatta et al., 2007 [Publ 2698], fig. 1A
| tT | bi | riS | oT | roT | roS | iD | |
| 16 | 16 | 10 | 1 | 0 | 0 | 0 | 30.5 |
1
Lopes-Lima et al., 2017 [Publ 3734], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 16 | 16 | 12 | 2 | 1 | 0 | 0 | 27 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wen et al., 2017 [Publ 3795], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 16 | 16 | 14 | 2 | 0 | 0 | 0 | 28.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Huang et al., 2018 [Publ 3828], fig. 6-M
| tT | bi | riS | oT | roT | roS | iD | |
| 16 | 16 | 14 | 2 | 1 | 1 | 1 | 29 |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Johnson et al., 2018 [Publ 3844], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 16 | 16 | 14 | 1 | 0 | 0 | 0 | 30.5 |
1
Walker et al., 2006 [Publ 2666], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 16 | 16 | 13 | 2 | 20 | 19 | 2 | 30.5 |
+-------------- Hyrioidea --+ | +----------- Etherioidea +--+ | +-------- Unioninae ** +--+ | +----- Margaritiferidae ** +--+ | +-- Gonideinae ** +--+ +-- Ambleminae **
Huang et al., 2013 [Publ 3629], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 16 | 16 | 14 | 2 | 0 | 0 | 0 | 28 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Inoue et al., 2020 [Publ 3941], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 16 | 16 | 14 | 1 | 0 | 0 | 0 | 30.5 |
1
Lyubas et al., 2019 [Publ 3963], fig. 10
| tT | bi | riS | oT | roT | roS | iD | |
| 16 | 16 | 12 | 1 | 0 | 0 | 0 | 30.5 |
1
Kongim et al., 2023 [Publ 4290], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 13 | 1 | 0 | 0 | 0 | 30 |
1
Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 13 | 1 | 0 | 0 | 0 | 28 |
1
Amaro et al., 2017 [Publ 3791], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 15 | 2 | 1 | 0 | 0 | 28.5 |
+-------- Margaritifera_margaritifera__2 ** --+ | +----- Unionidae ** +--+ | +-- Margaritifera_margaritifera__1 ** +--+ +-- Margaritifera_falcata **
Hoeh, 1990 [Publ 2717], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 2 | 1 | 0 | 0 | 0 | 29.5 |
1
Hou et al., 2024 [Publ 4400], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 13 | 1 | 0 | 0 | 0 | 30 |
1
Liu et al., 2017 [Publ 3939], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 12 | 1 | 0 | 0 | 0 | 28.5 |
1
Bolotov et al., 2022 [Publ 4250], fig. S5
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 12 | 1 | 0 | 0 | 0 | 30 |
1
Bolotov et al., 2022 [Publ 4250], fig. S2
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 13 | 1 | 0 | 0 | 0 | 29.5 |
1
Jeratthitikul et al., 2025 [Publ 4476], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 13 | 1 | 0 | 0 | 0 | 30 |
1
Hoeh, 1990 [Publ 2717], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 13 | 1 | 0 | 0 | 0 | 29.5 |
1
Hoeh, 1990 [Publ 2717], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 13 | 1 | 0 | 0 | 0 | 29.5 |
1
Xue et al., 2016 [Publ 3659], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 13 | 1 | 0 | 0 | 0 | 29 |
1
Porto-Hannes et al., 2021 [Publ 4227], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 12 | 1 | 0 | 0 | 0 | 28.5 |
1
Douda et al., 2024 [Publ 4364], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 13 | 2 | 0 | 0 | 0 | 27.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Dai et al., 2024 [Publ 4386], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 1 | 0 | 0 | 0 | 26.5 |
1
Wu et al., 2020 [Publ 4147], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 2 | 0 | 0 | 0 | 25.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Lydeard et al., 2000 [Publ 2654], fig. 2b
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 2 | 0 | 0 | 0 | 29.5 |
+-- Unioninae ** | --+-- Margaritiferidae ** | +-- Ambleminae **
Szumowski et al., 2012 [Publ 3569], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 1 | 0 | 0 | 0 | 29 |
1
Turner et al., 2000 [Publ 2722], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 2 | 0 | 0 | 0 | 28 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Reis et al., 2013 [Publ 3580], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 2 | 1 | 0 | 0 | 29 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Bolotov et al., 2016 [Publ 3743], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 2 | 16 | 13 | 2 | 26.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Unionidae ** +--+ +-- Margaritiferidae **
Feng et al., 2016 [Publ 3800], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 2 | 0 | 0 | 0 | 26.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Lydeard et al., 2000 [Publ 2654], fig. 2c
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 9 | 2 | 0 | 0 | 0 | 28.5 |
+----- Unioninae ** | +----- Margaritiferidae ** | --+----- Quadrulini ** | +----- Lampsilini ** | | +-- Amblemini ** +--+ +-- Pleurobemini **
Lopes-Lima et al., 2024 [Publ 4376], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 1 | 0 | 0 | 0 | 29.5 |
1
Froufe et al., 2016 [Publ 3738], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 1 | 0 | 0 | 0 | 27.5 |
1
Araujo et al., 2017 [Publ 3801], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 2 | 0 | 0 | 0 | 26 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Zhang et al., 2004 [Publ 2735], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 1 | 0 | 0 | 0 | 28 |
1
Bolotov et al., 2016 [Publ 3743], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 2 | 16 | 13 | 2 | 26.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Unionidae ** +--+ +-- Margaritiferidae **
Lydeard et al., 2000 [Publ 2654], fig. 2a
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 11 | 2 | 0 | 0 | 0 | 28.5 |
+----- Unioninae ** | +----- Margaritiferidae ** | --+----- Lampsilini ** | +----- Amblemini ** | | +-- Quadrulini ** +--+ +-- Pleurobemini **
Lydeard et al., 2000 [Publ 2654], fig. 2d
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 2 | 0 | 0 | 0 | 29.5 |
+-- Unioninae ** | --+-- Margaritiferidae ** | +-- Ambleminae **
Lopes-Lima et al., 2020 [Publ 4080], fig. 10
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 10 | 2 | 0 | 0 | 0 | 27.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Burzynski et al., 2017 [Publ 3774], fig. 1-bottom
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 1 | 0 | 0 | 0 | 28 |
1
Graf, 2002 [Publ 2651], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 2 | 5 | 4 | 1 | 27.5 |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Sayenko et al., 2021 [Publ 4189], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 1 | 0 | 0 | 0 | 27.5 |
1
Walker et al., 2006 [Publ 2666], fig. 6-M
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 2 | 1 | 1 | 1 | 25.5 |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Burzynski et al., 2017 [Publ 3774], fig. 1-top
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 1 | 0 | 0 | 0 | 28 |
1
Walker et al., 2006 [Publ 2666], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 2 | 2 | 1 | 1 | 27 |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Xiang et al., 2025 [Publ 4433], fig. 7
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 1 | 0 | 0 | 0 | 28 |
1
Walker et al., 2006 [Publ 2666], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 2 | 2 | 1 | 1 | 29 |
+----------------- Hyrioidea | --+ +-------------- Unioninae ** | | +--+ +----------- Lamprotulini ** | | +--+ +-- Gonideini ** | +-----+ | | +-- Pseudodontini ** +--+ | +----- Lampsilini ** +--+ | +-- Pleurobemini ** +--+ +-- Margaritiferidae **
Vikhrev et al., 2018 [Publ 3812], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 2 | 0 | 0 | 0 | 25.5 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Ouyang et al., 2011 [Publ 3408], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 10 | 1 | 0 | 0 | 0 | 28 |
1
Jeratthitikul & Sutcharit, 2023 [Publ 4289], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 1 | 0 | 0 | 0 | 29 |
1
Wu et al., 2024 [Publ 4379], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 10 | 1 | 0 | 0 | 0 | 29 |
1
Hauck et al., 2023 [Publ 4333], fig. 9
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 12 | 1 | 0 | 0 | 0 | 26.5 |
1
Bogan et al., 2023 [Publ 4295], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 10 | 2 | 0 | 0 | 0 | 28 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Manendo et al., 2008 [Publ 2669], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 1 | 0 | 0 | 0 | 28.5 |
1
Reis et al., 2013 [Publ 3580], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 2 | 1 | 0 | 0 | 27 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Walker et al., 2006 [Publ 2666], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 2 | 2 | 1 | 1 | 27 |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Pfarrer et al., 2022 [Publ 4305], fig. 2-5
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 10 | 1 | 0 | 0 | 0 | 26.5 |
1
Freeman et al., 2025 [Publ 4443], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 10 | 1 | 0 | 0 | 0 | 27 |
1
Zieritz et al., 2026 [Publ 4812], fig. 2a
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 10 | 1 | 0 | 0 | 0 | 28 |
1
Froufe et al., 2017 [Publ 3802], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 9 | 1 | 0 | 0 | 0 | 28 |
1
Konopleva et al., 2020 [Publ 4072], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 9 | 1 | 0 | 0 | 0 | 28 |
1
Lopes-Lima et al., 2020 [Publ 4080], fig. 11
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 10 | 2 | 0 | 0 | 0 | 26.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Araujo et al., 2016 [Publ 3746], fig. 1_2b
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 10 | 2 | 1 | 0 | 0 | 26.5 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Choi et al., 2020 [Publ 4169], fig. 2a
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 11 | 1 | 0 | 0 | 0 | 27.5 |
1
Wu et al., 2022 [Publ 4238], fig. 2a_S1
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 10 | 1 | 0 | 0 | 0 | 28 |
1
Smith et al., 2020 [Publ 4136], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 10 | 1 | 0 | 0 | 0 | 28 |
1
Froufe et al., 2015 [Publ 3664], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 10 | 1 | 0 | 0 | 0 | 26.5 |
1
Liu et al., 2024 [Publ 4380], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 10 | 1 | 0 | 0 | 0 | 28 |
1
Mabuchi & Nishida, 2020 [Publ 4120], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 12 | 1 | 0 | 0 | 0 | 39 |
1
Freeman et al., 2025 [Publ 4443], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 8 | 1 | 0 | 0 | 0 | 27 |
1
Sano et al., 2022 [Publ 4311], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 12 | 1 | 0 | 0 | 0 | 27 |
1
Zieritz et al., 2024 [Publ 4381], fig. 7A
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 10 | 1 | 0 | 0 | 0 | 28 |
1
Curole & Kocher, 2005 [Publ 2682], fig. 1-M
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 10 | 2 | 0 | 0 | 0 | 24.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Hou et al., 2025 [Publ 4432], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 9 | 1 | 0 | 0 | 0 | 27 |
1
Lopes-Lima et al., 2020 [Publ 4080], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 9 | 2 | 0 | 0 | 0 | 25.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Porto-Hannes et al., 2021 [Publ 4227], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 6 | 1 | 0 | 0 | 0 | 26.5 |
1
Pfeiffer et al., 2021 [Publ 4126], fig. 12
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 8 | 1 | 0 | 0 | 0 | 27 |
1
Curole & Kocher, 2005 [Publ 2682], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 9 | 2 | 0 | 0 | 0 | 25.5 |
+----- Ambleminae ** --+ | +-- Unioninae ** +--+ +-- Margaritiferidae **
Pfeiffer et al., 2021 [Publ 4126], fig. 27
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 7 | 1 | 0 | 0 | 0 | 27 |
1
Araujo et al., 2016 [Publ 3746], fig. 2a
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 9 | 1 | 0 | 0 | 0 | 26.5 |
1
Hou et al., 2025 [Publ 4401], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 9 | 1 | 0 | 0 | 0 | 27 |
1
Graf et al., 2014 [Publ 3591], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 13 | 2 | 0 | 0 | 0 | 30 |
+-- Margaritiferidae ** | --+-- Unioninae ** | +-- Parreysiinae **
Araujo et al., 2009 [Publ 2589], fig. 7B
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 10 | 2 | 0 | 0 | 0 | 26.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Freeman et al., 2025 [Publ 4443], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 9 | 1 | 0 | 0 | 0 | 26 |
1
Amaro et al., 2017 [Publ 3791], fig. 3B
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 9 | 2 | 1 | 0 | 0 | 24 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Amaro et al., 2017 [Publ 3791], fig. 3A
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 9 | 2 | 1 | 0 | 0 | 24 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Araujo et al., 2009 [Publ 2589], fig. 7A
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 7 | 2 | 0 | 0 | 0 | 25.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Schilling et al., 2025 [Publ 4737], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 9 | 1 | 0 | 0 | 0 | 25.5 |
1
Khalloufi et al., 2011 [Publ 2748], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 8 | 1 | 0 | 0 | 0 | 24.5 |
1
Roe, 2013 [Publ 3571], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 9 | 1 | 0 | 0 | 0 | 26.5 |
1
Klishko et al., 2019 [Publ 3886], fig. 9
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 8 | 2 | 0 | 0 | 0 | 24 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Morrison et al., 2021 [Publ 4362], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 9 | 1 | 0 | 0 | 0 | 27 |
1
Sonowal et al., 2021 [Publ 4221], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 1 | 6 | 3 | 1 | 26.5 |
+-- Hyrioidea --+ +-- Unionidae **
Hewitt et al., 2019 [Publ 3962], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 11 | 2 | 0 | 0 | 0 | 23.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Huff et al., 2004 [Publ 2697], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 6 | 2 | 2 | 0 | 0 | 23.5 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Schilling et al., 2025 [Publ 4737], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 9 | 1 | 0 | 0 | 0 | 23.5 |
1
Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 2 | 0 | 0 | 0 | 24 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Froufe et al., 2017 [Publ 3802], fig. 7
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 2 | 0 | 0 | 0 | 23 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Bespalaya et al., 2018 [Publ 3811], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 9 | 2 | 0 | 0 | 0 | 25.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Choi et al., 2020 [Publ 4169], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 1 | 0 | 0 | 0 | 25 |
1
Chase et al., 2018 [Publ 3861], fig. 1-M
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 11 | 9 | 2 | 1 | 1 | 1 | 25.5 |
+----- Unionidae ** --+ | +-- Margaritiferidae ** +--+ +-- Hyrioidea
Burzynski & Soroka, 2018 [Publ 3862], fig. 5b
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 1 | 0 | 0 | 0 | 24 |
1
Roe & Lydeard, 1998 [Publ 2693], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 5 | 1 | 0 | 0 | 0 | 24.5 |
1
Huff et al., 2004 [Publ 2697], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 7 | 2 | 1 | 0 | 0 | 23.5 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Vikhrev et al., 2017 [Publ 3810], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 2 | 0 | 0 | 0 | 24.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Huff et al., 2004 [Publ 2697], fig. 5-6
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 7 | 2 | 2 | 0 | 0 | 22.5 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Roe et al., 2002 [Publ 2689], fig. 3a
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 1 | 0 | 0 | 0 | 24.5 |
1
Stodola et al., 2021 [Publ 4266], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 7 | 1 | 0 | 0 | 0 | 26.5 |
1
Lopes-Lima et al., 2021 [Publ 4217], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 1 | 0 | 0 | 0 | 25.5 |
1
Lopes-Lima et al., 2020 [Publ 4080], fig. 13
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 2 | 0 | 0 | 0 | 24.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Keogh & Simons, 2019 [Publ 3935], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 1 | 0 | 0 | 0 | 24.5 |
1
Chase et al., 2018 [Publ 3861], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 2 | 1 | 1 | 1 | 23.5 |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 2 | 0 | 0 | 0 | 23 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Amaro et al., 2017 [Publ 3791], fig. 2-M
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 7 | 1 | 0 | 0 | 0 | 25 |
1
Morrison et al., 2021 [Publ 4362], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 1 | 0 | 0 | 0 | 26 |
1
Araujo et al., 2009 [Publ 2589], fig. 7C
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 2 | 0 | 0 | 0 | 24.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Kano et al., 2019 [Publ 4194], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 2 | 0 | 0 | 0 | 23.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Muanta et al., 2019 [Publ 3921], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 8 | 1 | 0 | 0 | 0 | 26 |
1
Zieritz et al., 2026 [Publ 4812], fig. 4a
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 1 | 0 | 0 | 0 | 25 |
1
Soroka, 2010 [Publ 3404], fig. 3b
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 5 | 1 | 0 | 0 | 0 | 22.5 |
1
Chen et al., 2025 [Publ 4448], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 1 | 0 | 0 | 0 | 25 |
1
Chase et al., 2018 [Publ 3847], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 10 | 8 | 2 | 1 | 1 | 1 | 25 |
+----- Unionidae ** --+ | +-- Margaritiferidae ** +--+ +-- Hyrioidea
Sonowal et al., 2021 [Publ 4221], fig. 3a
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 15 | 7 | 1 | 6 | 3 | 1 | 26.5 |
+-------- Parreysiinae ** | +-------- Gonideinae ** --+ | +-- Hyrioidea | +--+ | | +-- Trigonioidea +--+ | +-- Pleurobemini ** | | +--+-- Lampsilini ** | +-- Unioninae **
Martínez et al., 2026 [Publ 4820], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 1 | 7 | 7 | 1 | 25 |
+-- Etherioidea --+ +-- Unionidae **
Inoue et al., 2013 [Publ 3579], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 2 | 0 | 0 | 0 | 23 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Jeratthitikul et al., 2019 [Publ 3911], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 14 | 1 | 0 | 0 | 0 | 25 |
1
Wu et al., 2024 [Publ 4388], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 1 | 0 | 0 | 0 | 25 |
1
Takeuchi et al., 2015 [Publ 3661], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 2 | 1 | 0 | 0 | 23 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Bolotov et al., 2015 [Publ 3646], fig. 2A
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 6 | 2 | 0 | 0 | 0 | 23 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Sonowal et al., 2021 [Publ 4221], fig. 3b
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 0 | 1 | 6 | 3 | 1 | 22.5 |
+-- Hyrioidea --+ +-- Unionidae **
Kandl et al., 2001 [Publ 3384], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 1 | 0 | 0 | 0 | 23.5 |
1
Nagel et al., 1998 [Publ 1946], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 1 | 0 | 0 | 0 | 23.5 |
1
Lopes-Lima et al., 2020 [Publ 4080], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 2 | 0 | 0 | 0 | 23.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Takeuchi et al., 2015 [Publ 3661], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 2 | 1 | 0 | 0 | 23 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Bolotov et al., 2015 [Publ 3646], fig. 2B
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 2 | 0 | 0 | 0 | 23 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Fonseca et al., 2016 [Publ 3775], fig. 1-M
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 1 | 0 | 0 | 0 | 24 |
1
Soroka, 2010 [Publ 3404], fig. 3a
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 1 | 0 | 0 | 0 | 22.5 |
1
Hewitt et al., 2018 [Publ 3758], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 6 | 2 | 0 | 0 | 0 | 22 |
+-- Margaritiferidae ** --+ +-- Unionidae **
King et al., 1999 [Publ 2699], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 1 | 0 | 0 | 0 | 24.5 |
1
Pfeiffer et al., 2021 [Publ 4126], fig. 25
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 1 | 0 | 0 | 0 | 25 |
1
Bolotov et al., 2022 [Publ 4250], fig. S4
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 6 | 1 | 0 | 0 | 0 | 25 |
1
Zieritz et al., 2012 [Publ 3673], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 4 | 2 | 0 | 0 | 0 | 23 |
+----- Anodontina ** | --+ +-- Cristariina ** | | +--+-- Margaritiferidae ** | +-- Unionini **
Soroka, 2010 [Publ 3404], fig. 5a
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 6 | 1 | 0 | 0 | 0 | 22.5 |
1
Araujo et al., 2009 [Publ 2395], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 6 | 2 | 0 | 0 | 0 | 21.5 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Lopes-Lima et al., 2020 [Publ 4080], fig. 7
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 6 | 2 | 0 | 0 | 0 | 22.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Sheng et al., 2014 [Publ 3704], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 6 | 1 | 0 | 0 | 0 | 22 |
1
Klishko et al., 2018 [Publ 3945], fig. 8
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 5 | 1 | 0 | 0 | 0 | 23.5 |
1
Zieritz et al., 2026 [Publ 4812], fig. 5a
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 8 | 1 | 0 | 0 | 0 | 24 |
1
Lopes-Lima et al., 2020 [Publ 4080], fig. 12
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 6 | 2 | 0 | 0 | 0 | 22.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Lopes-Lima et al., 2020 [Publ 4080], fig. 16
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 7 | 2 | 0 | 0 | 0 | 22.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Inoue et al., 2014 [Publ 3635], fig. 3B
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 3 | 1 | 0 | 0 | 0 | 21.5 |
1
Zhang et al., 2024 [Publ 4397], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 6 | 1 | 0 | 0 | 0 | 24 |
1
Bolotov et al., 2016 [Publ 3732], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 7 | 2 | 0 | 0 | 0 | 23.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Tomilova et al., 2020 [Publ 4097], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 5 | 1 | 0 | 0 | 0 | 22.5 |
1
Marrone et al., 2019 [Publ 3907], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 4 | 1 | 0 | 0 | 0 | 23.5 |
1
Bauer et al., 2021 [Publ 4365], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 5 | 1 | 0 | 0 | 0 | 24 |
1
Soroka, 2010 [Publ 3404], fig. 5b
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 5 | 1 | 0 | 0 | 0 | 22.5 |
1
Hoeh et al., 2002 [Publ 2673], fig. 1-F
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 6 | 1 | 1 | 1 | 1 | 23 |
+-- Hyrioidea --+ +-- Unionidae **
Hoeh et al., 2002 [Publ 2673], fig. 1-M
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 6 | 1 | 1 | 1 | 1 | 23 |
+-- Hyrioidea --+ +-- Unionidae **
Froufe et al., 2015 [Publ 3664], fig. 1-M
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 6 | 1 | 0 | 0 | 0 | 22.5 |
1
Pfeiffer et al., 2021 [Publ 4126], fig. 18
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 7 | 1 | 0 | 0 | 0 | 25 |
1
Kondakov et al., 2018 [Publ 3914], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 6 | 1 | 0 | 0 | 0 | 24 |
1
Hou et al., 2025 [Publ 4445], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 1 | 0 | 0 | 0 | 23 |
1
Serb, 2006 [Publ 2678], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 7 | 1 | 0 | 0 | 0 | 22.5 |
1
Chen et al., 2012 [Publ 3624], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 1 | 0 | 0 | 0 | 21 |
1
Pfeiffer et al., 2018 [Publ 3827], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 3 | 1 | 0 | 0 | 0 | 23 |
1
Källersjö et al., 2005 [Publ 2684], fig. 1A
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 2 | 2 | 0 | 0 | 0 | 21 |
+-- Margaritiferidae ** | --+-- Anodontini ** | +-- Unionini **
Amaro et al., 2017 [Publ 3791], fig. 3A-M
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 2 | 0 | 0 | 0 | 20 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Buhay et al., 2002 [Publ 2661], fig. 2A
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 3 | 1 | 0 | 0 | 0 | 22.5 |
1
Bogan et al., 2023 [Publ 4295], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 2 | 0 | 0 | 0 | 22 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Lopes-Lima et al., 2021 [Publ 4217], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 1 | 0 | 0 | 0 | 22 |
1
Guerra et al., 2017 [Publ 3772], fig. 2a-M
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 4 | 2 | 1 | 1 | 1 | 20.5 |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Zieritz et al., 2024 [Publ 4381], fig. 8A
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 1 | 0 | 0 | 0 | 23 |
1
Buhay et al., 2002 [Publ 2661], fig. 2B
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 1 | 0 | 0 | 0 | 22.5 |
1
Serb & Barnhart, 2008 [Publ 2674], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 10 | 1 | 0 | 0 | 0 | 22.5 |
1
Smith et al., 2023 [Publ 4291], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 1 | 0 | 0 | 0 | 23 |
1
Serb, 2006 [Publ 2678], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 8 | 1 | 0 | 0 | 0 | 22.5 |
1
Goncalves et al., 2022 [Publ 4253], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 6 | 1 | 0 | 0 | 0 | 23 |
1
Soroka, 2010 [Publ 3404], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 4 | 1 | 0 | 0 | 0 | 21.5 |
1
Jones & Neves, 2010 [Publ 2646], fig. 16
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 1 | 0 | 0 | 0 | 23 |
1
Froufe et al., 2016 [Publ 3727], fig. 3_4
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 3 | 1 | 0 | 0 | 0 | 23 |
1
Soroka, 2010 [Publ 3404], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 1 | 0 | 0 | 0 | 21.5 |
1
Jones & Neves, 2010 [Publ 2646], fig. 17
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 1 | 1 | 0 | 0 | 0 | 23 |
1
Soroka & Burzynski, 2018 [Publ 3762], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 1 | 0 | 0 | 0 | 22 |
1
Hoeh et al., 1995 [Publ 3391], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 4 | 1 | 0 | 0 | 0 | 22.5 |
1
Amaro et al., 2017 [Publ 3791], fig. 3B-M
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 2 | 0 | 0 | 0 | 20 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Källersjö et al., 2005 [Publ 2684], fig. 1B-F
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 4 | 2 | 0 | 0 | 0 | 21 |
+-- Margaritiferidae ** | --+-- Anodontini ** | +-- Unionini **
Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 1 | 0 | 0 | 0 | 22 |
1
Serb, 2006 [Publ 2678], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 8 | 1 | 0 | 0 | 0 | 22.5 |
1
Huff et al., 2004 [Publ 2697], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 3 | 2 | 1 | 0 | 0 | 21.5 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Lopes-Lima et al., 2020 [Publ 4080], fig. 8
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 2 | 0 | 0 | 0 | 21.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Guerra et al., 2017 [Publ 3772], fig. 2b-M
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 4 | 2 | 1 | 1 | 1 | 20.5 |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Serb & Barnhart, 2008 [Publ 2674], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 9 | 1 | 0 | 0 | 0 | 22.5 |
1
Pfeiffer et al., 2021 [Publ 4126], fig. 30
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 1 | 0 | 0 | 0 | 23 |
1
Lopes-Lima et al., 2020 [Publ 4080], fig. 9
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 2 | 0 | 0 | 0 | 20.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Soroka, 2010 [Publ 3404], fig. 8b
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 1 | 0 | 0 | 0 | 20.5 |
1
Kandl et al., 2001 [Publ 3384], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 1 | 0 | 0 | 0 | 21 |
1
Choi et al., 2020 [Publ 4169], fig. 3a
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 5 | 1 | 0 | 0 | 0 | 21.5 |
1
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 2 | 0 | 0 | 0 | 19 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Keogh & Simons, 2019 [Publ 3935], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 6 | 1 | 0 | 0 | 0 | 22.5 |
1
Riccardi et al., 2020 [Publ 4089], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 1 | 0 | 0 | 0 | 22 |
1
Huang et al., 2013 [Publ 3629], fig. 3-M
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 1 | 0 | 0 | 0 | 20.5 |
1
Bogatov et al., 2018 [Publ 3859], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 2 | 0 | 0 | 0 | 19 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Freeman et al., 2025 [Publ 4443], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 5 | 1 | 0 | 0 | 0 | 22 |
1
Lopes-Lima et al., 2020 [Publ 4080], fig. 17
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 5 | 2 | 0 | 0 | 0 | 19.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Bogan et al., 2023 [Publ 4295], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 2 | 0 | 0 | 0 | 21 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Soroka, 2010 [Publ 3404], fig. 3a-M
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 1 | 0 | 0 | 0 | 20.5 |
1
Moyer & Díaz-Ferguson, 2012 [Publ 3375], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 5 | 1 | 0 | 0 | 0 | 20.5 |
1
Lopes-Lima et al., 2020 [Publ 4080], fig. 14
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 2 | 0 | 0 | 0 | 20.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Klishko et al., 2017 [Publ 3946], fig. 8
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 1 | 0 | 0 | 0 | 21.5 |
1
Wei et al., 2016 [Publ 3665], fig. 0
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Zieritz et al., 2016 [Publ 3804], fig. S2-1
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 3 | 1 | 0 | 0 | 0 | 22 |
1
Prie et al., 2012 [Publ 3487], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 2 | 0 | 0 | 0 | 20.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Nagel et al., 1998 [Publ 1946], fig. 9
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 2 | 0 | 0 | 0 | 18 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-M
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 1 | 0 | 0 | 0 | 21.5 |
1
Dai et al., 2024 [Publ 4358], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 1 | 0 | 0 | 0 | 22 |
1
King et al., 1999 [Publ 2699], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 3 | 1 | 0 | 0 | 0 | 22 |
1
Bashê, 2022 [Publ 4265], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 5 | 2 | 0 | 0 | 0 | 20.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Soroka, 2010 [Publ 3404], fig. 8a
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 1 | 0 | 0 | 0 | 20.5 |
1
Wen et al., 2017 [Publ 3795], fig. 3-M
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 1 | 0 | 0 | 0 | 20.5 |
1
Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 1 | 0 | 0 | 0 | 22 |
1
Soroka, 2010 [Publ 3404], fig. 3b-M
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 1 | 0 | 0 | 0 | 20.5 |
1
Bogatov et al., 2018 [Publ 3859], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 2 | 0 | 0 | 0 | 19 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Cyr et al., 2007 [Publ 2745], fig. 2A
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 20 |
1
Saxon et al., 2023 [Publ 4338], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 21 |
1
Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5_7
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 19.5 |
1
Mabuchi et al., 2021 [Publ 4233], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 21 |
1
Cyr et al., 2007 [Publ 2745], fig. 2B
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 4 | 1 | 0 | 0 | 0 | 21.5 |
1
Jones et al., 2006 [Publ 2681], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 4 | 1 | 0 | 0 | 0 | 21 |
1
Krebs, 2004 [Publ 2692], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 20 |
1
Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 2 | 0 | 0 | 0 | 19.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Krebs et al., 2013 [Publ 3515], fig. 2A
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 20.5 |
1
Zanatta & Harris, 2013 [Publ 3530], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 19 |
1
Soroka, 2005 [Publ 2726], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 20.5 |
1
Konopleva et al., 2017 [Publ 3807], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 21 |
1
Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 11 | 9 | 2 | 6 | 6 | 1 | 23 |
+-- Margaritiferidae ** | --+-- Unionidae ** | +-- Hyrioidea
Buhay et al., 2002 [Publ 2661], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 21.5 |
1
Lane et al., 2016 [Publ 3735], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 1 | 1 | 0 | 0 | 0 | 22 |
1
Cyr et al., 2007 [Publ 2745], fig. 2C
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 4 | 1 | 0 | 0 | 0 | 23.5 |
1
Soroka, 2010 [Publ 3404], fig. 9a
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 21 |
1
Nagel et al., 1996 [Publ 3400], fig. 351
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 4 | 1 | 0 | 0 | 0 | 21 |
1
Mulvey et al., 1997 [Publ 3390], fig. 1_3
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 20 |
1
Soroka, 2010 [Publ 3404], fig. 2-M
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 19.5 |
1
Nagel et al., 1998 [Publ 1946], fig. 7
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 4 | 1 | 0 | 0 | 0 | 21 |
1
Bolotov et al., 2022 [Publ 4250], fig. S3
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 21 |
1
Lopes-Lima et al., 2021 [Publ 4217], fig. 14
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 4 | 1 | 0 | 0 | 0 | 21 |
1
Burzynski & Soroka, 2018 [Publ 3862], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 19.5 |
1
Patterson et al., 2021 [Publ 4222], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Jiang et al., 2010 [Publ 3438], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 1 | 1 | 0 | 0 | 0 | 18.5 |
1
Froufe et al., 2017 [Publ 3802], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Gangloff et al., 2013 [Publ 3551], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Soroka, 2008 [Publ 3695], fig. 1-M
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 18.5 |
1
Cyr et al., 2007 [Publ 2745], fig. 2D
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20.5 |
1
Nagel et al., 1998 [Publ 1946], fig. 8
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 1 | 1 | 0 | 0 | 0 | 19.5 |
1
Pfeiffer et al., 2018 [Publ 3835], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Lopes-Lima et al., 2021 [Publ 4217], fig. 11
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Marshall et al., 2014 [Publ 3593], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 14 | 12 | 2 | 10 | 10 | 1 | 28 |
+-- Margaritiferidae **
+--+
| +-- Unioninae **
--+
| +-- Ambleminae **
+--+
+-- Hyrioidea
Bulakhova et al., 2023 [Publ 4325], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Combosch et al., 2017 [Publ 3806], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 10 | 8 | 2 | 11 | 9 | 2 | 25.5 |
+-------- Hyrioidea --+ | +----- Unionidae ** +--+ | +-- Margaritiferidae ** +--+ +-- Etherioidea
Soroka, 2010 [Publ 3404], fig. 9c
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 3 | 1 | 0 | 0 | 0 | 20 |
1
Zieritz et al., 2026 [Publ 4812], fig. 7a
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 1 | 1 | 0 | 0 | 0 | 20 |
1
Giribet & Distel, 2003 [Publ 3670], fig. 3f5
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 1 | 2 | 2 | 0 | 0 | 16 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Mezhzherin et al., 2014 [Publ 3705], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Soroka, 2010 [Publ 3404], fig. 9b
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Sayenko et al., 2017 [Publ 3760], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Machordom et al., 2003 [Publ 2688], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 2 | 0 | 0 | 0 | 18.5 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Curole & Kocher, 2002 [Publ 2712], fig. 1b-M
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 18.5 |
1
Combosch et al., 2017 [Publ 3806], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 2 | 11 | 9 | 2 | 17.5 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Curole & Kocher, 2002 [Publ 2712], fig. 1a-M
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 18.5 |
1
Mezhzherin et al., 2014 [Publ 3705], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Krebs, 2004 [Publ 2692], fig. 2-M
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 19 |
1
Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 1 | 1 | 0 | 0 | 0 | 18.5 |
1
Klishko et al., 2016 [Publ 3733], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Johnson et al., 2023 [Publ 4323], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Zhang et al., 2013 [Publ 4214], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 2 | 0 | 0 | 0 | 18.5 |
+-------- Cristaria ** --+ | +----- Sinanodonta ** +--+ | +-- Margaritiferidae ** +--+ +-- Gonideinae **
Soroka, 2008 [Publ 3695], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 18.5 |
1
Klishko et al., 2014 [Publ 3666], fig. 8
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Curole & Kocher, 2002 [Publ 2712], fig. 1-F
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 2 | 0 | 0 | 0 | 18 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Hinzmann et al., 2013 [Publ 3541], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 19 |
1
Hoeh et al., 1996 [Publ 2694], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 1 | 0 | 0 | 17.5 |
+----- Unionidae ** --+ | +-- None__1 +--+ +-- None__2
Hoeh et al., 1998 [Publ 2695], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 2 | 2 | 1 | 1 | 18.5 |
+-------- Etherioidea --+ | +----- Unioninae ** +--+ | +-- Margaritiferidae ** +--+ +-- Ambleminae **
Soroka, 2010 [Publ 3404], fig. 8b-M
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 17.5 |
1
Mock et al., 2013 [Publ 3598], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 18.5 |
1
Froufe et al., 2014 [Publ 3592], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 19 |
1
Kilikowska et al., 2020 [Publ 4335], fig. 4a
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 18.5 |
1
Hoeh et al., 1998 [Publ 2695], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 2 | 2 | 1 | 1 | 18.5 |
+-------- Etherioidea --+ | +----- Ambleminae ** +--+ | +-- Margaritiferidae ** +--+ +-- Unioninae **
Metzger et al., 2018 [Publ 3952], fig. 1C
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 18 |
1
Imsong & Murali, 2023 [Publ 4370], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 18.5 |
1
Xia et al., 2016 [Publ 3712], fig. 7B
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 19 |
1
Zieritz et al., 2024 [Publ 4381], fig. 6A
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 19 |
1
Smith & Johnson, 2020 [Publ 4229], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 19 |
1
Soroka, 2010 [Publ 3404], fig. 8a-M
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 17.5 |
1
Peng et al., 2012 [Publ 3689], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 19 |
1
Lopes-Lima et al., 2016 [Publ 3757], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 19 |
1
Shirai et al., 2010 [Publ 3353], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 19 |
1
Machordom et al., 2003 [Publ 2688], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 2 | 0 | 0 | 0 | 17.5 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Campbell, 2000 [Publ 2724], fig. 1-5
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 2 | 0 | 0 | 0 | 16 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wang et al., 2016 [Publ 3714], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 18.5 |
1
Krebs et al., 2019 [Publ 4092], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 18.5 |
1
Taylor et al., 2007 [Publ 2696], fig. 1-3
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 2 | 1 | 0 | 0 | 16 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Välilä et al., 2017 [Publ 3793], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 2 | 0 | 0 | 0 | 17.5 |
+-- Unionidae ** --+ +-- Margaritiferidae **
González & Giribet, 2015 [Publ 3632], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 4 | 0 | 2 | 4 | 2 | 2 | 19.5 |
+-- Etherioidea
+--+
| +-- Margaritiferidae **
--+
| +-- Unionidae **
+--+
+-- Hyrioidea
Mock et al., 2004 [Publ 2686], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 0 | 0 | 0 | -- |
1
González & Giribet, 2015 [Publ 3632], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 4 | 2 | 2 | -- |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Unionidae ** +--+ +-- Margaritiferidae **
Bieler et al., 2014 [Publ 3622], fig. 26_27_29_30
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 3 | 0 | 2 | 3 | 2 | 2 | 18 |
+-------- Hyrioidea --+ | +----- Unionidae ** +--+ | +-- Etherioidea +--+ +-- Margaritiferidae **
Soroka, 2010 [Publ 3404], fig. 4-M
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 0 | 0 | 0 | -- |
1
Klunzinger et al., 2021 [Publ 4086], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 4 | 4 | 1 | -- |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Lemer et al., 2019 [Publ 4093], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 4 | 2 | 1 | -- |
+----- Etherioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
González & Giribet, 2015 [Publ 3632], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 3 | 0 | 2 | 4 | 2 | 2 | 18 |
+-------- Hyrioidea --+ | +----- Unionidae ** +--+ | +-- Etherioidea +--+ +-- Margaritiferidae **
Soroka, 2010 [Publ 3404], fig. 5a-M
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 0 | 0 | 0 | -- |
1
Mock et al., 2004 [Publ 2686], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 0 | 0 | 0 | -- |
1
Soroka, 2010 [Publ 3404], fig. 5b-M
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 0 | 0 | 0 | -- |
1
González & Giribet, 2015 [Publ 3632], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 4 | 0 | 2 | 4 | 2 | 2 | 18.5 |
+----- Unionidae ** | --+----- Hyrioidea | | +-- Etherioidea +--+ +-- Margaritiferidae **
Plazzi & Passamonti, 2010 [Publ 3354], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 0 | 0 | 0 | -- |
1
Graf et al., 2015 [Publ 3619], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 18 | 17 | 2 | -- |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Unionidae ** +--+ +-- Margaritiferidae **
Xue et al., 2016 [Publ 3659], fig. 1-M
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 0 | 0 | 0 | -- |
1
Bieler et al., 2014 [Publ 3622], fig. 28
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 4 | 0 | 2 | 3 | 2 | 2 | 19.5 |
+-- Etherioidea
+--+
| +-- Margaritiferidae **
--+
| +-- Unionidae **
+--+
+-- Hyrioidea
Baker et al., 2003 [Publ 2690], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 7 | 7 | 1 | -- |
+-- Unionidae ** --+ +-- Hyrioidea
Baker et al., 2004 [Publ 2685], fig. 2-3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 8 | 8 | 1 | -- |
+-- Unionidae ** --+ +-- Hyrioidea
