MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Thu 19 March 2026
superfamily UNIONOIDEA Rafinesque, 1820: cladomics
Cladomics pages are experimental — even more so than the rest of this site!
The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.
The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.
Following the summary, the available published clades representing the taxon are listed.
For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!
Cladomics Summary
number of species = 822 (1123 total)
number of subtaxa (rank: family) = 2
analyzed in 585 cladograms in 324 publications
maximum number of terminals in an analysis = 581 (mean = 23.81, median = 14)
maximum number of subtaxa (family) = 2 (mean = 1.45, median = 1)
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 583
maximum number of included terminals = 581, (mean = 23.89 median = 14)
maximum number of subtaxa (family) = 2 (mean = 1.46, median = 1)
proportion of cladograms that support basic monophyly = 0.94
Rigorous Monophyly
number of cladograms that test rigorous monophyly = 94
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (family) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 581 (mean = 43.86, median = 23)
maximum number of subtaxa (family) = 2 (mean = 2, median = 2)
proportion of cladograms that support rigorous monophyly = 0.69
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Cladograms
Graf, 2000 [Publ 2647], fig. 1
** Top Monophyly Cladogram **
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 16 | 14 | 2 | 10 | 9 | 2 | 0 |
+-------------- Margaritiferidae ** | | +-------- Unionini ** --+ +--+ | | | +----- Pleurobemini ** | | +--+ | | | +-- Anodontini ** +--+ +--+ | +-- Lampsilini ** | | +-------- Parreysiini ** +--+ | +----- Coelaturini ** +--+ | +-- Hyrioidea +--+ +-- Etherioidea
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14
** Top Sister-Group Cladogram **
| tT | bi | riS | oT | roT | roS | iD | |
| 581 | 582 | 651 | 2 | 50 | 48 | 2 | 701.5 |
+----------------------------- Iridininae
|
+-----------------------------------------------------------------+ +-------------------------- Aspathariinae
| | |
| +--+ +----------------------- Etheria
| | |
| +--+ +-------------------- Anodontites_trapesialis
| | |
| +--+ +----------------- Mulleria
| | |
| +--+ +-------------- Lamproscapha
| | |
| | | +----- Monocondylaea
| +--+ +-----+
| | | | +-- Tamsiella_schroeteriana__2
| | | +--+
| +--+ +-- Tamsiella_tamsiana
| |
| | +-------- Mycetopodinae
| | |
| +--+ +-- Anodontites_guanarensis
| | +--+
| | | +-- Fossula
| +--+
| | +-- Anodontites_trigona
| +--+
| +-- Tamsiella_schroeteriana__1
|
| +-------------------------------------------------------------------------------------------- Hyrioidea
| |
| | +-------------------------------- Lamellidens **
| | |
| | +-----+ +-- Balwantia **
--+ | | | +--------------------------+
| | | | | +-- Trapezidens **
| | | +--+
| | | | +-------------------------- Leoparreysiini **
| | | | |
| | | +--+ +----------------------- Pseudomulleria
| | | | |
| | | | | +-- Parreysiini **
| | | +--+ +-----------------+
| | | | | +-- Coelaturini **
| | | | |
| | | +--+ +-- Pseudoscabies **
| | | | +--------------+
| | | | | +-- Indonaia **
| | | +--+
| | | | +-------------- Indochinella **
| | | | |
| | | +--+ +----------- Radiatula **
| | +--------------------------------------------------+ | |
| | | | +--+ +-- Songkhlanaia_songkhramensis__2 **
| | | | | +-----+
| | | | | | +-- Unionetta_fabagina__2 **
| | | | +--+
| | | | | +----- Harmandia **
| | | | +--+
| | | | | +-- Unionetta_fabagina__1 **
| | | | +--+
| | | | +-- Scabies **
| | | |
| | | | +----- Amuranodonta **
| | | | +-----------------------------+
| | | | | | +-- Anemina **
+--+ | | | +--+
| | | | +-- Buldowskia **
| | | |
| | | | +-- Beringiana **
| | | | +-----+
| | | | | +-- Pletholophus **
| | +--+ +-----------------------+
| | | | | +----- Mimo **
| | | | +--+
| | | | | +-- Acudonta **
| | | | +--+
| | | | +-- Sinanodonta **
| | | |
| | | | +----------------------------- Cristaria **
| | +--+ |
| | | | +----------------- Lepidodesmini **
| | | | |
| | | | | +----------- Lanceolariini **
| | | | | +--+
| | | | +--------+ | | +-------- Pseudoacuticosta_retiaria__2 **
| | | | | | | +--+
| | | | | | | | +----- Acuticosta_chinensis__2 **
| | +--+ | | | +--+
| | | | +--+ | +-- Acuticosta_chinensis__1 **
| | | | | +--+
| | | | | +-- Pseudoacuticosta_retiaria__1 **
| | | | |
| | | | | +----------- Globunio **
| | | | +--+
| | | | | +-------- Aculamprotulina **
| | | | +--+
| | +--+ | +----- Middendorffinaiina **
| | | +--+
| | | | +-- Unionina **
| | | +--+
| | | +-- Oxynaiina **
| | |
| | | +-------- Pseudodontoideus **
+--+ | +--------------+
| | | | +----- Lasmigona **
| | | +--+
| | | | +-- Pterosyna **
| | | +--+
| +--+ +-- Arcidens **
| |
| | +-------------------- Anodontina **
| | |
| | | +-------------- Utterbackia **
| | | +--+
| +--+ | | +----------- Pyganodon_grandis **
| | | +--+
| | | | +-------- Pyganodon_lacustris **
| | | +--+
| | | | +----- Pyganodon_cataracta__2 **
| +--+ +--+
| | | +-- Pyganodon_cataracta__1 **
| | +--+
| | +-- Potamilus_fragilis__3 **
| |
| | +-------------- Utterbackiana **
| | |
| +--+ +----------- Simpsonaias **
| | |
| | | +-- Alasminota **
| +--+ +-----+
| | | +-- Alasmidonta **
| | |
| +--+ +-- Prolasmidonta **
| | +--+
| | | +-- Platynaias **
| +--+
| | +-- Anodontoides **
| +--+
| +-- Strophitus **
|
| +-------------------------------------------------------------------------------------- Rectidentini **
| |
| | +----------------------------------------------------------------------------------- Margaritiferidae **
| | |
| | | +-------------------- Chamberlainiini **
| | | |
| | | +--+ +----------------- Parvasolenaia **
| | | | | |
| | | | +--+ +-------------- Pseudopostolata **
| | | | | |
| | | | +--+ +----------- Koreosolenaia **
| | | | | |
| | | | +--+ +-- Cosmopseudodon **
| | | | | +-----+
| | | | | | +-- Guiunio **
+--+ | | +--+
| | +--------+ | +----- Ptychorhynchus **
| | | | +--+
| | | | | +-- Obovalis **
| | | | +--+
| | | | +-- Postolata **
| | | |
| | | | +-- Caudiculatus **
| | | | +--------------+
| | | | | +-- Lamprotulini **
| | | | |
| | | +-----+ +-------------- Sinosolenaia_carinata **
| | | | |
| | | | | +----- Gonidea **
| | | +--+ +-----+
+--+ +-----------------------------------------------+ | | | +-- Microcondylaea **
| | | | | +--+
| | | +--+ +-- Leguminaia **
| | | |
| | | | +-------- Inversidens **
| | | +--+
| | | | +----- Sinosolenaia_recognita **
| | | +--+
| | | | +-- Sinosolenaia_iridinea **
| | | +--+
| | | +-- Sinosolenaia_oleivora **
| | |
| | | +----------------------------- Indopseudodontina **
| | | |
| | | | +-- Lannanaia **
| | +--+ +-----------------------+
| | | | +-- Pilsbryoconcha **
| | | |
| | | | +----- Songkhlanaia_tamodienica **
| | +--+ +-----------------+
| | | | | +-- Songkhlanaia_moreleti **
| | | | +--+
| | | | +-- Songkhlanaia_songkhramensis__1 **
| | | |
| | +--+ +-- Nyeinchanconcha **
| | | +-----------------+
| | | | +-- Sundadontina **
| | | |
+--+ | | +-- Isannaia **
| +--+ +--+
| | | +-- Namkongnaia **
| | +-----------+
| | | | +-- Thaiconcha **
| | | +--+
| +--+ +-- Bineurus **
| |
| | +-------------- Pseudodon_walpolei **
| | |
| +--+ +----------- Pseudodon_vagulus **
| | |
| | | +----- Pseudodon_vondembuschianus **
| +--+ +--+
| | | | +-- Pseudodon_mekongi **
| | | +--+
| +--+ +-- Schepmaniini **
| |
| | +----- Pseudodon_lenyanensis__2 **
| +--+
| | +-- Pseudodon_inoscularis **
| +--+
| +-- Pseudodon_lenyanensis__1 **
|
| +-- Quadrulini **
| +--------------------------------------------------------------------------+
| | +-- Pleurobemini **
| |
| | +----- Amblema **
+--+ +--------------------------------------------------------------------+
| | | +-- Nephronaias **
| | +--+
| | +-- Psoronaias **
| |
+--+ +----------------------------------------------------------------------- Reginaia **
| |
| | +----- Popenaias **
| | +--------------------------------------------------------------+
| | | | +-- Cyrtonaias **
+--+ | +--+
| | +-- Glebula **
| |
| | +----------------------------------------------------------------- Paetulunio **
| | |
+--+ | +----------------- Toxolasma_parvum **
| | |
| | | +----- Toxolasma_lividum__2 **
| | | +--------+
| | +--------------------------------------------+ | | +-- Toxolasma_cylindrellus **
| | | | | +--+
| | | | | +-- Toxolasma_lividum__1 **
| | | | |
+--+ | +--+ +-------- Toxolasma_texasiense__2 **
| | | +--+
| | | | | +----- Toxolasma_glans **
| | | | +--+
| | | | | +-- Toxolasma_paulum **
| | +--+ +--+
| | | +-- Toxolasma_texasiense__1 **
| | |
| | | +-------- Toxolasma__4 **
| | +--+
+--+ | +----- Toxolasma_granulatum **
| +--+
| | +-- Toxolasma__2 **
| +--+
| +-- Toxolasma__3 **
|
| +----- Toxolasma_pullus **
| +-----------------------------------------------------+
| | | +-- Toxolasma_germanum **
| | +--+
| | +-- Toxolasma__1 **
| |
| | +-- Obliquaria **
| | +-----------------------------------------------------+
| | | +-- Plectomerus **
+--+ |
| | +-------- Potamilus_inflatus **
| | +--------+
| | | | +----- Potamilus_ohiensis **
| | | +--+
| | | | +-- Potamilus_amphichaenus **
| | | +--+
| | +-----------------------------------+ +-- Potamilus_streckersoni **
| | | |
+--+ | | +----- Potamilus_fragilis__2 **
| | | +--------+
| | | | | +-- Potamilus_leptodon **
| | | | +--+
| | +--+ +-- Potamilus_fragilis__1 **
| | |
| | | +----------- Potamilus_metnecktayi **
| | | |
| | +--+ +-------- Potamilus_alatus__3 **
| | | |
| | +--+ +----- Potamilus_alatus__2 **
+--+ | |
| +--+----- Potamilus_purpuratus__2 **
| |
| | +-- Potamilus_alatus__1 **
| +--+
| +-- Potamilus_purpuratus__1 **
|
| +-- Villosa_villosa **
| +-----------------------------------------------+
| | +-- Atlanticoncha **
| |
| | +-- Cambarunio_nebulosus **
| | +--------------------------------------------+
+--+ | +-- Ligumia **
| |
| | +-------- Leaunio_umbrans **
| | +-----------------------------------+
| | | | +----- Cambarunio_taeniatus **
+--+ | +--+
| | | +-- Cambarunio_iris **
| | +--+
| | +-- Cambarunio_simus **
| |
| | +----------------------------------------- Hamiota **
| | |
+--+ | +----------------------- Pachynaias **
| | |
| | +--------------+ +-------------------- Sagittunio **
| | | | |
| | | | | +----- Ellipsaria **
| | | +--+ +-----------+
| | | | | | +-- Potamilus_capax **
| | | | | +--+
| | | | | +-- Truncilla **
+--+ | +--+
| | | +----- Medionidus_acutissimus **
| | | +--------+
| | | | | +-- Lemiox **
| | | | +--+
| | +--+ +-- Medionidus_penicillatus **
| | |
| | | +----------- Ptychobranchus **
| | | |
| | +--+ +-- Cyprogenia_aberti__2 **
| | | +-----+
+--+ | | +-- Dromus **
| +--+
| | +----- Cyprogenia_stegaria__2 **
| +--+
| | +-- Cyprogenia_aberti__1 **
| +--+
| +-- Cyprogenia_stegaria__1 **
|
| +-- Villosa_delumbis **
| +--------------------------------+
| | +-- Villosa_vibex **
| |
| | +-------------- Obovaria_olivaria **
| | |
| | +-----------------+ +-- Venustaconcha **
| | | | +--------+
| | | | | +-- Epioblasma **
| | | +--+
| | | | +-------- Obovaria_retusa **
+--+ | +--+
| | | +----- Obovaria_choctawensis **
| | +--+
| | | +-- Obovaria_unicolor **
| | +--+
| | +-- Obovaria_subrotunda **
| |
| | +-- Mesonaias **
| | +-----------------------+
| | | +-- Obovaria_arkansasensis **
| | |
+--+ +--+ +-- Lampsilis_splendida **
| | | +--------------------+
| | | | +-- Lampsilis_bracteata **
| | | |
| | +--+ +-- Lampsilis_fasciola **
| | | +-----------------+
| | | | +-- Ortmanniana_pectorosa **
| | +--+
| | | +----------------- Lampsilis_ornata **
| | | |
| | +--+ +-------------- Lampsilis_ovata__2 **
| | | |
| | +--+ +----------- Lampsilis_dolabraeformis **
| | | |
| | | | +----- Lampsilis_cardium__2 **
| | +--+ +--+
| | | | | +-- Lampsilis_cardium__1 **
+--+ | | +--+
| +--+ +-- Lampsilis_satura **
| |
| | +----- Lampsilis_cariosa__2 **
| +--+
| | +-- Lampsilis_ovata__1 **
| +--+
| +-- Lampsilis_cariosa__1 **
|
| +----------- Ortmanniana_ligamentina__2 **
| +--------------+
| | | +-------- Lampsilis_abrupta **
| | +--+
| | | +----- Lampsilis_higginsii__2 **
| | +--+
| | | +-- Ortmanniana_ligamentina__1 **
| | +--+
+--+ +-- Lampsilis_higginsii__1 **
|
| +----- Medionidus_conradicus **
| +-----------------+
| | | +-- Leaunio_lienosus **
| | +--+
| | +-- Leaunio_vanuxemensis **
| |
+--+ +-- Lampsilis_floridensis **
| +-----------------+
| | +-- Lampsilis_teres **
| |
| | +-- Lampsilis_siliquoidea__1 **
+--+ |
| +-- Lampsilis_siliquoidea__2 **
| +--------------+
| | +-- Lampsilis_powellii__1 **
| | |
+--+ +-- Lampsilis_powellii__2 **
|
| +-------------- Lampsilis_sietmani **
| |
+--+ +-- Lampsilis_straminea__2 **
| +--------+
| | +-- Lampsilis_virescens **
+--+
| +-------- Lampsilis_hydiana__2 **
+--+
| +----- Lampsilis_hydiana__1 **
+--+
| +-- Lampsilis_bergmanni **
+--+
+-- Lampsilis_straminea__1 **
Huang et al., 2019 [Publ 3843], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 174 | 174 | 154 | 2 | 26 | 24 | 2 | 190 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Huang et al., 2019 [Publ 3843], fig. S4
| tT | bi | riS | oT | roT | roS | iD | |
| 174 | 174 | 110 | 2 | 26 | 24 | 2 | 188 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Huang et al., 2019 [Publ 3843], fig. S3
| tT | bi | riS | oT | roT | roS | iD | |
| 172 | 172 | 169 | 2 | 26 | 24 | 2 | 186 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5
| tT | bi | riS | oT | roT | roS | iD | |
| 128 | 128 | 128 | 2 | 13 | 12 | 2 | 143.5 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Bolotov et al., 2017 [Publ 3764], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 124 | 124 | 71 | 1 | 0 | 0 | 0 | 141.5 |
1
Bolotov et al., 2017 [Publ 3761], fig. S2
| tT | bi | riS | oT | roT | roS | iD | |
| 123 | 136 | 122 | 2 | 15 | 13 | 2 | 149 |
+-- Chamberlainia **
+--------------------+
| +-- Margaritiferidae **
|
| +-- Etherioidea
--+--------------------+
| +-- Hyrioidea
|
| +-------------------- Parreysiinae **
+--+
| +----------------- Unioninae **
+--+
| +-------------- Rectidentini **
+--+
| +----------- Ambleminae **
+--+
| +-------- Pseudodontini **
+--+
| +----- Gonideini **
+--+
| +-- Sinohyriopsis **
+--+
+-- Lamprotulini **
Bolotov et al., 2017 [Publ 3761], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 123 | 136 | 136 | 2 | 15 | 13 | 2 | 150 |
+-------- Margaritiferidae ** --+ | +----- Unionidae ** +--+ | +-- Etherioidea +--+ +-- Hyrioidea
Bolotov et al., 2020 [Publ 4100], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 117 | 117 | 99 | 2 | 0 | 0 | 0 | 131 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
| tT | bi | riS | oT | roT | roS | iD | |
| 113 | 118 | 100 | 2 | 9 | 7 | 2 | 132.5 |
+-------- Hyrioidea --+ | +----- Margaritiferidae ** +--+ | +-- Etherioidea +--+ +-- Unionidae **
Bolotov et al., 2020 [Publ 4100], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 111 | 111 | 99 | 1 | 0 | 0 | 0 | 126.5 |
1
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c
| tT | bi | riS | oT | roT | roS | iD | |
| 105 | 105 | 63 | 2 | 0 | 0 | 0 | 202.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Campbell et al., 2005 [Publ 2660], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 102 | 102 | 103 | 2 | 0 | 0 | 0 | 125 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Campbell et al., 2005 [Publ 2660], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 102 | 102 | 92 | 2 | 0 | 0 | 0 | 124 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wu et al., 2026 [Publ 4811], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 95 | 95 | 95 | 2 | 0 | 0 | 0 | 108.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt
| tT | bi | riS | oT | roT | roS | iD | |
| 92 | 92 | 87 | 2 | 4 | 3 | 2 | 105.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa
| tT | bi | riS | oT | roT | roS | iD | |
| 92 | 92 | 90 | 2 | 4 | 3 | 2 | 105.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Wu et al., 2024 [Publ 4350], fig. 8a_S1
| tT | bi | riS | oT | roT | roS | iD | |
| 88 | 88 | 85 | 2 | 4 | 3 | 2 | 101.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Wu et al., 2024 [Publ 4350], fig. 8b_S2
| tT | bi | riS | oT | roT | roS | iD | |
| 88 | 88 | 84 | 2 | 4 | 3 | 2 | 101.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Dai et al., 2025 [Publ 4440], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 85 | 85 | 83 | 2 | 3 | 3 | 2 | 97.5 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Chen et al., 2025 [Publ 4448], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 83 | 84 | 82 | 2 | 3 | 3 | 2 | 97.5 |
+-------- Etherioidea --+ | +----- Unionidae ** +--+ | +-- Hyrioidea +--+ +-- Margaritiferidae **
Konopleva et al., 2023 [Publ 4262], fig. 8
| tT | bi | riS | oT | roT | roS | iD | |
| 82 | 82 | 79 | 1 | 0 | 0 | 0 | 98 |
1
Bolotov et al., 2018 [Publ 3826], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 82 | 82 | 50 | 1 | 0 | 0 | 0 | 96.5 |
1
Wu et al., 2021 [Publ 4366], fig. 2_I
| tT | bi | riS | oT | roT | roS | iD | |
| 81 | 81 | 79 | 2 | 10 | 8 | 2 | 92.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Wu et al., 2021 [Publ 4366], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 81 | 81 | 79 | 2 | 10 | 8 | 2 | 92.5 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2
| tT | bi | riS | oT | roT | roS | iD | |
| 76 | 76 | 74 | 2 | 0 | 0 | 0 | 88.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1
| tT | bi | riS | oT | roT | roS | iD | |
| 76 | 76 | 73 | 2 | 0 | 0 | 0 | 88.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Hou et al., 2024 [Publ 4400], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 76 | 76 | 74 | 2 | 0 | 0 | 0 | 88 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2
| tT | bi | riS | oT | roT | roS | iD | |
| 73 | 73 | 63 | 1 | 0 | 0 | 0 | 86 |
1
Campbell & Lydeard, 2012 [Publ 3309], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 73 | 73 | 67 | 1 | 0 | 0 | 0 | 86 |
1
Wu et al., 2022 [Publ 4238], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 71 | 71 | 69 | 2 | 3 | 3 | 2 | 83.5 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
| tT | bi | riS | oT | roT | roS | iD | |
| 71 | 77 | 71 | 2 | 9 | 8 | 2 | 88 |
+-------- Hyrioidea --+ | +----- Margaritiferidae ** +--+ | +-- Etherioidea +--+ +-- Unionidae **
Wu et al., 2021 [Publ 4366], fig. 3-BI
| tT | bi | riS | oT | roT | roS | iD | |
| 70 | 70 | 68 | 2 | 4 | 3 | 2 | 82.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Wu et al., 2021 [Publ 4366], fig. 3-ML
| tT | bi | riS | oT | roT | roS | iD | |
| 70 | 70 | 68 | 2 | 4 | 3 | 2 | 82.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Wu et al., 2021 [Publ 4366], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 65 | 65 | 63 | 2 | 4 | 3 | 2 | 78 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Wang et al., 2025 [Publ 4488], fig. 3B
| tT | bi | riS | oT | roT | roS | iD | |
| 65 | 65 | 63 | 2 | 0 | 0 | 0 | 76.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Bolotov et al., 2024 [Publ 4369], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 63 | 63 | 61 | 1 | 0 | 0 | 0 | 76.5 |
1
Bolotov et al., 2024 [Publ 4369], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 62 | 62 | 60 | 1 | 0 | 0 | 0 | 75.5 |
1
Bolotov et al., 2022 [Publ 4252], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 61 | 61 | 55 | 1 | 0 | 0 | 0 | 74.5 |
1
Bolotov et al., 2022 [Publ 4252], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 61 | 61 | 49 | 1 | 0 | 0 | 0 | 74.5 |
1
Jeratthitikul et al., 2024 [Publ 4387], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 59 | 59 | 57 | 1 | 0 | 0 | 0 | 74 |
1
Bolotov et al., 2022 [Publ 4239], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 58 | 58 | 52 | 1 | 0 | 0 | 0 | 71.5 |
1
Bolotov et al., 2022 [Publ 4239], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 58 | 58 | 54 | 1 | 0 | 0 | 0 | 71.5 |
1
Bolotov et al., 2022 [Publ 4239], fig. S2
| tT | bi | riS | oT | roT | roS | iD | |
| 58 | 58 | 54 | 1 | 0 | 0 | 0 | 71.5 |
1
Zanatta & Murphy, 2006 [Publ 2672], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 57 | 57 | 48 | 2 | 0 | 0 | 0 | 71 |
+-- Unioninae ** | --+-- Margaritiferidae ** | +-- Ambleminae **
Sano et al., 2020 [Publ 4138], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 57 | 57 | 60 | 2 | 11 | 9 | 2 | 70 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Sano et al., 2020 [Publ 4138], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 57 | 57 | 59 | 2 | 11 | 9 | 2 | 70 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Sano et al., 2020 [Publ 4138], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 57 | 57 | 60 | 2 | 11 | 9 | 2 | 70 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Zanatta & Murphy, 2006 [Publ 2672], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 56 | 56 | 51 | 2 | 0 | 0 | 0 | 69 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Konopleva et al., 2023 [Publ 4269], fig. S1_4
| tT | bi | riS | oT | roT | roS | iD | |
| 56 | 56 | 54 | 2 | 0 | 0 | 0 | 67.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wu et al., 2024 [Publ 4379], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 54 | 54 | 52 | 2 | 0 | 0 | 0 | 65.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Fukata & Iigo, 2020 [Publ 4106], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 54 | 54 | 52 | 2 | 0 | 0 | 0 | 67 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wu et al., 2018 [Publ 3836], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 53 | 53 | 45 | 2 | 0 | 0 | 0 | 65 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Hou et al., 2025 [Publ 4445], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 53 | 53 | 51 | 1 | 0 | 0 | 0 | 66.5 |
1
Dai et al., 2024 [Publ 4358], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 53 | 53 | 51 | 2 | 0 | 0 | 0 | 65.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wu et al., 2024 [Publ 4388], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 53 | 53 | 51 | 2 | 0 | 0 | 0 | 67 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Dai et al., 2023 [Publ 4285], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 53 | 53 | 51 | 2 | 0 | 0 | 0 | 65.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Huang et al., 2019 [Publ 3843], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 52 | 52 | 50 | 2 | 4 | 3 | 2 | 64.5 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Pfeiffer et al., 2019 [Publ 3937], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 52 | 53 | 50 | 2 | 5 | 4 | 2 | 69 |
+-------- Etherioidea --+ | +----- Unionidae ** +--+ | +-- Hyrioidea +--+ +-- Margaritiferidae **
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
| tT | bi | riS | oT | roT | roS | iD | |
| 52 | 52 | 50 | 2 | 5 | 4 | 2 | 65.5 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Konopleva et al., 2019 [Publ 3900], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 51 | 51 | 41 | 2 | 0 | 0 | 0 | 65.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Jeratthitikul et al., 2025 [Publ 4423], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 50 | 50 | 48 | 1 | 0 | 0 | 0 | 65 |
1
Bolotov et al., 2022 [Publ 4250], fig. 8
| tT | bi | riS | oT | roT | roS | iD | |
| 49 | 49 | 42 | 2 | 0 | 0 | 0 | 62 |
+----- Parreysiinae **
+--+
| | +-- Pseudodontini **
| +--+
| +-- Rectidentini **
|
--+ +-- Gibbosulinae **
| |
| +-- Margaritiferinae **
| |
| +-- Sinosolenaia **
| |
+-----+-- Leguminaia **
|
+-- Gonidea **
|
+-- Lamprotula **
|
+-- Potomida **
Chen et al., 2025 [Publ 4663], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 48 | 48 | 47 | 2 | 0 | 0 | 0 | 62.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5
| tT | bi | riS | oT | roT | roS | iD | |
| 48 | 48 | 46 | 2 | 0 | 0 | 0 | 60 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Huang et al., 2019 [Publ 3843], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 47 | 47 | 55 | 2 | 0 | 0 | 0 | 78.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Dai et al., 2025 [Publ 4441], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 47 | 47 | 46 | 2 | 0 | 0 | 0 | 61.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Dai et al., 2024 [Publ 4389], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 46 | 46 | 44 | 2 | 0 | 0 | 0 | 59.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wu et al., 2020 [Publ 4121], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 45 | 45 | 43 | 2 | 4 | 3 | 2 | 57 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Dai et al., 2025 [Publ 4440], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 45 | 45 | 38 | 2 | 0 | 0 | 0 | 57 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Dai et al., 2025 [Publ 4440], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 45 | 45 | 38 | 2 | 0 | 0 | 0 | 57 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Huang et al., 2019 [Publ 3843], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 45 | 45 | 44 | 2 | 0 | 0 | 0 | 59.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Dai et al., 2023 [Publ 4285], fig. 3-BI
| tT | bi | riS | oT | roT | roS | iD | |
| 45 | 45 | 41 | 2 | 0 | 0 | 0 | 58.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Zieritz et al., 2024 [Publ 4381], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 45 | 45 | 43 | 2 | 0 | 0 | 0 | 58.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Dai et al., 2023 [Publ 4285], fig. 3-ML
| tT | bi | riS | oT | roT | roS | iD | |
| 45 | 45 | 43 | 2 | 0 | 0 | 0 | 58.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Froufe et al., 2019 [Publ 3936], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 44 | 44 | 42 | 2 | 0 | 0 | 0 | 56 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Froufe et al., 2019 [Publ 3936], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 44 | 44 | 42 | 2 | 0 | 0 | 0 | 56 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Campbell & Lydeard, 2026 [Publ 4810], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 44 | 44 | 42 | 2 | 3 | 2 | 1 | 56 |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Wu et al., 2019 [Publ 3864], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 44 | 46 | 44 | 2 | 2 | 2 | 2 | 58.5 |
+-------- Unionidae ** --+ | +----- Margaritiferidae ** +--+ | +-- Hyrioidea +--+ +-- Etherioidea
Konopleva et al., 2025 [Publ 4446], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 44 | 44 | 42 | 1 | 0 | 0 | 0 | 59 |
1
Campbell & Lydeard, 2012 [Publ 3308], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 43 | 43 | 42 | 1 | 0 | 0 | 0 | 57 |
1
Inoue et al., 2020 [Publ 3941], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 42 | 42 | 42 | 1 | 0 | 0 | 0 | 56.5 |
1
Campbell & Lydeard, 2012 [Publ 3308], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 42 | 42 | 41 | 1 | 0 | 0 | 0 | 55 |
1
Campbell & Lydeard, 2012 [Publ 3308], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 42 | 42 | 36 | 1 | 0 | 0 | 0 | 55 |
1
Campbell & Lydeard, 2012 [Publ 3308], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 42 | 42 | 40 | 1 | 0 | 0 | 0 | 55 |
1
Dai et al., 2024 [Publ 4386], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 42 | 42 | 40 | 1 | 0 | 0 | 0 | 55.5 |
1
Smith et al., 2019 [Publ 3951], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 41 | 41 | 40 | 1 | 0 | 0 | 0 | 55.5 |
1
Whelan et al., 2023 [Publ 4287], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 41 | 41 | 42 | 1 | 0 | 0 | 0 | 57.5 |
1
Chen et al., 2025 [Publ 4487], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 41 | 41 | 39 | 2 | 0 | 0 | 0 | 53 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Burlakova et al., 2012 [Publ 3486], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 41 | 41 | 42 | 1 | 0 | 0 | 0 | 56 |
1
Chen et al., 2025 [Publ 4487], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 41 | 41 | 39 | 2 | 0 | 0 | 0 | 53 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Chen et al., 2023 [Publ 4302], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 41 | 41 | 32 | 2 | 0 | 0 | 0 | 53 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Inoue et al., 2020 [Publ 3941], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 41 | 41 | 41 | 1 | 0 | 0 | 0 | 55.5 |
1
Chen et al., 2023 [Publ 4302], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 41 | 41 | 37 | 2 | 0 | 0 | 0 | 53 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Bolotov et al., 2023 [Publ 4312], fig. S2
| tT | bi | riS | oT | roT | roS | iD | |
| 40 | 40 | 38 | 1 | 0 | 0 | 0 | 56 |
1
Aunins et al., 2018 [Publ 3863], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 40 | 40 | 38 | 2 | 0 | 0 | 0 | 52 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5
| tT | bi | riS | oT | roT | roS | iD | |
| 39 | 39 | 37 | 2 | 0 | 0 | 0 | 52 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wu et al., 2022 [Publ 4238], fig. 2d
| tT | bi | riS | oT | roT | roS | iD | |
| 39 | 39 | 37 | 1 | 0 | 0 | 0 | 52 |
1
Bolotov et al., 2023 [Publ 4312], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 38 | 38 | 36 | 1 | 0 | 0 | 0 | 53 |
1
Chase et al., 2018 [Publ 3847], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 38 | 39 | 38 | 2 | 1 | 1 | 1 | 53 |
+-------------- Margaritiferidae ** | --+ +----------- Unioninae ** | | +--+ +-------- Lampsilini ** | | +--+ +-- Hyrioidea | +--+ | | +-- Amblemini ** +--+ | +-- Pleurobemini ** +--+ +-- Quadrulini **
Boyer et al., 2011 [Publ 2718], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 38 | 38 | 33 | 1 | 0 | 0 | 0 | 51.5 |
1
Pfeiffer et al., 2016 [Publ 3660], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 38 | 38 | 34 | 1 | 0 | 0 | 0 | 52 |
1
Cha et al., 2018 [Publ 3916], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 38 | 38 | 36 | 2 | 0 | 0 | 0 | 50 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Huang et al., 2018 [Publ 3828], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 38 | 38 | 36 | 2 | 4 | 3 | 2 | 50 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Bolotov et al., 2023 [Publ 4312], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 38 | 38 | 36 | 1 | 0 | 0 | 0 | 53 |
1
Perkins et al., 2017 [Publ 3777], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 37 | 37 | 34 | 1 | 0 | 0 | 0 | 51.5 |
1
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 37 | 37 | 35 | 2 | 0 | 0 | 0 | 50 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Breton et al., 2011 [Publ 3376], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 37 | 37 | 36 | 2 | 1 | 1 | 1 | 50.5 |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Burlakova et al., 2012 [Publ 3486], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 36 | 36 | 26 | 1 | 0 | 0 | 0 | 52.5 |
1
Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 36 | 36 | 22 | 1 | 0 | 0 | 0 | 50.5 |
1
Perkins et al., 2025 [Publ 4447], fig. 2_mtDNA-ITS
| tT | bi | riS | oT | roT | roS | iD | |
| 36 | 36 | 37 | 1 | 0 | 0 | 0 | 53.5 |
1
Wu et al., 2024 [Publ 4350], fig. 5c
| tT | bi | riS | oT | roT | roS | iD | |
| 36 | 36 | 24 | 2 | 0 | 0 | 0 | 50.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wu et al., 2024 [Publ 4350], fig. 6-7
| tT | bi | riS | oT | roT | roS | iD | |
| 36 | 36 | 34 | 2 | 0 | 0 | 0 | 49.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wu et al., 2024 [Publ 4350], fig. 5b
| tT | bi | riS | oT | roT | roS | iD | |
| 36 | 36 | 34 | 2 | 0 | 0 | 0 | 49.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 36 | 36 | 20 | 1 | 0 | 0 | 0 | 50.5 |
1
Wu et al., 2024 [Publ 4350], fig. 5a
| tT | bi | riS | oT | roT | roS | iD | |
| 36 | 36 | 34 | 2 | 0 | 0 | 0 | 49.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wu et al., 2024 [Publ 4388], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 36 | 36 | 34 | 2 | 0 | 0 | 0 | 49.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Liu et al., 2024 [Publ 4380], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 36 | 36 | 34 | 2 | 0 | 0 | 0 | 49.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Bolotov et al., 2020 [Publ 4084], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 36 | 36 | 22 | 2 | 0 | 0 | 0 | 50 |
+----- Margaritiferidae ** | --+----- Ambleminae ** | | +-- Gonideinae ** +--+ +-- Unioninae **
Jeratthitikul et al., 2021 [Publ 4219], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 35 | 35 | 33 | 1 | 0 | 0 | 0 | 50 |
1
Boyer et al., 2011 [Publ 2718], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 35 | 35 | 26 | 1 | 0 | 0 | 0 | 48.5 |
1
Burlakova et al., 2012 [Publ 3486], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 35 | 35 | 30 | 1 | 0 | 0 | 0 | 53.5 |
1
Wu et al., 2025 [Publ 4442], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 35 | 35 | 33 | 1 | 0 | 0 | 0 | 49 |
1
Jeratthitikul et al., 2021 [Publ 4219], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 34 | 34 | 32 | 1 | 0 | 0 | 0 | 50 |
1
Dai et al., 2026 [Publ 4757], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 34 | 34 | 33 | 2 | 0 | 0 | 0 | 46.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Chen et al., 2020 [Publ 4170], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 34 | 34 | 32 | 2 | 0 | 0 | 0 | 49 |
+----- Prolasmidonta **
+--+
| | +-- Alasmidonta **
| +--+
+-----------------+ +-- Anemina **
| |
| | +----- Pseudodontini **
| +--+
| | +-- Lamprotula_caveata **
| +--+
| +-- Chamberlainia **
|
| +----- Oxynaiina **
| +-----+
| | | +-- Lepidodesmini **
--+ | +--+
| | +-- Aculamprotulina **
| +-----------+
| | | +----- Platynaias **
| | | +--+
| | | | | +-- Utterbackia_imbecillis **
| | | | +--+
| | +--+ +-- Pyganodon **
| | |
| | | +----- Sinanodonta **
| | +--+
| | | +-- Cristaria **
+--+ +--+
| +-- Anodontina **
|
| +-- Utterbackia_peninsularis **
| +-----+
| | +-- Potamilus_alatus **
| +-----------+
| | | +----- Lamprotula_leaii **
| | +--+
| | | +-- Sinohyriopsis **
+--+ +--+
| +-- Gonideini **
|
| +----------------- Margaritiferidae **
+--+
| +-------------- Quadrulini **
+--+
| +----------- Toxolasma **
+--+
| +-------- Lampsilis_ornata **
+--+
| +----- Potamilus_leptodon **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Lampsilis_powellii **
Chen et al., 2026 [Publ 4805], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 34 | 34 | 32 | 1 | 0 | 0 | 0 | 48 |
1
Huang et al., 2002 [Publ 2657], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 34 | 34 | 31 | 2 | 0 | 0 | 0 | 48 |
+-- Cumberlandia ** | --+-- Margaritifera ** | +-- Unionidae **
Konopleva et al., 2019 [Publ 4041], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 34 | 34 | 21 | 1 | 0 | 0 | 0 | 50 |
1
Huang et al., 2002 [Publ 2657], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 34 | 34 | 15 | 2 | 0 | 0 | 0 | 48 |
+-- Cumberlandia ** | --+-- Margaritifera ** | +-- Unionidae **
Burlakova et al., 2019 [Publ 3899], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 33 | 33 | 21 | 1 | 0 | 0 | 0 | 46 |
1
Krebs et al., 2003 [Publ 2691], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 33 | 33 | 29 | 2 | 0 | 0 | 0 | 46.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Sano et al., 2020 [Publ 4138], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 33 | 33 | 16 | 2 | 1 | 0 | 0 | 56.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 32 | 32 | 30 | 2 | 8 | 7 | 2 | 43 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Lopes-Lima et al., 2017 [Publ 3792], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 32 | 32 | 30 | 2 | 0 | 0 | 0 | 44 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Lyubas et al., 2022 [Publ 4255], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 22 | 1 | 0 | 0 | 0 | 43 |
1
Zieritz et al., 2021 [Publ 4218], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 29 | 2 | 0 | 0 | 0 | 44 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wang et al., 2018 [Publ 3865], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 29 | 1 | 0 | 0 | 0 | 44.5 |
1
Dai et al., 2024 [Publ 4385], fig. 3-BI
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 28 | 2 | 0 | 0 | 0 | 43.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Sano et al., 2017 [Publ 3799], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 32 | 2 | 2 | 0 | 0 | 45.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Fonseca et al., 2016 [Publ 3775], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 29 | 2 | 0 | 0 | 0 | 43 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Zieritz et al., 2020 [Publ 4102], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 27 | 1 | 0 | 0 | 0 | 46 |
1
Dai et al., 2024 [Publ 4385], fig. 3-ML
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 29 | 2 | 0 | 0 | 0 | 43.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Bogan et al., 2023 [Publ 4295], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 28 | 2 | 0 | 0 | 0 | 43.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Ouyang et al., 2015 [Publ 4493], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 38 | 1 | 0 | 0 | 0 | 54 |
1
Wang et al., 2018 [Publ 3846], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 29 | 1 | 0 | 0 | 0 | 44.5 |
1
Lyubas et al., 2022 [Publ 4255], fig. 7
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 28 | 1 | 0 | 0 | 0 | 43 |
1
Ouyang et al., 2015 [Publ 4493], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 29 | 1 | 0 | 0 | 0 | 46 |
1
Pfeiffer et al., 2019 [Publ 3866], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 31 | 31 | 30 | 1 | 0 | 0 | 0 | 44 |
1
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 30 | 31 | 29 | 2 | 1 | 1 | 1 | 44.5 |
+-- Margaritiferidae ** | --+-- Hyrioidea | +-- Unionidae **
Burdick & White, 2007 [Publ 2679], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 30 | 30 | 31 | 1 | 0 | 0 | 0 | 46 |
1
Wu et al., 2020 [Publ 4147], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 30 | 30 | 29 | 2 | 0 | 0 | 0 | 42.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Pfeiffer et al., 2021 [Publ 4126], fig. 9
| tT | bi | riS | oT | roT | roS | iD | |
| 29 | 29 | 27 | 1 | 0 | 0 | 0 | 44 |
1
Serb et al., 2003 [Publ 2659], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 29 | 29 | 20 | 1 | 0 | 0 | 0 | 42 |
1
Lopes-Lima et al., 2019 [Publ 3901], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 29 | 29 | 27 | 2 | 0 | 0 | 0 | 41.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Serb et al., 2003 [Publ 2659], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 29 | 29 | 27 | 1 | 0 | 0 | 0 | 42 |
1
Smith et al., 2020 [Publ 4129], fig. 2-3
| tT | bi | riS | oT | roT | roS | iD | |
| 28 | 28 | 26 | 1 | 0 | 0 | 0 | 41.5 |
1
Xiang et al., 2025 [Publ 4433], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 28 | 28 | 26 | 1 | 0 | 0 | 0 | 42 |
1
Hou et al., 2025 [Publ 4401], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 28 | 28 | 26 | 1 | 0 | 0 | 0 | 43 |
1
Smith et al., 2020 [Publ 4129], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 28 | 28 | 26 | 1 | 0 | 0 | 0 | 41.5 |
1
Lopes-Lima et al., 2019 [Publ 3901], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 28 | 28 | 25 | 2 | 0 | 0 | 0 | 40 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Hou et al., 2025 [Publ 4432], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 28 | 28 | 26 | 1 | 0 | 0 | 0 | 43 |
1
Pfeiffer et al., 2018 [Publ 3827], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 28 | 28 | 26 | 2 | 0 | 0 | 0 | 40 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Guerra et al., 2019 [Publ 4168], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 28 | 28 | 26 | 2 | 13 | 12 | 2 | 40.5 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Whelan et al., 2023 [Publ 4287], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 28 | 28 | 27 | 1 | 0 | 0 | 0 | 42.5 |
1
Guerra et al., 2019 [Publ 4168], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 28 | 28 | 26 | 2 | 13 | 12 | 2 | 40.5 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Neemuchwala et al., 2023 [Publ 4292], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 27 | 27 | 25 | 1 | 0 | 0 | 0 | 40.5 |
1
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A
| tT | bi | riS | oT | roT | roS | iD | |
| 27 | 27 | 21 | 2 | 3 | 2 | 2 | 38 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Lydeard et al., 1996 [Publ 2656], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 27 | 27 | 25 | 2 | 1 | 0 | 0 | 40 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B
| tT | bi | riS | oT | roT | roS | iD | |
| 27 | 27 | 25 | 2 | 3 | 2 | 2 | 38 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Whelan et al., 2023 [Publ 4287], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 27 | 27 | 25 | 1 | 0 | 0 | 0 | 41.5 |
1
Zhang et al., 2024 [Publ 4397], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 27 | 27 | 25 | 2 | 0 | 0 | 0 | 40.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Hoeh et al., 2009 [Publ 2716], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 27 | 28 | 26 | 2 | 20 | 19 | 2 | 41 |
+-------------------------- Hyrioidea | | +-------- Iridinidae --+ +--------------+ | | | +----- Etheria | | +--+ | | | +-- Mulleria +--+ +--+ | +-- Mycetopodidae | | +-------------------- Margaritiferidae ** +--+ | +----------------- Parreysiinae ** +--+ | +-------------- Pseudomulleria +--+ | +----------- Unioninae ** +--+ | +-------- Rectidentini ** +--+ | +----- Ambleminae ** +--+ | +-- Gonideini ** +--+ +-- Pseudodontini **
Lydeard et al., 1996 [Publ 2656], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 27 | 27 | 23 | 2 | 0 | 0 | 0 | 40 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Graf & Cummings, 2006 [Publ 1968], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 27 | 27 | 21 | 2 | 20 | 19 | 2 | 40 |
+-- Hyrioidea
+--+
| +-- Etherioidea
--+
| +-- Margaritiferidae **
+--+
+-- Unionidae **
Campbell et al., 2008 [Publ 2675], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 26 | 26 | 18 | 1 | 0 | 0 | 0 | 41.5 |
1
Lopes-Lima et al., 2019 [Publ 3901], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 26 | 26 | 24 | 2 | 0 | 0 | 0 | 38.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Konopleva et al., 2021 [Publ 4213], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 26 | 26 | 21 | 1 | 0 | 0 | 0 | 42 |
1
Hoeh et al., 2009 [Publ 2716], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 26 | 45 | 39 | 2 | 20 | 19 | 2 | 58 |
+----------------- Parreysiinae ** | | +----- Margaritiferidae ** --+ +--------+ | | | +-- Hyrioidea | | +--+ +--+ +-- Etherioidea | | +----------- Pseudodontini ** +--+ | +-------- Rectidentini ** +--+ | +----- Unioninae ** +--+ | +-- Gonideini ** +--+ +-- Ambleminae **
Hoeh et al., 2009 [Publ 2716], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 26 | 27 | 24 | 2 | 20 | 19 | 2 | 42 |
+-------------------------- Hyrioidea | | +-- Etheria | +--+ --+ | +-- Mulleria | +-----------------+ | | | +-- Mycetopodidae | | +--+ +--+ +-- Iridinidae | | +-------------------- Parreysiinae ** +--+ | +----------------- Pseudomulleria +--+ | +-------------- Margaritiferidae ** +--+ | +----------- Pseudodontini ** +--+ | +-------- Rectidentini ** +--+ | +----- Unioninae ** +--+ | +-- Gonideini ** +--+ +-- Ambleminae **
Whelan et al., 2011 [Publ 3380], fig. 1-MP
| tT | bi | riS | oT | roT | roS | iD | |
| 26 | 30 | 27 | 2 | 10 | 9 | 2 | 44.5 |
+-------- Etherioidea --+ | +----- Unionidae ** +--+ | +-- Margaritiferidae ** +--+ +-- Hyrioidea
Campbell et al., 2008 [Publ 2675], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 26 | 26 | 23 | 1 | 0 | 0 | 0 | 40.5 |
1
Whelan et al., 2011 [Publ 3380], fig. 1-ML
| tT | bi | riS | oT | roT | roS | iD | |
| 26 | 31 | 30 | 2 | 10 | 9 | 2 | 44 |
+-------- Hyrioidea --+ | +----- Margaritiferidae ** +--+ | +-- Etherioidea +--+ +-- Unionidae **
Campbell et al., 2008 [Publ 2675], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 25 | 25 | 6 | 1 | 0 | 0 | 0 | 40.5 |
1
Lopes-Lima et al., 2021 [Publ 4217], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 25 | 25 | 23 | 1 | 0 | 0 | 0 | 38.5 |
1
Campbell et al., 2008 [Publ 2675], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 25 | 25 | 9 | 1 | 0 | 0 | 0 | 39.5 |
1
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11
| tT | bi | riS | oT | roT | roS | iD | |
| 25 | 25 | 23 | 1 | 0 | 0 | 0 | 39.5 |
1
Liu et al., 2023 [Publ 4310], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 24 | 24 | 17 | 1 | 0 | 0 | 0 | 37 |
1
Liu et al., 2023 [Publ 4310], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 24 | 24 | 22 | 1 | 0 | 0 | 0 | 37 |
1
Wu et al., 2021 [Publ 4215], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 24 | 24 | 22 | 1 | 0 | 0 | 0 | 38 |
1
Perkins et al., 2017 [Publ 3777], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 24 | 24 | 10 | 1 | 0 | 0 | 0 | 41.5 |
1
Galal-Khallaf et al., 2024 [Publ 4459], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 24 | 28 | 29 | 1 | 4 | 4 | 1 | 53.5 |
+----- Parreysiinae ** --+ | +-- Etherioidea +--+ +-- Unioninae **
Zieritz et al., 2021 [Publ 4231], fig. 3-M_4
| tT | bi | riS | oT | roT | roS | iD | |
| 23 | 23 | 21 | 2 | 0 | 0 | 0 | 36 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wu et al., 2020 [Publ 4121], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 23 | 23 | 21 | 2 | 11 | 9 | 2 | 35.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Unionidae ** +--+ +-- Margaritiferidae **
Jeratthitikul et al., 2019 [Publ 3922], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 23 | 23 | 21 | 1 | 0 | 0 | 0 | 36.5 |
1
Soroka & Burzynski, 2017 [Publ 3837], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 23 | 23 | 21 | 1 | 0 | 0 | 0 | 36 |
1
Wu et al., 2020 [Publ 4121], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 23 | 23 | 21 | 2 | 11 | 9 | 2 | 35.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Unionidae ** +--+ +-- Margaritiferidae **
Konopleva et al., 2017 [Publ 3807], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 23 | 23 | 21 | 2 | 8 | 6 | 2 | 36.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Wu et al., 2024 [Publ 4379], fig. 4_6
| tT | bi | riS | oT | roT | roS | iD | |
| 23 | 23 | 21 | 2 | 0 | 0 | 0 | 35.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Inoue et al., 2020 [Publ 4128], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 23 | 23 | 16 | 1 | 0 | 0 | 0 | 39.5 |
1
Wu et al., 2023 [Publ 4296], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 23 | 23 | 21 | 1 | 0 | 0 | 0 | 36 |
1
Dai et al., 2024 [Publ 4386], fig. 5-ML
| tT | bi | riS | oT | roT | roS | iD | |
| 22 | 22 | 19 | 2 | 0 | 0 | 0 | 33 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Huang et al., 2019 [Publ 3843], fig. 3-M
| tT | bi | riS | oT | roT | roS | iD | |
| 22 | 22 | 20 | 2 | 1 | 1 | 1 | 34.5 |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG
| tT | bi | riS | oT | roT | roS | iD | |
| 22 | 22 | 20 | 1 | 0 | 0 | 0 | 35.5 |
1
Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE
| tT | bi | riS | oT | roT | roS | iD | |
| 22 | 22 | 20 | 1 | 0 | 0 | 0 | 35.5 |
1
Dai et al., 2024 [Publ 4386], fig. 5-BI
| tT | bi | riS | oT | roT | roS | iD | |
| 22 | 22 | 16 | 2 | 0 | 0 | 0 | 33 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Inoue et al., 2020 [Publ 4128], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 22 | 22 | 24 | 1 | 0 | 0 | 0 | 37.5 |
1
Wu et al., 2018 [Publ 4143], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 22 | 22 | 20 | 1 | 0 | 0 | 0 | 35.5 |
1
Jeratthitikul et al., 2021 [Publ 4212], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 21 | 21 | 19 | 1 | 0 | 0 | 0 | 36 |
1
Guerra et al., 2017 [Publ 3772], fig. 2a
| tT | bi | riS | oT | roT | roS | iD | |
| 21 | 21 | 19 | 2 | 4 | 3 | 2 | 33 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Chapman et al., 2008 [Publ 2667], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 21 | 21 | 19 | 1 | 0 | 0 | 0 | 34.5 |
1
Rosenberg et al., 1997 [Publ 2720], fig. 2A
| tT | bi | riS | oT | roT | roS | iD | |
| 21 | 22 | 4 | 2 | 1 | 0 | 0 | 31.5 |
+----- Cumberlandia ** | +----- None__2 --+ +----- Margaritifera ** | | +-- None__1 +--+ +-- Unionidae **
Guerra et al., 2017 [Publ 3772], fig. 2b
| tT | bi | riS | oT | roT | roS | iD | |
| 21 | 21 | 19 | 2 | 4 | 3 | 2 | 33 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Chapman et al., 2008 [Publ 2744], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 21 | 21 | 19 | 1 | 0 | 0 | 0 | 34.5 |
1
Rosenberg et al., 1997 [Publ 2720], fig. 2B
| tT | bi | riS | oT | roT | roS | iD | |
| 21 | 22 | 3 | 2 | 1 | 0 | 0 | 31.5 |
+-- Cumberlandia ** | +-- None__1 | --+-- None__2 | +-- Margaritifera ** | +-- Unionidae **
Wu et al., 2016 [Publ 3711], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 21 | 21 | 19 | 1 | 0 | 0 | 0 | 34.5 |
1
Zhou et al., 2007 [Publ 2658], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 21 | 21 | 19 | 2 | 0 | 0 | 0 | 33.5 |
+-------------------- Margaritiferinae ** | | +----------------- Anodontini ** --+ | | | +----------- Chamberlainiini ** | | +--+ +--+ | | +-------- Lamprotula_leaii ** | | +--+ | | | +----- Lamprotula_caveata ** | | +--+ | | | +-- Gibbosulinae ** +--+ +--+ | +-- Lamprotula_cornuumlunae ** | | +----- Aculamprotulina ** | +--+ | | | +-- Tchangsinaia ** | | +--+ +-----+ +-- Nodularia ** | | +----- Schistodesmus ** +--+ | +-- Lanceolariini ** +--+ +-- Acuticostini **
Matos et al., 2024 [Publ 4383], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 18 | 2 | 0 | 0 | 0 | 33.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Roe & Hoeh, 2003 [Publ 2655], fig. 4f2
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 16 | 2 | 11 | 10 | 2 | 31.5 |
+-- Etherioidea
+--+
| +-- Hyrioidea
--+
| +-- Margaritiferidae **
+--+
+-- Unionidae **
Inoue et al., 2014 [Publ 3635], fig. 3A
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 17 | 2 | 0 | 0 | 0 | 32 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Hoeh et al., 2001 [Publ 2652], fig. 14f2
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 30 | 18 | 2 | 11 | 10 | 2 | 40 |
+----------------- Margaritiferidae ** | | +-------------- Amblemini ** | | | +-------------- Parreysiinae ** | | | +-------------- Eurynia ** --+ | | +-------------- Fusconaia ** | | | +-------------- Gonideinae ** | | | +-------------- Pleurobema ** | | +--+-------------- Quadrulini ** | | +----------- Ortmanniana ** +--+ | | +-------- Glebula ** | +--+ | | +----- Potamilus ** | +--+ | | +-- Ligumia ** | +--+ | +-- Toxolasma ** | | +----------- Cyrtonaias ** +--+ | +-------- Unionini ** +--+ | +----- Anodontini ** +--+ | +-- Hyrioidea +--+ +-- Etherioidea
Hoeh et al., 2002 [Publ 2653], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 17 | 2 | 11 | 10 | 2 | 34.5 |
+----------------- Hyrioidea --+ | +-------------- Etherioidea +--+ | +----------- Parreysiinae ** +--+ | +-------- Margaritiferidae ** +--+ | +----- Ambleminae ** +--+ | +-- Gonideinae ** +--+ +-- Unioninae **
Hoeh et al., 2001 [Publ 2652], fig. 14f4
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 17 | 2 | 11 | 10 | 2 | 34.5 |
+----------------- Hyrioidea --+ | +-------------- Etherioidea +--+ | +----------- Parreysiinae ** +--+ | +-------- Margaritiferidae ** +--+ | +----- Gonideinae ** +--+ | +-- Unioninae ** +--+ +-- Ambleminae **
Matos et al., 2024 [Publ 4384], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 18 | 2 | 0 | 0 | 0 | 33.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Guerra et al., 2019 [Publ 4168], fig. 1-M
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 18 | 2 | 2 | 2 | 1 | 32.5 |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Zhou et al., 2016 [Publ 3710], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 18 | 2 | 0 | 0 | 0 | 34 |
+-------- Unioninae ** --+ | +----- Margaritiferidae ** +--+ | +-- Ambleminae ** +--+ +-- Gonideinae **
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 17 | 2 | 0 | 0 | 0 | 33 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Roe & Hoeh, 2003 [Publ 2655], fig. 4f3
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 30 | 28 | 2 | 11 | 10 | 2 | 40 |
+-------- Margaritiferidae ** --+ | +----- Unionidae ** +--+ | +-- Etherioidea +--+ +-- Hyrioidea
Guerra et al., 2019 [Publ 4168], fig. 2-M
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 18 | 2 | 2 | 2 | 1 | 32.5 |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Pieri et al., 2018 [Publ 3867], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 23 | 1 | 0 | 0 | 0 | 39 |
1
Smith et al., 2020 [Publ 4136], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 20 | 13 | 1 | 0 | 0 | 0 | 34.5 |
1
Bogan & Hoeh, 2000 [Publ 2650], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 20 | 21 | 14 | 2 | 14 | 13 | 2 | 38.5 |
+-------------------- Hyrioidea | | +-------- Iridinidae --+ +--------+ | | | +----- Etheria | | +--+ | | | +-- Mulleria +--+ +--+ | +-- Mycetopodidae | | +-------------- Parreysiinae ** | | +--+ +----------- Pseudomulleria | | +--+ +-------- Anodontini ** | | +--+ +----- Gonideinae ** | | +--+----- Unionini ** | | +-- Margaritiferidae ** +--+ +-- Ambleminae **
Rosenberg et al., 1994 [Publ 2719], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 19 | 19 | 5 | 2 | 0 | 0 | 0 | 29.5 |
+-- Cumberlandia ** | --+-- Margaritifera ** | +-- Unionidae **
Sano et al., 2017 [Publ 3799], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 19 | 19 | 22 | 2 | 2 | 0 | 0 | 34.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Froufe et al., 2019 [Publ 3936], fig. 3-M
| tT | bi | riS | oT | roT | roS | iD | |
| 19 | 19 | 17 | 2 | 0 | 0 | 0 | 32 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Riccardi et al., 2020 [Publ 4089], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 19 | 19 | 17 | 2 | 0 | 0 | 0 | 31 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 19 | 19 | 17 | 2 | 0 | 0 | 0 | 31.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Wu et al., 2023 [Publ 4339], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 19 | 19 | 17 | 2 | 0 | 0 | 0 | 31.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Froufe et al., 2019 [Publ 3936], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 19 | 19 | 17 | 2 | 0 | 0 | 0 | 32 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Zhou et al., 2007 [Publ 2658], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 18 | 18 | 16 | 1 | 0 | 0 | 0 | 33 |
1
Chong et al., 2008 [Publ 2668], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 18 | 18 | 10 | 1 | 0 | 0 | 0 | 31 |
1
Araujo et al., 2018 [Publ 3779], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 18 | 18 | 14 | 2 | 0 | 0 | 0 | 30.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Bolotov et al., 2019 [Publ 3974], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 18 | 18 | 16 | 1 | 0 | 0 | 0 | 33 |
1
Kim et al., 2021 [Publ 4197], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 18 | 18 | 16 | 1 | 0 | 0 | 0 | 32.5 |
1
Porto-Hannes et al., 2019 [Publ 3969], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 18 | 18 | 13 | 1 | 0 | 0 | 0 | 32.5 |
1
Sano et al., 2022 [Publ 4311], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 18 | 18 | 16 | 1 | 0 | 0 | 0 | 33 |
1
Wu et al., 2022 [Publ 4238], fig. 2e
| tT | bi | riS | oT | roT | roS | iD | |
| 18 | 18 | 16 | 1 | 0 | 0 | 0 | 31 |
1
Porto-Hannes et al., 2019 [Publ 3969], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 18 | 18 | 16 | 1 | 0 | 0 | 0 | 32.5 |
1
Huang et al., 2018 [Publ 3828], fig. 3_5
| tT | bi | riS | oT | roT | roS | iD | |
| 18 | 18 | 16 | 2 | 11 | 9 | 2 | 30.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Unionidae ** +--+ +-- Margaritiferidae **
Smith et al., 2018 [Publ 3790], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 18 | 18 | 17 | 1 | 0 | 0 | 0 | 32.5 |
1
Wu et al., 2026 [Publ 4811], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 17 | 17 | 15 | 2 | 0 | 0 | 0 | 30 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Smith, 2021 [Publ 4331], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 17 | 17 | 15 | 2 | 3 | 2 | 1 | 30 |
+----- Etherioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Lopes-Lima et al., 2018 [Publ 3829], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 17 | 17 | 15 | 2 | 11 | 9 | 2 | 28.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Unionidae ** +--+ +-- Margaritiferidae **
Prié & Puillandre, 2014 [Publ 3529], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 17 | 17 | 15 | 2 | 0 | 0 | 0 | 27.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Plazzi et al., 2011 [Publ 3676], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 17 | 18 | 11 | 1 | 1 | 1 | 1 | 32.5 |
+----------- Cristariina ** | | +-------- Lanceolariini ** | | --+ +-------- Anodonta ** | | | +-------- Unionini ** | | +--+ +-- Alasmidontina ** +-----+ | +-- Pseudanodonta ** | | +----- Ambleminae ** | | +--+ +-- Hyrioidea | | +--+-- Lamprotulini ** | +-- Chamberlainiini **
Wu et al., 2025 [Publ 4442], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 17 | 17 | 15 | 1 | 0 | 0 | 0 | 30.5 |
1
Inoue et al., 2013 [Publ 3528], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 17 | 17 | 13 | 1 | 0 | 0 | 0 | 31 |
1
Fukata & Iigo, 2019 [Publ 4108], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 17 | 17 | 15 | 1 | 0 | 0 | 0 | 31.5 |
1
Grobler et al., 2011 [Publ 3359], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 17 | 17 | 13 | 1 | 0 | 0 | 0 | 31.5 |
1
Lopes-Lima et al., 2018 [Publ 3829], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 17 | 17 | 15 | 2 | 11 | 9 | 2 | 28.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Unionidae ** +--+ +-- Margaritiferidae **
Chen et al., 2025 [Publ 4455], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 17 | 17 | 15 | 2 | 9 | 7 | 2 | 30 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Unionidae ** +--+ +-- Margaritiferidae **
Lyubas et al., 2019 [Publ 3963], fig. 10
| tT | bi | riS | oT | roT | roS | iD | |
| 16 | 16 | 12 | 1 | 0 | 0 | 0 | 30.5 |
1
Zanatta et al., 2007 [Publ 2698], fig. 1B
| tT | bi | riS | oT | roT | roS | iD | |
| 16 | 16 | 13 | 1 | 0 | 0 | 0 | 30.5 |
1
Szumowski et al., 2012 [Publ 3569], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 16 | 16 | 12 | 1 | 0 | 0 | 0 | 30.5 |
1
Walker et al., 2006 [Publ 2666], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 16 | 16 | 13 | 2 | 20 | 19 | 2 | 30.5 |
+-------------- Hyrioidea --+ | +----------- Etherioidea +--+ | +-------- Unioninae ** +--+ | +----- Margaritiferidae ** +--+ | +-- Gonideinae ** +--+ +-- Ambleminae **
Zhou et al., 2017 [Publ 3788], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 16 | 16 | 14 | 1 | 0 | 0 | 0 | 30.5 |
1
Wen et al., 2017 [Publ 3795], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 16 | 16 | 14 | 2 | 0 | 0 | 0 | 28.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Lopes-Lima et al., 2017 [Publ 3734], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 16 | 16 | 12 | 2 | 1 | 0 | 0 | 27 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Stewart et al., 2021 [Publ 4451], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 16 | 16 | 15 | 2 | 0 | 0 | 0 | 28 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Johnson et al., 2018 [Publ 3844], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 16 | 16 | 14 | 1 | 0 | 0 | 0 | 30.5 |
1
Inoue et al., 2020 [Publ 3941], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 16 | 16 | 14 | 1 | 0 | 0 | 0 | 30.5 |
1
Huang et al., 2013 [Publ 3629], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 16 | 16 | 14 | 2 | 0 | 0 | 0 | 28 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Huang et al., 2018 [Publ 3828], fig. 6-M
| tT | bi | riS | oT | roT | roS | iD | |
| 16 | 16 | 14 | 2 | 1 | 1 | 1 | 29 |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Zanatta et al., 2007 [Publ 2698], fig. 1A
| tT | bi | riS | oT | roT | roS | iD | |
| 16 | 16 | 10 | 1 | 0 | 0 | 0 | 30.5 |
1
Bolotov et al., 2022 [Publ 4250], fig. S2
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 13 | 1 | 0 | 0 | 0 | 29.5 |
1
Kongim et al., 2023 [Publ 4290], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 13 | 1 | 0 | 0 | 0 | 30 |
1
Hoeh, 1990 [Publ 2717], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 2 | 1 | 0 | 0 | 0 | 29.5 |
1
Jeratthitikul et al., 2025 [Publ 4476], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 13 | 1 | 0 | 0 | 0 | 30 |
1
Hoeh, 1990 [Publ 2717], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 13 | 1 | 0 | 0 | 0 | 29.5 |
1
Amaro et al., 2017 [Publ 3791], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 15 | 2 | 1 | 0 | 0 | 28.5 |
+-------- Margaritifera_margaritifera__2 ** --+ | +----- Unionidae ** +--+ | +-- Margaritifera_margaritifera__1 ** +--+ +-- Margaritifera_falcata **
Liu et al., 2017 [Publ 3939], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 12 | 1 | 0 | 0 | 0 | 28.5 |
1
Hou et al., 2024 [Publ 4400], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 13 | 1 | 0 | 0 | 0 | 30 |
1
Xue et al., 2016 [Publ 3659], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 13 | 1 | 0 | 0 | 0 | 29 |
1
Bolotov et al., 2022 [Publ 4250], fig. S5
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 12 | 1 | 0 | 0 | 0 | 30 |
1
Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 13 | 1 | 0 | 0 | 0 | 28 |
1
Hoeh, 1990 [Publ 2717], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 13 | 1 | 0 | 0 | 0 | 29.5 |
1
Porto-Hannes et al., 2021 [Publ 4227], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 15 | 15 | 12 | 1 | 0 | 0 | 0 | 28.5 |
1
Lydeard et al., 2000 [Publ 2654], fig. 2d
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 2 | 0 | 0 | 0 | 29.5 |
+-- Unioninae ** | --+-- Margaritiferidae ** | +-- Ambleminae **
Lydeard et al., 2000 [Publ 2654], fig. 2a
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 11 | 2 | 0 | 0 | 0 | 28.5 |
+----- Unioninae ** | +----- Margaritiferidae ** | --+----- Lampsilini ** | +----- Amblemini ** | | +-- Quadrulini ** +--+ +-- Pleurobemini **
Lopes-Lima et al., 2020 [Publ 4080], fig. 10
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 10 | 2 | 0 | 0 | 0 | 27.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Dai et al., 2024 [Publ 4386], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 1 | 0 | 0 | 0 | 26.5 |
1
Araujo et al., 2017 [Publ 3801], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 2 | 0 | 0 | 0 | 26 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Szumowski et al., 2012 [Publ 3569], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 1 | 0 | 0 | 0 | 29 |
1
Wu et al., 2020 [Publ 4147], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 2 | 0 | 0 | 0 | 25.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Reis et al., 2013 [Publ 3580], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 2 | 1 | 0 | 0 | 29 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Zhang et al., 2004 [Publ 2735], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 1 | 0 | 0 | 0 | 28 |
1
Turner et al., 2000 [Publ 2722], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 2 | 0 | 0 | 0 | 28 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Froufe et al., 2016 [Publ 3738], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 1 | 0 | 0 | 0 | 27.5 |
1
Douda et al., 2024 [Publ 4364], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 13 | 2 | 0 | 0 | 0 | 27.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Lydeard et al., 2000 [Publ 2654], fig. 2b
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 2 | 0 | 0 | 0 | 29.5 |
+-- Unioninae ** | --+-- Margaritiferidae ** | +-- Ambleminae **
Bolotov et al., 2016 [Publ 3743], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 2 | 16 | 13 | 2 | 26.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Unionidae ** +--+ +-- Margaritiferidae **
Feng et al., 2016 [Publ 3800], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 2 | 0 | 0 | 0 | 26.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Lydeard et al., 2000 [Publ 2654], fig. 2c
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 9 | 2 | 0 | 0 | 0 | 28.5 |
+----- Unioninae ** | +----- Margaritiferidae ** | --+----- Quadrulini ** | +----- Lampsilini ** | | +-- Amblemini ** +--+ +-- Pleurobemini **
Bolotov et al., 2016 [Publ 3743], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 2 | 16 | 13 | 2 | 26.5 |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Unionidae ** +--+ +-- Margaritiferidae **
Lopes-Lima et al., 2024 [Publ 4376], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 14 | 14 | 12 | 1 | 0 | 0 | 0 | 29.5 |
1
Manendo et al., 2008 [Publ 2669], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 1 | 0 | 0 | 0 | 28.5 |
1
Hauck et al., 2023 [Publ 4333], fig. 9
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 12 | 1 | 0 | 0 | 0 | 26.5 |
1
Walker et al., 2006 [Publ 2666], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 2 | 2 | 1 | 1 | 27 |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Wu et al., 2024 [Publ 4379], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 10 | 1 | 0 | 0 | 0 | 29 |
1
Burzynski et al., 2017 [Publ 3774], fig. 1-top
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 1 | 0 | 0 | 0 | 28 |
1
Ouyang et al., 2011 [Publ 3408], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 10 | 1 | 0 | 0 | 0 | 28 |
1
Xiang et al., 2025 [Publ 4433], fig. 7
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 1 | 0 | 0 | 0 | 28 |
1
Reis et al., 2013 [Publ 3580], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 2 | 1 | 0 | 0 | 27 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Burzynski et al., 2017 [Publ 3774], fig. 1-bottom
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 1 | 0 | 0 | 0 | 28 |
1
Walker et al., 2006 [Publ 2666], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 2 | 2 | 1 | 1 | 27 |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Bogan et al., 2023 [Publ 4295], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 10 | 2 | 0 | 0 | 0 | 28 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Graf, 2002 [Publ 2651], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 2 | 5 | 4 | 1 | 27.5 |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Vikhrev et al., 2018 [Publ 3812], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 2 | 0 | 0 | 0 | 25.5 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Sayenko et al., 2021 [Publ 4189], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 1 | 0 | 0 | 0 | 27.5 |
1
Jeratthitikul & Sutcharit, 2023 [Publ 4289], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 1 | 0 | 0 | 0 | 29 |
1
Walker et al., 2006 [Publ 2666], fig. 6-M
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 2 | 1 | 1 | 1 | 25.5 |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Walker et al., 2006 [Publ 2666], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 13 | 13 | 11 | 2 | 2 | 1 | 1 | 29 |
+----------------- Hyrioidea | --+ +-------------- Unioninae ** | | +--+ +----------- Lamprotulini ** | | +--+ +-- Gonideini ** | +-----+ | | +-- Pseudodontini ** +--+ | +----- Lampsilini ** +--+ | +-- Pleurobemini ** +--+ +-- Margaritiferidae **
Mabuchi & Nishida, 2020 [Publ 4120], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 12 | 1 | 0 | 0 | 0 | 39 |
1
Sano et al., 2022 [Publ 4311], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 12 | 1 | 0 | 0 | 0 | 27 |
1
Freeman et al., 2025 [Publ 4443], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 10 | 1 | 0 | 0 | 0 | 27 |
1
Araujo et al., 2016 [Publ 3746], fig. 1_2b
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 10 | 2 | 1 | 0 | 0 | 26.5 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Lopes-Lima et al., 2020 [Publ 4080], fig. 11
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 10 | 2 | 0 | 0 | 0 | 26.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Pfarrer et al., 2022 [Publ 4305], fig. 2-5
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 10 | 1 | 0 | 0 | 0 | 26.5 |
1
Zieritz et al., 2026 [Publ 4812], fig. 2a
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 10 | 1 | 0 | 0 | 0 | 28 |
1
Smith et al., 2020 [Publ 4136], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 10 | 1 | 0 | 0 | 0 | 28 |
1
Konopleva et al., 2020 [Publ 4072], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 9 | 1 | 0 | 0 | 0 | 28 |
1
Curole & Kocher, 2005 [Publ 2682], fig. 1-M
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 10 | 2 | 0 | 0 | 0 | 24.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Liu et al., 2024 [Publ 4380], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 10 | 1 | 0 | 0 | 0 | 28 |
1
Choi et al., 2020 [Publ 4169], fig. 2a
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 11 | 1 | 0 | 0 | 0 | 27.5 |
1
Froufe et al., 2015 [Publ 3664], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 10 | 1 | 0 | 0 | 0 | 26.5 |
1
Zieritz et al., 2024 [Publ 4381], fig. 7A
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 10 | 1 | 0 | 0 | 0 | 28 |
1
Freeman et al., 2025 [Publ 4443], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 8 | 1 | 0 | 0 | 0 | 27 |
1
Froufe et al., 2017 [Publ 3802], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 9 | 1 | 0 | 0 | 0 | 28 |
1
Wu et al., 2022 [Publ 4238], fig. 2a_S1
| tT | bi | riS | oT | roT | roS | iD | |
| 12 | 12 | 10 | 1 | 0 | 0 | 0 | 28 |
1
Roe, 2013 [Publ 3571], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 9 | 1 | 0 | 0 | 0 | 26.5 |
1
Hou et al., 2025 [Publ 4432], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 9 | 1 | 0 | 0 | 0 | 27 |
1
Araujo et al., 2009 [Publ 2589], fig. 7B
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 10 | 2 | 0 | 0 | 0 | 26.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Morrison et al., 2021 [Publ 4362], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 9 | 1 | 0 | 0 | 0 | 27 |
1
Klishko et al., 2019 [Publ 3886], fig. 9
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 8 | 2 | 0 | 0 | 0 | 24 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Amaro et al., 2017 [Publ 3791], fig. 3A
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 9 | 2 | 1 | 0 | 0 | 24 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Porto-Hannes et al., 2021 [Publ 4227], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 6 | 1 | 0 | 0 | 0 | 26.5 |
1
Amaro et al., 2017 [Publ 3791], fig. 3B
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 9 | 2 | 1 | 0 | 0 | 24 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Freeman et al., 2025 [Publ 4443], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 9 | 1 | 0 | 0 | 0 | 26 |
1
Hou et al., 2025 [Publ 4401], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 9 | 1 | 0 | 0 | 0 | 27 |
1
Pfeiffer et al., 2021 [Publ 4126], fig. 12
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 8 | 1 | 0 | 0 | 0 | 27 |
1
Araujo et al., 2016 [Publ 3746], fig. 2a
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 9 | 1 | 0 | 0 | 0 | 26.5 |
1
Lopes-Lima et al., 2020 [Publ 4080], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 9 | 2 | 0 | 0 | 0 | 25.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Graf et al., 2014 [Publ 3591], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 13 | 2 | 0 | 0 | 0 | 30 |
+-- Margaritiferidae ** | --+-- Unioninae ** | +-- Parreysiinae **
Curole & Kocher, 2005 [Publ 2682], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 9 | 2 | 0 | 0 | 0 | 25.5 |
+----- Ambleminae ** --+ | +-- Unioninae ** +--+ +-- Margaritiferidae **
Araujo et al., 2009 [Publ 2589], fig. 7A
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 7 | 2 | 0 | 0 | 0 | 25.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Schilling et al., 2025 [Publ 4737], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 9 | 1 | 0 | 0 | 0 | 25.5 |
1
Khalloufi et al., 2011 [Publ 2748], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 8 | 1 | 0 | 0 | 0 | 24.5 |
1
Pfeiffer et al., 2021 [Publ 4126], fig. 27
| tT | bi | riS | oT | roT | roS | iD | |
| 11 | 11 | 7 | 1 | 0 | 0 | 0 | 27 |
1
Huff et al., 2004 [Publ 2697], fig. 5-6
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 7 | 2 | 2 | 0 | 0 | 22.5 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Araujo et al., 2009 [Publ 2589], fig. 7C
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 2 | 0 | 0 | 0 | 24.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Hewitt et al., 2019 [Publ 3962], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 11 | 2 | 0 | 0 | 0 | 23.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Chase et al., 2018 [Publ 3861], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 2 | 1 | 1 | 1 | 23.5 |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Choi et al., 2020 [Publ 4169], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 1 | 0 | 0 | 0 | 25 |
1
Lopes-Lima et al., 2020 [Publ 4080], fig. 13
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 2 | 0 | 0 | 0 | 24.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Bespalaya et al., 2018 [Publ 3811], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 9 | 2 | 0 | 0 | 0 | 25.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Burzynski & Soroka, 2018 [Publ 3862], fig. 5b
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 1 | 0 | 0 | 0 | 24 |
1
Huff et al., 2004 [Publ 2697], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 6 | 2 | 2 | 0 | 0 | 23.5 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Sonowal et al., 2021 [Publ 4221], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 1 | 6 | 3 | 1 | 26.5 |
+-- Hyrioidea --+ +-- Unionidae **
Roe & Lydeard, 1998 [Publ 2693], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 5 | 1 | 0 | 0 | 0 | 24.5 |
1
Huff et al., 2004 [Publ 2697], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 7 | 2 | 1 | 0 | 0 | 23.5 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Schilling et al., 2025 [Publ 4737], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 9 | 1 | 0 | 0 | 0 | 23.5 |
1
Amaro et al., 2017 [Publ 3791], fig. 2-M
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 7 | 1 | 0 | 0 | 0 | 25 |
1
Vikhrev et al., 2017 [Publ 3810], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 2 | 0 | 0 | 0 | 24.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Lopes-Lima et al., 2021 [Publ 4217], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 1 | 0 | 0 | 0 | 25.5 |
1
Kano et al., 2019 [Publ 4194], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 2 | 0 | 0 | 0 | 23.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Keogh & Simons, 2019 [Publ 3935], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 1 | 0 | 0 | 0 | 24.5 |
1
Froufe et al., 2017 [Publ 3802], fig. 7
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 2 | 0 | 0 | 0 | 23 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 2 | 0 | 0 | 0 | 23 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Morrison et al., 2021 [Publ 4362], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 1 | 0 | 0 | 0 | 26 |
1
Chase et al., 2018 [Publ 3861], fig. 1-M
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 11 | 9 | 2 | 1 | 1 | 1 | 25.5 |
+----- Unionidae ** --+ | +-- Margaritiferidae ** +--+ +-- Hyrioidea
Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 2 | 0 | 0 | 0 | 24 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Stodola et al., 2021 [Publ 4266], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 7 | 1 | 0 | 0 | 0 | 26.5 |
1
Roe et al., 2002 [Publ 2689], fig. 3a
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 1 | 0 | 0 | 0 | 24.5 |
1
Bolotov et al., 2015 [Publ 3646], fig. 2A
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 6 | 2 | 0 | 0 | 0 | 23 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Muanta et al., 2019 [Publ 3921], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 8 | 1 | 0 | 0 | 0 | 26 |
1
Zieritz et al., 2026 [Publ 4812], fig. 4a
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 1 | 0 | 0 | 0 | 25 |
1
Nagel et al., 1998 [Publ 1946], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 1 | 0 | 0 | 0 | 23.5 |
1
Chase et al., 2018 [Publ 3847], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 10 | 8 | 2 | 1 | 1 | 1 | 25 |
+----- Unionidae ** --+ | +-- Margaritiferidae ** +--+ +-- Hyrioidea
Soroka, 2010 [Publ 3404], fig. 3a
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 1 | 0 | 0 | 0 | 22.5 |
1
Bolotov et al., 2015 [Publ 3646], fig. 2B
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 2 | 0 | 0 | 0 | 23 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Soroka, 2010 [Publ 3404], fig. 3b
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 5 | 1 | 0 | 0 | 0 | 22.5 |
1
Inoue et al., 2013 [Publ 3579], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 2 | 0 | 0 | 0 | 23 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Kandl et al., 2001 [Publ 3384], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 1 | 0 | 0 | 0 | 23.5 |
1
Takeuchi et al., 2015 [Publ 3661], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 2 | 1 | 0 | 0 | 23 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Bolotov et al., 2022 [Publ 4250], fig. S4
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 6 | 1 | 0 | 0 | 0 | 25 |
1
Wu et al., 2024 [Publ 4388], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 1 | 0 | 0 | 0 | 25 |
1
Chen et al., 2025 [Publ 4448], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 1 | 0 | 0 | 0 | 25 |
1
Fonseca et al., 2016 [Publ 3775], fig. 1-M
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 1 | 0 | 0 | 0 | 24 |
1
Takeuchi et al., 2015 [Publ 3661], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 2 | 1 | 0 | 0 | 23 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Hewitt et al., 2018 [Publ 3758], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 6 | 2 | 0 | 0 | 0 | 22 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Jeratthitikul et al., 2019 [Publ 3911], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 14 | 1 | 0 | 0 | 0 | 25 |
1
Sonowal et al., 2021 [Publ 4221], fig. 3a
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 15 | 7 | 1 | 6 | 3 | 1 | 26.5 |
+-------- Parreysiinae ** | +-------- Gonideinae ** --+ | +-- Hyrioidea | +--+ | | +-- Trigonioidea +--+ | +-- Pleurobemini ** | | +--+-- Lampsilini ** | +-- Unioninae **
Lopes-Lima et al., 2020 [Publ 4080], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 2 | 0 | 0 | 0 | 23.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Sonowal et al., 2021 [Publ 4221], fig. 3b
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 0 | 1 | 6 | 3 | 1 | 22.5 |
+-- Hyrioidea --+ +-- Unionidae **
King et al., 1999 [Publ 2699], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 1 | 0 | 0 | 0 | 24.5 |
1
Pfeiffer et al., 2021 [Publ 4126], fig. 25
| tT | bi | riS | oT | roT | roS | iD | |
| 9 | 9 | 7 | 1 | 0 | 0 | 0 | 25 |
1
Araujo et al., 2009 [Publ 2395], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 6 | 2 | 0 | 0 | 0 | 21.5 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Tomilova et al., 2020 [Publ 4097], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 5 | 1 | 0 | 0 | 0 | 22.5 |
1
Inoue et al., 2014 [Publ 3635], fig. 3B
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 3 | 1 | 0 | 0 | 0 | 21.5 |
1
Hoeh et al., 2002 [Publ 2673], fig. 1-F
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 6 | 1 | 1 | 1 | 1 | 23 |
+-- Hyrioidea --+ +-- Unionidae **
Zieritz et al., 2012 [Publ 3673], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 4 | 2 | 0 | 0 | 0 | 23 |
+----- Anodontina ** | --+ +-- Cristariina ** | | +--+-- Margaritiferidae ** | +-- Unionini **
Sheng et al., 2014 [Publ 3704], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 6 | 1 | 0 | 0 | 0 | 22 |
1
Soroka, 2010 [Publ 3404], fig. 5a
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 6 | 1 | 0 | 0 | 0 | 22.5 |
1
Pfeiffer et al., 2021 [Publ 4126], fig. 18
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 7 | 1 | 0 | 0 | 0 | 25 |
1
Hoeh et al., 2002 [Publ 2673], fig. 1-M
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 6 | 1 | 1 | 1 | 1 | 23 |
+-- Hyrioidea --+ +-- Unionidae **
Soroka, 2010 [Publ 3404], fig. 5b
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 5 | 1 | 0 | 0 | 0 | 22.5 |
1
Zieritz et al., 2026 [Publ 4812], fig. 5a
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 8 | 1 | 0 | 0 | 0 | 24 |
1
Klishko et al., 2018 [Publ 3945], fig. 8
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 5 | 1 | 0 | 0 | 0 | 23.5 |
1
Kondakov et al., 2018 [Publ 3914], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 6 | 1 | 0 | 0 | 0 | 24 |
1
Zhang et al., 2024 [Publ 4397], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 6 | 1 | 0 | 0 | 0 | 24 |
1
Lopes-Lima et al., 2020 [Publ 4080], fig. 12
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 6 | 2 | 0 | 0 | 0 | 22.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Froufe et al., 2015 [Publ 3664], fig. 1-M
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 6 | 1 | 0 | 0 | 0 | 22.5 |
1
Bolotov et al., 2016 [Publ 3732], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 7 | 2 | 0 | 0 | 0 | 23.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Lopes-Lima et al., 2020 [Publ 4080], fig. 7
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 6 | 2 | 0 | 0 | 0 | 22.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Bauer et al., 2021 [Publ 4365], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 5 | 1 | 0 | 0 | 0 | 24 |
1
Lopes-Lima et al., 2020 [Publ 4080], fig. 16
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 7 | 2 | 0 | 0 | 0 | 22.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Marrone et al., 2019 [Publ 3907], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 8 | 8 | 4 | 1 | 0 | 0 | 0 | 23.5 |
1
Amaro et al., 2017 [Publ 3791], fig. 3A-M
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 2 | 0 | 0 | 0 | 20 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Pfeiffer et al., 2021 [Publ 4126], fig. 30
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 1 | 0 | 0 | 0 | 23 |
1
Jones & Neves, 2010 [Publ 2646], fig. 17
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 1 | 1 | 0 | 0 | 0 | 23 |
1
Serb & Barnhart, 2008 [Publ 2674], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 9 | 1 | 0 | 0 | 0 | 22.5 |
1
Amaro et al., 2017 [Publ 3791], fig. 3B-M
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 2 | 0 | 0 | 0 | 20 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Bogan et al., 2023 [Publ 4295], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 2 | 0 | 0 | 0 | 22 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Soroka & Burzynski, 2018 [Publ 3762], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 1 | 0 | 0 | 0 | 22 |
1
Lopes-Lima et al., 2020 [Publ 4080], fig. 8
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 2 | 0 | 0 | 0 | 21.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Källersjö et al., 2005 [Publ 2684], fig. 1B-F
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 4 | 2 | 0 | 0 | 0 | 21 |
+-- Margaritiferidae ** | --+-- Anodontini ** | +-- Unionini **
Serb, 2006 [Publ 2678], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 7 | 1 | 0 | 0 | 0 | 22.5 |
1
Serb & Barnhart, 2008 [Publ 2674], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 10 | 1 | 0 | 0 | 0 | 22.5 |
1
Hou et al., 2025 [Publ 4445], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 1 | 0 | 0 | 0 | 23 |
1
Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 1 | 0 | 0 | 0 | 22 |
1
Serb, 2006 [Publ 2678], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 8 | 1 | 0 | 0 | 0 | 22.5 |
1
Chen et al., 2012 [Publ 3624], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 1 | 0 | 0 | 0 | 21 |
1
Soroka, 2010 [Publ 3404], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 4 | 1 | 0 | 0 | 0 | 21.5 |
1
Pfeiffer et al., 2018 [Publ 3827], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 3 | 1 | 0 | 0 | 0 | 23 |
1
Zieritz et al., 2024 [Publ 4381], fig. 8A
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 1 | 0 | 0 | 0 | 23 |
1
Froufe et al., 2016 [Publ 3727], fig. 3_4
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 3 | 1 | 0 | 0 | 0 | 23 |
1
Buhay et al., 2002 [Publ 2661], fig. 2A
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 3 | 1 | 0 | 0 | 0 | 22.5 |
1
Källersjö et al., 2005 [Publ 2684], fig. 1A
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 2 | 2 | 0 | 0 | 0 | 21 |
+-- Margaritiferidae ** | --+-- Anodontini ** | +-- Unionini **
Hoeh et al., 1995 [Publ 3391], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 4 | 1 | 0 | 0 | 0 | 22.5 |
1
Smith et al., 2023 [Publ 4291], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 1 | 0 | 0 | 0 | 23 |
1
Goncalves et al., 2022 [Publ 4253], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 6 | 1 | 0 | 0 | 0 | 23 |
1
Guerra et al., 2017 [Publ 3772], fig. 2b-M
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 4 | 2 | 1 | 1 | 1 | 20.5 |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Guerra et al., 2017 [Publ 3772], fig. 2a-M
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 4 | 2 | 1 | 1 | 1 | 20.5 |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Lopes-Lima et al., 2021 [Publ 4217], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 1 | 0 | 0 | 0 | 22 |
1
Jones & Neves, 2010 [Publ 2646], fig. 16
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 1 | 0 | 0 | 0 | 23 |
1
Serb, 2006 [Publ 2678], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 8 | 1 | 0 | 0 | 0 | 22.5 |
1
Huff et al., 2004 [Publ 2697], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 3 | 2 | 1 | 0 | 0 | 21.5 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Soroka, 2010 [Publ 3404], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 1 | 0 | 0 | 0 | 21.5 |
1
Buhay et al., 2002 [Publ 2661], fig. 2B
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 1 | 0 | 0 | 0 | 22.5 |
1
Bashê, 2022 [Publ 4265], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 5 | 2 | 0 | 0 | 0 | 20.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Nagel et al., 1998 [Publ 1946], fig. 9
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 2 | 0 | 0 | 0 | 18 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 2 | 0 | 0 | 0 | 19 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Prie et al., 2012 [Publ 3487], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 2 | 0 | 0 | 0 | 20.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Klishko et al., 2017 [Publ 3946], fig. 8
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 1 | 0 | 0 | 0 | 21.5 |
1
Bogatov et al., 2018 [Publ 3859], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 2 | 0 | 0 | 0 | 19 |
+-- Margaritiferidae ** --+ +-- Unionidae **
King et al., 1999 [Publ 2699], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 3 | 1 | 0 | 0 | 0 | 22 |
1
Zieritz et al., 2016 [Publ 3804], fig. S2-1
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 3 | 1 | 0 | 0 | 0 | 22 |
1
Lopes-Lima et al., 2020 [Publ 4080], fig. 14
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 2 | 0 | 0 | 0 | 20.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Soroka, 2010 [Publ 3404], fig. 3a-M
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 1 | 0 | 0 | 0 | 20.5 |
1
Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 1 | 0 | 0 | 0 | 22 |
1
Bogatov et al., 2018 [Publ 3859], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 2 | 0 | 0 | 0 | 19 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Kandl et al., 2001 [Publ 3384], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 1 | 0 | 0 | 0 | 21 |
1
Soroka, 2010 [Publ 3404], fig. 3b-M
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 1 | 0 | 0 | 0 | 20.5 |
1
Keogh & Simons, 2019 [Publ 3935], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 6 | 1 | 0 | 0 | 0 | 22.5 |
1
Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-M
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 1 | 0 | 0 | 0 | 21.5 |
1
Bogan et al., 2023 [Publ 4295], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 2 | 0 | 0 | 0 | 21 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Moyer & Díaz-Ferguson, 2012 [Publ 3375], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 5 | 1 | 0 | 0 | 0 | 20.5 |
1
Soroka, 2010 [Publ 3404], fig. 8a
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 1 | 0 | 0 | 0 | 20.5 |
1
Wen et al., 2017 [Publ 3795], fig. 3-M
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 1 | 0 | 0 | 0 | 20.5 |
1
Lopes-Lima et al., 2020 [Publ 4080], fig. 9
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 2 | 0 | 0 | 0 | 20.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Choi et al., 2020 [Publ 4169], fig. 3a
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 5 | 1 | 0 | 0 | 0 | 21.5 |
1
Riccardi et al., 2020 [Publ 4089], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 1 | 0 | 0 | 0 | 22 |
1
Freeman et al., 2025 [Publ 4443], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 5 | 1 | 0 | 0 | 0 | 22 |
1
Soroka, 2010 [Publ 3404], fig. 8b
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 1 | 0 | 0 | 0 | 20.5 |
1
Lopes-Lima et al., 2020 [Publ 4080], fig. 17
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 5 | 2 | 0 | 0 | 0 | 19.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Dai et al., 2024 [Publ 4358], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 1 | 0 | 0 | 0 | 22 |
1
Huang et al., 2013 [Publ 3629], fig. 3-M
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 4 | 1 | 0 | 0 | 0 | 20.5 |
1
Wei et al., 2016 [Publ 3665], fig. 0
| tT | bi | riS | oT | roT | roS | iD | |
| 6 | 6 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 11 | 9 | 2 | 6 | 6 | 1 | 23 |
+-- Margaritiferidae ** | --+-- Unionidae ** | +-- Hyrioidea
Buhay et al., 2002 [Publ 2661], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 21.5 |
1
Krebs et al., 2013 [Publ 3515], fig. 2A
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 20.5 |
1
Saxon et al., 2023 [Publ 4338], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 21 |
1
Soroka, 2010 [Publ 3404], fig. 9a
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 21 |
1
Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 2 | 0 | 0 | 0 | 19.5 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Mabuchi et al., 2021 [Publ 4233], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 21 |
1
Lane et al., 2016 [Publ 3735], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 1 | 1 | 0 | 0 | 0 | 22 |
1
Cyr et al., 2007 [Publ 2745], fig. 2C
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 4 | 1 | 0 | 0 | 0 | 23.5 |
1
Bolotov et al., 2022 [Publ 4250], fig. S3
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 21 |
1
Konopleva et al., 2017 [Publ 3807], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 21 |
1
Nagel et al., 1996 [Publ 3400], fig. 351
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 4 | 1 | 0 | 0 | 0 | 21 |
1
Lopes-Lima et al., 2021 [Publ 4217], fig. 14
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 4 | 1 | 0 | 0 | 0 | 21 |
1
Zanatta & Harris, 2013 [Publ 3530], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 19 |
1
Nagel et al., 1998 [Publ 1946], fig. 7
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 4 | 1 | 0 | 0 | 0 | 21 |
1
Cyr et al., 2007 [Publ 2745], fig. 2A
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 20 |
1
Soroka, 2005 [Publ 2726], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 20.5 |
1
Krebs, 2004 [Publ 2692], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 20 |
1
Jones et al., 2006 [Publ 2681], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 4 | 1 | 0 | 0 | 0 | 21 |
1
Soroka, 2010 [Publ 3404], fig. 2-M
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 19.5 |
1
Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5_7
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 19.5 |
1
Cyr et al., 2007 [Publ 2745], fig. 2B
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 4 | 1 | 0 | 0 | 0 | 21.5 |
1
Mulvey et al., 1997 [Publ 3390], fig. 1_3
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 1 | 0 | 0 | 0 | 20 |
1
Burzynski & Soroka, 2018 [Publ 3862], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 19.5 |
1
Klishko et al., 2016 [Publ 3733], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Soroka, 2008 [Publ 3695], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 18.5 |
1
Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 1 | 1 | 0 | 0 | 0 | 18.5 |
1
Patterson et al., 2021 [Publ 4222], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Soroka, 2008 [Publ 3695], fig. 1-M
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 18.5 |
1
Gangloff et al., 2013 [Publ 3551], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Lopes-Lima et al., 2021 [Publ 4217], fig. 11
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Curole & Kocher, 2002 [Publ 2712], fig. 1b-M
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 18.5 |
1
Zieritz et al., 2026 [Publ 4812], fig. 7a
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 1 | 1 | 0 | 0 | 0 | 20 |
1
Curole & Kocher, 2002 [Publ 2712], fig. 1-F
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 2 | 0 | 0 | 0 | 18 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Jiang et al., 2010 [Publ 3438], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 1 | 1 | 0 | 0 | 0 | 18.5 |
1
Pfeiffer et al., 2018 [Publ 3835], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Soroka, 2010 [Publ 3404], fig. 9c
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 3 | 1 | 0 | 0 | 0 | 20 |
1
Cyr et al., 2007 [Publ 2745], fig. 2D
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20.5 |
1
Klishko et al., 2014 [Publ 3666], fig. 8
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Curole & Kocher, 2002 [Publ 2712], fig. 1a-M
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 18.5 |
1
Giribet & Distel, 2003 [Publ 3670], fig. 3f5
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 1 | 2 | 2 | 0 | 0 | 16 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Froufe et al., 2017 [Publ 3802], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Sayenko et al., 2017 [Publ 3760], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Machordom et al., 2003 [Publ 2688], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 2 | 0 | 0 | 0 | 18.5 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Bulakhova et al., 2023 [Publ 4325], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Krebs, 2004 [Publ 2692], fig. 2-M
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 19 |
1
Nagel et al., 1998 [Publ 1946], fig. 8
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 1 | 1 | 0 | 0 | 0 | 19.5 |
1
Zhang et al., 2013 [Publ 4214], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 2 | 0 | 0 | 0 | 18.5 |
+-------- Cristaria ** --+ | +----- Sinanodonta ** +--+ | +-- Margaritiferidae ** +--+ +-- Gonideinae **
Soroka, 2010 [Publ 3404], fig. 9b
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Marshall et al., 2014 [Publ 3593], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 14 | 12 | 2 | 10 | 10 | 1 | 28 |
+-- Margaritiferidae **
+--+
| +-- Unioninae **
--+
| +-- Ambleminae **
+--+
+-- Hyrioidea
Mezhzherin et al., 2014 [Publ 3705], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Combosch et al., 2017 [Publ 3806], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 10 | 8 | 2 | 11 | 9 | 2 | 25.5 |
+-------- Hyrioidea --+ | +----- Unionidae ** +--+ | +-- Margaritiferidae ** +--+ +-- Etherioidea
Mezhzherin et al., 2014 [Publ 3705], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Johnson et al., 2023 [Publ 4323], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 1 | 0 | 0 | 0 | 20 |
1
Combosch et al., 2017 [Publ 3806], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 2 | 11 | 9 | 2 | 17.5 |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Machordom et al., 2003 [Publ 2688], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 2 | 0 | 0 | 0 | 17.5 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Hoeh et al., 1998 [Publ 2695], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 2 | 2 | 1 | 1 | 18.5 |
+-------- Etherioidea --+ | +----- Unioninae ** +--+ | +-- Margaritiferidae ** +--+ +-- Ambleminae **
Shirai et al., 2010 [Publ 3353], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 19 |
1
Wang et al., 2016 [Publ 3714], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 18.5 |
1
Zieritz et al., 2024 [Publ 4381], fig. 6A
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 19 |
1
Hinzmann et al., 2013 [Publ 3541], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 19 |
1
Mock et al., 2013 [Publ 3598], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 18.5 |
1
Soroka, 2010 [Publ 3404], fig. 8a-M
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 17.5 |
1
Kilikowska et al., 2020 [Publ 4335], fig. 4a
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 18.5 |
1
Hoeh et al., 1996 [Publ 2694], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 1 | 0 | 0 | 17.5 |
+----- Unionidae ** --+ | +-- None__1 +--+ +-- None__2
Soroka, 2010 [Publ 3404], fig. 8b-M
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 17.5 |
1
Froufe et al., 2014 [Publ 3592], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 19 |
1
Campbell, 2000 [Publ 2724], fig. 1-5
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 2 | 0 | 0 | 0 | 16 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Hoeh et al., 1998 [Publ 2695], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 2 | 2 | 1 | 1 | 18.5 |
+-------- Etherioidea --+ | +----- Ambleminae ** +--+ | +-- Margaritiferidae ** +--+ +-- Unioninae **
Imsong & Murali, 2023 [Publ 4370], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 18.5 |
1
Metzger et al., 2018 [Publ 3952], fig. 1C
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 18 |
1
Xia et al., 2016 [Publ 3712], fig. 7B
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 19 |
1
Peng et al., 2012 [Publ 3689], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 19 |
1
Lopes-Lima et al., 2016 [Publ 3757], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 19 |
1
Välilä et al., 2017 [Publ 3793], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 2 | 0 | 0 | 0 | 17.5 |
+-- Unionidae ** --+ +-- Margaritiferidae **
Taylor et al., 2007 [Publ 2696], fig. 1-3
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 2 | 1 | 0 | 0 | 16 |
+-- Margaritiferidae ** --+ +-- Unionidae **
Smith & Johnson, 2020 [Publ 4229], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 19 |
1
Krebs et al., 2019 [Publ 4092], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 1 | 0 | 0 | 0 | 18.5 |
1
González & Giribet, 2015 [Publ 3632], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 4 | 0 | 2 | 4 | 2 | 2 | 19.5 |
+-- Etherioidea
+--+
| +-- Margaritiferidae **
--+
| +-- Unionidae **
+--+
+-- Hyrioidea
Soroka, 2010 [Publ 3404], fig. 5a-M
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 0 | 0 | 0 | -- |
1
Lemer et al., 2019 [Publ 4093], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 4 | 2 | 1 | -- |
+----- Etherioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Bieler et al., 2014 [Publ 3622], fig. 28
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 4 | 0 | 2 | 3 | 2 | 2 | 19.5 |
+-- Etherioidea
+--+
| +-- Margaritiferidae **
--+
| +-- Unionidae **
+--+
+-- Hyrioidea
González & Giribet, 2015 [Publ 3632], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 3 | 0 | 2 | 4 | 2 | 2 | 18 |
+-------- Hyrioidea --+ | +----- Unionidae ** +--+ | +-- Etherioidea +--+ +-- Margaritiferidae **
Plazzi & Passamonti, 2010 [Publ 3354], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 0 | 0 | 0 | -- |
1
Graf et al., 2015 [Publ 3619], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 18 | 17 | 2 | -- |
+-------- Hyrioidea --+ | +----- Etherioidea +--+ | +-- Unionidae ** +--+ +-- Margaritiferidae **
Mock et al., 2004 [Publ 2686], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 0 | 0 | 0 | -- |
1
González & Giribet, 2015 [Publ 3632], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 4 | 2 | 2 | -- |
+-------- Etherioidea --+ | +----- Hyrioidea +--+ | +-- Unionidae ** +--+ +-- Margaritiferidae **
Klunzinger et al., 2021 [Publ 4086], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 4 | 4 | 1 | -- |
+----- Hyrioidea --+ | +-- Margaritiferidae ** +--+ +-- Unionidae **
Mock et al., 2004 [Publ 2686], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 0 | 0 | 0 | -- |
1
Bieler et al., 2014 [Publ 3622], fig. 26_27_29_30
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 3 | 0 | 2 | 3 | 2 | 2 | 18 |
+-------- Hyrioidea --+ | +----- Unionidae ** +--+ | +-- Etherioidea +--+ +-- Margaritiferidae **
González & Giribet, 2015 [Publ 3632], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 4 | 0 | 2 | 4 | 2 | 2 | 18.5 |
+----- Unionidae ** | --+----- Hyrioidea | | +-- Etherioidea +--+ +-- Margaritiferidae **
Soroka, 2010 [Publ 3404], fig. 4-M
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 0 | 0 | 0 | -- |
1
Xue et al., 2016 [Publ 3659], fig. 1-M
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 0 | 0 | 0 | -- |
1
Soroka, 2010 [Publ 3404], fig. 5b-M
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 1 | 0 | 0 | 0 | -- |
1
Baker et al., 2003 [Publ 2690], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 7 | 7 | 1 | -- |
+-- Unionidae ** --+ +-- Hyrioidea
Baker et al., 2004 [Publ 2685], fig. 2-3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 8 | 8 | 1 | -- |
+-- Unionidae ** --+ +-- Hyrioidea
