	Home \| About \| Database \| Mussel of the Month \| Publications & Presentations

MUSSELpdb

Taxonomy

Species & Genus Richness by
Taxonomy

Geography

Species Richness & Endemism by
Geography

Data last updated
Mon 1 Dec 2025

Publication Chen et al. (2025): summary

publication summary | cladograms

Full Citation

Chen, H., H.-Q. Xiang, Y.-T. Dai, Z.-G. Chen, Y.-M. He et al. 2025. A new genus and two new species of freshwater mussels (Bivalvia, Unionidae) from Sichuan, China: Overlooked cryptic endemism in the upper Yangtze River Basin. Zoosystematics and Evolution 101(4): 1459-1470. Available electronically.

Attributed Nominal Genera

Tongchuania Chen et al., 2025

Chen Xiang, He, Huang & Wu in Chen et al., 2025. Zoosyst. Evol. 101(4): 1464.
Type species: Tongchuania mingiana Chen et al., 2025 (orig. desig.) = Tongchuania mingiana

Attributed Nominal Species

Unio crassus Soroka, 2005 = Unio nanus

‘Philippson’ of Authors.
Soroka, 2005. Zool. Sci. 22: 1138, Table 1.
Soroka, 2010. Folia Malac. 18(4): Table 2. [in part]
Reis et al., 2013. Graellsia 69(1): 22, Table 2.
Prié & Puillandre, 2014. Hydrobiologia 735(1): online resource 1 [as ‘Unio crassus crassus’].
Douda et al., 2014. J. Appl. Ecol. 51: 1087. [in part]
Froufe et al., 2016. Org Divers Evol 16(3): 606, fig. 3. [in part]
Klishko et al., 2017. Zootaxa 4286(1): 98, Table 3.
Soroka & Burzynski, 2017. mtDNA B 2(2): 863.
Burzynski et al., 2017. MPE 107: 606.
Lopes-Lima et al., 2017. Biol. Rev. 92(1): Table S1 [as ‘Unio crassus crassus’]. [in part]
Araujo et al., 2018. J. Linn. Soc., Zool. 182(2): supplementary table. [in part]
Klishko et al., 2018. Systemat. Biodiv. 16(3): 291.
Burzynski & Soroka, 2018. PeerJ 6, e5573: 3.
Wang et al., 2018. Conserv. Genet. Resour. 10(4): 611.
Wang et al., 2018. Conserv. Genet. Resour. 11: 140.
Cha et al., 2018. mtDNA B 3(2): 946.
Aunins et al., 2018. mtDNA B 3(2): 1125.
Wu et al., 2018. PLoS One 13(8), e0200956: 6. [in part]
Klishko et al., 2019. Ruthenica 29(1): 57, Table 1.
Wu et al., 2019. ZooKeys 812: 29.
Huang et al., 2019. MPE 130: Table S5. [in part]
Fukata & Iigo, 2019. mtDNA B 4(2): 3489.
Guerra et al., 2019. BMC Evol. Biol. 19 (229): Table S1.
Froufe et al., 2019. Heredity 124: 185.
Soroka, 2020. J. Zool. Syst. Evol. Res. 58(2): Table 2.
Fukata & Iigo, 2020. mtDNA B 5(2): 1216.
Sano et al., 2020. Plankton Benthos Res. 15(2): 99.
Wu et al., 2020. Invert. Syst. 34: Table S2.
Kilikowska et al., 2020. Life 10(7): 1. [in part]
Lopes-Lima et al., 2021. MPE 163: fig. 6 [by voucher number]. [in part]
Sayenko et al., 2021. Systemat. Biodiv. 19: 58, Table 1.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 2.
Zieritz et al., 2021. Zool. J. Linn. Soc. 193(3): 1092.
Kim et al., 2021. mtDNA B 6(1): 80.
Pfarrer et al., 2022. Hydrobiologia 849: ESM1.
Wu et al., 2022. Zool. Scripta 51: Table S2.
Dai et al., 2023. Invert. Syst. 37(2): Table 2.
Chen et al., 2023. ZooKeys 1173: Table 1.
Wu et al., 2023. Org Divers Evol (2023): Table 2.
Dai et al., 2024. Zool. Scripta 53: Table 1.
Wu et al., 2024. J. Zool. Syst. Evol. Res. 2024, 1499508: Table 2.
Wu et al., 2024. Zool. Scripta 53(5): Table S3.
Dai et al., 2024. Invert. Syst. 38(6): Table 3. [in part]
Chen et al., 2025. Zoosyst. Evol. 101(4): Tables S2-S3.
Dai et al., 2025. Zoosyst. Evol. 101(2): Tables 1-2.
(misidentified reference to Unio crassus Retzius, 1788)

Nodularia douglasiae Choi et al., 2020 = Nodularia dualobtusa

‘Griffith & Pidgeon’ of Authors.
Cha et al., 2018. mtDNA B 3(2): 946. [in part]
Fukata & Iigo, 2019. mtDNA B 4(2): 3488. [in part]
Choi et al., 2020. Sci. Repts. 10: 5 [as ‘Nodularia douglasiae Clade A’]. [in part]
Sayenko et al., 2021. Systemat. Biodiv. 19: 57, Table S2. [in part]
Mabuchi et al., 2021. mtDNA B 6(4): 1536. [in part]
Kim et al., 2021. mtDNA B 6(1): 80. [in part]
Lopes-Lima et al., 2024. MPE 195: Table S2. [in part]
Dai et al., 2024. Invert. Syst. 38(6): fig. 3, Table S1. [in part]
Wu et al., 2024. Invert. Syst. 38, IS24014: Table S1. [in part]
Chen et al., 2025. Zoosyst. Evol. 101(4): Table S2. [in part]
(misidentified reference to Unio douglasiae Griffith & Pidgeon, 1833)

Chamberlainia hainesiana Kongim et al., 2015 = Chamberlainia somsakpanhai

‘Lea’ of Authors.
Kongim et al., 2015. ZooKeys 514: 98, fig. 2A.
Lopes-Lima et al., 2017. MPE 106: 177.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Froufe et al., 2019. Heredity 124: 185.
Huang et al., 2019. MPE 130: Table S4.
Chen et al., 2020. mtDNA B 5(3): 2800.
Teiga-Teixeira et al., 2020. mtDNA B 5(3): 2978.
Zieritz et al., 2020. Biodivers. Conserv. 29: 2244.
Bolotov et al., 2020. Sci. Repts. 10: Table S1.
Lopes-Lima et al., 2020. MPE 146: Table 1.
Zieritz et al., 2021. Zool. J. Linn. Soc. 193(3): Table 1.
Zieritz et al., 2021. Aquatic Conserv.: Mar. Freshw. Ecosyst. 31: 3176.
Jeratthitikul et al., 2021. Eur. J. Taxonomy 775: 125.
Wu et al., 2021. ZooKeys 1054: 87.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
Wu et al., 2022. Zool. Scripta 51: Table S2.
Pavan-Kumar et al., 2022. Mol. Biol. Rep. 49: Table S1.
Jeratthitikul et al., 2022. Sci. Repts. 12: Table S1.
Dai et al., 2023. Invert. Syst. 37(2): Tables 1-2.
Konopleva et al., 2023. Zoosyst. Evol. 99(1): Table S1.
Wu et al., 2023. Org Divers Evol (2023): Table 2.
Dai et al., 2024. Zool. Scripta 53: Table 1.
Dai et al., 2024. Invert. Syst. 38(6): Table 3.
Dai et al., 2024. Zoosyst. Evol. 100(4): Table 2.
Chen et al., 2025. Zoosyst. Evol. 101(4): Table S3.
Dai et al., 2025. Zoosyst. Evol. 101(2): Table 1.
Dai et al., 2025. Zoosyst. Evol. 101(2): Table 2.
(misidentified reference to Unio hainesianus Lea, 1856)

Cuneopsis heudei Lopes-Lima et al., 2017 = Cuneopsis szechenyii

‘Heude’ of Authors.
Liu et al., 2017. PLoS One 12(12 ), e0189737: Table S4.
Lopes-Lima et al., 2017. MPE 106: Table 2.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Wu et al., 2018. PLoS One 13(8), e0200956: 5.
Huang et al., 2019. MPE 130: Tables S1, S4. [in part]
Lopes-Lima et al., 2020. MPE 146: Table 1. [in part]
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1. [in part]
Wu et al., 2022. Zool. Scripta 51: Table S1. [in part]
Jeratthitikul et al., 2022. Sci. Repts. 12: Table S1.
Konopleva et al., 2023. Zoosyst. Evol. 99(1): Table S1.
Chen et al., 2023. ZooKeys 1173: Table 1.
Wu et al., 2023. ZooKeys 1166: Table 1.
Liu et al., 2023. ZooKeys 1179: Table 1.
Chen et al., 2025. Zoosyst. Evol. 101(4): Table S2. [in part]
Dai et al., 2025. Zoosyst. Evol. 101(2): Table 1. [in part]
(misidentified reference to Unio heudei Heude, 1874)

Schistodesmus lampreyanus Huang et al., 2002 = Schistodesmus xinyuensis

‘Baird & Adams’ of Authors.
Huang et al., 2002. J. Moll. Studies 68: Table 1.
Zhang et al., 2004. Acta Hydrobiol. Sinica 28(3): fig. 1.
Zhou et al., 2007. Acta Zool. Sinica 53(6): Table 1.
Szumowski et al., 2012. Amer. Malac. Bull. 30(1): Table S3.
Liu et al., 2017. PLoS One 12(12 ), e0189737: Table S4.
Lopes-Lima et al., 2017. MPE 106: Table 2.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Sano et al., 2017. J. Wat. Res. Prot. 9: Table 2.
Wu et al., 2018. PLoS One 13(8), e0200956: Table 1.
Wu et al., 2019. ZooKeys 812: 29.
Fukata & Iigo, 2019. mtDNA B 4(2): fig. 1.
Huang et al., 2019. MPE 130: Tables S1, S4. [in part]
Fukata & Iigo, 2020. mtDNA B 5(2): fig. 1.
Wu et al., 2020. Invert. Syst. 34: Table S2.
Wu et al., 2020. Malacologia 63(1): Table 1.
Lopes-Lima et al., 2020. MPE 146: Table 1.
Choi et al., 2020. Sci. Repts. 10: Table S5.
Sano et al., 2020. Plankton Benthos Res. 15(2): Table 1.
Kim et al., 2021. mtDNA B 6(1): fig. 1.
Wu et al., 2021. J. Lake Sci. 33(6): attached Tables 1-2.
Wu et al., 2022. Zool. Scripta 51: Tables S1-S2.
Pavan-Kumar et al., 2022. Mol. Biol. Rep. 49: Table S1.
Jeratthitikul et al., 2022. Sci. Repts. 12: Table S1.
Dai et al., 2023. Invert. Syst. 37(2): Table 2.
Wu et al., 2023. Org Divers Evol (2023): Table 2.
Wu et al., 2023. ZooKeys 1166: Table 1.
Liu et al., 2023. ZooKeys 1179: Table 1.
Bogan et al., 2023. Diversity 15(6), 710: Table 1.
Chen et al., 2023. ZooKeys 1173: Table 1.
Dai et al., 2024. Zool. Scripta 53: Tables 1, S2.
Wu et al., 2024. J. Zool. Syst. Evol. Res. 2024, 1499508: Table 2.
Wu et al., 2024. Zool. Scripta 53(5): Tables S2-S3.
Dai et al., 2024. Invert. Syst. 38(6): Tables 2-3.
Hou et al., 2024. Invert. Syst. 38, IS24071: Table 2.
Hou et al., 2025. Zoosyst. Evol. 101(1): Table S2.
Hou et al., 2025. Zoosyst. Evol. 101(1): Table S2.
Dai et al., 2025. Zoosyst. Evol. 101(2): Tables 1-2.
Chen et al., 2025. Zoosyst. Evol. 101(4): Table S3.
(misidentified reference to Unio (Dysnomia) lampreyanus Baird & Adams, 1867)

Tongchuania mingiana Chen et al., 2025 = Tongchuania mingiana

Chen, Xiang, He, Huang & Wu in Chen et al., 2025. Zoosyst. Evol. 101(4): 1464, figs. 4-5.

Pseudocuneopsis perflora Chen et al., 2025 = Pseudocuneopsis perflora

Chen, Xiang, He, Huang & Wu in Chen et al., 2025. Zoosyst. Evol. 101(4): 1464, fig. 6.
Type(s): HOLOTYPE (ORIG. DESIG.) NCUPP_250521, Bahe River, Dazhou City, Sichuan Province, China, 31.0978°N, 107.1856°E.

Taxonomic Opinions

There are 91 taxonomic opinions from this publication. Click here to see the full list.

"Making the world a better place, one mollusk at a time."
	The MUSSEL Project — Home Page https://musselpdb.org/. Site developed and maintained by Dan Graf & Kevin Cummings. Funded by The National Science Foundation.