Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Pleurobemini | Fusconaia
8 specimens + 1 GenBank voucher
Rafinesque, 1820. Ann. Gén. Sci. Phys. 5 (13): 308.
Rafinesque, 1864. [in] Binney & Tryon, reprint: 52.
(intro'd as intra-sp. of Obliquaria (Rotundaria) subrotunda Rafinesque, 1820)
Fusconaia subrotunda — Williams et al. (2008), IUCN (2012)
Fusconaia subrotunda subrotunda — Watters et al. (2009)
‘Lamarck’ Crouch, 1827. Illustr. Lamarck’s Conch.: 16, pl. 9, fig. 3.
(misidentified reference to Unio brevialis Lamarck, 1819)
Quadrula (sec. Fusconaia) subrotunda — Simpson (1900), Simpson (1914)
Fusconaia subrotunda — Burch (1975), Parmalee & Bogan (1998)

Lea, 1831. Trans. Amer. Phil. Soc. 4 [O. 1]: 117 [127], pl. 18, fig. 45.
Type(s): HOLOTYPE USNM_85832, Cincinnati, Ohio.
Quadrula (sec. Fusconaia) subrotunda — Simpson (1900), Simpson (1914)
Quadrula subrotunda — Ortmann (1909), Ortmann (1909), Ortmann (1909), Ortmann (1910)
Fusconaia subrotundus — Ortmann (1912), Turgeon et al. (1998)
Unio subrotundus — de Gregorio (1914), Henderson (1935)
Quadrula sintoxia — Vanatta (1915)
Fusconaia subrotunda — Ortmann (1919), Ortmann & Walker (1922), Ortmann (1924), Haas (1969), Burch (1975), ... FMCS (2021), Hopper et al. (2023), Hauck et al. (2023), FMCS (2025), Schilling et al. (2025)
Fusconaia subtrotunda [sic] — Ortmann (1925)
Quadrula (Fusconia [sic]) subrotunda — Frierson (1927)
Fusconaia polita — Morrison (1969)
Fusconaia ebena — Oesch (1984)
Fusconaia subrotunda subrotunda — Watters et al. (2009), ITIS (2012)

Lea, 1834. Trans. Amer. Phil. Soc. 5 [O. 1]: 98 [210], pl. 14, fig. 41.
Type(s): HOLOTYPE USNM_85815, Mahoning R., Ohio.
Quadrula (sec. Fusconaia) kirtlandiana — Simpson (1900), Simpson (1914)
Quadrula kirtlandiana — Ortmann (1909)
Quadrula subrotunda kirtlandiana — Ortmann (1909)
Quadrula subrotunda var. kirtlandiana — Ortmann (1910)
Fusconaia subrotundus — Ortmann (1912)
Fusconaia subrotunda kirtlandiana — Ortmann (1919)
Quadrula (Fusconia [sic]) kirtlandiana — Frierson (1927)
Fusconaia kirtlandiana — Haas (1969)
Fusconaia subrotunda — Burch (1975), R.I. Johnson (1980), Parmalee & Bogan (1998), Williams et al. (2008)
Fusconaia subrotunda subrotunda — Watters et al. (2009)
Fusconaia subrotunda kirklandiana [sic] — ITIS (2012)
‘Say’ Conrad, 1834. New Fresh-Water Shells: 71.
(misidentified reference to Unio personatus Say, 1829)
Quadrula (sec. Fusconaia) subrotunda — Simpson (1900), Simpson (1914)
Fusconaia subrotunda — Burch (1975), Parmalee & Bogan (1998), Williams et al. (2008)
Fusconaia subrotunda subrotunda — Watters et al. (2009)
Say, 1834. Amer. Conch. 6: ??. [nomen nudum]
Conrad, 1837. Monogr. Fam. Unionidae 1(8): 67, pl. 37, fig. 2.
Quadrula (sec. Fusconaia) subrotunda — Simpson (1900), Simpson (1914)
Quadrula (Fusconia [sic]) subrotunda — Frierson (1927)
Fusconaia subrotunda — Haas (1969), Burch (1975), Parmalee & Bogan (1998)
Fusconaia polita — Morrison (1969)
Fusconaia subrotunda subrotunda — Watters et al. (2009)
‘Lea’ of Authors.
Lea, 1838. Synopsis Fam. Naiades: 23.
Hanley, 1843. Illustr. Descr. Cat. Rec. Bivalve Shells: 203.
ITIS, 2012. ITIS: [web].
(error for Unio kirtlandianus Lea, 1834)
Quadrula (sec. Fusconaia) kirtlandiana — Simpson (1900), Simpson (1914)
Fusconaia subrotunda — Parmalee & Bogan (1998)
Fusconaia subrotunda subrotunda — Watters et al. (2009)

Lea, 1840. Proc. Amer. Phil. Soc. 1: 286.
Lea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 195 [33], pl. 8, fig. 6.
Type(s): HOLOTYPE USNM_85803, Holston R., Tenn.
Quadrula (sec. Fusconaia) pilaris — Simpson (1900), Simpson (1914)
Fusconaia pilaris lesueuriana — Ortmann (1918)
Quadrula (Fusconia [sic]) pilaris — Frierson (1927)
Fusconaia pilaris — Haas (1969)
Fusconaia subrotunda — Burch (1975), R.I. Johnson (1980), Parmalee & Bogan (1998), Williams et al. (2008)
Fusconaia subrotunda lesueuriana — Watters et al. (2009)
Fusconaia subrotunda lesueriana [sic] — ITIS (2012)

Lea, 1840. Proc. Amer. Phil. Soc. 1: 285.
Lea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 209 [47], pl. 14, fig. 24.
Type(s): HOLOTYPE USNM_85817, French Broad R., Tenn.
Quadrula (sec. Fusconaia) pilaris — Simpson (1900), Simpson (1914)
Fusconaia pilaris — Ortmann (1918), Haas (1969)
Quadrula (Fusconia [sic]) pilaris — Frierson (1927)
Fusconaia subrotunda — Burch (1975), R.I. Johnson (1980), Parmalee & Bogan (1998), Williams et al. (2008)
Fusconaia subrotunda pilaris — ITIS (2012)
‘Lea’ of Authors.
Conrad, 1853. Proc. Acad. Nat. Sci. 6: 251.
Frierson, 1927. Check List N. Amer. Naiades: 56.
(error for Unio lesueurianus Lea, 1840)
Lea, 1868. Proc. Acad. Nat. Sci. 20: 163.
Unio subrotundus — Henderson (1935)

Lea, 1871. Proc. Acad. Nat. Sci. 23: 191.
Lea, 1874. Jour. Acad. Nat. Sci. 8 [O. 13]: 5 [9], pl. 1, fig. 1.
Type(s): FIGURED SYNTYPE USNM_85820, Holston R., Tenn.
Quadrula (sec. Fusconaia) globata — Simpson (1900), Simpson (1914)
Fusconaia pilaris — Ortmann (1918), Haas (1969)
Quadrula (Fusconia [sic]) pilaris — Frierson (1927)
Fusconaia subrotunda — Burch (1975), Parmalee & Bogan (1998), Williams et al. (2008)
Call, 1894. Proc. Indiana Acad. Sci. 1893(3): ??.
(misidentified reference to Unio ebenus Lea, 1831)
Quadrula (Fusconia [sic]) subrotunda — Frierson (1927)

B.H. Wright, 1896. Nautilus 9(12): 133, pl. 3, fig. 2 [as ‘bursa-pastoris’].
Type(s): LECTOTYPE USNM_133427, Clinch R., Lee Co., Va.
Quadrula (sec. Fusconaia) bursa-pastoris — Simpson (1900), Simpson (1914)
Fusconaja [sic] bursa-pastoris — Ortmann (1913)
Fusconaia pilaris bursa-pastoris — Ortmann (1918)
Quadrula (Fusconia [sic]) bursa-pastoris — Frierson (1927)
Fusconaia bursapastoris — Haas (1969)
Fusconaia subrotunda — Burch (1975), R.I. Johnson (1980), Parmalee & Bogan (1998), Williams et al. (2008)
Simpson, 1900. Proc. Acad. Nat. Sci. 52: 83, pl. 2, fig. 8.
Simpson, 1900. Proc. U.S. Nat. Mus. 22: 790.
Simpson, 1914. Descr. Cat. Naiades: 887.
Quadrula (sec. Fusconaia) flexuosa — Simpson (1900), Simpson (1914)
Quadrula flexuosa — Simpson (1900)
Fusconaia pilaris lesueuriana — Ortmann (1918)
Quadrula (Fusconia [sic]) pilaris — Frierson (1927)
Fusconaia pilaris — Haas (1969)
Fusconaia subrotunda — Burch (1975), Parmalee & Bogan (1998), Williams et al. (2008)

Simpson, 1900. Proc. U.S. Nat. Mus. 22: 791.
Simpson, 1914. Descr. Cat. Naiades: 892.
(intro'd as intra-sp. of Unio kirtlandianus Lea, 1834)
Type(s): LECTOTYPE USNM_150155, Scott Co., Va.
Quadrula (sec. Fusconaia) kirtlandiana var. minor — Simpson (1900), Simpson (1914)
Fusconaia subrotunda — R.I. Johnson (1980), Parmalee & Bogan (1998), Williams et al. (2008)
Marsh, 1902. Nautilus 15(10): 115.
Marsh, 1902. Nautilus 16: 8, pl. 1, two upper figs. [as andrewsæ].
Quadrula (sec. Fusconaia) andrewsii — Simpson (1914)
Fusconaia pilaris — Ortmann (1918), Haas (1969)
Quadrula (Fusconia [sic]) pilaris — Frierson (1927)
Fusconaia subrotunda — Burch (1975), Parmalee & Bogan (1998), Williams et al. (2008)
Marsh, 1902. Nautilus 16: 7, pl. 1, lower two figs.
Quadrula (sec. Fusconaia) beauchampii — Simpson (1914)
Fusconaia pilaris — Ortmann (1918), Haas (1969)
Quadrula (Fusconia [sic]) pilaris — Frierson (1927)
Fusconaia subrotunda — Burch (1975), Parmalee & Bogan (1998), Williams et al. (2008)

Ortmann, 1913. Nautilus 27(8): 89.
(intro'd as intra-sp. of Unio subrotundus Lea, 1831)
Type(s): LECTOTYPE CM_61.5239 (by R.I. Johnson, 1977: 238), Elk River, Gassaway, Braxton County, West Virginia.
Fusconaja [sic] subrotunda leucogona — Ortmann (1913)
Fusconaia subrotunda — Burch (1975), R.I. Johnson (1980), Parmalee & Bogan (1998), Williams et al. (2008)
Fusconaia subrotunda subrotunda — Watters et al. (2009)
‘Lea’ Ferussac, 1835. Mag. Zool. 5(59-60): 29.
(error for Unio kirtlandianus Lea, 1834)
Marsh, 1902. Nautilus 16: 8, pl. 1, upper figs.
(error for Quadrula andrewsii Marsh, 1902)
Quadrula (sec. Fusconaia) andrewsii — Simpson (1914)
Quadrula (Fusconia [sic]) pilaris — Frierson (1927)
Fusconaia pilaris — Haas (1969)
Fusconaia subrotunda — Parmalee & Bogan (1998), Williams et al. (2008)
‘Lea’ Ortmann, 1925. Amer. Midl. Nat. 9: 328.
(error for Unio subrotundus Lea, 1831)
Fusconaia subrotunda — Burch (1975)
‘Lea’ ITIS, 2012. ITIS: [web].
(error for Unio lesueurianus Lea, 1840)
Quadrula (sec. Fusconaia) subrotunda — Simpson (1900), Simpson (1914)
Quadrula (sec. Fusconaia) kirtlandiana — Simpson (1900), Simpson (1914)
Quadrula (sec. Fusconaia) pilaris — Simpson (1900), Simpson (1914)
Quadrula (sec. Fusconaia) globata — Simpson (1900), Simpson (1914)
Quadrula (sec. Fusconaia) bursa-pastoris — Simpson (1900), Simpson (1914)
Quadrula (sec. Fusconaia) flexuosa — Simpson (1900), Simpson (1914)
Quadrula flexuosa — Simpson (1900)
Quadrula (sec. Fusconaia) kirtlandiana var. minor — Simpson (1900), Simpson (1914)
Quadrula subrotunda — Ortmann (1909), Ortmann (1909), Ortmann (1909), Ortmann (1910)
Quadrula kirtlandiana — Ortmann (1909)
Quadrula subrotunda kirtlandiana — Ortmann (1909)
Quadrula subrotunda var. kirtlandiana — Ortmann (1910)
Fusconaia subrotundus — Ortmann (1912), Turgeon et al. (1998)
Fusconaja [sic] bursa-pastoris — Ortmann (1913)
Fusconaja [sic] subrotunda leucogona — Ortmann (1913)
Unio subrotundus — de Gregorio (1914), Henderson (1935)
Quadrula (sec. Fusconaia) andrewsii — Simpson (1914)
Quadrula (sec. Fusconaia) beauchampii — Simpson (1914)
Quadrula sintoxia — Vanatta (1915)
Fusconaia pilaris lesueuriana — Ortmann (1918)
Fusconaia pilaris — Ortmann (1918), Haas (1969)
Fusconaia pilaris bursa-pastoris — Ortmann (1918)
Fusconaia subrotunda — Ortmann (1919), Ortmann & Walker (1922), Ortmann (1924), Haas (1969), Burch (1975), ... FMCS (2021), Hopper et al. (2023), Hauck et al. (2023), FMCS (2025), Schilling et al. (2025)
Fusconaia subrotunda kirtlandiana — Ortmann (1919)
Fusconaia subtrotunda [sic] — Ortmann (1925)
Quadrula (Fusconia [sic]) subrotunda — Frierson (1927)
Quadrula (Fusconia [sic]) kirtlandiana — Frierson (1927)
Quadrula (Fusconia [sic]) pilaris — Frierson (1927)
Quadrula (Fusconia [sic]) bursa-pastoris — Frierson (1927)
Fusconaia polita — Morrison (1969)
Fusconaia kirtlandiana — Haas (1969)
Fusconaia bursapastoris — Haas (1969)
Fusconaia ebena — Oesch (1984)
Fusconaia subrotunda subrotunda — Watters et al. (2009), ITIS (2012)
Fusconaia subrotunda lesueuriana — Watters et al. (2009)
Fusconaia subrotunda kirklandiana [sic] — ITIS (2012)
Fusconaia subrotunda lesueriana [sic] — ITIS (2012)
Fusconaia subrotunda pilaris — ITIS (2012)