Publication Campbell & Lydeard (2026): species taxonomic opinions

Campbell, D.C. & C. Lydeard. 2026. Three decades of mussel mitochondria. Malacologia 68(1/2): 237-294. https://doi.org/10.4002/040.068.0109.

Acudonta baitiaoensis Wu, Liu & Zhang in Wu et al., 2025

Acuticosta chinensis (Lea, 1868)

Acuticosta retiaria (Heude, 1883) = Pseudoacuticosta retiaria + Acuticosta ovata

+ Quadrula (Lamprotula) ovata Simpson, 1900

Alasmidonta raveneliana (Lea, 1834)

Alasmidonta uwharriensis Whelan, Perkins & Mays in Whelan et al., 2023

Alasmidonta varicosa (Lamarck, 1819)

Alasmidonta (Bullella) arcula (Lea, 1838) = Alasmidonta arcula + Alasmidonta triangulata

+ Margaritana triangulata Lea, 1858

Alasmidonta (Decurambis) marginata Say, 1818 = Alasmidonta marginata

Alasmidonta (s.s.) undulata (Say, 1817) = Alasmidonta undulata

Alasminota etowaensis (Conrad, 1849)

Alasminota holstonia (Lea, 1838)

Alathyria jacksoni Iredale, 1934

Alathyria pertexta Iredale, 1934

Alathyria profuga (Gould, 1850)

Amblema elliottii (Lea, 1857)

Amblema neislerii (Lea, 1858)

Amblema plicata (Say, 1817)

Amuranodonta kijaensis Moskvicheva, 1973

Anemina arcaeformis (Heude, 1877) = Anemina harlandi

Anodonta anatina (Linnaeus, 1758)

Anodonta cygnea (Linnaeus, 1758)

Anodonta exulcerata Porro, 1838

Anodonta seddoni Gürlek, Kebapçı & Lopes-Lima in Lopes-Lima et al., 2021 = Anodonta seddonae

Anodontites elongata (Swainson, 1823)

Anodontites guanarensis Marshall, 1927

Anodontites trapesialis (Lamarck, 1819)

Anodontites trigona (Spix in Wagner, 1827)

Anodontoides denigratus (Lea, 1852)

Anodontoides ferussacianus (Lea, 1834)

Arcidens confragosus (Say, 1829)

Arcidens wheeleri (Ortmann & Walker, 1912)

Aspatharia pfeifferiana (Bernardi, 1860)

Assamnaia involuta (Hanley, 1856)

Atlanticoncha ochracea (Say, 1817)

Balwantia soleniformis (Benson, 1836)

Beringiana beringiana (Middendorff, 1851)

Beringiana fukuharai San, Hattori & Kondo in Lopes-Lima et al., 2020

Beringiana gosannensis San, Hattori & Kondo in Lopes-Lima et al., 2020

Beringiana japonica (Clessin, 1874)

Bineurus anodontinus (Rochebrune, 1882) = Bineurus anodontinum

Bineurus exilis (Morelet, 1866)

Bineurus loeiensis Konopleva, Bolotov, Pfeiffer, Vikhrev, Kondakov, Gofarov, Tomilova, Tanmuangpak, & Tumpeesuwa, 2021

Bineurus mouhotii (Lea, 1863)

Bineurus panhai Jeratthitikul & Sutcharit, 2023

Brachyanodon californiensis (Lea, 1852)

Brachyanodon impurus (Say, 1829) = Brachyanodon impura

+ Anodonta chapalensis Crosse & Fischer, 1893

Brachyanodon nuttallianus (Lea, 1838) = Brachyanodon nuttalliana

Brachyanodon oregonensis (Lea, 1838)

Buldowskia flavotincta (Martens, 1905)

Buldowskia iwakawai (Suzuki, 1939)

Buldowskia kamiyai San, Hattori & Kondo in Lopes-Lima et al., 2020

Buldowskia shadini (Moskvicheva, 1973)

Buldowskia suifunica (Lindholm, 1925)

Callonaia duprei (Récluz, 1842)

Cambarunio dactylus (Lea, 1840)

Cambarunio hesperus Watters, 2018

Cambarunio iris (Lea, 1829)

Cambarunio jonesii (Lea, 1859)

Cambarunio nebulosus (Conrad, 1834)

Cambarunio simus (Lea, 1838)

Cambarunio taeniatus (Conrad, 1834)

Castalia ambigua Lamarck, 1819

Castalia stevensi (H.B. Baker, 1930)

Castaliella quadrata (G.B. Sowerby II, 1869)

Caudiculatus caudiculatus (Martens, 1867)

Chambardia rubens (Lamarck, 1819)

Chambardia wahlbergi (Krauss, 1848)

Chamberlainia duclerci (Rochebrune, 1882)

Chamberlainia hainesiana (Lea, 1856)

Coelatura aegyptiaca (Cailliaud, 1827)

Coelatura alluaudi (Dautzenberg, 1908)

Coelatura bakeri (H. Adams, 1866)

Coelatura briarti (Dautzenberg, 1901)

Coelatura choziensis (Preston, 1910)

Coelatura choziensis bangweolica Haas, 1936 = Coelatura choziensis

Coelatura gabonensis (Küster, 1862)

Coelatura hauttecoeuri (Bourguignat, 1883)

Coelatura hauttecoeuri kyogae Mandahl-Barth, 1954 = Coelatura hauttecoeuri

Coelatura horei (E.A. Smith, 1880)

Coelatura hypsiprymna (Martens, 1897)

Coelatura kunenensis (Mousson, 1887)

Coelatura leopoldvillensis (Putzeys, 1898)

Coelatura luapulaensis (Preston, 1913)

Coelatura mesafricana Pilsbry & Bequaert, 1927

Coelatura mossambicensis (Martens, 1860)

Coelatura ratidota (Charmes, 1885)

Coelatura stanleyvillensis Pilsbry & Bequaert, 1927

Coelatura stuhlmanni (Martens, 1897)

Cosmopseudodon resupinatus (Martens, 1902)

Cosmopseudodon wenshanensis Wu & Liu in Wu et al., 2024 = Cosmopseudodon resupinatus

Cristaria beirensis Liu & Zhang, 1982 = Cristaria plicata

Cristaria bellua (Morelet, 1866)

Cristaria clessini (Kobelt, 1879)

Cristaria plicata (Leach, 1814)

Cristaria radiata Simpson, 1900

Cristaria truncata Dang, Thai, & Pham, 1980

Ctenodesma bersinara Zieritz, Rahim, Prayogo & Lopes-Lima in Zieritz et al., 2024

Ctenodesma borneensis (Issel, 1874)

Ctenodesma mawonae Zieritz, Rahim, Lopes-Lima & Pfeiffer in Zieritz et al., 2024

Cucumerunio novaehollandiae (Gray, 1834)

Cumberlandia monodonta (Say, 1829)

Cuneopsis celtiformis (Heude, 1874)

Cuneopsis celtiosimilis Hou & Wu in Hou et al., 2025

Cuneopsis demangei Haas, 1929

Cuneopsis heudei (Heude, 1874)

Cuneopsis kiangsiensis Tchang & Li, 1965

Cuneopsis rufescens (Heude, 1874)

Cuneopsis szechenyii (Neumayr, 1899)

Cyclonaias tuberculata (Rafinesque, 1820)

Cyprogenia aberti (Conrad, 1850)

Cyprogenia stegaria (Rafinesque, 1820)

Cyrtonaias tampicoensis (Lea, 1838) = Mesonaias tampicoensis

Diaurora aurorea (Heude, 1883)

Diaurora laeve Chen, Dai, Huang & Wu in Chen et al., 2023

Diplodon chilensis (Gray, 1828)

Diplodon gratus (Lea, 1860)

Diplodon suavidicus (Lea, 1856) = Diplodon suavidicus + Guyanhyrida demeraraensis

+ Unio demeraraensis Lea, 1859

Discomya radulosa (Drouët & Chaper, 1892)

Dromus dromas (Lea, 1834)

Echyridella aucklandica (Gray, 1843)

Echyridella menziesii (Gray, 1843)

Echyridella onekaka Fenwick & Marshall, 2006

Ellipsaria lineolata (Rafinesque, 1820)

Elliptio angustata (Lea, 1831)

Elliptio arca (Conrad, 1834)

Elliptio arctata (Conrad, 1834)

Elliptio complanata (Lightfoot, 1786)

Elliptio congaraea (Lea, 1831)

Elliptio crassidens (Lamarck, 1819)

Elliptio dariensis (Lea, 1842)

Elliptio downiei (Lea, 1858)

Elliptio fisheriana (Lea, 1838)

Elliptio folliculata (Lea, 1838)

Elliptio fraterna (Lea, 1852)

Elliptio fumata (Lea, 1857)

Elliptio hopetonensis (Lea, 1838)

Elliptio icterina (Conrad, 1834)

Elliptio jayensis (Lea, 1838)

Elliptio mcmichaeli Clench & Turner, 1956

Elliptio mediocris (Lea, 1863)

Elliptio nasutidus (Simpson, 1900) [sic]

Elliptio nigella (Lea, 1852)

Elliptio producta (Conrad, 1836)

Elliptio pullata (Lea, 1857)

Elliptio purpurella (Lea, 1857)

Elliptio roanokensis (Lea, 1834)

Elliptio shepardiana (Lea, 1834)

Elliptio spinosa (Lea, 1836)

Elliptoideus sloatianus (Lea, 1840)

Ensidens dugasti (Morlet, 1892)

Ensidens ingallsianus (Lea, 1852)

Ensidens jaculus (Rochebrune, 1882)

Ensidens sagittarius (Lea, 1856)

Ensidens spiculus Pfeiffer, Graf, Cummings, & Page, 2021

Ensidens telus Pfeiffer, Graf, Cummings, & Page, 2021

Epioblasma brevidens (Lea, 1831)

Epioblasma capsaeformis (Lea, 1834)

Epioblasma obliquata (Rafinesque, 1820)

Epioblasma rangiana (Lea, 1838)

Epioblasma triquetra (Rafinesque, 1820)

Epioblasma walkeri (Wilson & Clark, 1914)

Etheria elliptica Lamarck, 1807

Eurynia dilatata (Rafinesque, 1820)

Fossula fossiculifera (d’Orbigny, 1835)

Fusconaia askewi (Marsh, 1896)

Fusconaia burkei (Walker in Ortmann & Walker, 1922)

Fusconaia cerina (Conrad, 1838)

Fusconaia chunii (Lea, 1862)

Fusconaia cor (Conrad, 1834)

Fusconaia cuneolus (Lea, 1840)

Fusconaia escambia Clench & Turner, 1956

Fusconaia flava (Rafinesque, 1820)

Fusconaia hebetata (Conrad, 1854) = Fusconaia flava

Fusconaia iheringi (B.H. Wright, 1898)

Fusconaia masoni (Conrad, 1834)

Fusconaia mitchelli (Simpson in Dall, 1896)

Fusconaia ozarkensis (Call, 1887)

Fusconaia subrotunda (Lea, 1831)

Gibbosula crassa (Wood, 1815)

Gibbosula laosensis (Lea, 1863)

Gibbosula rochechouartii (Heude, 1875)

Gibbosula sinensis Chen, Dai, Chen & Wu in Z.-G. Chen et al., 2025

Glebula rotundata (Lamarck, 1819)

Globunio mirificus Chen, Dai, Huang & Wu in Dai, Chen, Ouyang et al., 2025

Gonidea angulata (Lea, 1838)

Grandidieria burtoni (Woodward, 1859)

Guiunio rarus Chen, Li, Dai & Wu in Dai et al., 2025

Hamiota altilis (Conrad, 1834)

Hamiota australis (Simpson, 1900)

Hamiota perovalis (Conrad, 1834)

Hamiota subangulata (Lea, 1840)

Harmandia somboriensis Rochebrune, 1882

Hemistena lata (Rafinesque, 1820)

Huizhou xietianxiangi H. Chen, Z.-G. Chen, H.-Q. Xiang, Y.-M. He & X.-P. Wu in Chen et al., 2025

Hyridella australis (Lamarck, 1819)

Hyridella depressa (Lamarck, 1819)

Hyridella drapeta (Iredale, 1934)

Hyriopsis bialata Simpson, 1900

Hyriopsis delaportei (Crosse & Fischer, 1876)

Hyriopsis desowitzi Brandt, 1974

Hyriopsis khoratensis Pfeiffer, Graf, Cummings, & Page, 2021

Hyriopsis kratiensis Pfeiffer, Graf, Cummings, & Page, 2021

Hyriopsis myersiana (Lea, 1856)

Hyriopsis panhai Jeratthitikul, Paphatmethin, Zieritz, Lopes-Lima, & Ngor, 2021

Hyriopsis phuphaniensis Pfeiffer, Graf, Cummings, & Page, 2021

Hyriopsis sakhonensis Pfeiffer, Graf, Cummings, & Page, 2021

Indochinella pugio (Benson, 1862)

Indochinella pugio daweiensis Bolotov et al., 2019 = Indochinella pugio

Indochinella pugio paradoxa Bolotov et al., 2019 = Indochinella pugio

Indochinella pugio pugio (Benson, 1862) = Indochinella pugio

Indochinella pugio viridissima Bolotov et al., 2019 = Indochinella pugio

Indonaia andersoniana (Nevill, 1877)

Indonaia bonneaudii (Eydoux, 1838)

Indonaia caerulea (Lea, 1831)

Indonaia corbis (Hanley, 1856)

Indonaia cylindrica Annandale & Prashad, 1919

Indonaia gratiosa (Philippi, 1843)

Indonaia rugosa (Gmelin, 1791)

Indonaia shurtleffiana (Lea, 1856)

Indonaia subclathrata (Martens, 1899)

Indopseudodon bogani Bolotov, Kondakov & Konopleva in Bolotov et al., 2017

Indopseudodon crebristriatus (Anthony, 1865)

Indopseudodon indawgyiensis Bolotov, Konopleva, Kondakov & Vikhrev in Bolotov et al., 2023

Indopseudodon kayinensis Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan, & Kondakov, 2020

Indopseudodon salwenianus (Gould, 1844)

Inversidens brandtii (Kobelt, 1879)

Inversidens rentianensis Wu & Wu in Wu et al., 2021

Inversiunio jokohamensis (Ihering, 1893)

Inversiunio reinianus (Kobelt, 1879)

Inversiunio verrucosus Kondo, Hyun & Seung-Ho, 2007

Inversiunio yanagawensis (Kondo, 1982)

Isannaia fortunata Jeratthitikul, Sutcharit & Prasankok, 2024

Isannaia occultata Jeratthitikul, Sutcharit & Prasankok, 2024

Khairuloconcha lunbawangorum (Issel, 1874) [sic]

Khairuloconcha sahanae Zieritz, Jainih & Lopes-Lima in Zieritz et al., 2021

Koreosolenaia sitgyensis Lee, Kim, Lopes-Lima & Bogan in Lopes-Lima et al., 2020

Lamellidens chindwinensis Bolotov, Konopleva, Chan, Lunn, Win, Gofarov, Kondakov, Tomilova, & Vikhrev, 2022

Lamellidens corrianus (Lea, 1834)

Lamellidens ferrugineus (Annandale, 1918)

Lamellidens generosus (Gould, 1847)

Lamellidens jenkinsianus (Benson, 1862)

Lamellidens lamellatus (Lea, 1838)

Lamellidens mainwaringi Preston, 1912

Lamellidens marginalis (Lamarck, 1819)

Lamellidens nongyangensis Preston, 1912

Lamellidens savadiensis (Nevill, 1877)

Lamproscapha ensiformis (Spix in Wagner, 1827)

Lamprotula caveata (Heude, 1877)

Lamprotula cornuumlunae (Heude, 1883)

Lamprotula leaii (Gray in Griffith & Pidgeon, 1833)

Lamprotula seomjinensis Kondo, Hyun, & Seung-Ho, 2007

Lampsilis bergmanni Inoue & Randklev in Inoue et al., 2020

Lampsilis bracteata (Gould, 1855)

Lampsilis cardium Rafinesque, 1820

Lampsilis cariosa (Say, 1817)

Lampsilis dolabraeformis (Lea, 1838)

Lampsilis fasciola Rafinesque, 1820

Lampsilis hydiana (Lea, 1838)

Lampsilis ornata (Conrad, 1835)

Lampsilis ovata (Say, 1817)

Lampsilis pectorosa (Conrad, 1834) = Ortmanniana pectorosa

Lampsilis powellii (Lea, 1852)

Lampsilis radiata (Gmelin, 1791)

Lampsilis satura (Lea, 1852)

Lampsilis sietmani Keogh & Simons, 2019

Lampsilis siliquoidea (Barnes, 1823)

Lampsilis splendida (Lea, 1838)

Lampsilis straminea (Conrad, 1834)

Lampsilis straminea claibornensis (Lea, 1838) = Lampsilis straminea

Lampsilis straminea contrarius (Conrad, 1849) = Lampsilis straminea

Lampsilis virescens (Lea, 1858)

Lanceolaria acrorrhyncha (Martens, 1894)

Lanceolaria eucylindrica Lin, 1962 = Lanceolaria grayii

Lanceolaria gladiola (Heude, 1877)

Lanceolaria grayii (Griffith & Pidgeon, 1833)

Lanceolaria kihirai Kondo & Hattori, 2019

Lanceolaria lanceolata (Lea, 1856)

Lanceolaria oxyrhyncha (Martens, 1861)

Lanceolaria triformis (Heude, 1877)

Lannanaia kokensis Jeratthitikul, Sutcharit & Prasankok, 2024

Lasmigona (Pterosyna) complanata (Barnes, 1823) = Pterosyna complanata

Lasmigona (s.s.) costata (Rafinesque, 1820) = Lasmigona costata

Leaunio lienosus (Conrad, 1834)

Leaunio lienosus aquilonius Watters, 2018 = Leaunio lienosus

Leaunio lienosus caliginosus (Lea, 1845) = Leaunio lienosus

Leaunio lienosus lienosus (Conrad, 1834) = Leaunio lienosus

Leaunio lienosus unicostatus (B.H. Wright, 1899) = Leaunio lienosus

Leaunio ortmanni (Walker, 1925)

Leaunio pataecus Watters, 2018

Leaunio umbrans (Lea, 1857)

Leaunio vanuxemensis (Lea, 1838)

Leguminaia anatolica Gürlek, Kebapçı & Lopes-Lima in Lopes-Lima et al., 2021

Leguminaia saulcyi (Bourguignat, 1852)