species Nodularia douglasiae (Griffith & Pidgeon, 1833): synonymy

Unio douglasiae Griffith & Pidgeon, 1833

Griffith & Pidgeon, 1833. [in] Cuvier, Animal Kingdom 12(38): pl. 21, fig. 2 [as ‘douglasiæ’].
‘Gray’ Griffith & Pidgeon, 1834. [in] Cuvier, Animal Kingdom 12(39-40): 601 [as ‘douglasiæ’].

Nodularia (s.s.) douglasiæ — Simpson (1900)
Unio (Nodularia) douglasiae — Martens (1905), Zhadin (1938)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Nodularia (s.s.) douglasiae — Preston (1912), Haas (1924), Korniushin (1998)
Nodularia (s.s.) (Group of N. douglasiæ) douglasiæ — Simpson (1914)
Nodularia douglasiae — Haas (1920), Prozorova et al. (2005), Graf (2007), Graf & Cummings (2007), Bogan (2010), ... Campbell & Lydeard (2026), Wu et al. (2026), Wu et al. (2026), Wu et al. (2026), Li et al. (2026)
Unio douglasiae douglasiae — Haas (1930), Haas (1969)
Unio douglasiae — Zhadin (1965), Kil (1976), Habe (1977), Liu et al. (1979), Liu et al. (1980), ... Zhou et al. (2017), Araujo et al. (2018), Wang et al. (2018), Aunins et al. (2018), Wang et al. (2018)
Nodularidia douglasiae — He & Zhuang (2013)
“Unio douglasiae” ** — Ouyang et al. (2015)
“Nodularia douglasiae” ** — Liu et al. (2017), Klishko et al. (2019), Sayenko et al. (2021), Dai et al. (2024)
“Nodularia douglasiae” ** — Liu et al. (2017), Mabuchi et al. (2021)
“Nodularia douglasiae” ** — Cha et al. (2018), Fukata & Iigo (2019), Sayenko et al. (2021), Mabuchi et al. (2021), Kim et al. (2021), Lopes-Lima et al. (2024), Dai et al. (2024), Wu et al. (2024), Chen et al. (2025)
“Nodularia douglasiae” ** — Fukata & Iigo (2019), Sano et al. (2020), Kim et al. (2021), Pavan-Kumar et al. (2022), Dai et al. (2024)
“Nodularia cf. douglasiae” ** — Huang et al. (2019)
Nodularia douglasiae Clade C — Choi et al. (2020)
Nodularia douglasiae Clade B — Choi et al. (2020)
“Nodularia douglasiae Clade A” ** — Choi et al. (2020)
Nodularia douglasiae Clade A — Choi et al. (2020)
“Nodularia douglasiae Clade B” ** — Choi et al. (2020)
Nodularia douglasiae douglasiae — Bogatov (2022)
Nodularia sp. 2 — Dai et al. (2024)
“Nodularia douglasiae” ** — Klymus et al. (2025)

Unio murchisonianus Lea, 1834

Lea, 1834. Trans. Amer. Phil. Soc. 5 [O. 1]: 33, pl. 3, fig. 6.
HOLOTYPE (MONOTYPY) USNM_84156, China [“Calcutta, India” penciled out].

Nodularia (s.s.) douglasiæ — Simpson (1900)
Unio (Nodularia) douglasiae — Martens (1905)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Nodularia (s.s.) douglasiae — Preston (1912)
Nodularia (s.s.) (Group of N. douglasiæ) douglasiæ — Simpson (1914)
Unio douglasiae douglasiae — Haas (1969)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae — Lopes-Lima et al. (2020)

Unio osbeckii Philippi, 1844

Philippi, 1844. Zeitschr. f. Malakozool. 1: 164.
Philippi, 1847. Abbild. Beschr. Conch. 3: 45, pl. 3, fig. 1.

Nodularia (s.s.) douglasiæ — Simpson (1900)
Unio (Nodularia) douglasiae — Martens (1905)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Nodularia (s.s.) douglasiae — Preston (1912)
Nodularia (s.s.) (Group of N. douglasiæ) douglasiæ — Simpson (1914)
Unio douglasiae osbecki — Haas (1930)
Unio douglasiae douglasiae — Haas (1969)
Unio douglasiae — Liu et al. (1979)
Nodularia douglasiae — Prozorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)

Unio shanghaiensis Lea, 1859

Lea, 1859. Proc. Acad. Nat. Sci. 11: 153.
Lea, 1860. Jour. Acad. Nat. Sci. 4 [O. 7]: 242, pl. 36, fig. 121.
HOLOTYPE BMNH_1965171, Shanghai, China.

Nodularia (s.s.) douglasiæ var. shanghaiensis — Simpson (1900)
Unio (Nodularia) douglasiae — Martens (1905)
Nodularia (s.s.) douglasiae shanghaiensis — Haas (1911)
Nodularia (s.s.) douglasiae var. shanghaiensis — Preston (1912)
Nodularia (s.s.) (Group of N. douglasiæ) douglasiæ var. shanghaiensis — Simpson (1914)
Nodularia douglasiae — Haas (1920), Lopes-Lima et al. (2020)
Unio douglasiae douglasiae — Haas (1930), Haas (1969)
Unio douglasiae — Liu et al. (1979)
Nodularidia douglasiae — He & Zhuang (2013)

Unio wrightii Lea, 1865

Lea, 1865. Proc. Acad. Nat. Sci. 17: 75.
Lea, 1868. Jour. Acad. Nat. Sci. 6 [O. 12]: 283, pl. 39, fig. 97.
HOLOTYPE USNM_84129, China.

Nodularia (s.s.) douglasiæ — Simpson (1900)
Unio (Nodularia) douglasiae — Martens (1905)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Nodularia (s.s.) (Group of N. douglasiæ) douglasiæ — Simpson (1914)
Unio wrigtii [sic] — de Gregorio (1914)
Unio douglasiae douglasiae — Haas (1969)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae — Lopes-Lima et al. (2020)

Unio sculptus Deshayes, 1873

Deshayes, 1873. Nouvelles archives 9: 9, pl. 1, figs. 3-3a.
SYNTYPES MNHN-IM-2000-1779, Petchily, Chine Centrale.

Nodularia (s.s.) douglasiæ — Simpson (1900)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Nodularia (s.s.) douglasiae — Preston (1912)
Nodularia (s.s.) (Group of N. douglasiæ) douglasiæ — Simpson (1914)
Unio douglasiae osbecki — Haas (1930)
Unio douglasiae douglasiae — Haas (1969)
Unio douglasiae — Liu et al. (1979)
Unio sculptus — Yu et al. (2001)
Nodularia douglasiae — Prozorova et al. (2005), Schneider et al. (2013), Lopes-Lima et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)

Unio biwae Kobelt, 1879

Kobelt, 1879. Abh. Senckenb. Naturf. Ges. 11: 425, pl. 23, figs. 2-4.
LECTOTYPE SMF_3650 (by Zilch, 1967: 51), Biwa-See, Japan.

Nodularia (s.s.) biwæ — Simpson (1900)
Nodularia (s.s.) biwae — Haas (1911)
Nodularia (s.s.) (Group of N. douglasiæ) biwæ — Simpson (1914)
Unio biwae — Haas (1930), Haas (1969), Habe (1977), Habe (1991), Slugina (2006)
Unio douglasiae biwae — Kondo (1997), Kondo (2008)
Nodularia biwae — Bogan (2010)
Nodularidia douglasiae — He & Zhuang (2013)
“Nodularia douglasiae biwae” ** — Sano et al. (2017)
Nodularia douglasiae — Lopes-Lima et al. (2020), Mabuchi et al. (2021), Nagel (2025)
Nodularia douglasiae biwae — Mabuchi & Nishida (2020)

Unio douglasiae firmus Heude, 1885

(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833; homonym of Unio firmus Lea, 1866)
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65 bis, figs. 128b-b’.

Nodularia (s.s.) douglasiæ — Simpson (1900)
Unio douglasiae — Liu et al. (1979)
Nodularia douglasiae — Prozorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)

Unio douglasiae chloreus Heude, 1885

(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65 bis, fig. 128c.

Nodularia (s.s.) douglasiæ — Simpson (1900)
Unio douglasiae — Liu et al. (1979)
Nodularia douglasiae — Prozorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)

Unio douglasiae dactylinus Heude, 1885

(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65, fig. 128e.

Nodularia (s.s.) douglasiæ — Simpson (1900)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Nodularia (s.s.) douglasiae — Preston (1912)
Nodularia (s.s.) (Group of N. douglasiæ) douglasiæ — Simpson (1914)
Unio douglasiae douglasiae — Haas (1969)
Unio douglasiae — Liu et al. (1979)
Nodularia douglasiae — Prozorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)

Unio douglasiae hainanensis Heude, 1885

(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65, fig. 128f.

Unio douglasiae — Liu et al. (1979)
Nodularia douglasiae — Prozorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)

Unio douglasiae deltae Heude, 1885

(homonym of Unio favidens var. deltae Benson, 1862; intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65 bis, fig. 128d [as ‘deltæ’].

Unio douglasiae — Liu et al. (1979)
Nodularia douglasiae — Prozorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)

Unio jourdyi Morlet, 1886

Morlet, 1886. J. de Conch. 34: 76, pl. 13, figs. 5-5a.
SYNTYPES MNHN-IM-2000-33685, Bac Hat etang de la riviere Claire.

Nodularia (s.s.) jourdyi — Simpson (1900)
Unio jourdyi — Dautzenberg & Fischer (1905), Dautzenberg & Fischer (1905)
Oxynaia jourdyi — Haas (1913), Haas (1930), Thiele (1934), Haas (1969), Haas (1969), Dang et al. (1980), Graf & Cummings (2007), Bogan (2010), IUCN (2011), Zieritz et al. (2018), Do et al. (2018)
Nodularia (s.s.) (Group of N. pazii) jourdyi — Simpson (1914)
Nodularia jourdyi — Bolotov et al. (2018), Pfeiffer et al. (2019), Sayenko et al. (2021), Graf & Cummings (2021), Li et al. (2026)
Nodularia douglasiae — Wu et al. (2024)

Unio pictorum var. amurensis Mousson, 1887

(intro'd as intra-sp. of Mya pictorum Linnaeus, 1758)
Mousson, 1887. J. de Conch. 35: 26, pl. 1, fig. 8.

Nodularia (s.s.) douglasiæ var. shanghaiensis — Simpson (1900)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Nodularia (s.s.) (Group of N. douglasiæ) douglasiæ var. shanghaiensis — Simpson (1914)
Unio douglasiae amurensis — Haas (1930), Haas (1969)
Nodularia amurensis — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia (s.s.) amurensis — Zatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012)
Nodularia douglasiae — Graf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020), Sayenko et al. (2021)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae amurensis — Bogatov (2022)

Unio middendorffi Westerlund, 1890

Middendorff, 1851. Reise Nord. Ost. Sibiriens 2(1): 276-277, pl. 28, figs. 1-3 [as Unio pictorum var. longirostris].
Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 113.
NEOTYPE BIV229_5 (by Bolotov et al., 2020: Table S1), Gladkaya River, Primorye Region, Russia.

Unio douglasiae amurensis — Haas (1930), Haas (1969)
Unio pictorum middendorffi — Haas (1940)
Nodularia middendorffi — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia (s.s.) middendorffi — Zatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012)
Nodularia douglasiae — Graf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae middendorffi — Bogatov (2022)

Unio pliculosus Martens, 1894

Martens, 1894. S.-Ber. Ges. Naturf. Freunde zu Berlin 1894: 216.
Martens, 1905. Zool. Jb., Jena, Suppl. 8: 60, pl. 3, figs. 3-3b.
SYNTYPE SMF_3652, Okkwa Chhangphiöng, Korea.

Unio (Nodularia) pliculosus — Martens (1905)
Nodularia (s.s.) pliculosus — Haas (1911)
Nodularia (s.s.) (Group of N. douglasiæ) pliculosa — Simpson (1914)
Unio douglasiae pliculosus — Haas (1930), Haas (1969)
Unio douglasiae — Habe (1977)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae — Lopes-Lima et al. (2020), Nagel (2025)

Unio verrucifer Martens, 1894

Martens, 1894. S.-Ber. Ges. Naturf. Freunde zu Berlin 1894: 216.
Martens, 1905. Zool. Jb., Jena, Suppl. 8: 61, pl. 3, figs. 2-2b.
LECTOTYPE ZMB_MOLL_38.363 (by Nagel, 2025: 158), Hangang.

Unio (Nodularia) verrucifer — Martens (1905)
Nodularia (s.s.) verrucifera — Haas (1911)
Nodularia (s.s.) (Group of N. douglasiæ) verrucifera — Simpson (1914)
Unio douglasiae verrucifer — Haas (1930), Haas (1969)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae — Lopes-Lima et al. (2020), Nagel (2025)

Unio abbreviatus Westerlund, 1897

(homonym of Unio requienii var. abbreviatus de Joannis, 1859)
Westerlund, 1897. Ann. Mus. St. Pétersbourg 1897: 132.
PARATYPE SMF_3662, Ussuri fl. bei Noor, Amur gebiet.

Nodularia (s.s.) douglasiæ — Simpson (1900)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Nodularia (s.s.) douglasiae — Preston (1912)
Nodularia (s.s.) (Group of N. douglasiæ) douglasiæ — Simpson (1914)
Unio douglasiae amurensis — Haas (1930), Haas (1969)
Unio (Nodularia) douglasiae var. schrenki [sic] — Zhadin (1938)
Unio douglasiae var. abbreviatus — Zhadin (1965)
Nodularia abbreviata — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010)
Nodularia (s.s.) abbreviata — Zatravkin & Bogatov (1987), Starobogatov et al. (2004)
Nodularia douglasiae — Graf (2007), Lopes-Lima et al. (2020), Bolotov et al. (2020), Nagel (2025)
Nodularia (s.s.) schrencki — Bogatov (2012)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae schrencki — Bogatov (2022)

Unio schrencki Westerlund, 1897

Westerlund, 1897. Ann. Mus. St. Pétersbourg 1897: 131.
SYNTYPE SMF_3661, Nertscha-Fl. b-Nertschinsk, Amur-Gebiet.

Nodularia (s.s.) douglasiæ — Simpson (1900)
Nodularia (s.s.) (Group of N. douglasiæ) douglasiæ — Simpson (1914)
Unio douglasiae amurensis — Haas (1930), Haas (1969)
Unio (Nodularia) douglasiae var. schrenki [sic] — Zhadin (1938)
Unio douglasiae var. schrencki — Zhadin (1965)
Nodularia schrencki — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Bogatov & Prozorova (2017)
Nodularia (s.s.) schrencki — Zatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012)
Nodularia douglasiae — Graf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020), Sayenko et al. (2021), Nagel (2025)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae schrencki — Bogatov (2022)

Unio kobelti Neumayr, 1899

(homonym of Unio kobelti Kobelt & Rolle, 1895)
Neumayr, 1899. Wiss. Erg. Reise Szechény Ost-Asien 2: 645, pl. 3, fig. 1.

Nodularia douglasiae — Haas (1920)
Nodularia (s.s.) douglasiae — Haas (1924)
Unio douglasiae — Liu et al. (1979)
Nodularidia douglasiae — He & Zhuang (2013)

Unio boettgeri Neumayr, 1899

(homonym of Unio firmus var. boettgeri Ihering, 1893)
Neumayr, 1899. Wiss. Erg. Reise Szechény Ost-Asien 2: 645, pl. 2, fig. 3 [as ‘böttgeri’].

Nodularia (s.s.) (Group of N. douglasiæ) bœttgeri — Simpson (1914)
Nodularia douglasiae — Haas (1920)
Nodularia (s.s.) douglasiae — Haas (1924)
Nodularidia douglasiae — He & Zhuang (2013)

Nodularia douglasiae taiwanica Pilsbry & Hirase, 1905

(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Pilsbry & Hirase, 1905. Proc. Acad. Nat. Sci. 57: 750.
HOLOTYPE ANSP_58170, Formosa.

Nodularia (s.s.) (Group of N. douglasiæ) douglasiæ var. taiwanica — Simpson (1914)
Nodularia (s.s.) douglasiae taiwanica — Haas (1924)
Unio douglasiae taiwanicus — Haas (1930), Kuroda (1941), Haas (1969)
Unio douglasiae — Habe (1977), Liu et al. (1979), Wu (1982)
Nodularidia douglasiae — He & Zhuang (2013)

Unio jourdyi var. corrugata Dautzenberg & Fischer, 1905

(homonym of Unio corrugata Mauduyt, 1839; intro'd as intra-sp. of Unio jourdyi Morlet, 1886)
Dautzenberg & Fischer, 1905. J. de Conch. 53: 205.
SYNTYPE MNHN_MP_0149, Bas-Luc Nam.

Unio jourdyi var. corrugata — Dautzenberg & Fischer (1905)
Nodularia (s.s.) (Group of N. pazii) jourdyi var. corrugata — Simpson (1914)

Unio jourdyi var. ponderosa Dautzenberg & Fischer, 1905

(homonym of Mya ponderosa Dillwyn, 1817; intro'd as intra-sp. of Unio jourdyi Morlet, 1886)
Dautzenberg & Fischer, 1905. J. de Conch. 53: 453.
SYNTYPE MNHN_MP_0148, Tonkin, Grand Lac, Hanoi.

Unio jourdyi var. ponderosa — Dautzenberg & Fischer (1905)
Nodularia (s.s.) (Group of N. pazii) jourdyi var. ponderosa — Simpson (1914)

Nodularia douglasiae crassidens Haas, 1910

(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Haas, 1910. Ann. Mag. Nat. Hist. 6: 499.
Haas, 1910. Conch. Cab. 9 (pt. 2, h. 42): pl. 7, figs. 4-5.
Haas, 1911. Conch. Cab. 9 (pt. 2, h. 44): 75.
LECTOTYPE SMF_3659 (by Zilch, 1967: 51), Hunan, M. China.

Nodularia (s.s.) douglasiae crassidens — Haas (1911)
Nodularia (s.s.) (Group of N. douglasiæ) douglasiæ var. crassidens — Simpson (1914)
Unio douglasiae douglasiae — Haas (1930), Haas (1969)
Nodularia douglasiae crassidens — Dang et al. (1980), Do et al. (2018)
Nodularia crassidens — Prozorova et al. (2005)
Nodularia dorri — IUCN (2012)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae — Nagel (2025)

Nodularia douglasiae flavoviridis Haas, 1910

(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Haas, 1910. Conch. Cab. 9 (pt. 2, h. 41): 71, pl. 6, fig. 2.
LECTOTYPE SMF_3654 (by Zilch, 1967: 51), W. Ufer d. Sees Buirnor, O. Mongolei, Amur-Geb.

Unio douglasiae flavoviridis — Haas (1930)
Unio douglasiae amurensis — Haas (1969)
Nodularia flavoviridis — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010)
Nodularia (s.s.) flavoviridis — Zatravkin & Bogatov (1987), Starobogatov et al. (2004)
Nodularia douglasiae — Graf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020), Nagel (2025)
Nodularia (s.s.) schrencki — Bogatov (2012)
Nodularidia douglasiae — He & Zhuang (2013)

Nodularia denserugata Haas, 1910

Haas, 1910. Ann. Mag. Nat. Hist. 6: 496.
Haas, 1910. Conch. Cab. 9 (pt. 2, h. 42): pl. 8, figs. 3-4.
Haas, 1911. Conch. Cab. 9 (pt. 2, h. 44): 83, pl. 8, figs. 3-4.

Nodularia (s.s.) denserugata — Haas (1911)
Nodularia (s.s.) (Group of N. douglasiæ) denserugata — Simpson (1914)
Scabies denserugata — Haas (1930)
Unio denserugatus — Haas (1969)
Middendorffinaia (Sujfununio) denserugata — Moskvicheva & Starobogatov (1973)
Nodularia denserugata — Graf & Cummings (2007), Bogan (2010)
Ptychorhynchus denserugata — He & Zhuang (2013), Zieritz et al. (2018)
Ptychorhynchus denserugatum — Graf & Cummings (2021)
Nodularia douglasiae — Wu et al. (2024)

Nodularia verrucosa Haas, 1910

Haas, 1910. Nachr. Bl. Dtsch. Malak. Ges. 42: 99.
LECTOTYPE SMF_3660 (by Zilch, 1967: 51), Hunan, M. China.

Nodularia verrucosa — Haas (1910), Simpson (1914)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae — Nagel (2025)

Middendorffinaia (s.s.) arsenievi Moskvicheva & Starobogatov, 1973

(correct name for Middendorffinaia arsenjevi Moskvicheva & Starobogatov, 1973)
Moskvicheva & Starobogatov, 1973. Byull. Mosk. Obshch.. Ispyt. Prir. 78(2): 29, figs. 3м-п.

Middendorffinaia (s.s.) arsenievi — Moskvicheva & Starobogatov (1973), Zatravkin & Bogatov (1987), Starobogatov et al. (2004)
Middendorffinaia (s.s.) arsenjevi [sic] — Zatravkin (1983)
Middendorffinaia arsenievi — Kantor & Sysoev (2005), Kantor et al. (2010)
Unio crassus mongolicus — Graf (2007)
Middendorffinaia (s.s.) ussuriensis — Bogatov (2012), Vinarski & Kantor (2016)
Middendorffinaia mongolica — Klishko et al. (2019)
Nodularia douglasiae — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Middendorffinaia mongolica ussuriensis — Bogatov (2022)

Nodularia vladivostokensis Moskvicheva, 1973

Moskvicheva, 1973. Zool. Zhur. 52(10): 1463, figs. 1 п-р.

Nodularia vladivostokensis — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Klishko et al. (2018)
Nodularia (s.s.) vladivostokensis — Zatravkin & Bogatov (1987), Starobogatov et al. (2004)
Nodularia douglasiae — Graf (2007), Lopes-Lima et al. (2020), Bolotov et al. (2020), Sayenko et al. (2021)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae douglasiae — Bogatov (2022)

Nodularia sujfunica Moskvicheva, 1973

Moskvicheva, 1973. Zool. Zhur. 52(10): 1463, figs. 1 с-т.

Nodularia sujfunica — Moskvicheva (1973), Zatravkin (1983)
Nodularia (s.s.) sujfunica — Zatravkin & Bogatov (1987)
Nodularia (s.s.) vladivostokensis — Starobogatov et al. (2004)
Nodularia vladivostokensis — Kantor & Sysoev (2005)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia suifunica [sic] — Klishko et al. (2018)
Nodularia douglasiae — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularia douglasiae schrencki — Bogatov (2022)

Middendorffinaia arsenjevi Moskvicheva & Starobogatov, 1973

(incorrect original spelling of Middendorffinaia (s.s.) arsenievi Moskvicheva & Starobogatov, 1973)
Moskvicheva & Starobogatov, 1973. Byull. Mosk. Obshch.. Ispyt. Prir. 78(2): 22, fig. 1.
‘Moskv. et Star.’ Zatravkin, 1983. [in] I.M. Likharev, Molluscs: 41.

Nodularia lebedevi Zatravkin & Starobogatov, 1984

Zatravkin & Starobogatov, 1984. Zool. Zhur. 63(12): 1787, figs. 5-8.

Nodularia sp.n. — Zatravkin (1983)
Nodularia (Amurunio) lebedevi — Zatravkin & Bogatov (1987), Korniushin (1998), Starobogatov et al. (2004), Klishko et al. (2018)
Nodularia lebedevi — Kantor & Sysoev (2005), Kantor et al. (2010)
Nodularia douglasiae — Graf (2007), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularia (s.s.) middendorffi — Bogatov (2012)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia middendorffi — Vinarski & Kantor (2016)
Nodularia douglasiae middendorffi — Bogatov (2022)

Scabies longata Liu et al., 1991

Liu et al., 1991. Acta Zootax. Sinica 16(3): 279 [281], fig’d.

Scabies longata — Hu (2005), Graf & Cummings (2007), Bogan (2010), WoRMS (2017)
Scabies crispata — He & Zhuang (2013), Graf & Cummings (2021)
Nodularia douglasiae — Dai et al. (2024)

Nodularia (Magadaninaia) extremalis Martynov & Chernyshev, 1992

Martynov & Chernyshev, 1992. Zool. Zhur. 71: 19, figs. 1-3.

Nodularia (Magadaninaia) extremalis — Martynov & Chernyshev (1992), Starobogatov et al. (2004)
Nodularia extremalis — Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia douglasiae — Graf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae schrencki — Bogatov (2022)

Nodularia (s.s.) moskvichevae Bogatov & Starobogatov, 1992

Bogatov & Starobogatov, 1992. Zool. Zhur. 71: 134, fig’d.

Nodularia (s.s.) moskvichevae — Starobogatov et al. (2004)
Nodularia moskvichevae — Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia douglasiae — Graf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularia douglasiae middendorffi — Bogatov (2022)

Middendorffinaia (Middendorffinaia) ochotica Bogatov, 2000

Bogatov, 2000. Zool. Zhur. 79: 861, text fig. a-r.

Middendorffinaia (s.s.) ochotica — Starobogatov et al. (2004), Bogatov (2012), Vinarski & Kantor (2016), Bogatov & Prozorova (2017)
Middendorffinaia ochotica — Kantor & Sysoev (2005), Kantor et al. (2010)
Unio crassus mongolicus — Graf (2007)
Middendorffinaia mongolica — Klishko et al. (2019)
Nodularia douglasiae — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Middendorffinaia mongolica ochotica — Bogatov (2022)

Nodularia sakhalinensis Bogatov, 2001

Bogatov, 2001. Byull. Dal'nev. Malak.Obshch. 5: 75.

Nodularia (s.s.) sakhalinensis — Starobogatov et al. (2004)
Nodularia sakhalinensis — Kantor & Sysoev (2005), Kantor et al. (2010), Klishko et al. (2018)
Nodularia douglasiae — Graf (2007), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae schrencki — Bogatov (2022)

Acuticosta jianghanensis He & Zhuang, 2013

He & Zhuang, 2013. Freshwater Bivalves China: 23, fig. 47-48.

Acuticosta jianghanensis — He & Zhuang (2013), Zieritz et al. (2018), Liu et al. (2020)
Nodularia douglasiae — Bogatov & Prozorova (2017)

Nodularia douglasiae nipponensis ‘Martens’ of Authors

(misidentified reference to Unio nipponensis Martens, 1877)
‘Martens’ of Authors.
Sano et al., 2017. J. Wat. Res. Prot. 9: Table 2. [in part]

Nodularia douglasiae — Sano et al. (2020), Choi et al. (2020)

Unio föttgeri of Authors

of Authors.

Unio douglasiae — Liu et al. (1979)

Unio ezonis ‘Kuroda, MS’ in ANSP coll. of Authors

‘Kuroda, MS’ in ANSP coll. of Authors.

Nodularia suifunica ‘Moskvicheva’ of Authors

(error for Nodularia sujfunica Moskvicheva, 1973)
‘Moskvicheva’ of Authors.
Klishko et al., 2018. Systemat. Biodiv. 16(3): 288.
Bogatov, 2022. Lg. Bivalve Moll. Russia’s Fresh Waters: 59.

Margarita (Unio) merchisonianus Lea, 1836

(error for Unio murchisonianus Lea, 1834)
Lea, 1836. Synopsis Fam. Naiades: 14 [corrected in the errata].

Unio pictorum var. longirostris Middendorff, 1851

(misidentified reference to Unio longirostris Rossmässler, 1836)
Middendorff, 1851. Reise Nord. Ost. Sibiriens 2(1): 276-277, pl. 28, figs. 1-3.
Schrenck, 1867. Reise Forsch. Amurlande 2: 696.

Nodularia middendorffi — Moskvicheva (1973), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia (s.s.) middendorffi — Zatravkin & Bogatov (1987), Bogatov (2012)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae — Lopes-Lima et al. (2020)

Unio murchinsonianus Küster, 1862

(error for Unio murchisonianus Lea, 1834)
‘Lea’ Küster, 1862. Conchyl. Cab. 9(2): 309.

Unio pictorum Schrenck, 1867

(misidentified reference to Mya pictorum Linnaeus, 1758)
‘L.’ Schrenck, 1867. Reise Forsch. Amurlande 2: 696-699.

Nodularia middendorffi — Moskvicheva (1973)

Unio pictorum var. longirostris Westerlund, 1877

(misidentified reference to Unio longirostris Rossmässler, 1836)
Westerlund, 1877. Kongl. Svenska Vet. Akad. Handl. 14(12): 74.

Nodularia (s.s.) douglasiæ — Simpson (1900)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Nodularia (s.s.) douglasiae — Preston (1912)
Nodularia (s.s.) (Group of N. douglasiæ) douglasiæ — Simpson (1914)
Unio douglasiae amurensis — Haas (1930), Haas (1969)
Nodularia amurensis — Vinarski & Kantor (2016)

Unio nipponensis Kobelt, 1879

(misidentified reference to Unio nipponensis Martens, 1877)
Kobelt, 1879. Abh. Senckenb. Naturf. Ges. 11: 422, pl. 12, fig. 3.

Nodularia (s.s.) douglasiæ var. nipponensis — Simpson (1900)
Unio (Nodularia) nipponensis — Martens (1905)

Unio douglasiae var. osbeckii Heude, 1885

(reference to Unio osbeckii Philippi, 1844)
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65, fig. 128i.

Unio (Nodularia) douglasiae var. sinuolatus — Martens (1905)

Unio schanghaiensis Neumayr, 1899

(error for Unio shanghaiensis Lea, 1859)
‘Lea’ Neumayr, 1899. Wiss. Erg. Reise Szechény Ost-Asien 2: 645.

Unio wrigtii de Gregorio, 1914

(error for Unio wrightii Lea, 1865)
‘Lea’ de Gregorio, 1914. Il Nat. Siciliano 2: 40.

Unio (Nodularia) douglasiae var. schrenki Zhadin, 1938

(error for Unio schrencki Westerlund, 1897)
‘Westerlund’ Zhadin, 1938. Faune USSR Mollusques 4(1): 99.

Unio taiwanensis Habe, 1977

(error for Nodularia douglasiae taiwanica Pilsbry & Hirase, 1905)
‘Pilsbry’ Habe, 1977. Mollusc. Japan: 114.

Unio firnus Liu et al., 1979

(error for Unio douglasiae firmus Heude, 1885)
Liu et al., 1979. Econ. Fauna China: 74.

Unio douglasiae chlorens Liu et al., 1979

(error for Unio douglasiae chloreus Heude, 1885)
Liu et al., 1979. Econ. Fauna China: 74.

Unio douglasiae cleltae Liu et al., 1979

(error for Unio douglasiae deltae Heude, 1885)
Liu et al., 1979. Econ. Fauna China: 74.

Nodularia doctylina Liu et al., 1980

(error for Unio douglasiae dactylinus Heude, 1885)
Liu et al., 1980. Acta Zool. Sinica 26(4): 366.

Nodularia sachalinensis Bogatov, 2001

(error for Nodularia sakhalinensis Bogatov, 2001)
Bogatov, 2001. Byull. Dal'nev. Malak.Obshch. 5: 59.

Unio douglassiae Kantor & Sysoev, 2005

(error for Unio douglasiae Griffith & Pidgeon, 1833)
‘Griffith et Pidgeon’ Kantor & Sysoev, 2005. Cat. Moll. Russia: 337.

Nodularia jourdyi Bolotov et al., 2018

(reference to Unio jourdyi Morlet, 1886)
Bolotov et al., 2018. Sci. Repts. 8: 4.
Pfeiffer et al., 2019. MPE 137: 117.

Nodularia douglasiae — Lopes-Lima et al. (2020), Choi et al. (2020), Sayenko et al. (2021), Wu et al. (2024)

Unio osbreckii Lopes-Lima et al., 2020

(error for Unio osbeckii Philippi, 1844)
‘Philippi’ Lopes-Lima et al., 2020. MPE 146: Appendix 1.