Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Unionini | Aculamprotulina

genus Scriptolamprotula Modell, 1964: cladomics

9 Recent valid species (9 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 9 (9 total)
analyzed in 114 cladograms in 64 publications
maximum number of terminals in an analysis = 6 (mean = 2.68, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 67
maximum number of included terminals = 6, (mean = 3.87 median = 4)
proportion of cladograms that support basic monophyly = 1

Cladograms

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
6 6 5 7 625 0 0 0 
  +----------------- Scriptolamprotula_tortuosa **     
--+                                                 
  |  +-------------- Scriptolamprotula_tientsinensis **
  +--+                                              
     |  +----------- Scriptolamprotula_fibrosa **      
     +--+                                           
        |  +-------- Scriptolamprotula_coreana **      
        +--+                                        
           |  +----- Scriptolamprotula_scripta__2 **   
           +--+                                     
              |  +-- Scriptolamprotula_polysticta **   
              +--+                                  
                 +-- Scriptolamprotula_scripta__1 **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
5 5 3 5 44 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
5 5 3 5 48 0 0 6 
  +----------- Scriptolamprotula_tortuosa **     
--+                                           
  |  +-------- Scriptolamprotula_scripta **      
  +--+                                        
     |  +----- Scriptolamprotula_coreana **      
     +--+                                     
        |  +-- Scriptolamprotula_fibrosa **      
        +--+                                  
           +-- Scriptolamprotula_tientsinensis **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
5 5 3 5 136 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
5 5 3 5 91 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
5 5 3 5 69 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_scripta **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_polysticta **   
  +--+                                     
     |  +-- Scriptolamprotula_coreana **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
5 5 3 5 87 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
5 5 2 5 195 0 0 5 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+  +----- Scriptolamprotula_coreana **      
  |  |                                     
  +--+----- Scriptolamprotula_scripta **      
     |                                     
     |  +-- Scriptolamprotula_fibrosa **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
5 5 3 5 90 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Lopes-Lima et al., 2020 [Publ 4080], fig. 5

iTtTbiriSoTroTroSiD
5 5 3 5 4 0 0 2 
  +----------- Scriptolamprotula_tortuosa **     
--+                                           
  |  +-------- Scriptolamprotula_tientsinensis **
  +--+                                        
     |  +----- Scriptolamprotula_fibrosa **      
     +--+                                     
        |  +-- Scriptolamprotula_scripta **      
        +--+                                  
           +-- Scriptolamprotula_coreana **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
5 5 3 5 69 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_tientsinensis **
  |  +--+                                  
  |  |  +-- Scriptolamprotula_coreana **      
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
5 5 3 5 193 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_fibrosa **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
5 5 3 5 117 0 0 6 
  +----------- Scriptolamprotula_tortuosa **     
--+                                           
  |  +-------- Scriptolamprotula_scripta **      
  +--+                                        
     |  +----- Scriptolamprotula_coreana **      
     +--+                                     
        |  +-- Scriptolamprotula_fibrosa **      
        +--+                                  
           +-- Scriptolamprotula_tientsinensis **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
5 5 3 5 91 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
5 5 2 5 40 0 0 3 
  +-------- Scriptolamprotula_tortuosa **  
  |                                     
--+  +----- Scriptolamprotula_coreana **   
  |  |                                  
  +--+----- Scriptolamprotula_fibrosa **   
     |                                  
     |  +-- Scriptolamprotula_polysticta **
     +--+                               
        +-- Scriptolamprotula_scripta **

Hou et al., 2025 [Publ 4445], fig. 5

iTtTbiriSoTroTroSiD
5 5 3 5 48 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Huang et al., 2019 [Publ 3843], fig. S1

iTtTbiriSoTroTroSiD
5 5 3 5 42 0 0 2 
  +----------- Scriptolamprotula_tortuosa **     
--+                                           
  |  +-------- Scriptolamprotula_tientsinensis **
  +--+                                        
     |  +----- Scriptolamprotula_fibrosa **      
     +--+                                     
        |  +-- Scriptolamprotula_coreana **      
        +--+                                  
           +-- Scriptolamprotula_scripta **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
5 5 3 5 195 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_fibrosa **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
5 5 3 5 69 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_scripta **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_polysticta **   
  +--+                                     
     |  +-- Scriptolamprotula_coreana **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
5 5 3 5 71 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
5 5 3 5 87 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
5 5 3 5 37 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_polysticta **   
  |  +--+                                  
  |  |  +-- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_coreana **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
5 5 3 5 40 0 0 4 
  +-------- Scriptolamprotula_tortuosa **  
  |                                     
--+     +-- Scriptolamprotula_fibrosa **   
  |  +--+                               
  |  |  +-- Scriptolamprotula_coreana **   
  +--+                                  
     |  +-- Scriptolamprotula_scripta **   
     +--+                               
        +-- Scriptolamprotula_polysticta **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
5 5 3 5 64 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_scripta **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_polysticta **   
  +--+                                     
     |  +-- Scriptolamprotula_coreana **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
5 5 3 5 49 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
5 5 3 5 83 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_polysticta **   
  |  +--+                                  
  |  |  +-- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_coreana **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
5 5 3 5 81 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Wang et al., 2025 [Publ 4488], fig. 3B

iTtTbiriSoTroTroSiD
5 5 3 5 60 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
5 5 3 5 40 0 0 6 
  +----------- Scriptolamprotula_tortuosa **     
--+                                           
  |  +-------- Scriptolamprotula_scripta **      
  +--+                                        
     |  +----- Scriptolamprotula_coreana **      
     +--+                                     
        |  +-- Scriptolamprotula_fibrosa **      
        +--+                                  
           +-- Scriptolamprotula_tientsinensis **

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
5 5 3 5 34 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_tientsinensis **
  |  +--+                                  
  |  |  +-- Scriptolamprotula_coreana **      
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
4 4 2 4 52 0 0 5 
  +-------- Scriptolamprotula_tortuosa **     
--+                                        
  |  +----- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_tientsinensis **
     +--+                                  
        +-- Scriptolamprotula_coreana **

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 4 30 0 0 5 
  +-------- Scriptolamprotula_tortuosa **     
--+                                        
  |  +----- Scriptolamprotula_polysticta **   
  +--+                                     
     |  +-- Scriptolamprotula_fibrosa **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 4 42 0 0 5 
  +-------- Scriptolamprotula_tortuosa **     
--+                                        
  |  +----- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_tientsinensis **
     +--+                                  
        +-- Scriptolamprotula_coreana **

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
4 4 2 4 30 0 0 5 
  +-------- Scriptolamprotula_tortuosa **     
--+                                        
  |  +----- Scriptolamprotula_polysticta **   
  +--+                                     
     |  +-- Scriptolamprotula_tientsinensis **
     +--+                                  
        +-- Scriptolamprotula_fibrosa **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 4 52 0 0 5 
  +-------- Scriptolamprotula_tortuosa **     
--+                                        
  |  +----- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_tientsinensis **
     +--+                                  
        +-- Scriptolamprotula_coreana **

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 4 26 0 0 5 
  +-------- Scriptolamprotula_tortuosa **     
--+                                        
  |  +----- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_tientsinensis **
     +--+                                  
        +-- Scriptolamprotula_coreana **

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 4 18 0 0 5 
  +-------- Scriptolamprotula_tortuosa **     
--+                                        
  |  +----- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_coreana **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Wu et al., 2020 [Publ 4147], fig. 4

iTtTbiriSoTroTroSiD
4 4 2 4 10 0 0 5 
  +-------- Scriptolamprotula_tortuosa **     
--+                                        
  |  +----- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_tientsinensis **
     +--+                                  
        +-- Scriptolamprotula_fibrosa **

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 41 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_coreana **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 65 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_coreana **

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 20 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_coreana **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 65 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_coreana **

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
3 3 1 3 41 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_coreana **

Zhou et al., 2007 [Publ 2658], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 15 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_scripta **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 51 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_coreana **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
3 3 1 3 88 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_scripta **

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 35 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_coreana **      
  +--+                                  
     +-- Scriptolamprotula_tientsinensis **

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 37 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_coreana **      
  +--+                                  
     +-- Scriptolamprotula_tientsinensis **

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
3 3 1 3 39 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_coreana **      
  +--+                                  
     +-- Scriptolamprotula_tientsinensis **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 65 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_coreana **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 28 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_scripta **

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 18 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_coreana **      
  +--+                                  
     +-- Scriptolamprotula_tientsinensis **

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 18 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_scripta **

Wu et al., 2024 [Publ 4379], fig. 4_6

iTtTbiriSoTroTroSiD
3 3 1 3 20 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_scripta **      
  +--+                                  
     +-- Scriptolamprotula_tientsinensis **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
3 3 1 3 45 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_coreana **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
3 3 1 3 88 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_scripta **      
  +--+                                  
     +-- Scriptolamprotula_tientsinensis **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
2 2 0 2 37 0 0 -- 
  +-- Scriptolamprotula_tientsinensis **
--+                                  
  +-- Scriptolamprotula_tortuosa **

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 22 0 0 -- 
  +-- Scriptolamprotula_tientsinensis **
--+                                  
  +-- Scriptolamprotula_scripta **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 35 0 0 -- 
  +-- Scriptolamprotula_tientsinensis **
--+                                  
  +-- Scriptolamprotula_tortuosa **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 30 0 0 -- 
  +-- Scriptolamprotula_tortuosa **
--+                             
  +-- Scriptolamprotula_coreana **

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 32 0 0 -- 
  +-- Scriptolamprotula_coreana ** 
--+                             
  +-- Scriptolamprotula_tortuosa **

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 22 0 0 -- 
  +-- Scriptolamprotula_scripta **      
--+                                  
  +-- Scriptolamprotula_tientsinensis **

Zhou et al., 2017 [Publ 3788], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 14 0 0 -- 
  +-- Scriptolamprotula_tientsinensis **
--+                                  
  +-- Scriptolamprotula_tortuosa **

Wen et al., 2017 [Publ 3795], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 14 0 0 -- 
  +-- Scriptolamprotula_coreana ** 
--+                             
  +-- Scriptolamprotula_tortuosa **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 51 0 0 -- 
  +-- Scriptolamprotula_scripta **      
--+                                  
  +-- Scriptolamprotula_tientsinensis **

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 21 0 0 -- 
  +-- Scriptolamprotula_scripta **      
--+                                  
  +-- Scriptolamprotula_tientsinensis **

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 29 0 0 -- 
  +-- Scriptolamprotula_tortuosa **
--+                             
  +-- Scriptolamprotula_coreana **

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Scriptolamprotula_scripta **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
1 1 0 1 75 0 0 -- 
-- Scriptolamprotula_tortuosa **

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Scriptolamprotula_fibrosa **

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Scriptolamprotula_fibrosa **

Xue et al., 2016 [Publ 3659], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Scriptolamprotula_tortuosa **

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Scriptolamprotula_tientsinensis **

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Scriptolamprotula_tortuosa **

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Scriptolamprotula_fibrosa **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
1 1 0 1 137 0 0 -- 
-- Scriptolamprotula_tortuosa **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 123 0 0 -- 
-- Scriptolamprotula_tortuosa **

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Scriptolamprotula_tientsinensis **

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 6 0 0 -- 
-- Scriptolamprotula_tortuosa **

Wu et al., 2022 [Publ 4238], fig. 2e

iTtTbiriSoTroTroSiD
1 1 0 1 17 0 0 -- 
-- Scriptolamprotula_tortuosa **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
1 1 0 1 19 0 0 -- 
-- Scriptolamprotula_tortuosa **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Scriptolamprotula_tortuosa **

Wu et al., 2022 [Publ 4238], fig. 2a_S1

iTtTbiriSoTroTroSiD
1 1 0 1 11 0 0 -- 
-- Scriptolamprotula_tientsinensis **

Dai et al., 2024 [Publ 4386], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Scriptolamprotula_scripta **

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iTtTbiriSoTroTroSiD
1 1 0 1 12 0 0 -- 
-- Scriptolamprotula_tortuosa **

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
1 1 0 1 24 0 0 -- 
-- Scriptolamprotula_tortuosa **

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Scriptolamprotula_tientsinensis **

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iTtTbiriSoTroTroSiD
1 1 0 1 12 0 0 -- 
-- Scriptolamprotula_tortuosa **

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Scriptolamprotula_tortuosa **

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Scriptolamprotula_scripta **

Kim et al., 2021 [Publ 4197], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 17 0 0 -- 
-- Scriptolamprotula_tientsinensis **

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Scriptolamprotula_tortuosa **

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Scriptolamprotula_fibrosa **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
1 1 0 1 75 0 0 -- 
-- Scriptolamprotula_tortuosa **

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 46 0 0 -- 
-- Scriptolamprotula_tortuosa **

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
1 1 0 1 28 0 0 -- 
-- Scriptolamprotula_tientsinensis **

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 16 0 0 -- 
-- Scriptolamprotula_tientsinensis **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Scriptolamprotula_tortuosa **

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Scriptolamprotula_tortuosa **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Scriptolamprotula_tientsinensis **

Ouyang et al., 2015 [Publ 4493], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Scriptolamprotula_tortuosa **

Ouyang et al., 2011 [Publ 3408], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 12 0 0 -- 
-- Scriptolamprotula_tortuosa **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Scriptolamprotula_tortuosa **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
1 1 0 1 79 0 0 -- 
-- Scriptolamprotula_tortuosa **

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 15 0 0 -- 
-- Scriptolamprotula_tientsinensis **

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 0 0 -- 
-- Scriptolamprotula_tortuosa **

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Scriptolamprotula_tortuosa **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 137 0 0 -- 
-- Scriptolamprotula_tortuosa **

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 6 0 0 -- 
-- Scriptolamprotula_tortuosa **

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
1 1 0 1 24 0 0 -- 
-- Scriptolamprotula_tortuosa **

Ouyang et al., 2015 [Publ 4493], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Scriptolamprotula_tortuosa **

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Scriptolamprotula_tientsinensis **

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 22 0 0 -- 
-- Scriptolamprotula_tortuosa **

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
1 1 0 1 16 0 0 -- 
-- Scriptolamprotula_tortuosa **