Publication Graf & Cummings (2021): species taxonomic opinions

Graf, D.L. & K.S. Cummings. 2021. A ‘big data’ approach to global freshwater mussel diversity (Bivalvia: Unionoida), with an updated checklist of genera and species. Journal of Molluscan Studies 87(1). eyaa034 (36 pp.). https://doi.org/10.1093/mollus/eyaa034.

Nephronaias championi (Martens, 1900) = Nephronaias goascoranensis

Nephronaias copanensis (Martens, 1900)

Nephronaias dysonii (Lea, 1859)

Nephronaias goascoranensis (Lea, 1858)

Nephronaias gundlachi (Dunker, 1858) = Actinonaias gundlachi

Nephronaias hermanni (Haas, 1929) = Psoronaias guatemalensis

Nephronaias hjalmarsoni (Dunker, 1858)

Nephronaias lempensis Marshall, 1926 = Nephronaias goascoranensis

Nephronaias macnielii (Lea, 1869)

Nephronaias mellea (Lea, 1859)

Nephronaias oregonensis (Lea, 1852)

Nephronaias plicatula (Küster, 1856) = Nephronaias aztecorum

Nephronaias ravistella (Morelet, 1849)

Nephronaias rowellii (Lea, 1859)

Nephronaias rugulosa (Küster, 1856)

Nephronaias sajensis (Frierson, 1927) = Nephronaias dysonii

Nephronaias scamnata (Morelet, 1849)

Nephronaias sphenorhyncha (Crosse & Fischer, 1893)

Nephronaias tabascoensis (Küster, 1856)

Nephronaias tempisquensis Pilsbry, 1920

Nephronaias yzabalensis (Crosse & Fischer, 1892)

Nitia acuminata (H. Adams, 1866)

Nitia chefneuxi (Neuville & Anthony, 1906)

Nitia monceti (Bourguignat, 1883)

Nitia mutelaeformis (Germain, 1906)

Nitia teretiuscula (Philippi, 1847)

Nodularia breviconcha Lee, Kim, Bogan & Kondo in Lopes-Lima et al., 2020

Nodularia diespiter (Mabille, 1887)

Nodularia dorri (Wattebled, 1886)

Nodularia douglasiae (Griffith & Pidgeon, 1833)

Nodularia gladiator (Ancey, 1881)

Nodularia jourdyi (Morlet, 1886) = Nodularia douglasiae

Nodularia micheloti (Morlet, 1886)

Nodularia nipponensis (Martens, 1877)

Nodularia nuxpersicae (Dunker, 1849)

Nodularia persculpta Haas, 1910

Nyassunio nyassaensis (Lea, 1864)

Nyassunio ujijiensis (Crosse, 1881)

Nyeinchanconcha nyeinchani Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan & Kondakov, 2020

Obliquaria reflexa Rafinesque, 1820

Obovalis omiensis (Heimburg, 1884)

+ Pseudodon loomisi Simpson, 1900

Obovaria arkansasensis (Lea, 1862)

+ Unio (Obovaria) jacksoniana Frierson, 1912

Obovaria choctawensis (Athearn, 1964)

Obovaria haddletoni (Athearn, 1964)

Obovaria olivaria (Rafinesque, 1820)

Obovaria retusa (Lamarck, 1819)

Obovaria subrotunda (Rafinesque, 1820)

Obovaria unicolor (Lea, 1845)

Ortmanniana ligamentina (Lamarck, 1819)

+ Unio carinatus Barnes, 1823

Ortmanniana pectorosa (Conrad, 1834)

Pachynaias rugososulcata (Lea, 1866) = Pachynaias spheniopsis

Pachynaias spheniopsis (Morelet, 1849)

Paetulunio fabalis (Lea, 1831)

Parreysia annandalei Preston, 1912 = Parreysia marcens

Parreysia corbis (Hanley, 1856) = Indonaia corbis

Parreysia corrugata (Müller, 1774)

+ Unio multidentatus Philippi, 1847

Parreysia cylindrica Annandale & Prashad, 1919 = Indonaia cylindrica

Parreysia favidens (Benson, 1862) = Parreysia corrugata

Parreysia gowhattensis (Theobald, 1873) = Parreysia marcens

Parreysia rajahensis (Lea, 1841)

Parreysia rakhinensis Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan & Kondakov, 2020

Parreysia rugosa (Gmelin, 1791) = Indonaia rugosa

Parreysia sikkimensis (Lea, 1859) = Parreysia corrugata

Parreysia smaragdites (Benson, 1862)

Parreysia triembola (Benson, 1862) = Parreysia nagpoorensis

Parreysia viridula (Benson, 1862) = Parreysia corrugata

Parvasolenaia neotriangularis (He & Zhuang, 2013)

Parvasolenaia rivularis (Heude, 1877)

Parvasolenaia triangularis (Heude, 1885)

Parvaspina collina (Conrad, 1837)

Parvaspina steinstansana (Johnson & Clarke, 1983)

Pegias fabula (Lea, 1838)

Physunio cambodiensis (Lea, 1856) = Lens contradens

Physunio friersoni Simpson, 1914 = Lamellidens friersoni

Physunio inornatus (Lea, 1856) = Lens inornatus

Physunio micropterus (Morelet, 1866) = Lens micropterus

+ Unio semialatus Deshayes in Deshayes & Jullien, 1876

Physunio modelli Brandt, 1974

Physunio superbus (Lea, 1843)

+ Unio gravidus Lea, 1856

Pilsbryoconcha carinifera (Conrad, 1837)

Pilsbryoconcha compressa (Martens, 1860) = Pilsbryoconcha exilis

Pilsbryoconcha exilis (Lea, 1838)

Pilsbryoconcha expressa (Martens, 1900)

Pilsbryoconcha lemeslei (Morelet, 1875) = Namkongnaia lemeslei

Plectomerus dombeyanus (Valenciennes in Humboldt & Bonpland, 1827)

+ Unio trapezoides Lea, 1831

Pleiodon ovatus (Swainson, 1823)

+ Pleiodon macmurtriei Conrad, 1834

Pleiodon spekii (Woodward, 1859)

Plethobasus cicatricosus (Say, 1829)

Plethobasus cooperianus (Lea, 1834)

Plethobasus cyphyus (Rafinesque, 1820)

+ Unio aesopus Green, 1827

Pletholophus reiniana (Martens, 1875) = Pletholophus reinianus

Pletholophus tenuis (Griffith & Pidgeon, 1833)

+ Symphynota discoidea Lea, 1834

Pleurobema athearni Gangloff, Williams & Feminella, 2006

Pleurobema beadleianum (Lea, 1861)

Pleurobema clava (Lamarck, 1819)

+ Unio bournianus Lea, 1840

+ Pleurobema mytiloides Rafinesque, 1820

Pleurobema cordatum (Rafinesque, 1820)

Pleurobema curtum (Lea, 1859)

Pleurobema decisum (Lea, 1831)

+ Unio chattanoogaensis Lea, 1858

Pleurobema fibuloides (Lea, 1859)

Pleurobema georgianum (Lea, 1841)

+ Unio nucleopsis Conrad, 1849

+ Unio troschelianus Lea, 1852

Pleurobema hanleyianum (Lea, 1852)

Pleurobema hartmanianum (Lea, 1860)

Pleurobema marshalli Frierson, 1927

Pleurobema oviforme (Conrad, 1834)

Pleurobema perovatum (Conrad, 1834)

+ Unio johannis Lea, 1859

+ Unio flavidulus Lea, 1861

Pleurobema plenum (Lea, 1840)

Pleurobema pyriforme (Lea, 1857)

Pleurobema riddellii (Lea, 1862)

Pleurobema rubellum (Conrad, 1834)

+ Pleurobema avellana Simpson, 1900

+ Unio furvus Conrad, 1834

+ Unio (Pleurobema) hagleri Frierson, 1900

Pleurobema rubrum (Rafinesque, 1820) = Pleurobema sintoxia

Pleurobema sintoxia (Rafinesque, 1820)

Pleurobema stabile (Lea, 1861)

+ Unio murrayensis Lea, 1868

Pleurobema strodeanum (B.H. Wright, 1898)

Pleurobema taitianum (Lea, 1834)

Pleurobema verum (Lea, 1861)

Pleuronaia barnesiana (Lea, 1838)

Pleuronaia dolabelloides (Lea, 1840)

Pleuronaia gibber (Lea, 1838)

Popenaias metallica (Say, 1831)

Popenaias popeii (Lea, 1857)

Popenaias tehuantepecensis (Crosse & Fischer, 1893) = Nephronaias tehuantepecensis

Potamilus alatus (Say, 1817)

Potamilus amphichaenus (Frierson, 1898)

Potamilus capax (Green, 1832)

Potamilus inflatus (Lea, 1831)

Potamilus metnecktayi R.I. Johnson, 1998

Potamilus ohiensis (Rafinesque, 1820)

Potamilus purpuratus (Lamarck, 1819)

Potamilus streckersoni Smith, Johnson, Inoue, Doyle & Randklev, 2019

Potomida acarnanica (Kobelt, 1879)

Potomida littoralis (Cuvier, 1798)

+ Mysca (Potomida) corrugata Swainson, 1840

Potomida semirugata (Lamarck, 1819)

Pressidens exanthematicus (Küster, 1861)

Pressidens insularis (Drouet, 1894) = Pressidens exanthematicus

Pressidens moellendorffi Haas, 1910 = Pressidens exanthematicus

Prisodon obliquus Schumacher, 1817

Prisodon syrmatophorus (Gmelin, 1791)

Prisodontopsis aviculaeformis F.R. Woodward, 1991

+ Unio (Metaptera) johnstoni E.A. Smith, 1893

Prisodontopsis mweruensis (E.A. Smith, 1908)

Prohyriopsis stolata (Martens, 1900)

Pronodularia japanensis (Lea, 1859)

Protunio messageri (Bavay & Dautzenberg, 1901)

Pseudanodonta complanata (Rossmässler, 1835)

Pseudobaphia banggiangensis Bogan & Do, 2018

Pseudobaphia biesiana (Heude, 1877)

Pseudobaphia liuovatus (He & Zhuang, 2013) = Pseudocuneopsis liuovatus

+ Unio ovatus Liu, Duan & Wang in Liu et al., 1994

Pseudodon aureus Heude, 1885 = Pseudobaphia aurea

Pseudodon avae (Theobald, 1873) = Indopseudodon salwenianus

Pseudodon bogani Bolotov, Kondakov & Konopleva in Bolotov et al., 2017 = Indopseudodon bogani

Pseudodon crebristriatus (Anthony, 1865) = Indopseudodon crebristriatus

Pseudodon inoscularis (Gould, 1844)

Pseudodon kayinensis Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan & Kondakov, 2020 = Indopseudodon kayinensis

Pseudodon manueli Konopleva, Kondakov & Vikrev in Bolotov et al., 2017 = Indopseudodon salwenianus

Pseudodon nankingensis (Heude, 1874) = Simpsonasus nankingensis

Pseudodon peguensis (Anthony, 1865) = Indopseudodon salwenianus

Pseudodon pinchonianus (Heude, 1883) = Ptychorhynchus murinum

+ Pseudodon solidus Haas, 1911

Pseudodon resupinatus Martens, 1902 = Cosmopseudodon resupinatus

Pseudodon salwenianus (Gould, 1844) = Indopseudodon salwenianus

Pseudodon secundus Heude, 1877 = Simpsonasus nankingensis

Pseudodontoideus connasaugaensis (Lea, 1858)

+ Margaritana alabamensis Lea, 1861

Pseudodontoideus subvexus (Conrad, 1834)

Pseudodontopsis euphratica (Bourguignat, 1852)

Pseudomulleria dalyi (E.A. Smith, 1898)

Pseudospatha tanganyicensis (E.A. Smith, 1880)

Pseudunio auricularius (Spengler, 1793)

+ Unio sinuata Lamarck, 1819

Pseudunio homsensis (Lea, 1865)

Pseudunio marocanus (Pallary, 1918)

Psoronaias distincta (Crosse & Fischer, 1893)

Psoronaias guatemalensis (Simpson, 1900)

Psoronaias herrerae (Marshall, 1923) = Psoronaias semigranosa

Psoronaias kuxensis Frierson, 1917 = Psoronaias guatemalensis

Psoronaias martensi Frierson, 1927 = Psoronaias crocodilorum

Psoronaias ostreata (Morelet, 1849) = Psoronaias crocodilorum

Psoronaias percompressa (Martens, 1887) = Psoronaias crocodilorum

Psoronaias profunda (Simpson, 1914) = Psoronaias crocodilorum

Psoronaias quadrata (Simpson, 1914) = Psoronaias crocodilorum

Psoronaias salinarum (Haas, 1929) = Psoronaias crocodilorum

Psoronaias semigranosa (von dem Busch in Philippi, 1843) = Psoronaias semigranosa + Psoronaias crocodilorum

+ Unio crocodilorum Morelet, 1849

+ Unio morini Morelet, 1851

+ Quadrula rudis Simpson, 1900

+ Unio psoricus Morelet, 1851

Psoronaias usumasintae (Crosse & Fischer, 1892) = Psoronaias crocodilorum

Ptychobranchus fasciolaris (Rafinesque, 1820)

+ Unio phaseolus Hildreth, 1828

Ptychobranchus foremanianus (Lea, 1842)

Ptychobranchus greenii (Conrad, 1834)

Ptychobranchus jonesi (van der Schalie, 1934)

Ptychobranchus occidentalis (Conrad, 1836)

Ptychobranchus subtentus (Say, 1825)

Ptychorhynchus denserugatum (Haas, 1910) = Nodularia douglasiae

Ptychorhynchus murinum (Heude, 1883)

Ptychorhynchus pfisteri (Heude, 1874)

Ptychorhynchus schoedei (Haas, 1930)

Ptychorhynchus schomburgianum (Heude, 1885)

Pyganodon cataracta (Say, 1817)

Pyganodon fragilis (Lamarck, 1819)

Pyganodon gibbosa (Say, 1824)

Pyganodon grandis (Say, 1829) = Pyganodon grandis + Pyganodon globosa

+ Anodonta globosa Lea, 1841

Pyganodon lacustris (Lea, 1857)

Quadrula couchiana (Lea, 1860)

Quadrula fragosa (Conrad, 1835)

Quadrula quadrula (Rafinesque, 1820)

+ Unio apiculatus Say, 1829

+ Unio rumphianus Lea, 1852

Radiatula bonneaudii (Eydoux, 1838) = Indonaia bonneaudii

Radiatula chaudhurii (Preston, 1912)

Radiatula chindwinensis Bolotov, Konopleva, Vikrev, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Gofarov, Tomilova & Kondakov, 2019

Radiatula crispisulcata (Benson, 1862)

Radiatula mouhoti Vikhrev, Bolotov & Konopleva in Bolotov et al., 2017

Radiatula myitkyinae (Prashad, 1930)

Radiatula myitthanensis Bolotov, Konopleva, Vikrev, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Gofarov, Tomilova & Kondakov, 2019

Rectidens lingulatus (Drouet & Chaper, 1892) = Rectidens sumatrensis

+ Unio prolongatus Drouët, 1894

Rectidens sumatrensis (Dunker, 1852)

Reginaia apalachicola (Williams & Fradkin, 1999)

Reginaia ebenus (Lea, 1831)

Reginaia rotulata (B.H. Wright, 1899)

Reticulatus elvae (Walker, 1924) = Arotonaias reticulata

Reticulatus reticulatus (Simpson, 1900) = Arotonaias reticulata

Rhombuniopsis fultoni Moskvicheva & Starobogatov, 1973

Rhombuniopsis superstes (Neumayr, 1899) = Leoparreysia superstes

Rhombuniopsis tauriformis (Fulton, 1906)

Sagittunio aldermani Watters, 2018

Sagittunio nasutus (Say, 1817)

Sagittunio subrostratus (Say, 1831)

Sagittunio vaughanianus (Lea, 1838)

Scabiellus songkramensis (Kongim & Panha in Kongim et al., 2015)

Scabies anceps (Deshayes in Deshayes & Jullien, 1876)

Scabies crispata (Gould, 1843) = Scabies crispata + Nodularia nuxpersicae + Nodularia douglasiae

+ Scabies chinensis Liu et al., 1991

+ Scabies longata Liu et al., 1991

Scabies humilis (Lea, 1856)

Scabies mandarinus (Morelet, 1864)

Scabies nucleus (Lea, 1856)

Scabies phaselus (Lea, 1856)

Scabies pilata (Lea, 1866)

Scabies scobinatus (Lea, 1856)

Schepmania nieuwenhuisi (Schepman, 1898)

Schepmania parcesculpta (Martens, 1903)

Schistodesmus lampreyanus (Baird & Adams, 1867)

Schistodesmus spinosus (Simpson, 1900)

Simpsonaias ambigua (Say, 1825)

Simpsonella gracilis (Lea, 1850)

Simpsonella purpurea (Valenciennes, 1827) = Simpsonella burroughiana

Sinanodonta angula (Tchang, Li & Liu, 1965)

Sinanodonta calipygos (Kobelt, 1879)

Sinanodonta elliptica (Heude, 1878)

Sinanodonta jourdyi (Morlet, 1886)

Sinanodonta lauta (Martens, 1877)

Sinanodonta lucida (Heude, 1877)

Sinanodonta schrenkii (Lea, 1870)

Sinanodonta tumens (Haas, 1910)

+ Oguranodonta ogurae Kuroda & Habe, 1987

Sinanodonta woodiana (Lea, 1834)

Sinohyriopsis cumingii (Lea, 1852)

Sinohyriopsis goliath (Rolle, 1904)

Sinohyriopsis schlegelii (Martens, 1861)

Solenaia carinata (Heude, 1877) = Sinosolenaia carinata

Solenaia emarginata (Lea, 1860)

Solenaia iridinea (Heude, 1874) = Sinosolenaia iridinea

Solenaia khwaenoiensis Panha & Deein in Deein et al., 2004 = Solenaia emarginata

Sphenonaias callosa (Lea, 1841) = Ortmanniana ligamentina

Sphenonaias colorata (Küster, 1856)

Sphenonaias flucki (Bartsch, 1906) = Nephronaias flucki

Sphenonaias liebmanni (Philippi, 1847)

Sphenonaias mexicana (Philippi, 1847)

Sphenonaias microdon (Martens, 1887) = Mesonaias explicata

Strophitus pascagoulaensis Smith, Johnson, Pfeiffer & Gangloff, 2018

Strophitus radiatus (Conrad, 1834)

Strophitus undulatus (Say, 1817)

Strophitus williamsi Smith, Johnson, Pfeiffer & Gangloff, 2018

Sundadontina brandti Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan & Kondakov, 2020

Sundadontina cumingii (Lea, 1851)

+ Pseudodus chaperi Morgan, 1885

Sundadontina harmandi (Crosse & Fischer, 1876)

Sundadontina mabilli (Rochebrune, 1882)

Sundadontina moreleti (Crosse & Fischer, 1876) = Songkhlanaia moreleti

Sundadontina ponderosa (Preston, 1909)

Sundadontina sulcata (Rochebrune, 1882)

Sundadontina tanintharyiensis Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan & Kondakov, 2020

Sundadontina taskaevi Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan & Kondakov, 2020

Sundadontina tumida (Morelet, 1866)

Tamsiella amazonica Bonetto, 1972

Tamsiella schroeteriana (Lea, 1852)

Tamsiella tamsiana (Dunker, 1858)

Thaiconcha callifera (Martens, 1860) = Thaiconcha callifera + Songkhlanaia moreleti

+ Pseudodon thomsoni Morlet, 1884

+ Pseudodon ellipticum Conrad, 1865

Thaiconcha ovalis (Morlet, 1889) = Sundadontina ovalis

Theliderma cylindrica (Say, 1817)

Theliderma intermedia (Conrad, 1836)

Theliderma johnsoni Bogan & Lopes-Lima in Lopes-Lima & Bogan, 2019

Theliderma metanevra (Rafinesque, 1820)

+ Unio tuberosus Lea, 1840

Theliderma sparsa (Lea, 1841)

Theliderma stapes (Lea, 1831)

Toxolasma corvunculus (Lea, 1868)

Toxolasma cylindrellus (Lea, 1868)

Toxolasma lividum Rafinesque, 1831

Toxolasma parvum (Barnes, 1823)

Toxolasma paulum (Lea, 1840)

Toxolasma pullus (Conrad, 1838)

Toxolasma texasiense (Lea, 1857)

+ Lampsilis mearnsi Simpson, 1900

Trapezidens angustior (Hanley & Theobald, 1876)

Trapezidens dolichorhynchus (Tapparone Canefri, 1889)

Trapezidens exolescens (Gould, 1843)

Trapezidens scutum (Sowerby, 1868)

Trapezidens yeti Konopleva, Bolotov, Kondakov, Kononov, Gofarov, Tomilova, Lunn, Chan, Win & Vikhrev, 2020

Trapezoideus foliaceus (Gould, 1843)

Trapezoideus lenya Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan & Kondakov, 2020

Triplodon chodo Mansur & Pimpão, 2008

Triplodon corrugatus (Lamarck, 1819)

+ Triplodon rugosus Spix in Wagner, 1827

Tritogonia nobilis (Conrad, 1854) = Quadrula nobilis

Tritogonia verrucosa (Rafinesque, 1820) = Quadrula verrucosa

Truncilla cognata (Lea, 1860)