Bivalvia | ... | Unionoidea | Unionidae | Unioninae | Unionini | Oxynaiina | Nodularia
273 specimens + 234 Genbank vouchers
Griffith & Pidgeon, 1833. [in] Cuvier, Animal Kingdom 12(38): pl. 21, fig. 2 [as ‘douglasiæ’].
‘Gray’ Griffith & Pidgeon, 1834. [in] Cuvier, Animal Kingdom 12(39-40): 601 [as ‘douglasiæ’].
Nodularia (s.s.) douglasiae — Simpson (1900), Preston (1912), Simpson (1914), Haas (1924), Korniushin (1998)
Unio (Nodularia) douglasiae — Martens (1905), Zhadin (1938)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Nodularia douglasiae — Haas (1920), Prozorova et al. (2005), Graf (2007), Graf & Cummings (2007), Bogan (2010), ... Chen et al. (2025), Hou et al. (2025), Wang et al. (2025), Chen et al. (2025), Chen et al. (2025)
Unio douglasiae — Zhadin (1965), Kil (1976), Habe (1977), Liu et al. (1979), Liu et al. (1980), ... Zhou et al. (2017), Araujo et al. (2018), Wang et al. (2018), Aunins et al. (2018), Wang et al. (2018)
Unio douglasiae douglasiae — Haas (1969)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia cf. douglasiae — Huang et al. (2019)
Nodularia douglasiae Clade C — Choi et al. (2020)
Nodularia douglasiae Clade B — Choi et al. (2020)
Nodularia douglasiae Clade A — Choi et al. (2020)
Nodularia douglasiae douglasiae — Bogatov (2022)
Nodularia sp. 2 — Dai et al. (2024)

Lea, 1834. Trans. Amer. Phil. Soc. 5 [O. 1]: 33, pl. 3, fig. 6.
Type(s): HOLOTYPE (MONOTYPY) USNM_84156, China [“Calcutta, India” penciled out].
Nodularia (s.s.) douglasiae — Simpson (1900), Preston (1912), Simpson (1914)
Unio (Nodularia) douglasiae — Martens (1905)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Unio douglasiae douglasiae — Haas (1969)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae — Lopes-Lima et al. (2020)
Philippi, 1844. Zeitschr. f. Malakozool. 1: 164.
Philippi, 1847. Abbild. Beschr. Conch. 3: 45, pl. 3, fig. 1.
Nodularia (s.s.) douglasiae — Simpson (1900), Preston (1912), Simpson (1914)
Unio (Nodularia) douglasiae — Martens (1905)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Unio douglasiae douglasiae — Haas (1969)
Unio douglasiae — Liu et al. (1979)
Nodularia douglasiae — Prozorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)

Lea, 1859. Proc. Acad. Nat. Sci. 11: 153.
Lea, 1860. Jour. Acad. Nat. Sci. 4 [O. 7]: 242, pl. 36, fig. 121.
Type(s): HOLOTYPE BMNH_1965171, Shanghai, China.
Nodularia (s.s.) douglasiae var. shanghaiensis — Simpson (1900), Preston (1912), Simpson (1914)
Unio (Nodularia) douglasiae — Martens (1905)
Nodularia (s.s.) douglasiae shanghaiensis — Haas (1911)
Nodularia douglasiae — Haas (1920), Lopes-Lima et al. (2020)
Unio douglasiae douglasiae — Haas (1969)
Unio douglasiae — Liu et al. (1979)
Nodularidia douglasiae — He & Zhuang (2013)

Lea, 1865. Proc. Acad. Nat. Sci. 17: 75.
Lea, 1868. Jour. Acad. Nat. Sci. 6 [O. 12]: 283, pl. 39, fig. 97.
Type(s): HOLOTYPE USNM_84129, China.
Nodularia (s.s.) douglasiae — Simpson (1900), Simpson (1914)
Unio (Nodularia) douglasiae — Martens (1905)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Unio wrigtii [sic] — de Gregorio (1914)
Unio douglasiae douglasiae — Haas (1969)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae — Lopes-Lima et al. (2020)

Deshayes, 1873. Nouvelles archives 9: 9, pl. 1, figs. 3-3a.
Type(s): SYNTYPES MNHN-IM-2000-1779, Petchily, Chine Centrale.
Nodularia (s.s.) douglasiae — Simpson (1900), Preston (1912), Simpson (1914)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Unio douglasiae douglasiae — Haas (1969)
Unio douglasiae — Liu et al. (1979)
Unio sculptus — Yu et al. (2001)
Nodularia douglasiae — Prozorova et al. (2005), Schneider et al. (2013), Lopes-Lima et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)

Kobelt, 1879. Abh. Senckenb. Naturf. Ges. 11: 425, pl. 23, figs. 2-4.
Type(s): LECTOTYPE SMF_3650, Biwa-See, Japan.
Nodularia (s.s.) biwae — Simpson (1900), Haas (1911), Simpson (1914)
Unio biwae — Haas (1969), Habe (1977), Habe (1991), Slugina (2006)
Unio douglasiae biwae — Kondo (1997), Kondo (2008)
Nodularia biwae — Bogan (2010)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae biwae — Sano et al. (2017), Mabuchi & Nishida (2020)
Nodularia douglasiae — Lopes-Lima et al. (2020), Mabuchi et al. (2021)
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65 bis, fig. 128c.
(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Nodularia (s.s.) douglasiae — Simpson (1900)
Unio douglasiae — Liu et al. (1979)
Nodularia douglasiae — Prozorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65, fig. 128e.
(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Nodularia (s.s.) douglasiae — Simpson (1900), Preston (1912), Simpson (1914)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Unio douglasiae douglasiae — Haas (1969)
Unio douglasiae — Liu et al. (1979)
Nodularia douglasiae — Prozorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65 bis, fig. 128d [as ‘deltæ’].
(homonym of Unio favidens var. deltae Benson, 1862; intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Unio douglasiae — Liu et al. (1979)
Nodularia douglasiae — Prozorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65 bis, figs. 128b-b’.
(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833; homonym of Unio firmus Lea, 1866)
Nodularia (s.s.) douglasiae — Simpson (1900)
Unio douglasiae — Liu et al. (1979)
Nodularia douglasiae — Prozorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65, fig. 128f.
(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Unio douglasiae — Liu et al. (1979)
Nodularia douglasiae — Prozorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)

Morlet, 1886. J. de Conch. 34: 76, pl. 13, figs. 5-5a.
Type(s): SYNTYPES MNHN_MP_0150, Bac Hat etang de la riviere Claire.
Nodularia (s.s.) jourdyi — Simpson (1900), Simpson (1914)
Unio jourdyi — Dautzenberg & Fischer (1905), Dautzenberg & Fischer (1905)
Oxynaia jourdyi — Haas (1913), Thiele (1934), Haas (1969), Haas (1969), Dang et al. (1980), Graf & Cummings (2007), Bogan (2010), IUCN (2011), Zieritz et al. (2018), Do et al. (2018)
Nodularia jourdyi — Bolotov et al. (2018), Pfeiffer et al. (2019), Sayenko et al. (2021), Graf & Cummings (2021)
Nodularia douglasiae — Wu et al. (2024)
Mousson, 1887. J. de Conch. 35: 26, pl. 1, fig. 8.
(intro'd as intra-sp. of Mya pictorum Linnaeus, 1758)
Nodularia (s.s.) douglasiae var. shanghaiensis — Simpson (1900), Simpson (1914)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Unio douglasiae amurensis — Haas (1969)
Nodularia amurensis — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia (s.s.) amurensis — Zatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012)
Nodularia douglasiae — Graf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020), Sayenko et al. (2021)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae amurensis — Bogatov (2022)
Middendorff, 1851. Reise Nord. Ost. Sibiriens 2(1): 276-277, pl. 28, figs. 1-3 [as Unio pictorum var. longirostris].
Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 113.
Type(s): NEOTYPE BIV229_5 (by Bolotov et al., 2020: Table S1), Gladkaya River, Primorye Region, Russia.
Unio pictorum middendorffi — Haas (1940)
Unio douglasiae amurensis — Haas (1969)
Nodularia middendorffi — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia (s.s.) middendorffi — Zatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012)
Nodularia douglasiae — Graf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae middendorffi — Bogatov (2022)

Martens, 1894. S.-Ber. Ges. Naturf. Freunde zu Berlin 1894: 216.
Martens, 1905. Zool. Jb., Jena, Suppl. 8: 60, pl. 3, figs. 3-3b.
Type(s): TYPE ZMB_55620, Singei, Korea.
Unio (Nodularia) pliculosus — Martens (1905)
Nodularia (s.s.) pliculosus — Haas (1911)
Nodularia (s.s.) pliculosa — Simpson (1914)
Unio douglasiae pliculosus — Haas (1969)
Unio douglasiae — Habe (1977)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae — Lopes-Lima et al. (2020)

Martens, 1894. S.-Ber. Ges. Naturf. Freunde zu Berlin 1894: 216.
Martens, 1905. Zool. Jb., Jena, Suppl. 8: 61, pl. 3, figs. 2-2b.
Type(s): TYPE ZMB_55622, Hangang.
Unio (Nodularia) verrucifer — Martens (1905)
Nodularia (s.s.) verrucifera — Haas (1911), Simpson (1914)
Unio douglasiae verrucifer — Haas (1969)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae — Lopes-Lima et al. (2020)

Westerlund, 1897. Ann. Mus. St. Pétersbourg 1897: 132.
(homonym of Unio requienii var. abbreviatus de Joannis, 1859)
Type(s): PARATYPE SMF_3662, Ussuri fl. bei Noor, Amur gebiet.
Nodularia (s.s.) douglasiae — Simpson (1900), Preston (1912), Simpson (1914)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Unio (Nodularia) douglasiae var. schrenki [sic] — Zhadin (1938)
Unio douglasiae var. abbreviatus — Zhadin (1965)
Unio douglasiae amurensis — Haas (1969)
Nodularia abbreviata — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010)
Nodularia (s.s.) abbreviata — Zatravkin & Bogatov (1987), Starobogatov et al. (2004)
Nodularia douglasiae — Graf (2007), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularia (s.s.) schrencki — Bogatov (2012)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae schrencki — Bogatov (2022)

Westerlund, 1897. Ann. Mus. St. Pétersbourg 1897: 131.
Type(s): PARATYPE SMF_3663, Amur - Fluss.
Nodularia (s.s.) douglasiae — Simpson (1900), Simpson (1914)
Unio (Nodularia) douglasiae var. schrenki [sic] — Zhadin (1938)
Unio douglasiae var. schrencki — Zhadin (1965)
Unio douglasiae amurensis — Haas (1969)
Nodularia schrencki — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Bogatov & Prozorova (2017)
Nodularia (s.s.) schrencki — Zatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012)
Nodularia douglasiae — Graf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020), Sayenko et al. (2021)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae schrencki — Bogatov (2022)
Neumayr, 1899. Wiss. Erg. Reise Szechény Ost-Asien 2: 645, pl. 2, fig. 3 [as ‘böttgeri’].
(homonym of Unio firmus var. boettgeri Ihering, 1893)
Nodularia (s.s.) boettgeri — Simpson (1914)
Nodularia douglasiae — Haas (1920)
Nodularia (s.s.) douglasiae — Haas (1924)
Nodularidia douglasiae — He & Zhuang (2013)
Neumayr, 1899. Wiss. Erg. Reise Szechény Ost-Asien 2: 645, pl. 3, fig. 1.
(homonym of Unio kobelti Kobelt & Rolle, 1895)
Unio kobelti — Simpson (1914)
Nodularia douglasiae — Haas (1920)
Nodularia (s.s.) douglasiae — Haas (1924)
Unio douglasiae — Liu et al. (1979)
Nodularidia douglasiae — He & Zhuang (2013)

Pilsbry & Hirase, 1905. Proc. Acad. Nat. Sci. 57: 750.
(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Type(s): HOLOTYPE ANSP_58170, Formosa.
Nodularia (s.s.) douglasiae var. taiwanica — Simpson (1914)
Nodularia (s.s.) douglasiae taiwanica — Haas (1924)
Unio douglasiae taiwanicus — Kuroda (1941)
Unio douglasiae taiwanica — Haas (1969)
Unio douglasiae — Habe (1977), Liu et al. (1979), Wu (1982)
Nodularidia douglasiae — He & Zhuang (2013)

Dautzenberg & Fischer, 1905. J. de Conch. 53: 205.
(homonym of Unio corrugata Mauduyt, 1839; intro'd as intra-sp. of Unio jourdyi Morlet, 1886)
Type(s): SYNTYPE MNHN_MP_0149, Bas-Luc Nam.
Unio jourdyi var. corrugata — Dautzenberg & Fischer (1905)
Nodularia (s.s.) jourdyi var. corrugata — Simpson (1914)

Dautzenberg & Fischer, 1905. J. de Conch. 53: 453.
(homonym of Mya ponderosa Dillwyn, 1817; intro'd as intra-sp. of Unio jourdyi Morlet, 1886)
Type(s): SYNTYPE MNHN_MP_0148, Tonkin, Grand Lac, Hanoi.
Unio jourdyi var. ponderosa — Dautzenberg & Fischer (1905)
Nodularia (s.s.) jourdyi var. ponderosa — Simpson (1914)
Haas, 1910. Ann. Mag. Nat. Hist. 6: 496.
Haas, 1910. Conch. Cab. 9 (pt. 2, h. 42): pl. 8, figs. 3-4.
Haas, 1911. Conch. Cab. 9 (pt. 2, h. 44): 83, pl. 8, figs. 3-4.
Nodularia (s.s.) denserugata — Haas (1911), Simpson (1914)
Unio denserugatus — Haas (1969)
Middendorffinaia (Sujfununio) denserugata — Moskvicheva & Starobogatov (1973)
Nodularia denserugata — Graf & Cummings (2007), Bogan (2010)
Ptychorhynchus denserugata — He & Zhuang (2013), Zieritz et al. (2018)
Ptychorhynchus denserugatum — Graf & Cummings (2021)
Nodularia douglasiae — Wu et al. (2024)

Haas, 1910. Nachr. Bl. Dtsch. Malak. Ges. 42: 99.
Type(s): TYPE SMF_3660, Hunan, M. China.
Nodularia verrucosa — Haas (1910), Simpson (1914)
Nodularidia douglasiae — He & Zhuang (2013)

Haas, 1910. Ann. Mag. Nat. Hist. 6: 499.
Haas, 1910. Conch. Cab. 9 (pt. 2, h. 42): pl. 7, figs. 4-5.
Haas, 1911. Conch. Cab. 9 (pt. 2, h. 44): 75.
(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Type(s): TYPE SMF_3659, Hunan, M. China.
Nodularia (s.s.) douglasiae crassidens — Haas (1911)
Nodularia (s.s.) douglasiae var. crassidens — Simpson (1914)
Unio douglasiae douglasiae — Haas (1969)
Nodularia douglasiae crassidens — Dang et al. (1980), Do et al. (2018)
Nodularia crassidens — Prozorova et al. (2005)
Nodularia dorri — IUCN (2012)
Nodularidia douglasiae — He & Zhuang (2013)

Haas, 1910. Conch. Cab. 9 (pt. 2, h. 41): 71, pl. 6, fig. 2.
(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Type(s): TYPE SMF_3654, W. Ufer d. Sees Buirnor, O. Mongolei, Amur-Geb.
Unio douglasiae amurensis — Haas (1969)
Nodularia flavoviridis — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010)
Nodularia (s.s.) flavoviridis — Zatravkin & Bogatov (1987), Starobogatov et al. (2004)
Nodularia douglasiae — Graf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularia (s.s.) schrencki — Bogatov (2012)
Nodularidia douglasiae — He & Zhuang (2013)
Moskvicheva & Starobogatov, 1973. Byull. Mosk. Obshch.. Ispyt. Prir. 78(2): 22, fig. 1.
‘Moskv. et Star.’ Zatravkin, 1983. [in] I.M. Likharev, Molluscs: 41.
(incorrect original spelling of Middendorffinaia (s.s.) arsenievi Moskvicheva & Starobogatov, 1973)
Moskvicheva & Starobogatov, 1973. Byull. Mosk. Obshch.. Ispyt. Prir. 78(2): 29, figs. 3м-п.
(correct name for Middendorffinaia arsenjevi Moskvicheva & Starobogatov, 1973)
Middendorffinaia (s.s.) arsenievi — Moskvicheva & Starobogatov (1973), Zatravkin & Bogatov (1987), Starobogatov et al. (2004)
Middendorffinaia (s.s.) arsenjevi [sic] — Zatravkin (1983)
Middendorffinaia arsenievi — Kantor & Sysoev (2005), Kantor et al. (2010)
Unio crassus mongolicus — Graf (2007)
Middendorffinaia (s.s.) ussuriensis — Bogatov (2012), Vinarski & Kantor (2016)
Middendorffinaia mongolica — Klishko et al. (2019)
Nodularia douglasiae — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Middendorffinaia mongolica ussuriensis — Bogatov (2022)
Moskvicheva, 1973. Zool. Zhur. 52(10): 1463, figs. 1 с-т.
Nodularia sujfunica — Moskvicheva (1973), Zatravkin (1983)
Nodularia (s.s.) sujfunica — Zatravkin & Bogatov (1987)
Nodularia (s.s.) vladivostokensis — Starobogatov et al. (2004)
Nodularia vladivostokensis — Kantor & Sysoev (2005)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia suifunica [sic] — Klishko et al. (2018)
Nodularia douglasiae — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularia douglasiae schrencki — Bogatov (2022)
Moskvicheva, 1973. Zool. Zhur. 52(10): 1463, figs. 1 п-р.
Nodularia vladivostokensis — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Klishko et al. (2018)
Nodularia (s.s.) vladivostokensis — Zatravkin & Bogatov (1987), Starobogatov et al. (2004)
Nodularia douglasiae — Graf (2007), Lopes-Lima et al. (2020), Bolotov et al. (2020), Sayenko et al. (2021)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae douglasiae — Bogatov (2022)
Zatravkin & Starobogatov, 1984. Zool. Zhur. 63(12): 1787, figs. 5-8.
Nodularia sp.n. — Zatravkin (1983)
Nodularia (Amurunio) lebedevi — Zatravkin & Bogatov (1987), Korniushin (1998), Starobogatov et al. (2004), Klishko et al. (2018)
Nodularia lebedevi — Kantor & Sysoev (2005), Kantor et al. (2010)
Nodularia douglasiae — Graf (2007), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularia (s.s.) middendorffi — Bogatov (2012)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia middendorffi — Vinarski & Kantor (2016)
Nodularia douglasiae middendorffi — Bogatov (2022)
Liu et al., 1991. Acta Zootax. Sinica 16(3): 279 [281], fig’d.
Scabies longata — Hu (2005), Graf & Cummings (2007), Bogan (2010), WoRMS (2017)
Scabies crispata — He & Zhuang (2013), Graf & Cummings (2021)
Nodularia douglasiae — Dai et al. (2024)
Martynov & Chernyshev, 1992. Zool. Zhur. 71: 19, figs. 1-3.
Nodularia (Magadaninaia) extremalis — Martynov & Chernyshev (1992), Starobogatov et al. (2004)
Nodularia extremalis — Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia douglasiae — Graf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae schrencki — Bogatov (2022)
Bogatov & Starobogatov, 1992. Zool. Zhur. 71: 134, fig’d.
Nodularia (s.s.) moskvichevae — Starobogatov et al. (2004)
Nodularia moskvichevae — Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia douglasiae — Graf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularia douglasiae middendorffi — Bogatov (2022)
Bogatov, 2000. Zool. Zhur. 79: 861, text fig. a-r.
Middendorffinaia (s.s.) ochotica — Starobogatov et al. (2004), Bogatov (2012), Vinarski & Kantor (2016), Bogatov & Prozorova (2017)
Middendorffinaia ochotica — Kantor & Sysoev (2005), Kantor et al. (2010)
Unio crassus mongolicus — Graf (2007)
Middendorffinaia mongolica — Klishko et al. (2019)
Nodularia douglasiae — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Middendorffinaia mongolica ochotica — Bogatov (2022)
Bogatov, 2001. Byull. Dal'nev. Malak.Obshch. 5: 75.
Nodularia (s.s.) sakhalinensis — Starobogatov et al. (2004)
Nodularia sakhalinensis — Kantor & Sysoev (2005), Kantor et al. (2010), Klishko et al. (2018)
Nodularia douglasiae — Graf (2007), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae schrencki — Bogatov (2022)
He & Zhuang, 2013. Freshwater Bivalves China: 23, fig. 47-48.
Acuticosta jianghanensis — He & Zhuang (2013), Zieritz et al. (2018), Liu et al. (2020)
Nodularia douglasiae — Bogatov & Prozorova (2017)
‘Martens’ of Authors.
Sano et al., 2017. J. Wat. Res. Prot. 9: Table 2. [in part]
(misidentified reference to Unio nipponensis Martens, 1877)
Nodularia douglasiae — Sano et al. (2020), Choi et al. (2020)
‘Moskvicheva’ of Authors.
Klishko et al., 2018. Systemat. Biodiv. 16(3): 288.
Bogatov, 2022. Lg. Bivalve Moll. Russia’s Fresh Waters: 59.
(error for Nodularia sujfunica Moskvicheva, 1973)
‘Kuroda, MS’ in ANSP coll. of Authors.
Lea, 1836. Synopsis Fam. Naiades: 14 [corrected in the errata].
(error for Unio murchisonianus Lea, 1834)
Middendorff, 1851. Reise Nord. Ost. Sibiriens 2(1): 276-277, pl. 28, figs. 1-3.
Schrenck, 1867. Reise Forsch. Amurlande 2: 696.
(misidentified reference to Unio longirostris Rossmässler, 1836)
Nodularia middendorffi — Moskvicheva (1973), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia (s.s.) middendorffi — Zatravkin & Bogatov (1987), Bogatov (2012)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae — Lopes-Lima et al. (2020)
‘Lea’ Küster, 1862. Conchyl. Cab. 9(2): 309.
(error for Unio murchisonianus Lea, 1834)
‘L.’ Schrenck, 1867. Reise Forsch. Amurlande 2: 696-699.
(misidentified reference to Mya pictorum Linnaeus, 1758)
Nodularia middendorffi — Moskvicheva (1973)
Westerlund, 1877. Kongl. Svenska Vet. Akad. Handl. 14(12): 74.
(misidentified reference to Unio longirostris Rossmässler, 1836)
Nodularia (s.s.) douglasiae — Simpson (1900), Preston (1912), Simpson (1914)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Unio douglasiae amurensis — Haas (1969)
Nodularia amurensis — Vinarski & Kantor (2016)
Kobelt, 1879. Abh. Senckenb. Naturf. Ges. 11: 422, pl. 12, fig. 3.
(misidentified reference to Unio nipponensis Martens, 1877)
Nodularia (s.s.) douglasiae var. nipponensis — Simpson (1900)
Unio (Nodularia) nipponensis — Martens (1905)
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65, fig. 128i.
(reference to Unio osbeckii Philippi, 1844)
Unio (Nodularia) douglasiae var. sinuolatus — Martens (1905)
‘Lea’ Neumayr, 1899. Wiss. Erg. Reise Szechény Ost-Asien 2: 645.
(error for Unio shanghaiensis Lea, 1859)
‘Lea’ de Gregorio, 1914. Il Nat. Siciliano 2: 40.
(error for Unio wrightii Lea, 1865)
‘Westerlund’ Zhadin, 1938. Faune USSR Mollusques 4(1): 99.
(error for Unio schrencki Westerlund, 1897)
‘Pilsbry’ Habe, 1977. Mollusc. Japan: 114.
(error for Nodularia douglasiae taiwanica Pilsbry & Hirase, 1905)
Liu et al., 1979. Econ. Fauna China: 74.
(error for Unio douglasiae firmus Heude, 1885)
Liu et al., 1979. Econ. Fauna China: 74.
(error for Unio douglasiae chloreus Heude, 1885)
Liu et al., 1979. Econ. Fauna China: 74.
(error for Unio douglasiae deltae Heude, 1885)
Liu et al., 1980. Acta Zool. Sinica 26(4): 366.
(error for Unio douglasiae dactylinus Heude, 1885)
Bogatov, 2001. Byull. Dal'nev. Malak.Obshch. 5: 59.
(error for Nodularia sakhalinensis Bogatov, 2001)
‘Griffith et Pidgeon’ Kantor & Sysoev, 2005. Cat. Moll. Russia: 337.
(error for Unio douglasiae Griffith & Pidgeon, 1833)
Bolotov et al., 2018. Sci. Repts. 8: 4.
Pfeiffer et al., 2019. MPE 137: 117.
(reference to Unio jourdyi Morlet, 1886)
Nodularia douglasiae — Lopes-Lima et al. (2020), Choi et al. (2020), Sayenko et al. (2021), Wu et al. (2024)
‘Philippi’ Lopes-Lima et al., 2020. MPE 146: Appendix 1.
(error for Unio osbeckii Philippi, 1844)
Nodularia (s.s.) douglasiae — Simpson (1900), Preston (1912), Simpson (1914), Haas (1924), Korniushin (1998)
Nodularia (s.s.) douglasiae var. shanghaiensis — Simpson (1900), Preston (1912), Simpson (1914)
Nodularia (s.s.) biwae — Simpson (1900), Haas (1911), Simpson (1914)
Nodularia (s.s.) jourdyi — Simpson (1900), Simpson (1914)
Nodularia (s.s.) douglasiae var. nipponensis — Simpson (1900)
Unio (Nodularia) douglasiae — Martens (1905), Zhadin (1938)
Unio (Nodularia) douglasiae var. sinuolatus — Martens (1905)
Unio (Nodularia) nipponensis — Martens (1905)
Unio (Nodularia) pliculosus — Martens (1905)
Unio (Nodularia) verrucifer — Martens (1905)
Unio jourdyi — Dautzenberg & Fischer (1905), Dautzenberg & Fischer (1905)
Unio jourdyi var. corrugata — Dautzenberg & Fischer (1905)
Unio jourdyi var. ponderosa — Dautzenberg & Fischer (1905)
Nodularia verrucosa — Haas (1910), Simpson (1914)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Nodularia (s.s.) douglasiae shanghaiensis — Haas (1911)
Nodularia (s.s.) douglasiae crassidens — Haas (1911)
Nodularia (s.s.) pliculosus — Haas (1911)
Nodularia (s.s.) verrucifera — Haas (1911), Simpson (1914)
Nodularia (s.s.) denserugata — Haas (1911), Simpson (1914)
Oxynaia jourdyi — Haas (1913), Thiele (1934), Haas (1969), Haas (1969), Dang et al. (1980), Graf & Cummings (2007), Bogan (2010), IUCN (2011), Zieritz et al. (2018), Do et al. (2018)
Unio kobelti — Simpson (1914)
Nodularia (s.s.) douglasiae var. taiwanica — Simpson (1914)
Nodularia (s.s.) douglasiae var. crassidens — Simpson (1914)
Nodularia (s.s.) pliculosa — Simpson (1914)
Nodularia (s.s.) boettgeri — Simpson (1914)
Nodularia (s.s.) jourdyi var. corrugata — Simpson (1914)
Nodularia (s.s.) jourdyi var. ponderosa — Simpson (1914)
Unio wrigtii [sic] — de Gregorio (1914)
Nodularia douglasiae — Haas (1920), Prozorova et al. (2005), Graf (2007), Graf & Cummings (2007), Bogan (2010), ... Chen et al. (2025), Hou et al. (2025), Wang et al. (2025), Chen et al. (2025), Chen et al. (2025)
Nodularia (s.s.) douglasiae taiwanica — Haas (1924)
Unio (Nodularia) douglasiae var. schrenki [sic] — Zhadin (1938)
Unio pictorum middendorffi — Haas (1940)
Unio douglasiae taiwanicus — Kuroda (1941)
Unio douglasiae — Zhadin (1965), Kil (1976), Habe (1977), Liu et al. (1979), Liu et al. (1980), ... Zhou et al. (2017), Araujo et al. (2018), Wang et al. (2018), Aunins et al. (2018), Wang et al. (2018)
Unio douglasiae var. schrencki — Zhadin (1965)
Unio douglasiae var. abbreviatus — Zhadin (1965)
Unio douglasiae douglasiae — Haas (1969)
Unio douglasiae amurensis — Haas (1969)
Unio douglasiae pliculosus — Haas (1969)
Unio douglasiae taiwanica — Haas (1969)
Unio douglasiae verrucifer — Haas (1969)
Unio denserugatus — Haas (1969)
Unio biwae — Haas (1969), Habe (1977), Habe (1991), Slugina (2006)
Nodularia amurensis — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia middendorffi — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia abbreviata — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010)
Nodularia schrencki — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Bogatov & Prozorova (2017)
Nodularia flavoviridis — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010)
Nodularia vladivostokensis — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Klishko et al. (2018)
Nodularia sujfunica — Moskvicheva (1973), Zatravkin (1983)
Middendorffinaia (s.s.) arsenievi — Moskvicheva & Starobogatov (1973), Zatravkin & Bogatov (1987), Starobogatov et al. (2004)
Middendorffinaia (Sujfununio) denserugata — Moskvicheva & Starobogatov (1973)
Nodularia douglasiae crassidens — Dang et al. (1980), Do et al. (2018)
Middendorffinaia (s.s.) arsenjevi [sic] — Zatravkin (1983)
Nodularia sp.n. — Zatravkin (1983)
Nodularia (s.s.) middendorffi — Zatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012)
Nodularia (s.s.) abbreviata — Zatravkin & Bogatov (1987), Starobogatov et al. (2004)
Nodularia (s.s.) amurensis — Zatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012)
Nodularia (s.s.) schrencki — Zatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012)
Nodularia (s.s.) flavoviridis — Zatravkin & Bogatov (1987), Starobogatov et al. (2004)
Nodularia (s.s.) vladivostokensis — Zatravkin & Bogatov (1987), Starobogatov et al. (2004)
Nodularia (s.s.) sujfunica — Zatravkin & Bogatov (1987)
Nodularia (Amurunio) lebedevi — Zatravkin & Bogatov (1987), Korniushin (1998), Starobogatov et al. (2004), Klishko et al. (2018)
Nodularia (Magadaninaia) extremalis — Martynov & Chernyshev (1992), Starobogatov et al. (2004)
Unio douglasiae biwae — Kondo (1997), Kondo (2008)
Unio sculptus — Yu et al. (2001)
Middendorffinaia (s.s.) ochotica — Starobogatov et al. (2004), Bogatov (2012), Vinarski & Kantor (2016), Bogatov & Prozorova (2017)
Nodularia (s.s.) sakhalinensis — Starobogatov et al. (2004)
Nodularia (s.s.) moskvichevae — Starobogatov et al. (2004)
Middendorffinaia arsenievi — Kantor & Sysoev (2005), Kantor et al. (2010)
Middendorffinaia ochotica — Kantor & Sysoev (2005), Kantor et al. (2010)
Nodularia extremalis — Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia lebedevi — Kantor & Sysoev (2005), Kantor et al. (2010)
Nodularia moskvichevae — Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia sakhalinensis — Kantor & Sysoev (2005), Kantor et al. (2010), Klishko et al. (2018)
Nodularia crassidens — Prozorova et al. (2005)
Scabies longata — Hu (2005), Graf & Cummings (2007), Bogan (2010), WoRMS (2017)
Unio crassus mongolicus — Graf (2007)
Nodularia denserugata — Graf & Cummings (2007), Bogan (2010)
Nodularia biwae — Bogan (2010)
Middendorffinaia (s.s.) ussuriensis — Bogatov (2012), Vinarski & Kantor (2016)
Nodularia dorri — IUCN (2012)
Acuticosta jianghanensis — He & Zhuang (2013), Zieritz et al. (2018), Liu et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)
Ptychorhynchus denserugata — He & Zhuang (2013), Zieritz et al. (2018)
Scabies crispata — He & Zhuang (2013), Graf & Cummings (2021)
Nodularia douglasiae biwae — Sano et al. (2017), Mabuchi & Nishida (2020)
Nodularia douglasiae nipponensis — Sano et al. (2017)
Nodularia jourdyi — Bolotov et al. (2018), Pfeiffer et al. (2019), Sayenko et al. (2021), Graf & Cummings (2021)
Nodularia suifunica [sic] — Klishko et al. (2018)
Middendorffinaia mongolica — Klishko et al. (2019)
Nodularia cf. douglasiae — Huang et al. (2019)
Nodularia douglasiae Clade C — Choi et al. (2020)
Nodularia douglasiae Clade B — Choi et al. (2020)
Nodularia douglasiae Clade A — Choi et al. (2020)
Ptychorhynchus denserugatum — Graf & Cummings (2021)
Nodularia douglasiae douglasiae — Bogatov (2022)
Nodularia douglasiae amurensis — Bogatov (2022)
Nodularia douglasiae schrencki — Bogatov (2022)
Nodularia douglasiae middendorffi — Bogatov (2022)
Middendorffinaia mongolica ochotica — Bogatov (2022)
Middendorffinaia mongolica ussuriensis — Bogatov (2022)
Nodularia sp. 2 — Dai et al. (2024)