Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Lampsilini | Cambarunio
15 specimens + 2 Genbank vouchers

Lea, 1829. Trans. Amer. Phil. Soc. 3 [O. 1]: 439, pl. 11, fig. 18.
Type(s): SYNTYPES USNM_86035, Cincinnati, Ohio.
Lampsilis (sec. Eurynia) iris — Simpson (1900)
Unio (s.s.) iris — Call (1900)
Lampsilis iris — Ortmann (1909), Ortmann (1909), Ortmann (1909), Ortmann (1910)
Eurynia (Micromya) iris — Ortmann (1912), Utterback (1916)
Lampsilis (Eurynia) iris — Simpson (1914)
Unio iris — de Gregorio (1914)
Micromya iris — Ortmann (1924), Ortmann (1925)
Lampsilis (Ligumia) nervosa — Frierson (1927)
Ligumia nervosa — Haas (1969)
Villosa iris — Burch (1975), Gordon et al. (1979), Mathiak (1979), R.I. Johnson (1980), Mackie et al. (1980), ... Bolotov et al. (2020), Inoue et al. (2020), Klymus et al. (2020), Gasparini et al. (2020), Jeratthitikul et al. (2022)
Cambarunio iris — Watters (2018), Pfeiffer et al. (2019), Graf & Cummings (2021), FMCS (2021), Hopper et al. (2023), FMCS (2025)

Lea, 1836. Synopsis Fam. Naiades: 28. [nomen nudum]
Lea, 1838. Trans. Amer. Phil. Soc. 6 [O. 2]: 33, pl. 10, fig. 28.
Type(s): HOLOTYPE (MONOTYPY) USNM_85264, Tennessee.
Lampsilis (sec. Eurynia) iris — Simpson (1900)
Lampsilis (Eurynia) iris — Simpson (1914)
Eurynia nebulosus — Ortmann (1918)
Lampsilis (Ligumia) nervosa — Frierson (1927)
Ligumia nervosa — Haas (1969)
Villosa iris — Oesch (1984), Parmalee & Bogan (1998), Williams et al. (2008)
Cambarunio iris — Watters (2018)

Lea, 1836. Synopsis Fam. Naiades: 29. [nomen nudum]
Lea, 1838. Trans. Amer. Phil. Soc. 6 [O. 2]: 34, pl. 10, fig. 29.
Type(s): HOLOTYPE USNM_85266, Holston R., Tenn.
Lampsilis (sec. Eurynia) nebulosus — Simpson (1900)
Lampsilis (Eurynia) nebulosa — Simpson (1914)
Unio glaber, aff. — de Gregorio (1914)
Eurynia nebulosus — Ortmann (1918)
Lampsilis (Ligumia) nebulosa — Frierson (1927)
Ligumia nebulosa — Haas (1969)
Villosa iris — Parmalee & Bogan (1998), Williams et al. (2008)
Cambarunio iris — Watters (2018)

Lea, 1838. Trans. Amer. Phil. Soc. 6 [O. 2]: 104, pl. 24, fig. 114 [as ‘novi-eboraci’].
Type(s): LECTOTYPE USNM_86042 (by R.I. Johnson, 1974: 100), Oak Orchard Cr., N.Y.
Lampsilis (sec. Eurynia) iris — Simpson (1900)
Lampsilis (Eurynia) iris — Simpson (1914)
Micromya iris — Ortmann (1925)
Lampsilis (Ligumia) nervosa — Frierson (1927)
Ligumia (Micromya) iris novieboraci — F.C. Baker (1928)
Ligumia nervosa — Haas (1969)
Villosa iris — Oesch (1984), Strayer & Jirka (1997), Parmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009)
Cambarunio iris — Watters (2018)

Lea, 1836. Synopsis Fam. Naiades: 26. [nomen nudum]
Lea, 1838. Trans. Amer. Phil. Soc. 6 [O. 2]: 7, pl. 3, fig. 7.
Type(s): HOLOTYPE USNM_85128, Tennessee.
Lampsilis (sec. Eurynia) obscurus — Simpson (1900)
Lampsilis (Eurynia) obscura — Simpson (1914)
Unio obscurus — de Gregorio (1914)
Eurynia nebulosus — Ortmann (1918)
Micromya nebulosa — Ortmann (1924)
Lampsilis (Ligumia) vanuxemensis — Frierson (1927)
Ligumia vanuxemensis — Haas (1969)
Villosa nebulosa — Burch (1975)
Villosa iris — Parmalee & Bogan (1998), Williams et al. (2008)
Leaunio vanuxemensis — Watters (2018)

Lea, 1836. Synopsis Fam. Naiades: 26. [nomen nudum]
Lea, 1838. Trans. Amer. Phil. Soc. 6 [O. 2]: 32, pl. 10, fig. 27.
Type(s): HOLOTYPE USNM_85130, Cumberland R., Tenn.
Lampsilis (sec. Eurynia) obscurus — Simpson (1900)
Lampsilis (Eurynia) obscura — Simpson (1914)
Eurynia nebulosus — Ortmann (1918)
Lampsilis (Ligumia) zeigleriana — Frierson (1927)
Ligumia zeigleriana — Haas (1969)
Villosa iris — Parmalee & Bogan (1998), Williams et al. (2008)
Leaunio vanuxemensis — Watters (2018)

Lea, 1840. Proc. Amer. Phil. Soc. 1: 286 [as ‘amœnus’].
Lea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 200, pl. 10, fig. 12 [as ‘amœnus’].
Type(s): HOLOTYPE USNM_85070, Holston R., Tenn.
Lampsilis (sec. Eurynia) amœnus — Simpson (1900)
Lampsilis (Eurynia) amoena — Simpson (1914)
Eurynia nebulosus — Ortmann (1918)
Lampsilis (Ligumia) nervosa — Frierson (1927)
Ligumia nervosa — Haas (1969)
Villosa nebulosa — Burch (1975)
Villosa iris — Parmalee & Bogan (1998), Williams et al. (2008)
Cambarunio iris — Watters (2018)

Lea, 1840. Proc. Amer. Phil. Soc. 1: 287.
Lea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 201, pl. 11, fig. 14.
Type(s): HOLOTYPE USNM_86047, Holston R., Tenn.
Lampsilis (sec. Eurynia) fatuus — Simpson (1900)
Lampsilis (Eurynia) fatua — Simpson (1914)
Unio fatuus, var. — de Gregorio (1914)
Eurynia nebulosus — Ortmann (1918)
Lampsilis (Ligumia) nervosa — Frierson (1927)
Ligumia nervosa — Haas (1969)
Villosa iris — Parmalee & Bogan (1998), Williams et al. (2008)
Cambarunio iris — Watters (2018)

Lea, 1840. Proc. Amer. Phil. Soc. 1: 286.
Lea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 198, pl. 10, fig. 10.
Type(s): HOLOTYPE USNM_85026, Big Pigeon R., Tenn.
Lampsilis (sec. Eurynia) tener — Simpson (1900)
Lampsilis (Eurynia) tenera — Simpson (1914)
Eurynia nebulosus — Ortmann (1918)
Lampsilis (Ligumia) tenera — Frierson (1927)
Ligumia tenera — Haas (1969)
Villosa iris — Parmalee & Bogan (1998), Williams et al. (2008)
Cambarunio iris — Watters (2018)

Lea, 1841. Proc. Amer. Phil. Soc. 2: 82.
Lea, 1842. Trans. Amer. Phil. Soc. 8 [O. 3]: 243, pl. 25, fig. 59.
Type(s): HOLOTYPE USNM_85067, French Broad River, E Tenn.
Lampsilis (sec. Eurynia) tener — Simpson (1900)
Lampsilis (Eurynia) tenera — Simpson (1914)
Eurynia nebulosus — Ortmann (1918)
Lampsilis (Ligumia) tenera — Frierson (1927)
Ligumia tenera — Haas (1969)
Villosa iris — Parmalee & Bogan (1998), Williams et al. (2008)
Cambarunio iris — Watters (2018)
Haldeman, 1842. Jour. Acad. Nat. Sci. 8: 201.
Lampsilis (sec. Eurynia) puniceus — Simpson (1900)
Lampsilis (Eurynia) punicea — Simpson (1914)
Eurynia nebulosus — Ortmann (1918)
Lampsilis (Ligumia) vanuxemensis — Frierson (1927)
Ligumia vanuxemensis — Haas (1969)
Villosa iris — Parmalee & Bogan (1998), Williams et al. (2008)
Cambarunio iris — Watters (2018)
‘Lea’ of Authors.
Hanley, 1843. Illustr. Descr. Cat. Rec. Bivalve Shells: 193.
Conrad, 1853. Proc. Acad. Nat. Sci. 6: 260.
Lea, 1870. Synopsis Fam. Naiades: 45.
(error for Unio zeiglerianus Lea, 1838)

Lea, 1860. Proc. Acad. Nat. Sci. 12: 306.
Lea, 1860. Jour. Acad. Nat. Sci. 4 [O. 8]: 350, pl. 58, fig. 175.
Type(s): HOLOTYPE USNM_85034, Bull R., Tenn.
Lampsilis (sec. Eurynia) nebulosus — Simpson (1900)
Lampsilis (Eurynia) nebulosa — Simpson (1914)
Eurynia nebulosus — Ortmann (1918)
Lampsilis (Ligumia) nebulosa — Frierson (1927)
Ligumia nebulosa — Haas (1969)
Villosa iris — Parmalee & Bogan (1998), Williams et al. (2008)
Cambarunio iris — Watters (2018)

Lea, 1860. Proc. Acad. Nat. Sci. 12: 93.
Lea, 1860. Jour. Acad. Nat. Sci. 4 [O. 8]: 42, pl. 55, fig. 167.
Type(s): HOLOTYPE USNM_85084, Tuscumbia, Alabama.
Lampsilis (sec. Eurynia) nebulosus — Simpson (1900)
Lampsilis (Eurynia) nebulosa — Simpson (1914)
Eurynia nebulosus — Ortmann (1918)
Micromya nebulosa — Ortmann (1925)
Lampsilis (Ligumia) nebulosa — Frierson (1927)
Ligumia nebulosa — Haas (1969)
Villosa nebulosa — Burch (1975)
Villosa iris — Parmalee & Bogan (1998), Williams et al. (2008)
Cambarunio iris — Watters (2018)
Binney, 1864. Smithsonian Misc. Coll. (174): 57.
(error for Unio cresserus Hanley, 1843)
Ligumia nervosa — Haas (1969)
‘Say’ Reeve, 1864. Conch. Icon. 16: pl. 17, sp. 78; errata [changed to ‘U. patulus Lea’].
(misidentified reference to Unio subrostratus Say, 1831; error for Unio patulus Lea, 1829)
Lampsilis (sec. Eurynia) iris — Simpson (1900)
Lampsilis (Eurynia) iris — Simpson (1914)
Lampsilis (Ligumia) nervosa — Frierson (1927)

J.G. Anthony, 1866. Amer. J. Conch. 2(2): 146, pl. 7, fig. 2.
Type(s): HOLOTYPE MCZ_161893, Michigan.
Lampsilis (sec. Eurynia) iris — Simpson (1900)
Lampsilis (Eurynia) iris — Simpson (1914)
Lampsilis (Ligumia) fasciata-opalina — Frierson (1927)
Ligumia siliquoidea siliquoidea — Haas (1969)
Villosa iris — Parmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009)
Cambarunio iris — Watters (2018)

Lea, 1871. Proc. Acad. Nat. Sci. 23: 191.
Lea, 1874. Jour. Acad. Nat. Sci. 8 [O. 13]: 19 [23], pl. 6, fig. 16.
Type(s): FIGURED SYNTYPE USNM_85094, E Tennessee.
Lampsilis (sec. Eurynia) vanuxemensis — Simpson (1900)
Lampsilis (Eurynia) vanuxemensis — Simpson (1914)
Eurynia nebulosus — Ortmann (1918)
Lampsilis (Ligumia) vanuxemensis — Frierson (1927)
Ligumia vanuxemensis — Haas (1969)
Villosa iris — Parmalee & Bogan (1998), Williams et al. (2008)
Leaunio vanuxemensis — Watters (2018)
‘Rafinesque’ of Authors.
Frierson, 1927. Check List N. Amer. Naiades: 77, sp. 336.
Haas, 1969. Das Tierreich 88: 445.
(misidentified reference to Unio (Leptodea) nervosa Rafinesque, 1820)
Villosa iris — Parmalee & Bogan (1998), Watters et al. (2009)
Cambarunio iris — Watters (2018)
‘Lea’ Hanley, 1843. Illustr. Descr. Cat. Rec. Bivalve Shells: 196.
(error for Unio creperus Lea, 1838)
Lampsilis (sec. Eurynia) iris — Simpson (1900)
Lampsilis (Eurynia) iris — Simpson (1914)
Ligumia nervosa — Haas (1969)
Villosa iris — Parmalee & Bogan (1998)
‘Gmelin’ De Kay, 1843. Zool. New York 5: 189, pl. 18. fig. 236.
(misidentified reference to Mya radiata Gmelin, 1791)
Lampsilis (sec. Eurynia) iris — Simpson (1900)
Lampsilis (Eurynia) iris — Simpson (1914)
Villosa iris — Oesch (1984), Parmalee & Bogan (1998)
Lea, 1852. Synopsis Fam. Naiades: 29 [as ‘amænus’].
‘Lea’ Hanley, 1856. Illustr. Descr. Cat. Rec. Bivalve Shells: 386 [as ‘amænus’].
(error for Unio amoenus Lea, 1840)
‘Say’ Küster, 1861. Conchyl. Cab. 9(2): 203, pl. 67, fig. 3.
(misidentified reference to Unio subrostratus Say, 1831)
Lampsilis (sec. Eurynia) iris — Simpson (1900)
Lampsilis (Eurynia) iris — Simpson (1914)
Lampsilis (Ligumia) nebulosa — Frierson (1927)
Villosa iris — Parmalee & Bogan (1998), Watters et al. (2009)
‘Say’ Küster, 1861. Conchyl. Cab. 9(2): 203, pl. 68, fig. 1.
(misidentified reference to Unio subrostratus Say, 1831)
Lampsilis (sec. Eurynia) subrostratus — Simpson (1900)
Lampsilis (Eurynia) nebulosa — Simpson (1914)
Lampsilis (sec. Eurynia) puniceus — Simpson (1900)
Lampsilis (sec. Eurynia) vanuxemensis — Simpson (1900)
Lampsilis (sec. Eurynia) obscurus — Simpson (1900)
Lampsilis (sec. Eurynia) iris — Simpson (1900)
Lampsilis (sec. Eurynia) fatuus — Simpson (1900)
Lampsilis (sec. Eurynia) nebulosus — Simpson (1900)
Lampsilis (sec. Eurynia) amœnus — Simpson (1900)
Lampsilis (sec. Eurynia) tener — Simpson (1900)
Lampsilis (sec. Eurynia) subrostratus — Simpson (1900)
Unio (s.s.) iris — Call (1900)
Lampsilis iris — Ortmann (1909), Ortmann (1909), Ortmann (1909), Ortmann (1910)
Eurynia (Micromya) iris — Ortmann (1912), Utterback (1916)
Lampsilis (Eurynia) punicea — Simpson (1914)
Lampsilis (Eurynia) vanuxemensis — Simpson (1914)
Lampsilis (Eurynia) obscura — Simpson (1914)
Lampsilis (Eurynia) iris — Simpson (1914)
Lampsilis (Eurynia) fatua — Simpson (1914)
Lampsilis (Eurynia) nebulosa — Simpson (1914)
Lampsilis (Eurynia) amoena — Simpson (1914)
Lampsilis (Eurynia) tenera — Simpson (1914)
Unio obscurus — de Gregorio (1914)
Unio fatuus, var. — de Gregorio (1914)
Unio glaber, aff. — de Gregorio (1914)
Unio iris — de Gregorio (1914)
Eurynia nebulosus — Ortmann (1918)
Micromya nebulosa — Ortmann (1924), Ortmann (1925)
Micromya iris — Ortmann (1924), Ortmann (1925)
Lampsilis (Ligumia) fasciata-opalina — Frierson (1927)
Lampsilis (Ligumia) zeigleriana — Frierson (1927)
Lampsilis (Ligumia) vanuxemensis — Frierson (1927)
Lampsilis (Ligumia) tenera — Frierson (1927)
Lampsilis (Ligumia) nervosa — Frierson (1927)
Lampsilis (Ligumia) nebulosa — Frierson (1927)
Ligumia (Micromya) iris novieboraci — F.C. Baker (1928)
Ligumia siliquoidea siliquoidea — Haas (1969)
Ligumia zeigleriana — Haas (1969)
Ligumia vanuxemensis — Haas (1969)
Ligumia tenera — Haas (1969)
Ligumia nervosa — Haas (1969)
Ligumia nebulosa — Haas (1969)
Villosa iris — Burch (1975), Gordon et al. (1979), Mathiak (1979), R.I. Johnson (1980), Mackie et al. (1980), ... Bolotov et al. (2020), Inoue et al. (2020), Klymus et al. (2020), Gasparini et al. (2020), Jeratthitikul et al. (2022)
Villosa nebulosa — Burch (1975)
Cambarunio iris — Watters (2018), Pfeiffer et al. (2019), Graf & Cummings (2021), FMCS (2021), Hopper et al. (2023), FMCS (2025)
Leaunio vanuxemensis — Watters (2018)