Publication Campbell & Lydeard (2026): species taxonomic opinions

Campbell, D.C. & C. Lydeard. 2026. Three decades of mussel mitochondria. Malacologia 68(1/2): 237-294. https://doi.org/10.4002/040.068.0109.

Leguminaia saulcyi (Bourguignat, 1852)

Leguminaia wheatleyi (Lea, 1862)

Lemiox rimosus (Rafinesque, 1831)

Lens comptus (Deshayes in Deshayes & Jullien, 1876)

Lens contradens (Lea, 1838)

Lens eximius (Lea, 1856)

Lens inornatus (Lea, 1856)

Lens lugens (Drouët & Chaper, 1892)

Lens maenamensis Pfeiffer, Graf, Cummings, & Page, 2021

Lens micropterus (Morelet, 1866)

Lens misellus (Morelet, 1865)

Lens novoselovi Konopleva, Bolotov, Spitsyn, Kondakov, Gofarov, & Vikhrev, 2019

Lens pallegoixi (G.B. Sowerby II, 1867)

Lens rolfbrandti Jeratthitikul & Panha in Jeratthitikul et al., 2019

Leoparreysia burmana (Blanford, 1869)

Leoparreysia canefrii Vikhrev, Bolotov & Kondakov in Bolotov et al., 2017

Leoparreysia olivacea (Prashad, 1930)

Leoparreysia tavoyensis (Gould, 1843)

Leoparreysia whitteni Bolotov, Konopleva, Vikrev, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Gofarov, Tomilova, & Kondakov, 2019

Lepidodesma aligera (Heude, 1877)

Lepidodesma languilati (Heude, 1874)

Ligodonta obscura Perkins, DeVilbiss & Whelan in Perkins et al., 2025

Ligumia recta (Lamarck, 1819)

Lortiella rugata (G.B. Sowerby II, 1868)

Margaritifera dahurica (Middendorff, 1850)

Margaritifera falcata (Gould, 1850)

Margaritifera hembeli (Conrad, 1838)

Margaritifera laevis (Haas, 1910)

Margaritifera margaritifera (Linnaeus, 1758)

Margaritifera marrianae R.I. Johnson, 1983

Margaritifera middendorffi (Rosén, 1926)

Medionidus acutissimus (Lea, 1831)

Medionidus conradicus (Lea, 1834)

Medionidus penicillatus (Lea, 1857)

Megalonaias nervosa (Rafinesque, 1820)

Megalonaias nickliniana (Lea, 1834)

Microcondylaea bonellii (Férussac, 1827)

Microdontia anodontaeformis (Tapparone Canefri, 1883)

Middendorffinaia mongolica (Middendorff, 1851)

Mimo guoliangi Xiang, Lu, He, & Li, 2025

Moncetia anceyi Bourguignat, 1886

Monocondylaea minuana (d’Orbigny, 1835)

Monocondylaea parchappii (d’Orbigny, 1835)

Mulleria rivoli (Deshayes, 1827)

Mutela dubia (Gmelin, 1791)

Mutela hargeri E.A. Smith, 1908

Mutela rostrata (Rang, 1835)

Mycetopoda siliquosa (Spix in Wagner, 1827)

Mytella strigata (Hanley, 1843) = Mytella charruana

Namkongnaia inkhavilayi Jeratthitikul, Sutcharit, Ngor, & Prasankok, 2021

Namkongnaia lemeslei (Morelet, 1875)

Neotrigonia lamarckii (J.E. Gray, 1838)

Neotrigonia margaritacea (Lamarck, 1804)

Nephronaias tempisquensis Pilsbry, 1920

Nitia acuminata (H. Adams, 1866)

Nitia monceti (Bourguignat, 1883)

Nitia teretiuscula (Philippi, 1847)

Nodularia breviconcha Lee, Kim, Bogan & Kondo in Lopes-Lima et al., 2020

Nodularia dorri (Wattebled, 1886)

Nodularia douglasiae (Griffith & Pidgeon, 1833)

Nodularia dualobtusa Wu & Liu in Wu et al., 2024

Nodularia fusiformans Wu & Liu in Wu et al., 2024

Nodularia guiensis Hou & Wu in Hou, Liu et al., 2025

Nodularia hanensis Wu & Liu in Wu et al., 2024

Nodularia huana Wu & Liu in Wu et al., 2024

Nodularia micheloti (Morlet, 1886)

Nodularia nipponensis (Martens, 1877)

Nodularia nuxpersicae (Dunker, 1848)

Nyassunio nyassaensis (Lea, 1864)

Nyeinchanconcha nyeinchani Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan & Kondakov, 2020

Obliquaria reflexa Rafinesque, 1820

Obovalis omiensis (Heimburg, 1884)

Obovaria arkansasensis (Lea, 1862)

Obovaria choctawensis (Athearn, 1964)

Obovaria unicolor (Lea, 1845)

Obovaria (Pseudoon) olivaria (Rafinesque, 1820) = Obovaria olivaria

Obovaria (Rotundaria) subrotunda (Rafinesque, 1820) = Obovaria subrotunda

Obovaria (s.s.) retusa (Lamarck, 1819) = Obovaria retusa

Ortmanniana abrupta (Say, 1831) = Lampsilis abrupta

Ortmanniana higginsii (Lea, 1857) = Lampsilis higginsii

Ortmanniana ligamentina (Lamarck, 1819)

Pachynaias spheniopsis (Morelet, 1849)

Paetulunio fabalis (Lea, 1831)

Parreysia corrugata (Müller, 1774)

Parreysia keralaensis Bolotov, Pasupuleti & Subba Rao in Bolotov et al., 2022

Parreysia marcens (Benson, 1862)

Parreysia nagpoorensis (Lea, 1860)

Parreysia plagiosoma (Benson, 1862)

Parreysia rakhinensis Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan, & Kondakov, 2020

Parvasolenaia rivularis (Heude, 1877)

Parvaspina collina (Conrad, 1837)

Parvaspina lanceolata (Lea, 1828)

Parvaspina steinstansana (Johnson & Clarke, 1983)

Physunio massini (Morelet, 1864)

Physunio modelli Brandt, 1974

Physunio pellucidus Pfeiffer, Graf, Cummings, & Page, 2021

Physunio superbus (Lea, 1843)

Pilsbryoconcha acuta Jeratthitikul & Prasankok in Jeratthitikul et al., 2022

Pilsbryoconcha carinifera (Conrad, 1837)

Pilsbryoconcha compressa (Martens, 1860) = Pilsbryoconcha exilis

Pilsbryoconcha exilis (Lea, 1838)

Pilsbryoconcha hoikaab Jeratthitikul & Prasankok in Jeratthitikul et al., 2022

Pilsbryoconcha kittitati Jeratthitikul & Prasankok in Jeratthitikul et al., 2022

Pilsbryoconcha linguaeformis (Morelet, 1875)

Pilsbryoconcha mekongiana Jeratthitikul & Prasankok in Jeratthitikul et al., 2022

Pilsbryoconcha schomburgki (Martens, 1860)

Platynaias compressa (Lea, 1829)

Platynaias decorata (Lea, 1852)

Platynaias subviridis (Conrad, 1835)

Plectomerus dombeyanus (Valenciennes in Humboldt & Bonpland, 1827)

Plethobasus cooperianus (Lea, 1834)

Plethobasus cyphyus (Rafinesque, 1820)

Pletholophus guangzhouensis Dai, Chen, Huang & Wu in Dai et al., 2024

Pletholophus honglinhensis Bogan, Do, Froufe, & Lopes-Lima, 2023

Pletholophus reinianus (Martens, 1875)

Pletholophus tenuis (Griffith & Pidgeon, 1833)

Pleurobema athearni Gangloff, Williams, & Feminella, 2006

Pleurobema beadleianum (Lea, 1861)

Pleurobema clava (Lamarck, 1819)

Pleurobema cordatum (Rafinesque, 1820)

Pleurobema decisum (Lea, 1831)

Pleurobema georgianum (Lea, 1841)

Pleurobema hanleyianum (Lea, 1852)

Pleurobema oviforme (Conrad, 1834)

Pleurobema perovatum (Conrad, 1834)

Pleurobema plenum (Lea, 1840)

Pleurobema pyriforme (Lea, 1857)

Pleurobema riddellii (Lea, 1862)

Pleurobema rubellum (Conrad, 1834)

Pleurobema sintoxia (Rafinesque, 1820)

Pleurobema stabile (Lea, 1861)

Pleurobema strodeanum (B.H. Wright, 1898)

Pleurobema taitianum (Lea, 1834)

Pleuronaia barnesiana (Lea, 1838)

Pleuronaia dolabelloides (Lea, 1840)

Pleuronaia estabrookianus Lea, 1845 = Pleuronaia estabrookiana

Pleuronaia gibber (Lea, 1838)

Popenaias berezai Inoue, Cummings, Tiemann, Miller, Johnson, Smith, & Randklev, 2020

Popenaias popeii (Lea, 1857)

Popenaias semirasa (Pilsbry, 1910)

Postolata guangxiensis Dai, Huang, Guo & Wu in Dai et al., 2023

Postolata longjiangensis Liu & Wu in Liu et al., 2024

Potamilus alatus (Say, 1817)

Potamilus amphichaenus (Frierson, 1898)

Potamilus capax (Green, 1832)

Potamilus fragilis (Rafinesque, 1820)

Potamilus inflatus (Lea, 1831)

Potamilus leptodon (Rafinesque, 1820)

Potamilus metnecktayi R.I. Johnson, 1998

Potamilus ohiensis (Rafinesque, 1820)

Potamilus purpuratus (Lamarck, 1819)

Potamilus streckersoni Smith, Johnson, Inoue, Doyle, & Randklev, 2019

Potomida acarnanica (Kobelt, 1879)

Potomida littoralis (Cuvier, 1798)

Potomida semirugata (Lamarck, 1819)

Prisodon obliquus Schumacher, 1817

Prisodon syrmatophorus (Gmelin, 1791)

Prisodontopsis aviculaeformis F.R. Woodward, 1991

Prisodontopsis mweruensis (E.A. Smith, 1908)

Prolasmidonta heterodon (Lea, 1829)

Pronodularia japanensis (Lea, 1859)

Pseudanodonta complanata (Rossmässler, 1835)

Pseudocuneopsis capitata (Heude, 1874)

Pseudocuneopsis liuovatus (He & Zhuang, 2013)

+ Pseudocuneopsis heqing Wang & Wu in Wang et al., 2025

Pseudocuneopsis perflora Chen, Xiang, He, Huang & Wu in H. Chen et al., 2025

Pseudocuneopsis sichuanensis Huang, Dai, Chen & Wu in Wu et al., 2022

Pseudocuneopsis wuana Liu & Wu in Liu et al., 2023

Pseudocuneopsis yangshuoensis Wu & Liu in Wu et al., 2023

Pseudocuneopsis yemaoi Dai, Chen, Huang & Wu in Dai et al., 2024

Pseudodon aeneolus Drouët & Chaper, 1892 = Cosmopseudodon aeneolus

Pseudodon cambodjensis (Fischer, 1891) = Pseudodon vondembuschianus

Pseudodon inoscularis (Gould, 1844)

Pseudodon lenyanensis Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan, & Kondakov, 2020

Pseudodon mekongi Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan, & Kondakov, 2020

Pseudodon septuaginta Jeratthitikul, Phuhong et al., 2025

Pseudodon vagulus (Petit, 1865)

Pseudodon vondembuschianus (Lea, 1840)

Pseudodon vondembuschianus laosicus (Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan & Kondakov, 2020) = Pseudodon mekongi

Pseudodon vondembuschianus tapienicus Konopleva, Lheknim, Sriwoon, Kondakov, Vikhrev & Bolotov in Konopleva et al., 2023 = Pseudodon mekongi

Pseudodon vondembuschianus thasaenicus Konopleva, Lheknim, Sriwoon, Kondakov, Vikhrev & Bolotov in Konopleva et al., 2023 = Pseudodon mekongi

Pseudodontoideus connasaugaensis (Lea, 1858)

Pseudodontoideus subvexus (Conrad, 1834)

Pseudomulleria dalyi (E.A. Smith, 1898)

Pseudopostolata angula Dai, Chen, Huang & Wu in Dai et al., 2024

Pseudoscabies scobina (Hanley, 1856)

Pseudospatha tanganyicensis (E.A. Smith, 1880)

Pseudunio auricularius (Spengler, 1793)

Pseudunio homsensis (Lea, 1865)

Pseudunio marocanus (Pallary, 1918)

Psoronaias guatemalensis (Simpson, 1900)

Psoronaias percompressa (Martens, 1887) = Psoronaias crocodilorum

Psoronaias profunda (Simpson, 1914) = Psoronaias crocodilorum

Psoronaias semigranosa (von dem Busch in Philippi, 1843)

Psoronaias usumasintae (Crosse & Fischer, 1892) = Psoronaias crocodilorum

Ptychobranchus fasciolaris (Rafinesque, 1820)

Ptychobranchus greenii (Conrad, 1834)

Ptychorhynchus pfisteri (Heude, 1874)

Pustulosa infucata (Conrad, 1834)

Pustulosa kieneriana (Lea, 1852)

Pustulosa kleiniana (Lea, 1852)

Pustulosa necki (Burlakova, Karatayev, Lopes-Lima & Bogan in Burlakova et al., 2018)

Pustulosa nodulata (Rafinesque, 1820)

Pustulosa petrina (Gould, 1855)

Pustulosa pustulosa (Lea, 1831)

Pustulosa succissa (Lea, 1852)

Pyganodon cataracta (Say, 1817)

Pyganodon fragilis (Lamarck, 1819)

Pyganodon grandis (Say, 1829)

Pyganodon lacustris (Lea, 1857)

Quadrula fragosa (Conrad, 1836)

Quadrula nobilis (Conrad, 1854)

Quadrula quadrula (Rafinesque, 1820)

Quadrula verrucosa (Rafinesque, 1820)

Radiatula chaudhurii (Preston, 1912)

Radiatula chindwinensis Bolotov, Konopleva, Vikrev, Lopes-Lima, Bogan, Lunn, Chan,Win, Aksenova, Gofarov, Tomilova, & Kondakov, 2019

Radiatula crispisulcata (Benson, 1862)

Radiatula kachinensis Bolotov, Konopleva, Chan, Lunn, Win, Gofarov, Kondakov, Tomilova, & Vikhrev, 2022

Radiatula mouhoti Vikhrev, Bolotov & Konopleva in Bolotov et al., 2017

Radiatula mouhoti haungthayawensis Bolotov et al., 2019 = Radiatula mouhoti

Radiatula myitkyinae (Prashad, 1930)

Radiatula myitthanensis Bolotov, Konopleva, Vikrev, Lopes-Lima, Bogan, Lunn, Chan,Win, Aksenova, Gofarov, Tomilova, & Kondakov, 2019

Rectidens lauris Zieritz, Anwari, Diba & Pfeiffer in Zieritz et al., 2024

Rectidens sumatrensis (Dunker, 1852)

Reginaia ebenus (Lea, 1831)

Reginaia rotulata (B.H. Wright, 1899)

Sagittunio nasutus (Say, 1817)

Sagittunio subrostratus (Say, 1831)

Sagittunio vaughanianus (Lea, 1838)

Scabiellus songkramensis (Kongim & Panha in Kongim et al., 2015)

Scabies anceps (Deshayes in Deshayes & Jullien, 1876)

Scabies crispata (Gould, 1843)

Scabies humilis (Lea, 1856)

Scabies mandarinus (Morelet, 1864)

Scabies nucleus (Lea, 1856)

Scabies phaselus (Lea, 1856)

Scabies pilata (Lea, 1866)

Scabies scobinata (Lea, 1856) = Scabies scobinatus

Schepmania nieuwenhuisi (Schepman, 1898)

Schistodesmus lampreyanus (Baird & Adams, 1867)

Schistodesmus luqiaoensis Hou & Wu in Hou et al., 2025

Schistodesmus spinosus (Simpson, 1900)

Schistodesmus tongpenensis Hou & Wu in Hou et al., 2025

Schistodesmus xinyuensis Hou & Wu in Hou et al., 2025

Scriptolamprotula coreana (Martens, 1886)

Scriptolamprotula fibrosa (Heude, 1877)

Scriptolamprotula polysticta (Heude, 1877)

Scriptolamprotula scripta (Heude, 1875)

Scriptolamprotula tientsinensis (Crosse & Debeaux, 1863)

Scriptolamprotula tortuosa (Lea, 1865)

Simpsonaias ambigua (Say, 1825)

Simpsonunio floridensis (Lea, 1852) = Lampsilis floridensis

Simpsonunio teres (Rafinesque, 1820) = Lampsilis teres

+ Unio anodontoides Lea, 1831

Sinanodonta angula (Tchang, Li, & Liu, 1965)

Sinanodonta calipygos (Kobelt, 1879)

Sinanodonta elliptica (Heude, 1878)

Sinanodonta jourdyi (Morlet, 1886)

Sinanodonta kyushuensis Sano & Kondo, 2025 = Sinanodonta lauta

Sinanodonta lauta (Martens, 1877)

Sinanodonta lucida (Heude, 1877)

Sinanodonta pacifica (Heude, 1878)

Sinanodonta plana Sano & Kondo, 2025 = Sinanodonta calipygos

Sinanodonta schrenkii (Lea, 1870)

Sinanodonta tumens (Haas, 1910)

Sinanodonta woodiana (Lea, 1834)

Sinohyriopsis cumingii (Lea, 1852)

Sinohyriopsis goliath (Rolle, 1904)

Sinohyriopsis schlegelii (Martens, 1861)

Sinosolenaia carinata (Heude, 1877)

Sinosolenaia iridinea (Heude, 1874)

Sinosolenaia oleivora (Heude, 1877)

Sinosolenaia recognita (Heude, 1877)

Songkhlanaia moreleti (Crosse & Fischer, 1876)

Songkhlanaia songkhramensis Jeratthitikul, Sutcharit, & Prasankok, 2025

Songkhlanaia tamodienica Konopleva, Lheknim, Sriwoon, Kondakov, Vikhrev & Bolotov in Konopleva et al., 2023

Sphenonaias explicata (Morelet, 1849) = Mesonaias explicata

+ Unio microdon Martens, 1887

Strophitus howellsi Smith, Kiser, Johnson & Randklev, 2023

Strophitus pascagoulaensis Smith, Johnson, Pfeiffer, & Gangloff, 2018

Strophitus radiatus (Conrad, 1834)

Strophitus undulatus (Say, 1817)

Strophitus williamsi Smith, Johnson, Pfeiffer, & Gangloff, 2018

Sulcularia viridis (Rafinesque, 1820)

Sundadontina brandti Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan & Kondakov, 2020

Sundadontina cumingii (Lea, 1851)

Sundadontina harmandi (Crosse & Fischer, 1876)

Sundadontina plugpomenica Konopleva, Lheknim, Sriwoon, Kondakov, Vikhrev & Bolotov in Konopleva et al., 2023

Sundadontina sulcata (Rochebrune, 1882)

Sundadontina tanintharyiensis Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan, & Kondakov, 2020

Sundadontina taskaevi Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan, & Kondakov, 2020

Sundadontina tumida (Morelet, 1866)

Tamsiella schroeteriana (Lea, 1852)

Tamsiella tamsiana (Dunker, 1858)

Tchangsinaia pisciculus (Heude, 1874) = Tchangsinaia piscicula

Thaiconcha callifera (Martens, 1860)

Thaiconcha munelliptica Konopleva, Bolotov, Pfeiffer, Vikhrev, Kondakov, Gofarov, Tomilova, Tanmuangpak, & Tumpeesuwan, 2021