MUSSELp

MUSSELpdb

Introduction

Browse

d'basing

Search the MUSSELpdb

Taxonomy

all genera

Unionidae

Margaritiferidae

Hyriidae

Etheriidae

Mycetopodidae

Iridinidae


Species & Genus Richness by
Taxonomy

Geography

Summary

North America

Central America

South America

Afrotropics

North Eurasia

East Asia

Australasia


Species Richness & Endemism by
Geography

Data last updated
Mon 1 Dec 2025

Publication Lea (1840): summary

publication summary

Full Citation

Lea, I. 1840. Descriptions of new fresh water and land shells. Proceedings of the American Philosophical Society 1: 284-289. Available electronically.

Attributed Nominal Species

image of type lot

Unio amoenus Lea, 1840 = Cambarunio iris

Lea, 1840. Proc. Amer. Phil. Soc. 1: 286 [as ‘amœnus’].
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 200, pl. 10, fig. 12 [as ‘amœnus’].
Type(s): HOLOTYPE USNM_85070, Holston R., Tenn.

image of type lot

Anodonta argentea Lea, 1840 = Anodontoides denigratus

Lea, 1840. Proc. Amer. Phil. Soc. 1: 289.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 223, pl. 19, fig. 41.
Type(s): HOLOTYPE USNM_86538, Stones R., Tenn.

Unio atromarginatus Lea, 1840 = Elliptoideus sloatianus

Lea, 1840. Proc. Amer. Phil. Soc. 1: 288 [as ‘atro-marginatus’].
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 207 [45], pl. 13, fig. 21 [as ‘atro-marginatus’].

Unio biangularis Lea, 1840 = Ortmanniana pectorosa

Lea, 1840. Proc. Amer. Phil. Soc. 1: 288.
Lea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 197, pl. 9, fig. 8 [as ‘biangulatus’].

image of type lot

Unio biangulatus Lea, 1840 = Ortmanniana pectorosa

Lea, 1840. Proc. Amer. Phil. Soc. 1: 288 [as ‘biangularis’].
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 197, pl. 9, fig. 8.
(error for Unio biangularis Lea, 1840)
Type(s): HOLOTYPE USNM_86120, Caney Fork, Tennessee.

There are 46 available nominal species attributed to this publication (51 total).
image of type lot

Unio bournianus Lea, 1840 = Pleurobema clava

Lea, 1840. Proc. Amer. Phil. Soc. 1: 288.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 213 [51], pl. 15, fig. 28.
Type(s): HOLOTYPE USNM_84683, Scioto R., Chillicothe, Ohio.

image of type lot

Unio boydianus Lea, 1840 = Atlanticoncha ochracea

Lea, 1840. Proc. Amer. Phil. Soc. 1: 286.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 216 [54], pl. 16, fig. 32.
Type(s): LECTOTYPE USNM_86126, Oak Orchard, Orleans Co., N.Y.

image of type lot

Unio boykinianus Lea, 1840 = Megalonaias nervosa

Lea, 1840. Proc. Amer. Phil. Soc. 1: 288.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 208 [46], pl. 13, fig. 22.
Type(s): TOPOTYPE USNM_83903, Columbus, Georgia.

image of type lot

Unio cincinnatiensis Lea, 1840 = Epioblasma cincinnatiensis

Lea, 1840. Proc. Amer. Phil. Soc. 1: 285.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 194, pl. 8, fig. 4.
Type(s): HOLOTYPE USNM_84199, Cincinnati, Ohio.

Anodonta couperiana Lea, 1840 = Utterbackiana couperiana

Lea, 1840. Proc. Amer. Phil. Soc. 1: 289 [as ‘cowperiana’].
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 227, pl. 20, fig. 46.
(correct name for Anodonta cowperiana Lea, 1840)
Type(s): LECTOTYPE USNM_86673 (by R.I. Johnson, 1965: 3), [Altamaha River] Hopeton, near Darien [McIntosh Co.], Georgia.

Anodonta cowperiana Lea, 1840 = Utterbackiana couperiana

Lea, 1840. Proc. Amer. Phil. Soc. 1: 289.
(incorrect original spelling of Anodonta couperiana Lea, 1840)

image of type lot

Unio cucumoides Lea, 1840 = Cucumerunio novaehollandiae

Lea, 1840. Proc. Amer. Phil. Soc. 1: 285.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 192, pl. 7, fig. 2.
Type(s): HOLOTYPE (MONOTYPY) AMNH_30897, Hunters River, New South Wales, Australia.

image of type lot

Unio cuneolus Lea, 1840 = Fusconaia cuneolus

Lea, 1840. Proc. Amer. Phil. Soc. 1: 286.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 193 [31], pl. 7, fig. 3.
Type(s): HOLOTYPE USNM_84591, Holston R., Tenn.

image of type lot

Margaritana curreyiana Lea, 1840 = Pegias fabula

Lea, 1840. Proc. Amer. Phil. Soc. 1: 288.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 223, pl. 18, fig. 40 [on plate as ‘curreyana’].
Type(s): LECTOTYPE USNM_86227 (by Clarke, 1981: 9), Stones River, Tenn.

image of type lot

Unio dactylus Lea, 1840 = Cambarunio dactylus

Lea, 1840. Proc. Amer. Phil. Soc. 1: 287.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 196, pl. 9, fig. 7.
Type(s): HOLOTYPE USNM_85904, Cany [Caney] Fork, E. Tenn.

image of type lot

Unio delphinus Lea, 1840 = Hyriopsis bialata

Lea, 1840. Proc. Amer. Phil. Soc. 1: 285.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 218, pl. 17, fig. 35.
(homonym of Unio delphinus Spengler, 1793)
Type(s): HOLOTYPE USNM_83910, New Holland.

image of type lot

Unio dolabelloides Lea, 1840 = Pleuronaia dolabelloides

Lea, 1840. Proc. Amer. Phil. Soc. 1: 288.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 215 [53], pl. 15, fig. 31.
Type(s): HOLOTYPE USNM_85828, Holston R., Tenn.

image of type lot

Unio edgarianus Lea, 1840 = Fusconaia cor

Lea, 1840. Proc. Amer. Phil. Soc. 1: 288.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 214 [52], pl. 15, fig. 30.
Type(s): HOLOTYPE USNM_84685, Tenn R., Florence, Ala.

image of type lot

Unio exiguus Lea, 1840 = Villosa vibex

Lea, 1840. Proc. Amer. Phil. Soc. 1: 287.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 191 [29], pl. 7, fig. 1.
Type(s): HOLOTYPE USNM_84974, Chattahoochee R., Columbus, Georgia.

image of type lot

Unio fatuus Lea, 1840 = Cambarunio iris

Lea, 1840. Proc. Amer. Phil. Soc. 1: 287.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 201, pl. 11, fig. 14.
Type(s): HOLOTYPE USNM_86047, Holston R., Tenn.

image of type lot

Anodonta ferruginea Lea, 1840 = Anodontoides ferussacianus

Lea, 1840. Proc. Amer. Phil. Soc. 1: 289.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 225 [63], pl. 19, fig. 43.
Type(s): HOLOTYPE USNM_86539, Simons Creek, Indiana.

image of type lot

Anodonta footiana Lea, 1840 = Pyganodon grandis

Lea, 1840. Proc. Amer. Phil. Soc. 1: 289.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 225 [63], pl. 20, fig. 44.
Type(s): HOLOTYPE USNM_86468, Fort Winnebago.

image of type lot

Unio geddingsianus Lea, 1840 = Elliptio icterina

Lea, 1840. Proc. Amer. Phil. Soc. 1: 285.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 202 [40], pl. 11, fig. 15.
Type(s): HOLOTYPE USNM_85650, Congaree R., S.C.

image of type lot

Anodonta harpethensis Lea, 1840 = Pyganodon grandis

Lea, 1840. Proc. Amer. Phil. Soc. 1: 289.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 224 [62], pl. 19, fig. 42.
Type(s): HOLOTYPE USNM_86657, Harpeth R., Tenn.

image of type lot

Unio holstonensis Lea, 1840 = Pleurobema oviforme

Lea, 1840. Proc. Amer. Phil. Soc. 1: 288.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 212 [50], pl. 15, fig. 27.
Type(s): HOLOTYPE USNM_84641, Holston R., Tenn.

image of type lot

Unio incrassatus Lea, 1840 = Elliptio crassidens

Lea, 1840. Proc. Amer. Phil. Soc. 1: 286.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 217 [55], pl. 16, fig. 34.
Type(s): HOLOTYPE USNM_84537, Columbus, Ga.

image of type lot

Unio javanus Lea, 1840 = Lens contradens

Lea, 1840. Proc. Amer. Phil. Soc. 1: 285.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 220, pl. 18, fig. 37.
Type(s): HOLOTYPE USNM_85187, Java.

image of type lot

Unio lenior Lea, 1842 = Epioblasma lenior

Lea, 1840. Proc. Amer. Phil. Soc. 1: 286 [as ‘lenis’].
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 204, pl. 12, fig. 18.
(new name for Unio lenis Lea, 1840)
Type(s): HOLOTYPE USNM_86130, Stones R., Tenn.

Unio lenis Lea, 1840 = Epioblasma lenior

Lea, 1840. Proc. Amer. Phil. Soc. 1: 286.
(homonym of Unio lenis Conrad, 1838; renamed Unio lenior Lea, 1842)

image of type lot

Unio lesueurianus Lea, 1840 = Fusconaia subrotunda

Lea, 1840. Proc. Amer. Phil. Soc. 1: 286.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 195 [33], pl. 8, fig. 6.
Type(s): HOLOTYPE USNM_85803, Holston R., Tenn.

image of type lot

Unio lineatus Lea, 1840 = Lampsilis binominata

Lea, 1840. Proc. Amer. Phil. Soc. 1: 287.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 206 [44], pl. 12, fig. 20.
(homonym of Unio lineata Bory de St. Vincent, 1827; renamed Lampsilis (s.s.) binominatus Simpson, 1900)
Type(s): LECTOTYPE USNM_84884, Chattahoochee River, Columbus, [Muscogee County,] Georgia, [Chattahoochee River drainage].

Anodonta maryattana Lea, 1840 = Pyganodon grandis

Lea, 1840. Proc. Amer. Phil. Soc. 1: 289 [as ‘maryattiana’].
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 226, pl. 20, fig. 45.
(correct name for Anodonta maryattiana Lea, 1840)

Anodonta maryattiana Lea, 1840 = Pyganodon grandis

Lea, 1840. Proc. Amer. Phil. Soc. 1: 289.
(incorrect original spelling of Anodonta maryattana Lea, 1840)

image of type lot

Unio nitens Lea, 1840 = Leaunio vanuxemensis

Lea, 1840. Proc. Amer. Phil. Soc. 1: 288.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 205 [43], pl. 12, fig. 19.
Type(s): HOLOTYPE USNM_85132, Long Creek, Tenn.

image of type lot

Unio obtusus Lea, 1840 = Lampsilis straminea

Lea, 1840. Proc. Amer. Phil. Soc. 1: 287.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 201 [39], pl. 11, fig. 13.
(homonym of Unio obtusa d’Orbigny, 1835)
Type(s): HOLOTYPE USNM_86142, Georgia.

image of type lot

Unio orientalis Lea, 1840 = Elongaria orientalis

Lea, 1840. Proc. Amer. Phil. Soc. 1: 285.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 221, pl. 18, fig. 38.
Type(s): HOLOTYPE USNM_86058, Java.

image of type lot

Unio paulus Lea, 1840 = Toxolasma paulum

Lea, 1840. Proc. Amer. Phil. Soc. 1: 287.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 213 [51], pl. 15, fig. 29.
Type(s): HOLOTYPE (MONOTYPY) USNM_85274, Columbus, Ga.

image of type lot

Unio pilaris Lea, 1840 = Fusconaia subrotunda

Lea, 1840. Proc. Amer. Phil. Soc. 1: 285.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 209 [47], pl. 14, fig. 24.
Type(s): HOLOTYPE USNM_85817, French Broad R., Tenn.

image of type lot

Unio plenus Lea, 1840 = Pleurobema plenum

Lea, 1840. Proc. Amer. Phil. Soc. 1: 286.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 211 [49], pl. 14, fig. 26.
Type(s): HOLOTYPE USNM_84677, Cincinnati, Ohio.

image of type lot

Unio pusillus Lea, 1840 = Elliptio congaraea

Lea, 1840. Proc. Amer. Phil. Soc. 1: 286.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 220 [58], pl. 18, fig. 36.
Type(s): HOLOTYPE USNM_85241, Ogeechee R., Georgia.

image of type lot

Unio sloatianus Lea, 1840 = Elliptoideus sloatianus

Lea, 1840. Proc. Amer. Phil. Soc. 1: 287.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 217 [55], pl. 16, fig. 33.
Type(s): LECTOTYPE AMNH_56104, Chattahoochee River, Georgia.

image of type lot

Unio stonensis Lea, 1840 = Eurynia dilatata

Lea, 1840. Proc. Amer. Phil. Soc. 1: 286.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 195 [33], pl. 8, fig. 5.
Type(s): HOLOTYPE USNM_84969, Stone R., Tenn.

image of type lot

Unio striatus Lea, 1840 = Pleurobema pyriforme

Lea, 1840. Proc. Amer. Phil. Soc. 1: 287.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 203 [41], pl. 12, fig. 16.
(homonym of Unio striatus Goldfuss, 1837; renamed Pleurobema simpsoni Vanatta, 1915)
Type(s): HOLOTYPE USNM_84797, Columbus, Ga.

image of type lot

Unio strigosus Lea, 1840 = Elliptio arctata

Lea, 1840. Proc. Amer. Phil. Soc. 1: 287.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 198 [36], pl. 9, fig. 9.
Type(s): HOLOTYPE USNM_85890, Chattahoochee R., Columbus, Georgia.

image of type lot

Unio subangulatus Lea, 1840 = Hamiota subangulata

Lea, 1840. Proc. Amer. Phil. Soc. 1: 287 [as ‘sub-angulatus’].
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 209 [47], pl. 13, fig. 23.
Type(s): LECTOTYPE USNM_85081 (by Clench & Turner, 1956: 197), Chattahoochee R., Georgia.

image of type lot

Unio tener Lea, 1840 = Cambarunio iris

Lea, 1840. Proc. Amer. Phil. Soc. 1: 286.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 198, pl. 10, fig. 10.
Type(s): HOLOTYPE USNM_85026, Big Pigeon R., Tenn.

image of type lot

Unio tennesseensis Lea, 1840 = Cambarunio taeniatus

Lea, 1840. Proc. Amer. Phil. Soc. 1: 288.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 199, pl. 10, fig. 11.
Type(s): HOLOTYPE USNM_84961, Stone River, Tenn.

image of type lot

Unio tortivus Lea, 1840 = Elliptio arctata

Lea, 1840. Proc. Amer. Phil. Soc. 1: 287.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 204 [42], pl. 12, fig. 17.
Type(s): POSSIBLE SYNTYPE USNM_85674, Lee Co., Ga.

image of type lot

Unio tuberosus Lea, 1840 = Theliderma metanevra

Lea, 1840. Proc. Amer. Phil. Soc. 1: 286.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 210 [48], pl. 14, fig. 25.
Type(s): HOLOTYPE USNM_84219, Cumberland R., Middle Tenn.

image of type lot

Margaritana vondembuschiana Lea, 1840 = Pseudodon vondembuschianus

Lea, 1840. Proc. Amer. Phil. Soc. 1: 288 [as ‘vondenbuschiana’].
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 222 [60], pl. 18, fig. 39 [as ‘vondenbuschiana’].
Lea, 1852. Synopsis Fam. Naiades: 45 [as ‘vondenbuschiana’].
Lea, 1870. Synopsis Fam. Naiades: 73.
(correct name for Margaritana vondenbuschiana Lea, 1840; renamed Monodontina buschiana Conrad, 1853)
Type(s): FIGURED SYNTYPE USNM_86348, Java.

Margaritana vondenbuschiana Lea, 1840 = Pseudodon vondembuschianus

Lea, 1840. Proc. Amer. Phil. Soc. 1: 288.
specimen image from publicationLea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 222, pl. 18, fig. 39.
Lea, 1852. Synopsis Fam. Naiades: 45.
Adams & Adams, 1857. Gen. Rec. Moll. 2(31-32): 501.
(incorrect original spelling of Margaritana vondembuschiana Lea, 1840)

 
NSF icon MUSSEL icon
"Making the world a better place, one mollusk at a time."