MUSSELpdb
Taxonomy
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freshwater mussels
by Taxonomy
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freshwater bivalves
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freshwater mussels
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Data last updated
Tue 02 June 2026
Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Unionini | Aculamprotulina
genus Scriptolamprotula Modell, 1964: cladomics
9 Recent valid species (9 total species)
Cladomics pages are experimental — even more so than the rest of this site!
The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.
The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.
Following the summary, the available published clades representing the taxon are listed.
For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!
Cladomics Summary
number of species = 9 (9 total)
analyzed in 119 cladograms in 67 publications
maximum number of terminals in an analysis = 6 (mean = 2.5, median = 2)
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 70
maximum number of included terminals = 6, (mean = 3.56 median = 4)
proportion of cladograms that support basic monophyly = 1
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Cladograms
Chen et al., 2026 [Publ 4827], fig. 2
** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 6 | 6 | 4 | 6 | 6 | 0 | 0 | ** |
+-- Huizhou
+--------------+
| +-- Globunio
--+
| +-------------- Scriptolamprotula_tortuosa **
+--+
| +----------- Scriptolamprotula_tientsinensis **
+--+
| +-------- Scriptolamprotula_fibrosa **
+--+
| +----- Scriptolamprotula_scripta **
+--+
| +-- Scriptolamprotula_coreana **
+--+
+-- Scriptolamprotula_polysticta **
Wu et al., 2018 [Publ 3836], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 5 | 48 | 0 | 0 | 5.0 |
+----------- Scriptolamprotula_tortuosa **
+--+
| | +-------- Scriptolamprotula_scripta **
| +--+
| | +----- Scriptolamprotula_coreana **
| +--+
--+ | +-- Scriptolamprotula_fibrosa **
| +--+
| +-- Scriptolamprotula_tientsinensis **
|
| +-------- Unio
| |
+-----+ +-- Nodularia
| +--+
| | +-- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Huang et al., 2019 [Publ 3843], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 2 | 5 | 196 | 0 | 0 | 4.0 |
+-------- Scriptolamprotula_tortuosa **
|
+-----+ +----- Scriptolamprotula_coreana **
| | |
| +--+----- Scriptolamprotula_scripta **
| |
--+ | +-- Scriptolamprotula_fibrosa **
| +--+
| +-- Scriptolamprotula_tientsinensis **
|
| +----------- Unio
+--+
| +-------- Schistodesmus
+--+
| +----- Nodularia
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Huang et al., 2019 [Publ 3843], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 5 | 41 | 0 | 0 | 5.0 |
+----------- Scriptolamprotula_tortuosa **
+--------------+
| | +-------- Scriptolamprotula_scripta **
| +--+
| | +----- Scriptolamprotula_coreana **
| +--+
| | +-- Scriptolamprotula_fibrosa **
--+ +--+
| +-- Scriptolamprotula_tientsinensis **
|
| +-- Cristaria
| +--------------------+
| | +-- Lanceolaria
+--+
| +-------------------- Lepidodesma
+--+
| +----------------- Schistodesmus
+--+
| +-------------- Nodularia
+--+
| +----------- Cuneopsis_rufescens
+--+
| +-------- Tchangsinaia
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_szechenyii
+--+
+-- Cuneopsis_heudei
Huang et al., 2019 [Publ 3843], fig. S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 5 | 42 | 0 | 0 | 1.0 |
+-------- Nodularia
+-----+
| | +----- Tchangsinaia
| +--+
| | +-- Cuneopsis
--+ +--+
| +-- Schistodesmus
|
| +----------- Scriptolamprotula_tortuosa **
+--+
| +-------- Scriptolamprotula_tientsinensis **
+--+
| +----- Scriptolamprotula_fibrosa **
+--+
| +-- Scriptolamprotula_coreana **
+--+
+-- Scriptolamprotula_scripta **
Huang et al., 2019 [Publ 3843], fig. S3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 5 | 194 | 0 | 0 | 3.0 |
+-------- Scriptolamprotula_tortuosa **
|
+-----+ +-- Scriptolamprotula_coreana **
| | +--+
| | | +-- Scriptolamprotula_scripta **
| +--+
--+ | +-- Scriptolamprotula_fibrosa **
| +--+
| +-- Scriptolamprotula_tientsinensis **
|
| +----------- Unio
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Huang et al., 2019 [Publ 3843], fig. S4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 5 | 196 | 0 | 0 | 3.0 |
+----------- Acuticosta | +----------- Alasmidonta | +----------- Anodonta_anatina | +----------- Anemina | +----------- Anodonta_cygnea | +----------- Cristaria | +----------- Brachyanodon | +----------- Lasmigona | +----------- Lepidodesma | +----------- Pseudanodonta | +----------- Pyganodon | +----------- Simpsonaias | +----------- Strophitus | --+----------- Utterbackia | +----------- Schistodesmus | +----------- Unio | +----------- Lanceolaria | | +-- Pletholophus +--------+ | +-- Sinanodonta | | +-------- Scriptolamprotula_tortuosa ** | | +--+ +-- Scriptolamprotula_coreana ** | | +--+ | | | +-- Scriptolamprotula_scripta ** | +--+ | | +-- Scriptolamprotula_fibrosa ** | +--+ | +-- Scriptolamprotula_tientsinensis ** | | +----- Cuneopsis_celtiformis | | | +----- Tchangsinaia | | +-----+----- Cuneopsis_rufescens | +----- Nodularia | | +-- Cuneopsis_szechenyii +--+ +-- Cuneopsis_heudei
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 5 | 117 | 0 | 0 | 5.0 |
+----------- Scriptolamprotula_tortuosa **
+-----+
| | +-------- Scriptolamprotula_scripta **
| +--+
| | +----- Scriptolamprotula_coreana **
| +--+
--+ | +-- Scriptolamprotula_fibrosa **
| +--+
| +-- Scriptolamprotula_tientsinensis **
|
| +-------------- Middendorffinaia
| |
+--+ +----------- Unio
| |
+--+ +-------- Nodularia
| |
+--+ +----- Schistodesmus
| |
+--+----- Cuneopsis
|
| +-- Tchangsinaia
+--+
+-- Inversiunio
Lopes-Lima et al., 2020 [Publ 4080], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 5 | 4 | 0 | 0 | 1.0 |
+-------------- Nodularia --+ | +----------- Scriptolamprotula_tortuosa ** +--+ | +-------- Scriptolamprotula_tientsinensis ** +--+ | +----- Scriptolamprotula_fibrosa ** +--+ | +-- Scriptolamprotula_scripta ** +--+ +-- Scriptolamprotula_coreana **
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 3 | 5 | 627 | 0 | 0 | 1.0 |
+----------- Scriptolamprotula_tortuosa **
+--------+
| | +-------- Scriptolamprotula_tientsinensis **
| +--+
| | +----- Scriptolamprotula_fibrosa **
| +--+
--+ | +-- Scriptolamprotula_scripta **
| +--+
| +-- Scriptolamprotula_coreana **
|
| +----------------- Middendorffinaia
| |
+--+ +-------------- Unio
| |
| | +-- Schistodesmus
+--+ +--------+
| | +-- Nodularia
| |
+--+ +-- Tchangsinaia
| +-----+
| | +-- Inversiunio
+--+
| +----- Pseudocuneopsis
+--+
| +-- Diaurora
+--+
+-- Cuneopsis
Wu et al., 2018 [Publ 4143], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 18 | 0 | 0 | 4.0 |
+-- Unio
+--------+
| +-- Nodularia
--+
| +-------- Scriptolamprotula_tortuosa **
+--+
| +----- Scriptolamprotula_scripta **
+--+
| +-- Scriptolamprotula_coreana **
+--+
+-- Scriptolamprotula_tientsinensis **
Huang et al., 2019 [Publ 3843], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 52 | 0 | 0 | 4.0 |
+-------- Scriptolamprotula_tortuosa **
+--+
| | +----- Scriptolamprotula_scripta **
| +--+
| | +-- Scriptolamprotula_tientsinensis **
--+ +--+
| +-- Scriptolamprotula_coreana **
|
| +-------- Unio
+--+
| +----- Nodularia
+--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus
Wu et al., 2019 [Publ 3864], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 42 | 0 | 0 | 4.0 |
+-------- Scriptolamprotula_tortuosa **
+--------+
| | +----- Scriptolamprotula_scripta **
| +--+
| | +-- Scriptolamprotula_tientsinensis **
--+ +--+
| +-- Scriptolamprotula_coreana **
|
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Wu et al., 2020 [Publ 4121], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 30 | 0 | 0 | 4.0 |
+----------- Unio --+ | +-------- Scriptolamprotula_tortuosa ** +--+ | +----- Scriptolamprotula_scripta ** +--+ | +-- Scriptolamprotula_fibrosa ** +--+ +-- Scriptolamprotula_tientsinensis **
Wu et al., 2020 [Publ 4121], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 44 | 0 | 0 | 4.0 |
+-------- Scriptolamprotula_tortuosa **
+--------+
| | +----- Scriptolamprotula_scripta **
| +--+
| | +-- Scriptolamprotula_coreana **
--+ +--+
| +-- Scriptolamprotula_tientsinensis **
|
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Wu et al., 2020 [Publ 4121], fig. S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 30 | 0 | 0 | 4.0 |
+----------- Unio --+ | +-------- Scriptolamprotula_tortuosa ** +--+ | +----- Scriptolamprotula_scripta ** +--+ | +-- Scriptolamprotula_tientsinensis ** +--+ +-- Scriptolamprotula_fibrosa **
Wu et al., 2020 [Publ 4147], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 26 | 0 | 0 | 4.0 |
+-------- Scriptolamprotula_tortuosa **
+--------+
| | +----- Scriptolamprotula_scripta **
| +--+
| | +-- Scriptolamprotula_tientsinensis **
--+ +--+
| +-- Scriptolamprotula_coreana **
|
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Cuneopsis
+--+
+-- Tchangsinaia
Wu et al., 2020 [Publ 4147], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 10 | 0 | 0 | 4.0 |
+----------- Unio --+ | +-------- Scriptolamprotula_tortuosa ** +--+ | +----- Scriptolamprotula_scripta ** +--+ | +-- Scriptolamprotula_tientsinensis ** +--+ +-- Scriptolamprotula_fibrosa **
Wu et al., 2021 [Publ 4366], fig. 3-BI
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 69 | 0 | 0 | 4.0 |
+-------- Scriptolamprotula_tortuosa **
+--------+
| | +----- Scriptolamprotula_scripta **
| +--+
| | +-- Scriptolamprotula_coreana **
--+ +--+
| +-- Scriptolamprotula_tientsinensis **
|
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Cuneopsis
+--+
+-- Tchangsinaia
Wu et al., 2021 [Publ 4366], fig. 3-ML
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 69 | 0 | 0 | 4.0 |
+-------- Scriptolamprotula_tortuosa **
+--------+
| | +----- Scriptolamprotula_scripta **
| +--+
| | +-- Scriptolamprotula_coreana **
--+ +--+
| +-- Scriptolamprotula_tientsinensis **
|
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Wu et al., 2021 [Publ 4366], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 64 | 0 | 0 | 4.0 |
+-------- Scriptolamprotula_tortuosa **
+--------+
| | +----- Scriptolamprotula_scripta **
| +--+
| | +-- Scriptolamprotula_coreana **
--+ +--+
| +-- Scriptolamprotula_tientsinensis **
|
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wu et al., 2022 [Publ 4238], fig. 2d
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 34 | 0 | 0 | 4.0 |
+-------- Scriptolamprotula_tortuosa **
+-----------+
| | +----- Scriptolamprotula_scripta **
| +--+
| | +-- Scriptolamprotula_tientsinensis **
--+ +--+
| +-- Scriptolamprotula_coreana **
|
| +----------------- Unio
+--+
| +-------------- Pseudocuneopsis
+--+
| +----------- Nodularia
+--+
| +-------- Schistodesmus
+--+
| +----- Cuneopsis
+--+
| +-- Tchangsinaia
+--+
+-- Inversiunio
Wu et al., 2022 [Publ 4238], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 69 | 0 | 0 | 4.0 |
+-------- Scriptolamprotula_tortuosa **
+--------+
| | +----- Scriptolamprotula_scripta **
| +--+
| | +-- Scriptolamprotula_tientsinensis **
--+ +--+
| +-- Scriptolamprotula_coreana **
|
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Konopleva et al., 2023 [Publ 4269], fig. S1_4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 52 | 0 | 0 | 4.0 |
+-------- Scriptolamprotula_tortuosa **
+--+
| | +----- Scriptolamprotula_scripta **
| +--+
| | +-- Scriptolamprotula_tientsinensis **
--+ +--+
| +-- Scriptolamprotula_coreana **
|
| +-------- Unio
+--+
| +----- Nodularia
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 91 | 0 | 0 | 4.0 |
+-------- Scriptolamprotula_tortuosa **
+-----+
| | +----- Scriptolamprotula_scripta **
| +--+
| | +-- Scriptolamprotula_coreana **
--+ +--+
| +-- Scriptolamprotula_tientsinensis **
|
| +----------- Unio
| |
+--+ +-- Nodularia
| +-----+
| | +-- Pseudocuneopsis
+--+
| +----- Tchangsinaia
+--+
| +-- Schistodesmus
+--+
+-- Cuneopsis
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 91 | 0 | 0 | 4.0 |
+-------- Scriptolamprotula_tortuosa **
+-----------+
| | +----- Scriptolamprotula_scripta **
| +--+
| | +-- Scriptolamprotula_coreana **
--+ +--+
| +-- Scriptolamprotula_tientsinensis **
|
| +----------------- Unio
| |
+--+ +-------------- Pseudocuneopsis
| |
+--+ +----------- Nodularia
| |
+--+ +-- Schistodesmus_lampreyanus
| +-----+
| | +-- Cuneopsis
+--+
| +----- Tchangsinaia
+--+
| +-- Schistodesmus_xinyuensis
+--+
+-- Schistodesmus_spinosus
Wu et al., 2024 [Publ 4350], fig. 8a_S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 87 | 0 | 0 | 4.0 |
+-------- Scriptolamprotula_tortuosa **
+-----+
| | +----- Scriptolamprotula_scripta **
| +--+
| | +-- Scriptolamprotula_coreana **
--+ +--+
| +-- Scriptolamprotula_tientsinensis **
|
| +----------- Unio
| |
+--+ +-------- Nodularia
| |
+--+-------- Pseudocuneopsis
|
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wu et al., 2024 [Publ 4350], fig. 8b_S2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 87 | 0 | 0 | 4.0 |
+-------- Scriptolamprotula_tortuosa **
+-----+
| | +----- Scriptolamprotula_scripta **
| +--+
| | +-- Scriptolamprotula_coreana **
--+ +--+
| +-- Scriptolamprotula_tientsinensis **
|
| +----------- Unio
| |
+--+ +-------- Nodularia
| |
+--+-------- Pseudocuneopsis
|
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wu et al., 2024 [Publ 4379], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 49 | 0 | 0 | 4.0 |
+-------- Scriptolamprotula_tortuosa **
+--------+
| | +----- Scriptolamprotula_scripta **
| +--+
| | +-- Scriptolamprotula_coreana **
--+ +--+
| +-- Scriptolamprotula_tientsinensis **
|
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Dai et al., 2024 [Publ 4386], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 37 | 0 | 0 | 4.0 |
+-------- Scriptolamprotula_tortuosa **
+--------+
| | +----- Scriptolamprotula_scripta **
| +--+
| | +-- Scriptolamprotula_coreana **
--+ +--+
| +-- Scriptolamprotula_tientsinensis **
|
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Hou et al., 2024 [Publ 4400], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 71 | 0 | 0 | 4.0 |
+-------- Scriptolamprotula_tortuosa **
+-----------+
| | +----- Scriptolamprotula_scripta **
| +--+
| | +-- Scriptolamprotula_coreana **
--+ +--+
| +-- Scriptolamprotula_tientsinensis **
|
| +----------------- Unio
+--+
| +-------------- Middendorffinaia
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Dai et al., 2025 [Publ 4440], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 1 | 4 | 40 | 0 | 0 | 3.0 |
+----- Scriptolamprotula_tortuosa **
|
+--------------+ +-- Scriptolamprotula_scripta **
| | |
| +--+-- Scriptolamprotula_coreana **
| |
--+ +-- Scriptolamprotula_fibrosa **
|
| +----------------- Middendorffinaia
| |
+--+ +-------------- Nodularia
| |
+--+ +----------- Cuneopsis
| |
| | +-- Diaurora
+--+--------+
| +-- Schistodesmus
|
| +-------- Pseudocuneopsis
+--+
| +----- Unio
+--+
| +-- Tchangsinaia
+--+
+-- Inversiunio
Dai et al., 2025 [Publ 4440], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 40 | 0 | 0 | 4.0 |
+-------- Scriptolamprotula_tortuosa **
+--------+
| | +----- Scriptolamprotula_scripta **
| +--+
| | +-- Scriptolamprotula_fibrosa **
| +--+
--+ +-- Scriptolamprotula_coreana **
|
| +-- Middendorffinaia
| +-----------+
| | +-- Nodularia
| |
+--+ +----------- Cuneopsis
| |
| | +-- Diaurora
+--+--------+
| +-- Schistodesmus
|
| +-------- Pseudocuneopsis
+--+
| +----- Inversiunio
+--+
| +-- Tchangsinaia
+--+
+-- Unio
Dai et al., 2025 [Publ 4440], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 83 | 0 | 0 | 4.0 |
+-------- Scriptolamprotula_tortuosa **
+-----+
| | +----- Scriptolamprotula_scripta **
| +--+
| | +-- Scriptolamprotula_coreana **
--+ +--+
| +-- Scriptolamprotula_tientsinensis **
|
| +----------- Unio
| |
+--+ +-- Nodularia
| +-----+
| | +-- Pseudocuneopsis
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Hou et al., 2025 [Publ 4445], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 48 | 0 | 0 | 4.0 |
+-------- Scriptolamprotula_tortuosa **
+--------+
| | +----- Scriptolamprotula_scripta **
| +--+
| | +-- Scriptolamprotula_coreana **
--+ +--+
| +-- Scriptolamprotula_tientsinensis **
|
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Chen et al., 2025 [Publ 4448], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 81 | 0 | 0 | 4.0 |
+-------- Scriptolamprotula_tortuosa **
+--------+
| | +----- Scriptolamprotula_scripta **
| +--+
| | +-- Scriptolamprotula_coreana **
--+ +--+
| +-- Scriptolamprotula_tientsinensis **
|
| +-------------- Unio
| |
+--+ +----------- Pseudocuneopsis
| |
+--+ +-------- Nodularia
| |
+--+ +-- Tongchuania
| +--+
| | +-- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wang et al., 2025 [Publ 4488], fig. 3B
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 60 | 0 | 0 | 4.0 |
+-------- Scriptolamprotula_tortuosa **
+-----------+
| | +----- Scriptolamprotula_scripta **
| +--+
| | +-- Scriptolamprotula_coreana **
--+ +--+
| +-- Scriptolamprotula_tientsinensis **
|
| +----------------- Unio
+--+
| +-------------- Middendorffinaia
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 136 | 0 | 0 | 4.0 |
+-------- Scriptolamprotula_tortuosa **
+--------+
| | +----- Scriptolamprotula_scripta **
| +--+
| | +-- Scriptolamprotula_coreana **
| +--+
--+ +-- Scriptolamprotula_tientsinensis **
|
| +-------------- Globunio
| |
| | +-- Middendorffinaia
+--+ +--------+
| | +-- Unio
| |
+--+ +-- Tchangsinaia
| +-----+
| | +-- Cuneopsis
+--+
| +----- Pseudocuneopsis
+--+
| +-- Schistodesmus
+--+
+-- Nodularia
Wu et al., 2026 [Publ 4811], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 91 | 0 | 0 | 4.0 |
+-------- Scriptolamprotula_tortuosa **
+-----------+
| | +----- Scriptolamprotula_scripta **
| +--+
| | +-- Scriptolamprotula_coreana **
--+ +--+
| +-- Scriptolamprotula_tientsinensis **
|
| +----------------- Unio
+--+
| +-------------- Middendorffinaia
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wu et al., 2026 [Publ 4825], fig. 6B
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 4 | 4 | 2 | 4 | 61 | 0 | 0 | 4.0 |
+-------- Scriptolamprotula_tortuosa **
+-----------+
| | +----- Scriptolamprotula_scripta **
| +--+
| | +-- Scriptolamprotula_coreana **
--+ +--+
| +-- Scriptolamprotula_tientsinensis **
|
| +----------------- Unio
+--+
| +-------------- Middendorffinaia
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Zhou et al., 2007 [Publ 2658], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 18 | 0 | 0 | 3.0 |
+-- Tchangsinaia
+-----+
| +-- Nodularia
--+
| +----- Scriptolamprotula_tortuosa **
+--+
| +-- Scriptolamprotula_tientsinensis **
+--+
+-- Scriptolamprotula_scripta **
Zhou et al., 2007 [Publ 2658], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 15 | 0 | 0 | 3.0 |
+----- Scriptolamprotula_tortuosa **
+--+
| | +-- Scriptolamprotula_tientsinensis **
| +--+
--+ +-- Scriptolamprotula_scripta **
|
| +----- Acuticosta
+--+
| +-- Nodularia
+--+
+-- Cuneopsis
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 28 | 0 | 0 | 3.0 |
+-- Unio
+-----+
| +-- Nodularia
--+
| +----- Scriptolamprotula_tortuosa **
+--+
| +-- Scriptolamprotula_tientsinensis **
+--+
+-- Scriptolamprotula_scripta **
Wu et al., 2016 [Publ 3711], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 18 | 0 | 0 | 3.0 |
+-- Unio
+-----+
| +-- Nodularia
--+
| +----- Scriptolamprotula_tortuosa **
+--+
| +-- Scriptolamprotula_coreana **
+--+
+-- Scriptolamprotula_tientsinensis **
Soroka & Burzynski, 2017 [Publ 3837], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 20 | 0 | 0 | 3.0 |
+----- Scriptolamprotula_tortuosa **
+-----+
| | +-- Scriptolamprotula_tientsinensis **
| +--+
--+ +-- Scriptolamprotula_coreana **
|
| +-------- Unio
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Nodularia
Huang et al., 2018 [Publ 3828], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 39 | 0 | 0 | 3.0 |
+----- Scriptolamprotula_tortuosa **
+--+
| | +-- Scriptolamprotula_coreana **
| +--+
--+ +-- Scriptolamprotula_tientsinensis **
|
| +----- Unio
+--+
| +-- Tchangsinaia
+--+
+-- Nodularia
Aunins et al., 2018 [Publ 3863], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 37 | 0 | 0 | 3.0 |
+----- Scriptolamprotula_tortuosa **
+--------+
| | +-- Scriptolamprotula_coreana **
| +--+
--+ +-- Scriptolamprotula_tientsinensis **
|
| +----------- Unio
+--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Cha et al., 2018 [Publ 3916], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 35 | 0 | 0 | 3.0 |
+----- Scriptolamprotula_tortuosa **
+-----+
| | +-- Scriptolamprotula_coreana **
| +--+
--+ +-- Scriptolamprotula_tientsinensis **
|
| +-------- Unio
+--+
| +----- Schistodesmus
+--+
| +-- Nodularia
+--+
+-- Tchangsinaia
Froufe et al., 2019 [Publ 3936], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 41 | 0 | 0 | 3.0 |
+----- Scriptolamprotula_tortuosa **
+-----+
| | +-- Scriptolamprotula_tientsinensis **
| +--+
--+ +-- Scriptolamprotula_coreana **
|
| +-------- Unio
+--+
| +----- Nodularia
+--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus
Froufe et al., 2019 [Publ 3936], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 41 | 0 | 0 | 3.0 |
+----- Scriptolamprotula_tortuosa **
+-----+
| | +-- Scriptolamprotula_tientsinensis **
| +--+
--+ +-- Scriptolamprotula_coreana **
|
| +-------- Unio
+--+
| +----- Nodularia
+--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus
Fukata & Iigo, 2020 [Publ 4106], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 51 | 0 | 0 | 3.0 |
+----- Scriptolamprotula_tortuosa **
+--------+
| | +-- Scriptolamprotula_tientsinensis **
| +--+
--+ +-- Scriptolamprotula_coreana **
|
| +----------- Unio
| |
+--+ +-- Cuneopsis
| +-----+
| | +-- Schistodesmus
+--+
| +----- Pseudocuneopsis
+--+
| +-- Nodularia
+--+
+-- Tchangsinaia
Sano et al., 2020 [Publ 4138], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 66 | 0 | 0 | 3.0 |
+----- Scriptolamprotula_tortuosa **
+--------+
| | +-- Scriptolamprotula_tientsinensis **
| +--+
--+ +-- Scriptolamprotula_coreana **
|
| +----------- Unio
+--+
| +-------- Inversiunio
+--+
| +----- Nodularia
+--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus
Sano et al., 2020 [Publ 4138], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 66 | 0 | 0 | 3.0 |
+----- Scriptolamprotula_tortuosa **
+--------+
| | +-- Scriptolamprotula_tientsinensis **
| +--+
--+ +-- Scriptolamprotula_coreana **
|
| +----------- Unio
+--+
| +-------- Inversiunio
+--+
| +----- Nodularia
+--+
| +-- Schistodesmus
+--+
+-- Tchangsinaia
Sano et al., 2020 [Publ 4138], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 66 | 0 | 0 | 3.0 |
+----- Scriptolamprotula_tortuosa **
+--------+
| | +-- Scriptolamprotula_tientsinensis **
| +--+
--+ +-- Scriptolamprotula_coreana **
|
| +----------- Unio
+--+
| +-------- Nodularia
+--+
| +----- Inversiunio
+--+
| +-- Schistodesmus
+--+
+-- Tchangsinaia
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 45 | 0 | 0 | 3.0 |
+----- Scriptolamprotula_tortuosa **
+-----+
| | +-- Scriptolamprotula_tientsinensis **
| +--+
--+ +-- Scriptolamprotula_coreana **
|
| +-------- Unio
+--+
| +----- Nodularia
+--+
| +-- Schistodesmus
+--+
+-- Tchangsinaia
Wu et al., 2021 [Publ 4366], fig. 2_I
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 88 | 0 | 0 | 3.0 |
+----- Scriptolamprotula_tortuosa **
+--------+
| | +-- Scriptolamprotula_scripta **
| +--+
--+ +-- Scriptolamprotula_tientsinensis **
|
| +----------- Unio
+--+
| +-------- Nodularia
+--+
| +----- Cuneopsis
+--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus
Wu et al., 2021 [Publ 4366], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 88 | 0 | 0 | 3.0 |
+----- Scriptolamprotula_tortuosa **
+--------+
| | +-- Scriptolamprotula_tientsinensis **
| +--+
--+ +-- Scriptolamprotula_scripta **
|
| +----------- Unio
+--+
| +-------- Nodularia
+--+
| +----- Cuneopsis
+--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus
Wu et al., 2024 [Publ 4379], fig. 4_6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 20 | 0 | 0 | 3.0 |
+----- Scriptolamprotula_tortuosa **
+--+
| | +-- Scriptolamprotula_scripta **
| +--+
| +-- Scriptolamprotula_tientsinensis **
--+
| +-- Cristaria
| +--+
| | +-- Sinanodonta
+--+
| +-- Schistodesmus
+--+
+-- Nodularia
Fonseca et al., 2016 [Publ 3775], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 29 | 0 | 0 | -- |
+-- Scriptolamprotula_tortuosa **
+--------+
| +-- Scriptolamprotula_coreana **
--+
| +-------- Unio
+--+
| +----- Schistodesmus
+--+
| +-- Nodularia
+--+
+-- Tchangsinaia
Zhou et al., 2017 [Publ 3788], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 14 | 0 | 0 | -- |
+-- Nodularia
+--+
| +-- Tchangsinaia
--+
| +-- Scriptolamprotula_tientsinensis **
+--+
+-- Scriptolamprotula_tortuosa **
Lopes-Lima et al., 2017 [Publ 3792], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 30 | 0 | 0 | -- |
+-- Scriptolamprotula_tortuosa **
+--------+
| +-- Scriptolamprotula_coreana **
--+
| +-------- Unio
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Nodularia
Wen et al., 2017 [Publ 3795], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 14 | 0 | 0 | -- |
+-- Scriptolamprotula_coreana **
+--+
| +-- Scriptolamprotula_tortuosa **
--+
| +-- Lepidodesma
+--+
+-- Unio
Chen et al., 2020 [Publ 4170], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 32 | 0 | 0 | -- |
+----- Lepidodesma --+ | +-- Scriptolamprotula_coreana ** +--+ +-- Scriptolamprotula_tortuosa **
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 35 | 0 | 0 | -- |
+-- Scriptolamprotula_tientsinensis **
+--------+
| +-- Scriptolamprotula_tortuosa **
--+
| +-------- Unio
+--+
| +----- Schistodesmus
+--+
| +-- Nodularia
+--+
+-- Tchangsinaia
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 37 | 0 | 0 | -- |
+----- Lepidodesma --+ | +-- Scriptolamprotula_tientsinensis ** +--+ +-- Scriptolamprotula_tortuosa **
Wu et al., 2023 [Publ 4296], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 21 | 0 | 0 | -- |
+-- Scriptolamprotula_scripta **
+-----------+
| +-- Scriptolamprotula_tientsinensis **
--+
| +----------- Cuneopsis
| |
+--+ +-- Nodularia
| +-----+
| | +-- Schistodesmus
+--+
| +----- Tchangsinaia
+--+
| +-- Inversiunio
+--+
+-- Pseudocuneopsis
Liu et al., 2023 [Publ 4310], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 22 | 0 | 0 | -- |
+-- Scriptolamprotula_scripta **
+-----------+
| +-- Scriptolamprotula_tientsinensis **
--+
| +----------- Cuneopsis
| |
+--+ +-- Nodularia
| +-----+
| | +-- Schistodesmus
+--+
| +----- Tchangsinaia
+--+
| +-- Inversiunio
+--+
+-- Pseudocuneopsis
Liu et al., 2023 [Publ 4310], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 22 | 0 | 0 | -- |
+-------- Unio | | +-- Scriptolamprotula_tientsinensis ** +-----+ | +-- Scriptolamprotula_scripta ** --+ | +----- Cuneopsis | | | | +-- Nodularia | +--+ +--+ +-- Schistodesmus | | +-- Tchangsinaia | | +--+-- Inversiunio | +-- Pseudocuneopsis
Dai et al., 2024 [Publ 4358], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 51 | 0 | 0 | -- |
+-- Scriptolamprotula_scripta **
+--------------+
| +-- Scriptolamprotula_tientsinensis **
--+
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wu et al., 2026 [Publ 4825], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 17 | 0 | 0 | -- |
+----- Lanceolaria --+ | +-- Scriptolamprotula_scripta ** +--+ +-- Scriptolamprotula_tientsinensis **
Rosenberg et al., 1997 [Publ 2720], fig. 2B
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 21 | 0 | 0 | -- |
+-- Gonidea | +-- Pyganodon_grandis | +-- Utterbackia | +-- Pyganodon_cataracta | +-- Lampsilis_straminea | +-- Lampsilis_teres | +-- Megalonaias | +-- Obliquaria | --+-- Theliderma | +-- Quadrula | +-- Uniomerus | +-- Amblema | +-- Elliptio | +-- Fusconaia | +-- Plectomerus | +-- Scriptolamprotula_tortuosa ** | +-- Unio
Ouyang et al., 2011 [Publ 3408], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 12 | 0 | 0 | -- |
+-- Lanceolaria --+ +-- Scriptolamprotula_tortuosa **
Ouyang et al., 2015 [Publ 4493], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 30 | 0 | 0 | -- |
+-- Schistodesmus --+ +-- Scriptolamprotula_tortuosa **
Ouyang et al., 2015 [Publ 4493], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 30 | 0 | 0 | -- |
+-- Scriptolamprotula_tortuosa ** --+ +-- Acuticosta
Xue et al., 2016 [Publ 3659], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- |
+-------- Scriptolamprotula_tortuosa ** --+ | +----- Schistodesmus +--+ | +-- Nodularia +--+ +-- Unio
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 80 | 0 | 0 | -- |
+----------- Scriptolamprotula_tortuosa ** --+ | +-------- Nodularia +--+ | +----- Schistodesmus +--+ | +-- Tchangsinaia +--+ +-- Cuneopsis
Bolotov et al., 2017 [Publ 3761], fig. S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 138 | 0 | 0 | -- |
+----------- Scriptolamprotula_tortuosa ** --+ | +-------- Nodularia +--+ | +----- Cuneopsis +--+ | +-- Tchangsinaia +--+ +-- Schistodesmus
Bolotov et al., 2017 [Publ 3761], fig. S2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 138 | 0 | 0 | -- |
+-------- Scriptolamprotula_tortuosa ** | +-------- Schistodesmus --+ +-------- Nodularia | | +----- Cuneopsis_szechenyii +--+ | +-- Tchangsinaia +--+ +-- Cuneopsis_rufescens
Bolotov et al., 2017 [Publ 3764], fig. S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 124 | 0 | 0 | -- |
+-- Unio_crassus | +-- Unio_pictorum | +-- Unio_tumidus | +-- Scriptolamprotula_tortuosa ** | +-- Cuneopsis_szechenyii --+ +-- Tchangsinaia | +-- Cuneopsis_rufescens | +-- Nodularia_douglasiae | +-- Nodularia_nuxpersicae | +-- Schistodesmus
Guerra et al., 2017 [Publ 3772], fig. 2a
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 24 | 0 | 0 | -- |
+-- Scriptolamprotula_tortuosa ** --+ +-- Unio
Guerra et al., 2017 [Publ 3772], fig. 2b
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 24 | 0 | 0 | -- |
+-------------- Scriptolamprotula_tortuosa ** | --+ +-- Cristaria | +--------+ | | +-- Sinanodonta +--+ | +-------- Anodonta +--+ | +----- Platynaias +--+ | +-- Pyganodon +--+ +-- Utterbackia
Burzynski et al., 2017 [Publ 3774], fig. 1-bottom
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 12 | 0 | 0 | -- |
+-- Scriptolamprotula_tortuosa ** --+ +-- Unio
Burzynski et al., 2017 [Publ 3774], fig. 1-top
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 12 | 0 | 0 | -- |
+-- Scriptolamprotula_tortuosa ** --+ +-- Unio
Araujo et al., 2017 [Publ 3801], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- |
+-- Schistodesmus --+ +-- Scriptolamprotula_tortuosa **
Soroka & Burzynski, 2018 [Publ 3762], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 6 | 0 | 0 | -- |
+-- Unio --+ +-- Scriptolamprotula_tortuosa **
Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 6 | 0 | 0 | -- |
+-- Unio --+ +-- Scriptolamprotula_tortuosa **
Huang et al., 2018 [Publ 3828], fig. 3_5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 28 | 0 | 0 | -- |
+-- Unio --+ +-- Scriptolamprotula_tientsinensis **
Huang et al., 2018 [Publ 3828], fig. 6-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 16 | 0 | 0 | -- |
+-- Scriptolamprotula_tortuosa ** --+ +-- Unio
Wang et al., 2018 [Publ 3846], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 30 | 0 | 0 | -- |
+----------- Scriptolamprotula_tientsinensis ** --+ | +-------- Unio +--+ | +----- Nodularia +--+ | +-- Tchangsinaia +--+ +-- Cuneopsis
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 19 | 0 | 0 | -- |
+-- Scriptolamprotula_tortuosa ** --+ +-- Unio
Burzynski & Soroka, 2018 [Publ 3862], fig. 5b
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 9 | 0 | 0 | -- |
+-- Scriptolamprotula_tortuosa ** --+ +-- Unio
Wang et al., 2018 [Publ 3865], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 30 | 0 | 0 | -- |
+-------- Scriptolamprotula_tientsinensis ** --+ | +----- Unio +--+ | +-- Tchangsinaia +--+ +-- Nodularia
Huang et al., 2019 [Publ 3843], fig. 3-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 22 | 0 | 0 | -- |
+-- Scriptolamprotula_tortuosa ** --+ +-- Unio
Fukata & Iigo, 2019 [Publ 4108], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 16 | 0 | 0 | -- |
+----- Scriptolamprotula_tientsinensis ** --+ | +-- Acuticosta +--+ +-- Lanceolaria
Guerra et al., 2019 [Publ 4168], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 40 | 0 | 0 | -- |
+----- Scriptolamprotula_tortuosa ** --+ | +-- Schistodesmus +--+ +-- Unio
Guerra et al., 2019 [Publ 4168], fig. 1-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 21 | 0 | 0 | -- |
+-- Scriptolamprotula_tortuosa ** --+ +-- Unio
Guerra et al., 2019 [Publ 4168], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 40 | 0 | 0 | -- |
+-------------------- Scriptolamprotula_tortuosa ** | --+ +-- Schistodesmus | +--------------+ | | +-- Unio +--+ | +-------------- Lanceolaria +--+ | +----------- Sinanodonta +--+ | +-------- Anodonta +--+ | +----- Platynaias +--+ | +-- Pyganodon +--+ +-- Utterbackia
Guerra et al., 2019 [Publ 4168], fig. 2-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 21 | 0 | 0 | -- |
+----- Scriptolamprotula_tortuosa ** --+ | +-- Schistodesmus +--+ +-- Unio
Sano et al., 2020 [Publ 4138], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 33 | 0 | 0 | -- |
+-- Scriptolamprotula_tortuosa ** --+ +-- Acuticosta
Kim et al., 2021 [Publ 4197], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 17 | 0 | 0 | -- |
+-------------- Scriptolamprotula_tientsinensis ** | --+ +-- Acuticosta | +--------+ | | +-- Lanceolaria +--+ | +-------- Unio +--+ | +----- Nodularia +--+ | +-- Cuneopsis +--+ +-- Schistodesmus
Sonowal et al., 2021 [Publ 4221], fig. 3a
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- |
+-- Anodonta | +-- Unio_ravoisieri --+ +-- Unio_pictorum | +-- Scriptolamprotula_tientsinensis **
Sonowal et al., 2021 [Publ 4221], fig. 3b
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- |
+-- Scriptolamprotula_tientsinensis ** | +-- Lamprotula | +-- Lampsilis | +-- Potomida | --+-- Unio_pictorum | +-- Anodonta | +-- Unio_ravoisieri | +-- Lamellidens | +-- Elliptio
Sonowal et al., 2021 [Publ 4221], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 15 | 0 | 0 | -- |
+----------- Scriptolamprotula_tientsinensis ** --+ | +-------- Unio_pictorum +--+ | +----- Anodonta +--+ | +-- Unio_ravoisieri +--+ +-- Lamellidens
Wu et al., 2022 [Publ 4238], fig. 2a_S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 11 | 0 | 0 | -- |
+----------------- Scriptolamprotula_tientsinensis ** --+ | +-------------- Nodularia +--+ | +----------- Pseudocuneopsis +--+ | +-------- Schistodesmus +--+ | +----- Cuneopsis +--+ | +-- Tchangsinaia +--+ +-- Inversiunio
Wu et al., 2022 [Publ 4238], fig. 2e
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 17 | 0 | 0 | -- |
+----------------- Scriptolamprotula_tortuosa ** --+ | +-------------- Unio +--+ | +----------- Pseudocuneopsis +--+ | +-------- Nodularia +--+ | +----- Tchangsinaia +--+ | +-- Cuneopsis +--+ +-- Schistodesmus
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 75 | 0 | 0 | -- |
+-------------- Scriptolamprotula_tortuosa ** --+ | +----------- Nodularia +--+ | +-------- Schistodesmus +--+ | +----- Cuneopsis_szechenyii +--+ | +-- Tchangsinaia +--+ +-- Cuneopsis_rufescens
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 75 | 0 | 0 | -- |
+-------------- Scriptolamprotula_tortuosa ** --+ | +----------- Nodularia +--+ | +-------- Schistodesmus +--+ | +----- Cuneopsis_szechenyii +--+ | +-- Tchangsinaia +--+ +-- Cuneopsis_rufescens
Dai et al., 2023 [Publ 4285], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 52 | 0 | 0 | -- |
+-------------- Scriptolamprotula_tientsinensis ** | --+ +----------- Unio | | +--+ +-- Pseudocuneopsis | +-----+ | | +-- Nodularia +--+ | +----- Schistodesmus +--+ | +-- Cuneopsis +--+ +-- Tchangsinaia
Chen et al., 2023 [Publ 4302], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 40 | 0 | 0 | -- |
+----------------- Scriptolamprotula_fibrosa ** | | +-- Middendorffinaia --+ +-----------+ | | +-- Nodularia | | +--+ +----------- Cuneopsis | | | | +-- Diaurora +--+--------+ | +-- Schistodesmus | | +-------- Pseudocuneopsis +--+ | +----- Inversiunio +--+ | +-- Tchangsinaia +--+ +-- Unio
Chen et al., 2023 [Publ 4302], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 40 | 0 | 0 | -- |
+----------------- Scriptolamprotula_fibrosa ** | | +-- Middendorffinaia --+ +-----------+ | | +-- Nodularia | | +--+ +----------- Cuneopsis | | | | +-- Diaurora +--+--------+ | +-- Schistodesmus | | +-------- Pseudocuneopsis +--+ | +----- Inversiunio +--+ | +-- Tchangsinaia +--+ +-- Unio
Dai et al., 2024 [Publ 4386], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- |
+----- Scriptolamprotula_scripta ** --+ | +-- Anodonta +--+ +-- Lepidodesma
Dai et al., 2024 [Publ 4386], fig. 5-BI
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 21 | 0 | 0 | -- |
+-- Scriptolamprotula_scripta ** --+ +-- Lepidodesma
Dai et al., 2024 [Publ 4386], fig. 5-ML
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 21 | 0 | 0 | -- |
+----------------- Scriptolamprotula_scripta ** --+ | +-------------- Unio +--+ | +----------- Middendorffinaia +--+ | +-------- Nodularia +--+ | +----- Cuneopsis +--+ | +-- Inversiunio +--+ +-- Schistodesmus
Wu et al., 2024 [Publ 4388], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 52 | 0 | 0 | -- |
+----------------- Scriptolamprotula_tortuosa ** --+ | +-------------- Unio +--+ | +----------- Pseudocuneopsis +--+ | +-------- Nodularia +--+ | +----- Tchangsinaia +--+ | +-- Cuneopsis +--+ +-- Schistodesmus
Chen et al., 2025 [Publ 4487], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 40 | 0 | 0 | -- |
+----------------- Scriptolamprotula_fibrosa ** | --+ +-------------- Unio | | | | +-- Tchangsinaia +--+ +--------+ | | +-- Inversiunio | | +--+ +-- Cuneopsis | +-----+ | | +-- Pseudocuneopsis +--+ | +----- Nodularia +--+ | +-- Diaurora +--+ +-- Schistodesmus
Chen et al., 2025 [Publ 4487], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 40 | 0 | 0 | -- |
+-------------------- Scriptolamprotula_fibrosa ** | --+ +----------------- Unio | | +--+ +-- Tchangsinaia | +-----------+ | | +-- Inversiunio +--+ | +----------- Pseudocuneopsis +--+ | +-------- Cuneopsis +--+ | +----- Nodularia +--+ | +-- Diaurora +--+ +-- Schistodesmus
Klymus et al., 2025 [Publ 4814], fig. 3A
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 35 | 0 | 0 | -- |
+-------------- Scriptolamprotula_tortuosa ** | --+ +----------- Unio | | +--+ +-- Pseudocuneopsis | +-----+ | | +-- Schistodesmus +--+ | +----- Nodularia +--+ | +-- Tchangsinaia +--+ +-- Cuneopsis
Klymus et al., 2025 [Publ 4814], fig. 3B
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 35 | 0 | 0 | -- |
+-- Scriptolamprotula_tortuosa ** --+ +-- Lanceolaria
Campbell & Lydeard, 2026 [Publ 4810], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 46 | 0 | 0 | -- |
+----------------- Scriptolamprotula_tortuosa ** --+ | +-------------- Unio +--+ | +----------- Pseudocuneopsis +--+ | +-------- Nodularia +--+ | +----- Tchangsinaia +--+ | +-- Cuneopsis +--+ +-- Schistodesmus
