Bivalvia | Unionida

superfamily UNIONOIDEA Rafinesque, 1820: cladomics

2 families

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 816 (1114 total)
number of subtaxa (rank: family) = 2
analyzed in 575 cladograms in 320 publications
maximum number of terminals in an analysis = 175 (mean = 22.66, median = 14)
maximum number of subtaxa (family) = 2 (mean = 1.45, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 573
maximum number of included terminals = 175, (mean = 22.73 median = 14)
maximum number of subtaxa (family) = 2 (mean = 1.46, median = 1)
proportion of cladograms that support basic monophyly = 0.94

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 91
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (family) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 175 (mean = 37.07, median = 22)
maximum number of subtaxa (family) = 2 (mean = 2, median = 2)
proportion of cladograms that support rigorous monophyly = 0.69

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Unionoidea cladogram distances
PCoA Ordination

NJ Tree of Unionoidea cladogram distances
NJ Tree

Cladograms

Graf, 2000 [Publ 2647], fig. 1

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
7 16 14 2 10 9 2 0 
  +-------------- Margaritiferidae **
  |                               
  |     +-------- Unionini **        
--+  +--+                         
  |  |  |  +----- Pleurobemini **    
  |  |  +--+                      
  |  |     |  +-- Anodontini **      
  +--+     +--+                   
     |        +-- Lampsilini **      
     |                            
     |  +-------- Parreysiini **     
     +--+                         
        |  +----- Coelaturini **     
        +--+                      
           |  +-- Hyrioidea       
           +--+                   
              +-- Etherioidea

Huang et al., 2019 [Publ 3843], fig. S3

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
173 173 171 2 26 24 2 188 
  +-------- Hyrioidea       
--+                         
  |  +----- Etherioidea     
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
175 175 111 2 26 24 2 189 
  +-------- Etherioidea     
--+                         
  |  +----- Hyrioidea       
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
175 175 155 2 26 24 2 191 
  +-------- Hyrioidea       
--+                         
  |  +----- Etherioidea     
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
125 125 72 1 0 0 0 142.5 
1

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
124 137 137 2 15 13 2 151 
  +-------- Margaritiferidae **
--+                         
  |  +----- Unionidae **       
  +--+                      
     |  +-- Etherioidea     
     +--+                   
        +-- Hyrioidea

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
124 137 123 2 15 13 2 150 
                       +-- Chamberlainia **   
  +--------------------+                   
  |                    +-- Margaritiferidae **
  |                                        
  |                    +-- Etherioidea     
--+--------------------+                   
  |                    +-- Hyrioidea       
  |                                        
  |  +-------------------- Parreysiinae **    
  +--+                                     
     |  +----------------- Unioninae **       
     +--+                                  
        |  +-------------- Rectidentini **    
        +--+                               
           |  +----------- Ambleminae **      
           +--+                            
              |  +-------- Pseudodontini **   
              +--+                         
                 |  +----- Gonideini **       
                 +--+                      
                    |  +-- Sinohyriopsis **   
                    +--+                   
                       +-- Lamprotulini **

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
118 118 100 2 0 0 0 132 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
113 118 100 2 9 7 2 132.5 
  +-------- Hyrioidea       
--+                         
  |  +----- Margaritiferidae **
  +--+                      
     |  +-- Etherioidea     
     +--+                   
        +-- Unionidae **

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
112 112 100 1 0 0 0 127.5 
1

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iTtTbiriSoTroTroSiD
104 104 63 2 0 0 0 202.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Campbell et al., 2005 [Publ 2660], fig. 2

iTtTbiriSoTroTroSiD
102 102 92 2 0 0 0 124 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Campbell et al., 2005 [Publ 2660], fig. 1

iTtTbiriSoTroTroSiD
102 102 103 2 0 0 0 125 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
92 92 90 2 4 3 2 105.5 
  +-------- Hyrioidea       
--+                         
  |  +----- Etherioidea     
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
92 92 87 2 4 3 2 105.5 
  +-------- Hyrioidea       
--+                         
  |  +----- Etherioidea     
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
88 88 85 2 4 3 2 101.5 
  +-------- Hyrioidea       
--+                         
  |  +----- Etherioidea     
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
88 88 84 2 4 3 2 101.5 
  +-------- Hyrioidea       
--+                         
  |  +----- Etherioidea     
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
85 85 83 2 3 3 2 97.5 
  +-------- Etherioidea     
--+                         
  |  +----- Hyrioidea       
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
83 83 80 1 0 0 0 99 
1

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
83 83 50 1 0 0 0 96.5 
1

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
83 84 82 2 3 3 2 97.5 
  +-------- Etherioidea     
--+                         
  |  +----- Unionidae **       
  +--+                      
     |  +-- Hyrioidea       
     +--+                   
        +-- Margaritiferidae **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
81 81 79 2 10 8 2 92.5 
  +-------- Hyrioidea       
--+                         
  |  +----- Etherioidea     
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
81 81 79 2 10 8 2 92.5 
  +-------- Etherioidea     
--+                         
  |  +----- Hyrioidea       
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
76 76 74 2 0 0 0 88 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
76 76 73 2 0 0 0 88.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
76 76 74 2 0 0 0 88.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Campbell & Lydeard, 2012 [Publ 3309], fig. 3

iTtTbiriSoTroTroSiD
73 73 67 1 0 0 0 86 
1

Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2

iTtTbiriSoTroTroSiD
73 73 63 1 0 0 0 86 
1

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
71 77 71 2 9 8 2 88 
  +-------- Hyrioidea       
--+                         
  |  +----- Margaritiferidae **
  +--+                      
     |  +-- Etherioidea     
     +--+                   
        +-- Unionidae **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
71 71 69 2 3 3 2 83.5 
  +-------- Etherioidea     
--+                         
  |  +----- Hyrioidea       
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
70 70 68 2 4 3 2 82.5 
  +-------- Hyrioidea       
--+                         
  |  +----- Etherioidea     
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
70 70 68 2 4 3 2 82.5 
  +-------- Hyrioidea       
--+                         
  |  +----- Etherioidea     
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Wang et al., 2025 [Publ 4488], fig. 3B

iTtTbiriSoTroTroSiD
65 65 63 2 0 0 0 76.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
65 65 63 2 4 3 2 78 
  +-------- Hyrioidea       
--+                         
  |  +----- Etherioidea     
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Bolotov et al., 2024 [Publ 4369], fig. S1

iTtTbiriSoTroTroSiD
63 63 61 1 0 0 0 76.5 
1

Bolotov et al., 2024 [Publ 4369], fig. 2

iTtTbiriSoTroTroSiD
62 62 60 1 0 0 0 75.5 
1

Bolotov et al., 2022 [Publ 4252], fig. S1

iTtTbiriSoTroTroSiD
61 61 49 1 0 0 0 74.5 
1

Bolotov et al., 2022 [Publ 4252], fig. 2

iTtTbiriSoTroTroSiD
61 61 55 1 0 0 0 74.5 
1

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iTtTbiriSoTroTroSiD
60 60 59 1 0 0 0 76 
1

Bolotov et al., 2022 [Publ 4239], fig. S1

iTtTbiriSoTroTroSiD
58 58 52 1 0 0 0 71.5 
1

Bolotov et al., 2022 [Publ 4239], fig. S2

iTtTbiriSoTroTroSiD
58 58 54 1 0 0 0 71.5 
1

Bolotov et al., 2022 [Publ 4239], fig. 2

iTtTbiriSoTroTroSiD
58 58 54 1 0 0 0 71.5 
1

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
57 57 60 2 11 9 2 70 
  +-------- Hyrioidea       
--+                         
  |  +----- Etherioidea     
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
57 57 59 2 11 9 2 70 
  +-------- Hyrioidea       
--+                         
  |  +----- Etherioidea     
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
57 57 60 2 11 9 2 70 
  +-------- Hyrioidea       
--+                         
  |  +----- Etherioidea     
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Zanatta & Murphy, 2006 [Publ 2672], fig. 2

iTtTbiriSoTroTroSiD
57 57 48 2 0 0 0 71 
  +-- Unioninae **       
  |                   
--+-- Margaritiferidae **
  |                   
  +-- Ambleminae **

Zanatta & Murphy, 2006 [Publ 2672], fig. 1

iTtTbiriSoTroTroSiD
56 56 51 2 0 0 0 69 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
56 56 54 2 0 0 0 67.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
54 54 52 2 0 0 0 65.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
54 54 52 2 0 0 0 67 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
53 53 51 2 0 0 0 67 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
53 53 45 2 0 0 0 65 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Hou et al., 2025 [Publ 4445], fig. 5

iTtTbiriSoTroTroSiD
53 53 51 1 0 0 0 66.5 
1

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
53 53 51 2 0 0 0 65.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
53 53 51 2 0 0 0 65.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
52 52 50 2 4 3 2 64.5 
  +-------- Etherioidea     
--+                         
  |  +----- Hyrioidea       
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
52 53 50 2 5 4 2 69 
  +-------- Etherioidea     
--+                         
  |  +----- Unionidae **       
  +--+                      
     |  +-- Hyrioidea       
     +--+                   
        +-- Margaritiferidae **

Konopleva et al., 2019 [Publ 3900], fig. 1

iTtTbiriSoTroTroSiD
52 52 41 2 0 0 0 65.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
52 52 50 2 5 4 2 65.5 
  +-------- Etherioidea     
--+                         
  |  +----- Hyrioidea       
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Jeratthitikul et al., 2025 [Publ 4423], fig. 1

iTtTbiriSoTroTroSiD
51 51 49 1 0 0 0 66 
1

Chen et al., 2025 [Publ 4663], fig. 1

iTtTbiriSoTroTroSiD
49 49 47 2 0 0 0 62.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Bolotov et al., 2022 [Publ 4250], fig. 8

iTtTbiriSoTroTroSiD
49 49 42 2 0 0 0 62 
     +----- Parreysiinae **    
  +--+                      
  |  |  +-- Pseudodontini **   
  |  +--+                   
  |     +-- Rectidentini **    
  |                         
--+     +-- Gibbosulinae **    
  |     |                   
  |     +-- Margaritiferinae **
  |     |                   
  |     +-- Sinosolenaia **    
  |     |                   
  +-----+-- Leguminaia **      
        |                   
        +-- Gonidea **         
        |                   
        +-- Lamprotula **      
        |                   
        +-- Potomida **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
48 48 46 2 0 0 0 60 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
48 48 46 2 0 0 0 61.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
47 47 45 2 0 0 0 60.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Huang et al., 2019 [Publ 3843], fig. S1

iTtTbiriSoTroTroSiD
47 47 55 2 0 0 0 78.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
46 46 44 2 0 0 0 59.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Konopleva et al., 2025 [Publ 4446], fig. 4

iTtTbiriSoTroTroSiD
45 45 43 1 0 0 0 60 
1

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
45 45 41 2 0 0 0 58.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
45 45 38 2 0 0 0 57 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
45 45 38 2 0 0 0 57 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
45 45 44 2 0 0 0 59.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
45 45 43 2 0 0 0 58.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
45 45 43 2 4 3 2 57 
  +-------- Hyrioidea       
--+                         
  |  +----- Etherioidea     
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
44 44 42 2 0 0 0 56 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
44 46 44 2 2 2 2 58.5 
  +-------- Unionidae **       
--+                         
  |  +----- Margaritiferidae **
  +--+                      
     |  +-- Hyrioidea       
     +--+                   
        +-- Etherioidea

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
44 44 42 2 0 0 0 56 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Campbell & Lydeard, 2012 [Publ 3308], fig. 3

iTtTbiriSoTroTroSiD
43 43 42 1 0 0 0 57 
1

Inoue et al., 2020 [Publ 3941], fig. 3

iTtTbiriSoTroTroSiD
42 42 42 1 0 0 0 56.5 
1

Campbell & Lydeard, 2012 [Publ 3308], fig. 1

iTtTbiriSoTroTroSiD
42 42 41 1 0 0 0 55 
1

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
42 42 40 1 0 0 0 55.5 
1

Campbell & Lydeard, 2012 [Publ 3308], fig. 4

iTtTbiriSoTroTroSiD
42 42 40 1 0 0 0 55 
1

Campbell & Lydeard, 2012 [Publ 3308], fig. 2

iTtTbiriSoTroTroSiD
42 42 36 1 0 0 0 55 
1

Inoue et al., 2020 [Publ 3941], fig. 4

iTtTbiriSoTroTroSiD
41 41 41 1 0 0 0 55.5 
1

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
41 41 37 2 0 0 0 53 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
41 41 39 2 0 0 0 53 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Smith et al., 2019 [Publ 3951], fig. 2

iTtTbiriSoTroTroSiD
41 41 40 1 0 0 0 55.5 
1

Bolotov et al., 2023 [Publ 4312], fig. S2

iTtTbiriSoTroTroSiD
41 41 39 1 0 0 0 57 
1

Burlakova et al., 2012 [Publ 3486], fig. 1

iTtTbiriSoTroTroSiD
41 41 42 1 0 0 0 56 
1

Whelan et al., 2023 [Publ 4287], fig. 2

iTtTbiriSoTroTroSiD
41 41 42 1 0 0 0 57.5 
1

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
41 41 32 2 0 0 0 53 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
41 41 39 2 0 0 0 53 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
40 40 38 2 0 0 0 52 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
39 39 37 1 0 0 0 52 
1

Bolotov et al., 2023 [Publ 4312], fig. S1

iTtTbiriSoTroTroSiD
39 39 37 1 0 0 0 54 
1

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
39 39 37 2 0 0 0 52 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Bolotov et al., 2023 [Publ 4312], fig. 2

iTtTbiriSoTroTroSiD
39 39 38 1 0 0 0 55 
1

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
38 38 36 2 4 3 2 50 
  +-------- Etherioidea     
--+                         
  |  +----- Hyrioidea       
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
38 38 36 2 0 0 0 50 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Boyer et al., 2011 [Publ 2718], fig. 4

iTtTbiriSoTroTroSiD
38 38 33 1 0 0 0 51.5 
1

Pfeiffer et al., 2016 [Publ 3660], fig. 2

iTtTbiriSoTroTroSiD
38 38 34 1 0 0 0 52 
1

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
37 37 35 2 0 0 0 50 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
37 37 35 2 0 0 0 50.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Chase et al., 2018 [Publ 3847], fig. 4

iTtTbiriSoTroTroSiD
37 38 37 2 1 1 1 52 
  +-------------- Margaritiferidae **
  |                               
--+  +----------- Unioninae **       
  |  |                            
  +--+  +-------- Lampsilini **      
     |  |                         
     +--+     +-- Hyrioidea       
        |  +--+                   
        |  |  +-- Amblemini **       
        +--+                      
           |  +-- Pleurobemini **    
           +--+                   
              +-- Quadrulini **

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
37 37 35 2 0 0 0 50.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
37 37 35 2 0 0 0 50.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
37 37 35 2 0 0 0 50.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
37 37 25 2 0 0 0 51.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
37 37 35 2 0 0 0 50.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Burlakova et al., 2012 [Publ 3486], fig. 3

iTtTbiriSoTroTroSiD
36 36 26 1 0 0 0 52.5 
1

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 1

iTtTbiriSoTroTroSiD
36 36 22 1 0 0 0 50.5 
1

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 2

iTtTbiriSoTroTroSiD
36 36 20 1 0 0 0 50.5 
1

Breton et al., 2011 [Publ 3376], fig. 3

iTtTbiriSoTroTroSiD
36 36 35 2 1 1 1 49.5 
  +----- Hyrioidea       
--+                      
  |  +-- Margaritiferidae **
  +--+                   
     +-- Unionidae **

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iTtTbiriSoTroTroSiD
36 36 34 1 0 0 0 51 
1

Perkins et al., 2017 [Publ 3777], fig. 2

iTtTbiriSoTroTroSiD
36 36 34 1 0 0 0 51.5 
1

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
36 36 22 2 0 0 0 50 
  +----- Margaritiferidae **
  |                      
--+----- Ambleminae **      
  |                      
  |  +-- Gonideinae **      
  +--+                   
     +-- Unioninae **

Perkins et al., 2025 [Publ 4447], fig. 2_mtDNA-ITS

iTtTbiriSoTroTroSiD
36 36 37 1 0 0 0 53.5 
1

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iTtTbiriSoTroTroSiD
35 35 33 1 0 0 0 51 
1

Wu et al., 2025 [Publ 4442], fig. 2

iTtTbiriSoTroTroSiD
35 35 33 1 0 0 0 49 
1

Boyer et al., 2011 [Publ 2718], fig. 3

iTtTbiriSoTroTroSiD
35 35 26 1 0 0 0 48.5 
1

Burlakova et al., 2012 [Publ 3486], fig. 2

iTtTbiriSoTroTroSiD
35 35 30 1 0 0 0 53.5 
1

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
34 34 15 2 0 0 0 48 
  +-- Cumberlandia ** 
  |                
--+-- Margaritifera **
  |                
  +-- Unionidae **

Dai et al., 2026 [Publ 4757], fig. 3

iTtTbiriSoTroTroSiD
34 34 33 2 0 0 0 46.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
34 34 32 2 0 0 0 49 
                       +----- Prolasmidonta **           
                    +--+                              
                    |  |  +-- Alasmidonta **             
                    |  +--+                           
  +-----------------+     +-- Anemina **                 
  |                 |                                 
  |                 |  +----- Pseudodontini **           
  |                 +--+                              
  |                    |  +-- Lamprotula_caveata **      
  |                    +--+                           
  |                       +-- Chamberlainia **           
  |                                                   
  |                    +----- Oxynaiina **               
  |              +-----+                              
  |              |     |  +-- Lepidodesmini **           
--+              |     +--+                           
  |              |        +-- Aculamprotulina **         
  |  +-----------+                                    
  |  |           |     +----- Platynaias **              
  |  |           |  +--+                              
  |  |           |  |  |  +-- Utterbackia_imbecillis **  
  |  |           |  |  +--+                           
  |  |           +--+     +-- Pyganodon **               
  |  |              |                                 
  |  |              |  +----- Sinanodonta **             
  |  |              +--+                              
  |  |                 |  +-- Cristaria **               
  +--+                 +--+                           
     |                    +-- Anodontina **              
     |                                                
     |                    +-- Utterbackia_peninsularis **
     |              +-----+                           
     |              |     +-- Potamilus_alatus **        
     |  +-----------+                                 
     |  |           |  +----- Lamprotula_leaii **        
     |  |           +--+                              
     |  |              |  +-- Sinohyriopsis **           
     +--+              +--+                           
        |                 +-- Gonideini **               
        |                                             
        |  +----------------- Margaritiferidae **        
        +--+                                          
           |  +-------------- Quadrulini **              
           +--+                                       
              |  +----------- Toxolasma **               
              +--+                                    
                 |  +-------- Lampsilis_ornata **        
                 +--+                                 
                    |  +----- Potamilus_leptodon **      
                    +--+                              
                       |  +-- Lampsilis_siliquoidea **   
                       +--+                           
                          +-- Lampsilis_powellii **

Konopleva et al., 2019 [Publ 4041], fig. 3

iTtTbiriSoTroTroSiD
34 34 21 1 0 0 0 50 
1

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
34 34 31 2 0 0 0 48 
  +-- Cumberlandia ** 
  |                
--+-- Margaritifera **
  |                
  +-- Unionidae **

Burlakova et al., 2019 [Publ 3899], fig. 1

iTtTbiriSoTroTroSiD
33 33 21 1 0 0 0 46 
1

Krebs et al., 2003 [Publ 2691], fig. 3

iTtTbiriSoTroTroSiD
33 33 29 2 0 0 0 46.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
33 33 16 2 1 0 0 56.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
32 32 30 2 0 0 0 44 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
32 32 30 2 0 0 0 45 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
32 32 30 2 8 7 2 43 
  +-------- Hyrioidea       
--+                         
  |  +----- Etherioidea     
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Lyubas et al., 2022 [Publ 4255], fig. 7

iTtTbiriSoTroTroSiD
31 31 28 1 0 0 0 43 
1

Pfeiffer et al., 2019 [Publ 3866], fig. 1

iTtTbiriSoTroTroSiD
31 31 30 1 0 0 0 44 
1

Ouyang et al., 2015 [Publ 4493], fig. 2

iTtTbiriSoTroTroSiD
31 31 38 1 0 0 0 54 
1

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
31 31 29 1 0 0 0 44.5 
1

Lyubas et al., 2022 [Publ 4255], fig. S1

iTtTbiriSoTroTroSiD
31 31 22 1 0 0 0 43 
1

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
31 31 29 2 0 0 0 43 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Bogan et al., 2023 [Publ 4295], fig. 3

iTtTbiriSoTroTroSiD
31 31 28 2 0 0 0 43.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Dai et al., 2024 [Publ 4385], fig. 3-ML

iTtTbiriSoTroTroSiD
31 31 29 2 0 0 0 43.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
31 31 29 1 0 0 0 44.5 
1

Sano et al., 2017 [Publ 3799], fig. 2

iTtTbiriSoTroTroSiD
31 31 32 2 2 0 0 45.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Zieritz et al., 2020 [Publ 4102], fig. 4

iTtTbiriSoTroTroSiD
31 31 27 1 0 0 0 46 
1

Dai et al., 2024 [Publ 4385], fig. 3-BI

iTtTbiriSoTroTroSiD
31 31 28 2 0 0 0 43.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Ouyang et al., 2015 [Publ 4493], fig. 1

iTtTbiriSoTroTroSiD
31 31 29 1 0 0 0 46 
1

Burdick & White, 2007 [Publ 2679], fig. 3

iTtTbiriSoTroTroSiD
30 30 31 1 0 0 0 46 
1

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
30 30 29 2 0 0 0 42.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
30 31 29 2 1 1 1 44.5 
  +-- Margaritiferidae **
  |                   
--+-- Hyrioidea       
  |                   
  +-- Unionidae **

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

iTtTbiriSoTroTroSiD
29 29 27 2 0 0 0 41.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Pfeiffer et al., 2021 [Publ 4126], fig. 9

iTtTbiriSoTroTroSiD
29 29 27 1 0 0 0 44 
1

Serb et al., 2003 [Publ 2659], fig. 2

iTtTbiriSoTroTroSiD
29 29 20 1 0 0 0 42 
1

Serb et al., 2003 [Publ 2659], fig. 3

iTtTbiriSoTroTroSiD
29 29 27 1 0 0 0 42 
1

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

iTtTbiriSoTroTroSiD
28 28 25 2 0 0 0 40 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Pfeiffer et al., 2018 [Publ 3827], fig. 2

iTtTbiriSoTroTroSiD
28 28 26 2 0 0 0 40 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
28 28 26 2 13 12 2 40.5 
  +-------- Etherioidea     
--+                         
  |  +----- Hyrioidea       
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Smith et al., 2020 [Publ 4129], fig. 1

iTtTbiriSoTroTroSiD
28 28 26 1 0 0 0 41.5 
1

Hou et al., 2025 [Publ 4401], fig. 4

iTtTbiriSoTroTroSiD
28 28 26 1 0 0 0 43 
1

Whelan et al., 2023 [Publ 4287], fig. 3

iTtTbiriSoTroTroSiD
28 28 27 1 0 0 0 42.5 
1

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
28 28 26 2 13 12 2 40.5 
  +-------- Etherioidea     
--+                         
  |  +----- Hyrioidea       
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Smith et al., 2020 [Publ 4129], fig. 2-3

iTtTbiriSoTroTroSiD
28 28 26 1 0 0 0 41.5 
1

Xiang et al., 2025 [Publ 4433], fig. 6

iTtTbiriSoTroTroSiD
28 28 26 1 0 0 0 42 
1

Hou et al., 2025 [Publ 4432], fig. 4

iTtTbiriSoTroTroSiD
28 28 26 1 0 0 0 43 
1

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
27 27 21 2 20 19 2 40 
     +-- Hyrioidea       
  +--+                   
  |  +-- Etherioidea     
--+                      
  |  +-- Margaritiferidae **
  +--+                   
     +-- Unionidae **

Konopleva et al., 2021 [Publ 4213], fig. 1

iTtTbiriSoTroTroSiD
27 27 22 1 0 0 0 43 
1

Neemuchwala et al., 2023 [Publ 4292], fig. 2

iTtTbiriSoTroTroSiD
27 27 25 1 0 0 0 40.5 
1

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iTtTbiriSoTroTroSiD
27 27 25 2 3 2 2 38 
  +-------- Etherioidea     
--+                         
  |  +----- Hyrioidea       
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Whelan et al., 2023 [Publ 4287], fig. 4

iTtTbiriSoTroTroSiD
27 27 25 1 0 0 0 41.5 
1

Lydeard et al., 1996 [Publ 2656], fig. 4

iTtTbiriSoTroTroSiD
27 27 25 2 1 0 0 40 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
27 28 26 2 20 19 2 41 
  +-------------------------- Hyrioidea       
  |                                           
  |                 +-------- Iridinidae      
--+  +--------------+                         
  |  |              |  +----- Etheria         
  |  |              +--+                      
  |  |                 |  +-- Mulleria        
  +--+                 +--+                   
     |                    +-- Mycetopodidae   
     |                                        
     |  +-------------------- Margaritiferidae **
     +--+                                     
        |  +----------------- Parreysiinae **    
        +--+                                  
           |  +-------------- Pseudomulleria  
           +--+                               
              |  +----------- Unioninae **       
              +--+                            
                 |  +-------- Rectidentini **    
                 +--+                         
                    |  +----- Ambleminae **      
                    +--+                      
                       |  +-- Gonideini **       
                       +--+                   
                          +-- Pseudodontini **

Lydeard et al., 1996 [Publ 2656], fig. 3

iTtTbiriSoTroTroSiD
27 27 23 2 0 0 0 40 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iTtTbiriSoTroTroSiD
27 27 21 2 3 2 2 38 
  +-------- Etherioidea     
--+                         
  |  +----- Hyrioidea       
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Zhang et al., 2024 [Publ 4397], fig. 5

iTtTbiriSoTroTroSiD
27 27 25 2 0 0 0 40.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Campbell et al., 2008 [Publ 2675], fig. 6

iTtTbiriSoTroTroSiD
26 26 23 1 0 0 0 40.5 
1

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

iTtTbiriSoTroTroSiD
26 26 24 2 0 0 0 38.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Campbell et al., 2008 [Publ 2675], fig. 2

iTtTbiriSoTroTroSiD
26 26 18 1 0 0 0 41.5 
1

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
26 31 30 2 10 9 2 44 
  +-------- Hyrioidea       
--+                         
  |  +----- Margaritiferidae **
  +--+                      
     |  +-- Etherioidea     
     +--+                   
        +-- Unionidae **

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
26 45 39 2 20 19 2 58 
  +----------------- Parreysiinae **    
  |                                  
  |           +----- Margaritiferidae **
--+  +--------+                      
  |  |        |  +-- Hyrioidea       
  |  |        +--+                   
  +--+           +-- Etherioidea     
     |                               
     |  +----------- Pseudodontini **   
     +--+                            
        |  +-------- Rectidentini **    
        +--+                         
           |  +----- Unioninae **       
           +--+                      
              |  +-- Gonideini **       
              +--+                   
                 +-- Ambleminae **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
26 30 27 2 10 9 2 44.5 
  +-------- Etherioidea     
--+                         
  |  +----- Unionidae **       
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Hyrioidea

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
26 27 24 2 20 19 2 42 
  +-------------------------- Hyrioidea       
  |                                           
  |                       +-- Etheria         
  |                    +--+                   
--+                    |  +-- Mulleria        
  |  +-----------------+                      
  |  |                 |  +-- Mycetopodidae   
  |  |                 +--+                   
  +--+                    +-- Iridinidae      
     |                                        
     |  +-------------------- Parreysiinae **    
     +--+                                     
        |  +----------------- Pseudomulleria  
        +--+                                  
           |  +-------------- Margaritiferidae **
           +--+                               
              |  +----------- Pseudodontini **   
              +--+                            
                 |  +-------- Rectidentini **    
                 +--+                         
                    |  +----- Unioninae **       
                    +--+                      
                       |  +-- Gonideini **       
                       +--+                   
                          +-- Ambleminae **

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iTtTbiriSoTroTroSiD
25 25 23 1 0 0 0 39.5 
1

Lopes-Lima et al., 2021 [Publ 4217], fig. 1

iTtTbiriSoTroTroSiD
25 25 23 1 0 0 0 38.5 
1

Campbell et al., 2008 [Publ 2675], fig. 5

iTtTbiriSoTroTroSiD
25 25 6 1 0 0 0 40.5 
1

Campbell et al., 2008 [Publ 2675], fig. 4

iTtTbiriSoTroTroSiD
25 25 9 1 0 0 0 39.5 
1

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
24 24 22 1 0 0 0 37 
1

Wu et al., 2021 [Publ 4215], fig. 3

iTtTbiriSoTroTroSiD
24 24 22 1 0 0 0 38 
1

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

iTtTbiriSoTroTroSiD
24 28 29 1 4 4 1 53.5 
  +----- Parreysiinae **
--+                  
  |  +-- Etherioidea 
  +--+               
     +-- Unioninae **

Perkins et al., 2017 [Publ 3777], fig. 3

iTtTbiriSoTroTroSiD
24 24 10 1 0 0 0 41.5 
1

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
24 24 17 1 0 0 0 37 
1

Inoue et al., 2020 [Publ 4128], fig. 2

iTtTbiriSoTroTroSiD
23 23 16 1 0 0 0 39.5 
1

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
23 23 21 1 0 0 0 36 
1

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

iTtTbiriSoTroTroSiD
23 23 21 1 0 0 0 36.5 
1

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
23 23 21 2 0 0 0 36 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iTtTbiriSoTroTroSiD
23 23 21 1 0 0 0 36 
1

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
23 23 21 2 8 6 2 36.5 
  +-------- Hyrioidea       
--+                         
  |  +----- Etherioidea     
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
23 23 21 2 11 9 2 35.5 
  +-------- Hyrioidea       
--+                         
  |  +----- Etherioidea     
  +--+                      
     |  +-- Unionidae **       
     +--+                   
        +-- Margaritiferidae **

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
23 23 21 2 11 9 2 35.5 
  +-------- Hyrioidea       
--+                         
  |  +----- Etherioidea     
  +--+                      
     |  +-- Unionidae **       
     +--+                   
        +-- Margaritiferidae **

Wu et al., 2024 [Publ 4379], fig. 4_6

iTtTbiriSoTroTroSiD
23 23 21 2 0 0 0 35.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
22 22 20 1 0 0 0 35.5 
1

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

iTtTbiriSoTroTroSiD
22 22 20 1 0 0 0 35.5 
1

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
22 22 16 2 0 0 0 33 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
22 22 20 2 1 1 1 34.5 
  +----- Hyrioidea       
--+                      
  |  +-- Margaritiferidae **
  +--+                   
     +-- Unionidae **

Inoue et al., 2020 [Publ 4128], fig. 3

iTtTbiriSoTroTroSiD
22 22 24 1 0 0 0 37.5 
1

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
22 22 19 2 0 0 0 33 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

iTtTbiriSoTroTroSiD
22 22 20 1 0 0 0 35.5 
1

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
21 21 19 2 0 0 0 33.5 
  +-------------------- Margaritiferinae **       
  |                                            
  |  +----------------- Anodontini **             
--+  |                                         
  |  |     +----------- Chamberlainiini **        
  |  |  +--+                                   
  +--+  |  |  +-------- Lamprotula_leaii **       
     |  |  +--+                                
     |  |     |  +----- Lamprotula_caveata **     
     |  |     +--+                             
     |  |        |  +-- Gibbosulinae **           
     +--+        +--+                          
        |           +-- Lamprotula_cornuumlunae **
        |                                      
        |        +----- Aculamprotulina **        
        |     +--+                             
        |     |  |  +-- Tchangsinaia **           
        |     |  +--+                          
        +-----+     +-- Nodularia **              
              |                                
              |  +----- Schistodesmus **          
              +--+                             
                 |  +-- Lanceolariini **          
                 +--+                          
                    +-- Acuticostini **

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
21 21 19 2 4 3 2 33 
  +-------- Etherioidea     
--+                         
  |  +----- Hyrioidea       
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Rosenberg et al., 1997 [Publ 2720], fig. 2A

iTtTbiriSoTroTroSiD
21 22 4 2 1 0 0 31.5 
  +----- Cumberlandia ** 
  |                   
  +----- None__2      
--+                   
  +----- Margaritifera **
  |                   
  |  +-- None__1      
  +--+                
     +-- Unionidae **

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iTtTbiriSoTroTroSiD
21 22 3 2 1 0 0 31.5 
  +-- Cumberlandia ** 
  |                
  +-- None__1      
  |                
--+-- None__2      
  |                
  +-- Margaritifera **
  |                
  +-- Unionidae **

Chapman et al., 2008 [Publ 2667], fig. 1

iTtTbiriSoTroTroSiD
21 21 19 1 0 0 0 34.5 
1

Jeratthitikul et al., 2021 [Publ 4212], fig. 4

iTtTbiriSoTroTroSiD
21 21 19 1 0 0 0 36 
1

Chapman et al., 2008 [Publ 2744], fig. 2

iTtTbiriSoTroTroSiD
21 21 19 1 0 0 0 34.5 
1

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
21 21 19 2 4 3 2 33 
  +-------- Etherioidea     
--+                         
  |  +----- Hyrioidea       
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
21 21 19 1 0 0 0 34.5 
1

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
20 21 14 2 14 13 2 38.5 
  +-------------------- Hyrioidea       
  |                                     
  |           +-------- Iridinidae      
--+  +--------+                         
  |  |        |  +----- Etheria         
  |  |        +--+                      
  |  |           |  +-- Mulleria        
  +--+           +--+                   
     |              +-- Mycetopodidae   
     |                                  
     |  +-------------- Parreysiinae **    
     |  |                               
     +--+  +----------- Pseudomulleria  
        |  |                            
        +--+  +-------- Anodontini **      
           |  |                         
           +--+  +----- Gonideinae **      
              |  |                      
              +--+----- Unionini **        
                 |                      
                 |  +-- Margaritiferidae **
                 +--+                   
                    +-- Ambleminae **

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
20 20 17 2 11 10 2 34.5 
  +----------------- Hyrioidea       
--+                                  
  |  +-------------- Etherioidea     
  +--+                               
     |  +----------- Parreysiinae **    
     +--+                            
        |  +-------- Margaritiferidae **
        +--+                         
           |  +----- Gonideinae **      
           +--+                      
              |  +-- Unioninae **       
              +--+                   
                 +-- Ambleminae **

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
20 20 18 2 2 2 1 32.5 
  +----- Hyrioidea       
--+                      
  |  +-- Margaritiferidae **
  +--+                   
     +-- Unionidae **

Matos et al., 2024 [Publ 4384], fig. 3

iTtTbiriSoTroTroSiD
20 20 18 2 0 0 0 33.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
20 20 16 2 11 10 2 31.5 
     +-- Etherioidea     
  +--+                   
  |  +-- Hyrioidea       
--+                      
  |  +-- Margaritiferidae **
  +--+                   
     +-- Unionidae **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
20 30 18 2 11 10 2 40 
  +----------------- Margaritiferidae **
  |                                  
  |  +-------------- Amblemini **       
  |  |                               
  |  +-------------- Parreysiinae **    
  |  |                               
  |  +-------------- Eurynia **         
--+  |                               
  |  +-------------- Fusconaia **       
  |  |                               
  |  +-------------- Gonideinae **      
  |  |                               
  |  +-------------- Pleurobema **      
  |  |                               
  +--+-------------- Quadrulini **      
     |                               
     |  +----------- Ortmanniana **     
     +--+                            
     |  |  +-------- Glebula **         
     |  +--+                         
     |     |  +----- Potamilus **       
     |     +--+                      
     |        |  +-- Ligumia **         
     |        +--+                   
     |           +-- Toxolasma **       
     |                               
     |  +----------- Cyrtonaias **      
     +--+                            
        |  +-------- Unionini **        
        +--+                         
           |  +----- Anodontini **      
           +--+                      
              |  +-- Hyrioidea       
              +--+                   
                 +-- Etherioidea

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
20 20 17 2 11 10 2 34.5 
  +----------------- Hyrioidea       
--+                                  
  |  +-------------- Etherioidea     
  +--+                               
     |  +----------- Parreysiinae **    
     +--+                            
        |  +-------- Margaritiferidae **
        +--+                         
           |  +----- Ambleminae **      
           +--+                      
              |  +-- Gonideinae **      
              +--+                   
                 +-- Unioninae **

Zhou et al., 2016 [Publ 3710], fig. 1

iTtTbiriSoTroTroSiD
20 20 18 2 0 0 0 34 
  +-------- Unioninae **       
--+                         
  |  +----- Margaritiferidae **
  +--+                      
     |  +-- Ambleminae **      
     +--+                   
        +-- Gonideinae **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
20 30 28 2 11 10 2 40 
  +-------- Margaritiferidae **
--+                         
  |  +----- Unionidae **       
  +--+                      
     |  +-- Etherioidea     
     +--+                   
        +-- Hyrioidea

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
20 20 17 2 0 0 0 33 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
20 20 18 2 2 2 1 32.5 
  +----- Hyrioidea       
--+                      
  |  +-- Margaritiferidae **
  +--+                   
     +-- Unionidae **

Pieri et al., 2018 [Publ 3867], fig. 3

iTtTbiriSoTroTroSiD
20 20 23 1 0 0 0 39 
1

Smith et al., 2020 [Publ 4136], fig. 1

iTtTbiriSoTroTroSiD
20 20 13 1 0 0 0 34.5 
1

Inoue et al., 2014 [Publ 3635], fig. 3A

iTtTbiriSoTroTroSiD
20 20 17 2 0 0 0 32 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Matos et al., 2024 [Publ 4383], fig. 3

iTtTbiriSoTroTroSiD
20 20 18 2 0 0 0 33.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Froufe et al., 2019 [Publ 3936], fig. 4

iTtTbiriSoTroTroSiD
19 19 17 2 0 0 0 32 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Sano et al., 2017 [Publ 3799], fig. 1

iTtTbiriSoTroTroSiD
19 19 22 2 2 0 0 34.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iTtTbiriSoTroTroSiD
19 19 17 2 0 0 0 31.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Wu et al., 2023 [Publ 4339], fig. 5

iTtTbiriSoTroTroSiD
19 19 17 2 0 0 0 31.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Froufe et al., 2019 [Publ 3936], fig. 3-M

iTtTbiriSoTroTroSiD
19 19 17 2 0 0 0 32 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Riccardi et al., 2020 [Publ 4089], fig. 2

iTtTbiriSoTroTroSiD
19 19 17 2 0 0 0 31 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Rosenberg et al., 1994 [Publ 2719], fig. 3

iTtTbiriSoTroTroSiD
19 19 5 2 0 0 0 29.5 
  +-- Cumberlandia ** 
  |                
--+-- Margaritifera **
  |                
  +-- Unionidae **

Smith et al., 2018 [Publ 3790], fig. 1

iTtTbiriSoTroTroSiD
18 18 17 1 0 0 0 32.5 
1

Araujo et al., 2018 [Publ 3779], fig. 1

iTtTbiriSoTroTroSiD
18 18 14 2 0 0 0 30.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Chong et al., 2008 [Publ 2668], fig. 3

iTtTbiriSoTroTroSiD
18 18 10 1 0 0 0 31 
1

Porto-Hannes et al., 2019 [Publ 3969], fig. 3

iTtTbiriSoTroTroSiD
18 18 13 1 0 0 0 32.5 
1

Bolotov et al., 2019 [Publ 3974], fig. 1

iTtTbiriSoTroTroSiD
18 18 16 1 0 0 0 33 
1

Zhou et al., 2007 [Publ 2658], fig. 2

iTtTbiriSoTroTroSiD
18 18 16 1 0 0 0 33 
1

Kim et al., 2021 [Publ 4197], fig. 1

iTtTbiriSoTroTroSiD
18 18 16 1 0 0 0 32.5 
1

Sano et al., 2022 [Publ 4311], fig. 3

iTtTbiriSoTroTroSiD
18 18 16 1 0 0 0 33 
1

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
18 18 16 2 11 9 2 30.5 
  +-------- Hyrioidea       
--+                         
  |  +----- Etherioidea     
  +--+                      
     |  +-- Unionidae **       
     +--+                   
        +-- Margaritiferidae **

Wu et al., 2022 [Publ 4238], fig. 2e

iTtTbiriSoTroTroSiD
18 18 16 1 0 0 0 31 
1

Porto-Hannes et al., 2019 [Publ 3969], fig. 2

iTtTbiriSoTroTroSiD
18 18 16 1 0 0 0 32.5 
1

Wu et al., 2025 [Publ 4442], fig. 6

iTtTbiriSoTroTroSiD
17 17 15 1 0 0 0 30.5 
1

Grobler et al., 2011 [Publ 3359], fig. 2

iTtTbiriSoTroTroSiD
17 17 13 1 0 0 0 31.5 
1

Plazzi et al., 2011 [Publ 3676], fig. 5

iTtTbiriSoTroTroSiD
17 18 11 1 1 1 1 32.5 
  +----------- Cristariina **    
  |                           
  |  +-------- Lanceolariini **  
  |  |                        
--+  +-------- Anodonta **       
  |  |                        
  |  +-------- Unionini **       
  |  |                        
  +--+     +-- Alasmidontina **  
     +-----+                  
     |     +-- Pseudanodonta **  
     |                        
     |  +----- Ambleminae **     
     |  |                     
     +--+  +-- Hyrioidea      
        |  |                  
        +--+-- Lamprotulini **   
           |                  
           +-- Chamberlainiini **

Smith, 2021 [Publ 4331], fig. 1

iTtTbiriSoTroTroSiD
17 17 15 2 3 2 1 30 
  +----- Etherioidea     
--+                      
  |  +-- Margaritiferidae **
  +--+                   
     +-- Unionidae **

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iTtTbiriSoTroTroSiD
17 17 15 2 11 9 2 28.5 
  +-------- Hyrioidea       
--+                         
  |  +----- Etherioidea     
  +--+                      
     |  +-- Unionidae **       
     +--+                   
        +-- Margaritiferidae **

Inoue et al., 2013 [Publ 3528], fig. 3

iTtTbiriSoTroTroSiD
17 17 13 1 0 0 0 31 
1

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iTtTbiriSoTroTroSiD
17 17 15 1 0 0 0 31.5 
1

Prié & Puillandre, 2014 [Publ 3529], fig. 2

iTtTbiriSoTroTroSiD
17 17 15 2 0 0 0 27.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iTtTbiriSoTroTroSiD
17 17 15 2 11 9 2 28.5 
  +-------- Hyrioidea       
--+                         
  |  +----- Etherioidea     
  +--+                      
     |  +-- Unionidae **       
     +--+                   
        +-- Margaritiferidae **

Chen et al., 2025 [Publ 4455], fig. 1

iTtTbiriSoTroTroSiD
17 17 15 2 9 7 2 30 
  +-------- Hyrioidea       
--+                         
  |  +----- Etherioidea     
  +--+                      
     |  +-- Unionidae **       
     +--+                   
        +-- Margaritiferidae **

Johnson et al., 2018 [Publ 3844], fig. 2

iTtTbiriSoTroTroSiD
16 16 14 1 0 0 0 30.5 
1

Wen et al., 2017 [Publ 3795], fig. 3

iTtTbiriSoTroTroSiD
16 16 14 2 0 0 0 28.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
16 16 14 2 1 1 1 29 
  +----- Hyrioidea       
--+                      
  |  +-- Margaritiferidae **
  +--+                   
     +-- Unionidae **

Zanatta et al., 2007 [Publ 2698], fig. 1A

iTtTbiriSoTroTroSiD
16 16 10 1 0 0 0 30.5 
1

Szumowski et al., 2012 [Publ 3569], fig. 4

iTtTbiriSoTroTroSiD
16 16 12 1 0 0 0 30.5 
1

Inoue et al., 2020 [Publ 3941], fig. 5

iTtTbiriSoTroTroSiD
16 16 14 1 0 0 0 30.5 
1

Lopes-Lima et al., 2017 [Publ 3734], fig. 1

iTtTbiriSoTroTroSiD
16 16 12 2 1 0 0 27 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Stewart et al., 2021 [Publ 4451], fig. 1

iTtTbiriSoTroTroSiD
16 16 15 2 0 0 0 28 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Lyubas et al., 2019 [Publ 3963], fig. 10

iTtTbiriSoTroTroSiD
16 16 12 1 0 0 0 30.5 
1

Huang et al., 2013 [Publ 3629], fig. 3

iTtTbiriSoTroTroSiD
16 16 14 2 0 0 0 28 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Zhou et al., 2017 [Publ 3788], fig. 2

iTtTbiriSoTroTroSiD
16 16 14 1 0 0 0 30.5 
1

Jeratthitikul et al., 2025 [Publ 4476], fig. 2

iTtTbiriSoTroTroSiD
16 16 14 1 0 0 0 31 
1

Walker et al., 2006 [Publ 2666], fig. 6

iTtTbiriSoTroTroSiD
16 16 13 2 20 19 2 30.5 
  +-------------- Hyrioidea       
--+                               
  |  +----------- Etherioidea     
  +--+                            
     |  +-------- Unioninae **       
     +--+                         
        |  +----- Margaritiferidae **
        +--+                      
           |  +-- Gonideinae **      
           +--+                   
              +-- Ambleminae **

Zanatta et al., 2007 [Publ 2698], fig. 1B

iTtTbiriSoTroTroSiD
16 16 13 1 0 0 0 30.5 
1

Hoeh, 1990 [Publ 2717], fig. 4

iTtTbiriSoTroTroSiD
15 15 13 1 0 0 0 29.5 
1

Hoeh, 1990 [Publ 2717], fig. 3

iTtTbiriSoTroTroSiD
15 15 2 1 0 0 0 29.5 
1

Kongim et al., 2023 [Publ 4290], fig. 1

iTtTbiriSoTroTroSiD
15 15 13 1 0 0 0 30 
1

Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F

iTtTbiriSoTroTroSiD
15 15 13 1 0 0 0 28 
1

Bolotov et al., 2022 [Publ 4250], fig. S5

iTtTbiriSoTroTroSiD
15 15 12 1 0 0 0 30 
1

Hou et al., 2024 [Publ 4400], fig. 1

iTtTbiriSoTroTroSiD
15 15 13 1 0 0 0 30 
1

Hoeh, 1990 [Publ 2717], fig. 2

iTtTbiriSoTroTroSiD
15 15 13 1 0 0 0 29.5 
1

Amaro et al., 2017 [Publ 3791], fig. 2

iTtTbiriSoTroTroSiD
15 15 15 2 1 0 0 28.5 
  +-------- Margaritifera_margaritifera__2 **
--+                                       
  |  +----- Unionidae **                     
  +--+                                    
     |  +-- Margaritifera_margaritifera__1 **
     +--+                                 
        +-- Margaritifera_falcata **

Xue et al., 2016 [Publ 3659], fig. 1

iTtTbiriSoTroTroSiD
15 15 13 1 0 0 0 29 
1

Porto-Hannes et al., 2021 [Publ 4227], fig. 3

iTtTbiriSoTroTroSiD
15 15 12 1 0 0 0 28.5 
1

Liu et al., 2017 [Publ 3939], fig. 2

iTtTbiriSoTroTroSiD
15 15 12 1 0 0 0 28.5 
1

Bolotov et al., 2022 [Publ 4250], fig. S2

iTtTbiriSoTroTroSiD
15 15 13 1 0 0 0 29.5 
1

Dai et al., 2024 [Publ 4386], fig. 3

iTtTbiriSoTroTroSiD
14 14 12 1 0 0 0 26.5 
1

Reis et al., 2013 [Publ 3580], fig. 5

iTtTbiriSoTroTroSiD
14 14 12 2 1 0 0 29 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Bolotov et al., 2016 [Publ 3743], fig. 4

iTtTbiriSoTroTroSiD
14 14 12 2 16 13 2 26.5 
  +-------- Hyrioidea       
--+                         
  |  +----- Etherioidea     
  +--+                      
     |  +-- Unionidae **       
     +--+                   
        +-- Margaritiferidae **

Bolotov et al., 2016 [Publ 3743], fig. 2

iTtTbiriSoTroTroSiD
14 14 12 2 16 13 2 26.5 
  +-------- Hyrioidea       
--+                         
  |  +----- Etherioidea     
  +--+                      
     |  +-- Unionidae **       
     +--+                   
        +-- Margaritiferidae **

Lydeard et al., 2000 [Publ 2654], fig. 2b

iTtTbiriSoTroTroSiD
14 14 12 2 0 0 0 29.5 
  +-- Unioninae **       
  |                   
--+-- Margaritiferidae **
  |                   
  +-- Ambleminae **

Feng et al., 2016 [Publ 3800], fig. 2

iTtTbiriSoTroTroSiD
14 14 12 2 0 0 0 26.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Szumowski et al., 2012 [Publ 3569], fig. 5

iTtTbiriSoTroTroSiD
14 14 12 1 0 0 0 29 
1

Zhang et al., 2004 [Publ 2735], fig. 1

iTtTbiriSoTroTroSiD
14 14 12 1 0 0 0 28 
1

Lopes-Lima et al., 2024 [Publ 4376], fig. 2

iTtTbiriSoTroTroSiD
14 14 12 1 0 0 0 29.5 
1

Froufe et al., 2016 [Publ 3738], fig. 3

iTtTbiriSoTroTroSiD
14 14 12 1 0 0 0 27.5 
1

Wu et al., 2020 [Publ 4147], fig. 4

iTtTbiriSoTroTroSiD
14 14 12 2 0 0 0 25.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
14 14 12 2 0 0 0 26 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Lydeard et al., 2000 [Publ 2654], fig. 2d

iTtTbiriSoTroTroSiD
14 14 12 2 0 0 0 29.5 
  +-- Unioninae **       
  |                   
--+-- Margaritiferidae **
  |                   
  +-- Ambleminae **

Douda et al., 2024 [Publ 4364], fig. 1

iTtTbiriSoTroTroSiD
14 14 13 2 0 0 0 27.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Lydeard et al., 2000 [Publ 2654], fig. 2c

iTtTbiriSoTroTroSiD
14 14 9 2 0 0 0 28.5 
  +----- Unioninae **       
  |                      
  +----- Margaritiferidae **
  |                      
--+----- Quadrulini **      
  |                      
  +----- Lampsilini **      
  |                      
  |  +-- Amblemini **       
  +--+                   
     +-- Pleurobemini **

Lopes-Lima et al., 2020 [Publ 4080], fig. 10

iTtTbiriSoTroTroSiD
14 14 10 2 0 0 0 27.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Turner et al., 2000 [Publ 2722], fig. 2

iTtTbiriSoTroTroSiD
14 14 12 2 0 0 0 28 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Lydeard et al., 2000 [Publ 2654], fig. 2a

iTtTbiriSoTroTroSiD
14 14 11 2 0 0 0 28.5 
  +----- Unioninae **       
  |                      
  +----- Margaritiferidae **
  |                      
--+----- Lampsilini **      
  |                      
  +----- Amblemini **       
  |                      
  |  +-- Quadrulini **      
  +--+                   
     +-- Pleurobemini **

Manendo et al., 2008 [Publ 2669], fig. 1

iTtTbiriSoTroTroSiD
13 13 11 1 0 0 0 28.5 
1

Xiang et al., 2025 [Publ 4433], fig. 7

iTtTbiriSoTroTroSiD
13 13 11 1 0 0 0 28 
1

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iTtTbiriSoTroTroSiD
13 13 11 1 0 0 0 28 
1

Bogan et al., 2023 [Publ 4295], fig. 6

iTtTbiriSoTroTroSiD
13 13 10 2 0 0 0 28 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Walker et al., 2006 [Publ 2666], fig. 6-M

iTtTbiriSoTroTroSiD
13 13 11 2 1 1 1 25.5 
  +----- Hyrioidea       
--+                      
  |  +-- Margaritiferidae **
  +--+                   
     +-- Unionidae **

Walker et al., 2006 [Publ 2666], fig. 4

iTtTbiriSoTroTroSiD
13 13 11 2 2 1 1 27 
  +----- Hyrioidea       
--+                      
  |  +-- Margaritiferidae **
  +--+                   
     +-- Unionidae **

Walker et al., 2006 [Publ 2666], fig. 2

iTtTbiriSoTroTroSiD
13 13 11 2 2 1 1 29 
  +----------------- Hyrioidea       
  |                                  
--+  +-------------- Unioninae **       
  |  |                               
  +--+  +----------- Lamprotulini **    
     |  |                            
     +--+        +-- Gonideini **       
        |  +-----+                   
        |  |     +-- Pseudodontini **   
        +--+                         
           |  +----- Lampsilini **      
           +--+                      
              |  +-- Pleurobemini **    
              +--+                   
                 +-- Margaritiferidae **

Hauck et al., 2023 [Publ 4333], fig. 9

iTtTbiriSoTroTroSiD
13 13 12 1 0 0 0 26.5 
1

Walker et al., 2006 [Publ 2666], fig. 3

iTtTbiriSoTroTroSiD
13 13 11 2 2 1 1 27 
  +----- Hyrioidea       
--+                      
  |  +-- Margaritiferidae **
  +--+                   
     +-- Unionidae **

Wu et al., 2024 [Publ 4379], fig. 2

iTtTbiriSoTroTroSiD
13 13 10 1 0 0 0 29 
1

Sayenko et al., 2021 [Publ 4189], fig. 6

iTtTbiriSoTroTroSiD
13 13 11 1 0 0 0 27.5 
1

Reis et al., 2013 [Publ 3580], fig. 4

iTtTbiriSoTroTroSiD
13 13 11 2 1 0 0 27 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Vikhrev et al., 2018 [Publ 3812], fig. 3

iTtTbiriSoTroTroSiD
13 13 11 2 0 0 0 25.5 
  +-- Unionidae **       
--+                   
  +-- Margaritiferidae **

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iTtTbiriSoTroTroSiD
13 13 11 1 0 0 0 28 
1

Graf, 2002 [Publ 2651], fig. 2

iTtTbiriSoTroTroSiD
13 13 11 2 5 4 1 27.5 
  +----- Hyrioidea       
--+                      
  |  +-- Margaritiferidae **
  +--+                   
     +-- Unionidae **

Ouyang et al., 2011 [Publ 3408], fig. 1

iTtTbiriSoTroTroSiD
13 13 10 1 0 0 0 28 
1

Jeratthitikul & Sutcharit, 2023 [Publ 4289], fig. 1

iTtTbiriSoTroTroSiD
13 13 11 1 0 0 0 29 
1

Freeman et al., 2025 [Publ 4443], fig. 4

iTtTbiriSoTroTroSiD
12 12 8 1 0 0 0 27 
1

Freeman et al., 2025 [Publ 4443], fig. 3

iTtTbiriSoTroTroSiD
12 12 10 1 0 0 0 27 
1

Pfarrer et al., 2022 [Publ 4305], fig. 2-5

iTtTbiriSoTroTroSiD
12 12 10 1 0 0 0 26.5 
1

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

iTtTbiriSoTroTroSiD
12 12 12 1 0 0 0 39 
1

Choi et al., 2020 [Publ 4169], fig. 2a

iTtTbiriSoTroTroSiD
12 12 11 1 0 0 0 27.5 
1

Araujo et al., 2016 [Publ 3746], fig. 1_2b

iTtTbiriSoTroTroSiD
12 12 10 2 1 0 0 26.5 
  +-- Unionidae **       
--+                   
  +-- Margaritiferidae **

Sano et al., 2022 [Publ 4311], fig. 2

iTtTbiriSoTroTroSiD
12 12 12 1 0 0 0 27 
1

Smith et al., 2020 [Publ 4136], fig. 3

iTtTbiriSoTroTroSiD
12 12 10 1 0 0 0 28 
1

Froufe et al., 2017 [Publ 3802], fig. 6

iTtTbiriSoTroTroSiD
12 12 9 1 0 0 0 28 
1

Zieritz et al., 2024 [Publ 4381], fig. 7A

iTtTbiriSoTroTroSiD
12 12 10 1 0 0 0 28 
1

Wu et al., 2022 [Publ 4238], fig. 2a_S1

iTtTbiriSoTroTroSiD
12 12 10 1 0 0 0 28 
1

Froufe et al., 2015 [Publ 3664], fig. 1

iTtTbiriSoTroTroSiD
12 12 10 1 0 0 0 26.5 
1

Lopes-Lima et al., 2020 [Publ 4080], fig. 11

iTtTbiriSoTroTroSiD
12 12 10 2 0 0 0 26.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Liu et al., 2024 [Publ 4380], fig. 4

iTtTbiriSoTroTroSiD
12 12 10 1 0 0 0 28 
1

Curole & Kocher, 2005 [Publ 2682], fig. 1-M

iTtTbiriSoTroTroSiD
12 12 10 2 0 0 0 24.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Konopleva et al., 2020 [Publ 4072], fig. 2

iTtTbiriSoTroTroSiD
12 12 9 1 0 0 0 28 
1

Morrison et al., 2021 [Publ 4362], fig. 5

iTtTbiriSoTroTroSiD
11 11 9 1 0 0 0 27 
1

Pfeiffer et al., 2021 [Publ 4126], fig. 12

iTtTbiriSoTroTroSiD
11 11 8 1 0 0 0 27 
1

Amaro et al., 2017 [Publ 3791], fig. 3A

iTtTbiriSoTroTroSiD
11 11 9 2 1 0 0 24 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Araujo et al., 2009 [Publ 2589], fig. 7B

iTtTbiriSoTroTroSiD
11 11 10 2 0 0 0 26.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Khalloufi et al., 2011 [Publ 2748], fig. 2

iTtTbiriSoTroTroSiD
11 11 8 1 0 0 0 24.5 
1

Roe, 2013 [Publ 3571], fig. 4

iTtTbiriSoTroTroSiD
11 11 9 1 0 0 0 26.5 
1

Pfeiffer et al., 2021 [Publ 4126], fig. 27

iTtTbiriSoTroTroSiD
11 11 7 1 0 0 0 27 
1

Araujo et al., 2016 [Publ 3746], fig. 2a

iTtTbiriSoTroTroSiD
11 11 9 1 0 0 0 26.5 
1

Lopes-Lima et al., 2020 [Publ 4080], fig. 6

iTtTbiriSoTroTroSiD
11 11 9 2 0 0 0 25.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Araujo et al., 2009 [Publ 2589], fig. 7A

iTtTbiriSoTroTroSiD
11 11 7 2 0 0 0 25.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Klishko et al., 2019 [Publ 3886], fig. 9

iTtTbiriSoTroTroSiD
11 11 8 2 0 0 0 24 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Amaro et al., 2017 [Publ 3791], fig. 3B

iTtTbiriSoTroTroSiD
11 11 9 2 1 0 0 24 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Hou et al., 2025 [Publ 4401], fig. 3

iTtTbiriSoTroTroSiD
11 11 9 1 0 0 0 27 
1

Graf et al., 2014 [Publ 3591], fig. 3

iTtTbiriSoTroTroSiD
11 11 13 2 0 0 0 30 
  +-- Margaritiferidae **
  |                   
--+-- Unioninae **       
  |                   
  +-- Parreysiinae **

Porto-Hannes et al., 2021 [Publ 4227], fig. 4

iTtTbiriSoTroTroSiD
11 11 6 1 0 0 0 26.5 
1

Schilling et al., 2025 [Publ 4737], fig. 3

iTtTbiriSoTroTroSiD
11 11 9 1 0 0 0 25.5 
1

Freeman et al., 2025 [Publ 4443], fig. 5

iTtTbiriSoTroTroSiD
11 11 9 1 0 0 0 26 
1

Curole & Kocher, 2005 [Publ 2682], fig. 1

iTtTbiriSoTroTroSiD
11 11 9 2 0 0 0 25.5 
  +----- Ambleminae **      
--+                      
  |  +-- Unioninae **       
  +--+                   
     +-- Margaritiferidae **

Hou et al., 2025 [Publ 4432], fig. 3

iTtTbiriSoTroTroSiD
11 11 9 1 0 0 0 27 
1

Choi et al., 2020 [Publ 4169], fig. 5

iTtTbiriSoTroTroSiD
10 10 8 1 0 0 0 25 
1

Araujo et al., 2009 [Publ 2589], fig. 7C

iTtTbiriSoTroTroSiD
10 10 8 2 0 0 0 24.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Chase et al., 2018 [Publ 3861], fig. 1

iTtTbiriSoTroTroSiD
10 10 8 2 1 1 1 23.5 
  +----- Hyrioidea       
--+                      
  |  +-- Margaritiferidae **
  +--+                   
     +-- Unionidae **

Morrison et al., 2021 [Publ 4362], fig. 4

iTtTbiriSoTroTroSiD
10 10 8 1 0 0 0 26 
1

Stodola et al., 2021 [Publ 4266], fig. 4

iTtTbiriSoTroTroSiD
10 10 7 1 0 0 0 26.5 
1

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

iTtTbiriSoTroTroSiD
10 10 8 2 0 0 0 24 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Chase et al., 2018 [Publ 3861], fig. 1-M

iTtTbiriSoTroTroSiD
10 11 9 2 1 1 1 25.5 
  +----- Unionidae **       
--+                      
  |  +-- Margaritiferidae **
  +--+                   
     +-- Hyrioidea

Bespalaya et al., 2018 [Publ 3811], fig. 3

iTtTbiriSoTroTroSiD
10 10 9 2 0 0 0 25.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Kano et al., 2019 [Publ 4194], fig. 6

iTtTbiriSoTroTroSiD
10 10 8 2 0 0 0 23.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iTtTbiriSoTroTroSiD
10 10 8 1 0 0 0 24 
1

Roe & Lydeard, 1998 [Publ 2693], fig. 3

iTtTbiriSoTroTroSiD
10 10 5 1 0 0 0 24.5 
1

Hewitt et al., 2019 [Publ 3962], fig. 2

iTtTbiriSoTroTroSiD
10 10 11 2 0 0 0 23.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
10 10 8 1 6 3 1 26.5 
  +-- Hyrioidea
--+            
  +-- Unionidae **

Froufe et al., 2017 [Publ 3802], fig. 7

iTtTbiriSoTroTroSiD
10 10 8 2 0 0 0 23 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Roe et al., 2002 [Publ 2689], fig. 3a

iTtTbiriSoTroTroSiD
10 10 8 1 0 0 0 24.5 
1

Huff et al., 2004 [Publ 2697], fig. 4

iTtTbiriSoTroTroSiD
10 10 7 2 1 0 0 23.5 
  +-- Unionidae **       
--+                   
  +-- Margaritiferidae **

Schilling et al., 2025 [Publ 4737], fig. 4

iTtTbiriSoTroTroSiD
10 10 9 1 0 0 0 23.5 
1

Huff et al., 2004 [Publ 2697], fig. 2

iTtTbiriSoTroTroSiD
10 10 6 2 2 0 0 23.5 
  +-- Unionidae **       
--+                   
  +-- Margaritiferidae **

Vikhrev et al., 2017 [Publ 3810], fig. 4

iTtTbiriSoTroTroSiD
10 10 8 2 0 0 0 24.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Keogh & Simons, 2019 [Publ 3935], fig. S1

iTtTbiriSoTroTroSiD
10 10 8 1 0 0 0 24.5 
1

Amaro et al., 2017 [Publ 3791], fig. 2-M

iTtTbiriSoTroTroSiD
10 10 7 1 0 0 0 25 
1

Lopes-Lima et al., 2020 [Publ 4080], fig. 13

iTtTbiriSoTroTroSiD
10 10 8 2 0 0 0 24.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Lopes-Lima et al., 2021 [Publ 4217], fig. 2

iTtTbiriSoTroTroSiD
10 10 8 1 0 0 0 25.5 
1

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

iTtTbiriSoTroTroSiD
10 10 8 2 0 0 0 23 
  +-- Unionidae **       
--+                   
  +-- Margaritiferidae **

Huff et al., 2004 [Publ 2697], fig. 5-6

iTtTbiriSoTroTroSiD
10 10 7 2 2 0 0 22.5 
  +-- Unionidae **       
--+                   
  +-- Margaritiferidae **

Takeuchi et al., 2015 [Publ 3661], fig. 2

iTtTbiriSoTroTroSiD
9 9 7 2 1 0 0 23 
  +-- Unionidae **       
--+                   
  +-- Margaritiferidae **

Nagel et al., 1998 [Publ 1946], fig. 1

iTtTbiriSoTroTroSiD
9 9 7 1 0 0 0 23.5 
1

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
9 15 7 1 6 3 1 26.5 
  +-------- Parreysiinae **
  |                     
  +-------- Gonideinae **  
--+                     
  |     +-- Hyrioidea   
  |  +--+               
  |  |  +-- Trigonioidea
  +--+                  
     |  +-- Pleurobemini **
     |  |               
     +--+-- Lampsilini **  
        |               
        +-- Unioninae **

King et al., 1999 [Publ 2699], fig. 2

iTtTbiriSoTroTroSiD
9 9 7 1 0 0 0 24.5 
1

Chase et al., 2018 [Publ 3847], fig. 5

iTtTbiriSoTroTroSiD
9 10 8 2 1 1 1 25 
  +----- Unionidae **       
--+                      
  |  +-- Margaritiferidae **
  +--+                   
     +-- Hyrioidea

Kandl et al., 2001 [Publ 3384], fig. 5

iTtTbiriSoTroTroSiD
9 9 7 1 0 0 0 23.5 
1

Inoue et al., 2013 [Publ 3579], fig. 3

iTtTbiriSoTroTroSiD
9 9 7 2 0 0 0 23 
  +-- Unionidae **       
--+                   
  +-- Margaritiferidae **

Pfeiffer et al., 2021 [Publ 4126], fig. 25

iTtTbiriSoTroTroSiD
9 9 7 1 0 0 0 25 
1

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
9 9 0 1 6 3 1 22.5 
  +-- Hyrioidea
--+            
  +-- Unionidae **

Takeuchi et al., 2015 [Publ 3661], fig. 3

iTtTbiriSoTroTroSiD
9 9 7 2 1 0 0 23 
  +-- Unionidae **       
--+                   
  +-- Margaritiferidae **

Muanta et al., 2019 [Publ 3921], fig. 2

iTtTbiriSoTroTroSiD
9 9 8 1 0 0 0 26 
1

Hewitt et al., 2018 [Publ 3758], fig. 5

iTtTbiriSoTroTroSiD
9 9 6 2 0 0 0 22 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Fonseca et al., 2016 [Publ 3775], fig. 1-M

iTtTbiriSoTroTroSiD
9 9 7 1 0 0 0 24 
1

Lopes-Lima et al., 2020 [Publ 4080], fig. 5

iTtTbiriSoTroTroSiD
9 9 7 2 0 0 0 23.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Bolotov et al., 2015 [Publ 3646], fig. 2A

iTtTbiriSoTroTroSiD
9 9 6 2 0 0 0 23 
  +-- Unionidae **       
--+                   
  +-- Margaritiferidae **

Bolotov et al., 2015 [Publ 3646], fig. 2B

iTtTbiriSoTroTroSiD
9 9 7 2 0 0 0 23 
  +-- Unionidae **       
--+                   
  +-- Margaritiferidae **

Soroka, 2010 [Publ 3404], fig. 3a

iTtTbiriSoTroTroSiD
9 9 7 1 0 0 0 22.5 
1

Wu et al., 2024 [Publ 4388], fig. 4

iTtTbiriSoTroTroSiD
9 9 7 1 0 0 0 25 
1

Soroka, 2010 [Publ 3404], fig. 3b

iTtTbiriSoTroTroSiD
9 9 5 1 0 0 0 22.5 
1

Chen et al., 2025 [Publ 4448], fig. 2

iTtTbiriSoTroTroSiD
9 9 7 1 0 0 0 25 
1

Bolotov et al., 2022 [Publ 4250], fig. S4

iTtTbiriSoTroTroSiD
9 9 6 1 0 0 0 25 
1

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

iTtTbiriSoTroTroSiD
9 9 14 1 0 0 0 25 
1

Marrone et al., 2019 [Publ 3907], fig. 1

iTtTbiriSoTroTroSiD
8 8 4 1 0 0 0 23.5 
1

Araujo et al., 2009 [Publ 2395], fig. 3

iTtTbiriSoTroTroSiD
8 8 6 2 0 0 0 21.5 
  +-- Unionidae **       
--+                   
  +-- Margaritiferidae **

Lopes-Lima et al., 2020 [Publ 4080], fig. 12

iTtTbiriSoTroTroSiD
8 8 6 2 0 0 0 22.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Zhang et al., 2024 [Publ 4397], fig. 2

iTtTbiriSoTroTroSiD
8 8 6 1 0 0 0 24 
1

Tomilova et al., 2020 [Publ 4097], fig. 3

iTtTbiriSoTroTroSiD
8 8 5 1 0 0 0 22.5 
1

Bolotov et al., 2016 [Publ 3732], fig. 2

iTtTbiriSoTroTroSiD
8 8 7 2 0 0 0 23.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Hoeh et al., 2002 [Publ 2673], fig. 1-F

iTtTbiriSoTroTroSiD
8 8 6 1 1 1 1 23 
  +-- Hyrioidea
--+            
  +-- Unionidae **

Lopes-Lima et al., 2020 [Publ 4080], fig. 16

iTtTbiriSoTroTroSiD
8 8 7 2 0 0 0 22.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Bauer et al., 2021 [Publ 4365], fig. 2

iTtTbiriSoTroTroSiD
8 8 5 1 0 0 0 24 
1

Pfeiffer et al., 2021 [Publ 4126], fig. 18

iTtTbiriSoTroTroSiD
8 8 7 1 0 0 0 25 
1

Zieritz et al., 2024 [Publ 4381], fig. 8A

iTtTbiriSoTroTroSiD
8 8 6 1 0 0 0 24 
1

Inoue et al., 2014 [Publ 3635], fig. 3B

iTtTbiriSoTroTroSiD
8 8 3 1 0 0 0 21.5 
1

Kondakov et al., 2018 [Publ 3914], fig. 5

iTtTbiriSoTroTroSiD
8 8 6 1 0 0 0 24 
1

Soroka, 2010 [Publ 3404], fig. 5b

iTtTbiriSoTroTroSiD
8 8 5 1 0 0 0 22.5 
1

Hoeh et al., 2002 [Publ 2673], fig. 1-M

iTtTbiriSoTroTroSiD
8 8 6 1 1 1 1 23 
  +-- Hyrioidea
--+            
  +-- Unionidae **

Zieritz et al., 2012 [Publ 3673], fig. 4

iTtTbiriSoTroTroSiD
8 8 4 2 0 0 0 23 
  +----- Anodontina **      
  |                      
--+  +-- Cristariina **     
  |  |                   
  +--+-- Margaritiferidae **
     |                   
     +-- Unionini **

Soroka, 2010 [Publ 3404], fig. 5a

iTtTbiriSoTroTroSiD
8 8 6 1 0 0 0 22.5 
1

Froufe et al., 2015 [Publ 3664], fig. 1-M

iTtTbiriSoTroTroSiD
8 8 6 1 0 0 0 22.5 
1

Sheng et al., 2014 [Publ 3704], fig. 2

iTtTbiriSoTroTroSiD
8 8 6 1 0 0 0 22 
1

Lopes-Lima et al., 2020 [Publ 4080], fig. 7

iTtTbiriSoTroTroSiD
8 8 6 2 0 0 0 22.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Klishko et al., 2018 [Publ 3945], fig. 8

iTtTbiriSoTroTroSiD
8 8 5 1 0 0 0 23.5 
1

Amaro et al., 2017 [Publ 3791], fig. 3A-M

iTtTbiriSoTroTroSiD
7 7 5 2 0 0 0 20 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Hoeh et al., 1995 [Publ 3391], fig. 2

iTtTbiriSoTroTroSiD
7 7 4 1 0 0 0 22.5 
1

Smith et al., 2023 [Publ 4291], fig. 2

iTtTbiriSoTroTroSiD
7 7 5 1 0 0 0 23 
1

Goncalves et al., 2022 [Publ 4253], fig. 2

iTtTbiriSoTroTroSiD
7 7 6 1 0 0 0 23 
1

Amaro et al., 2017 [Publ 3791], fig. 3B-M

iTtTbiriSoTroTroSiD
7 7 5 2 0 0 0 20 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Jones & Neves, 2010 [Publ 2646], fig. 16

iTtTbiriSoTroTroSiD
7 7 5 1 0 0 0 23 
1

Soroka, 2010 [Publ 3404], fig. 4

iTtTbiriSoTroTroSiD
7 7 5 1 0 0 0 21.5 
1

Källersjö et al., 2005 [Publ 2684], fig. 1A

iTtTbiriSoTroTroSiD
7 7 2 2 0 0 0 21 
  +-- Margaritiferidae **
  |                   
--+-- Anodontini **      
  |                   
  +-- Unionini **

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

iTtTbiriSoTroTroSiD
7 7 5 1 0 0 0 22 
1

Serb & Barnhart, 2008 [Publ 2674], fig. 4

iTtTbiriSoTroTroSiD
7 7 9 1 0 0 0 22.5 
1

Lopes-Lima et al., 2020 [Publ 4080], fig. 8

iTtTbiriSoTroTroSiD
7 7 5 2 0 0 0 21.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Serb & Barnhart, 2008 [Publ 2674], fig. 3

iTtTbiriSoTroTroSiD
7 7 10 1 0 0 0 22.5 
1

Buhay et al., 2002 [Publ 2661], fig. 2A

iTtTbiriSoTroTroSiD
7 7 3 1 0 0 0 22.5 
1

Buhay et al., 2002 [Publ 2661], fig. 2B

iTtTbiriSoTroTroSiD
7 7 5 1 0 0 0 22.5 
1

Froufe et al., 2016 [Publ 3727], fig. 3_4

iTtTbiriSoTroTroSiD
7 7 3 1 0 0 0 23 
1

Pfeiffer et al., 2021 [Publ 4126], fig. 30

iTtTbiriSoTroTroSiD
7 7 5 1 0 0 0 23 
1

Guerra et al., 2017 [Publ 3772], fig. 2a-M

iTtTbiriSoTroTroSiD
7 7 4 2 1 1 1 20.5 
  +----- Hyrioidea       
--+                      
  |  +-- Margaritiferidae **
  +--+                   
     +-- Unionidae **

Bogan et al., 2023 [Publ 4295], fig. 4

iTtTbiriSoTroTroSiD
7 7 5 2 0 0 0 22 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Källersjö et al., 2005 [Publ 2684], fig. 1B-F

iTtTbiriSoTroTroSiD
7 7 4 2 0 0 0 21 
  +-- Margaritiferidae **
  |                   
--+-- Anodontini **      
  |                   
  +-- Unionini **

Pfeiffer et al., 2018 [Publ 3827], fig. 1

iTtTbiriSoTroTroSiD
7 7 3 1 0 0 0 23 
1

Guerra et al., 2017 [Publ 3772], fig. 2b-M

iTtTbiriSoTroTroSiD
7 7 4 2 1 1 1 20.5 
  +----- Hyrioidea       
--+                      
  |  +-- Margaritiferidae **
  +--+                   
     +-- Unionidae **

Hou et al., 2025 [Publ 4445], fig. 4

iTtTbiriSoTroTroSiD
7 7 5 1 0 0 0 23 
1

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

iTtTbiriSoTroTroSiD
7 7 5 1 0 0 0 22 
1

Serb, 2006 [Publ 2678], fig. 4

iTtTbiriSoTroTroSiD
7 7 8 1 0 0 0 22.5 
1

Serb, 2006 [Publ 2678], fig. 2

iTtTbiriSoTroTroSiD
7 7 7 1 0 0 0 22.5 
1

Chen et al., 2012 [Publ 3624], fig. 4

iTtTbiriSoTroTroSiD
7 7 5 1 0 0 0 21 
1

Serb, 2006 [Publ 2678], fig. 3

iTtTbiriSoTroTroSiD
7 7 8 1 0 0 0 22.5 
1

Huff et al., 2004 [Publ 2697], fig. 3

iTtTbiriSoTroTroSiD
7 7 3 2 1 0 0 21.5 
  +-- Unionidae **       
--+                   
  +-- Margaritiferidae **

Lopes-Lima et al., 2021 [Publ 4217], fig. 6

iTtTbiriSoTroTroSiD
7 7 5 1 0 0 0 22 
1

Soroka, 2010 [Publ 3404], fig. 2

iTtTbiriSoTroTroSiD
7 7 4 1 0 0 0 21.5 
1

Jones & Neves, 2010 [Publ 2646], fig. 17

iTtTbiriSoTroTroSiD
7 7 1 1 0 0 0 23 
1

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M

iTtTbiriSoTroTroSiD
6 6 4 2 0 0 0 19 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Choi et al., 2020 [Publ 4169], fig. 3a

iTtTbiriSoTroTroSiD
6 6 5 1 0 0 0 21.5 
1

Bashê, 2022 [Publ 4265], fig. 1

iTtTbiriSoTroTroSiD
6 6 5 2 0 0 0 20.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Huang et al., 2013 [Publ 3629], fig. 3-M

iTtTbiriSoTroTroSiD
6 6 4 1 0 0 0 20.5 
1

Moyer & Díaz-Ferguson, 2012 [Publ 3375], fig. 3

iTtTbiriSoTroTroSiD
6 6 5 1 0 0 0 20.5 
1

Nagel et al., 1998 [Publ 1946], fig. 9

iTtTbiriSoTroTroSiD
6 6 4 2 0 0 0 18 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Soroka, 2010 [Publ 3404], fig. 8b

iTtTbiriSoTroTroSiD
6 6 4 1 0 0 0 20.5 
1

Bogan et al., 2023 [Publ 4295], fig. 5

iTtTbiriSoTroTroSiD
6 6 4 2 0 0 0 21 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Klishko et al., 2017 [Publ 3946], fig. 8

iTtTbiriSoTroTroSiD
6 6 4 1 0 0 0 21.5 
1

Prie et al., 2012 [Publ 3487], fig. 2

iTtTbiriSoTroTroSiD
6 6 4 2 0 0 0 20.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Bogatov et al., 2018 [Publ 3859], fig. 1

iTtTbiriSoTroTroSiD
6 6 4 2 0 0 0 19 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

King et al., 1999 [Publ 2699], fig. 4

iTtTbiriSoTroTroSiD
6 6 3 1 0 0 0 22 
1

Soroka, 2010 [Publ 3404], fig. 3b-M

iTtTbiriSoTroTroSiD
6 6 4 1 0 0 0 20.5 
1

Soroka, 2010 [Publ 3404], fig. 3a-M

iTtTbiriSoTroTroSiD
6 6 4 1 0 0 0 20.5 
1

Bogatov et al., 2018 [Publ 3859], fig. 2

iTtTbiriSoTroTroSiD
6 6 4 2 0 0 0 19 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Lopes-Lima et al., 2020 [Publ 4080], fig. 9

iTtTbiriSoTroTroSiD
6 6 4 2 0 0 0 20.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Zieritz et al., 2016 [Publ 3804], fig. S2-1

iTtTbiriSoTroTroSiD
6 6 3 1 0 0 0 22 
1

Lopes-Lima et al., 2020 [Publ 4080], fig. 17

iTtTbiriSoTroTroSiD
6 6 5 2 0 0 0 19.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8

iTtTbiriSoTroTroSiD
6 6 4 1 0 0 0 22 
1

Lopes-Lima et al., 2020 [Publ 4080], fig. 14

iTtTbiriSoTroTroSiD
6 6 4 2 0 0 0 20.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Riccardi et al., 2020 [Publ 4089], fig. 1

iTtTbiriSoTroTroSiD
6 6 4 1 0 0 0 22 
1

Freeman et al., 2025 [Publ 4443], fig. 2

iTtTbiriSoTroTroSiD
6 6 5 1 0 0 0 22 
1

Wei et al., 2016 [Publ 3665], fig. 0

iTtTbiriSoTroTroSiD
6 6 2 1 0 0 0 20 
1

Kandl et al., 2001 [Publ 3384], fig. 4

iTtTbiriSoTroTroSiD
6 6 4 1 0 0 0 21 
1

Dai et al., 2024 [Publ 4358], fig. 3

iTtTbiriSoTroTroSiD
6 6 4 1 0 0 0 22 
1

Keogh & Simons, 2019 [Publ 3935], fig. 4

iTtTbiriSoTroTroSiD
6 6 6 1 0 0 0 22.5 
1

Wen et al., 2017 [Publ 3795], fig. 3-M

iTtTbiriSoTroTroSiD
6 6 4 1 0 0 0 20.5 
1

Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-M

iTtTbiriSoTroTroSiD
6 6 4 1 0 0 0 21.5 
1

Soroka, 2010 [Publ 3404], fig. 8a

iTtTbiriSoTroTroSiD
6 6 4 1 0 0 0 20.5 
1

Lopes-Lima et al., 2021 [Publ 4217], fig. 14

iTtTbiriSoTroTroSiD
5 5 4 1 0 0 0 21 
1

Buhay et al., 2002 [Publ 2661], fig. 1

iTtTbiriSoTroTroSiD
5 5 3 1 0 0 0 21.5 
1

Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5_7

iTtTbiriSoTroTroSiD
5 5 3 1 0 0 0 19.5 
1

Cyr et al., 2007 [Publ 2745], fig. 2A

iTtTbiriSoTroTroSiD
5 5 3 1 0 0 0 20 
1

Cyr et al., 2007 [Publ 2745], fig. 2B

iTtTbiriSoTroTroSiD
5 5 4 1 0 0 0 21.5 
1

Lane et al., 2016 [Publ 3735], fig. 3

iTtTbiriSoTroTroSiD
5 5 1 1 0 0 0 22 
1

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

iTtTbiriSoTroTroSiD
5 11 9 2 6 6 1 23 
  +-- Margaritiferidae **
  |                   
--+-- Unionidae **       
  |                   
  +-- Hyrioidea

Nagel et al., 1998 [Publ 1946], fig. 7

iTtTbiriSoTroTroSiD
5 5 4 1 0 0 0 21 
1

Mabuchi et al., 2021 [Publ 4233], fig. 1

iTtTbiriSoTroTroSiD
5 5 3 1 0 0 0 21 
1

Saxon et al., 2023 [Publ 4338], fig. 4

iTtTbiriSoTroTroSiD
5 5 3 1 0 0 0 21 
1

Cyr et al., 2007 [Publ 2745], fig. 2C

iTtTbiriSoTroTroSiD
5 5 4 1 0 0 0 23.5 
1

Soroka, 2005 [Publ 2726], fig. 3

iTtTbiriSoTroTroSiD
5 5 3 1 0 0 0 20.5 
1

Bolotov et al., 2022 [Publ 4250], fig. S3

iTtTbiriSoTroTroSiD
5 5 3 1 0 0 0 21 
1

Konopleva et al., 2017 [Publ 3807], fig. 5

iTtTbiriSoTroTroSiD
5 5 3 1 0 0 0 21 
1

Soroka, 2010 [Publ 3404], fig. 9a

iTtTbiriSoTroTroSiD
5 5 3 1 0 0 0 21 
1

Nagel et al., 1996 [Publ 3400], fig. 351

iTtTbiriSoTroTroSiD
5 5 4 1 0 0 0 21 
1

Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2

iTtTbiriSoTroTroSiD
5 5 3 2 0 0 0 19.5 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Krebs et al., 2013 [Publ 3515], fig. 2A

iTtTbiriSoTroTroSiD
5 5 3 1 0 0 0 20.5 
1

Zanatta & Harris, 2013 [Publ 3530], fig. 2

iTtTbiriSoTroTroSiD
5 5 3 1 0 0 0 19 
1

Soroka, 2010 [Publ 3404], fig. 2-M

iTtTbiriSoTroTroSiD
5 5 3 1 0 0 0 19.5 
1

Mulvey et al., 1997 [Publ 3390], fig. 1_3

iTtTbiriSoTroTroSiD
5 5 3 1 0 0 0 20 
1

Jones et al., 2006 [Publ 2681], fig. 2

iTtTbiriSoTroTroSiD
5 5 4 1 0 0 0 21 
1

Krebs, 2004 [Publ 2692], fig. 2

iTtTbiriSoTroTroSiD
5 5 3 1 0 0 0 20 
1

Curole & Kocher, 2002 [Publ 2712], fig. 1a-M

iTtTbiriSoTroTroSiD
4 4 2 1 0 0 0 18.5 
1

Nagel et al., 1998 [Publ 1946], fig. 8

iTtTbiriSoTroTroSiD
4 4 1 1 0 0 0 19.5 
1

Patterson et al., 2021 [Publ 4222], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 1 0 0 0 20 
1

Klishko et al., 2016 [Publ 3733], fig. 1

iTtTbiriSoTroTroSiD
4 4 2 1 0 0 0 20 
1

Soroka, 2008 [Publ 3695], fig. 1

iTtTbiriSoTroTroSiD
4 4 2 1 0 0 0 18.5 
1

Jiang et al., 2010 [Publ 3438], fig. 5

iTtTbiriSoTroTroSiD
4 4 1 1 0 0 0 18.5 
1

Mezhzherin et al., 2014 [Publ 3705], fig. 4

iTtTbiriSoTroTroSiD
4 4 2 1 0 0 0 20 
1

Lopes-Lima et al., 2021 [Publ 4217], fig. 11

iTtTbiriSoTroTroSiD
4 4 2 1 0 0 0 20 
1

Soroka, 2008 [Publ 3695], fig. 1-M

iTtTbiriSoTroTroSiD
4 4 2 1 0 0 0 18.5 
1

Mezhzherin et al., 2014 [Publ 3705], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 1 0 0 0 20 
1

Curole & Kocher, 2002 [Publ 2712], fig. 1-F

iTtTbiriSoTroTroSiD
4 4 2 2 0 0 0 18 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Soroka, 2010 [Publ 3404], fig. 9c

iTtTbiriSoTroTroSiD
4 4 3 1 0 0 0 20 
1

Combosch et al., 2017 [Publ 3806], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 2 11 9 2 17.5 
  +-------- Etherioidea     
--+                         
  |  +----- Hyrioidea       
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unionidae **

Giribet & Distel, 2003 [Publ 3670], fig. 3f5

iTtTbiriSoTroTroSiD
4 4 1 2 2 0 0 16 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Bulakhova et al., 2023 [Publ 4325], fig. 5

iTtTbiriSoTroTroSiD
4 4 2 1 0 0 0 20 
1

Sayenko et al., 2017 [Publ 3760], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 1 0 0 0 20 
1

Machordom et al., 2003 [Publ 2688], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 2 0 0 0 18.5 
  +-- Unionidae **       
--+                   
  +-- Margaritiferidae **

Gangloff et al., 2013 [Publ 3551], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 1 0 0 0 20 
1

Zhang et al., 2013 [Publ 4214], fig. 5

iTtTbiriSoTroTroSiD
4 4 2 2 0 0 0 18.5 
  +-------- Cristaria **       
--+                         
  |  +----- Sinanodonta **     
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Gonideinae **

Pfeiffer et al., 2018 [Publ 3835], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 1 0 0 0 20 
1

Curole & Kocher, 2002 [Publ 2712], fig. 1b-M

iTtTbiriSoTroTroSiD
4 4 2 1 0 0 0 18.5 
1

Klishko et al., 2014 [Publ 3666], fig. 8

iTtTbiriSoTroTroSiD
4 4 2 1 0 0 0 20 
1

Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3

iTtTbiriSoTroTroSiD
4 4 1 1 0 0 0 18.5 
1

Cyr et al., 2007 [Publ 2745], fig. 2D

iTtTbiriSoTroTroSiD
4 4 2 1 0 0 0 20.5 
1

Johnson et al., 2023 [Publ 4323], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 1 0 0 0 20 
1

Froufe et al., 2017 [Publ 3802], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 1 0 0 0 20 
1

Krebs, 2004 [Publ 2692], fig. 2-M

iTtTbiriSoTroTroSiD
4 4 2 1 0 0 0 19 
1

Marshall et al., 2014 [Publ 3593], fig. 1

iTtTbiriSoTroTroSiD
4 14 12 2 10 10 1 28 
     +-- Margaritiferidae **
  +--+                   
  |  +-- Unioninae **       
--+                      
  |  +-- Ambleminae **      
  +--+                   
     +-- Hyrioidea

Combosch et al., 2017 [Publ 3806], fig. 1

iTtTbiriSoTroTroSiD
4 10 8 2 11 9 2 25.5 
  +-------- Hyrioidea       
--+                         
  |  +----- Unionidae **       
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Etherioidea

Soroka, 2010 [Publ 3404], fig. 9b

iTtTbiriSoTroTroSiD
4 4 2 1 0 0 0 20 
1

Burzynski & Soroka, 2018 [Publ 3862], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 1 0 0 0 19.5 
1

Hoeh et al., 1998 [Publ 2695], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 2 2 1 1 18.5 
  +-------- Etherioidea     
--+                         
  |  +----- Ambleminae **      
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Unioninae **

Soroka, 2010 [Publ 3404], fig. 8b-M

iTtTbiriSoTroTroSiD
3 3 1 1 0 0 0 17.5 
1

Hoeh et al., 1998 [Publ 2695], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 2 2 1 1 18.5 
  +-------- Etherioidea     
--+                         
  |  +----- Unioninae **       
  +--+                      
     |  +-- Margaritiferidae **
     +--+                   
        +-- Ambleminae **

Hinzmann et al., 2013 [Publ 3541], fig. 5

iTtTbiriSoTroTroSiD
3 3 1 1 0 0 0 19 
1

Imsong & Murali, 2023 [Publ 4370], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 1 0 0 0 18.5 
1

Krebs et al., 2019 [Publ 4092], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 1 0 0 0 18.5 
1

Mock et al., 2013 [Publ 3598], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 1 0 0 0 18.5 
1

Välilä et al., 2017 [Publ 3793], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 2 0 0 0 17.5 
  +-- Unionidae **       
--+                   
  +-- Margaritiferidae **

Smith & Johnson, 2020 [Publ 4229], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 1 0 0 0 19 
1

Metzger et al., 2018 [Publ 3952], fig. 1C

iTtTbiriSoTroTroSiD
3 3 1 1 0 0 0 18 
1

Shirai et al., 2010 [Publ 3353], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 1 0 0 0 19 
1

Soroka, 2010 [Publ 3404], fig. 8a-M

iTtTbiriSoTroTroSiD
3 3 1 1 0 0 0 17.5 
1

Machordom et al., 2003 [Publ 2688], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 2 0 0 0 17.5 
  +-- Unionidae **       
--+                   
  +-- Margaritiferidae **

Wang et al., 2016 [Publ 3714], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 1 0 0 0 18.5 
1

Taylor et al., 2007 [Publ 2696], fig. 1-3

iTtTbiriSoTroTroSiD
3 3 1 2 1 0 0 16 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Hoeh et al., 1996 [Publ 2694], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 1 1 0 0 17.5 
  +----- Unionidae **
--+               
  |  +-- None__1  
  +--+            
     +-- None__2

Campbell, 2000 [Publ 2724], fig. 1-5

iTtTbiriSoTroTroSiD
3 3 1 2 0 0 0 16 
  +-- Margaritiferidae **
--+                   
  +-- Unionidae **

Xia et al., 2016 [Publ 3712], fig. 7B

iTtTbiriSoTroTroSiD
3 3 1 1 0 0 0 19 
1

Kilikowska et al., 2020 [Publ 4335], fig. 4a

iTtTbiriSoTroTroSiD
3 3 1 1 0 0 0 18.5 
1

Zieritz et al., 2024 [Publ 4381], fig. 6A

iTtTbiriSoTroTroSiD
3 3 1 1 0 0 0 19 
1

Peng et al., 2012 [Publ 3689], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 1 0 0 0 19 
1

Lopes-Lima et al., 2016 [Publ 3757], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 1 0 0 0 19 
1

Froufe et al., 2014 [Publ 3592], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 1 0 0 0 19 
1

Klunzinger et al., 2021 [Publ 4086], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 4 4 1 -- 
  +----- Hyrioidea       
--+                      
  |  +-- Margaritiferidae **
  +--+                   
     +-- Unionidae **

Bieler et al., 2014 [Publ 3622], fig. 28

iTtTbiriSoTroTroSiD
2 4 0 2 3 2 2 19.5 
     +-- Etherioidea     
  +--+                   
  |  +-- Margaritiferidae **
--+                      
  |  +-- Unionidae **       
  +--+                   
     +-- Hyrioidea

Lemer et al., 2019 [Publ 4093], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 4 2 1 -- 
  +----- Etherioidea     
--+                      
  |  +-- Margaritiferidae **
  +--+                   
     +-- Unionidae **

González & Giribet, 2015 [Publ 3632], fig. 6

iTtTbiriSoTroTroSiD
2 3 0 2 4 2 2 18 
  +-------- Hyrioidea       
--+                         
  |  +----- Unionidae **       
  +--+                      
     |  +-- Etherioidea     
     +--+                   
        +-- Margaritiferidae **

Plazzi & Passamonti, 2010 [Publ 3354], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 0 0 0 -- 
1

Soroka, 2010 [Publ 3404], fig. 4-M

iTtTbiriSoTroTroSiD
2 2 0 1 0 0 0 -- 
1

Bieler et al., 2014 [Publ 3622], fig. 26_27_29_30

iTtTbiriSoTroTroSiD
2 3 0 2 3 2 2 18 
  +-------- Hyrioidea       
--+                         
  |  +----- Unionidae **       
  +--+                      
     |  +-- Etherioidea     
     +--+                   
        +-- Margaritiferidae **

González & Giribet, 2015 [Publ 3632], fig. 2

iTtTbiriSoTroTroSiD
2 4 0 2 4 2 2 19.5 
     +-- Etherioidea     
  +--+                   
  |  +-- Margaritiferidae **
--+                      
  |  +-- Unionidae **       
  +--+                   
     +-- Hyrioidea

Soroka, 2010 [Publ 3404], fig. 5b-M

iTtTbiriSoTroTroSiD
2 2 0 1 0 0 0 -- 
1

Graf et al., 2015 [Publ 3619], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 18 17 2 -- 
  +-------- Hyrioidea       
--+                         
  |  +----- Etherioidea     
  +--+                      
     |  +-- Unionidae **       
     +--+                   
        +-- Margaritiferidae **

Mock et al., 2004 [Publ 2686], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 1 0 0 0 -- 
1

González & Giribet, 2015 [Publ 3632], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 4 2 2 -- 
  +-------- Etherioidea     
--+                         
  |  +----- Hyrioidea       
  +--+                      
     |  +-- Unionidae **       
     +--+                   
        +-- Margaritiferidae **

Mock et al., 2004 [Publ 2686], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 0 0 0 -- 
1

González & Giribet, 2015 [Publ 3632], fig. 3

iTtTbiriSoTroTroSiD
2 4 0 2 4 2 2 18.5 
  +----- Unionidae **       
  |                      
--+----- Hyrioidea       
  |                      
  |  +-- Etherioidea     
  +--+                   
     +-- Margaritiferidae **

Xue et al., 2016 [Publ 3659], fig. 1-M

iTtTbiriSoTroTroSiD
2 2 0 1 0 0 0 -- 
1

Soroka, 2010 [Publ 3404], fig. 5a-M

iTtTbiriSoTroTroSiD
2 2 0 1 0 0 0 -- 
1

Baker et al., 2003 [Publ 2690], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 7 7 1 -- 
  +-- Unionidae **
--+            
  +-- Hyrioidea

Baker et al., 2004 [Publ 2685], fig. 2-3

iTtTbiriSoTroTroSiD
1 1 0 1 8 8 1 -- 
  +-- Unionidae **
--+            
  +-- Hyrioidea