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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Gonideinae | Lamprotulini

genus Pronodularia Starobogatov, 1970: cladomics
     1 Recent valid species (1 total species)

Cladomics Summary

number of species = 1 (1 total)
analyzed in 118 cladograms in 71 publications
maximum number of terminals in an analysis = 1 (mean = 1, median = 1)

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
11013042--
  +----- Pronodularia_japanensis **
--+                             
  |  +-- Potomida               
  +--+                          
     +-- Lamprotula

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
11015232--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
110119842--
  +----- Lamprotula             
--+                             
  |  +-- Potomida               
  +--+                          
     +-- Pronodularia_japanensis **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
11014822--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iTtTbiriSoTroTroSiD
11013522--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
11012122--
  +----- Potomida               
--+                             
  |  +-- Lamprotula             
  +--+                          
     +-- Pronodularia_japanensis **

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
11014332--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Huang et al., 2013 [Publ 3629], fig. 3-M

iTtTbiriSoTroTroSiD
1101500--
-- Pronodularia_japanensis **

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
11013932--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
110119842--
  +----- Lamprotula             
--+                             
  |  +-- Potomida               
  +--+                          
     +-- Pronodularia_japanensis **

Froufe et al., 2015 [Publ 3664], fig. 1

iTtTbiriSoTroTroSiD
11011122--
  +----- Potomida               
--+                             
  |  +-- Lamprotula             
  +--+                          
     +-- Pronodularia_japanensis **

Guerra et al., 2017 [Publ 3772], fig. 2b-M

iTtTbiriSoTroTroSiD
1101700--
-- Pronodularia_japanensis **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
11016722--
  +----- Lamprotula             
--+                             
  |  +-- Potomida               
  +--+                          
     +-- Pronodularia_japanensis **

Wu et al., 2021 [Publ 4215], fig. 3

iTtTbiriSoTroTroSiD
11012332--
  +----- Lamprotula             
--+                             
  |  +-- Potomida               
  +--+                          
     +-- Pronodularia_japanensis **

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
11013911--
  +-- Pronodularia_japanensis **
--+                          
  +-- Lamprotula

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
11011322--
  +----- Potomida               
--+                             
  |  +-- Lamprotula             
  +--+                          
     +-- Pronodularia_japanensis **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
11013122--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
11019442--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
11013922--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Froufe et al., 2019 [Publ 3936], fig. 3-M

iTtTbiriSoTroTroSiD
11011811--
  +-- Potomida               
--+                          
  +-- Pronodularia_japanensis **

Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3

iTtTbiriSoTroTroSiD
1101300--
-- Pronodularia_japanensis **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
11013521--
  +-- Pronodularia_japanensis **
--+                          
  +-- Potomida

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
11014522--
  +----- Potomida               
--+                             
  |  +-- Lamprotula             
  +--+                          
     +-- Pronodularia_japanensis **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
110113832--
  +----- Lamprotula             
--+                             
  |  +-- Potomida               
  +--+                          
     +-- Pronodularia_japanensis **

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
11012400--
-- Pronodularia_japanensis **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
110112042--
  +----- Pronodularia_japanensis **
--+                             
  |  +-- Potomida               
  +--+                          
     +-- Lamprotula

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iTtTbiriSoTroTroSiD
11015722--
  +----- Pronodularia_japanensis **
--+                             
  |  +-- Lamprotula             
  +--+                          
     +-- Potomida

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
11017232--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Soroka, 2005 [Publ 2726], fig. 3

iTtTbiriSoTroTroSiD
1101400--
-- Pronodularia_japanensis **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

iTtTbiriSoTroTroSiD
1101911--
  +-- Potomida               
--+                          
  +-- Pronodularia_japanensis **

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
11014432--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Matos et al., 2024 [Publ 4383], fig. 3

iTtTbiriSoTroTroSiD
11011942--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iTtTbiriSoTroTroSiD
11011222--
  +----- Potomida               
--+                             
  |  +-- Lamprotula             
  +--+                          
     +-- Pronodularia_japanensis **

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
11012011--
  +-- Lamprotula             
--+                          
  +-- Pronodularia_japanensis **

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
11013311--
  +-- Lamprotula             
--+                          
  +-- Pronodularia_japanensis **

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
11012111--
  +-- Pronodularia_japanensis **
--+                          
  +-- Lamprotula

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
110119642--
  +----- Lamprotula             
--+                             
  |  +-- Potomida               
  +--+                          
     +-- Pronodularia_japanensis **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
11015532--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iTtTbiriSoTroTroSiD
11011222--
  +----- Lamprotula             
--+                             
  |  +-- Potomida               
  +--+                          
     +-- Pronodularia_japanensis **

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
110111742--
-- Pronodularia_japanensis **

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
11012831--
  +-- Pronodularia_japanensis **
--+                          
  +-- Lamprotula

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
11014632--
  +----- Lamprotula             
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Potomida

Sano et al., 2017 [Publ 3799], fig. 2

iTtTbiriSoTroTroSiD
11013211--
  +-- Pronodularia_japanensis **
--+                          
  +-- Lamprotula

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
11015342--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
11017232--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
11014022--
  +----- Pronodularia_japanensis **
--+                             
  |  +-- Lamprotula             
  +--+                          
     +-- Potomida

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
110113832--
  +----- Lamprotula             
--+                             
  |  +-- Potomida               
  +--+                          
     +-- Pronodularia_japanensis **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
11017932--
  +----- Lamprotula             
--+                             
  |  +-- Potomida               
  +--+                          
     +-- Pronodularia_japanensis **

Jeratthitikul et al., 2025 [Publ 4423], fig. 1

iTtTbiriSoTroTroSiD
11014922--
  +----- Potomida               
--+                             
  |  +-- Lamprotula             
  +--+                          
     +-- Pronodularia_japanensis **

Campbell et al., 2005 [Publ 2660], fig. 1

iTtTbiriSoTroTroSiD
110110111--
  +-- Potomida               
--+                          
  +-- Pronodularia_japanensis **

Plazzi et al., 2011 [Publ 3676], fig. 5

iTtTbiriSoTroTroSiD
11011700--
-- Pronodularia_japanensis **

Jiang et al., 2010 [Publ 3438], fig. 5

iTtTbiriSoTroTroSiD
1101300--
-- Pronodularia_japanensis **

Amaro et al., 2017 [Publ 3791], fig. 3B

iTtTbiriSoTroTroSiD
11011100--
-- Pronodularia_japanensis **

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
11012222--
  +----- Potomida               
--+                             
  |  +-- Lamprotula             
  +--+                          
     +-- Pronodularia_japanensis **

Guerra et al., 2017 [Publ 3772], fig. 2a-M

iTtTbiriSoTroTroSiD
1101700--
-- Pronodularia_japanensis **

Kongim et al., 2023 [Publ 4290], fig. 1

iTtTbiriSoTroTroSiD
11011422--
  +----- Lamprotula             
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Potomida

Amaro et al., 2017 [Publ 3791], fig. 3A

iTtTbiriSoTroTroSiD
11011100--
-- Pronodularia_japanensis **

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iTtTbiriSoTroTroSiD
11012400--
-- Pronodularia_japanensis **

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
11013022--
  +----- Potomida               
--+                             
  |  +-- Lamprotula             
  +--+                          
     +-- Pronodularia_japanensis **

Zhang et al., 2024 [Publ 4397], fig. 5

iTtTbiriSoTroTroSiD
11012742--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
11011622--
  +----- Potomida               
--+                             
  |  +-- Lamprotula             
  +--+                          
     +-- Pronodularia_japanensis **

Amaro et al., 2017 [Publ 3791], fig. 2

iTtTbiriSoTroTroSiD
11011500--
-- Pronodularia_japanensis **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
11013722--
  +----- Potomida               
--+                             
  |  +-- Lamprotula             
  +--+                          
     +-- Pronodularia_japanensis **

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
11014411--
  +-- Pronodularia_japanensis **
--+                          
  +-- Lamprotula

Walker et al., 2006 [Publ 2666], fig. 2

iTtTbiriSoTroTroSiD
11011400--
-- Pronodularia_japanensis **

Froufe et al., 2015 [Publ 3664], fig. 1-M

iTtTbiriSoTroTroSiD
1101711--
  +-- Potomida               
--+                          
  +-- Pronodularia_japanensis **

Amaro et al., 2017 [Publ 3791], fig. 3B-M

iTtTbiriSoTroTroSiD
1101600--
-- Pronodularia_japanensis **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
11017522--
  +----- Potomida               
--+                             
  |  +-- Lamprotula             
  +--+                          
     +-- Pronodularia_japanensis **

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
11014732--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
11012122--
  +----- Potomida               
--+                             
  |  +-- Lamprotula             
  +--+                          
     +-- Pronodularia_japanensis **

Chapman et al., 2008 [Publ 2667], fig. 1

iTtTbiriSoTroTroSiD
11012000--
-- Pronodularia_japanensis **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
11014732--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Amaro et al., 2017 [Publ 3791], fig. 2-M

iTtTbiriSoTroTroSiD
1101900--
-- Pronodularia_japanensis **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
11013521--
  +-- Pronodularia_japanensis **
--+                          
  +-- Potomida

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
11014432--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iTtTbiriSoTroTroSiD
11013522--
  +----- Potomida               
--+                             
  |  +-- Lamprotula             
  +--+                          
     +-- Pronodularia_japanensis **

Froufe et al., 2016 [Publ 3727], fig. 3_4

iTtTbiriSoTroTroSiD
1101642--
-- Pronodularia_japanensis **

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
11012400--
-- Pronodularia_japanensis **

Shirai et al., 2010 [Publ 3353], fig. 4

iTtTbiriSoTroTroSiD
1101200--
-- Pronodularia_japanensis **

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
11014332--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Froufe et al., 2019 [Publ 3936], fig. 4

iTtTbiriSoTroTroSiD
11011811--
  +-- Potomida               
--+                          
  +-- Pronodularia_japanensis **

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
11013911--
  +-- Pronodularia_japanensis **
--+                          
  +-- Lamprotula

Chapman et al., 2008 [Publ 2744], fig. 2

iTtTbiriSoTroTroSiD
11012000--
-- Pronodularia_japanensis **

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
11013732--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
11019442--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
11016722--
  +----- Lamprotula             
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Potomida

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
11019032--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
11012211--
  +-- Pronodularia_japanensis **
--+                          
  +-- Potomida

Sano et al., 2017 [Publ 3799], fig. 1

iTtTbiriSoTroTroSiD
11012000--
-- Pronodularia_japanensis **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
110112432--
-- Pronodularia_japanensis **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
11018732--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

iTtTbiriSoTroTroSiD
11011100--
-- Pronodularia_japanensis **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
11015532--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
11019032--
  +----- Pronodularia_japanensis **
--+                             
  |  +-- Potomida               
  +--+                          
     +-- Lamprotula

Plazzi & Passamonti, 2010 [Publ 3354], fig. 1

iTtTbiriSoTroTroSiD
1101100--
-- Pronodularia_japanensis **

Zhou et al., 2016 [Publ 3710], fig. 1

iTtTbiriSoTroTroSiD
11011911--
  +-- Lamprotula             
--+                          
  +-- Pronodularia_japanensis **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
11016722--
  +----- Lamprotula             
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Potomida

Kano et al., 2019 [Publ 4194], fig. 6

iTtTbiriSoTroTroSiD
1101900--
-- Pronodularia_japanensis **

Breton et al., 2011 [Publ 3376], fig. 3

iTtTbiriSoTroTroSiD
11013600--
-- Pronodularia_japanensis **

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
110111142--
  +-- Lamprotula_leaii       
  |                          
  +-- Lamprotula             
  |                          
--+-- Lamprotula_caveata     
  |                          
  +-- Potomida               
  |                          
  +-- Pronodularia_japanensis **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
11017522--
  +----- Lamprotula             
--+                             
  |  +-- Potomida               
  +--+                          
     +-- Pronodularia_japanensis **

Amaro et al., 2017 [Publ 3791], fig. 3A-M

iTtTbiriSoTroTroSiD
1101600--
-- Pronodularia_japanensis **

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
11013522--
-- Pronodularia_japanensis **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
11015232--
  +----- Lamprotula             
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Potomida

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
11019042--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Matos et al., 2024 [Publ 4384], fig. 3

iTtTbiriSoTroTroSiD
11011942--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Huang et al., 2013 [Publ 3629], fig. 3

iTtTbiriSoTroTroSiD
11011500--
-- Pronodularia_japanensis **

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
11014132--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
11011922--
  +----- Potomida               
--+                             
  |  +-- Lamprotula             
  +--+                          
     +-- Pronodularia_japanensis **

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
11014411--
  +-- Pronodularia_japanensis **
--+                          
  +-- Lamprotula

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
11014022--
  +----- Pronodularia_japanensis **
--+                             
  |  +-- Lamprotula             
  +--+                          
     +-- Potomida

Fonseca et al., 2016 [Publ 3775], fig. 1-M

iTtTbiriSoTroTroSiD
1101811--
  +-- Potomida               
--+                          
  +-- Pronodularia_japanensis **

Campbell et al., 2005 [Publ 2660], fig. 2

iTtTbiriSoTroTroSiD
110110111--
  +-- Potomida               
--+                          
  +-- Pronodularia_japanensis **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
11015122--
  +----- Potomida               
--+                             
  |  +-- Lamprotula             
  +--+                          
     +-- Pronodularia_japanensis **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
11017232--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
11019042--
  +----- Potomida               
--+                             
  |  +-- Pronodularia_japanensis **
  +--+                          
     +-- Lamprotula

Lopes-Lima et al., 2020 [Publ 4080], fig. 16

iTtTbiriSoTroTroSiD
1101731--
  +-- Pronodularia_japanensis **
--+                          
  +-- Lamprotula

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
11018222--
-- Pronodularia_japanensis **
 
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