species Nodularia douglasiae (Griffith & Pidgeon, 1833): synonymy

Unio douglasiae Griffith & Pidgeon, 1833

Griffith & Pidgeon, 1833. [in] Cuvier, Animal Kingdom 12(38): pl. 21, fig. 2 [as ‘douglasiæ’].
‘Gray’ Griffith & Pidgeon, 1834. [in] Cuvier, Animal Kingdom 12(39-40): 601 [as ‘douglasiæ’].

Nodularia (s.s.) douglasiæ — Simpson (1900)
Unio (Nodularia) douglasiae — Martens (1905), Zhadin (1938)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Nodularia (s.s.) douglasiae — Preston (1912), Simpson (1914), Haas (1924), Korniushin (1998)
Nodularia douglasiae — Haas (1920), Prozorova et al. (2005), Graf (2007), Graf & Cummings (2007), Bogan (2010), ... Chen et al. (2025), Klymus et al. (2025), Campbell & Lydeard (2026), Wu et al. (2026), Wu et al. (2026)
Unio douglasiae — Zhadin (1965), Kil (1976), Habe (1977), Liu et al. (1979), Liu et al. (1980), ... Zhou et al. (2017), Araujo et al. (2018), Wang et al. (2018), Aunins et al. (2018), Wang et al. (2018)
Unio douglasiae douglasiae — Haas (1969)
Nodularidia douglasiae — He & Zhuang (2013)
“Unio douglasiae” ** — Ouyang et al. (2015)
“Nodularia douglasiae” ** — Liu et al. (2017), Klishko et al. (2019), Sayenko et al. (2021), Dai et al. (2024)
“Nodularia douglasiae” ** — Liu et al. (2017), Mabuchi et al. (2021)
“Nodularia douglasiae” ** — Cha et al. (2018), Fukata & Iigo (2019), Sayenko et al. (2021), Mabuchi et al. (2021), Kim et al. (2021), Lopes-Lima et al. (2024), Dai et al. (2024), Wu et al. (2024), Chen et al. (2025)
“Nodularia douglasiae” ** — Fukata & Iigo (2019), Sano et al. (2020), Kim et al. (2021), Pavan-Kumar et al. (2022), Dai et al. (2024)
“Nodularia cf. douglasiae” ** — Huang et al. (2019)
Nodularia douglasiae Clade C — Choi et al. (2020)
Nodularia douglasiae Clade B — Choi et al. (2020)
“Nodularia douglasiae Clade A” ** — Choi et al. (2020)
Nodularia douglasiae Clade A — Choi et al. (2020)
“Nodularia douglasiae Clade B” ** — Choi et al. (2020)
Nodularia douglasiae douglasiae — Bogatov (2022)
Nodularia sp. 2 — Dai et al. (2024)
“Nodularia douglasiae” ** — Klymus et al. (2025)

Unio murchisonianus Lea, 1834

Lea, 1834. Trans. Amer. Phil. Soc. 5 [O. 1]: 33, pl. 3, fig. 6.
Type(s): HOLOTYPE (MONOTYPY) USNM_84156, China [“Calcutta, India” penciled out].

Nodularia (s.s.) douglasiæ — Simpson (1900)
Unio (Nodularia) douglasiae — Martens (1905)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Nodularia (s.s.) douglasiae — Preston (1912), Simpson (1914)
Unio douglasiae douglasiae — Haas (1969)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae — Lopes-Lima et al. (2020)

Unio osbeckii Philippi, 1844

Philippi, 1844. Zeitschr. f. Malakozool. 1: 164.
Philippi, 1847. Abbild. Beschr. Conch. 3: 45, pl. 3, fig. 1.

Nodularia (s.s.) douglasiæ — Simpson (1900)
Unio (Nodularia) douglasiae — Martens (1905)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Nodularia (s.s.) douglasiae — Preston (1912), Simpson (1914)
Unio douglasiae douglasiae — Haas (1969)
Unio douglasiae — Liu et al. (1979)
Nodularia douglasiae — Prozorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)

Unio shanghaiensis Lea, 1859

Lea, 1859. Proc. Acad. Nat. Sci. 11: 153.
Lea, 1860. Jour. Acad. Nat. Sci. 4 [O. 7]: 242, pl. 36, fig. 121.
Type(s): HOLOTYPE BMNH_1965171, Shanghai, China.

Nodularia (s.s.) douglasiæ var. shanghaiensis — Simpson (1900)
Unio (Nodularia) douglasiae — Martens (1905)
Nodularia (s.s.) douglasiae shanghaiensis — Haas (1911)
Nodularia (s.s.) douglasiae var. shanghaiensis — Preston (1912), Simpson (1914)
Nodularia douglasiae — Haas (1920), Lopes-Lima et al. (2020)
Unio douglasiae douglasiae — Haas (1969)
Unio douglasiae — Liu et al. (1979)
Nodularidia douglasiae — He & Zhuang (2013)

Unio wrightii Lea, 1865

Lea, 1865. Proc. Acad. Nat. Sci. 17: 75.
Lea, 1868. Jour. Acad. Nat. Sci. 6 [O. 12]: 283, pl. 39, fig. 97.
Type(s): HOLOTYPE USNM_84129, China.

Nodularia (s.s.) douglasiæ — Simpson (1900)
Unio (Nodularia) douglasiae — Martens (1905)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Nodularia (s.s.) douglasiae — Simpson (1914)
Unio wrigtii [sic] — de Gregorio (1914)
Unio douglasiae douglasiae — Haas (1969)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae — Lopes-Lima et al. (2020)

Unio sculptus Deshayes, 1873

Deshayes, 1873. Nouvelles archives 9: 9, pl. 1, figs. 3-3a.
Type(s): SYNTYPES MNHN_IM-2000-1779, Petchily, Chine Centrale.

Nodularia (s.s.) douglasiæ — Simpson (1900)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Nodularia (s.s.) douglasiae — Preston (1912), Simpson (1914)
Unio douglasiae douglasiae — Haas (1969)
Unio douglasiae — Liu et al. (1979)
Unio sculptus — Yu et al. (2001)
Nodularia douglasiae — Prozorova et al. (2005), Schneider et al. (2013), Lopes-Lima et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)

Unio biwae Kobelt, 1879

Kobelt, 1879. Abh. Senckenb. Naturf. Ges. 11: 425, pl. 23, figs. 2-4.
Type(s): LECTOTYPE SMF_3650, Biwa-See, Japan.

Nodularia (s.s.) biwæ — Simpson (1900)
Nodularia (s.s.) biwae — Haas (1911), Simpson (1914)
Unio biwae — Haas (1969), Habe (1977), Habe (1991), Slugina (2006)
Unio douglasiae biwae — Kondo (1997), Kondo (2008)
Nodularia biwae — Bogan (2010)
Nodularidia douglasiae — He & Zhuang (2013)
“Nodularia douglasiae biwae” ** — Sano et al. (2017)
Nodularia douglasiae — Lopes-Lima et al. (2020), Mabuchi et al. (2021)
Nodularia douglasiae biwae — Mabuchi & Nishida (2020)

Unio douglasiae chloreus Heude, 1885

(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65 bis, fig. 128c.

Nodularia (s.s.) douglasiæ — Simpson (1900)
Unio douglasiae — Liu et al. (1979)
Nodularia douglasiae — Prozorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)

Unio douglasiae dactylinus Heude, 1885

(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65, fig. 128e.

Nodularia (s.s.) douglasiæ — Simpson (1900)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Nodularia (s.s.) douglasiae — Preston (1912), Simpson (1914)
Unio douglasiae douglasiae — Haas (1969)
Unio douglasiae — Liu et al. (1979)
Nodularia douglasiae — Prozorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)

Unio douglasiae deltae Heude, 1885

(homonym of Unio favidens var. deltae Benson, 1862; intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65 bis, fig. 128d [as ‘deltæ’].

Unio douglasiae — Liu et al. (1979)
Nodularia douglasiae — Prozorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)

Unio douglasiae firmus Heude, 1885

(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833; homonym of Unio firmus Lea, 1866)
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65 bis, figs. 128b-b’.

Nodularia (s.s.) douglasiæ — Simpson (1900)
Unio douglasiae — Liu et al. (1979)
Nodularia douglasiae — Prozorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)

Unio douglasiae hainanensis Heude, 1885

(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65, fig. 128f.

Unio douglasiae — Liu et al. (1979)
Nodularia douglasiae — Prozorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)

Unio jourdyi Morlet, 1886

Morlet, 1886. J. de Conch. 34: 76, pl. 13, figs. 5-5a.
Type(s): SYNTYPES MNHN_MP_0150, Bac Hat etang de la riviere Claire.

Nodularia (s.s.) jourdyi — Simpson (1900), Simpson (1914)
Unio jourdyi — Dautzenberg & Fischer (1905), Dautzenberg & Fischer (1905)
Oxynaia jourdyi — Haas (1913), Thiele (1934), Haas (1969), Haas (1969), Dang et al. (1980), Graf & Cummings (2007), Bogan (2010), IUCN (2011), Zieritz et al. (2018), Do et al. (2018)
Nodularia jourdyi — Bolotov et al. (2018), Pfeiffer et al. (2019), Sayenko et al. (2021), Graf & Cummings (2021)
Nodularia douglasiae — Wu et al. (2024)

Unio pictorum var. amurensis Mousson, 1887

(intro'd as intra-sp. of Mya pictorum Linnaeus, 1758)
Mousson, 1887. J. de Conch. 35: 26, pl. 1, fig. 8.

Nodularia (s.s.) douglasiæ var. shanghaiensis — Simpson (1900)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Nodularia (s.s.) douglasiae var. shanghaiensis — Simpson (1914)
Unio douglasiae amurensis — Haas (1969)
Nodularia amurensis — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia (s.s.) amurensis — Zatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012)
Nodularia douglasiae — Graf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020), Sayenko et al. (2021)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae amurensis — Bogatov (2022)

Unio middendorffi Westerlund, 1890

Middendorff, 1851. Reise Nord. Ost. Sibiriens 2(1): 276-277, pl. 28, figs. 1-3 [as Unio pictorum var. longirostris].
Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 113.
Type(s): NEOTYPE BIV229_5 (by Bolotov et al., 2020: Table S1), Gladkaya River, Primorye Region, Russia.

Unio pictorum middendorffi — Haas (1940)
Unio douglasiae amurensis — Haas (1969)
Nodularia middendorffi — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia (s.s.) middendorffi — Zatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012)
Nodularia douglasiae — Graf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae middendorffi — Bogatov (2022)

Unio pliculosus Martens, 1894

Martens, 1894. S.-Ber. Ges. Naturf. Freunde zu Berlin 1894: 216.
Martens, 1905. Zool. Jb., Jena, Suppl. 8: 60, pl. 3, figs. 3-3b.
Type(s): TYPE ZMB_55620, Singei, Korea.

Unio (Nodularia) pliculosus — Martens (1905)
Nodularia (s.s.) pliculosus — Haas (1911)
Nodularia (s.s.) pliculosa — Simpson (1914)
Unio douglasiae pliculosus — Haas (1969)
Unio douglasiae — Habe (1977)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae — Lopes-Lima et al. (2020)

Unio verrucifer Martens, 1894

Martens, 1894. S.-Ber. Ges. Naturf. Freunde zu Berlin 1894: 216.
Martens, 1905. Zool. Jb., Jena, Suppl. 8: 61, pl. 3, figs. 2-2b.
Type(s): TYPE ZMB_55622, Hangang.

Unio (Nodularia) verrucifer — Martens (1905)
Nodularia (s.s.) verrucifera — Haas (1911), Simpson (1914)
Unio douglasiae verrucifer — Haas (1969)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae — Lopes-Lima et al. (2020)

Unio abbreviatus Westerlund, 1897

(homonym of Unio requienii var. abbreviatus de Joannis, 1859)
Westerlund, 1897. Ann. Mus. St. Pétersbourg 1897: 132.
Type(s): PARATYPE SMF_3662, Ussuri fl. bei Noor, Amur gebiet.

Nodularia (s.s.) douglasiæ — Simpson (1900)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Nodularia (s.s.) douglasiae — Preston (1912), Simpson (1914)
Unio (Nodularia) douglasiae var. schrenki [sic] — Zhadin (1938)
Unio douglasiae var. abbreviatus — Zhadin (1965)
Unio douglasiae amurensis — Haas (1969)
Nodularia abbreviata — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010)
Nodularia (s.s.) abbreviata — Zatravkin & Bogatov (1987), Starobogatov et al. (2004)
Nodularia douglasiae — Graf (2007), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularia (s.s.) schrencki — Bogatov (2012)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae schrencki — Bogatov (2022)

Unio schrencki Westerlund, 1897

Westerlund, 1897. Ann. Mus. St. Pétersbourg 1897: 131.
Type(s): PARATYPE SMF_3663, Amur - Fluss.

Nodularia (s.s.) douglasiæ — Simpson (1900)
Nodularia (s.s.) douglasiae — Simpson (1914)
Unio (Nodularia) douglasiae var. schrenki [sic] — Zhadin (1938)
Unio douglasiae var. schrencki — Zhadin (1965)
Unio douglasiae amurensis — Haas (1969)
Nodularia schrencki — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Bogatov & Prozorova (2017)
Nodularia (s.s.) schrencki — Zatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012)
Nodularia douglasiae — Graf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020), Sayenko et al. (2021)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae schrencki — Bogatov (2022)

Unio boettgeri Neumayr, 1899

(homonym of Unio firmus var. boettgeri Ihering, 1893)
Neumayr, 1899. Wiss. Erg. Reise Szechény Ost-Asien 2: 645, pl. 2, fig. 3 [as ‘böttgeri’].

Nodularia (s.s.) boettgeri — Simpson (1914)
Nodularia douglasiae — Haas (1920)
Nodularia (s.s.) douglasiae — Haas (1924)
Nodularidia douglasiae — He & Zhuang (2013)

Unio kobelti Neumayr, 1899

(homonym of Unio kobelti Kobelt & Rolle, 1895)
Neumayr, 1899. Wiss. Erg. Reise Szechény Ost-Asien 2: 645, pl. 3, fig. 1.

Unio kobelti — Simpson (1914)
Nodularia douglasiae — Haas (1920)
Nodularia (s.s.) douglasiae — Haas (1924)
Unio douglasiae — Liu et al. (1979)
Nodularidia douglasiae — He & Zhuang (2013)

Nodularia douglasiae taiwanica Pilsbry & Hirase, 1905

(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Pilsbry & Hirase, 1905. Proc. Acad. Nat. Sci. 57: 750.
Type(s): HOLOTYPE ANSP_58170, Formosa.

Nodularia (s.s.) douglasiae var. taiwanica — Simpson (1914)
Nodularia (s.s.) douglasiae taiwanica — Haas (1924)
Unio douglasiae taiwanicus — Kuroda (1941), Haas (1969)
Unio douglasiae — Habe (1977), Liu et al. (1979), Wu (1982)
Nodularidia douglasiae — He & Zhuang (2013)

Unio jourdyi var. corrugata Dautzenberg & Fischer, 1905

(homonym of Unio corrugata Mauduyt, 1839; intro'd as intra-sp. of Unio jourdyi Morlet, 1886)
Dautzenberg & Fischer, 1905. J. de Conch. 53: 205.
Type(s): SYNTYPE MNHN_MP_0149, Bas-Luc Nam.

Unio jourdyi var. corrugata — Dautzenberg & Fischer (1905)
Nodularia (s.s.) jourdyi var. corrugata — Simpson (1914)

Unio jourdyi var. ponderosa Dautzenberg & Fischer, 1905

(homonym of Mya ponderosa Dillwyn, 1817; intro'd as intra-sp. of Unio jourdyi Morlet, 1886)
Dautzenberg & Fischer, 1905. J. de Conch. 53: 453.
Type(s): SYNTYPE MNHN_MP_0148, Tonkin, Grand Lac, Hanoi.

Unio jourdyi var. ponderosa — Dautzenberg & Fischer (1905)
Nodularia (s.s.) jourdyi var. ponderosa — Simpson (1914)

Nodularia denserugata Haas, 1910

Haas, 1910. Ann. Mag. Nat. Hist. 6: 496.
Haas, 1910. Conch. Cab. 9 (pt. 2, h. 42): pl. 8, figs. 3-4.
Haas, 1911. Conch. Cab. 9 (pt. 2, h. 44): 83, pl. 8, figs. 3-4.

Nodularia (s.s.) denserugata — Haas (1911), Simpson (1914)
Unio denserugatus — Haas (1969)
Middendorffinaia (Sujfununio) denserugata — Moskvicheva & Starobogatov (1973)
Nodularia denserugata — Graf & Cummings (2007), Bogan (2010)
Ptychorhynchus denserugata — He & Zhuang (2013), Zieritz et al. (2018)
Ptychorhynchus denserugatum — Graf & Cummings (2021)
Nodularia douglasiae — Wu et al. (2024)

Nodularia verrucosa Haas, 1910

Haas, 1910. Nachr. Bl. Dtsch. Malak. Ges. 42: 99.
Type(s): TYPE SMF_3660, Hunan, M. China.

Nodularia verrucosa — Haas (1910), Simpson (1914)
Nodularidia douglasiae — He & Zhuang (2013)

Nodularia douglasiae crassidens Haas, 1910

(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Haas, 1910. Ann. Mag. Nat. Hist. 6: 499.
Haas, 1910. Conch. Cab. 9 (pt. 2, h. 42): pl. 7, figs. 4-5.
Haas, 1911. Conch. Cab. 9 (pt. 2, h. 44): 75.
Type(s): TYPE SMF_3659, Hunan, M. China.

Nodularia (s.s.) douglasiae crassidens — Haas (1911)
Nodularia (s.s.) douglasiae var. crassidens — Simpson (1914)
Unio douglasiae douglasiae — Haas (1969)
Nodularia douglasiae crassidens — Dang et al. (1980), Do et al. (2018)
Nodularia crassidens — Prozorova et al. (2005)
Nodularia dorri — IUCN (2012)
Nodularidia douglasiae — He & Zhuang (2013)

Nodularia douglasiae flavoviridis Haas, 1910

(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Haas, 1910. Conch. Cab. 9 (pt. 2, h. 41): 71, pl. 6, fig. 2.
Type(s): TYPE SMF_3654, W. Ufer d. Sees Buirnor, O. Mongolei, Amur-Geb.

Unio douglasiae amurensis — Haas (1969)
Nodularia flavoviridis — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010)
Nodularia (s.s.) flavoviridis — Zatravkin & Bogatov (1987), Starobogatov et al. (2004)
Nodularia douglasiae — Graf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularia (s.s.) schrencki — Bogatov (2012)
Nodularidia douglasiae — He & Zhuang (2013)

Middendorffinaia arsenjevi Moskvicheva & Starobogatov, 1973

(incorrect original spelling of Middendorffinaia (s.s.) arsenievi Moskvicheva & Starobogatov, 1973)
Moskvicheva & Starobogatov, 1973. Byull. Mosk. Obshch.. Ispyt. Prir. 78(2): 22, fig. 1.
‘Moskv. et Star.’ Zatravkin, 1983. [in] I.M. Likharev, Molluscs: 41.

Middendorffinaia (s.s.) arsenievi Moskvicheva & Starobogatov, 1973

(correct name for Middendorffinaia arsenjevi Moskvicheva & Starobogatov, 1973)
Moskvicheva & Starobogatov, 1973. Byull. Mosk. Obshch.. Ispyt. Prir. 78(2): 29, figs. 3м-п.

Middendorffinaia (s.s.) arsenievi — Moskvicheva & Starobogatov (1973), Zatravkin & Bogatov (1987), Starobogatov et al. (2004)
Middendorffinaia (s.s.) arsenjevi [sic] — Zatravkin (1983)
Middendorffinaia arsenievi — Kantor & Sysoev (2005), Kantor et al. (2010)
Unio crassus mongolicus — Graf (2007)
Middendorffinaia (s.s.) ussuriensis — Bogatov (2012), Vinarski & Kantor (2016)
Middendorffinaia mongolica — Klishko et al. (2019)
Nodularia douglasiae — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Middendorffinaia mongolica ussuriensis — Bogatov (2022)

Nodularia sujfunica Moskvicheva, 1973

Moskvicheva, 1973. Zool. Zhur. 52(10): 1463, figs. 1 с-т.

Nodularia sujfunica — Moskvicheva (1973), Zatravkin (1983)
Nodularia (s.s.) sujfunica — Zatravkin & Bogatov (1987)
Nodularia (s.s.) vladivostokensis — Starobogatov et al. (2004)
Nodularia vladivostokensis — Kantor & Sysoev (2005)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia suifunica [sic] — Klishko et al. (2018)
Nodularia douglasiae — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularia douglasiae schrencki — Bogatov (2022)

Nodularia vladivostokensis Moskvicheva, 1973

Moskvicheva, 1973. Zool. Zhur. 52(10): 1463, figs. 1 п-р.

Nodularia vladivostokensis — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Klishko et al. (2018)
Nodularia (s.s.) vladivostokensis — Zatravkin & Bogatov (1987), Starobogatov et al. (2004)
Nodularia douglasiae — Graf (2007), Lopes-Lima et al. (2020), Bolotov et al. (2020), Sayenko et al. (2021)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae douglasiae — Bogatov (2022)

Nodularia lebedevi Zatravkin & Starobogatov, 1984

Zatravkin & Starobogatov, 1984. Zool. Zhur. 63(12): 1787, figs. 5-8.

Nodularia sp.n. — Zatravkin (1983)
Nodularia (Amurunio) lebedevi — Zatravkin & Bogatov (1987), Korniushin (1998), Starobogatov et al. (2004), Klishko et al. (2018)
Nodularia lebedevi — Kantor & Sysoev (2005), Kantor et al. (2010)
Nodularia douglasiae — Graf (2007), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularia (s.s.) middendorffi — Bogatov (2012)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia middendorffi — Vinarski & Kantor (2016)
Nodularia douglasiae middendorffi — Bogatov (2022)

Scabies longata Liu et al., 1991

Liu et al., 1991. Acta Zootax. Sinica 16(3): 279 [281], fig’d.

Scabies longata — Hu (2005), Graf & Cummings (2007), Bogan (2010), WoRMS (2017)
Scabies crispata — He & Zhuang (2013), Graf & Cummings (2021)
Nodularia douglasiae — Dai et al. (2024)

Nodularia (Magadaninaia) extremalis Martynov & Chernyshev, 1992

Martynov & Chernyshev, 1992. Zool. Zhur. 71: 19, figs. 1-3.

Nodularia (Magadaninaia) extremalis — Martynov & Chernyshev (1992), Starobogatov et al. (2004)
Nodularia extremalis — Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia douglasiae — Graf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae schrencki — Bogatov (2022)

Nodularia (s.s.) moskvichevae Bogatov & Starobogatov, 1992

Bogatov & Starobogatov, 1992. Zool. Zhur. 71: 134, fig’d.

Nodularia (s.s.) moskvichevae — Starobogatov et al. (2004)
Nodularia moskvichevae — Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia douglasiae — Graf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularia douglasiae middendorffi — Bogatov (2022)

Middendorffinaia (Middendorffinaia) ochotica Bogatov, 2000

Bogatov, 2000. Zool. Zhur. 79: 861, text fig. a-r.

Middendorffinaia (s.s.) ochotica — Starobogatov et al. (2004), Bogatov (2012), Vinarski & Kantor (2016), Bogatov & Prozorova (2017)
Middendorffinaia ochotica — Kantor & Sysoev (2005), Kantor et al. (2010)
Unio crassus mongolicus — Graf (2007)
Middendorffinaia mongolica — Klishko et al. (2019)
Nodularia douglasiae — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Middendorffinaia mongolica ochotica — Bogatov (2022)

Nodularia sakhalinensis Bogatov, 2001

Bogatov, 2001. Byull. Dal'nev. Malak.Obshch. 5: 75.

Nodularia (s.s.) sakhalinensis — Starobogatov et al. (2004)
Nodularia sakhalinensis — Kantor & Sysoev (2005), Kantor et al. (2010), Klishko et al. (2018)
Nodularia douglasiae — Graf (2007), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae schrencki — Bogatov (2022)

Acuticosta jianghanensis He & Zhuang, 2013

He & Zhuang, 2013. Freshwater Bivalves China: 23, fig. 47-48.

Acuticosta jianghanensis — He & Zhuang (2013), Zieritz et al. (2018), Liu et al. (2020)
Nodularia douglasiae — Bogatov & Prozorova (2017)

Nodularia douglasiae nipponensis ‘Martens’ of Authors

(misidentified reference to Unio nipponensis Martens, 1877)
‘Martens’ of Authors.
Sano et al., 2017. J. Wat. Res. Prot. 9: Table 2. [in part]

Nodularia douglasiae — Sano et al. (2020), Choi et al. (2020)

Nodularia suifunica ‘Moskvicheva’ of Authors

(error for Nodularia sujfunica Moskvicheva, 1973)
‘Moskvicheva’ of Authors.
Klishko et al., 2018. Systemat. Biodiv. 16(3): 288.
Bogatov, 2022. Lg. Bivalve Moll. Russia’s Fresh Waters: 59.

Unio ezonis ‘Kuroda, MS’ in ANSP coll. of Authors

‘Kuroda, MS’ in ANSP coll. of Authors.

Unio föttgeri of Authors

of Authors.

Unio douglasiae — Liu et al. (1979)

Margarita (Unio) merchisonianus Lea, 1836

(error for Unio murchisonianus Lea, 1834)
Lea, 1836. Synopsis Fam. Naiades: 14 [corrected in the errata].

Unio pictorum var. longirostris Middendorff, 1851

(misidentified reference to Unio longirostris Rossmässler, 1836)
Middendorff, 1851. Reise Nord. Ost. Sibiriens 2(1): 276-277, pl. 28, figs. 1-3.
Schrenck, 1867. Reise Forsch. Amurlande 2: 696.

Nodularia middendorffi — Moskvicheva (1973), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia (s.s.) middendorffi — Zatravkin & Bogatov (1987), Bogatov (2012)
Nodularidia douglasiae — He & Zhuang (2013)
Nodularia douglasiae — Lopes-Lima et al. (2020)

Unio murchinsonianus Küster, 1862

(error for Unio murchisonianus Lea, 1834)
‘Lea’ Küster, 1862. Conchyl. Cab. 9(2): 309.

Unio pictorum Schrenck, 1867

(misidentified reference to Mya pictorum Linnaeus, 1758)
‘L.’ Schrenck, 1867. Reise Forsch. Amurlande 2: 696-699.

Nodularia middendorffi — Moskvicheva (1973)

Unio pictorum var. longirostris Westerlund, 1877

(misidentified reference to Unio longirostris Rossmässler, 1836)
Westerlund, 1877. Kongl. Svenska Vet. Akad. Handl. 14(12): 74.

Nodularia (s.s.) douglasiæ — Simpson (1900)
Nodularia (s.s.) douglasiae douglasiae — Haas (1911)
Nodularia (s.s.) douglasiae — Preston (1912), Simpson (1914)
Unio douglasiae amurensis — Haas (1969)
Nodularia amurensis — Vinarski & Kantor (2016)

Unio nipponensis Kobelt, 1879

(misidentified reference to Unio nipponensis Martens, 1877)
Kobelt, 1879. Abh. Senckenb. Naturf. Ges. 11: 422, pl. 12, fig. 3.

Nodularia (s.s.) douglasiæ var. nipponensis — Simpson (1900)
Unio (Nodularia) nipponensis — Martens (1905)

Unio douglasiae var. osbeckii Heude, 1885

(reference to Unio osbeckii Philippi, 1844)
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65, fig. 128i.

Unio (Nodularia) douglasiae var. sinuolatus — Martens (1905)

Unio schanghaiensis Neumayr, 1899

(error for Unio shanghaiensis Lea, 1859)
‘Lea’ Neumayr, 1899. Wiss. Erg. Reise Szechény Ost-Asien 2: 645.

Unio wrigtii de Gregorio, 1914

(error for Unio wrightii Lea, 1865)
‘Lea’ de Gregorio, 1914. Il Nat. Siciliano 2: 40.

Unio (Nodularia) douglasiae var. schrenki Zhadin, 1938

(error for Unio schrencki Westerlund, 1897)
‘Westerlund’ Zhadin, 1938. Faune USSR Mollusques 4(1): 99.

Unio taiwanensis Habe, 1977

(error for Nodularia douglasiae taiwanica Pilsbry & Hirase, 1905)
‘Pilsbry’ Habe, 1977. Mollusc. Japan: 114.

Unio firnus Liu et al., 1979

(error for Unio douglasiae firmus Heude, 1885)
Liu et al., 1979. Econ. Fauna China: 74.

Unio douglasiae chlorens Liu et al., 1979

(error for Unio douglasiae chloreus Heude, 1885)
Liu et al., 1979. Econ. Fauna China: 74.

Unio douglasiae cleltae Liu et al., 1979

(error for Unio douglasiae deltae Heude, 1885)
Liu et al., 1979. Econ. Fauna China: 74.

Nodularia doctylina Liu et al., 1980

(error for Unio douglasiae dactylinus Heude, 1885)
Liu et al., 1980. Acta Zool. Sinica 26(4): 366.

Nodularia sachalinensis Bogatov, 2001

(error for Nodularia sakhalinensis Bogatov, 2001)
Bogatov, 2001. Byull. Dal'nev. Malak.Obshch. 5: 59.

Unio douglassiae Kantor & Sysoev, 2005

(error for Unio douglasiae Griffith & Pidgeon, 1833)
‘Griffith et Pidgeon’ Kantor & Sysoev, 2005. Cat. Moll. Russia: 337.

Nodularia jourdyi Bolotov et al., 2018

(reference to Unio jourdyi Morlet, 1886)
Bolotov et al., 2018. Sci. Repts. 8: 4.
Pfeiffer et al., 2019. MPE 137: 117.

Nodularia douglasiae — Lopes-Lima et al. (2020), Choi et al. (2020), Sayenko et al. (2021), Wu et al. (2024)

Unio osbreckii Lopes-Lima et al., 2020

(error for Unio osbeckii Philippi, 1844)
‘Philippi’ Lopes-Lima et al., 2020. MPE 146: Appendix 1.