Bivalvia | Unionida | Hyrioidea | Hyriidae

subfamily VELESUNIONINAE Iredale, 1934: cladomics

6 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 17 (21 total)
number of subtaxa (rank: genus) = 6
analyzed in 74 cladograms in 41 publications
maximum number of terminals in an analysis = 11 (mean = 2.51, median = 1)
maximum number of subtaxa (genus) = 5 (mean = 1.91, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 30
maximum number of included terminals = 11, (mean = 4.73 median = 4)
maximum number of subtaxa (genus) = 5 (mean = 3.23, median = 2.5)
proportion of cladograms that support basic monophyly = 0.67

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 28
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 11 (mean = 4.68, median = 4)
maximum number of subtaxa (genus) = 5 (mean = 3.29, median = 2.5)
proportion of cladograms that support rigorous monophyly = 0.64

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Velesunioninae cladogram distances
PCoA Ordination

NJ Tree of Velesunioninae cladogram distances
NJ Tree

Cladograms

Graf et al., 2015 [Publ 3619], fig. 1

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
0.882 7 7 5 5 13 7 1 **
Cladomics cladogram HYRIINAE Westralunio Alathyria jacksoni Velesunio Alathyria profuga Alathyria pertexta Microdontia Lortiella

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

cst iT tT bi riS oT roT roS iD
0.682 6 20 20 5 15 14 1 24.5
Cladomics cladogram Neotrigonia Guyanhyrida HYRIINI CASTALIINI Westralunio Diplodon Hyridella drapeta Hyridella depressa Hyridella australis Cucumerunio Echyridella Velesunio ambiguus Microdontia Lortiella Alathyria Velesunio angasi

Santos-Neto et al., 2016 [Publ 3739], fig. 4

cst iT tT bi riS oT roT roS iD
0.682 6 13 11 5 15 14 1 12.5
Cladomics cladogram Guyanhyrida HYRIINI CASTALIINI Westralunio Diplodon Hyridella drapeta Hyridella depressa Echyridella Cucumerunio Hyridella australis Velesunio ambiguus Alathyria Velesunio angasi Lortiella Microdontia

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
0.882 7 14 12 5 194 7 1 7.0
Cladomics cladogram Westralunio HYRIINAE Alathyria jacksoni Alathyria profuga Velesunio Alathyria pertexta Lortiella Microdontia Anodontites IRIDININAE Etheria ASPATHARIINAE MYCETOPODINAE MONOCONDYLAEINAE Lamproscapha Gibbosula rochechouartii Gibbosula laosensis MARGARITIFERINAE PARREYSIINAE UNIONINAE LIMNOSCAPHINI AMBLEMINAE CHAMBERLAINIINI LAMPROTULINI GONIDEINI PSEUDODONTINI

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
0.882 7 14 12 5 192 7 1 9.0
Cladomics cladogram Westralunio HYRIINAE Velesunio Alathyria jacksoni Alathyria profuga Alathyria pertexta Lortiella Microdontia Anodontites IRIDININAE Etheria ASPATHARIINAE MYCETOPODINAE MONOCONDYLAEINAE Lamproscapha Gibbosula rochechouartii Gibbosula laosensis MARGARITIFERINAE PARREYSIINAE PSEUDODONTINI LIMNOSCAPHINI UNIONINAE CHAMBERLAINIINI AMBLEMINAE LAMPROTULINI GONIDEINI

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.824 7 14 9 5 194 7 1 9.0
Cladomics cladogram Echyridella Westralunio HYRIDELLINI RHIPIDODONTINI CASTALIINI HYRIINI Velesunio Alathyria jacksoni Alathyria profuga Alathyria pertexta Lortiella Microdontia Gibbosula rochechouartii Gibbosula laosensis MARGARITIFERINAE PARREYSIINAE Chamberlainia Megalonaias Uniomerus QUADRULINA Sinohyriopsis LAMPROTULINI GONIDEINI PSEUDODONTINI LIMNOSCAPHINI UNIONINAE Amblema Toxolasma PLEUROBEMINI Obliquaria Plectomerus Ellipsaria Potamilus leptodon Medionidus Potamilus alatus Ptychobranchus Truncilla Venustaconcha Cambarunio Leaunio Ortmanniana Lampsilis

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.636 10 29 34 5 623 19 1 36.0
Cladomics cladogram Westralunio Guyanhyrida HYRIINI CASTALIINI Diplodon Hyridella depressa Hyridella drapeta Cucumerunio Hyridella australis Echyridella Velesunio 1 Alathyria jacksoni 1 Velesunio ambiguus Alathyria profuga Velesunio 3 Microdontia Lortiella Velesunio angasi Alathyria pertexta 2 Velesunio 2 Alathyria jacksoni 2 Alathyria pertexta 1 UNIONINAE Lamellidens Balwantia Trapezidens LEOPARREYSIINI Pseudomulleria INDOCHINELLINI PARREYSIINI COELATURINI LIMNOSCAPHINI Pseudunio GIBBOSULINAE Cumberlandia Margaritifera AMBLEMINAE INDOPSEUDODONTINA Lannanaia Pilsbryoconcha Songkhlanaia Nyeinchanconcha Sundadontina Isannaia Namkongnaia Thaiconcha Bineurus Pseudodon walpolei Pseudodon vagulus Pseudodon vondembuschianus Pseudodon mekongi SCHEPMANIINI Pseudodon lenyanensis 2 Pseudodon inoscularis Pseudodon lenyanensis 1 CHAMBERLAINIINI Parvasolenaia Pseudopostolata Koreosolenaia Cosmopseudodon Guiunio Ptychorhynchus Obovalis Postolata Caudiculatus LAMPROTULINI Sinosolenaia carinata Gonidea Microcondylaea Leguminaia Inversidens Sinosolenaia recognita Sinosolenaia iridinea Sinosolenaia oleivora

Prié et al., 2026 [Publ 4844], fig. 2

cst iT tT bi riS oT roT roS iD
0.826 5 20 15 5 18 15 1 17.5
Cladomics cladogram MARGARITIFERINAE Echyridella Westralunio HYRIDELLINI Velesunio Alathyria Lortiella Microdontia Diplodon Guyanhyrida CASTALIINI Iridea HYRIINI

Lawrie et al., 2026 [Publ 4843], fig. 1

cst iT tT bi riS oT roT roS iD
0.733 11 11 16 5 7 6 1 15.0
Cladomics cladogram HYRIDELLINI CASTALIINI HYRIINI Westralunio Velesunio 6 Alathyria jacksoni 2 Alathyria profuga Velesunio ambiguus Velesunio 5 Alathyria pertexta 2 Velesunio 4 Alathyria jacksoni 1 Alathyria pertexta 1 Lortiella Velesunio 3 Microdontia Velesunio angasi Velesunio 1 Velesunio 2

Bolotov et al., 2016 [Publ 3743], fig. 2

cst iT tT bi riS oT roT roS iD
0.923 6 6 4 4 24 3 1 5.0
Cladomics cladogram HYRIINAE Microdontia Lortiella Alathyria pertexta Velesunio Alathyria jacksoni Alathyria profuga

Bolotov et al., 2016 [Publ 3743], fig. 4

cst iT tT bi riS oT roT roS iD
0.923 6 6 4 4 24 3 1 5.0
Cladomics cladogram HYRIINAE Microdontia Alathyria pertexta Lortiella Velesunio Alathyria jacksoni Alathyria profuga

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
0.923 6 6 4 4 133 3 1 3.0
Cladomics cladogram HYRIINAE Alathyria pertexta Microdontia Lortiella Velesunio Alathyria jacksoni Alathyria profuga

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
0.923 6 6 4 4 133 3 1 5.0
Cladomics cladogram HYRIINAE Velesunio Alathyria jacksoni Alathyria profuga Microdontia Alathyria pertexta Lortiella

Lawrie et al., 2026 [Publ 4843], fig. 2

cst iT tT bi riS oT roT roS iD
0.667 9 9 13 4 0 0 0 21.0
Cladomics cladogram Velesunio 9 Velesunio ambiguus Alathyria jacksoni 4 Alathyria profuga Velesunio 5 Velesunio 8 Microdontia Lortiella Velesunio angasi Velesunio 6 Velesunio 7 Velesunio 4 Alathyria jacksoni 1 Alathyria jacksoni 2 Alathyria jacksoni 3 Alathyria pertexta 1 Velesunio 1 Velesunio 2 Velesunio 3 Alathyria pertexta 2

Walker et al., 2006 [Publ 2666], fig. 6

cst iT tT bi riS oT roT roS iD
0.692 5 5 4 3 31 5 1 9.0
Cladomics cladogram CASTALIINI RHIPIDODONTINI Hyridella depressa Hyridella australis Echyridella Velesunio ambiguus Velesunio 3 Velesunio 2 Alathyria Velesunio 1 Velesunio angasi Lortiella

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

cst iT tT bi riS oT roT roS iD
0.900 2 2 0 2 32 3 1 --
Cladomics cladogram CASTALIINI RHIPIDODONTINI Echyridella Velesunio Lortiella

Hoeh et al., 2001 [Publ 2652], fig. 14f2

cst iT tT bi riS oT roT roS iD
0.800 2 5 0 2 29 3 1 10.0
Cladomics cladogram IRIDININAE Anodontites guanarensis Anodontites trigona MONOCONDYLAEINAE RHIPIDODONTINI Echyridella Lortiella CASTALIINI Velesunio

Hoeh et al., 2001 [Publ 2652], fig. 14f4

cst iT tT bi riS oT roT roS iD
0.900 2 2 0 2 29 3 1 --
Cladomics cladogram CASTALIINI RHIPIDODONTINI Echyridella Lortiella Velesunio

Hoeh et al., 2002 [Publ 2653], fig. 2

cst iT tT bi riS oT roT roS iD
0.900 2 2 0 2 29 3 1 --
Cladomics cladogram CASTALIINI RHIPIDODONTINI Echyridella Velesunio Lortiella

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

cst iT tT bi riS oT roT roS iD
0.800 2 5 0 2 29 3 1 10.0
Cladomics cladogram IRIDININAE MONOCONDYLAEINAE ANODONTITINAE CASTALIINI Velesunio Echyridella Lortiella RHIPIDODONTINI

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

cst iT tT bi riS oT roT roS iD
0.900 2 2 0 2 29 3 1 --
Cladomics cladogram CASTALIINI RHIPIDODONTINI Echyridella Velesunio Lortiella

Baker et al., 2004 [Publ 2685], fig. 2-3

cst iT tT bi riS oT roT roS iD
0.667 3 3 4 2 6 5 1 9.5
Cladomics cladogram Hyridella depressa Hyridella australis RHIPIDODONTINI Echyridella CASTALIINI Velesunio 3 Velesunio 4 Velesunio 2 Velesunio angasi Velesunio 1 Alathyria

Graf & Cummings, 2006 [Publ 1968], fig. 2

cst iT tT bi riS oT roT roS iD
0.917 3 3 1 2 44 6 1 7.0
Cladomics cladogram HYRIINAE Velesunio ambiguus Velesunio angasi Lortiella

Hoeh et al., 2009 [Publ 2716], fig. 3

cst iT tT bi riS oT roT roS iD
0.917 3 3 1 2 44 6 1 7.0
Cladomics cladogram HYRIINAE Velesunio ambiguus Velesunio angasi Lortiella

Hoeh et al., 2009 [Publ 2716], fig. 4

cst iT tT bi riS oT roT roS iD
0.917 3 3 1 2 43 6 1 7.0
Cladomics cladogram HYRIINAE Velesunio ambiguus Velesunio angasi Lortiella

Hoeh et al., 2009 [Publ 2716], fig. 5

cst iT tT bi riS oT roT roS iD
0.917 3 3 1 2 43 6 1 7.0
Cladomics cladogram HYRIINAE Velesunio ambiguus Velesunio angasi Lortiella

Marshall et al., 2014 [Publ 3593], fig. 1

cst iT tT bi riS oT roT roS iD
0.818 3 3 1 2 11 7 1 6.5
Cladomics cladogram Echyridella HYRIDELLINI RHIPIDODONTINI Velesunio ambiguus Velesunio Lortiella

Klunzinger et al., 2021 [Publ 4086], fig. 2

cst iT tT bi riS oT roT roS iD
0.857 3 4 2 2 3 1 1 7.0
Cladomics cladogram MARGARITIFERINAE UNIONINAE Westralunio Velesunio HYRIINAE

Klunzinger et al., 2022 [Publ 4259], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 1 1 1 6.0
Cladomics cladogram HYRIINAE Velesunio Westralunio

Graf, 2000 [Publ 2647], fig. 1

cst iT tT bi riS oT roT roS iD
0.900 1 1 0 1 16 3 1 --
Cladomics cladogram CASTALIINI RHIPIDODONTINI HYRIDELLINI Velesunio

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

cst iT tT bi riS oT roT roS iD
0.818 1 1 0 1 10 5 1 --
Cladomics cladogram Velesunio HYRIINAE

Graf, 2002 [Publ 2651], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 3 1 --
Cladomics cladogram Velesunio HYRIINAE

Baker et al., 2003 [Publ 2690], fig. 5

cst iT tT bi riS oT roT roS iD
0.727 2 2 0 1 6 5 1 --
Cladomics cladogram Hyridella australis RHIPIDODONTINI Hyridella depressa Velesunio Echyridella CASTALIINI

Whelan et al., 2011 [Publ 3380], fig. 1-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 3 1 --
Cladomics cladogram Velesunio HYRIINAE

Whelan et al., 2011 [Publ 3380], fig. 1-MP

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 3 1 --
Cladomics cladogram Velesunio HYRIINAE

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 39 3 1 --
Cladomics cladogram Velesunio HYRIINAE

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 80 1 1 --
Cladomics cladogram HYRIINAE Velesunio

Combosch et al., 2017 [Publ 3806], fig. 1

cst iT tT bi riS oT roT roS iD
0.875 1 1 0 1 212 2 1 --
Cladomics cladogram HYRIINI Velesunio HYRIDELLINI

Combosch et al., 2017 [Publ 3806], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 213 2 1 --
Cladomics cladogram Velesunio HYRIINAE

Huang et al., 2018 [Publ 3828], fig. 3_5

cst iT tT bi riS oT roT roS iD
0.875 1 1 0 1 28 2 1 --
Cladomics cladogram HYRIINI Velesunio HYRIDELLINI

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

cst iT tT bi riS oT roT roS iD
0.875 1 1 0 1 27 2 1 --
Cladomics cladogram HYRIINI Velesunio HYRIDELLINI

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

cst iT tT bi riS oT roT roS iD
0.875 1 1 0 1 27 2 1 --
Cladomics cladogram HYRIINI Velesunio HYRIDELLINI

Guerra et al., 2019 [Publ 4168], fig. 1

cst iT tT bi riS oT roT roS iD
0.909 1 1 0 1 40 4 1 --
Cladomics cladogram Echyridella Westralunio RHIPIDODONTINI CASTALIINI HYRIINI

Guerra et al., 2019 [Publ 4168], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 1 1 --
Cladomics cladogram HYRIINAE Westralunio

Guerra et al., 2019 [Publ 4168], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 4 1 --
Cladomics cladogram Westralunio HYRIINAE

Guerra et al., 2019 [Publ 4168], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 1 1 --
Cladomics cladogram HYRIINAE Westralunio

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 121 1 1 --
Cladomics cladogram HYRIINAE Velesunio

Wu et al., 2020 [Publ 4121], fig. 2

cst iT tT bi riS oT roT roS iD
0.875 1 1 0 1 33 2 1 --
Cladomics cladogram HYRIINI HYRIDELLINI Velesunio

Wu et al., 2020 [Publ 4121], fig. S1

cst iT tT bi riS oT roT roS iD
0.875 1 1 0 1 33 2 1 --
Cladomics cladogram HYRIINI HYRIDELLINI Velesunio

Sano et al., 2020 [Publ 4138], fig. 2

cst iT tT bi riS oT roT roS iD
0.875 1 1 0 1 68 2 1 --
Cladomics cladogram HYRIINI HYRIDELLINI Velesunio

Sano et al., 2020 [Publ 4138], fig. 3

cst iT tT bi riS oT roT roS iD
0.875 1 1 0 1 68 2 1 --
Cladomics cladogram HYRIINI HYRIDELLINI Velesunio

Sano et al., 2020 [Publ 4138], fig. 4

cst iT tT bi riS oT roT roS iD
0.875 1 1 0 1 68 2 1 --
Cladomics cladogram HYRIINI HYRIDELLINI Velesunio

Sonowal et al., 2021 [Publ 4221], fig. 3a

cst iT tT bi riS oT roT roS iD
0.875 1 1 0 1 14 2 1 --
Cladomics cladogram HYRIINI HYRIDELLINI Velesunio

Sonowal et al., 2021 [Publ 4221], fig. 3b

cst iT tT bi riS oT roT roS iD
0.875 1 1 0 1 14 2 1 --
Cladomics cladogram HYRIINI Velesunio HYRIDELLINI

Sonowal et al., 2021 [Publ 4221], fig. 4

cst iT tT bi riS oT roT roS iD
0.875 1 1 0 1 15 2 1 --
Cladomics cladogram HYRIINI Velesunio HYRIDELLINI

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 2 1 --
Cladomics cladogram Alathyria HYRIINAE

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 2 1 --
Cladomics cladogram Alathyria HYRIINAE

Gomes-dos-Santos et al., 2023 [Publ 4463], fig. 2a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 20 1 1 --
Cladomics cladogram HYRIINAE Westralunio

Gomes-dos-Santos et al., 2023 [Publ 4463], fig. 2b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 1 1 --
Cladomics cladogram Westralunio HYRIINAE

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 139 4 1 --
Cladomics cladogram Westralunio HYRIINAE

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 46 1 1 --
Cladomics cladogram HYRIINAE Westralunio

Plazzi et al., 2011 [Publ 3676], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 0 0 --
Cladomics cladogram Alathyria LAMPROTULINI CHAMBERLAINIINI

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 --
Cladomics cladogram MARGARITIFERINAE Velesunio PARREYSIINAE UNIONINAE AMBLEMINAE GONIDEINAE

Bieler et al., 2014 [Publ 3622], fig. 26

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram Velesunio UNIONINAE ASPATHARIINAE MARGARITIFERINAE

Bieler et al., 2014 [Publ 3622], fig. 27

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram Velesunio UNIONINAE ASPATHARIINAE MARGARITIFERINAE

Bieler et al., 2014 [Publ 3622], fig. 28

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram UNIONINAE Velesunio

Bieler et al., 2014 [Publ 3622], fig. 29

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram Velesunio UNIONINAE ASPATHARIINAE MARGARITIFERINAE

Bieler et al., 2014 [Publ 3622], fig. 30

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram Velesunio UNIONINAE ASPATHARIINAE MARGARITIFERINAE

Bieler et al., 2014 [Publ 3622], fig. 31

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 97 0 0 --
Cladomics cladogram Velesunio UNIONINAE MARGARITIFERINAE ASPATHARIINAE

González & Giribet, 2015 [Publ 3632], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram UNIONINAE Velesunio

González & Giribet, 2015 [Publ 3632], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram UNIONINAE Velesunio ASPATHARIINAE MARGARITIFERINAE

González & Giribet, 2015 [Publ 3632], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram Velesunio UNIONINAE MARGARITIFERINAE

González & Giribet, 2015 [Publ 3632], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram Velesunio UNIONINAE ASPATHARIINAE MARGARITIFERINAE

Elderkin et al., 2016 [Publ 3693], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 2 0 0 --
Cladomics cladogram Etheria ASPATHARIINAE Velesunio