MUSSELp

MUSSELpdb

Introduction

Browse

d'basing

Search the MUSSELpdb

Taxonomy

all genera

Unionidae

Margaritiferidae

Hyriidae

Etheriidae

Mycetopodidae

Iridinidae


Species & Genus Richness by
Taxonomy

Geography

Summary

North America

Central America

South America

Afrotropics

North Eurasia

East Asia

Australasia


Species Richness & Endemism by
Geography

Data last updated
Mon 1 Dec 2025

Publication Bolotov et al. (2020): summary

publication summary | cladograms

Full Citation

Bolotov, I.N., E.S. Konopleva, I.V. Vikhrev, M.Yu. Gofarov, M. Lopes-Lima et al. 2020. New freshwater mussel taxa discoveries clarify biogeographic division of Southeast Asia. Scientific Reports 10. 6616 (22 pp.). Available electronically.

Attributed Nominal Genera

Nyeinchanconcha Bolotov et al., 2020

Bolotov et al., 2020. Sci. Repts. 10: 11.
Type species: Nyeinchanconcha nyeinchani Bolotov et al., 2020 (orig. desig.) = Nyeinchanconcha nyeinchani

Scabiellus Bolotov et al., 2020

Bolotov et al., 2020. Sci. Repts. 10: 5.
Type species: Scabies songkramensis Kongim et al., 2015 (orig. desig.) = Scabiellus songkramensis

Sundadontina Bolotov et al., 2020

Bolotov et al., 2020. Sci. Repts. 10: 12.
Type species: Anodonta cumingii Lea, 1851 (orig. desig.) = Sundadontina cumingii

Thaiconcha Bolotov et al., 2020

Bolotov et al., 2020. Sci. Repts. 10: 14.
Type species: Anodonta callifera Martens, 1860 (orig. desig.) = Thaiconcha callifera

Attributed Nominal Species

Yaukthwa avaensis Bolotov et al., 2020 = Yaukthwa avaensis

Bolotov et al., 2020. Sci. Repts. 10: 8, fig. 2F.

Balwantia baniensis Bolotov et al., 2020 = Yaukthwa baniensis

Bolotov et al., 2020. Sci. Repts. 10: 5, fig. 2C.

Sundadontina brandti Bolotov et al., 2020 = Sundadontina brandti

Bolotov et al., 2020. Sci. Repts. 10: 12, fig. 4B.

Monodontina cambodiensis Bolotov et al., 2020 = Pseudodon cambodjensis

‘Petit’ of Authors.
Bolotov et al., 2020. Sci. Repts. 10: Table S1.
Konopleva et al., 2021. Sci. Repts. 11: Table S1.
Jeratthitikul et al., 2021. Eur. J. Taxonomy 775: 123.
(error for Monocondylea cambodjensis Petit de la Saussaye, 1865)

image of type lot

Pseudodon cambodjensis Pfeiffer & Graf, 2013 = Pseudodon cambodjensis

‘Petit’ of Authors.
Pfeiffer & Graf, 2013. J. Moll. Studies 79: 251.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Zieritz et al., 2016. Sci. Total Environ. 571: 1071.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Monodontina cambodjensis’].
Lopes-Lima et al., 2017. MPE 106: 177.
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Monodontina cambodjensis’].
Razak et al., 2018. Conserv. Genet. Resour. 11: 459.
Huang et al., 2019. MPE 130: Table S4 [as ‘Monodontina cambodjensis’].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Monodontina cambodjensis’].
Bolotov et al., 2020. Sci. Repts. 10: Table S1 [as ‘Monodontina cambodiensis’ (sic)].
Konopleva et al., 2021. Sci. Repts. 11: Table S1 [as ‘Monodontina cambodiensis’ (sic)].
Jeratthitikul et al., 2021. Eur. J. Taxonomy 775: 123 [as ‘Monodontina cambodiensis’ (sic)].
Pfeiffer et al., 2021. Invert. Syst. 35: 453 [as ‘Monodontina cambodjensis’].
Wu et al., 2021. ZooKeys 1054: 87 [as ‘Monodontina cambodjensis’].
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
Konopleva et al., 2023. Diversity 15(1), 10: Table S2 [as ‘Monodontina cambodjensis’].
Dai et al., 2023. Invert. Syst. 37(2): Table 1 [as ‘Monodontina cambodjensis’].
Dai et al., 2024. Zoosyst. Evol. 100(4): Table 2.
Dai et al., 2025. Zoosyst. Evol. 101(2): Table 1.
(reference to Monocondylea cambodjensis Petit de la Saussaye, 1865)
Type(s): SEQUENCED SPECIMEN UMMZ_304350, Pursat River, just downstream (north) of Highway 5 bridge, Pursat City, Cambodia.

There are 12 available nominal species attributed to this publication (36 total).

Contradens contradens Pfeiffer & Graf, 2015 = Lens micropterus

‘Lea’ of Authors.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Konopleva et al., 2019. Ecologica Montenegrina 24: Appendix 1 [as ‘Contradens aff. contradens sp. 1’].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Huang et al., 2019. MPE 130: Table S4 [as ‘Contradens aff. contradens sp. 1’].
Bolotov et al., 2020. Sci. Repts. 10: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Konopleva et al., 2023. Diversity 15(1), 10: Table S2 [as ‘Lens aff. contradens’].
(misidentified reference to Unio contradens Lea, 1838)

Parreysia corrugata Bolotov et al., 2017 = Parreysia nagpoorensis

‘Müller’ of Authors.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Parreysia corrugata sp. 2’]. [in part]
Bolotov et al., 2017. Sci. Repts. 7: Table S1. [in part]
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Parreysia corrygata sp. 2’].
Pfeiffer et al., 2018. J. Moll. Studies 84(4): Table 1. [in part]
Huang et al., 2019. MPE 130: Table S4 [as ‘Parreysia corrugata sp. 2’].
Bolotov et al., 2020. Sci. Repts. 10: Table S1 [as ‘Parreysia corrugata sp. 3’].
Bolotov et al., 2020. Sci. Repts. 10: Table S1 [as ‘Parreysia corrugata sp. 2’].
Imsong & Murali, 2023. Uttar Pradesh J. Zool. 44(20): Table 1.
(misidentified reference to Mya corrugata Müller, 1774)

Parreysia corrugata Bolotov et al., 2017 = Parreysia keralaensis

‘Müller’ of Authors.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Parreysia corrugata sp. 1’].
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Parreysia corrugata sp. 1’].
Pfeiffer et al., 2018. J. Moll. Studies 84(4): Table 1. [in part]
Huang et al., 2019. MPE 130: Table S4 [as ‘Parreysia corrugata sp. 1’].
Jeratthitikul et al., 2019. Tropical Nat. Hist. 19(1): 25.
Bolotov et al., 2020. Sci. Repts. 10: Table S1 [as ‘Parreysia corrugata sp. 1’].
(misidentified reference to Mya corrugata Müller, 1774)

Contradens crossei Jeratthitikul et al., 2019 = Lens maenamensis

‘Deshayes’ of Authors.
Jeratthitikul et al., 2019. Systemat. Biodiv. 17: 140, figs. 1.7-1.9.
Konopleva et al., 2019. Ecologica Montenegrina 24: 30.
Bolotov et al., 2020. Sci. Repts. 10: Table S1.
Graf & Cummings, 2021. J. Moll. Studies 87(1): 25.
(misidentified reference to Unio crossei Deshayes & Jullien, 1876)

Yaukthwa dalliana Konopleva et al., 2019 = Yaukthwa rectangularis

‘Frierson’ of Authors.
Bolotov et al., 2019. Sci. Repts. 9: Table S1.
Konopleva et al., 2019. Sci. Repts. 9: 7.
Konopleva et al., 2019. Ecologica Montenegrina 24: Appendix 1.
Zieritz et al., 2020. Biodivers. Conserv. 29: 2243.
Bolotov et al., 2020. Sci. Repts. 10: Table S1.
Pfeiffer et al., 2021. Invert. Syst. 35: Table S2.
(misidentified reference to Parreysia dalliana Frierson, 1913)

Anodontites elongata Whelan et al., 2011 = Tamsiella schroeteriana

‘Swainson’ of Authors.
Whelan et al., 2011. MPE 61: 506.
Graf et al., 2015. MPE 85: 3.
Bolotov et al., 2016. MPE 103: 108.
Lopes-Lima et al., 2017. MPE 106: 178.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 208.
Combosch et al., 2017. MPE 107: 195.
Bolotov et al., 2018. Sci. Repts. 8: Table S1.
Huang et al., 2018. MPE 120: 302.
Lopes-Lima et al., 2018. MPE 127: 103.
Huang et al., 2019. MPE 130: Table S4.
Bolotov et al., 2019. Sci. Repts. 9: Table S1.
Pfeiffer et al., 2019. MPE 137: 116.
Guerra et al., 2019. BMC Evol. Biol. 19 (229): 3.
Bolotov et al., 2020. Sci. Repts. 10: Table S1.
Sano et al., 2020. Plankton Benthos Res. 15(2): 101.
Wu et al., 2020. Invert. Syst. 34: 195.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
Chen et al., 2025. Zoosyst. Evol. 101(4): Table 1.
(misidentified reference to Anodon elongatus Swainson, 1823)

Parreysia favidens Pfeiffer et al., 2018 = Parreysia plagiosoma

‘Benson’ of Authors.
Pfeiffer et al., 2018. J. Moll. Studies 84(4): Table 1.
Bolotov et al., 2020. Sci. Repts. 10: Table S1 [as ‘Parreysia favidens sp. 1’].
(misidentified reference to Unio favidens Benson, 1862)

Parreysia favidens Bolotov et al., 2020 = Parreysia corrugata

‘Benson’ Bolotov et al., 2020. Sci. Repts. 10: Table S1 [as ‘Pareysia favidens sp. 2’].
(reference to Unio favidens Benson, 1862)

Hyriopsis gracilis Konopleva et al., 2019 = Hyriopsis khoratensis

‘Haas’ of Authors.
Konopleva et al., 2019. Sci. Repts. 9: Table S1.
Konopleva et al., 2019. Ecologica Montenegrina 24: Appendix 1.
Bolotov et al., 2020. Sci. Repts. 10: Table S1.
Zieritz et al., 2020. Biodivers. Conserv. 29: 2243.
(misidentified reference to Hyriopsis gracilis Haas, 1910)

Chamberlainia hainesiana Kongim et al., 2015 = Chamberlainia somsakpanhai

‘Lea’ of Authors.
Kongim et al., 2015. ZooKeys 514: 98, fig. 2A.
Lopes-Lima et al., 2017. MPE 106: 177.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Froufe et al., 2019. Heredity 124: 185.
Huang et al., 2019. MPE 130: Table S4.
Chen et al., 2020. mtDNA B 5(3): 2800.
Teiga-Teixeira et al., 2020. mtDNA B 5(3): 2978.
Zieritz et al., 2020. Biodivers. Conserv. 29: 2244.
Bolotov et al., 2020. Sci. Repts. 10: Table S1.
Lopes-Lima et al., 2020. MPE 146: Table 1.
Zieritz et al., 2021. Zool. J. Linn. Soc. 193(3): Table 1.
Zieritz et al., 2021. Aquatic Conserv.: Mar. Freshw. Ecosyst. 31: 3176.
Jeratthitikul et al., 2021. Eur. J. Taxonomy 775: 125.
Wu et al., 2021. ZooKeys 1054: 87.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
Wu et al., 2022. Zool. Scripta 51: Table S2.
Pavan-Kumar et al., 2022. Mol. Biol. Rep. 49: Table S1.
Jeratthitikul et al., 2022. Sci. Repts. 12: Table S1.
Dai et al., 2023. Invert. Syst. 37(2): Tables 1-2.
Konopleva et al., 2023. Zoosyst. Evol. 99(1): Table S1.
Wu et al., 2023. Org Divers Evol (2023): Table 2.
Dai et al., 2024. Zool. Scripta 53: Table 1.
Dai et al., 2024. Invert. Syst. 38(6): Table 3.
Dai et al., 2024. Zoosyst. Evol. 100(4): Table 2.
Chen et al., 2025. Zoosyst. Evol. 101(4): Table S3.
Dai et al., 2025. Zoosyst. Evol. 101(2): Table 1.
Dai et al., 2025. Zoosyst. Evol. 101(2): Table 2.
(misidentified reference to Unio hainesianus Lea, 1856)

Ensidens ingallsianus Muanta et al., 2019 = Ensidens telus

‘Lea’ of Authors.
Muanta et al., 2019. J. Moll. Studies 85(2): 227 [as ‘Clade A2’].
Konopleva et al., 2019. Ecologica Montenegrina 24: Appendix 1.
Bolotov et al., 2020. Sci. Repts. 10: Table S1.
Jeratthitikul et al., 2021. Raffles Bull. Zool. 69: 129. [in part]
(misidentified reference to Unio ingallsianus Lea, 1852)

Pseudodon inoscularis Bolotov et al., 2020 = Indopseudodon indawgyiensis

‘Gould’ of Authors.
Bolotov et al., 2020. Sci. Repts. 10: Tables 3, S1.
Konopleva et al., 2021. Sci. Repts. 11: Table S1.
Jeratthitikul et al., 2021. Eur. J. Taxonomy 775: 124.
Bolotov et al., 2022. Hindawi 6600359: Table S1.
Jeratthitikul et al., 2022. Sci. Repts. 12: Table S1.
Konopleva et al., 2023. Diversity 15(1), 10: Table S2.
Kongim et al., 2023. Tropical Nat. Hist. 7: Table 1.
Jeratthitikul & Sutcharit, 2023. Tropical Nat. Hist. 7: Table 1.
(misidentified reference to Anodon inoscularis Gould, 1844)

Pseudodon kayinensis Bolotov et al., 2020 = Indopseudodon kayinensis

Bolotov et al., 2020. Sci. Repts. 10: 14, fig. 3G.

Radiatula khadakvaslaensis Bolotov et al., 2017 = Indonaia shurtleffiana

‘Ray’ of Authors.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Radiatula khadakvaslaensis sp. 1’].
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Indonaia khadakvaslaensis sp. 1’].
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Indonaia khadakvaslaensis sp. 1’].
Huang et al., 2019. MPE 130: Table S4 [as ‘Indonaia khadakvaslaensis sp. 1’].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Indonaia khadakvaslaensis sp. 1’].
Bolotov et al., 2020. Sci. Repts. 10: Table S1 [as ‘Indonaia khadakvaslaensis’].
Bolotov et al., 2022. Hindawi 6600359: Table S1 [as ‘Indonaia khadakvaslaensis’].
(reference to Indonaia khadakvaslaensis Ray, 1966)

image of type lot

Monodontina laosica Bolotov et al., 2020 = Pseudodon vondembuschianus

Bolotov et al., 2020. Sci. Repts. 10: 11, fig. 3C.
Type(s): HOLOTYPE (ORIG. DESIG.) UMMZ_304650, Houai Pin, about 300 m upstream of mouth, ca. 4 km..., Attapeu Province, Laos. 14.7944, 106.48417.

Pilsbryoconcha lemeslei Bolotov et al., 2020 = Pilsbryoconcha acuta

‘Morelet’ of Authors.
Bolotov et al., 2020. Sci. Repts. 10: Table S1.
Konopleva et al., 2021. Sci. Repts. 11: Table S1.
(misidentified reference to Anodonta lemeslei Morelet, 1875)

Trapezoideus lenya Bolotov et al., 2020 = Trapezoideus lenya

Bolotov et al., 2020. Sci. Repts. 10: 7, fig. 2E.

Monodontina lenyanensis Bolotov et al., 2020 = Pseudodon lenyanensis

Bolotov et al., 2020. Sci. Repts. 10: 11, fig. 3D.

Indonaia lima Bolotov et al., 2017 = Indonaia rugosa

‘Simpson’ of Authors.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2018. Sci. Repts. 8: Table S1.
Huang et al., 2019. MPE 130: Table S4.
Konopleva et al., 2019. Sci. Repts. 9: Table S1.
Jeratthitikul et al., 2019. Tropical Nat. Hist. 19(1): 25.
Bolotov et al., 2020. Sci. Repts. 10: Table S1.
Bolotov et al., 2022. Hindawi 6600359: Table S1.
(misidentified reference to Nodularia (sec. Radiatula) lima Simpson, 1900)

Potomida littoralis Whelan et al., 2011 = Potomida semirugata

‘Cuvier’ of Authors.
Whelan et al., 2011. MPE 61: 506. [in part]
Bolotov et al., 2020. Sci. Repts. 10: Table S1.
Konopleva et al., 2021. Sci. Repts. 11: Table S1.
Jeratthitikul et al., 2021. Eur. J. Taxonomy 775: 125.
Bolotov et al., 2022. Hindawi 6600359: Table S1.
Jeratthitikul et al., 2022. Sci. Repts. 12: Table S1.
Konopleva et al., 2023. Diversity 15(1), 10: Table S2.
Konopleva et al., 2023. Zoosyst. Evol. 99(1): Table S1.
Dai et al., 2023. Invert. Syst. 37(2): Table 1. [in part]
Kongim et al., 2023. Tropical Nat. Hist. 7: Table 1.
Jeratthitikul et al., 2024. Zoosyst. Evol. 100(4): Table S1.
Dai et al., 2024. Zoosyst. Evol. 100(4): Table 2.
Jeratthitikul et al., 2025. ZooKeys 1224: Table S1.
(misidentified reference to Unio littoralis Cuvier, 1798)

Lamellidens marginalis Bolotov et al., 2017 = Lamellidens mainwaringi

‘Lamarck’ of Authors.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Lamellidens marginalis sp. 2’ and ‘sp. 3’].
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Lamellidens marginalis sp. 2’ and ‘sp. 3’].
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207. [in part]
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Lamellidens marginalis sp. 3’].
Huang et al., 2019. MPE 130: Table S4 [as ‘Lamellidens marginalis sp. 2’ and ‘sp. 3’].
Jeratthitikul et al., 2019. Tropical Nat. Hist. 19(1): 25.
Konopleva et al., 2020. Ecologica Montenegrina 27: 47.
Bolotov et al., 2020. Sci. Repts. 10: Table S1.
(misidentified reference to Unio marginalis Lamarck, 1819)

Monodontina mekongi Bolotov et al., 2020 = Pseudodon mekongi

Bolotov et al., 2020. Sci. Repts. 10: 11, fig. 3E.

Sundadontina myanmarensis Bolotov et al., 2020 = Sundadontina tanintharyiensis

Bolotov et al., 2020. Sci. Repts. 10: fig. S1.
(error for Sundadontina tanintharyiensis Bolotov et al., 2020)

Balwantia myanmarensis Bolotov et al., 2020 = Yaukthwa baniensis

Bolotov et al., 2020. Sci. Repts. 10: fig. S1.
(error for Balwantia baniensis Bolotov et al., 2020)

image of type lot

Nyeinchanconcha nyeinchani Bolotov et al., 2020 = Nyeinchanconcha nyeinchani

Bolotov et al., 2020. Sci. Repts. 10: 12, fig. 3F.
Type(s): HOLOTYPE (ORIG. DESIG.) NCSM_84884, Asia, Mekong Basin/Mekong R. Laos, Khammouan, Thakhek Co. Houay Karnbeung, small stream arising at cave near Ban Khounephavang [Kouanphavang], Nam Done drainage, 14 [air] km from Thakhek road to Gnommalath, [0.92 air miles WNW center Xianliab]. 17.45781° N, 104.9263° E.

Solenaia oleivora Huang et al., 2019 = Sinosolenaia recognita

‘Heude’ of Authors.
Huang et al., 2019. MPE 130: Table S4 [as ‘Solenaia oleivora sp. 2’].
Lopes-Lima et al., 2020. MPE 146: Table 1 [as ‘Solenaia cf. oleivora 3’].
Bolotov et al., 2020. Sci. Repts. 10: Table S1 [as ‘Solenaia oleivora sp. 2’].
(misidentified reference to Mycetopus oleivorus Heude, 1877)

Solenaia oleivora Huang et al., 2019 = Sinosolenaia iridinea

‘Heude’ of Authors.
Huang et al., 2019. MPE 130: Table S4 [as ‘Solenaia oleivora SDCFM189’].
Chen et al., 2020. mtDNA B 5(3): 2800 [as ‘Solenaia oleivora MG’]. [in part]
Lopes-Lima et al., 2020. MPE 146: Table 1 [as ‘Solenaia cf. oleivora 1’].
Bolotov et al., 2020. Sci. Repts. 10: Table S1 [as ‘Solenaia oleivora sp. 1’].
Sano et al., 2020. Plankton Benthos Res. 15(2): 101.
Bolotov et al., 2022. Hindawi 6600359: Table S1.
(misidentified reference to Mycetopus oleivorus Heude, 1877)

Parreysia rakhinensis Bolotov et al., 2020 = Parreysia rakhinensis

Bolotov et al., 2020. Sci. Repts. 10: 4, figs. 2A-B.

Ensidens sagittarius Muanta et al., 2019 = Ensidens dugasti

‘Lea’ of Authors.
Muanta et al., 2019. J. Moll. Studies 85(2): 227 [as ‘Clade B2’].
Konopleva et al., 2019. Ecologica Montenegrina 24: Appendix 1.
Bolotov et al., 2020. Sci. Repts. 10: Table S1.
(misidentified reference to Unio sagittarius Lea, 1856)

Sundadontina tanintharyiensis Bolotov et al., 2020 = Sundadontina tanintharyiensis

Bolotov et al., 2020. Sci. Repts. 10: 12, fig. 4C.

Sundadontina taskaevi Bolotov et al., 2020 = Sundadontina taskaevi

Bolotov et al., 2020. Sci. Repts. 10: 13, fig. 4D.

Taxonomic Opinions

There are 171 taxonomic opinions from this publication. Click here to see the full list.

 
NSF icon MUSSEL icon
"Making the world a better place, one mollusk at a time."