Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Unionini | Aculamprotulina

genus Scriptolamprotula Modell, 1964: cladomics

9 Recent valid species (9 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 9 (9 total)
analyzed in 116 cladograms in 65 publications
maximum number of terminals in an analysis = 6 (mean = 2.66, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 67
maximum number of included terminals = 6, (mean = 3.87 median = 4)
proportion of cladograms that support basic monophyly = 1

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Scriptolamprotula cladogram distances
PCoA Ordination

NJ Tree of Scriptolamprotula cladogram distances
NJ Tree

Cladograms

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
6 6 5 7 626 0 0 0 
  +----------------- Scriptolamprotula_tortuosa **     
--+                                                 
  |  +-------------- Scriptolamprotula_tientsinensis **
  +--+                                              
     |  +----------- Scriptolamprotula_fibrosa **      
     +--+                                           
        |  +-------- Scriptolamprotula_coreana **      
        +--+                                        
           |  +----- Scriptolamprotula_scripta__2 **   
           +--+                                     
              |  +-- Scriptolamprotula_polysticta **   
              +--+                                  
                 +-- Scriptolamprotula_scripta__1 **

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
5 5 3 5 49 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Hou et al., 2025 [Publ 4445], fig. 5

iTtTbiriSoTroTroSiD
5 5 3 5 48 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
5 5 3 5 64 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_scripta **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_polysticta **   
  +--+                                     
     |  +-- Scriptolamprotula_coreana **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
5 5 3 5 40 0 0 6 
  +----------- Scriptolamprotula_tortuosa **     
--+                                           
  |  +-------- Scriptolamprotula_scripta **      
  +--+                                        
     |  +----- Scriptolamprotula_coreana **      
     +--+                                     
        |  +-- Scriptolamprotula_fibrosa **      
        +--+                                  
           +-- Scriptolamprotula_tientsinensis **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
5 5 3 5 91 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
5 5 3 5 69 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_scripta **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_polysticta **   
  +--+                                     
     |  +-- Scriptolamprotula_coreana **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
5 5 3 5 193 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_fibrosa **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
5 5 3 5 117 0 0 6 
  +----------- Scriptolamprotula_tortuosa **     
--+                                           
  |  +-------- Scriptolamprotula_scripta **      
  +--+                                        
     |  +----- Scriptolamprotula_coreana **      
     +--+                                     
        |  +-- Scriptolamprotula_fibrosa **      
        +--+                                  
           +-- Scriptolamprotula_tientsinensis **

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
5 5 3 5 71 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
5 5 3 5 69 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_tientsinensis **
  |  +--+                                  
  |  |  +-- Scriptolamprotula_coreana **      
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
5 5 3 5 195 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_fibrosa **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
5 5 3 5 44 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
5 5 3 5 40 0 0 4 
  +-------- Scriptolamprotula_tortuosa **  
  |                                     
--+     +-- Scriptolamprotula_fibrosa **   
  |  +--+                               
  |  |  +-- Scriptolamprotula_coreana **   
  +--+                                  
     |  +-- Scriptolamprotula_scripta **   
     +--+                               
        +-- Scriptolamprotula_polysticta **

Huang et al., 2019 [Publ 3843], fig. S1

iTtTbiriSoTroTroSiD
5 5 3 5 42 0 0 2 
  +----------- Scriptolamprotula_tortuosa **     
--+                                           
  |  +-------- Scriptolamprotula_tientsinensis **
  +--+                                        
     |  +----- Scriptolamprotula_fibrosa **      
     +--+                                     
        |  +-- Scriptolamprotula_coreana **      
        +--+                                  
           +-- Scriptolamprotula_scripta **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
5 5 3 5 91 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
5 5 3 5 91 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
5 5 2 5 40 0 0 3 
  +-------- Scriptolamprotula_tortuosa **  
  |                                     
--+  +----- Scriptolamprotula_coreana **   
  |  |                                  
  +--+----- Scriptolamprotula_fibrosa **   
     |                                  
     |  +-- Scriptolamprotula_polysticta **
     +--+                               
        +-- Scriptolamprotula_scripta **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
5 5 3 5 87 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
5 5 3 5 69 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_scripta **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_polysticta **   
  +--+                                     
     |  +-- Scriptolamprotula_coreana **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
5 5 3 5 34 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_tientsinensis **
  |  +--+                                  
  |  |  +-- Scriptolamprotula_coreana **      
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
5 5 3 5 37 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_polysticta **   
  |  +--+                                  
  |  |  +-- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_coreana **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
5 5 3 5 87 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
5 5 3 5 48 0 0 6 
  +----------- Scriptolamprotula_tortuosa **     
--+                                           
  |  +-------- Scriptolamprotula_scripta **      
  +--+                                        
     |  +----- Scriptolamprotula_coreana **      
     +--+                                     
        |  +-- Scriptolamprotula_fibrosa **      
        +--+                                  
           +-- Scriptolamprotula_tientsinensis **

Lopes-Lima et al., 2020 [Publ 4080], fig. 5

iTtTbiriSoTroTroSiD
5 5 3 5 4 0 0 2 
  +----------- Scriptolamprotula_tortuosa **     
--+                                           
  |  +-------- Scriptolamprotula_tientsinensis **
  +--+                                        
     |  +----- Scriptolamprotula_fibrosa **      
     +--+                                     
        |  +-- Scriptolamprotula_scripta **      
        +--+                                  
           +-- Scriptolamprotula_coreana **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
5 5 3 5 83 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_polysticta **   
  |  +--+                                  
  |  |  +-- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_coreana **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
5 5 2 5 195 0 0 5 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+  +----- Scriptolamprotula_coreana **      
  |  |                                     
  +--+----- Scriptolamprotula_scripta **      
     |                                     
     |  +-- Scriptolamprotula_fibrosa **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
5 5 3 5 81 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Wang et al., 2025 [Publ 4488], fig. 3B

iTtTbiriSoTroTroSiD
5 5 3 5 60 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
5 5 3 5 136 0 0 4 
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
4 4 2 4 52 0 0 5 
  +-------- Scriptolamprotula_tortuosa **     
--+                                        
  |  +----- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_tientsinensis **
     +--+                                  
        +-- Scriptolamprotula_coreana **

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 4 18 0 0 5 
  +-------- Scriptolamprotula_tortuosa **     
--+                                        
  |  +----- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_coreana **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Wu et al., 2020 [Publ 4147], fig. 4

iTtTbiriSoTroTroSiD
4 4 2 4 10 0 0 5 
  +-------- Scriptolamprotula_tortuosa **     
--+                                        
  |  +----- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_tientsinensis **
     +--+                                  
        +-- Scriptolamprotula_fibrosa **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 4 42 0 0 5 
  +-------- Scriptolamprotula_tortuosa **     
--+                                        
  |  +----- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_tientsinensis **
     +--+                                  
        +-- Scriptolamprotula_coreana **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 4 52 0 0 5 
  +-------- Scriptolamprotula_tortuosa **     
--+                                        
  |  +----- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_tientsinensis **
     +--+                                  
        +-- Scriptolamprotula_coreana **

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
4 4 2 4 30 0 0 5 
  +-------- Scriptolamprotula_tortuosa **     
--+                                        
  |  +----- Scriptolamprotula_polysticta **   
  +--+                                     
     |  +-- Scriptolamprotula_tientsinensis **
     +--+                                  
        +-- Scriptolamprotula_fibrosa **

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 4 26 0 0 5 
  +-------- Scriptolamprotula_tortuosa **     
--+                                        
  |  +----- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_tientsinensis **
     +--+                                  
        +-- Scriptolamprotula_coreana **

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 4 30 0 0 5 
  +-------- Scriptolamprotula_tortuosa **     
--+                                        
  |  +----- Scriptolamprotula_polysticta **   
  +--+                                     
     |  +-- Scriptolamprotula_fibrosa **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Zhou et al., 2007 [Publ 2658], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 15 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_scripta **

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 41 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_coreana **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 65 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_coreana **

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
3 3 1 3 41 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_coreana **

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 20 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_coreana **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 28 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_scripta **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 51 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_coreana **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
3 3 1 3 88 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_scripta **

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 35 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_coreana **      
  +--+                                  
     +-- Scriptolamprotula_tientsinensis **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
3 3 1 3 45 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_coreana **

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 37 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_coreana **      
  +--+                                  
     +-- Scriptolamprotula_tientsinensis **

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
3 3 1 3 39 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_coreana **      
  +--+                                  
     +-- Scriptolamprotula_tientsinensis **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 65 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_coreana **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
3 3 1 3 88 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_scripta **      
  +--+                                  
     +-- Scriptolamprotula_tientsinensis **

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 18 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_coreana **      
  +--+                                  
     +-- Scriptolamprotula_tientsinensis **

Wu et al., 2024 [Publ 4379], fig. 4_6

iTtTbiriSoTroTroSiD
3 3 1 3 20 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_scripta **      
  +--+                                  
     +-- Scriptolamprotula_tientsinensis **

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 18 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_scripta **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 65 0 0 4 
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_coreana **

Wen et al., 2017 [Publ 3795], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 14 0 0 -- 
  +-- Scriptolamprotula_coreana ** 
--+                             
  +-- Scriptolamprotula_tortuosa **

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 29 0 0 -- 
  +-- Scriptolamprotula_tortuosa **
--+                             
  +-- Scriptolamprotula_coreana **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 30 0 0 -- 
  +-- Scriptolamprotula_tortuosa **
--+                             
  +-- Scriptolamprotula_coreana **

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 21 0 0 -- 
  +-- Scriptolamprotula_scripta **      
--+                                  
  +-- Scriptolamprotula_tientsinensis **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
2 2 0 2 37 0 0 -- 
  +-- Scriptolamprotula_tientsinensis **
--+                                  
  +-- Scriptolamprotula_tortuosa **

Zhou et al., 2017 [Publ 3788], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 14 0 0 -- 
  +-- Scriptolamprotula_tientsinensis **
--+                                  
  +-- Scriptolamprotula_tortuosa **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 51 0 0 -- 
  +-- Scriptolamprotula_scripta **      
--+                                  
  +-- Scriptolamprotula_tientsinensis **

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 32 0 0 -- 
  +-- Scriptolamprotula_coreana ** 
--+                             
  +-- Scriptolamprotula_tortuosa **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 35 0 0 -- 
  +-- Scriptolamprotula_tientsinensis **
--+                                  
  +-- Scriptolamprotula_tortuosa **

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 22 0 0 -- 
  +-- Scriptolamprotula_tientsinensis **
--+                                  
  +-- Scriptolamprotula_scripta **

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 22 0 0 -- 
  +-- Scriptolamprotula_scripta **      
--+                                  
  +-- Scriptolamprotula_tientsinensis **

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 22 0 0 -- 
-- Scriptolamprotula_tortuosa **

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 15 0 0 -- 
-- Scriptolamprotula_tientsinensis **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 137 0 0 -- 
-- Scriptolamprotula_tortuosa **

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iTtTbiriSoTroTroSiD
1 1 0 1 12 0 0 -- 
-- Scriptolamprotula_tortuosa **

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iTtTbiriSoTroTroSiD
1 1 0 1 12 0 0 -- 
-- Scriptolamprotula_tortuosa **

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Scriptolamprotula_tientsinensis **

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Scriptolamprotula_fibrosa **

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
1 1 0 1 28 0 0 -- 
-- Scriptolamprotula_tientsinensis **

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Scriptolamprotula_tortuosa **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Scriptolamprotula_tientsinensis **

Xue et al., 2016 [Publ 3659], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Scriptolamprotula_tortuosa **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
1 1 0 1 137 0 0 -- 
-- Scriptolamprotula_tortuosa **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
1 1 0 1 79 0 0 -- 
-- Scriptolamprotula_tortuosa **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
1 1 0 1 19 0 0 -- 
-- Scriptolamprotula_tortuosa **

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Scriptolamprotula_tortuosa **

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Scriptolamprotula_tientsinensis **

Klymus et al., 2025 [Publ 4814], fig. 3B

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Scriptolamprotula_tortuosa **

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Scriptolamprotula_tortuosa **

Wu et al., 2022 [Publ 4238], fig. 2e

iTtTbiriSoTroTroSiD
1 1 0 1 17 0 0 -- 
-- Scriptolamprotula_tortuosa **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 123 0 0 -- 
-- Scriptolamprotula_tortuosa **

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Scriptolamprotula_tortuosa **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Scriptolamprotula_tortuosa **

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 6 0 0 -- 
-- Scriptolamprotula_tortuosa **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
1 1 0 1 75 0 0 -- 
-- Scriptolamprotula_tortuosa **

Ouyang et al., 2015 [Publ 4493], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Scriptolamprotula_tortuosa **

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Scriptolamprotula_tientsinensis **

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Scriptolamprotula_scripta **

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
1 1 0 1 24 0 0 -- 
-- Scriptolamprotula_tortuosa **

Wu et al., 2022 [Publ 4238], fig. 2a_S1

iTtTbiriSoTroTroSiD
1 1 0 1 11 0 0 -- 
-- Scriptolamprotula_tientsinensis **

Kim et al., 2021 [Publ 4197], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 17 0 0 -- 
-- Scriptolamprotula_tientsinensis **

Dai et al., 2024 [Publ 4386], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Scriptolamprotula_scripta **

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Scriptolamprotula_fibrosa **

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
1 1 0 1 24 0 0 -- 
-- Scriptolamprotula_tortuosa **

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Scriptolamprotula_scripta **

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 16 0 0 -- 
-- Scriptolamprotula_tientsinensis **

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Scriptolamprotula_fibrosa **

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Scriptolamprotula_fibrosa **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Scriptolamprotula_tortuosa **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Scriptolamprotula_tortuosa **

Ouyang et al., 2015 [Publ 4493], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Scriptolamprotula_tortuosa **

Ouyang et al., 2011 [Publ 3408], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 12 0 0 -- 
-- Scriptolamprotula_tortuosa **

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 0 0 -- 
-- Scriptolamprotula_tortuosa **

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Scriptolamprotula_tientsinensis **

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 46 0 0 -- 
-- Scriptolamprotula_tortuosa **

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
1 1 0 1 16 0 0 -- 
-- Scriptolamprotula_tortuosa **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
1 1 0 1 75 0 0 -- 
-- Scriptolamprotula_tortuosa **

Klymus et al., 2025 [Publ 4814], fig. 3A

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Scriptolamprotula_tortuosa **

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Scriptolamprotula_tortuosa **

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 6 0 0 -- 
-- Scriptolamprotula_tortuosa **