Bivalvia | Unionida | Hyrioidea | Hyriidae

subfamily VELESUNIONINAE Iredale, 1934: cladomics

6 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 17 (21 total)
number of subtaxa (rank: genus) = 6
analyzed in 65 cladograms in 38 publications
maximum number of terminals in an analysis = 10 (mean = 2.38, median = 1)
maximum number of subtaxa (genus) = 5 (mean = 1.86, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 27
maximum number of included terminals = 10, (mean = 4.33 median = 3)
maximum number of subtaxa (genus) = 5 (mean = 3.07, median = 2)
proportion of cladograms that support basic monophyly = 0.67

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 26
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 10 (mean = 4.42, median = 3)
maximum number of subtaxa (genus) = 5 (mean = 3.15, median = 2)
proportion of cladograms that support rigorous monophyly = 0.65

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Velesunioninae cladogram distances
PCoA Ordination

NJ Tree of Velesunioninae cladogram distances
NJ Tree

Cladograms

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
10 28 34 5 620 18 1 0 
  +----------------------------------- Westralunio **           
  |                                                          
  |                             +----- Diplodon_suavidicus__2
  |  +--------------------------+                            
--+  |                          |  +-- Hyriini               
  |  |                          +--+                         
  |  |                             +-- Castaliini            
  |  |                                                       
  |  |                             +-- Hyridella_depressa    
  +--+                       +-----+                         
     |                       |     +-- Hyridella_drapeta     
     |  +--------------------+                               
     |  |                    |  +----- Diplodon_suavidicus__1
     |  |                    +--+                            
     |  |                       |  +-- Diplodon_chilensis    
     |  |                       +--+                         
     +--+                          +-- Diplodon_gratus       
        |                                                    
        |                          +-- Cucumerunio           
        |  +-----------------------+                         
        |  |                       +-- Hyridella_australis   
        |  |                                                 
        +--+  +----------------------- Echyridella           
           |  |                                              
           |  |              +-------- Velesunio__1 **          
           +--+  +-----------+                               
              |  |           |  +----- Alathyria_jacksoni__1 ** 
              |  |           +--+                            
              |  |              |  +-- Velesunio_ambiguus **    
              +--+              +--+                         
                 |                 +-- Alathyria_profuga **     
                 |                                           
                 |  +----------------- Velesunio__3 **          
                 |  |                                        
                 +--+  +-------------- Microdontia **           
                    |  |                                     
                    +--+           +-- Lortiella **             
                       |  +--------+                         
                       |  |        +-- Velesunio_angasi **      
                       +--+                                  
                          |  +-------- Alathyria_pertexta__2 ** 
                          +--+                               
                             |  +----- Velesunio__2 **          
                             +--+                            
                                |  +-- Alathyria_jacksoni__2 ** 
                                +--+                         
                                   +-- Alathyria_pertexta__1 **

Graf et al., 2015 [Publ 3619], fig. 1

iTtTbiriSoTroTroSiD
7 7 5 5 13 7 1 36 
  +-------------- Hyriinae          
  |                                 
--+  +----------- Westralunio **       
  |  |                              
  |  |     +----- Alathyria_jacksoni **
  +--+  +--+                        
     |  |  |  +-- Velesunio **         
     |  |  +--+                     
     +--+     +-- Alathyria_profuga ** 
        |                           
        |  +----- Alathyria_pertexta **
        +--+                        
           |  +-- Microdontia **       
           +--+                     
              +-- Lortiella **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
7 14 9 5 194 7 1 38 
  +----------- Echyridella       
  |                              
  +----------- Westralunio **       
  |                              
  +----------- Hyridellini       
  |                              
  |     +----- Rhipidodontini    
  +-----+                        
--+     |  +-- Castaliini        
  |     +--+                     
  |        +-- Hyriini           
  |                              
  |     +----- Velesunio **         
  |  +--+                        
  |  |  |  +-- Alathyria_jacksoni **
  |  |  +--+                     
  +--+     +-- Alathyria_profuga ** 
     |                           
     |  +----- Alathyria_pertexta **
     +--+                        
        |  +-- Lortiella **         
        +--+                     
           +-- Microdontia **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
7 14 12 5 192 7 1 41 
           +-- Westralunio **       
  +--------+                     
  |        +-- Hyriinae          
  |                              
--+     +----- Velesunio **         
  |  +--+                        
  |  |  |  +-- Alathyria_jacksoni **
  |  |  +--+                     
  +--+     +-- Alathyria_profuga ** 
     |                           
     |  +----- Alathyria_pertexta **
     +--+                        
        |  +-- Lortiella **         
        +--+                     
           +-- Microdontia **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
7 14 12 5 194 7 1 39 
           +-- Westralunio **       
  +--------+                     
  |        +-- Hyriinae          
  |                              
--+     +----- Alathyria_jacksoni **
  |  +--+                        
  |  |  |  +-- Alathyria_profuga ** 
  |  |  +--+                     
  +--+     +-- Velesunio **         
     |                           
     |  +----- Alathyria_pertexta **
     +--+                        
        |  +-- Lortiella **         
        +--+                     
           +-- Microdontia **

Santos-Neto et al., 2016 [Publ 3739], fig. 4

iTtTbiriSoTroTroSiD
6 13 11 5 14 13 1 31.5 
                    +----- Diplodon_suavidicus__2
  +-----------------+                            
  |                 |  +-- Hyriini               
  |                 +--+                         
  |                    +-- Castaliini            
  |                                              
  |              +-------- Westralunio **           
--+              |                               
  |  +-----------+     +-- Hyridella_drapeta     
  |  |           |  +--+                         
  |  |           |  |  +-- Hyridella_depressa    
  |  |           +--+                            
  |  |              |  +-- Diplodon_suavidicus__1
  +--+              +--+                         
     |                 +-- Diplodon_gratus       
     |                                           
     |  +----------------- Echyridella           
     |  |                                        
     +--+              +-- Cucumerunio           
        |  +-----------+                         
        |  |           +-- Hyridella_australis   
        +--+                                     
           |  +----------- Velesunio_ambiguus **    
           +--+                                  
              |  +-------- Alathyria **             
              +--+                               
                 |  +----- Velesunio_angasi **      
                 +--+                            
                    |  +-- Lortiella **             
                    +--+                         
                       +-- Microdontia **

Bolotov et al., 2016 [Publ 3743], fig. 4

iTtTbiriSoTroTroSiD
6 6 4 4 24 3 1 39 
  +----------- Hyriinae          
  |                              
  |     +----- Microdontia **       
--+  +--+                        
  |  |  |  +-- Alathyria_pertexta **
  |  |  +--+                     
  +--+     +-- Lortiella **         
     |                           
     |  +----- Velesunio **         
     +--+                        
        |  +-- Alathyria_jacksoni **
        +--+                     
           +-- Alathyria_profuga **

Bolotov et al., 2016 [Publ 3743], fig. 2

iTtTbiriSoTroTroSiD
6 6 4 4 24 3 1 39 
  +----------- Hyriinae          
  |                              
  |     +----- Microdontia **       
--+  +--+                        
  |  |  |  +-- Lortiella **         
  |  |  +--+                     
  +--+     +-- Alathyria_pertexta **
     |                           
     |  +----- Velesunio **         
     +--+                        
        |  +-- Alathyria_jacksoni **
        +--+                     
           +-- Alathyria_profuga **

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

iTtTbiriSoTroTroSiD
6 19 20 5 14 13 1 22.5 
                 +----- Diplodon_suavidicus__1
  +--------------+                            
  |              |  +-- Hyriini               
  |              +--+                         
  |                 +-- Castaliini            
  |                                           
  |           +-------- Westralunio **           
  |           |                               
  +-----------+     +-- Hyridella_drapeta     
  |           |  +--+                         
--+           |  |  +-- Hyridella_depressa    
  |           +--+                            
  |              |  +-- Diplodon_gratus       
  |              +--+                         
  |                 +-- Diplodon_suavidicus__2
  |                                           
  |                 +-- Hyridella_australis   
  |  +--------------+                         
  |  |              +-- Cucumerunio           
  +--+                                        
     |  +-------------- Echyridella           
     +--+                                     
        |  +----------- Velesunio_ambiguus **    
        +--+                                  
           |  +-------- Microdontia **           
           +--+                               
              |  +----- Lortiella **             
              +--+                            
                 |  +-- Alathyria **             
                 +--+                         
                    +-- Velesunio_angasi **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
6 6 4 4 133 3 1 39 
  +----------- Hyriinae          
  |                              
  |     +----- Velesunio **         
--+  +--+                        
  |  |  |  +-- Alathyria_jacksoni **
  |  |  +--+                     
  +--+     +-- Alathyria_profuga ** 
     |                           
     |  +----- Microdontia **       
     +--+                        
        |  +-- Alathyria_pertexta **
        +--+                     
           +-- Lortiella **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
6 6 4 4 133 3 1 39 
  +----------- Hyriinae          
  |                              
  |     +----- Alathyria_pertexta **
--+  +--+                        
  |  |  |  +-- Microdontia **       
  |  |  +--+                     
  +--+     +-- Lortiella **         
     |                           
     |  +----- Velesunio **         
     +--+                        
        |  +-- Alathyria_jacksoni **
        +--+                     
           +-- Alathyria_profuga **

Walker et al., 2006 [Publ 2666], fig. 6

iTtTbiriSoTroTroSiD
5 5 4 3 31 5 1 30 
  +----------------- Castaliini         
  |                                     
  +----------------- Rhipidodontini     
--+                                     
  |  +-------------- Hyridella_depressa 
  |  |                                  
  |  +-------------- Hyridella_australis
  +--+                                  
     |  +----------- Echyridella        
     |  |                               
     |  |        +-- Velesunio_ambiguus ** 
     +--+  +-----+                      
        |  |     +-- Velesunio__3 **       
        |  |                            
        +--+  +----- Velesunio__2 **       
           |  |                         
           |  |  +-- Alathyria **          
           +--+--+                      
              |  +-- Velesunio__1 **       
              |                         
              |  +-- Velesunio_angasi **   
              +--+                      
                 +-- Lortiella **

Klunzinger et al., 2022 [Publ 4259], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 2 1 1 1 33 
  +----- Hyriinae   
--+                 
  |  +-- Velesunio **  
  +--+              
     +-- Westralunio **

Baker et al., 2004 [Publ 2685], fig. 2-3

iTtTbiriSoTroTroSiD
3 3 4 2 6 5 1 32 
              +----- Hyridella_depressa 
  +-----------+                         
  |           |  +-- Hyridella_australis
  |           +--+                      
  |              +-- Rhipidodontini     
--+                                     
  |              +-- Echyridella        
  |  +-----------+                      
  |  |           +-- Castaliini         
  |  |                                  
  +--+           +-- Velesunio__3 **       
     |  +--------+                      
     |  |        +-- Velesunio__4 **       
     +--+                               
        |  +-------- Velesunio__2 **       
        +--+                            
           |  +----- Velesunio_angasi **   
           +--+                         
              |  +-- Velesunio__1 **       
              +--+                      
                 +-- Alathyria **

Klunzinger et al., 2021 [Publ 4086], fig. 2

iTtTbiriSoTroTroSiD
3 4 2 2 3 1 1 32 
  +----- Westralunio **
--+                 
  |  +-- Velesunio **  
  +--+              
     +-- Hyriinae

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 2 44 6 1 33 
  +-------- Hyriinae          
--+                           
  |  +----- Velesunio_ambiguus **
  +--+                        
     |  +-- Velesunio_angasi **  
     +--+                     
        +-- Lortiella **

Marshall et al., 2014 [Publ 3593], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 2 11 7 1 34.5 
  +----------- Echyridella       
  |                              
--+        +-- Hyridellini       
  |  +-----+                     
  |  |     +-- Rhipidodontini    
  +--+                           
     |  +----- Velesunio_ambiguus **
     +--+                        
        |  +-- Velesunio **         
        +--+                     
           +-- Lortiella **

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
3 3 1 2 43 6 1 33 
  +-------- Hyriinae          
--+                           
  |  +----- Velesunio_ambiguus **
  +--+                        
     |  +-- Velesunio_angasi **  
     +--+                     
        +-- Lortiella **

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 2 43 6 1 33 
  +-------- Hyriinae          
--+                           
  |  +----- Velesunio_ambiguus **
  +--+                        
     |  +-- Velesunio_angasi **  
     +--+                     
        +-- Lortiella **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 2 44 6 1 33 
  +-------- Hyriinae          
--+                           
  |  +----- Velesunio_ambiguus **
  +--+                        
     |  +-- Velesunio_angasi **  
     +--+                     
        +-- Lortiella **

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
2 2 0 2 29 3 1 -- 
        +-- Castaliini    
  +-----+                 
  |     +-- Rhipidodontini
--+                       
  |  +----- Echyridella   
  +--+                    
     |  +-- Lortiella **     
     +--+                 
        +-- Velesunio **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
2 5 0 2 29 3 1 35 
  +----- Rhipidodontini
  |                    
  +----- Echyridella   
--+                    
  +----- Lortiella **     
  |                    
  |  +-- Castaliini    
  +--+                 
     +-- Velesunio **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
2 2 0 2 29 3 1 -- 
        +-- Castaliini    
  +-----+                 
  |     +-- Rhipidodontini
--+                       
  |  +----- Echyridella   
  +--+                    
     |  +-- Velesunio **     
     +--+                 
        +-- Lortiella **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
2 5 0 2 29 3 1 36 
     +-- Castaliini    
  +--+                 
  |  +-- Velesunio **     
--+                    
  |  +-- Echyridella   
  |  |                 
  +--+-- Lortiella **     
     |                 
     +-- Rhipidodontini

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 32 3 1 -- 
        +-- Castaliini    
  +-----+                 
  |     +-- Rhipidodontini
--+                       
  |  +----- Echyridella   
  +--+                    
     |  +-- Velesunio **     
     +--+                 
        +-- Lortiella **

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 29 3 1 -- 
        +-- Castaliini    
  +-----+                 
  |     +-- Rhipidodontini
--+                       
  |  +----- Echyridella   
  +--+                    
     |  +-- Velesunio **     
     +--+                 
        +-- Lortiella **

Baker et al., 2003 [Publ 2690], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 1 6 5 1 -- 
           +-- Hyridella_australis
  +--------+                      
  |        +-- Rhipidodontini     
--+                               
  |  +-------- Hyridella_depressa 
  +--+                            
     |  +----- Velesunio **          
     +--+                         
        |  +-- Echyridella        
        +--+                      
           +-- Castaliini

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 15 2 1 -- 
  +----- Hyriini    
--+                 
  |  +-- Velesunio **  
  +--+              
     +-- Hyridellini

Combosch et al., 2017 [Publ 3806], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 14 2 1 -- 
  +----- Hyriini    
--+                 
  |  +-- Velesunio **  
  +--+              
     +-- Hyridellini

Elderkin et al., 2016 [Publ 3693], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 2 0 0 -- 
-- Velesunio_ambiguus **

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 1 1 -- 
  +-- Hyriinae   
--+              
  +-- Westralunio **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
1 1 0 1 35 3 1 -- 
  +-- Velesunio **
--+            
  +-- Hyriinae

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 33 2 1 -- 
  +----- Hyriini    
--+                 
  |  +-- Hyridellini
  +--+              
     +-- Velesunio **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
1 1 0 1 121 1 1 -- 
  +-- Hyriinae 
--+            
  +-- Velesunio **

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

iTtTbiriSoTroTroSiD
1 1 0 1 10 5 1 -- 
  +-- Velesunio **
--+            
  +-- Hyriinae

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 39 3 1 -- 
  +-- Velesunio **
--+            
  +-- Hyriinae

González & Giribet, 2015 [Publ 3632], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Velesunio_ambiguus **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
1 1 0 1 90 2 1 -- 
  +-- Alathyria **
--+            
  +-- Hyriinae

González & Giribet, 2015 [Publ 3632], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Velesunio_ambiguus **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 4 1 -- 
  +-- Westralunio **
--+              
  +-- Hyriinae

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 46 1 1 -- 
  +-- Hyriinae   
--+              
  +-- Westralunio **

Bieler et al., 2014 [Publ 3622], fig. 28

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Velesunio_ambiguus **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 68 2 1 -- 
  +----- Hyriini    
--+                 
  |  +-- Hyridellini
  +--+              
     +-- Velesunio **

Graf, 2002 [Publ 2651], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 17 3 1 -- 
  +-- Velesunio **
--+            
  +-- Hyriinae

Bieler et al., 2014 [Publ 3622], fig. 26_27_29_30

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Velesunio_ambiguus **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
1 1 0 1 139 4 1 -- 
  +-- Westralunio **
--+              
  +-- Hyriinae

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
1 1 0 1 35 3 1 -- 
  +-- Velesunio **
--+            
  +-- Hyriinae

González & Giribet, 2015 [Publ 3632], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Velesunio_ambiguus **

Plazzi et al., 2011 [Publ 3676], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 17 0 0 -- 
-- Alathyria_jacksoni **

Combosch et al., 2017 [Publ 3806], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 2 1 -- 
  +-- Velesunio **
--+            
  +-- Hyriinae

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 33 2 1 -- 
  +----- Hyriini    
--+                 
  |  +-- Hyridellini
  +--+              
     +-- Velesunio **

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
1 1 0 1 14 2 1 -- 
  +-- Hyriini    
  |              
--+-- Velesunio **  
  |              
  +-- Hyridellini

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 4 1 -- 
        +-- Echyridella   
  +-----+                 
  |     +-- Westralunio **   
--+                       
  |  +----- Rhipidodontini
  +--+                    
     |  +-- Castaliini    
     +--+                 
        +-- Hyriini

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
1 1 0 1 28 2 1 -- 
  +----- Hyriini    
--+                 
  |  +-- Velesunio **  
  +--+              
     +-- Hyridellini

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 27 2 1 -- 
  +----- Hyriini    
--+                 
  |  +-- Velesunio **  
  +--+              
     +-- Hyridellini

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 90 2 1 -- 
  +-- Alathyria **
--+            
  +-- Hyriinae

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Velesunio_ambiguus **

Graf, 2000 [Publ 2647], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 16 3 1 -- 
  +-------- Castaliini    
--+                       
  |  +----- Rhipidodontini
  +--+                    
     |  +-- Hyridellini   
     +--+                 
        +-- Velesunio **

González & Giribet, 2015 [Publ 3632], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Velesunio_ambiguus **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
1 1 0 1 80 1 1 -- 
  +-- Hyriinae 
--+            
  +-- Velesunio **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 68 2 1 -- 
  +----- Hyriini    
--+                 
  |  +-- Hyridellini
  +--+              
     +-- Velesunio **

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 1 1 -- 
  +-- Hyriinae   
--+              
  +-- Westralunio **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 68 2 1 -- 
  +----- Hyriini    
--+                 
  |  +-- Hyridellini
  +--+              
     +-- Velesunio **

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
1 1 0 1 14 2 1 -- 
  +-- Hyriini    
  |              
--+-- Hyridellini
  |              
  +-- Velesunio **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 27 2 1 -- 
  +----- Hyriini    
--+                 
  |  +-- Velesunio **  
  +--+              
     +-- Hyridellini