Publication Graf & Cummings (2021): species taxonomic opinions

Graf, D.L. & K.S. Cummings. 2021. A ‘big data’ approach to global freshwater mussel diversity (Bivalvia: Unionoida), with an updated checklist of genera and species. Journal of Molluscan Studies 87(1). eyaa034 (36 pp.). https://doi.org/10.1093/mollus/eyaa034.

Acostaea rivoli (Deshayes, 1827) = Mulleria rivoli

+ Acostaea guaduasana d’Orbigny, 1851

Actinonaias computata (Crosse & Fischer, 1893) = Actinonaias sapotalensis

Actinonaias coyensis (Pilsbry, 1910)

Actinonaias medellina (Lea, 1838)

Actinonaias moctezumensis (Pilsbry, 1910) = Disconaias fimbriata

Actinonaias sapotalensis (Lea, 1841)

Actinonaias signata (Pilsbry, 1910)

Actinonaias undivaga (Pilsbry, 1910) = Actinonaias signata

Aculamprotula coreana (Martens, 1886) = Scriptolamprotula coreana

Aculamprotula fibrosa (Heude, 1877) = Scriptolamprotula fibrosa

Aculamprotula grandidens (Lea, 1862) = Scriptolamprotula grandidens

Aculamprotula nodulosa (Wood, 1815) = Scriptolamprotula nodulosa

Aculamprotula scripta (Heude, 1875) = Scriptolamprotula scripta

Aculamprotula tientsinensis (Crosse & Debeaux, 1863) = Scriptolamprotula tientsinensis

Aculamprotula tortuosa (Lea, 1865) = Scriptolamprotula tortuosa

Aculamprotula zonata (Heude, 1883) = Scriptolamprotula zonata

Acuticosta chinensis (Lea, 1868)

Acuticosta retiaria (Heude, 1883) = Pseudoacuticosta retiaria + Acuticosta ovata

+ Quadrula (Lamprotula) ovata Simpson, 1900

Acuticosta sichuanica Zeng & Liu, 1989 = Inversidens sichuanica

Acuticosta trisulcata (Heude, 1883) = Inversidens trisulcata

Alasmidonta arcula (Lea, 1838)

Alasmidonta atropurpurea (Rafinesque, 1831)

Alasmidonta heterodon (Lea, 1829) = Prolasmidonta heterodon

Alasmidonta marginata Say, 1818

Alasmidonta mccordi Athearn, 1964

Alasmidonta raveneliana (Lea, 1834)

Alasmidonta robusta Clarke, 1981

Alasmidonta triangulata (Lea, 1858)

Alasmidonta undulata (Say, 1817)

Alasmidonta varicosa (Lamarck, 1819)

Alasmidonta viridis (Rafinesque, 1820) = Sulcularia viridis

Alasmidonta wrightiana (Walker, 1901)

Alathyria condola Iredale, 1943

Alathyria jacksoni Iredale, 1934

Alathyria pertexta Iredale, 1934

Alathyria profuga (Gould, 1850)

Amblema elliottii (Lea, 1857)

Amblema neislerii (Lea, 1858)

Amblema plicata (Say, 1817)

+ Amblema costata Rafinesque, 1820

Amuranodonta kijaensis Moskvicheva, 1973

Anemina arcaeformis (Heude, 1877) = Anemina harlandi

Anemina euscaphys (Heude, 1879) = Anemina harlandi

Anemina fluminea (Heude, 1877) = Anemina harlandi

Anemina globosula (Heude, 1878) = Anemina harlandi

Anodonta anatina (Linnaeus, 1758)

Anodonta californiensis Lea, 1852 = Brachyanodon californiensis

+ Anodonta dejecta Lewis, 1875

Anodonta cygnea (Linnaeus, 1758)

Anodonta cyrea Drouët, 1881 = Anodonta anatina

Anodonta exulcerata Porro, 1838

Anodonta impura Say, 1829 = Brachyanodon impura

Anodonta kennerlyi Lea, 1860 = Brachyanodon kennerlyi

Anodonta lurulenta (Morelet, 1849) = Anodontites lurulenta

Anodonta nuttalliana Lea, 1838 = Brachyanodon nuttalliana

Anodonta oregonensis Lea, 1838 = Brachyanodon oregonensis

Anodonta pseudodopsis Locard, 1883 = Anodonta anatina

Anodonta vescoiana Bourguignat, 1856

Anodontites aroana H.B. Baker, 1930

Anodontites carinata (Dunker, 1858) = Anodontites tortilis

Anodontites colombiensis Marshall, 1922

Anodontites crispata Bruguière, 1792

Anodontites cylindracea (Lea, 1838)

Anodontites depexa (Martens, 1900) = Anodontites tehuantepecensis

Anodontites elongata (Swainson, 1823)

Anodontites ferrarisii (d’Orbigny, 1835)

Anodontites guanarensis Marshall, 1927

Anodontites iheringi (Clessin, 1882)

Anodontites infossa H.B. Baker, 1930

Anodontites leotaudi (Guppy, 1866)

Anodontites lucida (d’Orbigny, 1835)

Anodontites montezuma (Lea, 1841)

Anodontites moricandii (Lea, 1860)

Anodontites obtusa (Spix in Wagner, 1827)

Anodontites patagonica (Lamarck, 1819)

Anodontites pittieri Marshall, 1922 = Anodontites tehuantepecensis

Anodontites schomburgiana (Sowerby, 1870)

Anodontites solenidea (Sowerby, 1867)

Anodontites tehuantepecensis (Crosse & Fischer, 1893)

Anodontites tenebricosa (Lea, 1834)

Anodontites tortilis (Lea, 1852)

Anodontites trapesialis (Lamarck, 1819)

Anodontites trapezea (Spix in Wagner, 1827)

Anodontites trigona (Spix in Wagner, 1827)

Anodontoides denigratus (Lea, 1852)

Anodontoides ferussacianus (Lea, 1834)

Arcidens confragosus (Say, 1829)

Arcidens wheeleri (Ortmann & Walker, 1912)

Arcidopsis footei (Theobald, 1876)

+ Trapezoideus prashadi Haas, 1922

Arotonaias cyrenoides (Philippi, 1847)

Arotonaias gabbianus (Lea, 1868) = Arotonaias gabbiana

+ Unio nicaraguensis Lea, 1868

Arotonaias imbricata (Mörch, 1861) = Arotonaias encarpa

Aspatharia chaiziana (Rang, 1835)

+ Anodonta tawaii Rang, 1835

Aspatharia dahomeyensis (Lea, 1859)

Aspatharia divaricata (Martens, 1897)

Aspatharia droueti (Chaper, 1885)

Aspatharia ganciniensis (Rochebrune, 1886)

+ Spatha semicorrugata Preston, 1909

Aspatharia marnoi (Jickeli, 1874)

Aspatharia pangallensis (Rochebrune, 1882)

+ Spatha mabillei Jousseaume, 1886

Aspatharia pfeifferiana (Bernardi, 1860)

Aspatharia rochebrunei (Jousseaume, 1886)

+ Spatha tristis Jousseaume, 1886

Aspatharia rugifera (Dunker, 1858)

Aspatharia subreniformis (Sowerby, 1867)

Balwantia baniensis Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan & Kondakov, 2020 = Yaukthwa baniensis

Balwantia elongatula (Bolotov, Konopleva, Vikhrev, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Gofarov, Tomilova & Kondakov, 2019) = Yaukthwa elongatula

Balwantia soleniformis (Benson, 1836)

Bartlettia stefanensis (Moricand, 1856)

Barynaias caldwellii (Lea, 1858) = Nephronaias caldwellii

Barynaias opacata (Crosse & Fischer, 1893) = Sphenonaias opacata

Barynaias pigerrima (Crosse & Fischer, 1893) = Psoronaias crocodilorum

Barynaias plexus (Conrad, 1838) = Sphenonaias plexus

Barynaias sallei (Crosse & Fischer, 1893) = Nephronaias sallei

Beringiana beringiana (Middendorff, 1851)

Beringiana fukuharai San, Hattori & Kondo in Lopes-Lima et al., 2020

Beringiana gosannensis San, Hattori & Kondo in Lopes-Lima et al., 2020

Beringiana japonica (Clessin, 1874)

Bineurus exilis (Morelet, 1866)

Bineurus mouhotii (Lea, 1863)

Brazzaea anceyi Bourguignat, 1886

Buldowskia flavotincta (Martens, 1905)

Buldowskia iwakawai (Suzuki, 1939)

Buldowskia kamiyai San, Hattori & Kondo in Lopes-Lima et al., 2020

Buldowskia shadini (Moskvicheva, 1973)

Buldowskia suifunica (Lindholm, 1925)

Callonaia duprei (Récluz, 1842)

Cambarunio dactylus (Lea, 1840)

Cambarunio hesperus Watters, 2018

Cambarunio iris (Lea, 1829)

Cambarunio jonesii (Lea, 1859)

Cambarunio nebulosus (Conrad, 1834)

Cambarunio simus (Lea, 1838)

Cambarunio taeniatus (Conrad, 1834)

Castalia ambigua Lamarck, 1819

Castalia cordata Swainson, 1840

Castalia crosseana Hidalgo, 1865

Castalia ecarinata Mousson, 1869

Castalia inflata d’Orbigny, 1835

Castalia martensi (Ihering, 1891)

Castalia multisulcata Hupé, 1857

Castalia nehringi (Ihering, 1891)

Castalia orbignyi (Deville & Hupé, 1850)

Castalia orinocensis Morrison, 1943

Castalia psammoica (d’Orbigny, 1835)

Castalia stevensi (H.B. Baker, 1930)

Castalia undosa Martens, 1885

Castaliella quadrata (Sowerby, 1869)

Castaliella schombergiana (Sowerby, 1869)

Castaliella sulcata (Krauss, 1848)

Caudiculatus caudiculatus (Martens, 1867)

Chambardia bourguignati (Bourguignat, 1886)

Chambardia flava (Pilsbry & Bequaert, 1927)

+ Aspatharia (Spathopsis) dautzenbergi Haas, 1936

Chambardia letourneuxi Bourguignat in Servain, 1890

Chambardia moutai (Dartevelle, 1939)

Chambardia nyassaensis (Lea, 1864)

Chambardia petersi (Martens, 1860)

Chambardia rubens (Lamarck, 1819)

Chambardia trapezia (Martens, 1897)

Chambardia wahlbergi (Krauss, 1848)

Chambardia welwitschii (Morelet, 1868)

Chambardia wissmanni (Martens, 1883)

Chamberlainia hainesiana (Lea, 1856)

+ Unio paviei Morlet, 1891

Chelidonopsis hirundo (Martens, 1881)

Coelatura aegyptiaca (Cailliaud, 1827)

+ Nodularia (Cælatura) cariei Germain, 1919

Coelatura alluaudi (Dautzenberg, 1908)

Coelatura bakeri (H. Adams, 1866)

Coelatura briarti (Dautzenberg, 1901)

Coelatura choziensis (Preston, 1910)

Coelatura cridlandi Mandahl-Barth, 1954

Coelatura disciformis (Rochebrune, 1886)

+ Caelatura (Laevirostris?) rotula Pilsbry & Bequaert, 1927

Coelatura essoensis (Chaper, 1885)

Coelatura gabonensis (Küster, 1862)

Coelatura hauttecoeuri (Bourguignat, 1883)

Coelatura horei (E.A. Smith, 1880)

Coelatura hypsiprymna (Martens, 1897)

Coelatura kipopoensis Mandahl-Barth, 1968

Coelatura kunenensis (Mousson, 1887)

Coelatura leopoldvillensis (Putzeys, 1898)

Coelatura lobensis (Frierson, 1913)

Coelatura luapulaensis (Preston, 1913)

Coelatura mesafricana Pilsbry & Bequaert, 1927

Coelatura mossambicensis (Martens, 1860)

Coelatura ratidota (Charmes, 1885)

Coelatura rothschildi (Neuville & Anthony, 1906)

Coelatura stagnorum (Dautzenberg, 1890)

Coelatura stanleyvillensis Pilsbry & Bequaert, 1927

Coelatura stuhlmanni (Martens, 1897)

Contradens comptus (Deshayes in Deshayes & Jullien, 1876) = Lens comptus

+ Harmandia munensis Brandt, 1974

Contradens contradens (Lea, 1838) = Lens contradens

Contradens crossei (Deshayes in Deshayes & Jullien, 1876) = Lens micropterus

Contradens eximius (Lea, 1856) = Lens eximius

Contradens fultoni Haas, 1930 = Lens fultoni

Contradens fulvaster (Drouet & Chaper, 1892) = Lens lugens

Contradens gratiosus (Philippi, 1843) = Indonaia gratiosa

Contradens inaequalis (Rochebrune, 1882) = Lens micropterus

Contradens misellus (Morelet, 1865) = Lens misellus

Contradens novoselovi Konopleva, Bolotov, Spitsyn, Kondakov, Gofarov & Vikhrev, 2019 = Lens novoselovi

Contradens pallegoixi (Sowerby, 1867) = Lens pallegoixi

Contradens peninsularis (Simpson, 1900) = Lens contradens

Contradens rolfbrandti Jeratthitikul & Panha in Jeratthitikul et al., 2019 = Lens rolfbrandti

Contradens semmelinki (Martens, 1891) = Lens semmelinki

Contradens subcircularis (Brandt, 1974) = Leoparreysia subcircularis

Cristaria beirensis Liu & Zhang, 1982 = Cristaria plicata

Cristaria clessini (Kobelt, 1879)

Cristaria plicata (Leach, 1814)

+ Cristaria tuberculata Schumacher, 1817

Cristaria radiata Simpson, 1900

Cristaria truncata Dang, Thai & Pham, 1980

Ctenodesma borneensis (Issel, 1874)

Ctenodesma scheibeneri Haas, 1927

Cucumerunio novaehollandiae (Gray, 1834)

Cumberlandia monodonta (Say, 1829)

Cuneopsis capitatus (Heude, 1874) = Pseudocuneopsis capitata

Cuneopsis celtiformis (Heude, 1874)

Cuneopsis demangei Haas, 1929

Cuneopsis heudei (Heude, 1874)

Cuneopsis kiangsiensis Tchang & Li, 1965

Cuneopsis pisciculus (Heude, 1874) = Tchangsinaia piscicula

Cuneopsis rufescens (Heude, 1874)

Cyclonaias infucata (Conrad, 1834) = Pustulosa infucata

Cyclonaias kieneriana (Lea, 1852) = Pustulosa kieneriana

+ Unio asperatus Lea, 1861

Cyclonaias kleiniana (Lea, 1852) = Pustulosa kleiniana

Cyclonaias necki Burlakova, Karatayev, Lopes-Lima & Bogan in Burlakova et al., 2018 = Pustulosa necki

Cyclonaias nodulata (Rafinesque, 1820) = Pustulosa nodulata

Cyclonaias petrina (Gould, 1855) = Pustulosa petrina

Cyclonaias pustulosa (Lea, 1831) = Pustulosa pustulosa + Pustulosa kieneriana

+ Quadrula archeri Frierson, 1905

+ Unio aureus Lea, 1859

+ Unio houstonensis Lea, 1859

+ Unio refulgens Lea, 1868

Cyclonaias succissa (Lea, 1852) = Pustulosa succissa

Cyclonaias tuberculata (Rafinesque, 1820)

Cyprogenia aberti (Conrad, 1850)

Cyprogenia stegaria (Rafinesque, 1820)

+ Unio irroratus Lea, 1828

Cyrtonaias explicata (Morelet, 1849) = Mesonaias explicata

+ Lampsilis (Proptera) lividus Simpson, 1900

Cyrtonaias sapperi (Ihering, 1901) = Mesonaias explicata

Cyrtonaias tampicoensis (Lea, 1838) = Mesonaias tampicoensis + Cyrtonaias berlandierii

+ Unio saladoensis Lea, 1860

+ Unio berlandierii Lea, 1857

Cyrtonaias umbrosa (Lea, 1856) = Mesonaias umbrosa + Mesonaias explicata

+ Unio (Lampsilis) rovirosai Pilsbry, 1900

Delphinonaias largillierti (Philippi, 1847)

+ Unio delphinulus Morelet, 1849

+ Unio paludosus Morelet, 1849

+ Unio planivalvis Morelet, 1851

Delphinonaias scutulata (Morelet, 1849)

Diaurora aurorea (Heude, 1883)

Diplodon aethiops (Lea, 1860)

Diplodon berthae Ortmann, 1921

Diplodon burroughianus (Lea, 1834)

Diplodon caipira (Ihering, 1893)

Diplodon charruanus (d’Orbigny, 1835)

Diplodon chilensis (Gray, 1828)

+ Unio patagonica d’Orbigny, 1835

Diplodon delodontus (Lamarck, 1819)

Diplodon ellipticus Spix in Wagner, 1827

+ Unio besckeanus Dunker, 1849

Diplodon fontainianus (d’Orbigny, 1835)

Diplodon funebralis (Lea, 1860)

Diplodon garbei Ihering, 1910

Diplodon granosus (Bruguière, 1792)

Diplodon gratus (Lea, 1860)

Diplodon greeffeanus (Ihering, 1893)

Diplodon guaporensis Bonetto & Tassara, 1987

Diplodon guaranianus (d’Orbigny, 1835)

Diplodon hildae Ortmann, 1921

Diplodon hylaeus (d’Orbigny, 1835)

Diplodon iheringi Simpson, 1914

Diplodon imitator Ortmann, 1921

Diplodon koseritzi (Clessin, 1888)

Diplodon losadae Haas, 1966

Diplodon martensi (Ihering, 1893)

Diplodon multistriatus (Lea, 1831)

+ Unio expansus Küster, 1856

Diplodon obsolescens F. Baker, 1913

Diplodon parallelopipedon (Lea, 1834)

Diplodon paranensis (Lea, 1834)

Diplodon parodizi Bonetto, 1962

Diplodon paulista (Ihering, 1893) = Diplodon paulistus

Diplodon peraeformis (Lea, 1860)

Diplodon pfeifferi (Dunker, 1849)

Diplodon piceus (Lea, 1860)

Diplodon rhombeus Spix in Wagner, 1827

Diplodon rhuacoicus (d’Orbigny, 1835)

Diplodon rotundus Spix in Wagner, 1827

Diplodon solidulus (Philippi, 1869)

Diplodon solisianus (d’Orbigny, 1835)

Diplodon suavidicus (Lea, 1856)

+ Diplodon flucki Morrison, 1943

Diplodon uruguayensis (Lea, 1860)

Diplodon variabilis (Maton, 1811)

Diplodon vicarius Ortmann, 1921

Diplodon voltzi Vernhout, 1914

Diplodon wymanii (Lea, 1860)

Diplodontites cookei Marshall, 1922

Diplodontites olssoni Pilsbry, 1933

Diplodontites pilsbryana Olsson & Wurtz, 1951

Discomya radulosa (Drouet & Chaper, 1892)

Disconaias conchos Taylor, 1997

Disconaias disca (Lea, 1838)

+ Unio panacoensis von dem Busch in Philippi, 1843

Disconaias fimbriata (Frierson, 1907)

Disconaias novileonis (Pilsbry, 1910) = Friersonia iridella

Disconaias purpuriata (Say, 1831)

Dromus dromas (Lea, 1834)

Echyridella aucklandica (Gray, 1843)

+ Diplodon websteri Simpson, 1902

Echyridella menziesii (Gray, 1843)

+ Diplodon (Hyridella) menziesi lucasi Suter, 1905

Echyridella onekaka Fenwick & Marshall, 2006

Ellipsaria lineolata (Rafinesque, 1820)

+ Obliquaria (Ellipsaria) ellipsaria Rafinesque, 1820

Elliptio ahenea (Lea, 1843)

+ Unio waltoni B.H. Wright, 1888

Elliptio angustata (Lea, 1831)

Elliptio arca (Conrad, 1834)

Elliptio arctata (Conrad, 1834)

Elliptio chipolaensis (Walker, 1905)

Elliptio cistellaeformis (Lea, 1863)

Elliptio complanata (Lightfoot, 1786)

Elliptio congaraea (Lea, 1831)

+ Unio waccamawensis Lea, 1863

Elliptio crassidens (Lamarck, 1819)

+ Unio (Elliptio) nigra Rafinesque, 1820