MUSSELp

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Unionidae

Margaritiferidae

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Data last updated
Thu 18 Dec 2025

Bivalvia | Unionoida | Unionoidea | Unionidae | Gonideinae | Pseudodontini

subtribe PSEUDODONTINA Frierson, 1927: cladomics

10 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 51 (52 total)
number of subtaxa (rank: genus) = 10
analyzed in 95 cladograms in 57 publications
maximum number of terminals in an analysis = 39 (mean = 7.57, median = 3)
maximum number of subtaxa (genus) = 10 (mean = 3.68, median = 3)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 85
maximum number of included terminals = 39, (mean = 8.34 median = 4)
maximum number of subtaxa (genus) = 10 (mean = 4, median = 3)
proportion of cladograms that support basic monophyly = 0.92

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 46
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 39 (mean = 12.65, median = 9.5)
maximum number of subtaxa (genus) = 10 (mean = 5.28, median = 5.5)
proportion of cladograms that support rigorous monophyly = 0.85

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Pseudodontina cladogram distances
PCoA Ordination

NJ Tree of Pseudodontina cladogram distances
NJ Tree

Cladograms

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
35 35 36 10 23 5 1 0 
  +----------------- Indopseudodontina
  |                                   
  |  +-------------- Pseudodon **        
--+  |                                
  |  |     +-------- Sundadontina **     
  |  |  +--+                          
  +--+  |  |  +----- Nyeinchanconcha **  
     |  |  +--+                       
     |  |     |  +-- Thaiconcha **       
     |  |     +--+                    
     +--+        +-- Bineurus **         
        |                             
        |        +-- Isannaia **         
        |  +-----+                    
        |  |     +-- Pilsbryoconcha **   
        +--+                          
           |  +----- Namkongnaia **      
           +--+                       
              |  +-- Lannanaia **        
              +--+                    
                 +-- Songkhlanaia **

Konopleva et al., 2025 [Publ 4446], fig. 4

iTtTbiriSoTroTroSiD
39 39 38 10 5 5 1 23 
  +----------------------- Indopseudodontina
  |                                         
--+  +-------------------- Lannanaia **        
  |  |                                      
  +--+  +----------------- Pilsbryoconcha **   
     |  |                                   
     +--+  +-------------- Songkhlanaia **     
        |  |                                
        |  |        +----- Nyeinchanconcha **  
        +--+  +-----+                       
           |  |     |  +-- Thaiconcha **       
           |  |     +--+                    
           +--+        +-- Bineurus **         
              |                             
              |  +-------- Sundadontina **     
              +--+                          
                 |  +----- Pseudodon **        
                 +--+                       
                    |  +-- Namkongnaia **      
                    +--+                    
                       +-- Isannaia **

Jeratthitikul et al., 2025 [Publ 4423], fig. 1

iTtTbiriSoTroTroSiD
38 38 37 10 12 5 1 24 
  +-------------------- Indopseudodontina
  |                                      
  |                 +-- Lannanaia **        
--+  +--------------+                    
  |  |              +-- Pilsbryoconcha **   
  |  |                                   
  +--+  +-------------- Songkhlanaia **     
     |  |                                
     |  |        +----- Nyeinchanconcha **  
     +--+  +-----+                       
        |  |     |  +-- Thaiconcha **       
        |  |     +--+                    
        +--+        +-- Bineurus **         
           |                             
           |  +-------- Sundadontina **     
           +--+                          
              |  +----- Isannaia **         
              +--+                       
                 |  +-- Namkongnaia **      
                 +--+                    
                    +-- Pseudodon **

Bolotov et al., 2023 [Publ 4312], fig. S2

iTtTbiriSoTroTroSiD
36 36 34 8 5 5 1 33 
  +----------------- Indopseudodontina
  |                                   
--+  +-------------- Songkhlanaia **     
  |  |                                
  |  |           +-- Nyeinchanconcha **  
  +--+  +--------+                    
     |  |        +-- Sundadontina **     
     |  |                             
     +--+        +-- Pseudodon **        
        |  +-----+                    
        |  |     +-- Pilsbryoconcha **   
        +--+                          
           |  +----- Namkongnaia **      
           +--+                       
              |  +-- Thaiconcha **       
              +--+                    
                 +-- Bineurus **

Bolotov et al., 2023 [Publ 4312], fig. 2

iTtTbiriSoTroTroSiD
34 34 33 8 5 5 1 26 
  +-------------------- Indopseudodontina
  |                                      
--+  +----------------- Pilsbryoconcha **   
  |  |                                   
  +--+  +-------------- Songkhlanaia **     
     |  |                                
     +--+           +-- Namkongnaia **      
        |  +--------+                    
        |  |        +-- Pseudodon **        
        +--+                             
           |  +-------- Sundadontina **     
           +--+                          
              |  +----- Nyeinchanconcha **  
              +--+                       
                 |  +-- Thaiconcha **       
                 +--+                    
                    +-- Bineurus **

Bolotov et al., 2023 [Publ 4312], fig. S1

iTtTbiriSoTroTroSiD
34 34 32 8 5 5 1 26 
  +----------------- Indopseudodontina
  |                                   
--+  +-------------- Pilsbryoconcha **   
  |  |                                
  |  |        +----- Nyeinchanconcha **  
  +--+  +-----+                       
     |  |     |  +-- Thaiconcha **       
     |  |     +--+                    
     +--+        +-- Bineurus **         
        |                             
        |  +-------- Songkhlanaia **     
        +--+                          
           |  +----- Sundadontina **     
           +--+                       
              |  +-- Namkongnaia **      
              +--+                    
                 +-- Pseudodon **

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
26 26 25 8 56 4 1 24 
  +----------------------- Indopseudodontina
--+                                         
  |  +-------------------- Pilsbryoconcha **   
  +--+                                      
     |  +----------------- Songkhlanaia **     
     +--+                                   
        |  +-------------- Namkongnaia **      
        +--+                                
           |  +----------- Pseudodon **        
           +--+                             
              |  +-------- Sundadontina **     
              +--+                          
                 |  +----- Nyeinchanconcha **  
                 +--+                       
                    |  +-- Thaiconcha **       
                    +--+                    
                       +-- Bineurus **

Konopleva et al., 2021 [Publ 4213], fig. 1

iTtTbiriSoTroTroSiD
19 19 17 7 8 4 1 26 
  +-------------- Indopseudodontina       
  |                                       
  |        +----- Pilsbryoconcha_exilis__2 **
--+  +-----+                              
  |  |     |  +-- Pilsbryoconcha_exilis__1 **
  |  |     +--+                           
  |  |        +-- Namkongnaia **             
  +--+                                    
     |        +-- Pseudodon **               
     |  +-----+                           
     |  |     +-- Sundadontina **            
     +--+                                 
        |  +----- Nyeinchanconcha **         
        +--+                              
           |  +-- Thaiconcha **              
           +--+                           
              +-- Bineurus **

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iTtTbiriSoTroTroSiD
18 18 16 7 18 4 1 38 
  +----------------- Indopseudodontina    
  |                                       
  |  +-------------- Pilsbryoconcha **       
--+  |                                    
  |  |           +-- Nyeinchanconcha **      
  |  |        +--+                        
  +--+        |  +-- Sundadontina_cumingii **
     |  +-----+                           
     |  |     |  +-- Thaiconcha **           
     |  |     +--+                        
     |  |        +-- Bineurus **             
     +--+                                 
        |        +-- Namkongnaia **          
        |  +-----+                        
        |  |     +-- Pseudodon **            
        +--+                              
           |  +----- Sundadontina_tumida **  
           +--+                           
              |  +-- Sundadontina_brandti ** 
              +--+                        
                 +-- Sundadontina_taskaevi **

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iTtTbiriSoTroTroSiD
18 18 15 7 18 4 1 32 
  +-------------- Indopseudodontina
  |                                
--+  +----------- Pilsbryoconcha **   
  |  |                             
  |  |     +----- Nyeinchanconcha **  
  +--+  +--+                       
     |  |  |  +-- Thaiconcha **       
     |  |  +--+                    
     +--+     +-- Bineurus **         
        |                          
        |  +----- Sundadontina **     
        +--+                       
           |  +-- Namkongnaia **      
           +--+                    
              +-- Pseudodon **

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
17 17 16 6 101 4 1 33 
  +-------------- Indopseudodontina
  |                                
--+  +----------- Bineurus **         
  |  |                             
  +--+  +-------- Nyeinchanconcha **  
     |  |                          
     +--+     +-- Thaiconcha **       
        |  +--+                    
        |  |  +-- Pilsbryoconcha **   
        +--+                       
           |  +-- Pseudodon **        
           +--+                    
              +-- Sundadontina **

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
17 17 16 6 95 4 1 35 
  +-------------- Indopseudodontina
  |                                
--+  +----------- Bineurus **         
  |  |                             
  +--+  +-------- Nyeinchanconcha **  
     |  |                          
     +--+     +-- Thaiconcha **       
        |  +--+                    
        |  |  +-- Pilsbryoconcha **   
        +--+                       
           |  +-- Pseudodon **        
           +--+                    
              +-- Sundadontina **

Jeratthitikul et al., 2025 [Publ 4476], fig. 2

iTtTbiriSoTroTroSiD
16 16 14 5 0 0 0 32 
  +----------- Pilsbryoconcha **
--+                          
  |  +-------- Pseudodon **     
  +--+                       
     |  +----- Namkongnaia **   
     +--+                    
        |  +-- Isannaia **      
        +--+                 
           +-- Sundadontina **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
15 15 13 7 61 2 1 39 
  +----------- Indopseudodontina
  |                             
  |     +----- Pilsbryoconcha **   
--+  +--+                       
  |  |  |  +-- Namkongnaia **      
  |  |  +--+                    
  |  |     +-- Pseudodon **        
  +--+                          
     |     +-- Nyeinchanconcha **  
     |  +--+                    
     |  |  +-- Sundadontina **     
     +--+                       
        |  +-- Thaiconcha **       
        +--+                    
           +-- Bineurus **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
15 15 13 7 61 2 1 39 
  +----------- Indopseudodontina
  |                             
  |     +----- Pilsbryoconcha **   
--+  +--+                       
  |  |  |  +-- Namkongnaia **      
  |  |  +--+                    
  |  |     +-- Pseudodon **        
  +--+                          
     |     +-- Nyeinchanconcha **  
     |  +--+                    
     |  |  +-- Sundadontina **     
     +--+                       
        |  +-- Thaiconcha **       
        +--+                    
           +-- Bineurus **

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
14 14 11 6 31 3 1 33 
  +-------- Indopseudodontina
  |                          
  |     +-- Nyeinchanconcha **  
--+  +--+                    
  |  |  +-- Bineurus **         
  |  |                       
  |  |  +-- Thaiconcha **       
  +--+--+                    
     |  +-- Sundadontina **     
     |                       
     |  +-- Pilsbryoconcha **   
     +--+                    
        +-- Pseudodon **

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
14 14 12 6 31 3 1 34 
  +----------- Indopseudodontina
  |                             
  |        +-- Nyeinchanconcha **  
--+  +-----+                    
  |  |     +-- Bineurus **         
  |  |                          
  +--+     +-- Thaiconcha **       
     |  +--+                    
     |  |  +-- Sundadontina **     
     +--+                       
        |  +-- Pilsbryoconcha **   
        +--+                    
           +-- Pseudodon **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
13 14 13 6 126 1 1 38 
  +-------------- Pilsbryoconcha **            
  |                                         
  |  +----------- Indopseudodontina         
--+  |                                      
  |  |        +-- Thaiconcha **                
  |  +--------+                             
  |  |        +-- Bineurus **                  
  +--+                                      
     |     +----- Pseudodon_cambodjensis **    
     |  +--+                                
     |  |  |  +-- Pseudodon_mekongi **         
     |  |  +--+                             
     |  |     +-- Pseudodon_vondembuschianus **
     +--+                                   
        |     +-- Pseudodon_inoscularis **     
        |  +--+                             
        |  |  +-- Sundadontina_tumida **       
        +--+                                
           |  +-- Nyeinchanconcha **           
           +--+                             
              +-- Sundadontina_cumingii **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
12 12 12 6 127 1 1 36 
  +-------------- Indopseudodontina    
  |                                    
--+  +----------- Pilsbryoconcha **       
  |  |                                 
  |  |        +-- Sundadontina_tumida **  
  +--+  +-----+                        
     |  |     +-- Pseudodon **            
     |  |                              
     +--+     +-- Nyeinchanconcha **      
        |  +--+                        
        |  |  +-- Sundadontina_cumingii **
        +--+                           
           |  +-- Thaiconcha **           
           +--+                        
              +-- Bineurus **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
12 12 13 6 187 3 1 36 
  +-------------- Indopseudodontina    
  |                                    
--+  +----------- Pilsbryoconcha **       
  |  |                                 
  |  |        +-- Sundadontina_tumida **  
  +--+  +-----+                        
     |  |     +-- Pseudodon **            
     |  |                              
     +--+     +-- Nyeinchanconcha **      
        |  +--+                        
        |  |  +-- Sundadontina_cumingii **
        +--+                           
           |  +-- Thaiconcha **           
           +--+                        
              +-- Bineurus **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
12 12 6 6 189 3 1 34 
  +----- Indopseudodontina     
  |                            
  |  +-- Nyeinchanconcha **       
  |  |                         
--+  +-- Thaiconcha **            
  |  |                         
  |  +-- Sundadontina_cumingii ** 
  |  |                         
  |  +-- Sundadontina_tumida__1 **
  +--+                         
     +-- Sundadontina_tumida__2 **
     |                         
     +-- Pilsbryoconcha **        
     |                         
     +-- Pseudodon **             
     |                         
     +-- Bineurus **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
12 12 12 6 189 3 1 33 
  +-------------- Indopseudodontina
  |                                
--+  +----------- Pilsbryoconcha **   
  |  |                             
  +--+        +-- Pseudodon **        
     |  +-----+                    
     |  |     +-- Sundadontina **     
     +--+                          
        |  +----- Nyeinchanconcha **  
        +--+                       
           |  +-- Thaiconcha **       
           +--+                    
              +-- Bineurus **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
12 12 10 6 113 3 1 36 
  +-------- Indopseudodontina    
  |                              
  |  +----- Bineurus **             
--+  |                           
  |  |  +-- Nyeinchanconcha **      
  |  +--+                        
  |  |  +-- Sundadontina_cumingii **
  +--+                           
     |  +-- Sundadontina_tumida **  
     +--+                        
     |  +-- Pseudodon **            
     |                           
     |  +-- Thaiconcha **           
     +--+                        
        +-- Pilsbryoconcha **

Jeratthitikul & Sutcharit, 2023 [Publ 4289], fig. 1

iTtTbiriSoTroTroSiD
11 11 9 7 2 2 1 39 
  +-------------- Indopseudodontina
  |                                
--+  +----------- Bineurus **         
  |  |                             
  |  |     +----- Namkongnaia **      
  +--+  +--+                       
     |  |  |  +-- Sundadontina **     
     |  |  +--+                    
     +--+     +-- Nyeinchanconcha **  
        |                          
        |  +----- Pseudodon **        
        +--+                       
           |  +-- Pilsbryoconcha **   
           +--+                    
              +-- Thaiconcha **

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
11 11 9 6 36 0 0 29 
  +-------------- Pilsbryoconcha ** 
--+                              
  |  +----------- Pseudodon **      
  +--+                           
     |  +-------- Sundadontina **   
     +--+                        
        |  +----- Nyeinchanconcha **
        +--+                     
           |  +-- Bineurus **       
           +--+                  
              +-- Thaiconcha **

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
11 11 9 6 37 0 0 31 
  +-------------- Pilsbryoconcha ** 
--+                              
  |  +----------- Pseudodon **      
  +--+                           
     |  +-------- Sundadontina **   
     +--+                        
        |  +----- Nyeinchanconcha **
        +--+                     
           |  +-- Bineurus **       
           +--+                  
              +-- Thaiconcha **

Zieritz et al., 2024 [Publ 4381], fig. 8A

iTtTbiriSoTroTroSiD
8 8 6 2 0 0 0 34 
  +-- Namkongnaia **
--+              
  +-- Pseudodon **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
8 8 0 3 75 3 1 37 
  +----- Indopseudodontina         
  |                                
  |  +-- Bineurus_mouhotii **         
  |  |                             
--+  +-- Pseudodon_cambodjensis **    
  |  |                             
  |  +-- Pseudodon_mekongi **         
  |  |                             
  |  +-- Pilsbryoconcha_exilis **     
  +--+                             
     +-- Pilsbryoconcha_carinifera ** 
     |                             
     +-- Bineurus_loeiensis **        
     |                             
     +-- Bineurus_exilis **           
     |                             
     +-- Pseudodon_vondembuschianus **

Konopleva et al., 2019 [Publ 3900], fig. 1

iTtTbiriSoTroTroSiD
8 8 0 3 44 3 1 37 
  +----- Indopseudodontina         
  |                                
  |  +-- Bineurus_mouhotii **         
  |  |                             
--+  +-- Pseudodon_cambodjensis **    
  |  |                             
  |  +-- Pseudodon_mekongi **         
  |  |                             
  |  +-- Pilsbryoconcha_exilis **     
  +--+                             
     +-- Pilsbryoconcha_carinifera ** 
     |                             
     +-- Bineurus_loeiensis **        
     |                             
     +-- Bineurus_exilis **           
     |                             
     +-- Pseudodon_vondembuschianus **

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
8 8 6 5 38 2 1 34 
  +----------- Indopseudodontina
  |                             
--+        +-- Pilsbryoconcha **   
  |  +-----+                    
  |  |     +-- Pseudodon **        
  +--+                          
     |  +----- Sundadontina **     
     +--+                       
        |  +-- Thaiconcha **       
        +--+                    
           +-- Bineurus **

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
7 7 5 5 25 2 1 33 
  +-------------- Indopseudodontina
--+                                
  |  +----------- Pilsbryoconcha **   
  +--+                             
     |  +-------- Pseudodon **        
     +--+                          
        |  +----- Sundadontina **     
        +--+                       
           |  +-- Thaiconcha **       
           +--+                    
              +-- Bineurus **

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
6 6 4 4 31 2 1 34 
  +----------- Indopseudodontina
--+                             
  |  +-------- Pilsbryoconcha **   
  +--+                          
     |  +----- Pseudodon **        
     +--+                       
        |  +-- Thaiconcha **       
        +--+                    
           +-- Bineurus **

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
6 6 4 4 31 2 1 34 
  +----------- Indopseudodontina
--+                             
  |  +-------- Pilsbryoconcha **   
  +--+                          
     |  +----- Pseudodon **        
     +--+                       
        |  +-- Thaiconcha **       
        +--+                    
           +-- Bineurus **

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
6 6 4 4 31 2 1 34 
  +----------- Indopseudodontina
--+                             
  |  +-------- Pilsbryoconcha **   
  +--+                          
     |  +----- Pseudodon **        
     +--+                       
        |  +-- Thaiconcha **       
        +--+                    
           +-- Bineurus **

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
6 6 4 5 25 0 0 36 
  +-------- Sundadontina **   
  |                        
--+     +-- Nyeinchanconcha **
  |  +--+                  
  |  |  +-- Bineurus **       
  +--+                     
     |  +-- Pilsbryoconcha ** 
     +--+                  
        +-- Pseudodon **

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
6 6 3 4 31 2 1 37 
  +----------- Indopseudodontina   
  |                                
--+  +-------- Pilsbryoconcha **      
  |  |                             
  +--+  +----- Bineurus_anodontinum **
     |  |                          
     +--+----- Bineurus_loeiensis **  
        |                          
        |  +-- Thaiconcha **          
        +--+                       
           +-- Pseudodon **

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
6 6 4 4 31 2 1 36 
  +----------- Indopseudodontina
--+                             
  |  +-------- Pilsbryoconcha **   
  +--+                          
     |  +----- Bineurus **         
     +--+                       
        |  +-- Thaiconcha **       
        +--+                    
           +-- Pseudodon **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
6 6 3 5 116 2 1 36 
  +----------- Indopseudodontina
  |                             
--+  +-------- Pilsbryoconcha **   
  |  |                          
  +--+  +----- Pseudodon **        
     |  |                       
     +--+  +-- Nyeinchanconcha **  
        |  |                    
        +--+-- Bineurus **         
           |                    
           +-- Sundadontina **

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
6 6 4 4 32 2 1 34 
  +----------- Indopseudodontina
--+                             
  |  +-------- Pilsbryoconcha **   
  +--+                          
     |  +----- Pseudodon **        
     +--+                       
        |  +-- Thaiconcha **       
        +--+                    
           +-- Bineurus **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
6 6 4 5 34 0 0 34 
  +----------- Pilsbryoconcha ** 
--+                           
  |  +-------- Pseudodon **      
  +--+                        
     |  +----- Sundadontina **   
     +--+                     
        |  +-- Nyeinchanconcha **
        +--+                  
           +-- Bineurus **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
5 5 3 4 75 0 0 35 
  +-------- Pilsbryoconcha ** 
--+                        
  |  +----- Pseudodon **      
  +--+                     
     |  +-- Bineurus **       
     +--+                  
        +-- Nyeinchanconcha **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
4 4 2 4 53 1 1 38 
  +----------- Indopseudodontina
--+                             
  |  +-------- Pilsbryoconcha **   
  +--+                          
     |  +----- Bineurus **         
     +--+                       
        |  +-- Pseudodon **        
        +--+                    
           +-- Sundadontina **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
4 4 2 4 53 1 1 38 
  +----------- Indopseudodontina
--+                             
  |  +-------- Bineurus **         
  +--+                          
     |  +----- Pilsbryoconcha **   
     +--+                       
        |  +-- Pseudodon **        
        +--+                    
           +-- Sundadontina **

Bolotov et al., 2022 [Publ 4239], fig. S2

iTtTbiriSoTroTroSiD
3 4 0 3 55 1 1 37 
  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina
  |                    
  +-- Sundadontina **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
3 3 1 2 88 3 1 37 
  +-------- Indopseudodontina        
--+                                  
  |  +----- Pilsbryoconcha_exilis **    
  +--+                               
     |  +-- Pseudodon **                
     +--+                            
        +-- Pilsbryoconcha_carinifera **

Bolotov et al., 2022 [Publ 4252], fig. S1

iTtTbiriSoTroTroSiD
3 4 0 3 58 1 1 37 
  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina
  |                    
  +-- Sundadontina **

Bolotov et al., 2022 [Publ 4239], fig. S1

iTtTbiriSoTroTroSiD
3 4 0 3 55 1 1 37 
  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina
  |                    
  +-- Sundadontina **

Wu et al., 2021 [Publ 4215], fig. 3

iTtTbiriSoTroTroSiD
3 19 17 3 21 2 1 51 
                 +----- Chamberlainia    
  +--------------+                       
  |              |  +-- Sundadontina **     
  |              +--+                    
  |                 +-- Sinohyriopsis    
  |                                      
--+              +----- Indopseudodontina
  |  +-----------+                       
  |  |           |  +-- Pseudodon **        
  |  |           +--+                    
  |  |              +-- Pilsbryoconcha **   
  |  |                                   
  +--+        +-------- Sinosolenaia     
     |  +-----+                          
     |  |     |  +----- Lamprotula       
     |  |     +--+                       
     |  |        |  +-- Potomida         
     |  |        +--+                    
     +--+           +-- Pronodularia     
        |                                
        |           +-- Ptychorhynchus   
        |  +--------+                    
        |  |        +-- Parvasolenaia    
        +--+                             
           |  +-------- Inversidens      
           +--+                          
              |  +----- Gonidea          
              +--+                       
                 |  +-- Microcondylaea   
                 +--+                    
                    +-- Leguminaia

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
3 3 1 2 88 3 1 37 
  +-------- Indopseudodontina        
--+                                  
  |  +----- Pilsbryoconcha_exilis **    
  +--+                               
     |  +-- Pseudodon **                
     +--+                            
        +-- Pilsbryoconcha_carinifera **

Bolotov et al., 2022 [Publ 4252], fig. 2

iTtTbiriSoTroTroSiD
3 4 0 3 58 1 1 37 
  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina
  |                    
  +-- Sundadontina **

Bolotov et al., 2022 [Publ 4239], fig. 2

iTtTbiriSoTroTroSiD
3 4 0 3 55 1 1 37 
  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina
  |                    
  +-- Sundadontina **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 51 0 0 -- 
  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 86 0 0 -- 
  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 44 0 0 -- 
  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Zhang et al., 2024 [Publ 4397], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 26 0 0 -- 
  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 52 0 0 -- 
  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
2 2 0 2 94 0 0 -- 
  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 84 0 0 -- 
  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Matos et al., 2024 [Publ 4384], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 18 0 0 -- 
  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Graf, 2002 [Publ 2651], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 16 0 0 -- 
  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
2 2 0 2 94 0 0 -- 
  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 42 0 0 -- 
  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 32 0 0 -- 
  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 2 52 0 0 -- 
  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 35 0 0 -- 
  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
2 2 0 2 72 0 0 -- 
  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Froufe et al., 2019 [Publ 3936], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 17 0 0 -- 
  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 72 0 0 -- 
  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Matos et al., 2024 [Publ 4383], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 18 0 0 -- 
  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 44 0 0 -- 
  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
2 2 0 2 72 0 0 -- 
  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 51 0 0 -- 
  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 45 0 0 -- 
  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 2 42 0 0 -- 
  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Kongim et al., 2023 [Publ 4290], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 13 1 1 -- 
  +----- Indopseudodontina
--+                       
  |  +-- Namkongnaia **      
  +--+                    
     +-- Pilsbryoconcha **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
2 2 0 2 90 0 0 -- 
  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
2 2 0 2 46 0 0 -- 
  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
2 2 0 2 90 0 0 -- 
  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 29 0 0 -- 
  +-- Pseudodon **   
--+               
  +-- Sundadontina **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 45 0 0 -- 
  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Bolotov et al., 2022 [Publ 4250], fig. 8

iTtTbiriSoTroTroSiD
2 2 0 2 47 4 1 -- 
  +----- Indopseudodontina
--+                       
  |  +-- Pseudodon **        
  +--+                    
     +-- Sundadontina **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
2 2 0 2 37 0 0 -- 
  +-- Pseudodon **     
--+                 
  +-- Pilsbryoconcha **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 67 0 0 -- 
  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Froufe et al., 2019 [Publ 3936], fig. 3-M

iTtTbiriSoTroTroSiD
2 2 0 2 17 0 0 -- 
  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
2 2 0 2 21 0 0 -- 
  +-- Pilsbryoconcha **
--+                 
  +-- Pseudodon **

Walker et al., 2006 [Publ 2666], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Pseudodon_vondembuschianus **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Pseudodon_vondembuschianus **

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Pilsbryoconcha_exilis **

Walker et al., 2006 [Publ 2666], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Pseudodon_vondembuschianus **

Walker et al., 2006 [Publ 2666], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Pseudodon_vondembuschianus **

Zieritz et al., 2020 [Publ 4102], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 30 1 1 -- 
  +-- Pilsbryoconcha **   
--+                    
  +-- Indopseudodontina

Walker et al., 2006 [Publ 2666], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Sundadontina_tumida **

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 8 0 0 -- 
-- Pilsbryoconcha_exilis **

Walker et al., 2006 [Publ 2666], fig. 6-M

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Sundadontina_tumida **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Pseudodon_vondembuschianus **
 
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